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Revision of the European species of the genus Astiphromma Forster, 1869

Authors:
  • Klinik Fallingbostel

Abstract and Figures

The European species of the genus Astiphromma Forster, 1869 are revised. 26 taxa are accepted as valid and keyed, described, and figured. One further European species described by Schwenke, Astiphromma peltolatum, has not been studied, because the type could not be located for the revision. Two species and one subspecies are described as new: Astiphromma flagella for spec. nov., Astiphromma flavoventrale spec. nov., and Astiphromma nigriceps asiaticiis subspec. nov. The male of Astiphromma uliginosum Schwenke, 1999 is described for the first time here. The following new synonyms are established: Astiphromma barbatum Schwenke, 1999 and Astiphromma caecum Schwenke, 1999 are new synonyms of Astiphromma aggressor (Fabricius, 1804); Astiphromma dispersum Schwenke, 1999 and Astiphromma laricis Schwenke, 1999 are new synonyms of Astiphromma alpinum Roman, 1909; Mesochorus (Astiphrommus) hamulus Thomson, 1886 and Astiphromma consertum Schwenke, 1999 are new synonyms of Astiphromma buccatum (Thomson, 1886); Mesothorus (Astiphrommus) graniger Thomson, 1886 is a new synonym of Astiphromma hirsutton (Bridgman, 1883); Astiphromma contum Schwenke, 1999 is a new synonym of Astiphromnia italicum Schwenke, 1999; Astiphromma intermedium Constantineanu Mustata, 1976 is a new synonym of Astiphromma scutellatum (Gravenhorst, 1829); Mesochorus (Astiphrommus) mandibularis Thomson, 1886 is a new synonym of Astiphromma striation (Brischke, 1880); and Astiphromma trimaculosum Schwenke, 2004 is a new synonym of Astiphromma uliginosum Schwenke, 1999.
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SPIXIANA 38 185-132 München, August 2015 ISSN 0341-8391
Revision of the European species of the genus
Astiphromma Förster, 1869
(Hymenoptera, Ichneumonidae, Mesochorinae)
Matthias Riedel
Riedel, M. 2015. Revision of the European species of the genus Astiphromma
Förster, 1869 (Hymenoptera, Ichneumonidae, Mesochorinae). Spixiana 38 (1): 85-
132.
The European species of the genus Astiphromma Förster, 1869 are revised. 26 taxa
are accepted as valid and keyed, described, and figured. One further European
species described by Schwenke, Astiphromma peltolatum, has not been studied, be-
cause the type could not be located for the revision.
Two species and one subspecies are described as new: Astiphromma flagellator
spec. nov., Astiphromma flavoventrale spec. nov., and Astiphromma nigriceps asiaticus
subspec. nov. The male of Astiphromma uliginosum Schwenke, 1999 is described for
the first time here.
The following new synonyms are established: Astiphromma barbatum Schwenke,
1999 and Astiphromma caecum Schwenke, 1999 are new synonyms of Astiphromma
aggressor (Fabricius, 1804); Astiphromma dispersum Schwenke, 1999 and Astiphromma
laricis Schwenke, 1999 are new synonyms of Astiphromma alpinum Roman, 1909;
Mesochorus (Astiphrommus) hamulus Thomson, 1886 and Astiphromma consertum
Schwenke, 1999 are new synonyms of Astiphromma buccatum (Thomson, 1886);
Mesochorus (Astiphrommus) graniger Thomson, 1886 is a new synonym of Astiphr omma
hirsutum (Bridgman, 1883); Astiphromma contum Schwenke, 1999 is a new synonym
of Astiphromma italicum Schwenke, 1999; Astiphromma intermedium Constantineanu
& Mustata, 1976 is a new synonym of Astiphromma scutellatum (Gravenhorst, 1829);
Mesochorus (Astiphrommus) mandibularis Thomson, 1886 is a new synonym of
Astiphromma striatum (Brischke, 1880); and Astiphromma trimaculosum Schwenke,
2004 is a new synonym of Astiphromma uliginosum Schwenke, 1999.
In dieser Arbeit werden die europäischen Arten der Gattung Astiphromma Förs-
ter, 1869 revidiert. 26 Taxa werden als valide betrachtet, beschrieben und abgebil-
det. Außerdem wird ein Bestimmungsschlüssel für diese Arten gegeben. Eine
weitere europäische, von Schwenke beschriebene Art, Astiphromma peltolatum,
konnte nicht untersucht werden, da das Typenmaterial nicht auffindbar war.
Zwei Arten und eine Unterart werden neu beschrieben: Astiphromma flagellator
spec. nov., Astiphromma flavoventrale spec. nov., und Astiphromma nigriceps asiaticus
subspec. nov., außerdem das bisher unbekannte Männchen von Astiphromma uli-
ginosum Schwenke, 1999.
Folgende Synonyme werden festgestellt: Astiphromma barbatum Schwenke, 1999
und Astiphromma caecum Schwenke, 1999 sind neue Synonyme zu Astiphromma
aggressor (Fabricius, 1804); Astiphromma dispersum Schwenke, 1999 und Astiphromma
laricis Schwenke, 1999 sind neue Synonyme zu Astiphromma alpinum Roman, 1909;
Mesochorus (Astiphrommus) hamulus Thomson, 1886 and Astiphromma consertum
Schwenke, 1999 sind neue Synonyme zu Astiphromma buccatum (Thomson, 1886);
Mesochorus (Astiphrommus) graniger Thomson, 1886 ist ein neues Synonym zu Asti-
phromma hirsutum (Bridgman, 1883); Astiphromma contum Schwenke, 1999 ist ein
86
neues Synonym zu Astiphromma italicum Schwenke, 1999; Astiphromma intermedium
Constantineanu & Mustata, 1976 ist ein neues Synonym zu Astiphromma scutellatum
(Gravenhorst, 1829); Mesochorus (Astiphrommus) mandibularis Thomson, 1886 ist ein
neues Synonym zu Astiphromma striatum (Brischke, 1880); und Astiphr omma trima-
culosum Schwenke, 2004 ist ein neues Synonym zu Astiphromma uliginosum Schwen-
ke, 1999.
Matthias Riedel, Amselweg 9A, 29683 Bad Fallingbostel, Germany;
e-mail: mamaflo.riedel@t-online.de
Introduction
The European species of the Mesochorinae genus
Astiphromma Förster, 1869 have recently been revised
by Schwenke (1999) with some additional material
that he published in 2004. Although his work repre-
sents a landmark study for the taxonomy of European
Mesochorinae, the determination keys and descrip-
tions of taxa are very short and not supplemented
by illustrations. Since some taxa were described
from one or very few specimens, the intraspecific
variability of structure and colour patterns could
not been addressed sufficiently. For several species,
only one sex has been described, and the opposite
sex is unknown yet.
Although Horstmann (2006) has re-described
some European taxa and corrected their nomen-
clature, it seemed necessary to study the European
material of Astiphromma again and to present a new
and illustrated key for a reliable identification of the
different species in the Western Palearctic region.
Generic classification of European Mesochorinae
In accordance to the traditional concept of European
entomologists, Schwenke (1999) differentiated seven
genera of European Mesochorinae and described
an additional genus, Planochorus in a subsequent
publication (Schwenke 2004). In a cladistic analysis
of the world genera of Mesochorinae, Wahl (1993)
placed the genus Stictopisthus back as a synonym
of Mesochorus, and synonymized Mesochorella and
Dolichochorus with Astiphromma considering the pres-
ence of the dorsolateral carina as an autapomorphy
of Astiphromma. Although Schwenke (1999, 2000)
stated that the dorsolateral carina of first tergite is
lacking in Mesochorella, this carina is visible at least
behind the spiracle and/or at the apical margin in
my available material (see Fig. 3). It was suggested
by Broad (2011) that his preliminary results of coding
A. longiceps for Wahls characters give some hints for
a polyphyletic Astiphromma if this taxon is included.
Here, I propose three subgenera Astiphromma
s. str., Mesochorella, and Dolichochorus within the
genus Astiphromma sensu lato with the common char-
acter of a partly or completely developed dorsolateral
carina on the postpetiole. Also, the nervellus in the
hind wing is intercepted in all species, though the dis-
coidella is not always visible. However, a closer clad-
istic analysis including larval characters and molecu-
lar data will certainly give more information on the
phylogenetic relationship of the mentioned genera.
I have not seen and studied any material of
the genus Planochorus which was established by
Schwenke (2004). His type material is not deposited
in the ZSM nor in the collection of Zwakhals (as it was
erroneously published in the original description).
According to his description, the characteristics of
this genus are a plane dorsal surface of the meso-
soma, a long coxa III, and whitish orbits of the head.
Additionally, the lower mandibular tooth is larger
than the upper one, and the nervellus of hind wing
is not clearly intercepted (e. g. discoidella not devel-
oped). Whether this species belongs to Astiphromma
in the wider sense or not, cannot be decided yet.
Biology of Astiphromma
All known European species of Astiphromma are
hyperparasitoids of different hosts. The primary
hosts are usually larvae of Lepidoptera and Sym-
phyta (Hymenoptera). However, at least one taxon,
Astiphromma nigriceps (Brischke) has been found
in adult Coleoptera (Chrysomelidae) (Schwenke
2000) but the primary parasitoide is unknown for
this taxon.
The dorsale group of Astiphromma characterized
by the pointed scutellum was found in combination
of Tachinidae (Diptera) as primary parasitoids where
the specialized form of scutellum is used to emerge
from the Tachinid pupa. The other Astiphromma
species from Europe have been reared together with
several Braconidae and Ichneumonidae as primary
parasitoids.
Variability of Astiphromma species
Some species of the genus Astiphromma, such as A. ag-
gressor (Fabricius) and A. splenium (Curtis) show a
large variability in size, colour, and structural details.
Therefore, many morphological features used for
separation of different taxa by former authors (e. g.
Lee 1992, Schwenke 1999, 2004) are not reliable if
87
one studies a larger assemblage of specimens. In my
opinion, the species concepts defined by just a single
holotype were sometimes separating individual
variants (e. g. A. caecum, A. barbatum, A. consertum,
A. laricis, A. trimaculosum, all described by Schwenke)
of common taxa only.
One the other hand, the above mentioned widely
variable species of Astiphromma might represent in
fact species groups which have to be closer defined by
rearing experiments and molecular data in the future.
Material and methods
Measurements and indices
FEI (flagellar-eye index): relation of length of first flagel-
lomere (without anellus) to length of compound eye.
MI (malar index): relation of length of malar space to
the width of mandibular base.
OOD (ocular-ocellar distance): relation of the distance
of the lateral ocellus to the compound eye to its di-
ameter.
RI (radial index): relation of length of anterior margin
of radial cell to length of pterostigma.
TEI (temple-eye index): relation of the width of temple
to the width of compound eye (seen from lateral).
Material
For the distributional records below I used the perso-
nally studied material only, new records are marked by
asterisks. The following collections have been studied:
Coll. Artmann, Arsten/Switzerland (CA), Coll. Giovan-
ni, Verona/Italy (CG), Coll. Jussila, Paattinen/Finland
(CJ), Coll. Riedel, Bad Fallingbostel/Germany (CR),
Coll. Schwarz, Linz/Austria (CS), Coll. Schnee, Mark-
kleeberg/Germany (SCHNEE), Coll. Varkonyi, Kuh-
mo/Finland (CV), Coll. Zwakhals/The Netherlands
(CZ), National Museums of Scotland, Edinburgh/Scot-
land (EDI), University Erzurum/Turkey (ERZ), Musee
de Zoologie Lausanne/Switzerland (LAU), Biozentrum
Linz/Austria (LINZ), Natural History Museum, Lon-
don/England (NHM), Swedish Museum of Natural
History, Stockholm/Sweden (STOCK), Museum of
Zoology, Warszawa/Poland (WARS), Zoologische
Staatssammlung Munich/Germany (ZSM).
Identification key of European
Astiphromma Förster, 1869
Key for WW
1. Scutellum pointed apically (rarely scutellum
slightly pointed or seemingly rounded). Dorso-
lateral carina of postpetiole running ventral to
spiracle (Fig. 1). Occipital carina dipped medi-
ally, occiput here more or less impressed. .........
....................................................... dorsale group 2.
Scutellum rounded apically. Dorsolateral carina
of postpetiole running at the same level or dor-
sal to spiracle (Fig. 2). Occipital carina not dipped
medially, occiput not impressed. .................... 3.
2. Body stouter. Ovipositor sheath 5.0-7.5 × as long
as wide (Fig. 73). Hind leg stout: femur III 3.8-
5.1 × as long as wide, metatarsus III 6-7.5 × as
long as wide and 0.40-0.42 × as long as tibia III.
Lower half of mesopleurum usually with very
scattered to scattered punctation (rarely with
denser punctures) (Fig. 105). Tarsus III often
infuscate. Claw III with strongly curved tip
(Fig. 61). Flagellum often infuscate or brownish-
red. .......................... A. dorsale (Holmgren, 1860)
Body slender. Ovipositor sheath slender, stab-
shaped, 7.5-9.5 × as long as wide (Fig. 74). Hind
leg slender: femur III 5.0-5.6
× as long as wide,
metatarsus III 8-9 × as long as wide and 0.45-0.5 ×
as long as tibia III. Lower half of mesopleurum
usually with scattered to dense punctation
(Fig. 106). Tarsus III usually yellowish-brown.
Claw III with less curved tip (Fig. 58). Flagellum
often completely or basally yellowish. ..............
.............................. A. hirsutum (Bridgman, 1883)
3. Inner spur of tibia III very short, about 0.25-0.3 ×
as long as metatarsus III. Head and mesosoma
with coarse and dense punctures. Lower man-
dibular tooth much larger than the upper one
(Fig. 49). Temple seen from dorsal with parallel
sides behind the eye (Fig. 17). Dorsolateral ca-
rina of postpetiole developed in part only (usu-
ally visible apically) (Fig. 3). ................................
.................................. A. nigriceps (Brischke, 1880)
Inner spur of tibia III at least 0.4 × as long as
metatarsus III. Punctation of head and meso-
soma less coarse and dense. Mandibular teeth
often of equal size. Dorsolateral carina of post-
petiole strong (Fig. 2). ........................................ 4.
4. Head seen from frontal almost triangular
(Fig. 35). Malar space very long, slightly longer
than the width of mandibular base (Fig. 35).
Flagellum long and slender, first flagellomere
about 1.0-1.1 × as long as the eye. Face cream-
yellow, with a median w-sized blackish spot. ..
...................................... A. longiceps (Strobl, 1904)
Head seen from frontal more or less quadran-
gular. Malar space shorter than the width of
mandibular base. Flagellum usually less slender,
first flagellomere at most 0.8 × as long as eye.
Face with different coloration. ......................... 5.
5. Postpetiole completely with longitudinal stria-
tion or rugae (rarely the striation is indistinct)
(Figs 122, 123). Second tergite without distinct
88
thyridium or small thyridium at basal margin
only. ..................................................................... 6.
Postpetiole at most with some striae at its apical
margin. If the striation is more pronounced (in
A. leucogrammum and A. tenuicorne, Figs 124,
125), then face completely yellow and mesopleu-
rum two-coloured: reddish-yellow below and
black above. Thyridium of second tergite usu-
ally distinct, at some distance from the basal
margin. ................................................................. 9.
6. Mesopleurum mainly reddish, often with yellow
stripes or spots. Lower mandibular tooth longer
than the upper one (as Fig. 66). Hypostomal
carina elevated. Nervulus more or less interstitial.
Head and mesopleurum densely and coarsely
punctate (Fig. 39). Temples parallel behind the
eyes. ............................ A. pictum (Brischke, 1880)
Mesopleurum mainly blackish, often with a small
yellowish spot below. Mandibular teeth of the
equal size. Hypostomal carina not elevated. . 7.
7. Tergites 2 and 3 more or less reddish. Face yel-
lowish except a central blackish spot (Fig. 40).
Head and mesopleurum with coarse and dense
punctation, temple with distinct punctures.
Postpetiole strongly widened (Fig. 124). Temple
broad behind eye, TEI 1.0-1.1. .............................
.................................. A. rimosum Schwenke, 1999
Metasoma blackish, with transverse yellowish
stripes on hind margins of tergites 2-7. Head
with superficial puncture, temple with superfi-
cial punctures, granulate. ................................. 8.
8. Face mainly black, only clypeus, lower facial
orbits and lower gena yellowish (Fig. 36). Meso-
pleurum ventrally with dense but fine puncta-
tion. Mesosternum black, at most with a yellow-
ish spot. Temple narrower, TEI 0.8-0.9. .............
................................... A. luridum Schwenke, 1999
Face yellow, with a large rectangular median
black spot below antenna (Fig. 31). Mesopleurum
rugulose-punctate ventrally, matt. Mesosternum
yellow (Fig. 107). Temple wider, TEI 1.1. ..........
..................................... A. flavoventrale spec. nov.
9. Face completely yellowish (Figs 26, 27, 34, 44).
............................................................................. 10.
Face black, sometimes with yellow colour pattern
in the lower half. .............................................. 14.
10. Lower mandibular tooth much larger than the
upper one. Metatarsus III with apical rings of
stout bristles (Fig. 50). Trochantellus III ven-
trally flattened and with edged sides (Fig. 51). .
.................................. A. striatum (Brischke, 1880)
Mandibular teeth of equal size. If lower tooth
slightly larger (in A. tenuicorne), then metatarsus
III without apical ring of bristles. Trochantellus
III not flattened ventrally, normally rounded. .
............................................................................. 11.
11. Mesopleurum black. ..............................................
..................................... A. anale (Holmgren, 1860)
Mesopleurum with red and/or yellow colour
pattern. ............................................................... 12.
12. Mesopleurum with 3 or 4 different colours:
ivory, yellow, reddish and more or less black in
spot-like distribution, the black colour covers
the dorsal 0.2 only. Femur III stouter, less than
5 × as long as wide. Ovipositor sheath stout,
about 6-7 × as long as wide, reddish (Fig. 87).
Propodeum with faint costula. ............................
.................................. A. simplex (Thomson, 1886)
Mesopleurum two-coloured: black above and
reddish below, with sharp border between both
colours. Ovipositor sheath slender, more than
8 × as long as wide (Figs 78, 89). Postpetiole with
some longitudinal striae apically (Figs 124, 125).
Femur III slender, more than 5 × as long as wide.
Nervulus interstitial. ....................................... 13.
13. Striation of postpetiole more extensive, usually
in more than the apical half (Fig. 125). Coxa III
often with a brownish dorsal spot at apex. Ptero-
stigma brown. Radial cell shorter, the anterior
margin at most 1.1 × as long as the pterostigma.
Costula usually faint. ............................................
.................... A. leucogrammum (Holmgren, 1860)
Striation of postpetiole less extended, distinct in
the apical third only (Fig. 124). Coxa III reddish-
yellow, without a brownish spot dorsally. Ptero-
stigma hyaline-brownish. Radial cell longer, its
anterior margin distinctly longer than the ptero-
stigma (1.2-1.4 ×). Costula usually distinct. ......
............................. A. tenuicorne (Thomson, 1886)
14. Clypeus completely and at least the lower third
of face more or less yellow-red or red (Figs 24,
46, 47). Preapical flagellomeres short (about 1.5 ×
as long as wide), the apical one usually blunt. .
............................................................................. 15.
Clypeus partly, and face mainly black, at most
around mouth narrowly yellowish or reddish.
Preapical flagellomeres often slender (about 2 ×
as long as wide). ............................................... 16.
15. Lateral cream-yellow spots on face usually long,
about 2
/3 as long as the face (Fig. 46), but some-
times less developed (Fig. 47). Tergite 3 with
yellow median stripe, following tergites dark-
89
ened basally and light apically (vespoid colour
pattern). ............. A. tridentatum Schwenke, 1999
Lateral cream-yellow spots shorter, usually about
1
/3 as long as the face (Fig. 24). Tergites mainly
black, tergite 3 without median yellowish stripe,
apical tergites black. .............................................
........................................ A. alpinum Roman, 1909
(here probably runs A. peltolatum Schwenke,
2004: close to dispersum, but larger (7 mm), Lat-
eral margins of face divergent ventrally, clypeus
reddish, face completely black. Mesosoma black,
tergites black with narrow yellowish apical
margins. Wing venes blackish. OOD 2.0. Ovi-
positor sheath shorter than metatarsus III.)
16. Tergites 4-7 often completely red. If apical ter-
gites more or less darkened (var. sericans), then
femur III diffusely darkened in the apical half
(Fig. 21). Claw III with strong teeth, usually the
basal teeth longer than the apical ones. .............
...................................... A. splenium (Curtis, 1833)
Tergites 4-7 black or more or less reddish. ......
............................................................................. 17.
17. Coxa III more or less blackened, femur III brown
or blackish. ........................................................ 18.
Coxa III red, at most with a brownish dorsal
tinge. If coxa III almost completey brownish (in
A. flagellator spec. nov.), then femur III com-
pletely reddish-yellow. ................................... 19.
18. Smaller, body length 5-6 mm. Ovipositor sheath
stab-like, slightly bent dorsally at apex (Fig. 83).
Scutellum not carinate. Nervulus interstitial. ...
............................... A. nigrocoxatum (Strobl, 1904)
Larger, body length 8-11 mm. Ovipositor sheath
wider (Fig. 68). Scutellum with strong lateral
carina, at least in the basal half (Fig. 63). Nervu-
lus postfurcal. ........ A. albitarse (Brischke, 1880)
19. Genal carina joining the hypostomal one at
mandibular base or very close to it (at most 1
/3
of malar length) (Fig. 5). Lower mandibular tooth
larger than the upper one (Fig. 66). .............. 20.
Genal carina joining the hypostomal one more
or less distant from mandibular base (at least
half of malar length) (Fig. 4). Lower mandibular
tooth at most slightly larger than the upper one
or both teeth of equal size. ............................. 21.
20. Temple strongly narrowed behind eye, TEI 0.6
(Fig. 7). Lower half of mesopleurum with scat-
tered fine puncture. Postpetiole not punctate,
smooth. Femur III stout, 3.3-4.0 × as long as wide.
Scutellum often with lateral carina in the basal
0.3. ............................ A. italicum Schwenke, 1999
Temple wider, parallel or widened behind eyes,
TEI 0.9-1.1 (Fig. 10). Lower half of mesopleurum
very densely punctate. Postpetiole with fine
punctures. Femur III slender, 4.1-4.7 × as long
as wide. Scutellum not carinate. .........................
............................... A. buccatum (Thomson, 1886)
21. Femur III dark brown to black. Ocelli large, OOD
0.8-1.0 (Fig. 20). ..... A. varipes (Holmgren, 1860)
Femur III reddish. Ocelli smaller, OOD usually
> 1.0. ................................................................... 22.
22. Coxae II and III more or less brownish. Flagellum
very short, preapical flagellomeres at most
slightly longer than wide (Fig. 65). ......................
......................................... A. flagellator spec. nov.
Coxae II and III reddish or yellowish-red, at most
coxa III with dorsal brownish spot. Flagellum
often slender, preapical flagellomeres distinctly
(usually 1.5-2 ×) longer than wide. ............... 23.
23. Mesosternum, mesopleurum and scutellum more
or less reddish. Tarsus III, ovipositor sheath and
basal flagellomeres usually blackish. Nervulus
of front wing usually interstitial or slightly
postfurcal. ...... A. scutellatum Gravenhorst, 1829
Mesosoma mainly black, at most lower meso-
pleurum and scutellum with diffuse dark reddish
suffusion (Fig. 104). Tarsus III often reddish,
nervulus more or less postfurcal. .................. 24.
24. Apical tergites with vespoid (dark/pale) colour
pattern (Fig. 118). Smaller, body length 4-6 mm.
Mesopleurum smooth. Frons, vertex and temple
with scattered and superficial puncture. Radial
cell longer, about 1.2 × as long as pterostigma at
anterior margin. Ovipositor sheath long and
slender, 7.7-8.4 × as long as wide (Fig. 91). area
superomedia behind costula with narrowed sides
apically. .............. A. uliginosum Schwenke, 1999
Apical tergites more or less blackish. Ovipositor
sheath usually stouter (not more than 7 × as long
as wide) (Figs 67, 72). ...................................... 25.
25. Temple slightly narrowed behind eye, TEI 0.8-
0.9. Tarsus III usually infuscate. ..... A. diversum
Schwenke, 1999
Temple strongly narrowed behind eye, TEI 0.7-
0.8. Tarsus III usually pale brownish or reddish,
Ovipositor sheath blackish. .................................
................................ A. aggressor (Fabricius, 1804)
90
Key for MM:
(MM of A. peltolatum, A. flavoventrale, A. flagellator and
A. longiceps are unknown.)
1. Scutellum pointed apically (rarely scutellum
seemingly rounded). Dorsolateral carina of
postpetiole situated ventrally of the spiracle
(Fig. 1) Occipital carina medially dipped, occiput
here more or less impressed. ...............................
....................................................... dorsale group 2.
Scutellum rounded apically. Dorsolateral carina
of postpetiole situated at the same level or dor-
sally to the spiracle (Fig. 2). Occipital carina
medially not dipped, occiput not impressed. ...
.............................................................................. 3.
2. Mesopleurum with very scattered weak punc-
ture. Face wider, its minimal width about 0.85-
0.95 × eye length. Flagellum often infuscate. .....
.................................. A. dorsale (Holmgren, 1860)
Mesopleurum with scattered to dense puncta-
tion. Face narrower, its minimal width 0.8-0.85 ×
eye length. Flagellum usually yellowish or
reddish-brown. ......................................................
.............................. A. hirsutum (Bridgman, 1883)
3. Clypeus and face black. Head and mesosoma
with coarse and dense punctation. Lower man-
dibular tooth much larger than the upper one.
Temple seen from dorsal with parallel sides
behind the eye. Inner spur of tibia III very short,
about 0.25-0.3 × as long as metatarsus III. .........
.................................. A. nigriceps (Brischke, 1880)
Clypeus and face completely or mainly yellow.
Lower mandibular tooth sometimes larger than
the upper one or equally sized. Inner spur of
tibia III at least 0.4 × as long as metatarsus III.
............................................................................... 4.
4. Postpetiole with more or less strong longitudinal
striation (if in A. luridum sometimes obsolete,
then pronotum completely yellowish) (Figs 122,
123). ...................................................................... 5.
Postpetiole usually smooth, at most with some
fine striae at its apical half (if in A. leucogrammum
and A. tenuicorne with some striation, then pro-
notum partly blackish) (Figs 124, 125). ........... 7.
5. Tergites 3 and 4 mainly reddish. .........................
.................................. A. rimosum Schwenke, 1999
Tergites 3 and 4 black, with narrow ivory apical
margins. ............................................................... 6.
6. Mesopleurum mainly reddish. ............................
..................................... A. pictum (Brischke, 1880)
Mesopleurum yellowish ventrally and black
dorsally (as Fig. 108). ............................................
................................... A. luridum Schwenke, 1999
7. Lower mandibular tooth larger than the upper
one. Genal carina joining the hypostomal one
distant from mandibular base (Fig. 4). Metatarsus
III with apical ring of stout bristles (Fig. 50).
Trochantellus III ventrally flattened, with edged
sides (Fig. 51). ........ A. striatum (Brischke, 1880)
Mandibular teeth of equal size. If lower tooth
slightly larger than the upper one (A. italicum
and A. buccatum), then genal carina joining the
hypostomal one at or close to mandibular base
(Fig. 5). Metatarsus III usually without apical
ring of stout bristles. Trochantellus III rounded
ventrally. ............................................................. 8.
8. Genal carina joining the hypostomal one at or
close to mandibular base (Fig. 5). Lower man-
dibular tooth slightly larger than the upper
one. ....................................................................... 9.
Genal carina joining the hypostomal one far from
the mandibular base (Fig. 6). Mandibular teeth
usually of equal size. ....................................... 10.
9. Mesosoma mainly black, only pronotum reddish
or reddish-brown. Femur III stout, 3.3-4.0 × as
long as wide. ........... A. italicum Schwenke, 1999
Mesoscutum anterolateral with large yellowish
hook-like spots. Femur III slenderer, 4.1-4.7 × as
long as wide. ....... A. buccatum (Thomson, 1886)
10. Coxa III mainly blackish or dark brown. ..... 11.
Coxa III yellowish-red. .................................... 12.
11. Larger species, body length 8-11 mm. Clasper
long and stab-like, about as long as the second
and half of the third tarsomere III together (as
in Fig. 101). Scutellum often with lateral carina
in the basal half (Fig. 63). Tarsus III cream-yel-
low. ........................... A. albitarse (Brischke, 1880)
Smaller species, body length 5-6.5 mm. Clasper
shorter, about as long as the second tarsomere
III, apically slightly thickened (Fig. 99). Scutellum
not carinate. Tarsus III reddish or infuscate. ....
.............................. A. nigrocoxatum (Strobl, 1904)
(here runs probably also the unknown M of
A. flagellator spec. nov.)
12. Femur III mainly brownish or black. Ocelli large,
OOD 0.8-1.0. .......... A. varipes (Holmgren, 1860)
Femur III reddish or yellowish-red, at most with
brownish suffusion dorsally or apically. Ocelli
smaller, OOD > 1.0. .......................................... 13.
91
13. Clasper long, distinctly longer than second tar-
somere III (Fig. 101). Often larger species, up to
10 mm. ............................................................... 14.
Clasper shorter, at most as long as the second
tarsomere III. Usually smaller species, body
length 5-6 mm. ................................................. 15.
14. Tarsus III reddish-yellow or brownish. Mesos-
ternum and sometimes scutellum black, at most
with some diffuse dark-reddish suffusion. ........
................................ A. aggressor (Fabricius, 1804)
Tarsus III usually blackish. Mesosternum, and
mesopleurum mainly red or brown. Scutellum
red, at least in its basal half. ................................
...................... A. scutellatum (Gravenhorst, 1829)
15. Mesopleurum partly or completely reddish or
yellowish. .......................................................... 16.
Mesopleurum mainly or completely black. . 20.
16. Apex of clasper widened, club-shaped (Fig. 97).
Pronotum and mesopleurum two-coloured:
blackish dorsally and reddish or yellowish ven-
trally, usually with a sharp border between both
colours (Fig. 108). Coxa III reddish-yellow, usu-
ally with infuscate dorsal spot at apex. Postpe-
tiole with fine longitudinal striation in its apical
half (Fig. 125). .........................................................
.................... A. leucogrammum (Holmgren, 1860)
Apex of clasper less clubbed apically, or stab-like.
Coxa III yellowish-red, usually without darker
dorsal spot. Postpetiole smooth or with fine
striation in apical third only (Fig. 124). ........ 17.
17. Lateral margins of face convergent ventrally
(Fig. 57). Clasper stab-like. Mesopleurum main-
ly yellowish or reddish-yellow. ..................... 18.
Lateral margins of face parallel or divergent
ventrally (Figs 53, 54, 56). Apex of clasper slight-
ly clavate (Fig. 102). Mesopleurum usually two-
or three-coloured: yellow, red, and black. ........
............................................................................. 19.
Figs 1-3. Dorsolateral carinae of first tergites of: 1. A. dorsale (Holmgren) W; 2. A. splenium (Curtis) W; 3. A. nigriceps
(Brischke).
1
2
3
92
18. Smaller, body length about 5 mm. Flagellum
with 28-32 flagellomeres. Mesopleurum almost
smooth. Face strongly convergent ventrally,
minimal width about 0.60-0.62 × as length of
eye. Usually parasitoids of Microgastrinae (Bra-
conidae) via Lepidoptera. .....................................
....... A. splenium var. plagiatum (Thomson, 1886)
Larger, body length 5.5-9 mm. Flagellum usu-
ally with more than 32 flagellomeres. Mesopleu-
rum often with scattered puncture. Face less
strongly convergent ventrally, minimal width
about 0.68-0.70 × of length of eye. Parasitoids of
various Ichneumonidae and Braconidae via
Lepidoptera. ...........................................................
........................... A. splenium s. str. (Curtis, 1833)
19. Mesopleurum two-coloured: red and black, usu-
ally with a sharp border between the colours.
Tibia III apically and tarsus III usually dark
brown or blackish. .................................................
.............................. A. tenuicorne (Thomson, 1886)
Mesopleurum three-coloured: yellowish ven-
trally, reddish centrally, with black margin
dorsally. Tibia III apically and tarsus III yellow-
ish or pale brown. .................................................
.................................. A. simplex (Thomson, 1886)
20. Temple as wide as or wider than the eye. Claw
III with strong teeth. .............................................
................................. A. diversum Schwenke, 1999
Temple narrower than the eye. Claw III with
small basal teeth. .............................................. 21.
21. Apical tergites with vespoid colour pattern:
brown basally and yellowish apically. ...............
.............................. A. uliginosum Schwenke, 1999
Apical tergites blackish, at most with narrow
yellowish apical margin. ................................. 22.
22. Pronotum reddish with yellowish margins,
mesosternum with a diffuse yellowish or reddish
stripe. Costula of propodeum usually faint or
lacking. Frontal orbit above antenna black. .......
........................................ A. alpinum Roman, 1909
– Pronotum usually blackish, pronotum and
mesopleurum at most with dark reddish suffu-
sion. Costula of propodeum usually present.
Frontal orbit above antenna usually with yel-
lowish spot (Fig. 53). ............................................
..................................... A. anale (Holmgren, 1860)
Description of species
Astiphromma (Astiphromma) aggressor
(Fabricius, 1804)
Figs 22, 67, 109
Ophion aggressor Fabricius, 1804: 132
Lectotype W: O. aggressor e Siellandia, Coll. Tønder Lund
(KOPENHAVN), designated by Horstmann 2001: 13,
not studied.
Mesochorus marginellus Holmgren, 1860: 121, synonymized
by Horstmann (2001: 13).
Lectotype W: Hlm, Bhn (STOCK), studied.
Astiphromma caecum Schwenke, 1999: 24, W syn. nov.
Holotype W: I, VI, Ricoaro, Campogrosso, 1500 m, 1.7.85
(ZSM), studied.
Astiphromma barbatulum Schwenke, 1999: 24, syn. nov.
Holotype W: Campi, Riva s. Garda, 1400 m. E/7.7.66 Hbth
(ZSM), studied.
Taxonomical remark. This taxon seems to be highly
variable in body size and metasomal colour pattern.
Therefore, A. caecum Schwenke and A. barbatulum
Schwenke described by single specimens are small
but typical examples of A. aggressor in my opinion.
A. caecum is similar to the lectotype of A. marginellus
with almost completely blackish tergites, A. bar-
batulum shows the commoner feature of metasomal
colour of A. aggressor, e. g. apical margin of tergite
2, median sandclock-shaped stripe of tergite 3 and
often stripe of tergite 4 yellowish-red.
Description of W
A variable species. Body length 5.0-9.5 mm. Flagel-
lum with (29)33-42 segments, slender, FEI 0.65-0.7,
preapical flagellomeres 1.5-2 × as long as wide, the
apical one blunt. Mandibular teeth of equal size.
Face punctate and granulate, lateral margins paral-
lel or slightly divergent ventrally, its minimal width
1.0-1.1 × eye length (Fig. 22). Temples strongly nar-
rowed behind eyes, TEI 0.65-0.8. Frons and vertex
without punctures. OOD 1.4-1.5. MI 0.6-0.7. Genal
carina joining the hypostomal one far from man-
dibular base.
Lower mesopleurum with fine scattered hair-
punctures. Scutellum not carinate. Area superomedia
1.8-2 × as long as wide, its lateral carinae more or
less narrowed apically, but often faint. Area petiolaris
slightly wider than long, about 0.3 × as long as pro-
podeum. Femur III 3.9-4.8 x as long as wide, inner
spur of tibia III 0.45-0.55 × as long as metatarsus III,
metatarsus III 0.5-0.6 × as long as tibia III and about
7.5 × as long as wide. Claws III with strong teeth.
Nervulus postfurcal (0.3-0.5 its length). Nervel-
lus of hind wing intercepted in the apical 0.2-0.25.
93
Pterostigma slender, 3.3-4.3 × as long as wide, RI
1.2-1.3, radius behind areolet straight or slightly
sinusoid (Fig. 109).
First tergite 2.3-2.8 × and postpetiole 1.2-1.4 × as
long as wide. Second tergite 0.9 × as long as wide, its
thyridium roundish or transverse. Ovipositor sheath
5.8-7.0
× as long as wide, narrowed in the apical
0.4, 0.8-1.2 × as long as the metatarsus III (Fig. 67).
Colour: Black. Ventral margin of face, clypeus
in its apical half and usually whole malar space
reddish-yellow, rarely almost completely blackish.
Flagellum brown dorsally and reddish or yellowish
ventrally. Coxa I and trochanter I yellowish, legs
otherwise reddish-yellow, coxa III sometimes with
reddish-brown spots, femur III often with an dorsal
infuscate mark apically, tibia III yellowish, at base
narrowly and at apex (0.15-0.25) infuscate. Tarsus
III reddish or brownish. Pterostigma pale-brown.
Apical margin of tergite 2 and often median stripe of
tergite 3, sometimes also of tergite 4 yellowish-red,
apical tergites sometimes more or less vespoid. In
the lectotype of A. marginellum and some other small
specimens metasoma blackish, only with narrow
apical yellowish margin of tergite 2. Median folds of
sternites yellow, lateral parts brownish. Ovipositor
sheath black, with pale tip.
Description of M
Body length 5.0-9.0 mm. Minimal width of face
about 0.85-0.9 × eye length. Flagellum with (26)33-41
segments. Lateral margins of face slightly divergent.
Femur III (4.1)4.6-4.9 × as long as wide. Metatarsus III
about 10 × as long as wide. Lateral carinae of area su-
peromedia often obsolete in the apical third. Clasper
about 1.4-1.6 × as long as the second tarsomere III.
Claw III with small basal teeth.
Colour: Black. Clypeus, face and lower 0.2
of temple yellowish. Flagellum yellowish below,
infuscate at apex. Mesosternum with dark reddish
suffusion, scutellum black. Tergite 2 with a large
triangular spot at apical margin, tergite 3 with a
sand-clock shaped yellowish-red median stripe.
Femur III sometimes with infuscate apex. Tarsus III
yellowish or brownish. Otherwise coloured as the W.
Hosts. Craesus latipes (Villaret, 1832) (Hym., Ten-
thredinidae) (Constantineanu & Mustata 1981),
Periclista lineolata (Klug, 1816) (Hym., Tenthredini-
dae) (Ozols 1961).
Material. 67 WW, 55 MM.
Distribution. Austria (CS, LINZ), Bulgaria (ZSM),
Byelorussia* (ZSM), Czechia (LINZ), Finland (CJ),
France (LAU), Germany (SCHNEE, ZSM), Great
Britain* (EDI, NHM), Italy (ZSM), Norway (CR),
Poland (WARS), Sweden (STOCK), Switzerland*
(LAU), Turkey* (ZSM).
Astiphromma (Astiphromma) albitarse
(Brischke, 1880)
Figs 23, 52, 63, 68
Mesochorus albitarsus Brischke, 1880: 191, M, type lost.
Astiphromma nigrum Pfankuch, 1921: 156, W, synonymized
by Schwenke (1999: 21).
Holotype W: Bremen 22.6.16 (BERLIN), not studied.
Astiphromma heydeni Habermehl, 1923: 270, W, synony-
mized by Schwenke (1999: 21), type not studied.
Pseudoacoenitus transsylvanicus Kiss, 1924: 96, W, syno-
nymized by Mócsár (1968: 183),
Holotype W: Hadad Dr. Kiss (BUDAPEST), not studied.
Description of W
Body length 8-11 mm. Flagellum with 40-45 flagel-
lomeres, FEI about 0.6, preapical flagellomeres 1.5-
2 × as long as wide, the apical one pointed. Temple
strongly narrowed behind eye, TEI 0.6. Face densely
punctate and granulate, lateral margins parallel, its
minimal width 0.9 × eye length (Fig. 23). Frons and
vertex smooth or with scattered fine punctation and/
or granulation, temple punctured. OOD 1.3-1.5.
MI 0.35-0.4. Genal carina joining the hypostomal
one far from mandibular base, hypostomal carina
not elevated.
Lower half of mesopleurum with fine and scat-
tered to dense puncture. Scutellum with distinct
lateral carina in more than the basal 0.5, usually
completely carinate (Fig. 63). Areae basalis and su-
peromedia confluent, area superomedia broad (about
1.3-1.5 × as long as wide), area petiolaris large, about
0.4 × as long as propodeum, hexagonal, somewhat
wider than petiolar base. Femur III 4.1-4.5 × as
long as wide. Inner spur of tibia III 0.5 × as long as
metatarsus III, metatarsus III about 0.56 × as long
as tibia III and 9.5 × as long as wide. Claw III with
strong teeth.
Nervulus strongly postfurcal (0.4-0.6 its length).
Nervellus of hind wing intercepted in the apical 0.25.
Second recurrent vein joining areolet in its middle.
Pterostigma 3.6-4.3 × as long as wide, RI 1.3, radius
behind areolet sinusoid.
First tergite 2.2-2.4 × and postpetiole 1.2-1.35 × as
long as wide, the latter with longitudinal impression
medially. Tergite 2 about 1.0-1.1 × as long as wide, its
thyridium small, roundish or transverse. Ovipositor
sheath about 6.9-7.1 × as long as wide and as long as
metatarsus III, narrowed in the apical 0.4 (Fig. 68).
94
Colour: Black. Yellowish are palps and mandible
except teeth. Yellowish-red are tegula, and hind
edge of pronotum. Legs yellowish-red, coxae I-II
more or less brownish, coxa III mainly dark brown.
Femur III brown, paler at base. Tibia III infuscate
in the apical 0.3-0.4, tarsus III infuscate. Metasoma
black, tergite 2 with a narrow yellow hind margin,
sternites yellowish medially and brownish laterally,
hypopygium brown. Pterostigma brown. Ovipositor
sheath black, with pale tip.
Description of M
Flagellum with 40-42 flagellomeres. Lateral facial
margins convergent ventrally (Fig. 52). OOD 1.2.
Frons more or less smooth and shining. Femur III
4.9 × as long as wide. Pterostigma 4.1 × as long as
wide. Nervulus postfurcal (0.3 its length). Claw III
with short basal teeth. Clasper about 1.5-1.7 × as long
as the second tarsomere III, stab-shaped.
Colour: Black. Yellow are face, clypeus, man-
dible except teeth, palps, malar space, lower 0.2
of temple, tegula, hind edge of pronotum, and
subtegular ridge. Cream-yellowish are coxae I-II,
trochanters, trochantelli and tarsi. Legs except
coxae and trochanters reddish, tibia III yellowish to
cream-yellow. Coxa III, femur III and apical 0.25-0.3
of tibia III brownish to blackish, Coxa III ventrally
and sometimes femur III basally yellowish. Tergite
2 with reddish-yellow apical margin or triangular
apical spot and usually tergite 3 with reddish-yellow
sandclock-shaped stripe. Clasper black, tip yellow-
ish.
Hosts. Unknown.
Material. 31 WW, 19 MM.
Distribution. Austria (LINZ), Bulgaria* (CR), Ger-
many (CR, NHM, ZSM), Great Britain* (EDI, NHM),
Italy* (CG), Netherlands* (CZ), Sweden* (NHM,
STOCK), Switzerland* (LAU).
Astiphromma (Astiphromma) alpinum
Roman, 1909
Figs 4, 9, 24, 69, 115
Holotype W: LnL [back side] 1904 (STOCK), studied.
Astiphromma dispersum Schwenke, 1999: 20, W, syn. nov.
Holotype W: Kvikkjokk/S, 11.7.64, R. Hinz (ZSM), stu-
died.
Astiphromma laricis Schwenke, 1999: 24, M, syn. nov.
Holotype M: 60 L. 49, ex. L. laricis 1960, H. Pschorn-
Walcher (ZSM), studied.
Description of W
Body length 5.5-6 mm. Flagellum with 29-35 flagel-
lomeres, FEI 0.6-0.7, preapical flagellomeres about
1.5 × as long as wide, the apical one blunt. Face
wide, its minimal width 1.0 × eye length, punctate
and finely granulated, matt, lateral margins parallel
(Fig. 24). Frons with fine granulation, without dis-
tinct punctation. Temples strongly narrowed behind
eyes, with scattered punctation, TEI 0.7-0.9 (Fig. 9).
MI 0.7. OOD 1.3-1.7. Genal carina joining the hypo-
stomal one far from mandibular base (1
/2-2
/3 of malar
Figs 4-5. Joint of genal and hypostomal carinae (arrow): 4. A. alpinum Roman W; 5. A. italicum Schwenke W.
4
5
95
space) (Fig. 4), hypostomal carina slightly elevated.
Lower half of mesopleurum with fine scattered
hair-punctures or almost smooth. Scutellum not cari-
nate. Propodeum with carination, but costula usually
faint or lacking. Area superomedia narrow, 2.5-3 ×
as long as wide, behind costulae with almost parallel
sides. Area petiolaris about as long as wide, 0.35 ×
as long as propodeum. Femur III 4.0-4.7 × as long as
wide. Inner spur of tibia III about 0.53 × as long as
metatarsus III, metatarsus III 0.45 × as long as tibia
III and 8.5 × as long as wide. Claw III with 4 teeth.
Nervulus postfurcal (0.1-0.4 × its length). Nervel-
lus of hind wing intercepted in the apical 0.25-0.4,
discoidella distinct. Second recurrent vein joining
areolet in its middle. Pterostigma 3.2-3.6 × as long
as wide, RI 1.15-1.2, radius behind areolet almost
straight.
First tergite 2.1 × and postpetiole about 1.25 ×
as long as wide. Second tergite 0.8-1.0 × as long as
wide, its thyridium large and transverse. Ovipositor
6 7
8
10
9
11
1312
Figs 6-13. Head of WW from dorsal: 6. A. luridum Schwenke; 7. A. italicum Schwenke; 8. A. nigrocoxatum (Strobl);
9. A. alpinum Roman; 10. A. buccatum (Thomson); 11. A. dorsale (Holmgren); 12. A. hirsutum (Bridgman); 13. A. leu-
cogrammum (Holmgren).
96
sheath about 6.2-7.7 × as long as wide and 0.8-1.0 ×
as long as metatarsus III, narrowed in the apical
half (Fig. 69).
Colour: Black. Yellowish are mandible except
teeth, clypeus, ventral 1
/3-1
/4 of facial orbit, hind
edge of pronotum and tegula. Flagellum reddish to
brown. Tergite 2 in the apical 0.2 yellowish, base of
tergite 3 with a narrow yellowish band or triangular
spot. Coxae I-II and all trochanters yellowish. Legs
otherwise reddish to reddish-yellow. Tibia III more
yellowish and slightly infuscate in apical 0.15-0.2.
Tarsus III more or less infuscate. Tergites black-
ish, hind margin of tergite 2 and sometimes basal
margin of tergite 3 with narrow yellowish bands.
Epipleura and sides of sternites brownish, sternites
yellow medially (Fig. 115). Pterostigma brownish or
yellowish-brown. Ovipositor sheath black, reddish
in the apical 0.2.
Description of M
Body length 5.5-6 mm. Flagellum with 31 flagello-
meres, FEI 0.63. Face punctate and granulate, lateral
margins of face divergent ventrally. Frons granulate
laterally and smooth medially, vertex with very scat-
tered superficial punctation. Temple granulate, only
slightly narrowed, TEI 0.9. OOD 1.3. Genal carina
meets the hypostomal one far from mandibular base.
Mesopleurum with fine very scattered punc-
ture. Mesoscutum finely granulate, with rugulose-
punctate area in front of the prescutellar groove.
Scutellum not carinate. Area superomedia narrow,
costula faint. Area petiolaris as long as wide. Femur
III 4.4 × as long as wide. Claw III with basal teeth.
Nervulus postfurcal (0.4 of its length), nervellus
of hind wing intercepted in the apical 0.25. Pter-
ostigma 3.6 × as long as wide, radius behind areolet
straight, RI 1.1.
First tergite 2.4 × and postpetiole 1.3-1.4 × as long
as wide. Postpetiole smooth. Second tergite 1.0-1.1 ×
as long as wide, its thyridium roundish. Clasper
retracted, about as long as the second tarsomere III.
Colour: Black. Scape and pedicellus yellow
ventrally. Face, clypeus, lower 0.2 of temple, pro-
pleurum, pronotum reddish, its lower and hind
edges mainly yellow. Mesosternum with a diffuse
brownish tinge or yellowish stripe. Tergite 2 in the
apical 0.2, and tergite 3 with a sand-clock shaped
yellowish stripe. Sternites 1-3 yellow medially, with
brown lateral spots, sternites 4-7 brownish, with nar-
row yellow apical margins. Pterostigma brownish-
yellow. Legs yellowish. Femur III weakly infuscate
at apex. Tibia III in the apical 0.15 slightly infuscate,
metatarsus III in the apical 0.3, and tarsomeres III
completely infuscate.
Hosts. Pristiphora laricis (Hartig, 1837) (Hym., Ten-
thredinidae) (NHM, ZSM), Pachynematus imperfectus
(Zaddach, 1876) (Hym., Tenthredinidae) (NHM).
Material. 18 WW, 6 MM.
Distribution. Germany (CR, ZSM), Great Britain*
(NHM), Italy (ZSM), Poland (ZSM), Sweden (STOCK,
ZSM), Switzerland* (NHM).
Astiphromma (Astiphromma) anale
(Holmgren, 1860)
Figs 26, 27, 53, 70
Mesochorus analis Holmgren, 1860: 120, WM, types not
studied.
Description of W
Body length 6 mm. Flagellum with 26-31 flagel-
lomeres, FEI 0.65-0.7, preapical segments 1.2-2 × as
long as wide. Mandibular teeth of equal size. Face
punctate and granulate, matt, lateral margins parallel
ventrally, its minimal width about 0.9-1.0 × as the
length of eye (Fig. 26). TEI 0.55-0.65. Lower frons
granulate laterally, smooth and shining medially.
OOD 1.4-1.6. MI 0.6-0.7. Genal carina reaches the
hypostomal one far from mandibular base, hypos-
tomal carina slightly elevated.
Lower half of mesopleurum with very fine and
scattered hair-puncture. Scutellum not carinate.
Propodeum with almost complete carination, areae
basalis and superomedia confluent, area supero-
media about 2 × as long as wide. Area petiolaris
large, wider than the petiolar base, 0.35 × as long
as propodeum. Femur III 3.8-4.3 × as long as wide,
inner spur of tibia III 0.6 × as long as metatarsus III,
metatarsus III 0.45 × as long as tibia III and 9-9.5 ×
as long as wide. Claws III with about 3 teeth in the
basal 2
/3.
Nervulus postfurcal of 0.3-0.5 its length. Pter-
ostigma 3.8 × as long as wide, RI 1.2-1.25, radius
behind areolet straight.
First tergite 2.4 × and postpetiole about 1.6 × as
long as wide apically, usually with a median longi-
tudinal impression. Thyridium of tergite 2 roundish.
Ovipositor sheath about 6-7.1 × as long as wide
and 0.9 × as long as metatarsus III, narrowed in the
apical fourth, tip sometimes slightly bent towards
dorsal (Fig. 70).
Colour: Black. Cream-yellow are palps and
mandible except teeth, clypeus, face (Figs 26, 27),
usually a small spot on frontal orbit above antenna
(Fig. 53), gena, tegula, hind edge of pronotum and
subtegular ridge, coxae I-II and all trochanters.
Scutellum rarely with reddish suffusion. Scape and
97
pedicellus yellowish, flagellum pale brownish or
yellowish. Legs yellowish-red, tibia III yellowish, at
apex indistinctly infuscate. Pterostigma yellow, with
darker margins. Tergite 2 with reddish band in apical
0.2, tergite 2 with sandclock-shaped reddish median
stripe, tergite 4 reddish with brownish sides, follow-
ing tergites reddish or yellowish-red, rarely tergite
4-7 dark brownish. Sternites reddish or yellowish,
sometimes with brown spots laterally. Ovipositor
sheath yellowish-red or brownish.
Description of M
Flagellum with 27-34 flagellomeres. TEI 0.5-0.65.
OOD 1.3-1.5. MI about 0.5. Nervulus slightly to dis-
tinctly postfurcal. Femur III 3.9-4.5 × as long as wide.
Clasper 1.0-1.2 × as long as the second tarsomere III.
Colour: Black. Clasper yellowish. Vertex rarely
with yellowish spot. Sometimes pronotum and/or
mesosternum with dark reddish-brown suffusion.
Femur III sometimes with narrowly infuscate apex.
Tibia III yellowish-red, sometimes apical 0.25 infus-
cate. Otherwise as in female.
Hosts. Unknown.
Material. 48 WW, 65 MM.
Distribution. Austria (CS, NHM, ZSM), Byelorus-
sia* (ZSM), Estonia* (CJ), Finland (CJ, NHM, ZSM),
France* (LINZ, NHM. ZSM), Germany (CR, ZSM),
Great Britain* (NHM), Netherlands* (CZ), Norway
(CR), Poland (WARS), Russia* (LINZ, STOCK),
Switzerland (NHM, ZSM), Sweden (STOCK, ZSM).
Astiphromma (Astiphromma) buccatum
(Thomson, 1886)
Figs 10, 25, 54, 64, 71, 110
Mesochorus (Astiphrommus) buccatus Thomson, 1886: 329,
WM.
Lectotype W: Tve 6/78 (LUND), designated by Fitton,
1982: 52, studied.
Mesochorus (Astiphrommus) hamulus Thomson, 1886: 330,
syn. nov.
Holotypus (herewith designated) M: Strandm Drewsen,
Danmark ex coll. Schiødte (KOPENHAVN), not
studied.
Astiphromma consertum Schwenke, 1999: 23, W, syn. nov.
Holotype W: Leyman/Alsace/F, 6.60/4.61, F. Daniel
(ZSM), studied.
Taxonomical remark. Thomson (1886: 329) in his
original publication did not mention how many
specimens he had for his description but arranged
A. hamulum within the group of yellow-faced fe-
males. Discussing the type material of A. hamulum,
Fitton (1982: 53) stated that there is no direct evidence
that the specimen in Copenhagen is a syntype and
that no typical material was found in the Thomson
collection in Lund. The specimen of Thomson from
Lund, which I studied, was labelled as a lectotype by
Schwenke, but is in fact a male of Astiphromma anale
(Holmgren). So far, only males of A. hamulum have
been found in all the examined European collections,
and the available literature mentioned males only
(Smits van Burgst 1918, Morley 1915, Jussila 2010).
I therefore suggest that Thomson made a lapsus
to include the species into the group of yellow-faced
Astiphromma females. Since A. hamulum shows many
structural details (larger lower mandibular tooth,
genal carina reaching mandibular base, strongly
developed temple) in common with the females
of A. buccatum (Thomson), I herewith synonymize
both taxa.
Description of W
Body length 6-7 mm. Flagellum with 32-35 flagel-
lomeres, FEI 0.64-0.72, preapical segments 1.5 ×
as long as wide. Apical flagellomere blunt. Lower
mandibular tooth slightly larger than the upper one.
Clypeus apically blunt and slightly protruding. Face
with coarse and dense puncture, lateral margins
usually divergent ventrally, its minimal width 1.1 ×
eye length (Fig. 25). Temples parallel behind eyes,
slightly narrowed apically, punctate, TEI 0.9-1.3
(Fig. 10). Frons and vertex granulate and with fine
punctures, temple with distinct punctation. Frons
sometimes with transverse striae laterally. OOD 1.4-
1.5. MI 0.65. Genal carina joining the hypostomal one
at or close to mandibular base, hypostomal carina
slightly elevated.
Lower half of mesopleurum with dense puncta-
tion. Scutellum not carinate. Propodeum with almost
complete carination. Area superomedia confluent
with area basalis, 2.5-3 × as long as wide, slightly
widened apically or with parallel sides. Area peti-
olaris large, wider than the petiolar base, about 0.3 ×
as long as propodeum. Femur III 4.1-4.7 × as long as
wide, inner spur of tibia III 0.5 × as long as metatarsus
III, metatarsus 0.48 × as long as tibia III and 9.5 × as
long as wide. Claw III with strong teeth.
Nervulus interstitial or slightly postfurcal (0.2
its length). second recurrent vein joining areolet in
its middle, pterostigma 3.3-3.6 × as long as wide, RI
0.9-1.1, radius behind areolet straight. Nervellus of
hind wing intercepted in the apical 0.2, discoidella
more or less distinct.
First tergite short, 2.1-2.3 × and postpetiole 1.0-
1.2 × as long as wide. Postpetiole with superficial
punctation. Second tergite about as long as wide, its
thyridium small, transverse or oblique. Ovipositor
98
sheath short, 5.2-6.2 × as long as wide and 0.8-1.3 ×
as long as metatarsus III (Fig. 71).
Colour: Black. Scape and pedicellus often, hind
edge of pronotum and tegula always yellow, fla-
gellum reddish basally, apically infuscate. Clypeus
completely black or narrowly reddish, ventral
margin of face beside clypeus narrowly reddish or
yellowish. Tergite 2 with a narrow reddish apical
margin, tergite 3 black or brownish, following ter-
gites black with narrow yellowish apical margins.
Sometimes all tergites except postpetiole with narrow
pale apical margins. Sternites yellowish with brown
lateral spots. Legs reddish-yellow, trochanter III
and tibia III more or less yellowish, tibia III usually
infuscate in the apical 0.1-0.2, tarsus III brownish
or infuscate. Pterostigma brown. Ovipositor sheath
black, with reddish tip.
14
16
18
20
19
21
17
15
Figs 14-20. Head of WW from dorsal: 14. A. flagellator spec. nov.; 15. A. flavoventrale spec. nov.; 16. A. striatum (Brisch-
ke); 17. A. nigriceps (Brischke); 18. A. scutellatum (Gravenhorst); 19. A. tenuicorne (Thomson); 20. A. varipes (Holmgren).
Fig. 21. Hind femur of A. splenium (Curtis) W.
99
Description of M
Body length 6-9 mm. Flagellum with (31)33-40
flagellomeres, FEI 0.6-0.65, preapical segments about
1.5 × as long as wide. Apical flagellomere blunt.
Lower mandibular tooth slightly larger than or equal
to the upper one. Face with coarse and dense punc-
tation, shining, lateral margins divergent ventrally
(Fig. 54). Temples parallel behind eyes, moderately
narrowed behind, TEI (0.7) 1.0-1.1. Frons, vertex, and
temple with scattered to dense, distinct punctures,
shining. OOD 1.2-1.6. MI 0.55-0.65. Genal carina
joining the hypostomal one close to mandibular base,
hypostomal carina slightly to moderately elevated.
Lower half of mesopleurum with fine and
22 23
25
27
24
26
Figs 22-27. Face of WW from frontal: 22. A. aggressor (Fabricius); 23. A. albitarse (Brischke); 24. A. alpinum Roman;
25. A. buccatum (Thomson); 26-27. A. anale (Holmgren).
100
moderately dense (or dense) puncture. Scutellum
not carinate. Propodeum with complete carination,
area superomedia usually strongly narrowed api-
cally, 2.5-3.5 × as long as wide. Area petiolaris much
wider than long, kidney-shaped. Femur III 4.3-4.8 ×
as long as wide. Inner spur of tibia III 0.5 × as long
as metatarsus III, metatarsus III 0.45-0.5 × as long as
tibia III. Claw III with about 5 strong teeth.
Nervulus interstitial to slightly postfurcal (0.2 its
length). second recurrent vein joining areolet in or
basal to its middle. Pterostigma 3.3-3.6 × as long as
wide, RI 1.0, radius behind areolet straight.
First tergite 2.6-2.9 × and postpetiole 1.4-1.5 × as
long as wide. Postpetiole smooth, often with a more
or less distinct longitudinal median impression. Sec-
ond tergite 1.2-1.5 × as long as wide, its thyridium
transverse or oblique, small. Clasper stab-like, about
1.0-1.2 × as long as the second tarsomere III.
Colour: Black. Scape below, pedicellus and ba-
sal flagellomeres on outer side yellowish. Clypeus,
face, lower frontal orbit, malar space, lower third
of gena, mandible except teeth, palps, coxae I-II
and trochanters I-II, all trochantelli cream-yellow.
Propleurum, pronotum mainly, mesosternum and
lower half of mesopleurum yellowish, sometimes
mesosternum partly reddish. Mesoscutum with
four yellowish longitudinal stripes laterally and
submedially (the submedial ones sometimes short),
confluent at anterior margin (Fig. 64). Tergites 2-6
with narrow reddish-yellow apical margins, tergite
3 with a sand-clock shaped reddish-yellow median
stripe. Epipleura and sternites yellow, sometimes
sternites brownish laterally. Legs reddish-yellow,
tibia III infuscate in the apical 0.1, metatarsus III in
the apical 0.3, and following tarsomeres III more
or less infuscate. Pterostigma yellowish-brown to
brown.
Hosts. Periclista sp. (Hym., Tenthredinidae) (ZSM),
Pachynematus sp. (Hym., Tenthredinidae) (Schwenke,
1999), Diprion sp. (Hym., Diprionidae) (Schwenke,
1999).
Material. 8 WW, 6 MM.
Distribution. Bulgaria* (ZSM), Germany (ZSM),
Great Britain (NHM), Morocco* (LINZ), Netherlands
(CZ), Spain (LINZ, WARS).
Astiphromma (Astiphromma) diversum
Schwenke, 1999
Figs 28, 59, 72
Astiphromma diversum Schwenke, 1999: 24, WM.
Holotypus W: Dessau/D, 4,4 km SW, 17.5.93, H. Zoerner
(SDEI), not studied.
Taxonomical remark. Schwenke had determined
some females and males as A. buccatum Thomson
(material in ZSM), but A. diversum can easily be
differentiated by the form of genal carina and the
equally sized mandibular teeth.
Description of W
Body length 5.5-6.5 mm. Flagellum with 34-37 flagel-
lomeres, short, FEI 0.67-0.72, preapical flagellomeres
1.5-2 × as long as wide, apical flagellomere pointed.
Mandibular teeth of almost equal size, mandible with
a strong ventral lamella. Face strongly punctate, with
fine granulation, but more or less shining, its width
1.1 × eye length, lateral margins divergent ventrally
(Fig. 28). Frons granulate, and more or less punctate
laterally. Temple more or less narrowed behind eye,
punctate, TEI 0.8-0.9. OOD about 1.5. MI 0.6-0.7.
Genal carina joining the hypostomal one distant
from mandibular base (about 1
/3-1
/2 of malar length),
hypostomal carina slightly elevated.
Lower half of mesopleurum with fine but dense
punctures. Scutellum not carinate. Propodeum with
complete carination, area superomedia slender,
2.5-2.7 × as long as wide, with almost parallel sides.
Area petiolaris large, hexagonal, 0.35 × as long as
propodeum. Femur III 4.0-4.8 × as long as wide. In-
ner spur of tibia III 0.5-0.52 × as long as metatarsus
III, metatarsus III about 0.45 times as long as tibia
III. Claw III with 4 strong teeth (Fig. 59).
Nervulus postfurcal (0.2-0.5 its length), nervellus
in hind wing intercepted almost in the middle or
apical third, discoidella distinct. Pterostigma about
3.9 × as long as wide, RI 1.15, radius behind areolet
straight. Second recurrent vein reaching areolet in
or beyond its middle.
First tergite 2.2 × and postpetiole 1.2-1.3 × as long
as wide. Second tergite 0.85-0.9 × as long as wide,
its thyridium large, roundish. Ovipositor sheath
about 6.5 × as long as wide and 0.85-0.9 × as long as
metatarsus III, slightly lanceolate, narrowed in the
apical half (Fig. 72).
Colour: Black. Ventral margins of face narrowly
reddish, malar space partly yellowish, clypeus al-
most completely black. Tegula and hind edge of
pronotum cream-yellow. Legs reddish. tibia III
more yellowish-red, slightly infuscate in the apical
0.1, tarsus III more or less infuscate. Tergites black,
only with very narrow paler hind margins, sternites
medially yellow, laterally black. Pterostigma brown.
Ovipositor sheath black, with pale tip.
Description of M
Body length 5-6.5 mm. Flagellum with 31-35 flagel-
lomeres. Facial sides divergent ventrally or parallel.
Temples slightly narrowed behind eyes, TEI 0.9-1.1.
101
Vertex with scattered puncture. OOD 1.1-1.5. Lower
mesopleurum with small but dense or scattered
punctation. Area petiolaris wider than petiolar base.
Nervulus slightly postfurcal (0.1-0.2), nervellus in
hind wing intercepted in its apical 0.2-0.6. Pter-
ostigma 3.5 × as long as wide, RI 1.25. Femur III 4.2
times as long as wide. Inner spur of tibia III 0.5 × as
long as metatarsus III, metatarsus III 0.47 × long as
tibia III. Claw III with 4 teeth. Clasper about 1.1-1.3
times as long as the second tarsomere III.
Colour: Black. Face, clypeus, mandibles except
teeth, palps, malar space, gena in the lower 0.2, hind
pronotal edge yellowish. Legs yellow-red, coxae
and trochanters I-II (III) more yellowish. Tibia III
slightly darkened at apex. Tarsus III more or less
infuscate. Bases of tergites 3 and 4 with a median
yellowish spot or tergite 3 with a median reddish
stripe. Pterostigma brownish.
Host. Pristiphora sp. (Hym., Tenthredinidae) via Trem-
atopygus sp. (Hym., Ichneumonidae) (Schwenke 1999).
Material. 2 WW, 14 MM.
Distribution. Austria (LINZ, NHM, ZSM), Bulgaria*
(ZSM), Czechia (LINZ), Germany (CR, SCHNEE),
Netherlands* (CZ), Sweden* (STOCK), Switzerland
(CA).
Astiphromma (Astiphromma) dorsale
(Holmgren, 1860)
Figs 1, 11, 29, 61, 73, 105, 126, 127
Mesochorus dorsalis Holmgren, 1860: 117, W M, types not
studied.
Mesochorus scutellatum Brischke, 1880: 180, praeocc. by
M. scutellatum Gravenhorst, 1829, W, synonymized
by Schmiedeknecht (1910: 1946).
Taxonomical remark. Lee (1992) published a revi-
sion of the world species of Astiphromma having
an apical tubercle or granule on scutellum (group
B of Nakanishi (1969)). He differentiated the taxa
A. dorsale and A. granigerum (recte A. hirsutum) by the
structure of the second tergite and clypeal foveae,
and the density of mesopleural punctures. However,
a detailed analysis of the European material of both
taxa revealed a large variability of all these charac-
ters. In my opinion, the morphological features used
by Lee (1992) are not reliable for a clear differenta-
tion of the European material. It seems that he used
a mix of both taxa for his description of A. dorsale.
Here, the form and length of ovipositor sheath,
of the hind femur and hind metatarsus have been
used to differentiate both taxa (see Figs 126-127).
However, some intermediate specimens could not
be determined doubtlessly.
Description of W
Body length very variable (6-11 mm). Flagellum
with 35-41 flagellomeres, FEI 0.6-0.65, preapical
segments about 2 × as long as wide. Mandibular
teeth equally sized. Facial width 1.0 × eye length.
Lateral facial margins more or less parallel, face
punctate and granulate, matt, with some oblique
striation beside clypeus (Fig. 29). Frons with fine
granulation. Temple punctate, TEI 0.9-1.0 (Fig. 11).
Genal carina joining the hypostomal one far from
mandibular base.
Lower part of mesopleurum with usually fine
scattered (rarely dense) punctation or almost smooth
(Fig. 105). Scutellum not carinate, pointed apically.
Propodeum short, apical transverse carina of area
superomedia usually faint or lacking, carination
otherwise complete. Area superomedia 1.3-1.6 × as
long as wide. Area petiolaris very large, wider than
long, about 0.45-0.5 × as long as propodeum. Femur
III stout, 3.8-5.1 × as long as wide. Metatarsus III
stout, about 6-7.5 × as long as wide and 0.40-0.45 ×
as long as tibia III. Inner spur of tibia III 0.6-0.7 × as
long as hind metatarsus. Hind claw strongly curved
apically (90°) and with strong and long teeth (Fig. 61).
Nervulus slightly postfurcal (0.3 × its length).
Second recurrent vein joining areolet basal of its mid-
dle. Pterostigma 3.6-3.8 × as long as wide, RI 1.2-1.3.
First tergite 2.3 × and postpetiole 1.0-1.1 × as long
as wide. Second tergite 0.8-1.1 × as long as wide, its
thyridium large and roundish. Ovipositor sheath
about 5.2-7.5 × as long as wide and 0.95-1.3 × as
long as the hind metatarsus (Fig. 73).
Colour: Black. Flagellum usually infuscate.
Yellowish are clypeus and malar space, face beside
clypeus, tegula, hind edge of pronotum and sub-
tegular ridge, coxae and trochanters I-II. Scutellum,
and sometimes mesoscutum, mesopleurum and
metapleurum more or less reddish. Legs reddish-
yellow, tibia III infuscate basally and apically, tarsus
III usually blackish. Pterostigma brown. Ovipositor
sheath black, with pale tip.
Variety: 1 specimen from Byelorussia (ZSM)
with second tergite 1.3 × as long as wide.
Description of M
Body length 6-9 mm. Flagellum with 34-37 flagel-
lomeres, FEI 0.6. Minimal width of face 0.8-0.9 ×
eye length. TEI 0.75-0.9. OOD1.0. Frons beneath
ocelli with fine puncture. Lower mesopleurum with
scattered punctation or almost smooth. Femur III
4.7-5.0 × as long as wide. Metatarsus III 0.45 × as
long as tibia III. First tergite 2.35 × and postpetiole
1.2 × as long as wide. Second tergite 0.95 × as long as
wide. Clasper about 1.2-1.3 × as long as the second
tarsomere III.
102
Colour: Black. Flagellum yellowish basally
and infuscate apically. Cream-yellow are mandible
except teeth, palps, clypeus, face, gena, lower third
of temple, hind edge of pronotum, subtegular ridge,
tegula, coxae I-II and all trochanters and trochantelli.
Propleurum, pronotum, mesopleurum, sometimes
mesoscutum, and scutellum more or less reddish.
Legs yellowish-red, tibia III in the apical 0.1-0.2,
and often tarsus III more or less infuscate. Metasoma
blackish, tergite 2 with a narrow yellowish apical
margin. Tergite 3 often diffusely reddish medially.
Sternites brownish, the frontal ones with yellow
median membranous areas. Pterostigma yellowish-
brown to brownish.
Figs 28-33. Face of WW from frontal: 28. A. diversum Schwenke; 29. A. dorsale (Holmgren); 30. A. flagellator spec.
nov.; 31. A. flavoventrale spec. nov.; 32. A. hirsutum (Bridgman); 33. A. italicum Schwenke.
28 29
31
33
30
32
103
Hosts. Panolis flammea (Denis & Schiffermüller,
1775) (Lep., Noctuidae) via Tachinidae (Diptera)
(Schwenke, 1999). Schwenke (1999) mentioned also
Meteorus sp. (Hym., Braconidae) as primary parasi-
toide, this report is very questionable.
Material. 64 WW, 27 MM.
Distribution. Austria (CS, CZ), Bulgaria (ZSM),
Byelorussia* (ZSM), Czechia (LINZ), France (CZ),
Germany (CR, CZ), Italy (CZ, ZSM), Netherlands
(CZ), Norway (CR), Russia (Moscow reg.) (LINZ),
Russia (Siberia) (CR), Switzerland* (LAU), Sweden
(STOCK).
A. dorsale (Holmgren) has not been found in
Britain so far, all specimens seen from Britain (EDI,
NHM) belong to A. hirsutum (Bridgman).
34 35
37
39
36
38
Figs 34-39. Face of WW from frontal: 34. A. leucogrammum (Holmgren); 35. A. longiceps (Strobl); 36. A. luridum
Schwenke; 37. A. striatum (Brischke); 38. A. nigrocoxatum (Strobl); 39. A. pictum (Brischke).
104
Astiphromma (Astiphromma) flagellator
spec. nov.
Figs 14, 30, 65, 75
Holotype W: Erlangen 10.3.1953, Mesochorus sp. n. R.
Bauer W, Holotypus, Astiphromma flagellator n. sp. det.
Riedel 2013 (ZSM).
Paratype W: NETHERLANDS: Wijster (Dr.) 28.v-11.vi.76
C. v. Achterberg (NHM).
Taxonomical remark. This new taxon resembles
A. buccatum (Thomson). It differs by the form of genal
carina, shorter flagellomeres, short first tergite, and
darkened coxa III.
Description of W
Body length 5.5-6.0 mm. Flagellum with 28 flagel-
lomeres, short, FEI 0.51, preapical flagellomeres only
slightly longer than wide (Fig.65). Mandibular teeth
equally sized or lower tooth slightly larger than
the upper one. Face, temple and frons with coarse
punctation (Fig. 30). Temple slightly narrowed be-
hind eye, TEI 0.9. Lateral margins of face divergent
ventrally, its minimal width 0.9 × eye length. OOD
1.5. MI 0.6. Genal carina joining the hypostomal one
more or less away from mandibular base, hypos-
tomal carina not elevated.
Lower half of mesopleurum with superficial but
dense punctation. Scutellum not carinate. Propo-
deum with complete carination, area superomedia
short, 1.5-1.7 × as long as wide, area petiolaris very
large, hexagonal, about 0.4 × as long as the propo-
deum. Femur III stout, 3.6-3.7 × as long as wide.
Metatarsus III about 0.45 × as long as tibia III and
9.5 × as long as wide. Claw III with 5 strong teeth.
Nervulus almost interstitial, nervellus in hind
wing intercepted in the apical 0.25, discoidella
weakly sclerotized. Pterostigma about 3.3 × as long as
wide, radius behind areolet slightly bent to anterior
wing margin, RI about 0.95.
Petiolus short, first tergite 2.1 × and postpetiole
1.1-1.2 × as long as wide, postpetiole in paratype
with median longitudinal impression. Second tergite
about 0.8 × as long as wide, thyridium very small,
slightly transverse. Ovipositor sheath 5.5-6.4 × as
long as wide and 0.9 × as long as metatarsus III,
narrowed in the apical half (Fig. 75).
Colour: Black. Flagellum infuscate. Clypeus
black, ventral margin of face and malar space
yellowish. Tegula, hind edge of pronotum and
subtegular ridge yellow. Coxa II basally and coxa
III dorsally brown, paler ventrally. Legs otherwise
reddish-yellow, tibia III yellowish, infuscate in
apical 0.15, tarsus III brownish. Tergite 2 (or 2-3)
with narrow yellowish apical margin. Sternites
yellow medially and more or less brown laterally.
Pterostigma brown. Ovipositor sheath blackish, with
pale tip.
M unknown.
Hosts. Unknown.
Material. 2 WW.
Distribution. Germany (ZSM), Netherlands (NHM).
Astiphromma (Astiphromma) flavoventrale
spec. nov.
Figs 15, 31, 76, 107, 119
Holotype W: Itl. [back] 1908, 22/7, Astiphr. ? incidens Thm
W var. (STOCK)
Paratype W: 15/7, Torne Tr. Malaise (STOCK), head
missing
Taxonomical remark. This new taxon can easily
be identified by its characteristic colour of face and
mesosoma. Unfortunately, the head of paratype is
missing.
Description of W
Body length 5mm. Flagellum stout, filiform, with 28
flagellomeres, FEI 0.8, preapical flagellomeres about
1.5 × as long as wide. Mandibular teeth equally sized.
Head granulate, matt. Face with superficial punc-
tation, lateral margins parallel ventrally (Fig. 31).
Temples slightly narrowed behind eyes, TEI 1.1
(Fig. 15). OOD about 1.3. MI 0.7. Genal carina join-
ing the hypostomal one far from mandibular base.
Hypostomal carina not elevated.
Lower part of mesopleurum with very dense
fine rugulose punctures, matt (Fig. 107). Upper
part punctate, speculum smooth. Scutellum not
carinate. Propodeum with complete carination, area
superomedia slender, 2.3-2.5 × as long as wide, with
costula in the basal third. Area petiolaris short, about
as long as wide. Coxa III rugulose, matt. Femur III
5.3 × as long as wide. Metatarsus III about 0.5 × as
long as tibia III. Inner spur of tibia III 0.43 × as long
as metatarsus III. Claw III with distinct teeth.
Nervulus interstitial. Nervellus of hind wing
intercepted in the lower 0.2, but discoidella not devel-
oped. Pterostigma about 3.3 × as long as wide, RI 1.0.
First tergite 2.1 × and postpetiole 1.25 × as long as
wide. Postpetiole with fine rugulo-striation. Second
tergite about 0.95 × as long as wide, without distinct
thyridium. Ovipositor sheath short, 5.5 × as long as
wide, and 0.8 × as long as metatarsus III, slightly
bent upwards (Fig. 76).
Colour: Black. Yellowish are clypeus, malar space,
mandible except teeth, lower 0.3 of temple, face
except a median rectangular black spot below the
105
antenna (Fig. 31), often subtegular ridge, lateral spot
and hind edge or upper margin of pronotum, tegula,
complete mesosternum, coxae and trochanters I-II,
ventral apical spot on coxa III, apical transverse
stripes of tergites 2
/3-7. Legs yellowish-red, coxa III
and trochanter III with a diffuse blackish stripe dor-
sally, tibia III indistinctly infuscate at apex, tarsus III
brownish. Pterostigma yellowish, margins brownish.
Ovipositor sheath reddish-brown.
M unknown.
Hosts. Unknown.
Material. 2 WW.
Distribution. Sweden (STOCK).
Astiphromma (Astiphromma) hirsutum
(Bridgman, 1883)
Figs 12, 32, 58, 74, 106, 126, 127
Mesochorus hirsutus Bridgman, 1883: 168.
Paralectotypes 1W, 2M: Mesochorus hirsutus Bridgman
1883 det. M. G. Fitton 1975 (glued on card) (NHM),
studied.
Mesochorus (Astiphrommus) graniger Thomson, 1886: 328,
syn. nov.
Lectotype W: Ört. (LUND), locus typicus: Sweden, Pålsö,
designated by Fitton, 1982: 53, studied.
Taxonomical remark. The species is closely related
to A. dorsale (Holmgren). In females the ovipositor
sheath and femur III are slenderer (Figs 126-127),
the flagellum usually paler, and the punctation
of lower mesopleurum denser. In most cases, the
specimens are distinguishable by using a combina-
tion of these morphological characters, but there
are some intermediate specimens regarding to all
of these characters.
For the males I have not found clearly distinc-
tive features. In my opinion, it is not impossible
that both species belong to a widely variable taxon
where the variation might depend on the primary
Tachinid parasitoid or other developmental factors.
On the other hand, the dorsale group might contain
several closely related taxa not distinguishable by
morphological characters yet.
Description of W
Body length 4.5-10.0 mm. Flagellum with 34-41
flagellomeres, FEI 0.7, preapical flagellomeres about
2 × as long as wide, apical one pointed. Mandibular
teeth of equal size. Temple moderately to strongly
narrowed behind eye, TEI 0.8-0.9 (Fig. 12). Face
densely punctured and granulated, with lateral
margins convergent ventrally, its minimal width
0.9-0.95 × eye length (Fig. 32). Frons with fine granu-
lation, but shining, vertex and temple punctate. OOD
1.0. Malar space finely striate, MI 0.7. Genal carina
joining the hypostomal one far from mandibular
base, hypostomal carina not elevated.
Lower half of mesopleurum with scattered but
coarse puncture (Fig. 106). Scutellum not carinate,
more or less pointed apically (in 1 specimen from
Britain scutellum virtually rounded). Propodeum
with distinct carinae, area superomedia narrowed
apically, about 1.8 × as long as wide, apical transverse
carina usually distinct, costula in the frontal half of
area superomedia. Area petiolaris large, as long as
wide, wider than the base of tergite 1, 0.4 × as long
as propodeum. Femur III 4.6-5.6 × as long as wide.
Inner spur of tibia III 0.46 × as long as metatarsus
III. Metatarsus III about 0.45-0.5 × as long as tibia
III. Claw III with strong teeth, the third tooth much
longer than the others.
Nervulus slightly postfurcal (0.2 its length).
Nervellus in hind wing intercepted in apical 0.25,
discoidella distinct. Pterostigma slender, about 3.8-
3.9 × as long as wide.
First tergite 2.4-2.8 × and postpetiole 1.1-1.35 × as
long as wide, smooth. Second tergite 1.0-1.1 × as long
as wide, its thyridium large, transverse. Ovipositor
sheath 7.3-9.8 × as long as wide and 1.1-1.3 × as long
as metatarsus III, slightly narrowed in the apical half
(Figs 74, 126, 127).
Colour: Black. Flagellum reddish or more or
less infuscate. Cream-yellow are mandibles except
teeth, palps, clypeus and wide ventral margin of face,
malar space, hind edges of pronotum, tegula, coxae
I-II and all trochanters. Mesopleurum and apex of
scutellum diffusely reddish. Legs yellowish-red, tibia
III infuscate in the apical 0.1-0.15. Tergite 2 with a
narrow yellow apical margin. Sternites yellowish,
with brownish sides, Pterostigma pale brownish.
Ovipositor sheath black, with pale tip.
Variety: 1 specimen from Britain (EDI): meso-
soma completely black.
Description of M
Body length 5.5-8.0 mm. Flagellum with 34-39 seg-
ments, FEI 0.6-0.65, preapical flagellomere about 2 ×
as long as wide, the apical one pointed. Mandibular
teeth equally sized. Face wide, its minimal width
about 0.80-0.95 × the eye length, with more or less
parallel lateral margins. Face punctate, frons with
fine granulation. Temple punctate, TEI 0.75. OOD
about 1.0. MI 0.5. Genal carina joining the hypostomal
one far from mandibular base. Mesopleurum with
scattered to dense punctation. Scutellum not cari-
nate. Propodeum usually with complete carination.
Area petiolaris hexagonal, somewhat wider than
106
long, Apical transverse carina of area superomedia
often present. Nervulus almost interstitial. Areolet
quadrangular, receiving second recurrent vein in
its middle. Pterostigma wide, about 3.2 × as long as
wide, RI 1.1.5. Femur III 4.8-5.3 × as long as wide.
Claw III with 3-4 strong teeth. First tergite 2.3 ×, and
postpetiole 1.25 × as long as wide, smooth. Second
tergite 0.9-0.95 × as long as wide, with roundish
thyridium. Clasper about 1.2-1.3 times as long as
second tarsomere III, with knob-like thickened apex.
Colour: Black. Flagellum usually pale brownish.
Cream-yellow are mandible except teeth, clypeus,
face, gena, lower third of temple, hind edge of pro-
notum, subtegular ridge, tegula, coxae I-II and tro-
chanters I-II. Propleurum, pronotum, meso pleurum,
mesoscutum sometimes, and scutellum more or less
40 41
43
45
42
44
Figs 40-45. Face of WW from frontal: 40. A. rimosum Schwenke; 41. A. scutellatum (Gravenhorst); 42. A. simplex
(Thomson); 43. A. splenium (Curtis); 44. A. tenuicorne (Thomson); 45. A. uliginosum Schwenke.
107
reddish. Legs yellowish-red, tibia III in the apical 0.1,
and tarsus III infuscate. Metasoma blackish, tergite
2 with a narrow yellowish apical margin. Sternites
brownish, the frontal ones with yellow median
membranous areas. Pterostigma brownish.
Variety: 1 specimen from Britain (EDI): scutel-
lum not pointed.
Hosts. Amphipyra pyramidea (Linnaeus, 1758) (Lep.,
Noctuidae) (ZSM) via Tachinidae (Diptera); Alsophila
aceraria (Denis & Schiffermüller, 1775) (Lep., Geo-
metridae) via Phorocera obscura (Fallén, 1810) (Dipt.,
Tachinidae) (ZSM).
Material. 38 WW, 29 MM.
Distribution. Czechia (LINZ), Great Britain (EDI,
NHM), Ireland* (NHM), Netherlands (CZ), Poland
(WARS), Spain* (CS), Switzerland* (LAU).
46 47
49
51
48
50
Figs 46-49. Face of WW from frontal: 46-47. A. tridentatum Schwenke; 48. A. varipes (Holmgren); 49. A. nigriceps
(Brischke).
Figs 50-51. A. striatum (Brischke) W: 50. Hind metatarsus; 51. Hind trochanter.
108
Astiphromma (Astiphromma) italicum
Schwenke, 1999
Figs 5, 7, 33, 66, 77
Astiphromma italicum Schwenke, 1999: 22, WM.
Holotype W: Pizzighettone/I, 12.5.73, F. Frilli (AEI).
Holotype not seen, paratypes (ZSM) studied.
Astiphromma contum Schwenke, 1999: 24, syn. nov.
Holotype W: Ovindoli/I, Dint. 23.6.88, A. Scaramozzino
(Scaramozzino). Holotype not seen, paratypes (ZSM)
studied.
Taxonomical remark. From the Collection Au-
bert in Lausanne I received a female which is
labelled as: Paratype, Astiphromma tenuicaudatus W
Sch, Astiphromma tenuicaudatum n. sp. W Schwenke,
1972 PARATYPUS, J. F. Aubert 5.6.1955 Bois de
Vincennes-PARIS. The studied specimen belongs
to A. italicum. The name A. tenuicaudatus has never
been published by Schwenke, and I do not know if
there is a holotype.
Description of W
Body length 5-6 mm. Flagellum with 31-35 flag-
ellomeres, FEI 0.55-0.57, preapical flagellomeres
about 1.3-1.5 × as long as wide, apical one blunt or
pointed. Lower mandibular tooth slightly larger than
the upper one (Fig. 66). Temple strongly narrowed
behind eye, TEI 0.55-0.65 (Fig. 7). Face very wide,
lateral margins parallel or convergent ventrally, its
minimal width 0.90-1.0 × eye length (Fig. 33). Frons
and vertex finely granulate, but shining, temple
punctate. OOD 1.2-1.6. Malar space finely striate,
MI 0.5. Genal carina joining the hypostomal one at
or very close to mandibular base (Fig. 5), hypostomal
carina slightly elevated.
Lower half of mesopleurum with fine and very
scattered punctures. Scutellum usually with short
lateral carina in the basal 0.3-0.5. Carinae of propo-
deum weak, apical half of area superomedia often
with faint lateral carinae. Area superomedia about 2 ×
as long as wide. Costula often faint. Area petiolaris
wider than long, 0.3 × as long as propodeum. Femur
III stout, 3.3-4.0 × as long as wide. Inner spur of tibia
III 0.55-0.60 × as long as metatarsus III. Metatarsus
III 6.0-6.5 × as long as wide and 0.42-0.5 × as long
as tibia III. Claw III with short basal teeth.
Nervulus strongly postfurcal (0.5-0.7 × its
length). Nervellus in hind wing intercepted in api-
cal 0.2-0.3. second recurrent vein joining areolet in its
middle. Pterostigma slender, about 3.5-3.8 × as long
as wide, RI 1.1-1.2, radius behind areolet straight or
slightly sinusoid.
First tergite 2.7-3.5 × and postpetiole 1.3-1.7 ×
as long as wide, the latter sometimes with a median
longitudinal rim. Second tergite 0.85-0.9 × (in one
specimen 1.4 × – due to preparation?) as long as wide,
its thyridium large, longer than wide or roundish.
Ovipositor sheath 6.4-7.6 × as long as wide and 0.95-
1.0 × as long as metatarsus III, stab-shaped, slightly
narrowed apically (Fig. 77).
Colour: Black. Flagellum infuscate. Narrow
ventral margin of clypeus and face and malar space
mainly reddish-yellow. Tegula and small spot at
hind edge of pronotum yellowish. Metasoma black,
reddish-yellow are narrow hind margin of tergite
2, sandclock-shaped median stripe of tergite 3, and
small basal spot on tergite 4. Sternites yellow, sides
brown. Pterostigma brownish. Legs reddish-yellow,
femur III more reddish, at apex slightly infuscate
dorsally. Tibia III infuscate in the apical 0.2-0.25,
tarsus III more or less infuscate. Ovipositor sheath
black, with pale tip.
Description of M
Body length 5-6 mm. Flagellum with 30-34 seg-
ments. Temple slightly narrowed behind eye, TEI
0.5-0.55. OOD 1.2-1.3. Lateral margins of face con-
vergent ventrally. Scutellum with lateral carina in
the basal 0.3-0.5. Lateral carinae of area superomedia
slightly narrowed apically, costula lacking. Femur
III 3.8-3.9 × as long as wide. Postpetiole often with
median rim. Clasper about 1.0-1.2 × as long as the
second tarsomere III.
Colour: Black. Flagellum infuscate. Clypeus,
face, malar space, lower 0.1-0.2 of temple, scapus
and pedicellus ventrally, tegula, subtegular ridge,
and hind edge of pronotum yellowish. Often pro-
pleurum and usually pronotum reddish-brown or
reddish-yellow. Coxae I-II and all trochanters and
trochantelli cream-yellow. Legs otherwise reddish-
yellow, apical 0.15-0.2 of tibia III, and tarsus III
infuscate. Metasoma black. Yellowish-red are narrow
median stripe or triangular apical spot of tergite 2,
sandclock-shaped median stripe of tergite 3, and
basal third of tergite 4. Sternites 2-4 yellow, the
following ones blackish. Pterostigma brown.
Hosts. Unknown.
Material. 13 WW, 15 MM.
Distribution. Austria* (LINZ), Bulgaria* (ZSM),
Czechia* (LINZ), France* (CJ, LAU), Germany*
(CR, ZSM), Great Britain* (EDI, NHM), Italy (ZSM),
Netherlands* (CZ).
109
Astiphromma (Astiphromma) leucogrammum
(Holmgren, 1860)
Figs 13, 34, 78, 97, 108, 125
Mesochorus leucogrammus Holmgren, 1860: 121, WM.
Lectotype W: Upl, Hfgn (STOCK), studied.
Description of W
Body length 5.5-6.0 mm. Flagellum slender, with
29-32 flagellomeres, FEI 0.75-0.9, preapical seg-
ments about 2 × as long as wide, apical flagellomere
pointed. Mandibular teeth equally sized. Face wide,
punctate, matt, lateral margins divergent ventrally
(Fig. 34). Frons granulate, matt laterally. Temple
slightly narrowed behind eye, TEI 0.8-1.0, without
distinct punctures (Fig. 13). OOD 1.5-1.7. MI 0.7.
Genal carina joining the hypostomal one far from
mandibular base (distance as malar length), hypos-
tomal carina not elevated.
Lower part of mesopleurum almost smooth.
Scutellum not carinate. Area superomedia with al-
most parallel lateral carinae, about 1.8-2 × as long as
wide. Area basalis and costula usually faint or lack-
ing. Area petiolaris small, distinctly wider than long,
0.2-0.25 × as long as propodeum. Femur III 4.8-5.8 ×
as long as wide. Inner spur of tibia III about 0.42 ×
as long as metatarsus III. Metatarsus III about 0.52 ×
as long as tibia III. Claw III with small basal teeth.
Nervulus interstitial. Nervellus intercepted in the
apical 0.2-0.3, discoidella indistinct. second recurrent
vein joining the areolet in its middle. Pterostigma
about 3.3-3.8 × as long as wide. RI 1.0-1.1.
First tergite 2.5-2.7 × and postpetiole 1.3-1.6 × as
long as wide. Postpetiole with distinct longitudinal
striae or rugae at least in the apical half (Fig. 125).
Second tergite as long as wide, its thyridium round-
ish, large. Ovipositor slender, bent upwards, 8.7-10 ×
as long as wide and 0.7-0.8 × as long as metatarsus
III (Fig. 78).
Colour: Black. Cream-yellow are lower 0.5 of
temple, face and clypeus, outer side of antenna,
mandible except teeth, coxae and trochanters I-II,
propleurum and pronotum laterally, and mesos-
ternum. Mesopleurum reddish ventrally and black
dorsally, with a sharp border between both colours
(Fig. 108). Legs yellowish-red, Apex of coxa III
with infuscate dorsal spot. Tibiae often with nar-
row infuscate base. Tibia III and sometimes femur
III with a more or less clear brownish stripe on the
outer sides. Tergites 1-2 black, tergite 2 with a small
triangular yellowish spot at apex, tergites 3-4 with
yellowish median stripe, brown laterally. Following
tergites brown with yellowish hind margins (vespoid
colour pattern), their epipleura and sternites yellow.
Ovipositor sheath black, with pale tip.
Variety: One British W (EDI) has a completely
blackish metasoma with ivory apical margins of
tergites 2-7.
Description of M
Body length 6 mm. Flagellum with 30 segments,
preapical flagellomeres about 1.5 times as long as
wide. Face wide, lateral margins more or less paral-
lel. MI about 0.5. OOD 1.4. Pterostigma wide, about
3.2 × as long as wide. Femur III about 4.6 × as long
as wide. Claw III with short basal teeth. Clasper
about 1.0-1.2 × as long as second tarsomere III, with
thickened apex (Fig. 97).
Colour: Black. Flagellum reddish basally, infus-
cate apically. Yellowish are mandible except teeth,
clypeus, face, gena, lower half of temple, propleu-
rum, anterior margin and hind edge of pronotum,
tegula, subtegular ridge, mesosternum and lower
half of mesopleurum, metapleurum ventrally. Ter-
gite 2 with a narrow apical yellow margin, tergites
3-4 with central yellow stripe, tergites 5-7 blackish,
sometimes with ivory apical margins. Tibia III with
narrow dark base. Pterostigma brown. Otherwise
like the female.
Hosts. Eupithecia sp. (Lep., Geometridae) via vari-
ous Braconidae (Hym.) and Ichneumonidae (Hym.)
(Schwenke, 1999).
Material. 9 WW, 3 MM.
Distribution. Austria (NHM), France (NHM), Ger-
many (ZSM), Sweden (STOCK).
Astiphromma (Dolichochorus) longiceps
(Strobl, 1904)
Figs 35, 79, 113, 121
Dolichochorus longiceps Strobl, 1904: 108, W. Types lost.
Description of W
Body length 5-6 mm. Flagellum with 29 flagellomer-
es, long and slender, FEI 1.0-1.1, preapical segments
2 × as long as wide, the apical flagellomere pointed.
Mandibular teeth equally sized. Face granulate,
without distinct punctures, its minimal width 0.9 ×
eye length (Fig. 35). Gena, frons and temple shining
and with almost smooth surface. TEI 1.0. OOD 1.6.
MI 1.2. Genal carina joining the hypostomal one far
from mandibular base.
Mesopleurum almost smooth ventrally. Scutel-
lum not carinate. Area superomedia wide basally
and strongly narrowed apically, about 1.8 × as long
as wide. Costula and area basalis faint. Area peti-
olaris about 0.25 × as long as propodeum. Coxa III
granulate, matt. Femur III 5.6 × as long as wide.
110
Inner spur of tibia III 0.4 × as long as metatarsus
III, metatarsus 0.54 × as long as tibia III. Claw III
without distinct teeth.
Nervulus of front wing interstitial, nervellus
of hind wing usually not intercepted, sometimes
slightly intercepted in the apical 0.2, but without
discoidella (Fig. 113). second recurrent vein joining
areolet basal to its middle. Pterostigma about 3.5 ×
as long as wide.
First tergite 2.2 × and postpetiole 1.3 × as long
as wide, the latter with strong longitudinal stria-
tion (Fig. 121). Second tergite as long as wide, its
thyridium small, strongly transverse. Ovipositor
sheath about 8-8.6 × as long as wide and 0.67-0.73 ×
Figs 52-57. Face of MM from frontal: 52. A. albitarse (Brischke); 53. A. anale (Holmgren); 54. A. buccatum (Thomson);
55. A. nigrocoxatum (Strobl); 56. A. simplex (Thomson); 57. A. splenium (Curtis).
52 53
55
57
54
56
111
as long as metatarsus III, slightly bent towards
dorsal (Fig. 79).
Colour: Black. Cream-yellow are palps, man-
dibles except teeth, clypeus and face except a
large w-shaped median spot (Fig. 35), malar space,
temple in the lower 0.3, and coxae and trochanters
I-II. Flagellum brownish, yellowish below. Legs
yellowish-red, coxa III more or less infuscate, femur
III brownish-yellow dorsally, tibia III infuscate on
outer side and apical 0.2-0.25, tarsus III brownish.
Pterostigma brownish, with paler base. Tergite 2
with a yellow band in the apical 0.25, tergites 3-4
with median sandglass-shaped yellow stripes and
with brownish sides, following tergites brownish.
Sternites and hypopygium yellow, with brownish
sides. Ovipositor sheath black, reddish in the apical
0.2.
M unknown.
Hosts. Unknown.
Material. 7 WW.
Distribution. Austria (ZSM), Germany (ZSM), Great
Britain (EDI), Sweden (STOCK)
Figs 58-62. Hind claws of WW: 58. A. hirsutum (Bridgman); 59. A. diversum Schwenke; 60. A. tenuicorne (Thomson);
61. A. dorsale (Holmgren); 62. A. nigriceps nigriceps (Brischke).
Figs 63-64. 63. Scutellum of A. albitarse (Brische) W, Arrow shows lateral carina; 64. Mesoscutum of A. buccatum
(Thomson) M.
Figs 65-66. 65. Apical flagellomeres of A. flagellator spec. nov.; 66. Mandible of A. italicum Schwenke W, Arrow shows
larger lower tooth.
58 59 61 62
6463
60
65 66
112
Astiphromma (Astiphromma) luridum
Schwenke, 1999
Figs 36, 80, 98
Astiphromma luridum Schwenke, 1999: 18, WM.
Holotype W: Wolfsegg a.H./Oberösterr., 15.5.87, M.
Schwarz (ZSM), holotype and paratypes studied.
Taxonomical remark. In some MM of A. luridum
Schwenke, the characteristic striation on postpetiole
is superficial and indistinct. These specimens resem-
ble MM of A. tenuicorne (Thomson), but the latter ones
have a reddish colour on lower mesopleurum, partly
blackish sides of pronotum, and large transverse
thyridia on their second tergites.
Description of W
Body length 5-6 mm. Flagellum stout, filiform, with
36-37 flagellomeres, FEI 0.8, preapical flagellomeres
about 1.7 × as long as wide. Mandibular teeth of the
equal size. Face with superficial puncture, granulate
and matt, its minimal width 1.0 × eye length. Lateral
margins of face parallel ventrally (Fig. 36). Temple
narrowed behind eye, TEI 0.8-0.9. OOD 1.5. Genal
carina joining the hypostomal one far from mandibu-
lar base. Hypostomal carina not elevated.
Lower part of mesopleurum with dense but
fine punctation. Scutellum not carinate. Propodeum
with almost complete carination, area superomedia
slender, 2.5-3 × as long as wide, confluent with area
basalis, costula in its basal third. Area petiolaris short,
wider than long, about 0.2 × as long as propodeum.
Femur III 4.8-5.0 × as long as wide. Metatarsus III
about 0.5 × as long as tibia III. Inner spur of tibia III
0.5 × as long as metatarsus III. Claw III with small
basal teeth.
Nervulus interstitial or slightly postfurcal. Ner-
vellus intercepted in the lower 0.2. Pterostigma about
3.8 × as long as wide.
First tergite 2.2-2.3 × and postpetiole 1.12-1.25 ×
as long as wide, the latter with fine longitudinal
striation or rugulo-striation. Second tergite about
0.95 × as long as wide, with indistinct thyridium at
frontal margin. Ovipositor sheath short, 5.5-6.0 × as
long as wide, and 0.66-0.7 × as long as metatarsus
III, slightly bent upwards (Fig. 80).
Colour: Black. Yellowish are clypeus, malar
space, mandibles except teeth, lower 0.2 of temple,
lower 0.2 of facial orbit (Fig. 36), subtegular ridge,
hind edge of pronotum, tegula, small spot on lower
mesopleurum, coxae and trochanters I-II, apical mar-
gin of coxa III, sternites, apical transverse stripes of
tergites 2-7. Legs reddish, tibia III slightly infuscate
in the apical third, tarsus III brownish. Coxa III with
infuscate stripe dorsally. Pterostigma pale brownish.
Ovipositor sheath reddish.
Description of M
Body length 5-6 mm. Flagellum with 35-38 flagel-
lomeres, FEI 0.8. preapical segments about 2 × as long
as wide. Apical margin of clypeus rounded, clypeus
not protruding. Lateral margins of face parallel. Head
granulate, face matt, with superficial punctation.
TEI 0.9-1.1. OOD 1.3-1.4. Mesopleurum matt, with
dense but superficial puncture. Area superomedia
wide, its sides slightly narrowed apically. Femur III
5.3-5.4 × as long as wide. Pterostigma wide, about
3.1-3.2 × as long as wide. RI 0.9. First tergite 2.0 ×
and postpetiole 1.25 × as long as wide. Striation of
postpetiole sometimes indistinct. Second tergite 0.9 ×
as long as wide, without distinct thyridium. Clasper
about 0.8 × as long as second tarsomere III, slightly
clubbed (Fig. 98).
Colour: Black. Cream-yellowish are mandible
except teeth, palps, clypeus, face, lower frontal orbit,
temple in the lower 0.25-0.4, scape, pedicellus and
first flagellomere ventrally, propleurum, pronotum
(sometimes only hind edge and anterior margin),
often ventral half of mesopleurum, mesosternum,
subtegular ridge, tegula, sometimes anterolateral
markings on mesoscutum, coxae, trochanters and
trochantelli I-II, often apical margin of coxa III,
trochantellus III, apical transverse bands and epi-
pleura of tergites 3-7. Tergite 2 with reddish trian-
gular spot in the apical 0.2, tergite 3 with a reddish
median stripe. Sternites reddish, with brownish
lateral spots. Coxa III, and often trochanter III with
infuscate dorsal stripes. Legs otherwise reddish, tibia
III in the apical 0.2, and tarsus III (sometimes also
tarsus II) more or less diffusely infuscate.
Hosts. Unknown.
Material. 4 WW, 6 MM (ZSM).
Distribution. Austria (LINZ), Germany (CR, ZSM),
Norway* (NHM).
Astiphromma (Mesochorella) nigriceps nigriceps
(Brischke, 1880)
Figs 17, 49, 62, 82, 111, 117
Mesochorus nigriceps Brischke, 1880: 192, M, types lost.
Taxonomical remark. Schwenke (1999, 2000) stated
that the dorsolateral carina of first tergite is lacking
in Mesochorella. However, in the examined material
the carina is faint in larger parts, but usually vis-
ible at least behind the spiracle and/or at the apical
margin (Fig. 3). A. nigriceps (Brischke) is the only
Astiphromma species parasitizing Coleoptera, but the
primary parasitoid remains unknown.
113
Description of W
Mesosoma coarsely and densely punctate. Scutellum
not carinate. Area basalis pointed apically, Area
superomedia pentagonal, short, about 1.7 × as long
as wide. Area petiolaris very large, hexagonal, about
0.4 × as long as propodeum. Femur III 3.9 × as long
as wide. Inner spur of tibia III very short, 0.25-0.3 ×
as long as metatarsus III, metatarsus III about 0.4 ×
as long as tibia III. Claw III strongly curved apically,
with long blackish broadened teeth (Fig. 62).
Nervulus almost interstitial. Nervellus in hind
wing intercepted at apical 0.33. Pterostigma very
wide, about 2.5 × as long as wide, RI 1.15, second
recurrent vein usually joining the areolet basal to
its middle (Fig. 111).
First tergite 2.6 × and postpetiole 1.3 × as long as
wide, the latter with weak and partly obsolete dorso-
lateral carina, a strong longitudinal rim centrally and
some fine striae apically. Second tergite about 0.8 ×
as long as wide, its thyridium roundish. Ovipositor
sheath 8.1 × as long as wide, about 1.5 × as long as
metatarsus III, slightly lanceolate, widened beyond
middle and narrowed apically (Fig. 82).
Colour: Black. Flagellum reddish basally, in-
fuscate apically. Legs except coxae and trochanters
reddish-yellow. Pterostigma yellowish. Tergite 1
with a triangular reddish spot apically, following
tergites reddish. Tergite 2 with large, and tergite
3 with small brown spots basolaterally. Sternites
reddish, sometimes with lateral brownish spots.
Ovipositor sheath black.
Description of M
Body length 5 mm. Flagellum with 30-31 flagel-
lomeres, FEI 0.65, preapical segments about 1.5 × as
long as wide. Head with coarse and dense puncture.
Temples parallel behind eyes, TEI 1.0. OOD 1.4-1.6.
Mesosoma coarsely and densely punctured. Scutel-
lum not carinate. Area petiolaris very large, hexago-
nal. Nervulus almost interstitial or slightly postfurcal
(0.2 its length). Nervellus in hind wing intercepted at
apical 0.4. Pterostigma very wide, 2.8-3.3 × as long
as wide. Femur III 4.2 × as long as wide. Postpetiole
with weak and partly obsolete dorsolateral carina,
a strong longitudinal rim centrally and some fine
striae apically. Clasper about as long as the second
tarsomere III and slightly bent upwards. Claw III
strongly curved at apex, with long blackish teeth.
Colour: Black. Flagellum brown, apically black-
ish. Face black. Mandibles except teeth and tegula
yellow. Pterostigma yellowish, paler centrally.
All coxae black, narrowly reddish apically. Legs
reddish-brown, tarsi brownish. Tergite 2 with
triangular reddish stripe apically, tergites 3-7 red-
dish, side of tergite 3 with brown spots, sternites
yellow-red, sternites 2-4 with brown lateral spots
(Fig.117).
Hosts. Colaphellus sophiae (Schaller, 1783) (Col.,
Chrysomelidae) (Schwenke 2000).
Material. 12 WW, 8 MM.
Distribution. Bulgaria (LINZ, ZSM), Greece* (ZSM),
Romania (LINZ), Serbia* (ZSM), Slovakia (LINZ),
Turkey* (ZSM).
Astiphromma (Mesochorella) nigriceps asiaticus
ssp. nov.
Fig. 116
Holotypus W: Usbekistan 80 km SO Karschi 31.4.91 Ha-
lada (LINZ)
Paratypes W: Usbekistan 80 km SO Karschi 31.4.91 Hala-
da (KR), W: Kasachstan mer. Chagir 42,4N 68,8E 40 km
S Aris leg. J. Halada 6.5.94 (LINZ), W: Kasachstan
mer. 10 km E Ddjambal 31.5.94 leg. J. Halada (LINZ);
W: Cruciferae, Dagestan Novo Biryuzyak, 20.v.1961
B. A. Vorobyov (NHM); W: On Cruciferae, Dagestan
Novo Biryuzyak, 24.iv.1961 B. A. Vorobyov (NHM)
Taxonomical remark. This subspecies from the
Caucasus and Central Asia differs from the Euro-
pean subspecies by a lower number of flagellomeres
(25-26) and different coloration of the metasoma: all
tergites blackish, with narrow yellowish or reddish
apical margins. Femur III 3.9 × as long as wide,
ovipositor sheath 8.0 × as long as wide and 1.5 × as
long as metatarsus III. Structure and colour pattern
otherwise as in A. nigriceps nigriceps (Brischke).
Description of W
Body length 5-5.5 mm. Flagellum with 25-26 flag-
ellomeres, FEI 0.5-0.55, preapical segments about
1.5 × as long as wide, apical flagellomere pointed.
Head with coarse, dense punctures, face with rugose
punctation, matt, but clypeus apically almost smooth
and shining. Lower mandibular tooth much larger
than the upper one (Fig. 49). Temple about as wide
as the eye and seen from dorsal slightly widened
behind eyes, punctate and shining, TEI 1.0-1.1.
Ocelli very small, OOD 2.0. MI 0.3. Genal carina
joining the hypostomal one far from mandibular
base, hypostomal carina not elevated.
Mesosoma coarsely and densely punctured.
Scutellum not carinate. Area basalis pointed api-
cally. Area superomedia short, about 1.5-1.7 × as
long as wide. Area petiolaris very large, hexagonal,
0.4-0.45 × as long as propodeum. Femur III 3.9 × as
long as wide. Inner spur of tibia III very short, 0.25-
0.3 × as long as metatarsus III, metatarsus III about
0.35-0.4 × as long as tibia III. Claw III strongly curved
114
apically, with long blackish broadened teeth (Fig. 62).
Nervulus slightly postfurcal (0.2 its length).
Nervellus in hind wing intercepted at apical 0.3.
Pterostigma wide, about 2.9 × as long as wide, RI 1.1,
second recurrent vein joining areolet in its middle.
First tergite 2.2 × and postpetiole 1.1-1.2 × as long
as wide, the latter with weak and partly obsolete
dorsolateral carina, and some fine striae apically.
Second tergite about 0.75 × as long as wide, its thy-
ridium small, roundish. Ovipositor sheath about 8.0 ×
as long as wide, about 1.5 × as long as metatarsus
III, slightly lanceolate, widened beyond middle and
narrowed apically.
Colour: Black. Flagellum reddish basally, in-
fuscate apically. Palps and mandible except teeth
yellowish. Legs except coxae and trochanters red-
dish-yellow. Pterostigma yellowish. Tergite 1 black,
following tergites with a more or less distinct yellow-
ish apical margin (vespoid colour pattern). Sternites
blackish. Ovipositor sheath black.
M unknown.
Material. 6 WW.
Distribution. Kasachstan* (LINZ), Uzbekistan*
(LINZ), Russia (Dagestan)* (NHM)
67
73
69
71
75
77
70
76
78
68
72
74
Figs 67-78. Ovipositor sheaths of WW from lateral: 67. A. aggressor (Fabricius); 68. A. albitarse (Brischke); 69. A. al-
pinum Roman; 70. A. anale (Holmgren); 71. A. buccatum (Thomson); 72. A. diversum Schwenke; 73. A. dorsale (Holm-
gren); 74. A. hirsutum (Bridgman); 75. A. flagellator spec. nov.; 76. A. flavoventrale spec. nov.; 77. A. italicum Schwen-
ke; 78. A. leucogrammum (Holmgren).
115
Astiphromma (Astiphromma) nigrocoxatum
(Strobl, 1904)
Figs 8, 38, 83, 99
Astiphromma anale var. nigrocoxatum Strobl, 1904: 103, M,
types lost.
Demophorellus mimulus Hedwig, 1955: 379, M, synonym-
ized by Horstmann (1981: 72).
Lectotype M: Gifhorn 8.5.52, K. HEDWIG (ZSM), not
studied.
Astiphromma interstitiale Constantineanu & Mustafa, 1969:
314-315, M, synonymized by Schwenke (1999: 21),
type not studied.
Description of W
Body length 5-6.5 mm. Flagellum with 25-30 flagel-
lomeres, FEI 0.6, preapical flagellomeres 1.3-1.5 × as
long as wide, apical one blunt to pointed. Mandibu-
lar teeth of equal size. Face punctate, matt, lateral
margins parallel or slightly divergent ventrally, its
79
81
83 84
82
80
86
88
92
85
87
89
91
90
Figs 79-92. Ovipositor sheaths of WW from lateral: 79. A. longiceps (Strobl); 80. A. luridum Schwenke; 81. A. striatum
(Brischke); 82. A. nigriceps (Brischke); 83. A. nigrocoxatum (Strobl); 84. A. pictum (Brischke); 85. A. rimosum Schwen-
ke; 86. A. scutellatum (Gravenhorst); 87. A. simplex (Thomson); 88. A. splenium (Curtis); 89. A. tenuicorne (Thom-
son); 90. A. tridentatum Schwenke; 91. A. uliginosum Schwenke; 92. A. varipes (Holmgren).
116
minimal width 1.0-1.1 × eye length. Temple strongly
narrowed behind eye, granulate and with superficial
punctures, TEI 1.0-1.1 (Fig. 8). Vertex finely granu-
late, matt. OOD 1.7-2.0. MI 0.7. Genal carina joining
the hypostomal one distant from mandibular base (by
half malar length), hypostomal carina not elevated.
Mesoscutum with superficial punctation, granu-
late and matt. Lower part of mesopleurum with
moderately dense and coarse punctures. Scutellum
not carinate. Propodeum with complete carination.
Area superomedia short, about 1.5 × as long as wide.
Area petiolaris large, hexagonal, about 0.4 × as long
as propodeum. Femur III 4.2-4.3 × as long as wide.
Inner spur of tibia III 0.5 × as long as metatarsus III.
Metatarsus III 7-8 × as long as wide and 0.4 × as long
as tibia III, claw III with short basal teeth.
Nervulus usually interstitial, sometimes slightly
ante- or postfurcal. Nervellus of hind wing inter-
cepted in the apical 0.25-0.3. Pterostigma wide,
3.1-3.2 × as long as wide, RI 1.1, radius behind areolet
straight or slightly bent to anterior wing margin.
second recurrent vein joining areolet in its middle.
First tergite 2.5 × and postpetiole about 1.5 ×
as long as wide, the latter sometimes with median
longitudinal impression and/or fine striae api-
cally. Second tergite 0.8-0.85 × as long as wide, its
thyridium small, roundish. Ovipositor sheath 8.5 ×
as long as wide, about 1.2 × as long as metatarsus
III, slightly narrowed in the apical 0.3 and bent
upwards (Fig. 83).
Colour: Black. Ventral margin of face and cly-
peus black (Fig. 38). Pedicellus and basal flagel-
lomeres more or less reddish, flagellum apically or
completely infuscate. Hind edge of pronotum, and
tegula yellow. Coxae I-II brownish, apically yellow-
ish. Coxa III and trochanter III brown to black. Legs
otherwise yellow, femur III reddish-brown to brown,
often narrowly yellow at base and apex. Tarsus
III infuscate. Tergite 2 with yellow apical margin,
and tergite 3 with a sandclock-shaped reddish or
yellowish stripe, sometimes more or less black, fol-
lowing tergites blackish, apical membranous parts
more or less yellowish. Sternites yellow medially
and apically, brown laterally. Pterostigma yellow-
ish with darker margins. Ovipositor sheath black,
with pale tip.
Description of M
Body length 4.5-6.5 mm. Flagellum with 27-29 flag-
ellomeres, preapical segments about 1.5 × as long as
wide. TEI 1.0-1.1. OOD 1.7-2.0. Lateral margins of
face slightly divergent. Lower part of mesopleurum
with fine scattered punctation or almost smooth.
Pterostigma wide, about 3 × as long as wide. Femur
III 4.4-4.5 × as long as wide. Claw III with basal teeth.
Clasper 1.0-1.2 × as long as the second tarsomere III,
with fine longitudinal striation on outer side, inner
side smooth and shining (Fig. 99).
Colour: Black. Yellowish are palps, mandibles,
clypeus, malar space and face. Clypeal foveae and
short median longitudinal stripe of face between
antennal scrobes brown to black. Coxae more or less
brown in the basal parts, apically yellowish. Legs
yellowish-brown, all trochanters and trochantelli
yellow. Femur III brown, tibia III often infuscate
on outer side and/or at apex, tarsus III usually
brownish. Tergite 2 yellowish in the apical 0.2, ter-
gite 3 yellowish, with brown sides, or blackish with
sandclock-shaped yellowish stripe. Sternites brown
or black, sternites 2-3(-4) with yellow median stripe.
Pterostigma yellowish-brown.
Hosts. Unknown.
Material. 31 WW, 66 MM.
Distribution. Austria (CS), Bulgaria* (ZSM),
Czechia* (LINZ), France* (CZ, NHM, ZSM), Ger-
many (STOCK, ZSM), Great Britain (NHM), Swit-
zerland (CS, LAU, NHM), Turkey* (LINZ).
Astiphromma peltolatum Schwenke, 2004
Taxonomical remark. Schwenke (2004: 83) stated in
his original description that the holotype is deposited
in the Horniman Museum/London. According to the
personal communication of the curator, no material
of the taxon has been found there. The Finnish mate-
rial determined as A. peltolatum by Jussila (2010: 6),
which I studied, belongs to A. alpinum Roman.
Therefore, no material of this taxon was available
for this revision, and the taxon is unknown to me.
Description of W
(translation of Schwenke, 2004: 83)
Body length 7.0 mm. Facial margins divergent ven-
trally, TEI about 1.0. Ocelli very small, OOD about
2.0. Ovipositor sheath shorter than the metatarsus
III, apically slightly narrowed and bent to dorsal.
Colour: Clypeus red, face black. Pterostigma
dark brownish-grey. Meso- and metasoma black,
wing veins blackish, legs yellow, tergites with nar-
row yellow hind margins.
M unknown.
Distribution. Finland.
117
Astiphromma (Astiphromma) pictum
(Brischke, 1880)
Figs 39, 84, 122
Mesochorus pictus Brischke, 1880: 186, M, types lost.
Mesochorus (Astiphrommus) incidens Thomson, 1886: 331,
WM, locus typicus England, synonymized by Schmie-
deknecht (1910: 1952), types lost (Fitton, 1982: 53).
Description of W
Body length 5.2-8.0 mm. Flagellum stout, filiform,
with 31-35 flagellomeres, FEI 0.75, preapical flag-
ellomeres 1.1-1.5 × as long as wide, apical one
blunt. Head including frons with coarse and dense
punctures and smooth intervals (Fig. 39). Lower
mandibular tooth larger than the upper one. Temples
parallel behind the eyes, TEI 1.0. Minimal width of
face 0.9-0.95 × eye length, lateral margins divergent
ventrally. OOD 1.3. MI 0.4-0.5. Genal carina joining
the hypostomal one far from mandibular base, hy-
postomal carina strongly elevated (as the diameter
of mandibular palpomeres).
Notaulus strongly impressed. Ventral part of
mesopleurum with dense coarse punctation. Scutel-
lum not carinate. Propodeum with complete carina-
tion. Are superomedia slender, about 2.3 × as long
as wide, its sides strongly narrowed apically. Area
petiolaris short, wider than long, about 0.25 × as
long as propodeum. Femur III 4.5 × as long as wide.
Metatarsus III about 0.45 × as long as tibia III. In-
ner spur of tibia III 0.43 × as long as metatarsus III.
Metatarsus III with apical rings of short bristles (as
in A. striatum). Claw III with distinct teeth.
Nervulus interstitial or slightly antefurcal.
Nervellus of hind wing intercepted in the apical
0.2. Pterostigma about 3.3-3.5 × as long as wide. RI
1.1, radius behind areolet straight. second recurrent
vein joining areolet apical of its middle.
First tergite 2.4 ×, and postpetiole 1.1 × as long as
wide, the latter with more or less strong longitudinal
striation (Fig. 122). Second tergite about as long as
wide, without distinct thyridium. Ovipositor sheath
short, 5.4 × as long as wide, about 0.75 × as long as
metatarsus III, slightly bent upwards (Fig. 84).
Colour: Black. Cream-yellow are sides of (or
rarely entire) clypeus, malar space, mandible ex-
cept teeth (Fig. 39), pedicellus and basal flagellar
segments on outer side, lateral spot on the anterior
margin and upper margin of pronotum, tegula, sub-
tegular ridge, spot on mesosternum, coxae I-II and
all trochanters, transverse stripes on hind margins
of tergites 3-7, epipleura and sternites. Sternites 2-3
with brownish spots laterally. Legs reddish-yellow,
tibia III more or less darkened in the apical 0.2, tarsus
III infuscate, with narrow yellow bases of segments.
Coxa III with a dorsal blackish stripe. Pronotum in
part, mesopleurum mainly and mesoscutum entirely
reddish. Mesoscutum with two more or less distinct
yellowish stripes. Scutellum yellow, with a diffuse
reddish spot on anterior margin. Pterostigma yel-
lowish. Ovipositor sheath black.
Variety 1: One specimen from Finland (CV) has
the mesosternum, anterior margin and hind edge
of pronotum, spot on subtegular ridge, tegula, and
facial orbits widely cream-yellow. Mesosoma other-
wise blackish, hind leg reddish-brown.
Description of M
Body length 5-7 mm. Hypostomal carina elevated.
Malar index 0.6. Pterostigma 3.2 × as long as wide.
Propodeum rugulose-punctate. Clasper about 0.8 ×
as long as second tarsomere III.
Colour: Black. Cream-yellow are face, clypeus,
malar space, lower 0.2 of temple, mandible except
teeth, palps, scapus, pedicellus and basal flagellom-
eres ventrally, coxae, trochanters I-II, all trochantelli,
and tibia III on outer side. Pronotum yellow, blackish
on collare. Mesopleurum often tricoloured: yellow-
ish ventrally, reddish centrally, and black dorsally.
Scutellum yellow, often with reddish basal spot.
Metapleurum and propodeum blackish. Epipleura
and sternites yellow. Pterostigma pale brownish,
margins darker.
Hosts. Apethymus abdominalis (Lepeletier, 1823)
and A. serotinus (Müller, 1776) (Hym., Tenthredi-
nidae) (ZSM), Periclista sp. (Hym., Tenthredinidae)
(Schwenke 1999).
Material. 27 WW, 28 MM.
Distribution. Finland (ZSM), Germany (CR, ZSM),
Great Britain (EDI, NHM), Poland (EDI, WARS),
Sweden* (STOCK), Switzerland* (LAU).
Astiphromma (Astiphromma) rimosum
Schwenke, 1999
Figs 40, 85, 100, 103, 123
Astiphromma rimosum Schwenke, 1999: 17, WM.
Holotype W: Dessau, 5 km NE, 13.5.88, Alopecurus-Wiese,
H. Zörner (SDEI), not studied.
Description of W
Body length 5.5-6 mm. Flagellum with 30 flagellom-
eres, FEI 0.65-0.7, preapical segments about 1.5 × as
long as wide, apical flagellomere pointed or blunt.
Mandibular teeth of equal size. Temples almost
parallel behind eyes, densely punctate, TEI 1.0-1.1.
Frons more or less rugose-punctate. Face matt,
lateral margins parallel or divergent ventrally, its
minimal width 0.95-1.0 × eye length (Fig. 49). OOD
118
1.6. MI 0.3-0.4. Genal carina joining the hypostomal
one far from mandibular base, hypostomal carina
not elevated.
Lower half of mesopleurum with very dense and
coarse punctation, partly rugose (Fig. 103). Scutellum
not carinate. Propodeum with complete carination.
Area superomedia slender, about 2.3-2.5 × as long
as wide. Area petiolaris short, distinctly wider than
long, 0.25 × as long as propodeum. Femur III about
4.3 × as long as wide. Metatarsus III 7.5-8 × as long
as wide and about 0.42 × as long as tibia III, with
ring of spines at apex. Inner spur of tibia III 0.52 ×
as long as metatarsus III. Claw III with some small
basal teeth.
Nervulus interstitial or slightly postfurcal (0.2 of
its length), pterostigma 3.6-4.0 × as long as wide. RI
1.1-1.2. Nervellus of hind wing intercepted in the
apical 0.3, discoidella distinct. Second recurrent vein
joining areolet in its middle.
First tergite 2.0
× and postpetiole 1.1-1.2 × as
long as wide, the latter with longitudinal rugae
and/or striation (Fig. 123). Second tergite 0.85
× as
long as wide, without distinct thyridium. Ovipositor
sheath about 5.5 × as long as wide and 0.9 × as long
as metatarsus III, narrowed in the apical 0.2 (Fig. 85).
Colour: Black. Flagellum reddish. Yellowish
are palps and mandible except teeth, clypeus, face
except a central blackish spot (Fig. 40), gena, tegula,
hind edge of pronotum and spot on subtegular ridge.
Scapus and pedicellus brownish dorsally. Scutellum
sometimes reddish. Coxae, trochanters and trochan-
telli I-II, and apical margin of coxa III yellowish. Legs
otherwise reddish, tibia III not distinctly infuscate
at apex. Postpetiole apically and tergites 2-3(4)
94
102
100
98
96
101
99
97
93
95
Figs 93-102. Claspers of MM from lateral: 93. A. italicum Schwenke; 94. A. splenium (Curtis); 95. A. uliginosum
Schwenke; 96. A. varipes (Holmgren); 97. A. leucogrammum (Holmgren); 98. A. luridum Schwenke; 99. A. nigrocoxa-
tum (Strobl); 100. A. rimosum Schwenke; 101. A. scutellatum (Gravenhorst); 102. A. simplex (Thomson).
119
reddish, the following ones blackish, Tergites 3-7
with yellow transversal stripes apically, sternites
yellowish. Pterostigma pale brownish. Ovipositor
sheath reddish.
Description of M
Body length 6-7 mm. Flagellum with 30 flagellom-
eres, FEI 0.67. Head with superfical punctation and
fine granulation, face and frons matt, temple slightly
shining. Lateral margins of face parallel. OOD 1.8.
MI 0.4. Lower mesopleurum with coarse and dense
punctures. Claws without distinct teeth. Clasper
short, 0.7 × as long as the second segment of hind
tarsus, apically thickened (Fig. 100).
Colour: Black. Sides of pronotum and mesosternum
with small yellowish spots, flagellum reddish, infus-
cate dorsally. Trochantellus III yellow. First tergite
103
105
107 108
104
106
Figs 103-108. Mesosoma of WW from lateral: 103. A. rimosum Schwenke; 104. A. aggressor (Fabricius); 105. A. dorsa-
le (Holmgren); 106. A. hirsutum (Bridgman); 107. A. flavoventrale spec. nov.; 108. A. leucogrammum (Holmgren).
120
black, postpetiole partly reddish-brown. Tergites 2-4
reddish, with brown lateral spots and yellow hind
margins, tergites 5-7 reddish-brown, with yellow
hind margins. Otherwise like the female.
Hosts. Unknown.
Material. 7 WW, 1 M.
Distribution. Byelorussia* (ZSM), Germany (ZSM),
Norway (CR).
Astiphromma (Astiphromma) scutellatum
(Gravenhorst, 1829)
Figs 18, 41, 86, 101
Mesochorus scutellatus Gravenhorst, 1829: 973, M.
Holotype M: without label (WROCLAW), locus typicus
Scheidnich/Poland, designated by Horstmann 2003:
33, not studied.
Mesochorus festivus Holmgren, 1860: 121, WM, synony-
mized by Schmiedeknecht (1910: 1950), not studied.
Astiphromma intermedium Constantineanu & Mustata,
1976: 110-111, syn. nov. Type not studied, interpre-
tation based on the description from Constantineanu
& Mustata 1981: 24-25.
Taxonomical remark. Astiphromma intermedium has
been described and compared with A. scutellum by
Constantineanu & Mustata (1976). For separation of
both taxa, they used the carination of propodeum,
position of nervulus, and length of tergites 2-3.
However, studying a larger sample of A. scutellatum,
the structural characteristics of A. intermedium are
within the variability of A. scutellatum. I suggest that
both species are synonyms, though I was not able
to study the types.
Description of W
Body length 6-8 mm. Flagellum with 36-41 flagel-
lomeres, FEI 0.65, preapical flagellomeres 1.5-1.7 ×
as long as wide. Mandibular teeth of equal size.
Face punctate and with fine granulation, shining,
lateral margins slightly divergent below, its minimal
width 0.9-0.95 × eye length. Frons smooth, vertex
with scattered punctation, temple with fine and
denser punctures. OOD 1.3. Temple strongly nar-
rowed behind eye, TEI 0.6-0.7 (Fig. 18). MI about
0.6. Genal carina joining the hypostomal one far
from mandibular base.
Lower half of mesopleurum with fine and scat-
tered punctation. Scutellum not carinate. Area basalis
sometimes faint or obsolete. Area superomedia
centrally broad, strongly narrowed apically, 1.8-2 ×
as long as wide, confluent with area basalis. Area
petiolaris large, wider than long, 0.35 × as long as
propodeum. Femur III 4.1-4.5 × as long as wide.
Inner spur of tibia III 0.5 × as long as metatarsus III.
Metatarsus III 10 × as long as wide and about 0.5 × as
long as tibia III. Claw III with about 3-4 strong teeth.
Nervulus postfurcal (0.2-0.4 its length). Nervel-
lus of hind wing intercepted in its apical 0.25.
Pterostigma about 4.2 × as long as wide, RI 1.0-1.2,
radius behind areolet straight or slightly sinusoid.
First tergite 2.6 × and postpetiole about 1.5 × as
long as wide. Second tergite 0.9-1.0 × as long as wide,
its thyridium small, roundish. Ovipositor sheath 6.3 ×
as long as wide and 0.9 × as long as metatarsus III,
narrowed in the apical third (Fig. 86).
Colour: Black. Flagellum brownish, sometimes
reddish below. Ventral margins of clypeus and face
narrowly reddish (sometimes completely black),
malar space largely yellowish (Fig. 41). Mesoster-
num, mesopleurum, and scutellum mainly reddish.
Pterostigma yellowish. Legs reddish-yellow. Tibia III
yellow, darkened in the apical 0.2, tarsus III mainly
infuscate, sometimes metatarsus III pale basally.
Metasoma blackish, tergite 2 often with a narrow
yellow hind margin, and tergite 3 often with an indis-
tinct reddish median stripe. Ovipositor sheath black.
Description of M
Body length 6-8 mm. Flagellum with 36-38 flagel-
lomeres. Lateral facial margins more or less parallel.
Frons smooth and shining. TEI 0.7-0.8. OOD 1.3. MI
0.5. Femur III 3.9-4.3 × as long as wide. Clasper 1.3-
1.6 × as long as the second tarsomere III (Fig. 101).
Colour: Black. Clypeus, face, malar space, and
lower temple yellow. Propleurum, pronotum in
larger part, mesosternum, mesopleurum and basal
half of scutellum red. Legs reddish, coxae and tro-
chanters I-II and tibia III more yellowish, tibia III
infuscate in the apical 0.2, tarsus III except base of
metatarsus infuscate. Colour otherwise as in females.
Hosts. Sparganothis pilleriana (Denis & Schiffer-
müller, 1775) (Lep., Tortricidae) (Constantineanu
& Mustata, 1976), Diprion sp. (Hym., Diprionidae)
(Schwenke 1999).
Material. 16 WW, 14 MM.
Distribution. Austria* (CS), Bulgaria (ZSM), France
(EDI, LAU), Germany (CR, ZSM), Greece* (ZSM),
Romania (Constantineanu & Mustata, 1976), Swit-
zerland* (LAU).
121
Astiphromma (Astiphromma) simplex
(Thomson, 1886)
Figs 42, 56, 87, 102
Mesochorus (Astiphrommus) simplex Thomson, 1886: 334,
WM.
Lectotype W: Yd (LUND), locus typicus Sweden: Yddinge,
designated by Fitton, 1982: 54, studied.
Taxonomical remark. This taxon represents one
of the smallest known European Astiphromma spe-
cies and seems to be very rare. The M of A. simplex
Thomson was misidentified by Schwenke (1999).
His description and determined material runs to
A. splenium var. plagiatum (Thomson).
Description of W
Body length 4.5-5.5 mm. Flagellum with 29-30 flagel-
lomeres, FEI 0.8, preapical segments 1.7 × as long as
wide. Apical flagellomere pointed. Mandibular teeth
of the same size. Lateral margins of face parallel or
divergent ventrally. Temples moderately narrowed
behind eyes, TEI 0.8-0.9. Face with dense punctation.
Frons with granulation laterally, temple with fine
granulation, but shining, without punctures. OOD
1.9. MI 0.5. Genal carina joining the hypostomal
one far from mandibular base, hypostomal carina
not elevated.
Lower half of mesopleurum smooth. Scutellum
not carinate. Propodeum with area basalis and
superomedia confluent, with almost parallel sides
apically, costula lacking. Area petiolaris small, wider
than long and slightly narrower than the petiolar
base. Femur III 4.6-4.8 × as long as wide. Inner spur
of tibia III 0.4 × as long as metatarsus III. Claw III
with small basal teeth.
Nervulus slightly postfurcal (0.1-0.2 its length).
Pterostigma 3.7 times as long as wide.
First tergite 2.8 × as long as wide, postpetiole
smooth, with median rim and some fine striae
apically, about 1.35 × as long as wide apically. Thy-
ridium of second tergite roundish. Ovipositor sheath
about 6.1-7 × as long as wide and 0.85 × as long as
metatarsus III, slightly narrowed in the apical third
(Fig. 87).
Colour: Black. Cream-yellowish are mandibles
except teeth, palps, face, clypeus, frontal orbits
above antennal scrobes, gena and temples in the
ventral 0.4 (Fig. 42), scapus below, propleurum,
anterior margin and hind edge of pronotum, tegula,
subtegular ridge, mesosternum and lower part of
mesopleurum. Mesopleurum yellowish-red centrally
and blackish in the dorsal fifth. Legs yellowish.
Hind leg more reddish-yellow, dorsal side of coxa
III with brownish spot apically. Tibia III narrowly
darkened basally. Tarsus III brownish. Postpetiole
with a small triangular yellow spot apically, tergite
2 with yellow hind margin (0.2 of length), tergites
3-5 yellow, with brownish halfmoon-sized spots
laterally, apical tergites with yellowish apical bands.
Epipleura and sternites yellowish. Pterostigma pale
brownish. Ovipositor sheath yellowish-red.
Variety: One specimen from Austria (LINZ) with
ovipositor sheath 8.4 × longer than wide.
Description of M
Body length 6 mm. Flagellum with 30 flagellomeres,
FEI 0.7, preapical segments 2 × as long as wide. Apical
flagellomere pointed. Mandibular teeth of the same
size. Apical margin of clypeus almost straight, not
rounded. Lateral margins of face divergent ventrally
(Fig. 56). Temples moderately narrowed behind eyes,
TEI 0.8-1.0. Face with dense punctation and granula-
tion. Frons with lateral granulation and superfical
punctures. Temple finely granulate, but shining,
without punctures. OOD 1.2. MI 0.3. Genal carina
joining the hypostomal one far from mandibular
base, hypostomal carina not elevated.
Lower half of mesopleurum smooth. Scutellum
not carinate. Propodeum with area basalis and
superomedia confluent, with almost parallel sides,
costula lacking. Area petiolaris small, heart-shaped,
wider than long and slightly narrower than the
petiolar base. Femur III about 4.8 × as long as wide,
inner spur of tibia III 0.4 × as long as metatarsus
III, metatarsus III 0.5 × as long as tibia III. Claw III
without distinct teeth.
Nervulus almost interstitial. Pterostigma 3.5
times as long as wide (anterior margin somewhat
rolled ventrally). Second recurrent vein joining
areolet in its middle.
First tergite 2.3 × as long as wide, postpetiole
smooth, with median rim and some fine striae api-
cally, about 1.5 × as long as wide apically. Second
tergite about as long as wide, its thyridium oblique.
Clasper with slightly thickened apex, about 0.9 × as
long as second tarsomere III (Fig. 102).
Colour: Black. Cream-yellowish are mandibles
except teeth, palps, face, clypeus, frontal orbits above
antennal scrobes, gena and temples in the ventral
0.3, propleurum, anterior margin and hind edge of
pronotum, tegula, subtegular ridge, mesosternum
and lower part of mesopleurum. Mesopleurum
yellowish-red centrally and blackish in the dorsal
fifth. Legs yellowish. Hind leg more reddish-yellow,
femur III slightly brownish dorsally. Tibia III not
distinctly darkened apically. Tergite 2 with yellow-
ish triangular apical mark, tergite 3 with yellowish
sandclock-shaped median spot, tergite 4 with yel-
lowish basal stripe. Pterostigma pale brownish.
122
Hosts. Unknown. The hosts mentioned by Schwen-
ke (1999) might belong to A. splenium var. plagiatum
(Thomson) (see taxonomical remark).
Material. 4 WW, 2 MM.
Distribution. Austria (NHM), Finland (CJ), Sweden
(LUND, STOCK).
109
111 112
113 114
110
Figs 109-112. Front wings of WW: 109. A. aggressor (Fabricius); 110. A. buccatum (Thomson); 111. A. nigriceps nigriceps
(Brischke); 112. A. varipes (Holmgren).
Figs 113-114. Nervellus of hind wing of WW: 113. A. longiceps (Strobl); 114. A. tenuicorne (Thomson).
Astiphromma (Astiphromma) splenium
(Curtis, 1833)
Figs 21, 43, 57, 88, 94
Mesochorus splenium Curtis, 1833: 424, M.
Mesochorus sericans Curtis, 1833: plate 464, W, syn-
onymized with Astiphromma splenium (Curtis) by
Carlson (1979: 705), re-established as a different
taxon by Horstmann (2006: 1451-1452), here the
synonymy of Carlson is confirmed.
Lectotype W: M (DUBLIN), designated by Fitton (1976:
323). Type not studied.
123
115 116 117 118
119
120
Figs 115-118. Metasoma from dorsal: 115. A. dispersum Schwenke W; 116. A. nigriceps asiaticus ssp. nov. W; 117. A. nigri-
ceps nigriceps (Brischke) M; 118. A. uliginosum Schwenke W.
Figs 119-120. Metasoma of WW: 119. A. flavoventrale spec. nov.; 120. A. uliginosum Schwenke (Holotype of A. trima-
culosum).
124
Mesochorus strenuus Holmgren, 1860: 199, WM, syn-
onymized by Muesebeck (1977: 705).
Lectotype W: Bl., Bhn (STOCK), studied.
Mesochorus (Astiphrommus) plagiatus Thomson, 1886:
332, M.
Lectotype M: Hbg (LUND), locus typicus: Sweden, Eng-
land, studied and herewith designated.
Taxonomical remark. This taxon is very common in
Europe and seems to be widely variable in size and
colour pattern which had given space for the descrip-
tion of several different species in the past. Meso-
chorus sericans Curtis, 1833 was synonymized with
A. splenium (Curtis) by Carlson (1979: 705), but re-
established as a different taxon by Horstmann (2006),
mainly by different colour patterns of the metasoma.
I have seen large series of A. splenium (Curtis)
from Scotland and Byelorussia where most WW were
typical A. splenium whereas some of them showed
the colour pattern of A. sericeus. In the material
from Edinburgh, both forms have been reared from
Scirtetes robustus (Woldstedt, 1877) (Ichneumonidae)
via Orthosia gracilis (Denis & Schiffermüller, 1776)
(Noctuidae) collected from the same place. There-
fore, I confirm the synonymizing of both taxa by
Carlson (1979).
Astiphromma plagiatum Thomson was synonym-
ized with Astiphromma strenuum Holmgren (= sple-
nium Curtis) by Schwenke 1999: 20. The lectotype
which I have studied is very small, but its structure
and coloration is typical for A. splenium. The smaller
specimens of A. splenium that have been determined
as A. plagiatum were usually reared from lepidop-
teran hosts with Microgasterinae as primary para-
sitoids, whereas the larger and typical A. splenium
specimens have been reared from Lepidoptera with
Meteorus (Braconidae) and several Ichneumonidae
as primary parasitoids. I suggest that the size of the
A. splenium imago is mainly influenced by some
(nutritional?) factors related to the primary parasi-
toid in the host.
Description of W
Size and colour very variable. Body length 5-9.5 mm.
Flagellum with 31-39 flagellomeres, FEI 0.65-0.70,
preapical flagellomeres 1.5-2 × as long as wide, apical
flagellomere pointed (rarely blunt). Lateral margins
of face parallel or slightly divergent or convergent
ventrally, its minimal width 0.85-0.9 × eye length
(Fig. 43). Mandibular teeth of equal size. Frons and
vertex with fine granulation and very scattered punc-
tation laterally, Temples with punctures, strongly
narrowed behind eyes, TEI 0.5-0.55. OOD 1.1-1.5.
MI 0.5-0.6. Genal carina joining the hypostomal
one far from mandibular base, hypostomal carina
not elevated.
Lower part of mesopleurum with scattered to
moderately dense puncture. Scutellum not carinate.
Propodeum with almost complete carination. Area
superomedia confluent with area basalis, about 2.5 ×
as long as wide, lateral carinae narrowed apically,
often more or less faint. Area petiolaris about as long
as wide, wider than the petiolar base, 0.3-0.35 × as
long as propodeum. Femur III 4.3-4.6 × as long as
wide. Inner spur of tibia III 0.45-0.5 × as long as
metatarsus III, metatarsus III 0.55-0.6 × as long as
tibia III and about 10.5 × as long as wide. Claw III
with long strong teeth.
Nervulus postfurcal of 0.4-0.6 of its length.
Nervellus intercepted in the lower 0.2-0.25. Ptero-
stigma 3.8-3.9 × as long as wide, RI 1.1-1.2, radius
behind areolet straight. second recurrent vein joining
areolet in its middle.
First tergite 2.3-2.4 × and postpetiole 1.15-1.3 × as
long as wide, smooth. Second tergite as long as wide.
its thyridium large, roundish. Ovipositor sheath
6.2-6.5 × as long as wide, about 0.9-1.0 × as long as
metatarsus III, narrowed in the apical 0.4 (Fig. 88).
Colour: black. Ventral margins of face and cly-
peus, and malar space completely yellowish-red
(Fig. 43). Flagellum infuscate. Yellow are hind edge of
pronotum, subtegular ridge, tegula, front and middle
coxae and trochanters. Scutellum with a diffuse red-
dish mark. Legs reddish-yellow, femur III reddish,
apical half more or less brownish (Fig. 21), tibia III
yellowish, at narrow base and apical 0.2-0.3 infus-
cate. Tarsus III reddish-brown. Pterostigma brown.
Colour of var. sericans: black. Ventral margins
of clypeus and face reddish, malar space more yel-
lowish. Flagellum reddish. Tegula, subtegular ridge
and hind edge of pronotum yellowish. All coxae,
trochanters, and trochantelli I-II cream-yellow,
legs otherwise reddish-yellow. Femur III with weak
apical infuscation, apical 0.25 of tibia III infuscate,
tarsus III reddish. Tergite 1 black, tergite 2 with large
median triangular spot at apex, tergites 3-4 yellow-
ish medially, brown laterally, following tergites
brown basally and yellowish apically (vespoid) or
completely brownish. Sternites 2-3 yellow, following
sternites reddish-yellow. Pterostigma pale brown.
Ovipositor sheath black, with pale tip.
Description of M
Body length 5-8 mm. Flagellum with 27-37 seg-
ments, FEI 0.67, preapical flagellomeres 2 × as long as
wide. Mandibular teeth of equal size. Lateral margins
of face strongly convergent ventrally (Fig. 57), face
narrow, minimal width about 0.6-0.75 × as length of
compound eye. Temples strongly narrowed behind
eyes, TEI 0.55. Face with dense punctation, matt.
Frons and vertex granulate, temple with superficial
125
punctures. OOD 1.5. MI 0.25. Genal carina joining
the hypostomal one far from mandibular base, hy-
postomal carina not elevated.
Lower half of mesopleurum smooth. Scutellum
not carinate. Propodeum with complete carination.
Sides of area superomedia narrowed apically, cos-
tula distinct. Area petiolaris small, wider than long.
Femur III 4.7-4.9 × as long as wide, inner spur of
tibia III 0.5 × as long as metatarsus III, metatarsus
III 0.50-0.55 × as long as tibia III. Claw III with teeth.
Nervulus slightly postfurcal (0.1-0.2 its length),
nervellus of hind wing intercepted in the apical 0.2,
but discoidella indistinct. Second recurrent vein join-
ing areolet basal of its middle. Pterostigma 3.8-3.9 ×
as long as wide, RI 1.2, radius behind areolet straight.
First tergite 2.8 × and postpetiole 1.3-1.4 × as long
as wide apically. Second tergite 1.0-1.1 × as long
as wide, its thyridium large, roundish or slightly
transverse. Clasper about 1.1-1.4 × as long as the
second tarsomere III (Fig. 94).
Colour: Black. Cream-yellowish are mandibles
except teeth, palps, face, clypeus, frontal orbit above
antennal scrobe, gena and temple in the ventral 0.4,
coxa I and trochanteres I-III. Propleurum, pronotum,
mesosternum and mesopleurum reddish, often with
yellowish stripes or spots. Mesopleurum often in-
fuscate below subtegular ridge. Metapleurum and
scutellum sometimes reddish. Legs reddish-yellow.
Tibia III yellow infuscate in the apical 0.2. Tarsus
III more or less brownish. Tergite 2 with large yel-
low triangular spot in the apical half, tergite 3 with
sandclock-shaped yellowish stripe, tergite 4 with
yellowish spot basally. Pterostigma pale brownish.
Hosts. Eupithecia actaeata (Wallerdorf, 1869) (Lep.,
Geometridae) via Dusona angustifrons Förster, 1868
(Hym., Ichneumonidae) (ZSM); Panolis flammea
(Denis & Schiffermüller, 1775) (Lep., Noctuidae)
via ?Scirtetes robustus (Woldstedt, 1877) (Hym.,
Ichneumonidae) (ZSM); Campoletis rapax (Graven-
horst, 1829) (Hym., Ichneumonidae) (EDI, ZSM);
Panolis flammea (Denis & Schiffermüller, 1775) (Lep.,
Noctuidae) via Meteorus versicolor (Wesmael, 1838)
(Hym., Braconidae) (NHM); ?Aerhalnia punctu-
lata via Dusona sp. (Hy,., Ichneumonidae) (EDI);
Zygaena filipendulae or laricerae (Lep., Zygaenidae)
via Charops cantator (DeGeer, 1778) (Hym., Ichneu-
monidae) (EDI); Orthosia incerta (Hufnagel, 1766)
(Lep., Noctuidae) (ZSM); Orthosia cerasi (Fabricius,
1775) (Lep., Noctuidae) (NHM); Thera variata (Denis
& Schiffermüller, 1775) (Lep., Geometridae) via
Protapanteles pinicola (Lyle, 1917) (Hym., Braconi-
dae) (NHM); Eupithecia sp. (Lep., Geometridae) via
Campoplegine sp. (Hym., Ichneumonidae) (NHM);
Taenocampa stabilis (?) via Aleiodes circumcriptus (Nees,
1834) (Hym., Braconidae) (NHM); Meteorus pulchri-
cornis (Wesmael, 1835) (Hym., Braconidae) (NHM);
Thera variata Denis & Schiffermüller, 1775 (Lep.,
Geometridae) via Meteorus melanostictus (Capron,
1887) (Hym., Braconidae) (NHM); Macaria liturata
(Clerck, 1759) (Lep., Geometridae) via Protapanteles
fulvipes (Haliday, 1834) (Hym., Braconidae) (ZSM);
Erannis defoliaria (Clerck, 1759) (Lep., Geometridae)
via Microgaster sp. (Hym., Braconidae); Orthosia sp.
(Lep., Noctuidae) via Microgaster sp. (Hym., Braco-
nidae) (ZSM); Odontopera bidentata (Clerck, 1759)
(Lep., Geometridae) via Apanteles s.lat. sp. (Hym.,
Braconidae) (Thomson, 1886); Amphipyra pyramidea
(Linnaeus, 1758) (Lep., Noctuidae) via Apanteles s.lat.
sp. (Hym., Braconidae) (ZSM).
Material. 252 WW, 133 MM.
Distribution. Austria (LINZ, NHM), Bulgaria (LINZ,
ZSM), Byelorussia (ZSM), Czechia (LINZ), Finland
(CJ), France (LINZ, ZSM), Germany (ZSM, CR), Great
Britain (EDI, NHM), Greece* (LINZ, ZSM), Hungary
(ZSM), Ireland (NHM), Italy* (CG, NHM, WARS,
ZSM), Netherlands (CZ, NHM), Montenegro (NHM),
Norway (CR), Poland (ZSM), European Russia (CR,
LINZ)), Serbia* (NHM), Siberia (CR), Slovakia (LINZ),
Slovenia* (LINZ), Sweden (EDI, CR, NHM, STOCK),
Switzerland (CZ), Ukraine (CR), Turkey (ZSM).
Astiphromma (Astiphromma) striatum
(Brischke, 1880)
Figs 16, 37, 50, 51, 81
Mesochorus (Astiphromma) striatus Brischke, 1880: 192,
types lost.
Mesochorus (Astiphrommus) mandibularis Thomson, 1886:
330, WM, syn. nov.
Lectotype W: Yd (LUND), locus typicus Sweden, Yddinge,
designated by Townes et al. 1965: 340, studied.
Taxonomical remark. Brischke (1880) described
both W and M of A. striatum, but his type material is
lost. However, his detailed description of the colour
patterns of both sexes corresponds exactly with the
one found in A. mandibularis. Although Brischke did
not mention the differently sized mandibular teeth, I
have no doubt that both taxa are synonyms.
Description of W
Body length 5-8 mm. Flagellum with 38-41 flagel-
lomeres, FEI 0.6-0.65, preapical segments about 2 ×
as long as wide, apical flagellomere pointed. Lower
mandibular tooth much larger than the upper one.
Temple narrowed behind eye, with scattered punc-
ture, TEI 0.7-0.75 (Fig. 16). Minimal width of face
1.0 × eye length, lateral margins parallel or slightly
divergent ventrally. Frons and vertex finely granu-
126
late, with very scattered punctation, OOD 1.1-1.4. MI
0.4. Genal carina joining the hypostomal one far from
mandibular base, hypostomal carina not elevated.
Lower part of mesopleurum with scattered to
dense puncture. Scutellum sometimes with lateral
carina in the basal 0.5. Propodeum with complete
carination. Area superomedia about 2 × as long as
wide, strongly narrowed apically. Area petiolaris
wider than long, about as wide as petiolar base,
0.35 × as long as propodeum. Femur III 3,8-4,2 × as
long as wide. Trochantellus III ventrally flattened,
the flat area with more or less sharp edges laterally
and medially (Fig. 51). Metatarsus III with apical
rings of small spines, about 0.42 × as long as the tibia
III (Fig. 50). Inner spur of tibia III 0.55-0.6 × as long
as metatarsus III. Claw III with short basal teeth.
Nervulus strongly postfurcal (0.3-0.6 its length),
nervellus of hind wing intercepted in the apical 0.2.
Second recurrent vein joining areolet in or basal to
its middle. Pterostigma about 3.7 × as long as wide,
RI about 1.2, radius behind areolet straight.
First tergite 3.0-3.2 × and postpetiole 1.6-1.8 ×
as long as wide. Second tergite usually as long as
wide (rarely 1.4 × as long as wide – artificially?), its
thyridium oval or roundish, slightly as long as wide.
Ovipositor sheath narrow, 8.1-8.9 × as long as wide,
about 1.1 × as long as metatarsus III, narrowed in the
apical third (Fig. 81).
Colour: Black. Flagellum reddish or yellowish,
dorsally infuscate. Clypeus, face, malar space, lower
0.2 of temple and lower half of frontal orbit yellow,
sometimes face with brown median stripe (Fig. 37).
Tegula, hind edge of pronotum and subtegular
ridge yellow. Mesosoma often with reddish and/
or yellowish suffusion, especially on mesoscutum,
mesosternum, and scutellum. Coxae I-II and all
trochanters and trochantelli yellowish. Legs other-
wise reddish-yellow, coxa III sometimes with pos-
terodorsal brownish spot. Tibia III infuscate in the
apical 0.1. Tergite 2 at apex with large reddish-yellow
triangular median spot, sometimes with a complete
median stripe, tergite 3 with a sandclock-shaped
reddish-yellow stripe, following tergites usually
more or less vespoid, sternites yellowish with brown
sides. Pterostigma yellowish, with darker margins.
Ovipositor sheath black, with reddish tip.
Description of M
Body length 7-9 mm. Flagellum with 37-40 flagel-
lomeres. Lateral margins of face slightly conver-
gent ventrally Vertex and temple with scattered
punctures. TEI 0.9. OOD about 1.0. Lower part of
mesopleurum with scattered punctures. Femur III
3.8-4.2 × as long as wide. Metatarsus III 0.4 × as
long as tibia III. Clasper about 1.0-1.2 × as long as
the second tarsomere III. Trochantellus III ventrally
flattened with more or less sharp edges laterally and
medially. Tibia III with numerous spines on outer
side. Claw III with long basal teeth.
Colour: Black. Coxa III sometimes with a black-
ish spot dorsal-apical. Femur III with narrow infusca-
tion apically. Sometimes apical tarsomere brownish.
Sternites 5-7 blackish. Otherwise coloured as the W.
Hosts. Mamestra brassicae (Linnaeus, 1758) and
Lacanobia oleacea (Linnaeus, 1758) (Lep., Noctuidae)
(Constantineanu & Mustata, 1981), Geometridae
larva (Lep.) (Schwenke 1999).
Material. 103 WW, 57 MM.
Distribution. Austria (CS, CZ, LINZ, NHM), Bul-
garia (ZSM), Byelorussia* (ZSM), Czechia (LINZ),
Finland (CJ), France* (ZSM), Germany (CR, LAU,
NHM, SCHNEE, ZSM), Poland (WARS), Russia
(LINZ), Sweden (NHM, STOCK, ZSM), Switzerland*
(LAU), Ukraine (CR).
121 122 123 124 125
Figs 121-125. Postpetiolus of WW: 121. A. longiceps (Strobl); 122. A. pictum (Brischke); 123. A. rimosum Schwenke;
124. A. tenuicorne (Thomson); 125. A. leucogrammum (Holmgren).
127
Astiphromma (Astiphromma) tenuicorne
(Thomson, 1886)
Figs 19, 44, 89
Mesochorus (Astiphrommus) tenuicornis Thomson, 1886:
332, WM.
Lectotype W: Pål (LUND), locus typicus Sweden: Pålsö,
designated by Fitton 1982: 54, studied.
Description of W
Body length 5-7.5 mm. Flagellum very slender,
with 36-39 flagellomeres, FEI 0.8-0.9, preapical
flagellomeres more than 2 × as long as wide. Lower
mandibular tooth often slightly larger than the
upper one. Temples strongly narrowed behind
eyes, with very superficial punctures, TEI 0.8-0.9
(Fig. 19). Face punctate, matt, its lateral margins
divergent ventrally, minimal width 0.9-1.0 × eye
length (Fig. 44). Frons and vertex often granulate,
Fig. 126. Relation of the size of femur III and ovipositor sheath in A. dorsale (Holmgren) and A. hirsutum (Bridgman).
Fig. 127. Relation of the size of ovipositor sheath and the length of metatarsus III in A. dorsale (Holmgren) and
A. hirsutum (Bridgman).
6.0
5.5
5.0
4.5
4.0
3.5
ı ı ı ı ı ı
5 6 7 8 9 10
A. dorsale
A. hirsutum
A. hirsutum Paralectotype
A. granigerum Lectotype
Length/height of ovipositor sheath
Length/height of femur III
1.40
1.35
1.30
1.25
1.20
1.15
1.10
1.05
1.10
0.95
0.90
ı ı ı ı ı
5 6 7 8 9
Length/height of ovipositor sheath
Length of ovipositor sheat
h/
length of metatarsus III
A. dorsale
A. hirsutum
A. hirsutum Paralectotype
A. granigerum Lectotype
128
with fine superficial punctation. OOD about 1.2.
MI 0.6. Genal carina joining the hypostomal one far
from mandibular base (by malar length).
Lower part of mesopleurum almost smooth or
with fine scattered punctation. Scutellum not cari-
nate. Propodeum usually with complete carination,
but areae basalis and supermedia often confluent.
Costula weak. Area superomedia very slender, 3.3-
3.7 × as long as wide, parallel or slightly convergent
apically. Area petiolaris small, about as long as wide,
0.2-0.25 × as long as propodeum. Femur III 5.1-5.6 ×
as long as wide. Inner spur of tibia III about 0.45 ×
as long as metatarsus III. Metatarsus III about 10 ×
as long as wide and 0.50-0.55 × as long as tibia III.
Claw III with small basal teeth.
Nervulus more or less interstitial. Nervellus in-
tercepted in the lower 0.2, but discoidella indistinct.
Pterostigma about 3.5-4.0 times as long as wide, RI
about 1.2. Second recurrent vein joining areolet in
its middle.
First tergite 2.6 × and postpetiole 1.3-1.7 × as
long as wide, the latter with fine longitudinal stria-
tion in the apical third. Second tergite 0.9 × as long
as wide, its thyridium large, transverse. Ovipositor
sheath 8.2-8.7 × as long as wide, 0.85-0.9 × as long
as metatarsus III, moderately narrowed and bent
upwards apically (Fig. 89).
Colour: Black. Face yellowish (Fig. 44). Cream-
yellow are lower 0.4 of temple, mandible except
teeth, all coxae and trochanters, trochantelli I-II,
mesosternum, propleurum and anterior border of
pronotum, often also hind edge of pronotum, tegula
and subtegular ridge. Mesopleurum black dorsally,
reddish ventrally with a sharp border of both colours
(as Fig. 108), mesosternum more or less yellowish
and/or mesopleurum ventrally with yellow colour.
Metapleurum and scutellum sometimes with dark
red suffusion. Legs reddish-yellow, dorsal part of
femur III often with diffuse brownish suffusion,
tibia III only slightly darkened apically. Tergites
1-2 black, tergite 2 with a triangular yellowish spot
at apex, tergites 3-5 yellowish, brownish laterally,
the following tergites yellowish or slightly vespoid,
epipleura and sternites yellowish, sternites 2-3
often with brown spots laterally. Ovipositor sheath
yellowish-brown.
Description of M
Body length 6-7 mm. Flagellum with 36 flagel-
lomeres, FEI about 0.7. Face wide, minimal width
0.9 × of eye length, its margins divergent ventrally.
MI 0.5. Temples moderately narrowed behind
eyes, with very superficial punctures, TEI 0.7-0.8.
Face punctate, matt, its lateral margins divergent
ventrally. OOD 1.2. Femur III 4.6 × as long as wide.
Postpetiole with fine longitudinal striation in the
apical half. Claw III long, apex moderately curved,
without distinct teeth. Clasper 0.8 × as long as second
metatarsus III, slightly clavate apically.
Colour: Black. Flagellum infuscate. Yellowish
are lower 0.3 of temples, mandibles except teeth,
clypeus and face except a dark longitudinal median
stripe above, coxae I-II, all trochanters and trochan-
telli, propleurum, anterior border of pronotum, hind
edge of pronotum, and tegula. Mesopleurum black
above, reddish below (usually with sharp border),
mesosternum more yellowish. Legs reddish-yellow,
coxa III sometimes with a darkened dorsal spot api-
cally. Femur III slightly darkened apically. Tibia III
more yellowish, infuscate in the apical 0.2, tarsus III
infuscate. Tergites 1-2 black, tergite 2 with a yellow-
ish band or triangular spot at apex, tergites 3-4 with
a sandclock-shaped yellowish median stripe, the
following more or less blackish, sternites yellowish,
with brown spots laterally. Pterostigma brown.
Hosts. Dolerus genucinctus Zaddach, 1859 (Hym.,
Tenthredinidae) (ZSM), Sioblia sturmii (Klug, 1817)
(Hym., Tenthredinidae) (ZSM), Monophadnus spinolae
(Klug, 1816) (Hym., Tenthredinidae) (ZSM).
Material. 59 WW, 3 MM.
Distribution. Austria (CS, ZSM), Bulgaria (ZSM),
Czechia (LINZ), Finland (CJ), Germany (CR, ZSM),
Great Britain (EDI, NHM), Netherlands* (CZ), Nor-
way (CR), Sweden (STOCK, ZSM), Switzerland*
(LAU, NHM).
Astiphromma (Astiphromma) tridentatum
Schwenke, 1999
Figs 46, 47, 90
Astiphromma tridentatum Schwenke, 1999: 20, W.
Holotype W: Rila-Kloster/Bulgar. E 232100/N 420900,
9.7.74, R. Hinz (ZSM), studied.
Taxonomical remark. The extent of pale facial
coloration seems to be quite variable. There are two
specimens – found at the same place and date as the
holotype – with the yellowish colour on the clypeus
and the lower face beside the clypeus only (Fig. 47, as
in A. alpinum). However, they are otherwise similar
to the holotype of A. tridentatum.
Description of W
Body length 6 mm. Flagellum with 33-34 flagel-
lomeres, FEI 0.58-0.61, preapical flagellomeres about
1.5 × as long as wide, apical flagellomere more or
less blunt. Mandible with strong lamella ventrally
and with equally sized teeth. Face punctate and
129
granulate, with fine oblique striation beside clypeus,
its minimal width 1.0 × eye length. Frons granulate,
laterally matt, without punctures. MI 0.6. Temples
strongly narrowed behind eyes, with very superficial
punctation, TEI 0.6-0.7. OOD 1.5-1.7. Genal carina
joining the hypostomal one far from mandibular
base, hypostomal carina not elevated.
Lower part of mesopleurum almost smooth
or with scattered fine punctation. Scutellum not
carinate. Propodeum with complete carination. Area
superomedia confluent with area basalis and slightly
narrowed apically. Costula faint laterally. Area
petiolaris wider than long and slightly wider than
petiolar base, 0.35 × as long as propodeum. Femur
III 3.9-4.2 × as long as wide. Inner spur of tibia III
0.5-0.55 × as long as metatarsus III. Metatarsus III
8.5 × as long as wide and about 0.45 × as long as the
tibia III. Claw III with quite short teeth.
Nervulus postfurcal (0.3-0.5 its length). Pter-
ostigma 3.2-3.5 × as long as wide, RI 1.1-1.3, radius
behind areolet straight. Second recurrent vein joining
areolet in its middle.
First tergite 2.5-3.0 × and postpetiole 1.4-1.6 ×
as long as wide. Postpetiole smooth. Second tergite
0.9-1.0 × as long as wide, its thyridium roundish.
Ovipositor sheath 7.5-9 × as long as wide, about
0.9-1.2 × as long as metatarsus III, in the apical third
slightly bent upwards (Fig. 90).
Colour: Black. Face with three ivory more or less
long stripes (Figs 46, 47). Flagellum brownish, paler
in the basal third, sometimes scapus and pedicellus
yellowish below. Scutellum dark reddish. Front and
middle legs yellowish-red, hind leg reddish-brown,
tibia III infuscate in the apical 0.15-0.25. Tergite 2
with a triangular yellowish spot in the apical third
or with a yellowish band in the apical 0.2, tergite
3 with a sandclock-sized yellowish median stripe,
following tergites reddish-yellow, sometimes with
lateral darkening and paler hind margins (vespoid).
Episterna and sternites yellow, sternites 2-3 with
brown lateral spots. Ovipositor sheath black, with
reddish tip.
M unknown.
Hosts. Unknown.
Material. 5 WW.
Distribution. Austria* (CS), Bulgaria (ZSM), Ger-
many* (ZSM), Sweden (NHM)
Astiphromma (Astiphromma) uliginosum
Schwenke, 1999
Figs 45, 91, 95, 118, 120
Astiphromma uliginosum Schwenke, 1999: 23, W.
Holotype W: ex Kokon an Weiden Gröbenz. [back side]
Schw. Eschenried 7.90 (ZSM), locus typicus Gröben-
zell/D, studied.
Astiphromma trimaculosum Schwenke, 2004: 83, syn. nov.
Holotype W: Horsell Common, Surrey 6/8-14/8/00
Malaise trap, J Pontin (Horniman Museum), studied.
Taxonomical remark. Schwenke (2004) compared
A. trimaculosum with A. uliginosum in his description,
but his proposed differences seem to be individual
variations. Studying both types and additional Eu-
ropean material convinces me to synonymize the
species.
Description of W
Body length 4.0-7.0 mm. Flagellum with 30-35
flagellomeres, slender, FEI 0.6-0.75, preapical flagel-
lomeres about 1.5-2 × as long as wide, apical flag-
ellomere pointed. Mandibular teeth of equal size.
Face punctate and granulate, but shining, lateral
margins parallel or slightly convergent or divergent
ventrally, minimal width 0.95 × eye length (Fig. 45).
Frons and vertex with granulation and very scattered
punctures. Temples strongly narrowed behind eyes,
with superficial puncture, TEI 0.5-0.6. OOD 1.3-1.7.
MI 0.6-0.7. Genal carina joining the hypostomal
one far from mandibular base, hypostomal carina
not elevated.
Mesopleurum with very scattered fine hair-
punctures. Scutellum not carinate. Propodeum
with complete carination, but carinae of area basalis
sometimes faint. Area superomedia about 2.2 × as
long as wide, convergent apically. Area petiolaris
large, wider than long, 0.3 × as long as propodeum.
Femur III 4.5-4.9 × as long as wide. Inner spur of tibia
III 0.45-0.5 × as long as metatarsus III. Metatarsus
III 9-10 × as long as wide and 0.52-0.55 × as long as
tibia III. Claw III with about 4-5 strong teeth, basal
teeth larger than the apical ones.
Nervulus strongly postfurcal (0.4-0.7 its length).
Nervellus of hind wing slightly intercepted in the
apical 0.2, discoidella indistinct. Pterostigma 3.9-4 ×
as long as wide, RI 1.1-1.2, radius behind areolet
straight or slightly sinusoid. Second recurrent vein
joining areolet in its middle.
First tergite 2.4-2.6 × and postpetiole 1.1-1.2 ×
as long as wide, smooth. Second tergite 0.8-1.0 ×
as long as wide, its thyridium large, transverse or
roundish. Ovipositor sheath 7.1-8.4 × as long as wide
and 0.85-1.0 × as long as metatarsus III, narrowed
in the apical 0.4-0.5 (Fig. 91).
130
Colour: Black. Narrow ventral margins of cl-
ypeus and face yellow. Clypeus and face otherwise
black. Flagellum reddish-brown, infuscate apically.
Cream-yellowish are malar space, hind edge of
pronotum, tegula, subtegular ridge, coxae I-II, and
all trochanters. Sometimes pronotum, mesopleurum
ventrally and scutellum with diffuse reddish-brown
suffusion. Legs except coxae and trochanters reddish-
yellow, femur III diffusely darkened at apex, tibia
III infuscate on apical 0.15-0.2. Pterostigma brown
or yellowish-brown. Tergite 1 black, sometimes
with a small yellowish median spot at apex, tergite
2 black, with large triangular reddish spot at apex,
tergites 3-(4) reddish-yellow, with dark brown half-
moon sized sides, following tergites brownish, with
yellowish apical margins (vespoid) (Figs 118, 120).
Sternites mainly cream-yellow, with brownish sides.
Ovipositor sheath brown or blackish, with pale tip.
Description of M (new male)
Body length 5 mm. Mandibular teeth of equal size.
Lateral margins of face divergent ventrally. Frons lat-
erally granulate, medially smooth. Temples strongly
narrowed behind eyes, TEI 0.6. OOD 1.4. Genal carina
joining the hypostomal one far from mandibular
base, hypostomal carina not elevated. Mesopleurum
with very scattered, fine hair-punctures. Scutellum
not carinate. Propodeum with complete carination.
Area superomedia long, slightly narrowed apically.
Area petiolaris wider than long. Femur III 4.1 × as
long as wide. Claw III with basal teeth. Nervulus
postfurcal (0.3 its length). Nervellus of hind wing
not clearly intercepted, discoidella indistinct. Pter-
ostigma 3.4 × as long as wide, RI 1.05, radius behind
areolet straight. second recurrent vein joining areolet
in the middle. First tergite 2,4 × and postpetiole 1.4 ×
as long as wide, smooth. Second tergite 0.9 × as long
as wide, thyridium large, transverse. Clasper about
1.1 × as long as second tarsomere III (Fig. 95).
Colour: Black. Cream-yellowish are mandibles
except teeth, palps, clypeus, face, gena, temple in the
lower 0.3, scapus and pedicellus below, hind edge of
pronotum, tegula, subtegular ridge, tibia III infus-
cate on apical 0.2. Tarsus III brownish. Pterostigma
brown. Tergite 1-2 black, tergite 2 with large trian-
gular yellowish spot apically, tergite 3 yellowish,
with brownish sides, tergites 4-7 brownish, with
yellowish apical margins (vespoid). Sternites 1-3
yellowish, with brown sides, following sternites
brown, with yellow apical margins.
Host. Pseudoips prasinana (Linnaeus 1758) (Lep.,
Noctuidae) (NHM), Epione sp. (Lep., Geometridae)
(Schwenke 1999).
Material. 15 WW, 1 M.
Distribution. Austria* (LINZ), Bulgaria* (ZSM),
Germany (ZSM), Great Britain* (NHM), Greece*
(ZSM), Norway* (CR).
Astiphromma (Astiphromma) varipes
(Holmgren, 1860)
Figs 20, 48, 92, 112
Mesochorus varipes Holmgren, 1860: 120, W.
Holotype W: Sm; Bhn (STOCK), studied.
Description of W
Body length 7-10 mm (rarely smaller, one specimen
5.5 mm). Flagellum with 36-49 flagellomeres, FEI
0.7-0.72, preapical flagellomeres 1.5-2 × as long as
wide, apical flagellomere pointed. Mandibular teeth
of equal size. Face punctate, shining, lateral margins
parallel or divergent ventrally, minimal width 0.9 ×
eye length (Fig. 48). Temples strongly narrowed
behind eyes, with superficial punctation, TEI 0.55
(Fig. 20). Frons smooth centrally, with fine granula-
tion laterally, without clear punctures. OOD 0.8-1.0.
MI 0.7. Genal carina joining the hypostomal one far
from mandibular base (by malar length), hypostomal
carina not elevated.
Lower part of mesopleurum with moderately
dense punctures. Scutellum not carinate. Propo-
deum with complete carination. Area superomedia
moderately narrowed apically, about 2 × as long
as wide, usually confluent with area basalis. Area
petiolaris large, about as long as wide, wider than
petiolar base, 0.35 × as long as propodeum. Femur
III 4.4-4.8 × as long as wide. Inner spur of tibia III
about 0.53 × as long as metatarsus III. Metatarsus III
9 × as long as wide and 0.55-0.6 × as long as tibia III.
Claws III with medium sized teeth.
Nervulus strongly postfurcal (0.3-0.5 its length).
Pterostigma about 4.3 × as long as wide. Radial vein
behind areolet distinctly sinusoid, RI 1.3 (Fig. 112).
First tergite 2.3 ×, and postpetiole 1.3-1.4 × as long
as wide, smooth. Second tergite 0.9-0.95 × as long
as wide, its thyridium roundish. Ovipositor sheath
5.3 × as long as wide, 0.8-0.9 × as long as metatarsus
III, in the apical third slightly narrowed (Fig. 92).
Colour: Black. Gena, mandible except teeth,
palps, coxa I, trochanter I, tegula, hind edge of pro-
notum and spot on subtegular ridge cream-yellow.
Reddish-yellow are apical margin of clypeus and
legs. Coxa III more reddish, femur III brown to
black, tibia III yellowish, basally (0.15) and apically
(0.3) infuscate. Tarsus III infuscate. Pterostigma dark
brown. Sternites brown laterally, medially yellow.
Ovipositor sheath black, with pale tip.
131
Description of M
Body length 6-9 mm. Flagellum with 40-44 flagello-
meres. MI 0.75. TEI 0.5-0.6. Genal carina joining the
hypostomal one far from mandibular base. Radial
vein behind areolet sinusoid. Femur III about 4.3 × as
long as wide. Clasper about 1.2-1.3 × as long as the
second segment of tarsus III, not narrowed apically.
Colour: Black. Mesosternum and mesopleurum
with diffuse yellowish or reddish marks. Cream-
yellow are face, clypeus, gena, mandible except teeth,
palps, coxae I-II, all trochanters, tegula, hind edge of
pronotum and spot on subtegular ridge. Tergites 2-3
more or less yellowish medially or apically. Tibia III
yellowish, with or without a narrow infuscate base,
infuscate in the apical 0.2. Tarsus III cream-yellow
or infuscate.
Hosts. Panolis flammea (Denis & Schiffermüller, 1775)
(Lep., Noctuidae) via Meteorus sp. (Hym., Braconi-
dae) (NHM); Bupalus sp. (Lep., Geometridae) via
Campoplex sp. (Hym., Ichneumonidae) (Schwenke
1999).
Material. 28 WW, 18 MM.
Distribution. Bulgaria (ZSM), Germany (CR, SCH-
NEE, ZSM), Great Britain* (EDI, NHM), Norway
(CR), Sweden (STOCK), Switzerland* (LAU).
Acknowledgements
I would like to thank all persons who cordially helped
me by sending material or supported my work in their
institutes, namely (in alphabetical order) G. Broad (Lon-
don/England), S. Coruh (Erzurum/Turkey), E. Diller
(Munich/Germany), F. Giovanni (Verona/Italy), A.
Freitag (Lausanne/Switzerland), C. Hansson (Lund/
Sweden), J. Hatton (London/England), K. Horstmann
(†) (Würzburg/Germany), R. Jussila (Paattinen/Fin-
land), J. Kolarov (Plovdiv/Bulgaria), S. Schmidt (Mu-
nich/Germany), H. Schnee (Markleeberg/Germany),
M. Schwarz (Linz/Austria), M. R. Shaw (Edinburgh/
Scotland), H. Vardal (Stockholm/Sweden). G. Varkonyi
(Kuhmo/Finland) and K. Zwakhals (Arkel/Nether-
lands).
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Contents
Ruthensteiner, B. & T. Heinzeller: Zum 80. Geburtstag von Dr. Hubert Fechter ................... 1
Alf, A. & K. Kreipl: A new species of the family Turbinidae Rafinesque, 1815 from Saint
Brandon, Western Indian Ocean (Mollusca, Gastropoda, Vetigastropoda, Turbini-
dae) ......................................................................................................................... 3-10
Schmidt-Petersen, J., E. Schwabe & G. Haszprunar: Acanthochitona pilosa spec. nov., a
new species of Acanthochitona Gray, 1821 from the Mediterranean (Mollusca,
Polyplacophora) ...................................................................................................... 11-20
Barros, J. C. N., C. A. S. Santana & S. F. B. Lima: Three new species of Anacithara from
the Southwestern Atlantic Ocean, Brazil (Mollusca, Neogastropoda, Conoidea,
Horaiclavidae) ......................................................................................................... 21-28
Kurt, K. & Ö. K. Erman: Harvestmen fauna of Gümüshane and Bayburt in Turkey (Arachnida,
Opiliones) ................................................................................................................ 29-38
Häussermann, V., C. Spano, M. Thiel & K. B. Lohrmann: First record of the sea anemone
Diadumene lineata (Verrill, 1869) from the Chilean coast (Cnidaria, Anthozoa,
Actiniaria) ................................................................................................................ 39-42
Baehr, M.: A new species of the genus Aristolebia Bates from North Vietnam (Coleoptera,
Carabidae, Lebiini) ................................................................................................. 43-47
Schlatter, R., V. Häussermann & G. Försterra: Some like it hot – hummingbirds making use
of hot springs in Chilean Patagonia (Neornithes, Apodiformes, Trochilidae) ........ 48
Assmann, T., E. Boutaud, J. Buse, V. Chikatunov, C. Drees, A.-L.-L. Friedman, W. Härdtle,
K. Homburg, T. Marcus, I. Renan & D. W. Wrase: The ground beetle tribe Cyclo-
somini s. l. in Israel (Coleoptera, Carabidae) ......................................................... 49-69
Tokár, Z. & T. Jaworski: Batia hilszczanskii spec. nov. from Cyprus (Lepidoptera, Oecopho-
ridae) ...................................................................................................................... 71-76
Hilszczañski, J., R. Plewa, T. Jaworski & A. Sierpiñski: Microrhagus pyrenaeus Bonvouloir,
1872 – a false click beetle new for the fauna of Poland with faunistic and eco-
logical data on Eucnemidae (Coleoptera, Elateroidea) ......................................... 77-84
Riedel, M.: Revision of the European species of the genus Astiphromma Förster, 1869
(Hymenoptera, Ichneumonidae, Mesochorinae) .................................................... 85-132
Schönitzer, K.: Faunistik (Grundlagen, Status, Ausblick) ....................................................... 133-138
Cunha, C. M., R. B. Salvador & L. R. L. Simone: The terrestrial microgastropods of Trinda-
de Island, Brazil (Gastropoda, Pulmonata) ............................................................ 139-143
Lange, T., J. Brehm & T. Moritz: A practical key for the identification of large fish rostra
(Pisces) ................................................................................................................... 145-160
Buchbesprechungen ............................................................................................................. 2, 70, 144
... The Mesochorinae is a globally distributed, moderately species-rich (about 880 described species; Yu et al. 2016) subfamily of specialised hyperparasitoids of, usually, the larvae of other ichneumonoids (Braconidae and Ichneumonidae), with some species-groups apparently specialising on Tachinidae (Diptera) (Dasch 1971;Riedel 2015;Broad et al. 2018). There are competing generic classifications for the subfamily. ...
... Originally described in the monotypic subgenus Dolichochorus of Mesochorus Gravenhorst 1829 (Strobl 1904), the holotype of D. longiceps is at the moment difficult to access and we have been unable to examine it. However, J.F. Perkins (who identified a specimen of D. longiceps in NHMUK and left a rich legacy of identifications based in many cases on examination of European type specimens), W. Schwenke (Schwenke 1999) and M. Riedel (Riedel 2015) have all interpreted the species in the same way and we see no reason not to follow this interpretation. Townes (1971), however, illustrated a typical species of Astiphromma, labeled as A. longipes, a name that does not exist, to our knowledge. ...
... Schwenke (1999) regarded Dolichochorus Strobl 1904 as a valid genus whereas Wahl (1993) had listed it as a synonym of Astiphromma, following Townes et al. (1965), although, as noted above, it is likely that H.K. Townes had not seen reliably identified specimens of D. longiceps. The only originally included species, Mesochorus (Dolichochorus) longiceps Strobl 1904, was also included as a species of Astiphromma by Riedel (2015) in his revision of European Astiphromma, although separated as a subgenus, Astiphromma (Dolichochorus). ...
Article
Dolichochorus Strobl 1904, is shown to be a valid genus of the ichneumonid subfamily Mesochorinae, and a senior synonym of Thamester Wahl 1993 (syn. nov.). Dolichochorus japonicus (Wahl 1993) is a new combination. Dolichochorus longiceps (Strobl 1904) is shown to be a senior synonym of Astiphromma elongatum Nakanishi 1969, syn. nov. The genus Dolichochorus is defined, a revised diagnosis provided for the genus and the two included species are provisionally separated.
... . Western Palearctic region, known from Russia [Riedel, 2015] including Karelia [Humala, 1997]. ...
... , known from Russia [Lee, 1992; Riedel, 2015] including Karelia [Humala, 1997]. ...
... in the Holarctic region, known from Russia [Riedel, 2015] including Karelia [Humala, 1997] (as A. strenuum (Holmgren, 1860)). ...
Article
Full-text available
In this faunistic survey, we give distributional records of 33 species of Mesochorinae (Hymenoptera: Ichneumonidae) which have recently been collected in Northwestern Russia. Of these, 13 taxa are new for the Russian fauna, namely Mesochorus curvicauda Thomson, 1886, M. declinans Habermehl, 1922, M. diluvius Schwenke, 1999, M. errabundus Hartig, 1838, M. frondosus Schwenke, 1999, M. fulgurator Horstmann, 2006, M. haeselbarthi Schwenke, 1999, M. laricis Hartig, 1838, M. perticatus Schwenke, 1999, M. suomensis Schwenke, 1999, and M. tetricus Holmgren, 1860. Two unknown males of Astiphromma flagellator Riedel, 2015 and A. flavoventrale Riedel, 2015 are also described.
... After studying the Schwenke collection in the Zoologische Staatssammlung Munich, Germany, additional informations on the Mesochorus species described by Schwenke and descriptions of some new European Mesochorus species were recently published by Riedel (2018aRiedel ( , 2018bRiedel ( , 2019. The European species of Cidaphus Förster had been revised by Fitton (1985) and the Western Palaearctic Astiphromma Förster by Riedel (2015). Recently, Dolichochorus Strobl was again raised to a valid genus separate from Astiphromma Förster and its members described by Broad & Watanabe (2019). ...
Article
For this study, a part of the Swedish material of the subfamily Mesochorinae (Hymenoptera: Ichneumonidae) collected by the Swedish Malaise Trap Project (SMTP) was determined. In the available material 97 different species of Mesochorinae were identified, 31 of them are new records for the Swedish fauna. Four species are new to science: Mesochorus albofacies sp. nov., Mesochorus longivalvator nov.sp., Mesochorus nigritor sp. nov. and Mesochorus nigroclypeatus sp. nov. The new taxa are described in detail and illustrated here. For Mesochorus frigidus Schwenke, 1999 and Mesochorus macrophyae Schwenke, 1999, the previously unknown males are described. Mesochorus skanensis Vikberg, 2017 is synonymized with Mesochorus terebratus Schwenke, 1999 and Mesochorus acutus Schwenke, 1999 is re-established as a separate species from Mesochorus suomiensis Schwenke, 1999. Stictopisthus Thomson is regarded as a subgenus of Mesochorus Gravenhorst. This is the third contribution to the fauna of Ichneumonidae from the Swedish Malaise Trap Project, following Riedel & Magnusson (2014) and Riedel & Magnusson (2017).