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An index of names of recent Asteroidea. Part 2: Valvatida

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... COMPOSITION: 2 species (Clark, 1993 (Perrier, 1894); "Challenger" (1973)(1974)(1975)(1976)(1977)(1978)(1979)(1980)(1981)(1982), AT 167 ; "Shackleton" (1977), AT 121 ; "Challenger" (1984)(1985), ES 266, AT 267 (Harvey et al., 1988). " Challenger", St. 50304, 51110;"Discovery" (1979), St. 10112 ...
... COMPOSITION: 2 species (Clark, 1993). Type species: Porania grandis Verrill, 1878. ...
... COMPOSITION: 1 species (Clark, 1993). Type species: Tylaster willei Danielssen et Koren, 1881. ...
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An annotated check-list of Asteroidea species occurring deeper than 2000 m in the seas bordering Europe is given. The check-list is based on published data. The list includes 66 species. For each species synonymy, data on localities in European seas and general species distribution are provided. Station data are presented separately in the present thematic issue.
... Los crinoideos y asteroideos recolectados se encuentran depositados en la Colección Nacional de Equinodermos (CNE) del Instituto de Ciencias del Mar y Limnología (ICML), UNAM, México, están preservados en alcohol etílico al 70 % aunque varios asteroideos se encuentran deshidratados en gavetas de la colección seca. Para el arreglo taxonómico del listado se adoptaron los criterios de los siguientes autores, para la clase Crinoidea: Clark y Clark (1967), Meyer et al., (1978), Messing y Dearborn (1990); para la clase Asteroidea: Clark y Downey (1992); Clark (1989), Clark (1993), Clark (1996); Blake (1987), Blake (1989) y Mah y Blake (2012) con algunas corroboraciones en la base datos WoRMS (Messing, 2020;World Register of Marine Species, 2023). ...
... Persephonaster echinulatus fue la especie más abundante con 145 ejemplares y más ampliamente distribuida en la zona de estudio (34 estaciones), así también Pseudarchaster gracilis gracilis con 75 ejemplares presentes en 22 estaciones de muestreo. Se registraron cambios en la sistemática y clasificación taxonómica de la familia Pseudarchasteridae mantenida en el orden Valvatida (Clark & Downey, 1992;Clark, 1993) ahora en el orden Paxillosida (Mah & Blake, 2012). ...
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Introducción: Las especies de las clases Crinoidea y Asteroidea se distribuyen en gran variedad de hábitats en todos los océanos desde la zona intermareal hasta grandes profundidades, existen pocos registros en aguas mexicanas, a profundidades superiores a los 200 m, los escasos registros existentes datan de documentos históricos de la literatura especializada y los ejemplares se resguardan en colecciones científicas extranjeras. Objetivo: Contribuir a la recopilación de información de los ejemplares resguardados en la Colección Nacional de Equinodermos (CNE) del Instituto de Ciencias del Mar y Limnología (ICML), UNAM y proveer un inventario de las especies de crinoideos y asteroideos que habitan en zonas profundas del talud de la Península de Yucatán, México. Métodos: Del año 2005 al 2014 se recolectaron ejemplares de crinoideos y asteriodeos en cuatro cruceros en el B/O “Justo Sierra” BIOREPES 1 y 2 y COBERPES 2 y 6. Se realizaron arrastres con red camaronera en 80 estaciones de muestreo. Resultados: El listado taxonómico comprende 1146 ejemplares, de los cuales 204 son crinoideos del orden Comatulida, distribuidos en tres familias tres géneros y tres especies y 942 asteroideos distribuidos en seis órdenes, 11 familias, 21 géneros y 28 especies. Se obtuvieron los registros nuevos para la CNE: Democrinus rawsonii y Atelecrinus balanoides en crinoideos. Astropecten alligator, Hymenaster modestus, Calyptraster personatus, Pteraster militarioides militarioides, Sclerasterias contorta en asteroideos. Con esta información se incrementan los registros de crinoideos y asteroideos (de profundidad mayor a 200 m) para los estados de Yucatán (16) y Quintana Roo (16).
... Downey (1975) described the eastern Pacific subspecies, namely N. gracilis malpeloensis. This subspecies was subsequently accepted without further analysis or comment (Clark, 1993;Cohen-Rengifo et al., 2009;Pérez-Ruzafa et al., 2013). ...
... The validity of N. gracilis malpeloensis has not been discussed by subsequent authors to date. Clark (1993) The re-examination of 44 individuals of N. gracilis (38 assigned to N. gracilis gracilis and six to N. gracilis malpeloensis) found no morphological support for N. gracilis malpeloensis. Specimens previously attributed to N. gracilis gracilis with a R greater than 100 mm, rarely present pedicellariae. ...
Article
Since the taxonomic revision of the Ophidiasteridae by H.L. Clark (1921), in which 20 genera were recognized as valid, several new genera and species were included in this family, and the boundaries between some of them became largely arbitrary. This also applies to Narcissia Gray, 1840, whose morphology and taxonomy are still very poorly understood today. The main goals of this research were: review the taxonomy of the genus Narcissia, from morphological studies; redescribe N. trigonaria Sladen, 1889 s. str.; characterize N. canariensis (d'Orbigny, 1839) and N. gracilis A.H. Clark, 1916, the two valid species most similar to N. trigonaria; review the validity of N. ahearnae Pawson, 2007 and N. trigonaria var. helenae Mortensen, 1933; review the validity of N. gracilis malpeloensis Downey, 1975. Altogether 370 specimens were analysed. All morphological structures of taxonomic importance have been illustrated, and, for the first time, the internal characters of this group were studied, from electron microscopy and microtomography. A new diagnosis for the genus Narcissia was constructed. Narcissia trigonaria was redescribed from a large specimen, from the type locality (Bahia, Brazil). Narcissia ahearnae is considered a valid species, and N. canariensis and N. trigonaria distinct entities. Narcissia trigonaria var. helenae is placed in the synonymy of N. trigonaria, and N. gracilis malpeloensis into the synonymy of N. gracilis. The dataset obtained in this work is by far the most complete acquired for the genus Narcissia.
... Subsequently, (Rowe 1977) hypothesised that P. horrida was potentially present in the Peruvian coast, which was later confirmed by Hooker and Solís-Marín (2011), who collected an R= 84mm specimen at a depth of 33 m on a rocky reef in Punta Sal, northern Peru. Furthermore, A.M. (Clark 1993) incorporated some localities in his checklist suggesting Baja California, Mexico, as the northernmost distributional limit of the species, also including the localities of Isla Clarión and Isla del Coco, both located in Costa Rica. Finally, (Maluf 1987) broadened the distribution of the species confirming the Galapagos Islands as part of the distributional range ofP. ...
... Distribution map of the sea star Paulia horrida along the Pacific coast of America showing historical records of the species and the new reported locality. Letters in the figure represent records made by the following authors: (A)(Clark 1910(Clark ,1993;(B)(Hooker and Solís-Marín 2011);(C) Present study ...
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We report the capture of a single specimen of the species Paulia horrida Gray, 1840, off the coast of Arica, in the Arica and Parinacota Region, Northern Chile. This finding extends the currently known distributional range of the species by 2,434 km towards the south, since the southernmost known area for this species corresponds to Punta Sal, Peru. We discuss the importance of finding this relatively unknown species in Chile and the need to continue sampling to confirm the continuous distributional range of the species between the south of Peru and the north of Chile.
... Ceramaster patagonicus Sladen (1889) is a widely occurring species, and recorded from the Aleutian Islands and the North Pacific as well as New Zealand and further represented by subspecies in South Africa, Brazil and the Okhotsk Sea. Downey in Clark and Downey (1992) relegated C. patagonicus to a subspecies of Ceramaster grenadensis but this was not met with agreement by Clark (1993) or by Mah (2011) and is considered herein as a separate species complex in accord with O'Hara (1998) and Clark and McKnight (2001). ...
... Ceramaster japonicus' full morphological range is incompletely documented. Clark (1993) lists the full bathymetric range of this species as 194-1410 m. Although Fisher (1911) has briefly covered the morphological range for this species, variation along its bathymetric range is poorly understood and characters such as pedicellariae and furrow spine number might be environmentally related phenotypic variation. ...
Article
Our recent morphological studies on the echinoderm collection of the National Museum of Nature and Science, Tokyo (NMST), indicated that the goniasterid starfishes Ceramaster japonicus (Sladen, 1889) and Ceramaster patagonicus (Sladen, 1889) are distributed in the Pacific Oceanside of Japan. In this study, we studied the NMST samples of C. japonicus and C. patagonicus from Japan, by using two mitochondrial DNA genes, the COI and 12S-rRNA, as markers to test relationships between these species in Japan. C. patagonicus sequences from GenBank were mined and included in the analyses. Results of phylogenetic and haplotype network analyses of both genes (final sequence lengths: COI = 317 bp, 12S = 477 bp) suggested that "Ceramaster patagonicus" and "Ceramaster japonicus" from Japanese waters are almost certainly synonymous without any population structure inside Japan.
... Systematic arrangements follow the criteria of A.M. Clark (1989Clark ( , 1993Clark ( , 1996, A.M. Clark and Mah (2001) and Mah (2014) for the Asteroidea; Smith et al. (1995), Stöhr et al. (2014) for the Ophiuroidea; Mortensen (1928Mortensen ( , 1935Mortensen ( , 1943aMortensen ( , 1943bMortensen ( , 1948Mortensen ( , 1950Mortensen ( , 1951, Kroh and Smith (2010) and Kroh and Mooi (2014) for the Echinoidea; Pawson and Fell (1965), Rowe (1969), Solís-Marín et al. (2009), Smirnov (2012 and WoRMS (2014) for the Holothuroidea. The taxonomic status of all species (valid names and synonyms) was tracked in the literature back to the original description, and was constructed with the help of systematics experts. ...
... During the 19th century, only six works were written; in the 20th century 24 references were published and during the current one, nine works have been published (Fig. 1A). The authors who contributed with the highest number of works were H.L. Clark (1907bClark ( , 1920Clark ( , 1940, Caso (1944Caso ( , 1945Caso ( , 1948bCaso ( , 1970Caso ( , 1977Caso ( , 1979 and A.M. Clark (1989Clark ( , 1993Clark ( , 1996. ...
Article
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The echinoderms of the Southern Mexican Pacific have been studied for three centuries, but discrepancies in the nomenclature of some species have pervaded through time. The objective of this work is to present the first updated checklist of all valid species and synonyms, and a historical review of the study of the echinoderms of the Southern Mexican Pacific is also presented. The checklist is based on an exhaustive published literature search and records of specimens deposited in museum and curated reference collections. There are 162 species of echinoderms in the Southern Mexican Pacific from 96 genera, 54 families and 20 orders. The State of Guerrero presented a total of 135 species, Oaxaca 94 and Chiapas 15. We updated the list and added five new records of the species (Microphiopholis platydisca, Ophiostigma tenue, Arbacia stellata, Thyone bidentata, Chiridota rigida) for the Southern Mexican Pacific and one for the Mexican Pacific (Encope laevis). This checklist expands the number of species known for the study area; nevertheless studies suggest that still more species are yet to be discovered
... The small body size of this sea star raises doubts about the validity of the species. Yamaguchi (1975) suspected that this species represents a juvenile form of Choriaster granulatus Lütken, 1869. Clark (1993 pointed out that the validity of the genus Bothriaster and the species B. primigenius was doubtful because the diagnostic characteristics were attributable to the immaturity of the lost holotype specimen. Liao & Clark (1995) reported that young individuals of C. granulatus have been described under other generic names, implying C. granu ...
Article
One of the oreasterid sea stars, Bothriaster primigenius, is a rarely encountered species in the tropical Indo-West Pacific. The taxonomic position of this species, and more specifically, whether to recognize this sea star as a species, remains unresolved. To resolve the question, we assessed the specific validity of this sea star based on morphological observations and molecular phylogenetic analysis. The results revealed that this species is a young individual of another oreasterid sea star, Choriaster granulatus, and is placed as a junior synonym for the same.
... Posteriormente, se examinaron ejemplares preservados en seco y alcohol etílico (70 %) depositados en dichas colecciones. La corroboración de la identidad taxonómica de los ejemplares se realizó con las siguientes claves y descripciones: Perrier (1875), Sladen (1891), Verrill (1899;1915), Caso (1961), Ummels (1963), Halpern (1970), Downey (1973) El listado taxonómico sigue la clasificación propuesta por A. M. Clark y Downey (1992) y Mah y Foltz (2011), así como los cambios nomenclaturales revisados en Halpern (1970), A. M. Clark (1989Clark ( , 1993Clark ( , 1996, A. M. Clark y Mah (2001) y Mah (2018. ...
... Results from Foltz et al. (2013) listed several previously accepted species as synonyms of H. phrygiana (Parelius, 1768), the type name for species within Hippasteria (Clark, 1993;Mah & Hansson, 2012). Sampled taxa that were shown to be synonyms of H. phrygiana by Foltz et al. (2013) include H. spinosa , H. armata (Fisher, 1911, H. kurilensis , H. armata Fisher, 1911, and H. aleutica Clark & Jewett, 2011 further removed any doubt regarding synonymy of the Patagonian H. hyadesi Perrier, 1891 or the New Zealand H. trojana Fell, 1958 into H. phrygiana. ...
... The poorly studied family Poraniidae from the order Val vatida is distributed throughout the Atlantic, Indian, Pacific, and Southern Oceans (Clark, 1984(Clark, , 1993. Recently, the classification of this group has been revised based on mor phological and molecular phylogenetic data (Mah and Foltz, 2014). ...
Article
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A sea star was collected from the Korea Strait in the waters adjacent to eastern Jeju Island, Korea (33°39′86″N, 127°33′12″E) at a depth of 92 m on November 5, 2016. This specimen was identified as Marginaster paucispinus Fisher, 1913, from the family Poraniidae of the order Valvatida, based on morphological characteristics. The genus Marginaster Perrier, 1881 and M. paucispinus, which were first reported in the South China Sea, are new to the Korean fauna. Partial sequences of mitochondrial COI and 16S ribosomal RNA of M. paucispinus were have been determined for the first time and were deposited in GenBank. They are the first molecular records for the genus Marginaster.
... As suggested in Hart et al. (1997) these include a revision of P. gunnii, splitting the east and western forms. The combination Patiriella pseudoexigua pacifica (Hayashi) is listed as "new comb" by Clark (1993) and so this species need not be referred to Asterina, a change supported by the molecular data. A further recommendation from the present study is a reassignment of the nominal species P. pseudoexigua from Taiwan. ...
... Asterina pseudoexigua pacifica Hayashi, 1977 Asterina pseudoexigua pacifica Komatsu et al., 1990 Patiriella pseudoexigua pacifica AM Clark, 1993 Material examined: Kushimoto, Wakayama Prefecture, coll. Mioko Matsubara, 20 Nov. 2002 Distribution: Known from Kushimoto, Wakayama Prefecture, Japan. ...
Article
Dartnall, Alan J., Byrne, Maria, Collins, John, Hart, Michael W (2003): A new viviparous species of asterinid (Echinodermata, Asteroidea, Asterinidae) and a new genus to accommodate the species of pantropical exiguoid sea stars. Zootaxa 359: 1-14, DOI: 10.5281/zenodo.156371
... Among them, 33 species of asteroids were recorded from Mudasalodai landing centre [23]. The wide distribution of echinoderms was reported and 18 genera and 117 species of the Asteroidea were listed [24]. Eight genera and 37 species of asteroids from the Australian region, including 10 Astrina sp and 10 Patiriella sp were recorded [25]. ...
... Among them, 33 species of asteroids were recorded from Mudasalodai landing centre [23]. The wide distribution of echinoderms was reported and 18 genera and 117 species of the Asteroidea were listed [24]. Eight genera and 37 species of asteroids from the Australian region, including 10 Astrina sp and 10 Patiriella sp were recorded [25]. ...
Article
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Objective: To study the echinoderms diversity in Mudasalodai and Pazhayar landing centers during November 2012 to December 2014. Methods: All the collected echinoderm samples were stored at 4 °C until further use. Results: As many as 19 species of echinoderms were recorded in both stations and it belongs to 10 families, three classes such as Asteroidea, Echinoidea and Ophiuroidea. The percentage composition varied from 0.15% to 73%. The species diversity ranged from 1.031 to 2.6; species richness was from 1.02 to 4.06 and species evenness ranged from 0.93 to 0.99. K-dominance curve drawn paralleled the trend of diversity indices. Cluster and multidimensional scaling showed the similarity in echinoderms composition within the stations. Conclusions: The present investigation is evident that the echinoderm occurrence was predominant during the summer season and proved statistically by using SPSS Primer 6.0.
... Para la organización taxonómica general del listado, se siguieron los criterios de los siguientes autores: para la clase Asteroidea: A.M. Clark (1989Clark ( , 1993Clark ( , 1996; para la clase Ophiuroidea: Mastumoto (1915), Fell (1960); para la clase Echinoidea: Mortensen (1928Mortensen ( , 1935Mortensen ( , 1940Mortensen ( , 1943Mortensen ( , 1948Mortensen ( , 1950Mortensen ( , 1951; para la clase Holothuroidea: Pawson y Fell (1965), Smiley y Pawson (1991). ...
Article
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Echinoderms (Echinodermata) from the Mexican Pacific. A systematic list of echinoderms of the Mexican Pacific, based on museum specimens of the Coleccion Nacional de Equinodermos, Instituto de Ciencias del Mar y Limnologia, Universidad Nacional Autonoma de Mexico and the National Museum of Natural History, Smithsonian Institution, Washington, D.C., is presented. A total of 196 echinoderm species is recorded, distributed in 112 genera, 56 families and 20 orders. Eight new records for the Mexican Pacific are presented: one for Class Crinoidea: Hyocrinus foelli; six for Class Asteroidea: Echinaster (Echinaster) parvispinus, Henricia nana, Henricia seminudus, Rathbunaster californicus and Leptasterias pusilla, and one for Class Ophiuroidea: Amphiodia tabogae. Rev. Biol. Trop. 56 (Suppl. 3): 57-73. Epub 2009 January 05.
... Para la organización taxonómica general del listado presentado se siguieron los criterios de los siguientes autores: para la Clase Crinoidea : Clark, (1947: Clark, ( , 1950; para la Clase Asteroidea: Clark y Downey (1992), Clark (1989Clark ( , 1993Clark ( , 1996; para la Clase Ophiuroidea: Matsumoto (1915), Fell (1960; para la Clase Echinoidea: Mortensen (1928Mortensen ( , 1935Mortensen ( , 1940Mortensen ( , 1943Mortensen ( , 1948Mortensen ( , 1950Mortensen ( , 1951; para la Clase Holothuroidea: Deichmann (1954), Pawson y Fell (1965). ...
... Para la organización taxonómica general del listado se adoptaron los criterios de los siguientes autores: para la clase Crinoidea: Clark (1947Clark ( , 1950, Clark y Clark (1967); para la Clase Asteroidea: Clark y Downey (1992), Clark (1989Clark ( , 1993Clark ( , 1996, O'Loughlin y Waters (2004); para la clase Ophiuroidea: Matsumoto (1915), Fell (1960; para la clase Echinoidea: Mortensen (1928Mortensen ( , 1935Mortensen ( , 1940Mortensen ( , 1943Mortensen ( , 1948Mortensen ( , 1950Mortensen ( , 1951; y para la clase Holothuroidea: Deichmann (1954), Pawson y Fell (1965). ...
Article
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We present a systematic list of the echinoderms from the Central American Caribbean, including the archipelago of San Andres, based on specimens deposited at the National Museum of Natural History, Smithsonian Institution, Washington D.C., the Invertebrate Zoology and Geology collections of the California Academy of Sciences, San Francisco, the Museo de Zoología, Universidad de Costa Rica, San José, Costa Rica, the literature and internet. A total of 253 echinoderm species are recorded, distributed in 142 genera, 63 families and 27 orders. Ophiuroidea (79 species) and Asteroidea (56 species) are the most diverse classes. Panama (154 species) and Belize (131 species) have the highest reported echinoderm diversities.
... All species in this paper are described and illustrated in Lambert (1981). Classification follows Clark (1989Clark ( , 1993Clark ( , 1996. (Fisher 1928 (Fisher 1930). ...
Article
Based on museum specimens, the known northern limit of three species of sea stars is extended north to localines in the Gulf of Alaska. The maximum depth limit of six species and the minimum depth limit of eight species are also extended.
... En la sección de resultados, los nuevos registros están señalados con un asterisco (*). Para la organización taxonómica general del listado, se siguieron los criterios de los siguientes autores: Clase Asteroidea: A. M. Clark (1989, 1993, 1996) Clark y Downey (1992, y Clase Echinoidea: Mortensen (1928Mortensen ( , 1935Mortensen ( , 1940Mortensen ( , 1943Mortensen ( , 1948Mortensen ( , 1950Mortensen ( , 1951 De los tres ordenes de la clase Asteroidea representados en la zona, el orden Valvatida fue el más diverso con cuatro familias. La familia más representada fue Ophidiasteridae con tres géneros. ...
Article
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We present a systematic list of the echinoderms (Asteroidea, Echinoidea) from the coast of Oaxaca in the Southern Mexican Pacific, based on museum specimens of the Colección Nacional de Equinodermos, Instituto de Ciencias del Mar y Limnología, Universidad Nacional Autónoma de México, National Museum of Natural History, Smithsonian Institution, Washington, D.C., and fieldwork. A total of 27 echinoderm species is recorded: 23 genera, 17 families and 9 orders. We include 11 new records for the state of Oaxaca, Mexico.
... Halityle Fisher, 1913, is a monotypic genus. Hubert Lyman Clark (1946) transferred Culcitaster anamesus H. L. Clark, 1914, to Halityle, but it is regarded as a junior synonym of H. regularis (Baker & Marsh 1976;A.M. Clark 1993;Döderlein 1935). ...
Article
Sea stars were collected at depths ranging from 91 to 160 m around Kumejima Island situated in the middle of the Ryukyu Islands chain in the Western Pacific. Sampling was carried out using a dredge or beam trawl in November 2009, as part of the KUMEJIMA 2009 Expedition. The specimens included an undescribed species of the genus Neoferdina, and three species that have never been reported in Japanese waters. Features that characterise this new species, Neoferdina longibrachia, include: extremely long and narrow arms, no alternating arrangement of large and small superomarginal plates, body surface (except for the marginal plates) completely covered with granules, and an adambulacral armature with 3 furrow spines.
... Classification of Asteroidea adopted herein follows Clark and Downey (1992). To establish synonymies, original literature dealing with descriptions and records of deep-water Asteroidea in the East Pacific was consulted, in addition to important reviews by A. M. Clark (1989Clark ( , 1993Clark ( , 1996 and A. M. Clark and Mah (2001). Other sources are indicated in the text where appropriate. ...
Article
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During a series of cruises aboard the R/V "EI Puma" aimed at collecting the deep-water benthic and pelagic fauna off the Pacific coast of Mexico, in the eastern Pacific, samples of Asteroidea were collected below 500 m depth (587-1 526 m). A total of 335 specimens were collected, belonging to 18 species, 14 identified to species, 3 to genus, and 1 previously undescribed species. New records are provided for Dipsacaster laetmophilus Fisher, 1910, Myxoderma sacculatum (Fisher, 1905), Peribolaster biserialis Fisher, 1905, Ampheraster chiroplus Fisher, 1928, Ampheraster hyperoncus (H. L. Clark, 1913), Anteliaster coscinactis Fisher, 1923, Nearchaster aciculosiis (Fisher, 1910), Ceramaster leptoceramus (Fisher, 1905), Mediaster transfuga Ludwig, 1905, and Lophaster furcilliger Fisher, 1905. All species were collected below the oxygen minimum zone that extends throughout the central and southern Gulf of California, or within the threshold zone where severe to mild hypoxic conditions prevail. Epibenthic dissolved oxygen concentrations associated with the capture of the specimens show support for strong tolerance to severe hypoxia (<1.0 ml O 2/l) for most species, but only mild hypoxia for Ctenodiscus crispatus (Retzius, 1805), and Nymphaster diomedeae Ludwig, 1905. A checklist of all species of Asteroidea occurring below 500 m depth off the Pacific coast of Mexico is included.
... This book is divided in 16 chapters (including this one) and two taxonomic lists (Appendix) (one from the Pacific Ocean and one from the Caribbean Sea and Atlantic Ocean including the Canary Islands) with information about presence/ absence, bathymetric range and the type of habitat or substrate where the species have been found. These taxonomic lists were built following the criteria of Clark (1931Clark ( , 1947 for Crinoidea; Clark (1989Clark ( , 1993Clark ( , 1996, Clark and Downey (1992), Clark and Mah (1999) for Asteroidea; Matsumoto (1915), Stöhr and O'Hara (2007) for Ophiuroidea ;Smith (1984) for Echinoidea and Panning (1929Panning ( , 1934aPanning ( ,b, 1935, Pawson and Fell (1965) for Holothuroidea. The World Register of Marine Species (Appeltans et al. 2011) was also consulted for all species. ...
Chapter
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Latin America (LA) includes all countries in the Americas that speak Spanish or Portuguese; in total it includes 22 countries including Mexico, Central America, the islands of the Caribbean and South America and Haiti (Hispaniola Island, shared with Dominican Republic). This book is divided in 16 chapters and two taxonomic lists with information about presence/absence, bathymetric range and the habitat or substrate where the species have been found. Fourteen chapters deal with studies done in countries, regions or islands. The final chapter has a biogeographical and diversity analysis of the two taxonomic lists. Each country chapter contains six sections. 1. An introduction. 2. A historic account of the investigations on echinoderms. 3. Distributional patterns and diversity. 4. Fishery or aquaculture activities. 5. Threats to echinoderms, and 6. A final section with general conclusions from the review and recommendations for future research to meet the needs of each country. Latin American echinoderm studies started a long time ago, as far back as the 13th century. The most common threats to echinoderms are bad coastal planning, coastal pollution, illegal fisheries, inefficient fisheries management, lack of protection, and lack of baseline studies. A total of 1,539 species (21.8 % of the worldwide species) are included in the taxonomic lists (appendix), with 717 species in the Pacific and 926 in the Caribbean and Atlantic. The richest class is Ophiuroidea followed by Asteroidea. All classes have more species in the Caribbean and Atlantic than the Pacific.
... Halityle Fisher, 1913, is a monotypic genus. Hubert Lyman Clark (1946) transferred Culcitaster anamesus H. L. Clark, 1914, to Halityle, but it is regarded as a junior synonym of H. regularis (Baker & Marsh 1976;A.M. Clark 1993;Döderlein 1935). ...
Article
Sea stars were collected at depths ranging from 91 to 160 m around Kumejima Island situated in the middle of the Ryukyu Islands chain in the Western Pacific. Sampling was carried out using a dredge or beam trawl in November 2009, as part of the KUMEJIMA 2009 Expedition. The specimens included an undescribed species of the genus Neoferdina, and three species that have never been reported in Japanese waters. Features that characterise this new species, Neoferdina longi-brachia, include: extremely long and narrow arms, no alternating arrangement of large and small superomarginal plates, body surface (except for the marginal plates) completely covered with granules, and an adambulacral armature with 3 furrow spines.
... The Jamaican isolates are broadly consistent with the single Atlantic species of Nymphaster, N. arenatus (Perrier, 1881), recognized by Clark & Downey (1992). Approximately 17 species of Nymphaster are recognized worldwide however (Clark, 1993), and all would need to be compared in any comprehensive evaluation. Complexities are further illustrated through reference to Fisher (1919), who treated many goniasterid genera -at least five (Rosaster Perrier, 1894;Paragonaster Sladen, 1889;Ateloris Fisher, 1911;Lithosoma Fisher, 1911;and Iconaster Sladen, 1889) -including species with simple marginal ossicle morphology superficially similar to that of the Antillean specimens. ...
Article
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The Antillean Cenozoic fossil record of asteroids comprises mainly dissociated ossicles. Most common among isolates from upper Oligocene deposits of Jamaica and Antigua are marginal ossicles of an extinct, indeterminate species of Pycinaster. This is the youngest known occurrence of the genus and the first from beyond Europe. A number of relatively complete fossils have been assigned to Pycinaster and (sub)familial status proposed for it together with Phocidaster. The latter proposition is based solely on a few marginals, but available diagnoses are judged insufficient to justify such recognition. The taxon Pycinasteridae is here synonymized with the Goniasteridae, although future study of added features (such as the ventral surface) might justify recognition at a higher taxonomic level. In addition to ossicles assigned to Pycinaster, many marginals are tentatively assigned to the surviving goniasterid Nymphaster. Numerous generic and many species names have been based on asteroid isolates, but the practice demands assumptions that are not readily justified. Linkage of discrete isolates under a single taxon name assumes derivation from a single source, an inference that can be verified only rarely (if ever), therefore reducing names to the single holotype ossicle. Availability of only isolates encourages comparison with extant taxa and biogeography, biasing interpretations with a Holocene overlay. Because of these constraints, a new nominal species of Pycinaster is not justified and assignment of ossicles to Nymphaster is tentative. However, given the importance of asteroids in marine communities, we emphasize the significance, largely ignored, of their presence in Cenozoic deposits of the wider Caribbean.
... , D. longispina (H. L. Clark, 1938), originally considered as a species of the separate genus Manasterina H. L. Clark, 1938, has no uncalci ed skin (H. L. Clark 1938O'Loughlin and Waters 2004). In the latest revision of the genera of Asterinidae, O' Loughlin and Waters (2004) revised Disasterina, which thus currently has ve valid species (see also A. M. Clark 1993;O'Loughlin 2009;Mah 2010). All of the species are distributed in the Indo-West Paci c region (Döderlein 1888;Koehler 1910;H. L. Clark 1916H. L. Clark , 1938Livingstone 1933;Mortensen 1933;A. M. Clark and Rowe 1971;Liao 1980). No species of Disasterina has ever before been found in Japanese waters. ...
Article
A new asterinid, Disasterina akajimaensis sp. nov., is described from a specimen collected at Akajima Island, the Ryukyu Islands, Japan. It is clearly distinguishable from its five congeners by the following diagnostic characters: body relatively low and flat, uncalcified interradial areas conspicuous on ventral interradii, papulae numerous in center of disc, abactinal plates mostly lacking spinelets, inferomarginal plates with 2-4 spinelets, actinolateral plates with 1 spine, oral plates with 4-5 oral spines and no suboral spine, adambulacral plates with 1-3 furrows and 1 subambulacral spine. This is the first report of this genus from Japanese waters. Notes on the genus Disasterina and a tabular summary of the characters of the currently known species are also given.
... Main publications Anthozoa (Hexacorallia) Daniellsen, 1890;Carlgren, 1913Carlgren, , 1942Jensen, 1992;Fautin, 2006 Maxilopoda (Scalpelliformes) Sars, 1886;Broch, 1953;Tarasov, Zevina, 1957 Malacostraca (Decapoda, Euphausiacea, Isopoda, Mysida, Amphipoda) Sars, 1886;Hansen, 1916;Stephensen, 1923Stephensen, , 1925Stephensen, , 1931Stephensen, , 1944Gurjanova, 1946Gurjanova, , 1951Gurjanova, , 1964Barnard et al., 1962;Bulycheva, 1964;Lomakina, 1964;Dahl, 1979;Kussakin, 1979Kussakin, , 1982Kussakin, , 1988Kussakin, , 1999Kussakin, , 2003Just, 1980a, b;Petrashov, 1993Petrashov, , 2004Svavarsson et al., 1993;Malyutina, Kussakin, 1996a, b, c;Brandt, 1997Brandt, , 2005Sokolov, 2000;Kamenskaya, 2001;Malyutina, Weisshappel, 2001;Vader et al., 2005;Brix, Svavarsson, 2010Scaphopoda Bouchet, Warén, 1979Ivanov, Zarubina, 2004Bivalvia Bernard, 1979Bouchet, Warén, 1979;Knudsen, 1985;Richling, 2000Crinoidea Clark, 1970Echinoidea Mortensen, 1903, 1907Mironov, 1995;Holothurioidea Madsen, Hansen, 1994;Rogacheva, 2007;Rogacheva et al., 2013Asteroidea Danielssen, Koren, 1884Clark, 1989Clark, , 1993Clark, , 1996Clark, Mah, 2001Ophiuroidea Mortensen, 1933Djakonov, 1954;Stöhr, OHara, 2012Ascidiacea Hartmeyer, 1923Millar, 1966;Monniot, Monniot, 1979 Several classes Gorbunov, 1946;Baranova, 1964;Koltun, 1964;Afanasjev, Filatova, 1980;Smirnov, 1994;Kröncke, 1998;Deubel, 2000;Sirenko, 2001;Sirenko et al., 2004;Smirnov, Smirnov, 2006;Budaeva et al., 2008;Bergman et al., 2009;Soltwedel et al 2009;MacDonald et al., 2010;Appeltans et al., 2012 whereas 43 species occur both shallower and deeper than 2000 m. Hence, endemics of the abyssal zone in the Arctic Ocean comprise 19% of the Arctic abyssal species fauna. ...
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A list of genera represented in the abyssal (>2000 m) of the Arctic Ocean was compiled for 8 classes (Scaphopoda, Bivalvia, Crinoidea, Echinoidea, Holothuroidea, Asteroidea, Ophiuroidea and Ascidiacea) and some orders from the classes Anthozoa, Maxillopoda and Malacostraca. This list includes 92 genera and 133 species. Species endemic to the Arctic Ocean comprise 51% whereas species endemic to the Arctic abyssal comprise 19%. Only two small genera are endemic to the Arctic Ocean. Genera with a worldwide distribution (60 genera or 65%) dominate the Arctic abyssal fauna. Most genera (55 or 60%) can be considered as deep-sea specialists, with half or more of the species in each genus occurring deeper than 2000 m. About one third of genera (37%) are known outside the Arctic in the hadal zone (>6000 m). The share of genera known from the hadal is probably higher in the Arctic abyssal than in the abyssal of other oceans. For many genera, the worldwide minimum or maximum of their depth ranges is found in the Arctic Ocean. Distribution patterns of genera suggest that many deep-sea Arctic species derive from their congeners distributed in geographically distant regions (primarily the Southern Ocean and the Indo-West Pacific). The deep-sea North Atlantic was the main transit region on the pathway to the Arctic. There is no firm evidence of the presence of derivatives of the pre-Pliocene deep-sea Arctic fauna in the modern Arctic fauna. Twelve genera (13%) with a worldwide distribution are recognized as the most reliable examples of the emergence of the abyssal fauna in the Arctic Ocean. Two genera (2%) are characterized by the following distribution and ecological patterns: (1) distribution is limited to the Northern Hemisphere, (2) wide bathymetrical range (from shelf to abyssal), (3) abyssal records only within the Arctic Ocean, and (4) very wide ranges of habitats. Arctic submergence of the shelf fauna to the abyssal zone is suggested for these genera. Some higher taxa, not considered in the present work in detail, may have a high share of genera with similar distribution patterns. For instance the share of genera having submerged from the Arctic shelf to the abyssal is about 25% in the abyssal Arctic amphipod fauna. KEY WORDS: distribution patterns, genera, Arctic Ocean, abyssal, faunal submergence, faunal emergence. Îñîáåííîñòè ãëîáàëüíîãî ðàñïðîñòðàíåíèÿ ðîäîâ, îáèòàþùèõ â Ñåâåðíîì Ëåäîâèòîì îêåàíå íà ãëóáèíàõ áîëåå 2000 ì
... We examined 760 asteroid specimens from samplings of Agassiz trawls. Their taxonomic classifica-tion according to Koehler (1906), Clark (1962Clark ( , 1989Clark ( , 1993Clark ( , 1996, and Bernasconi (1964) is shown in table II. The resulting taxonomic list included 12 families and 27 genera. ...
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Frequency and distribution of asteroid genera in the South Shetlands zone are analysed, based on data from 24 Agassiz trawls carried out during the Bentart 95 Survey. Asteroids were collected at more than 90 % of the stations, on all types of bottoms, with richness per station ranging from 0-10 genera. The specimens belonged to 27 genera and 12 families, with Astropectinidae, Goniasteridae and, above all, Asteriidae having the highest number of representatives (3 and 7 genera, respectively). Odontaster Verrill, 1880 and Labidiaster Lütken, 1872, followed by Psilaster Sladen, 1885 and Diplastrias Perrier, 1888, were found to have the widest distribution and highest occurrence. Odontaster was more frequent to a depth of 100 m, and Labidiaster between 200 and 300 m. The rarest genera were Notioceramus Fisher, 1940 and Chitonaster Sladen, 1889, endemic to Antarctic waters. Stations showing the highest richness (10 genera) were those located near the Antarctic Peninsula and north of Livingston Island. Asteroid poverty was noticeable inside Deception Island, where a mere 3 genera were recorded. As shown by similarity and grouping analysis based on presence-absence data, stations are mainly grouped in relation to genera richness when double absence is considered. These results show the distintiveness of the stations around Deception Island, probably due to its special geological characteristics.
... Distribution patterns have emerged from the analysis of the data compiled for this chapter, which have been compared to information available from the literature (see e.g. Hedgpeth 1969, Clark andDowney 1992). There are more records available from the literature from the last decade, which were purposedly not considered in this study, as they are not published in recognised biodiversity information systems, such as biodiversity.aq, ...
... The localities referred to for all specimens in this study are shown inFigure 1. Material: Six specimens (DNSM ECH16), Mntafufu River mouth; 11 specimens (DNSM ECH17), Ballito; five specimens (DNSM ECH18), Park Rynie. Distribution: South Africa, from Mozambique west to Namibia; St Helena, Amsterdam and St Paul islands, South Indian Ocean and southern Australia; depth range 0–10 m (Clark 1993). Remarks: De Lamarck (1816) described this species from specimens from 'Les mers d'Amerique'. ...
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The genus Heliaster (Gray, 1840) includes a remarkable group of seven species that inhabit rocky intertidal and shallow subtidal habitats and are morphologically characterized by a large disc and a large number of rays. Geographically, the genus is restricted along the tropical, subtropical, and temperate coasts of the Eastern Pacific. Four species inhabit mainland littoral sites: Heliaster helianthus (Lamarck, 1816) in the tropical, subtropical and temperate west coasts of South America; Heliaster polybranchius H. L. CLARK, 1907, in the tropical coast of Peru; Heliaster kubiniji XANTUS, 1860 and Heliaster microbrachius Xantus, 1860, in the west coasts of Mexico and central America. Three species inhabit off shore archipelagos: Heliaster canopus Perrier, 1875, in Chile's Juan Fernández Archipelago; and Heliaster solaris A. H. Clark, 1920, and Heliaster cumingii (Gray, 1840) in the Galápagos Archipelago of Ecuador (see J. E. Gray 1840, H. L. Clark 1907, A. H. Clark 1920, A. M. Clark and Mah 2001). © 2013 The Johns Hopkins University Press. All rights reserved.
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A review of high-latitude Goniasteridae south of 50°S is presented, including a key to species, figures and taxonomic descriptions of all species with a brief taxonomic summary for all genera. Nineteen species in 11 genera are reviewed. Era-tosaster jenae nov. gen and sp., Chitonaster trangae nov. sp., and Pillsburiaster calvus nov. sp. are described. Several taxa, including Chitonaster, have not been reviewed since their initial description and revision of the group has clarified taxonomic boundaries between species. The genus Pentoplia is a junior synonym of Chitonaster. Rarely encountered species, synonyms, and range extensions are reported for multiple species.
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Seven asteroids are described from the Tlayua quarries (18° 35´ N - 97° 56´ W), recovered from the Middle Member of the Tlayua Formation. Tlayua quarries are 94 km southeast of Puebla and 2 km northeast of the town of Tepexi de Rodriguez in the Eastern part of central Mexico. The Middle Member is fossiliferous and composed mainly of a micritic lithographic limestone with layers having packs that consist of as much as 30 % hematite. The fossils are confined to these iron rich layers. These Middle Member are about 28 m thick. Due to the abundance, variety and conservation of its paleobiota, the Tlayua quarry is one of the most important fossiliferous localities from North America’s Cretaceous. About 5 000 samples of invertebrate and vertebrate fossils have been collected from this locality and have been studied. The seven specimens belong to the Cretaceous strata and are believed to be of Lower Cretaceous (Albian) age. The asteroids described are referred to the genus Astropecten, Plutonaster, Tamaria and Echinaster. This is the first report of Cretaceous asteroids for Mexico.
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Twenty-seven species of asteroids have been collected over the period 1991-1995 by dredging, SCUBA-diving, and hand-collecting in the territorial waters of Singapore. They include fourteen species newly recorded for the area: Luidia hardwicki (Gray), Astropecten novaeguineae Döderlein, Asterina coronata von Martens, Nepanthia belcheri (Perrier), Nepanthia maculata Gray, Patiriella pseudoexigua Dartnall, Tegulaster ceylanica (Döderlein), Anthenea aspera Döderlein, Gymnanthenea laevis H.L. Clark, Fromia monilis Perrier, Ophidiaster granifer Lütken, Tamaria fusca Gray, Echinaster callosus von Marenzeller, Echinaster stereosomus Fisher. A total of thirty-one species, belonging to nine families, are known presently from off Singapore. A distribution table and an illustrated identification key to the species are provided. A new synonymy is proposed: Asterina spinigera Koehler becoming a junior synonym of Asterina coronata von Martens.
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