Taxonomic revision of the longicorn beetle genus Uracanthus Hope 1833 (Coleoptera: Cerambycidae: Cerambycinae: Uracanthini) from Australia

Abstract

A thorough taxonomic revision of the Australian Uracanthus is provided, including a redefinition of the scope of the genus, descriptions and illustrations of new and previously known species, and a key to all species. The revised Australian Uracanthus includes 39 species, eight of which are established as new to science: U. pseudogigas sp. nov., U. maculatus sp. nov., U. griseus sp. nov., U. bicoloratus sp. nov., U. perthensis sp. nov., U. punctulatus sp. nov., U. quadristriolatus sp. nov., and U. bistriolatus sp. nov. Six new synonyms are proposed (senior synonyms last): U. multilineatus McKeown with U. ventralis Lea, U. dentiapicalis McKeown with U. parvus Lea, U. marginellus Hope and U. inermis Lea (not Aurivillius) with U. bivitta Newman, U. fuscostriatus McKeown with U. lateroalbus Lea, and U. daviumbus Gressitt with U. longicornis Lea. Dorsal views of all species are presented as photographs, terminalia of both sexes illustrated, and distributions mapped. The larvae of this genus are borers of at least 31 genera of trees and parasitic plants in 15 families (Asteraceae, Betulaceae, Casuarinaceae, Cupressaceae, Fabaceae, Loranthaceae, Myrtaceae, Pittosporaceae, Proteaceae, Rhamnaceae, Rosaceae, Rutaceae, Sapindaceae, Sterculiaceae, and Xanthorrhoeaceae), including some economically important crops such as citrus, lychee (or litchi), peach, plum, and apricot. Several species are important pests of orchards. Uracanthus adults visit flowers of various tree species and are attracted to the light.

Full text

ZOOTAXA
Taxonomic revision of the longicorn beetle genus Uracanthus Hope
1833 (Coleoptera: Cerambycidae: Cerambycinae: Uracanthini)
from Australia
DUANGRAT THONGPHAK & QIAO WANG
Magnolia Press
Auckland, New Zealand
1569

THONGPHAK & WANG
2 · Zootaxa 1569 © 2007 Magnolia Press
DUANGRAT THONGPHAK & QIAO WANG
Taxonomic revision of the longicorn beetle genus Uracanthus Hope 1833 (Coleoptera: Ceram-
bycidae: Cerambycinae: Uracanthini) from Australia
(Zootaxa 1569)
139 pp.; 30 cm.
31 August 2007
ISBN 978-1-86977-133-1 (paperback)
ISBN 978-1-86977-134-8 (Online edition)
FIRST PUBLISHED IN 2007 BY
Magnolia Press
P.O. Box 41-383
Auckland 1346
New Zealand
e-mail: zootaxa@mapress.com
http://www.mapress.com/zootaxa/
© 2007 Magnolia Press
All rights reserved.
No part of this publication may be reproduced, stored, transmitted or disseminated, in any form, or by any
means, without prior written permission from the publisher, to whom all requests to reproduce copyright
material should be directed in writing.
This authorization does not extend to any other kind of copying, by any means, in any form, and for any purpose
other than private research use.
ISSN 1175-5326 (Print edition)
ISSN 1175-5334 (Online edition)

Accepted by C.N.Duckett: 25 May 2007; published: 31 Aug. 2007 3
ZOOTAXA
ISSN 1175-5326 (print edition)
ISSN 1175-5334 (online edition)
Copyright © 2007 · Magnolia Press
Zootaxa 1569: 1–139 (2007)
www.mapress.com/zootaxa/
Taxonomic revision of the longicorn beetle genus Uracanthus Hope 1833
(Coleoptera: Cerambycidae: Cerambycinae: Uracanthini) from Australia
DUANGRAT THONGPHAK & QIAO WANG1
Entomology and IPM Laboratory, Institute of Natural Resources, Massey University, Private Bag 11222, Palmerston North, New
Zealand.
1To whom correspondence should be addressed: Q.Wang@massey.ac.nz
Table of contents
Abstract . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .4
Introduction . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4
Materials and Methods . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5
Taxonomy . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .6
Genus Uracanthus Hope, 1833 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6
Key to the species of Uracanthus Hope . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7
Uracanthus bicoloratus, sp. nov. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 10
Uracanthus maculatus, sp. nov. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .12
Uracanthus perthensis, sp. nov. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .15
Uracanthus griseus, sp. nov. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 16
Uracanthus glabrilineatus Lea, 1917a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .17
Uracanthus triangularis Hope, 1833 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 18
Uracanthus pallens Hope, 1841 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 22
Uracanthus regalis McKeown, 1948 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .24
Uracanthus dubius Lea, 1916 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 25
Uracanthus simulans Pascoe, 1866 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .27
Uracanthus fuscus Lea, 1916 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .29
Uracanthus cryptophagus Olliff, 1892 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 30
Uracanthus fuscocinereus White, 1855 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 32
Uracanthus quadristriolatus, sp. nov. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .33
Uracanthus insignis Lea, 1916 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 35
Uracanthus punctulatus, sp. nov. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .36
Uracanthus maleficus Lea, 1917a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .37
Uracanthus gigas Lea, 1916 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 38
Uracanthus pseudogigas, sp. nov. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 40
Uracanthus acutus Blackburn, 1889 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 41
Uracanthus loranthi Lea, 1916 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .43
Uracanthus cupressianus Rondonuwu and Austin, 1988 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 45
Uracanthus minatus Pascoe, 1866 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 46
Uracanthus longicornis Lea, 1916 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 47
Uracanthus albatus Lea, 1916 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 49
Uracanthus ventralis Lea, 1917a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 52
Uracanthus strigosus Pascoe, 1875 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 53
Uracanthus parallelus Lea, 1916 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 56
Uracanthus froggatti Blackburn, 1894 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 57
Uracanthus tropicus Lea, 1916 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .58
Uracanthus parvus Lea, 1916 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 59
Uracanthus pertenuis Lea, 1916 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .61
Uracanthus bivitta Newman, 1838 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 63
Uracanthus discicollis Lea, 1916 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 66
Uracanthus corrugicollis Lea, 1917a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 69
Uracanthus lateroalbus Lea 1916 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .70
Uracanthus bistriolatus, sp.nov. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 71
Uracanthus suturalis Lea, 1916 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 73
Uracanthus ater Lea, 1917a . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 73
Acknowledgements . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 74
References . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 74

THONGPHAK & WANG
4 · Zootaxa 1569 © 2007 Magnolia Press
Abstract
A thorough taxonomic revision of the Australian Uracanthus is provided, including a redefinition of the scope of the
genus, descriptions and illustrations of new and previously known species, and a key to all species. The revised Austra-
lian Uracanthus includes 39 species, eight of which are established as new to science: U. pseudogigas sp. nov., U. macu-
latus sp. nov., U. griseus sp. nov., U. bicoloratus sp. nov., U. perthensis sp. nov., U. punctulatus sp. nov., U.
quadristriolatus sp. nov., and U. bistriolatus sp. nov. Six new synonyms are proposed (senior synonyms last): U. multi-
lineatus McKeown with U. ventralis Lea, U. dentiapicalis McKeown with U. parvus Lea, U. marginellus Hope and U.
inermis Lea (not Aurivillius) with U. bivitta Newman, U. fuscostriatus McKeown with U. lateroalbus Lea, and U. davi-
umbus Gressitt with U. longicornis Lea. Dorsal views of all species are presented as photographs, terminalia of both
sexes illustrated, and distributions mapped. The larvae of this genus are borers of at least 31 genera of trees and parasitic
plants in 15 families (Asteraceae, Betulaceae, Casuarinaceae, Cupressaceae, Fabaceae, Loranthaceae, Myrtaceae,
Pittosporaceae, Proteaceae, Rhamnaceae, Rosaceae, Rutaceae, Sapindaceae, Sterculiaceae, and Xanthorrhoeaceae),
including some economically important crops such as citrus, lychee (or litchi), peach, plum, and apricot. Several species
are important pests of orchards. Uracanthus adults visit flowers of various tree species and are attracted to the light.
Key words: Cerambycidae, Cerambycinae, Uracanthus, Australia, revision
Introduction
The genus Uracanthus is a large group of longicorn beetles found in the Australian Region, predominantly
distributed on the Australian mainland and Tasmania. The larvae of this genus are borers of at least 31 genera
of trees and parasitic plants, including some economically important crops such as citrus, lychee (or litchi),
peach, plum, and apricot. Adults visit flowers of various tree species and are attracted to artificial light. As
several species are important pests of fruit trees in Australia, this genus may represent a biosecurity risk for
countries that trade with Australia.
Uracanthus was proposed by Hope (1833) under the family Stenochoridae with U. triangularis as the
type species (by monotypy). During the nineteenth century, eleven species were described either in Uracan-
thus or in other genera and later transferred to Uracanthus (Boisduval 1835, Hope 1834, 1841a, 1841b, 1844,
Gahan 1893, Laporte 1840, Newman 1838, White 1855, Pascoe 1866, 1875, Blackburn 1889, 1894 & Olliff
1892).
The first half of the twentieth century saw a great activity in taxonomic work on Uracanthus by Aurivil-
lius (1917), Lea, (1916, 1917), and McKeown (1938, 1940, 1942, 1947 & 1948). Among those workers, Lea
(1916 & 1917) made the most significant contribution to the genus. In 1916, he revised the genus, described
fourteen new species and provided a key to 22 known species. One year later, he added six new species to the
genus. During this period, species descriptions were mainly based on the shape of the elytral apex, and the
pubescence pattern on the elytral disc and prothorax.
Duffy (1963) was the first to deal with immature stages of Uracanthus. In 1963, he described immature
stages of three species and summarised the distribution and biology of twelve species in Uracanthus. Ron-
donuwu and Austin’s (1988) work is the most recent taxonomic treatment of this genus, where they described
a new species, U. cupressianus, from South Australia. In this work, these authors used many more characters
including genitalic characters.
Prior to the current study, 37 species were recognized from Australia with a number of variants proposed
(Lea 1916 & 1917a, b, McKeown 1947 & 1948, Rondonuwu & Austin 1988), and four from New Guinea
(Gressitt 1951 & 1959). However, the taxonomy of this genus is still very unsettled, making species identifi-
cation, pest management and risk analysis, and biogeographic evaluation difficult. Therefore, a comprehen-
sive taxonomic revision of the Australian specise of Uracanthus was necessary.
We have examined all available Australian Uracanthus species and found five new synonyms and eight
new species; we propose these here. We have also seen types of two New Guinean species, U. daviumbus

Zootaxa 1569 © 2007 Magnolia Press · 5
REVISION OF URACANTHUS
Gressitt 1951 and U. albopleuron Gressitt 1959. The first species is found to be a synonym of U. longicornis
Lea 1916. The second one appears very different from the rest of Uracanthus species and its generic status
needs further investigation. The types or any other specimen of another two New Guinean species, U. declivis
Gressitt 1951 and U. stueberi Gressitt 1959, could not be located. Even though Gressittt (1951, 1959) indi-
cated that the type of U. declivis was in Taiwan Agricultural Research Institute, and that of U. stueberi in
Leiden, the Netherland, we could not find them. Therefore, the three New Guinean ‘Uracanthus’ species were
excluded in this study.
This study aims to provide a thorough taxonomic revision of the Australian Uracanthus, including a
redefinition of the scope of the genus, descriptions and illustrations of new and previously known species, and
a key to all Australian species. The revised Australian Uracanthus consists of 39 species, of which 31 are
redescribed and eight described as new. Six new synonymies are also proposed.
Materials and Methods
Specimens examined
Thousands of Uracanthus specimens were examined during the course of this study. Types of all species
were examined. Specimens were borrowed from following institutions, abbreviations of which were shown in
text:
AM Australian Museum, Sydney
AMNH American Museum of Natural History, New York
ANIC Australian National Insect Collection, CSIRO, Canberra
ASCU Agricultural Scientific Collections Unit, Orange Agricutural Institute, Orange
BMNH The Natural History Museum, London
FCNI Forestry Commission of New South Wales Insect Collection, Beecroft
HMO Hope Museum, Oxford, London
MAM Macleay Museum, Sydney
NTM Museum and Art Gallery of the Northern Territory, Darwin
NMV Museum of Victoria, Melbourne
QM Queensland Museum, Brisbane
QDPI Queensland Department of Primary Industries Insect Collection, Brisbane
SAM South Australian Museum, Adelaide
UQIC University of Queensland Insect Collection, Brisbane
WADA Western Australian Department of Agriculture Insect Collection, Perth
WAM Western Australian Museum, Perth
WINC Waite Insect and Nematode Collection, Adelaide
VAIC Victorian Agricultural Insect Collection, Melbourne
Measurement and illustrations
External morphological measurements (Fig. 1) were made under a stereomicroscope (Leica M212), using
a digimatic-meter (Mitutoyo, Japan). Measurements of terminalia were made under a compound microscope
(Olympus, CH), using ocular micrometers. Drawings of terminalia were made under the compound micro-
scopes with the aid of camera lucida attachments. Dorsal views of each species were photographed using a
Nikon Coolpix4500® digital camera (Tokyo, Japan) with a Nikon Coolight SL (Tokyo, Japan) for illumina-
tion.

THONGPHAK & WANG
6 · Zootaxa 1569 © 2007 Magnolia Press
Dissection of terminalia
The terminalia were prepared by soaking the whole beetle in a water bath set at 80°C for 10–20 mins
depending on the size of the beetles. The softened beetle was then fastened onto a dissecting board using
insect pins, and the terminalia were carefully removed with a pair of tiny sharp forceps, without removing the
abdomen. Terminalia were then cleared in 10% KOH at room temperature for 24 hrs. The terminalia were
then washed with distilled water and placed on a glass slide for measurement, drawing and photographing.
Dissected terminalia of all species except U. gigas Lea and U. pseudogigas sp. nov. were placed in glycerine
in small plastic vials (6mm dia x 12 mm long, BioQuip Inc., USA), pinned with the specimens from which
they originated. Terminalia of U. gigas and U. pseudogigas were mounted in Euparal as permenant slides. All
slides bear terminalia slide numbers that match the specimens from which they originated.
Geographic distribution and biological records
Distribution records were made from collection labels of each specimen examined and from the literature.
The latitude and longitude of distribution localities were confirmed using the online Australian atlas (http://
www.ga.gov.au/map/names/).
Terminology and descriptions
Morphological terminology in this study follows that of Lawrence and Britton (1991), Bense (1995),
Wang (1998), Wang and Lu (2004) and Lu and Wang (2005a & b). Terminology for terminalia partially fol-
lows that of Sharp and Muir (1912), Ehara (1954) and Wang (1993, 1998).
Taxonomy
Genus Uracanthus Hope, 1833
Uracantha Hope, 1833: 64.
Uracanthus Hope, 1834: 108; Laporte, 1840: 425; Thomson, 1860: 143, 151; 1864: 155, 413; Lacordaire, 1869: 390;
Aurivillius, 1912: 147; Lea, 1916: 368; McKeown, 1947: 62. [Type species: U. triangularis Hope, 1833 (by mono-
typy)].
Diagnosis
Genae long and parallel; head with transverse wrinkles on ventral side; prothorax constricted anteriorly;
elytra long, subparalell, gradually tapering towards apex, with longitudinal glabrous stripes and/or sub-basal,
glabrous (or sparsely pubescent) and subtriangular marks, or with dense pubescence throughout; tibia with 2
spurs at apex; hind tarsal segment 1 as long as or longer than sum of segments 2 and 3.
Description
Body size. Male body length: 11.81–53.78 mm; width: 1.71–10.83 mm. Female body length, 11.11–60.56
mm; width, 1.88–12.30 mm.
Colour (Figs 8–9). Body reddish to blackish brown with golden or whitish pubescence; many species with
1 or 2 longitudinal stripes of dense pubescence on each side of pronotal disc; elytron pubescent throughout, or
pubescent with 2–3 longitudinal glabrous stripes and/or a large sub-basal, glabrous (or sparsely pubescent)
and subtriangular mark.
Head (Figs 2–4). Head narrower than prothorax; postclypeus very large, triangular (Fig. 2A) or semicircu-
lar (Fig. 2B) in shape, flattened to convex, with coarse punctures and pubescence; distance between lower
lobes of eyes less than 2.5 × as long as distance between upper lobes of eyes; distance between upper lobes of
eyes about as long as 0.6 × or more than 0.6 × as long as distance between eyes on ventral side (Fig. 2C);
genal length less than 0.9 × as long as head width immediately below eyes (Figs 2B and 2D). Antennae slen-

Zootaxa 1569 © 2007 Magnolia Press · 7
REVISION OF URACANTHUS
der (Fig. 3), composed of 11 segments and filiform with the first segment (scape) most robust; several flagel-
lal segments may be produced at apex (Figs 3b and 4a) or fringed (Fig. 4b) in some species. Antennae may be
longer or shorter than body. Apical ¼ to 1/3 of segment 11 in males usually thinner than basal part, making the
segment appear to be 2 segments (Fig. 4c).
Thorax and abdomen (Figs 5–11). Prothorax longer than wide, constricted at anterior margin in most spe-
cies (Fig. 5); sides rounded or angular (Fig. 6); pronotal disc binodulose in middle and finely punctate in most
species; disc and sides transversely rugose in most species (Figs 7, 8). Elytra (Fig. 9) subparallel, gradually
tapering towards apex, less than 7.0 × as long as prothorax and less than 5.0 × as long as shoulder width; disc
with dense, fine punctures at basal half, gradually diminishing towards apex; disc of each elytron with 2–3
longitudinal carinae in most species; apex rounded, bispinose, unispined at suture, truncate or pointed (Fig. 9).
Legs (Fig. 10) slender, femora slightly thickened near apical 1/3; upper surface of femora and tibia with dense
pubescence; basal segment of hind tarsus as long as or longer than the sum of segments 2 and 3. Abdomen
(Fig. 11a) slender, with dense uniform pubescence but in males of U. albatus, U. ventralis and U. strigosus,
each sternite also with medial tuft of dense hairs (Fig. 11b); each sternite slightly longer than wide; apex of
terminal sternite truncate, round or round with notch.
Male terminalia (Figs 12–16). Aedeagus consisting of median lobe, median strut and internal sac (Fig.
12a). Apex of ventral lobe pointed (Fig. 12b), rounded (Fig. 12c), truncate (Fig. 12d) or emarginate (Fig. 12e);
dorsal lobe longer or shorter than ventral lobe. Internal sac (Fig. 12a) divided into three regions: basal
unspined region, spined region and terminal region. Spined region with 1–4 forms of spines or processes (Fig.
13); terminal region long and narrow, without any spine; sum of unspined and spined regions shorter than ter-
minal region. Tegmen with two parameres (Fig. 14), paramere long and slender or short and robust, cylindri-
cal in shape, apex rounded with long and short setae. Eighth sternite obliquely truncate (Fig. 15b) or rounded
(Fig. 15a, c, d) at side; shallowly or strongly emarginate at apex; ventral surface with 1–2 kinds of
microspines or cloud-like processes. Eighth tergite (Fig. 16) with 1–3 forms of spines on ventral surface; apex
rounded (Fig. 16a), pointed (Fig. 16b), truncate (Fig. 16c) or emarginate (Fig. 16d).
Ovipositor and spermatheca (Figs 17–18). Ovipositor usually very short; styli arising terminally (Fig. 17).
Spermatheca curved to different extent; spermathecal gland arising near (Fig. 18a) or at base (Fig. 18b).
Distribution
This group of beetles is widely distributed in all states of Australia, mainly along the coast.
Biology
The biology of species of the genus Uracanthus is largely unknown. According to fragmentary observa-
tions (reviewed by McKeown, 1947, Duffy, 1963, and Matthews, 1997; observed by Moore, 1972; Ron-
donuwu & Austin, 1988; Hawkeswood, 2002) and labels of specimens examined for this study, at least 31
plant genera are recorded as host plants of Uracanthus: Acacia, Aster, Amyema, Amygdalus, Armeniaca,
Banksia, Boronia, Callitris, Cassinia, Casuarina, Citrus, Correa, Corylus, Cupressus, Cytisus, Eucalyptus,
Eriostemon, Grevillea, Hakea, Helichrysum, Jacksonia, Lasiopetalum, Litchi, Lomatia, Loranthus, Mela-
leuca, Olearia, Pittosporum, Pomaderris, Pultenaea, and Ulex. Adults visit (and probably feed on) flowers of
Angophora, Eucalyptus, Leptospermum, Melaleuca and Xanthorrhoea. Larvae of many species bore inside
the branches and stems of living trees, and some are pests of citrus, lichi, peach, plum, and apricot.
Key to the species of Uracanthus Hope
1 Genal length at least 0.2 × as long as distance between genae .............................................................2
Genal length about 0.16 × as long as distance between genae; pubescence on elytra short and uniform;

THONGPHAK & WANG
8 · Zootaxa 1569 © 2007 Magnolia Press
apex of elytra more or less rounded at margin and with a small spine at suture ....................................
........................................................................................................U. corrugicollis Lea (Fig. 53; p. 69)
2(1) Each elytron with 3 or more distinct longitudinal lines of denser pubescence ................................... 3
Elytra without such distinct pubescent lines ........................................................................................ 8
3(2) Elytral apex spined at suture and rounded at margin; body blackish; pronotal disc with white pubes-
cence and 5 glabrous areas (2 more or less round on each side and 1 oval near base ............................
.............................................................................................................. U. suturalis Lea (Fig. 56; p. 73)
Elytral apex bispinose, if only spined at suture, then angled at margin; body reddish or reddish brown
............................................................................................................................................................... 4
4(3) Pronotal disc with 4 longitudinal stripes of dense or fairly dense pubescence .....................................5
Pronotal disc without 4 such pubescent stripes ....................................................................................6
5(4) Sides of pronotum almost rounded; each elytron with 3 raised longitudinal carinae on which pubes-
cence is denser; males without tufts of dense brownish hairs on sternites .............................................
....................................................................................................U. cryptophagus Olliff (Fig. 30; p. 30)
Sides of pronotum with a strongly raised, rounded or conical process; elytral disc without distinct lon-
gitudinal carinae; male sternites 1–3 with a medial tuft of very dense brownish hairs ..........................
.........................................................................................................U. strigosus Pascoe (Fig. 45; p. 53)
6(4) Each elytron with a large subtriangular glabrous mark, extending from shoulder to basal 1/3, some
part of the mark covered with sparse pubescence........................U. glabrilineatus Lea (Fig. 23; p. 17)
Elytra without the above subtriangular mark.........................................................................................7
7(6) Elytral apex distinctly bispinose.............................................................U. albatus Lea (Fig. 43; p. 49)
Elytral apex spined at suture and angled or slightly produced at margin................................................
...............................................................................................................U. ventralis Lea (Fig. 44; p. 52)
8(2) Body very large, 39–60 mm long ......................................................................................................... 9
Body usually much smaller; if large, shorter than 39 mm ..................................................................10
9 (8) Elytral apex bispinose or truncate with processes at both suture and margin; pronotal disc without a
ψ- or ‡-shaped glabrous mark ...................................................................U. gigas Lea (Fig. 36; p. 38)
Elytral apex spinose at suture and rounded at margin; pronotal disc with a ψ- or ‡-shaped glabrous
mark ........................................................................................ U. pseudogigas, sp. nov. (Fig. 37; p. 40)
10(8) Each elytron with 1 sutural and 1 marginal longitudinal stripes of dense yellowish or whitish pubes-
cence for entire length of elytron, stripe width various ......................................................................11
Elytra without such stripes ................................................................................................................. 13
11(10) Elytral pubescent stripes narrow, sum of sutural and marginal stripe width narrower than the area
between stripes at mid elytron; elytral apex strongly bispinose with the spine at margin distinctly
longer than that at suture .....................................................................U. pertenuis Lea (Fig. 50; p. 61)
Elytral pubescent stripes wide, sum of sutural and marginal strip width wider than the area between
stripes at mid elytron; elytral apex not strongly biospinose ................................................................12
12(11) Pronotal disc strongly rugose transversely; each side of pronotum with 2 spots of dense yellowish
pubescence; elytral apex emarginate, if weakly bispinose, then spine at suture distinctly longer than
that at margin ...................................................................................U. bivitta Newman (Fig. 51; p. 63)
Pronotal disc mostly smooth; each side of pronotum without 2 pubescent spots; elytral apex spined or
angled at suture and rounded at margin ............................................. U. discicollis Lea (Fig. 52; p. 66)
13(10) Each elytron with a large glabrous or sparsely pubescent subtriangular or semicircular mark, extend-
ing from shoulder to about basal 1/3 of elytra ....................................................................................14
Elytra without such marks ................................................................................................................. 23
14 (13) Subtriangular or semicircular marks on elytra completely or largely covered with sparse pubescence
..............................................................................................................................................................15

Zootaxa 1569 © 2007 Magnolia Press · 9
REVISION OF URACANTHUS
Subtriangular or semicircular marks on elytra completely or largely glabrous ..................................16
15(14) Elytral apex distinctly bispinose; body usually longer than 25 mm........U. dubius Lea (Fig. 27; p. 25)
Elytral apex emarginate or truncate with a spine or process at suture; body usually shorter than 25 mm
...................................................................................................................U. fuscus Lea (Fig. 29; p. 29)
16(14) Elytral apex with a small spine at suture ....................................... U. simulans Pascoe (Fig. 28; p. 27)
Elytral apex distinctly emarginate and bispinose ............................................................................... 17
17(16) Apical area of elytron distinctly glabrous, margined with a distinct line of dense white pubescence 18
Apical area of elytron pubescent .........................................................................................................19
18(17) Pronotal disc rugose transversely mainly anteriorly and posteriorly, with 2 distinct longitudinal stripes
of white pubescence; prothorax more slender ..................................... U. pallens Hope (Fig. 25; p. 22)
Pronotal disc transversely rugose throughout, with 2 vague or less defined longitudinal stripes of
white pubescence; prothorax more robust......................................... triangularis Hope (Fig. 24; p. 18)
19(17) Pubescence on elytral disc uneven, with a number of denser pubescence clusters among less dense
pubescence ................................................................................U. maculatus, sp. nov. (Fig. 20; p. 12)
Pubescence on elytral disc more or less even, without the pubescence patterns mentioned above ...20
20(19) Dorsal view of body greyish with basal half of elytra whitish ...........................................................21
Dorsal view of body brownish ............................................................................................................22
21(20) Golden or brownish pubescence present on the following areas: pronotal disc between 2 stripes of
white pubescence, scutellum, and a narrow stripe along suture of elytron extending from basal 1/5 to
apex ............................................................................................ U. regalis McKeown (Fig. 26; p. 24)
Pronotum and elytra with more or less uniform whitish pubescence......................................................
..........................................................................................................U. griseus, sp. nov. (Fig. 22; p. 16)
22(20) Pronotal disc with 2 clearly defined longitudinal stripes of white pubescence and golden or brownish
pubescence between the stripes; pubescence on elytral disc along suture golden or brown and that on
the remaining elytra whitish......................................................U. bicoloratus, sp. nov. (Fig. 19; p. 10)
Pronotal disc with 2 vague longitudinal stripes of white pubescence; pubescence on both pronotum
and elytra whitish ...................................................................... U. perthensis, sp. nov. (Fig. 21; p. 15)
23(13) Pronotal disc with a wide, clearly defined longitudinal glabrous area, if this area covered with sparse
pubescence, then each elytron with 4 short, longitudinal glabrous marks at base; elytral apex dis-
tinctly and strongly bispinose ..............................................................................................................24
Pronotal disc without above mentioned glabrous area; elytral apex various ......................................28
24(23) Wide longitudinal sub-glabrous area on pronotal disc covered with sparse pale pubescence; basal 1/3
of elytra with dense white pubescence and remaining elytra with golden pubescence; each elytron
with 4 basal, longitudinal glabrous marks, with the one along suture the longest...................................
...................................................................................................U. fuscocinereus White (Fig. 31; p. 32)
Wide longitudinal area on pronotal disc completely or almost completely glabrous .........................25
25(24) Anterior end of longitudinal glabrous area on pronotal disc distinctly wider than posterior end; apical
area of elytra almost completely glabrous .............................................U. insignis Lea (Fig. 33; p. 35)
Anterior end of longitudinal glabrous area on pronotal disc narrower than posterior end; apical area of
elytra covered with pubescence ..........................................................................................................26
26(25) Sides of pronotum covered with dense white or pale yellowish pubescence .........................................
...................................................................................................U. punctulatus, sp. nov. (Fig. 34; p. 36)
Each side of pronotum with a more or less distinct, longitudinal glabrous stripe ..............................27
27 (26) Pronotum wider than long; elytra much longer than 4.5 × as long as prothorax ....................................
.............................................................................................................U. maleficus Lea (Fig. 35; p. 37)
Pronotum longer than wide; elytra distinctly shorter than 4.5 × as long as prothorax ...........................
............................................................................................U. quadristriolatus, sp. nov. (Fig. 32; p. 33)

THONGPHAK & WANG
10 · Zootaxa 1569 © 2007 Magnolia Press
28 (23) Elytral apex very sharply pointed ................................................ U. acutus Blackburn (Fig. 38; p. 41)
Elytral apex rounded, truncate or emarginate, with or without spines ...............................................29
29( 28) Elytral apex rounded or slightly pointed, without spines .................................................................. 30
Elytral apex truncate or emarginate, with or without spines .............................................................. 31
30(29) Elytral apex rounded; elytral disc with very dense and long golden pubescence; antennae, legs, metas-
ternite and abdomen blackish brown and remaining body yellowish brown; ventral side of femora
with dense, long golden hairs in males ............................................U. minatus Pascoe (Fig. 41; p. 46)
Elytral apex slightly pointed; elytral disc with fairly dense and short golden pubescence; body reddish
brown in general, without above mentioned sharp colour contrast; male femora not as above .............
...........................................................................U. cupressianus Rondonuwu & Austin (Fig. 40; p. 45)
31(29) Pronotum with a clearly defined longitudinal stripe of dense golden pubescence on each side of disc;
elytral apex truncate or slightly emarginate...........................................U. loranthi Lea (Fig. 39; p. 43)
Pronotum without clearly defined pubescent stripes mentioned above ............................................. 32
32(31) Pronotal disc smooth or weakly rugose transversely.......................................................................... 33
Pronotal disc strongly rugose transversely ..........................................................................................35
33(32) Elytral apex truncate, with very dense white pubescence; pubescence on elytral disc unevenly distrib-
uted, giving an appearance of irregular clustered patterns ........U. froggatti Blackburn (Fig. 47; p. 57)
Elytral apex emarginate, truncate or spinose but without very dense white pubescence; pubescence on
elytral disc more or less evenly distributed ........................................................................................ 34
34(33) Elytral apex bispinose or truncate; each elytron with a wide submarginal, longitudinal glabrous mark
ranging from shoulder extending to 1/3 to 2/3 of elytron length, if the mark obscure or absent, then
elytral apex truncate ..............................................................................U. tropicus Lea (Fig. 48; p. 58)
Elytral apex emarginate and bispinose; elytral shoulder with a small longitudinal glabrous mark ........
............................................................................................................ U. parallelus Lea (Fig. 46; p. 56)
35(32) Elytral apex emarginate and clearly bispinose, or at least with a distinct spine at margin..................36
Elytral apex truncate or slightly emarginate, with a distinct spine at suture ......................................37
36(35) Body usually longer than 24 mm; pronotal disc binodulose.....U. bistriolatus, sp. nov. (Fig. 55; p. 71)
Body usually shorter than 21 mm; pronotal disc without any distinct nodules.......................................
..................................................................................................................U. parvus Lea (Fig. 49; p. 59)
37(35) ..Elytra with relatively sparse, evenly distributed, white pubescence; each side of pronotum without a
distinct longitudinal stripe of very dense white pubescence ....................... U. ater Lea (Fig. 57; p. 73)
Elytra with relatively dense whitish or yellowish pubescence, clearly denser near margin; each side of
pronotum near ventral side with a distinct longitudinal stripe of very dense white pubescence ........38
38(37) Antennal segment 3 shorter than segment 4; basal half of elytra with dense, coarse and shallow punc-
tures; male antennae distinctly longer than body............................. U. longicornis Lea (Fig. 42; p. 47)
Antennal segment 3 longer than or as long as segment 4; basal half of elytra without above mentioned
punctures; antennae shorter than body..............................................U. lateroalbus Lea (Fig. 54; p. 70)
Uracanthus bicoloratus, sp. nov.
(Figs 19, 58, 94)
Description
Male. Body length: 23.26–33.12 mm; width: 4.55–6.97 mm.
Colour (Fig. 19). Head, thorax, basal elytra, and legs dark reddish brown; antennae and remaining part of
elytra reddish brown. Frons and clypeus with dense golden pubescence; vertex with dense white pubescence
except a V-shaped glabrous area extending from medial frontal groove to posterior edge of vertex; legs and
abdomen with fairly dense white pubescence. Pronotum with 2 distinct longitudinal stripes of dense white

Zootaxa 1569 © 2007 Magnolia Press · 11
REVISION OF URACANTHUS
pubescence on each side: 1 on disc and 1 near ventral side; the remaining part of disc with fairly dense golden
pubescence. Elytra with dense golden and white pubescence; each elytron with a large sub-basal triangular
glabrous mark extending from shoulder to basal 1/3, margined with a very vague line of white pubescence;
apical area with sparse pubescence, margined with dense white pubescence. Body colour varies from reddish
brown to blackish brown, the sub-basal glabrous mark may be variable in size, either reduced or expanded.
Head. Postclypeus semicircular to subtriangular in shape and slightly convex, with dense coarse punc-
tures; frontoclypeal suture deep and narrow in middle; median frontal groove wide and deep; distance
between lower lobes of eyes 1.67–2.0 × as long as distance between upper lobes of eyes; distance between
upper lobes of eyes 1.2–1.4 × as long as distance between eyes on ventral side; genal length 0.36–0.43 × as
long as head width immediately below eyes. Antennae shorter than body; 5–10 flattened and produced on one
side at apex.
Thorax and abdomen. Pronotum about 1.11–1.32 × as long as wide, rounded with a very feeble process at
each side; posterior margin about 1.16–1.33 × as wide as anterior margin; pronotal disc feebly binodulose in
middle and strongly rugose transversely. Scutellum semicircular, with dense pubescence. Elytra 4.45–5.07 ×
as long as prothorax and 3.44–3.82 × as long as shoulder width; each elytron with three feeble longitudinal
carinae; basal elytra finely punctate; apex widely emarginate and bispinose. Abdomen slender; apex of termi-
nal sternite truncate.
Male terminalia. Apex of ventral lobe narrowly emarginate and apex of dorsal lobe pointed; dorsal lobe
shorter than ventral lobe; spined region of internal sac 3.45–4.04 × as long as basal unspined region; spined
region divided into 2 sections with unspined gap between sections: first section bearing dense long simple
spines, with fairly dense multi-branched spines near sides; second section with sparse simple spines (Fig.
58a). Eighth sternite strongly obliquely truncate at side, shallowly emarginate at apex, with long and fairly
long setae arising terminally; ventral side with very sparse multi-branched spines and fairly dense cloud-like
processes (Fig. 58b). Eighth tergite rounded at apex, with fairly dense short simple spines and multi-branched
spines on dorsal surface (Fig. 58c). Paramere short, 1.46–1.53 x as long as wide, cylindrical in shape; apex
rounded with long and short setae (Fig. 58d).
Female. Body length, 17.17–34.18 mm; width, 3.24–7.19 mm
Similar to male but differs in having shorter antennae and legs, and more robust abdomen; apex of termi-
nal sternite rounded; elytra 3.50–3.97 × as long as shoulder width and 4.42–5.17 × as long as prothorax.
Ovipositor and spermatheca. Ovipositor short; styli arising terminally with short hairs (Fig. 58e). Sper-
matheca clearly curved; spermathecal gland long and arising near base of spermatheca (Fig. 58f).
Distribution
Southeastern Australia (Fig. 94).
Biology
Banksia, Correa, Olearia and Helichrysum were recorded as host plants of this species. Adults were col-
lected from October to April, May to July and attracted to MV light traps.
Comments
Lea (1916) considered this new species as a variant (Var A) of U. triangularis. Having examined all avail-
able specimens, we found that the ‘var A’ is different from U. triangularis at species level. It resembles U. tri-
angularis but differs in having the sub-basal glabrous marks on elytra without a distinct marginal line of white
pubescence; apical areas of elytra covered with sparse pubescence, and dorsal lobe of aedeagus shorter than
ventral lobe.
Material examined
Holotype. , SA: Wooley Lake, near Beachport, ii.1965, at MV light; terminalia tube pinned with the
specimen; bearing a red holotype label (SAM).

THONGPHAK & WANG
12 · Zootaxa 1569 © 2007 Magnolia Press
Paratypes. 29 , 40 . All paratypes bear blue paratype labels. QLD: 1 , Stanthorpe, xii.1964, E. Sutton,
E. Sutton Collection (AM); 1 , Lamington National Park, 17-12.ii.1964, G. Monteith and H. A. Rose
(UQIC); 1 , 1 , Kilcoy District, 18.ii.1954, S. Gum (QM); 1 , 520 m, Wonga Hills site, 11.xii.2001, MV
light, vine scrub, Monteith, Cook & Wright, Coll. no. 10257 (QM); 1 , Beerwah, 20.ix.1935, H. E. Young
(QDPI); 1 , Brisbane, (no abdomen) (QM); 1 , same locality as above but IIIidge (UQIC); 1 , Biarraville,
QLD, 20.i.1956, J. C. Le Sovef (ANIC); 1 , Carnarvon Range, 10.i.1939, N. Geary (AM). NSW: 1 ,
Corong (SAM); 1 , Kosciusko, i.–ii.1940, F. H. Taylor (AM); 1 , Ballina, 1.ii.1894, Froggatt (ASCU); 1 ,
Mellabulla, F. H. Taylor (AM); 1 , National Park, 16.x.1921, A. Musgrave, Coll. no. K44380 (AM); 1 ,
Botany, no abdomen (QM); 1 , W of Kyogle, Toonumbar Dam, 11–12.xii.1996, MV lamp, D. J. Scambler &
J. A. Macdonald (ASCU); 1 , 1420 m, Dawsons Spring, Mt Kaputar National Park, 9.xii.1987, at Mercury
Vapour lamp, G. R. Brown (ASCU); 1 , 1180m, Bark Hut Camping area, Mt. Kaputar National Park,
10.xii.1987, at mercury vapour lamp, G. R. Brown and S & B. Underwood (ASCU); 1 , Mittagong,
24.iii.1968, R. H. Mulder Coll.(AM); 1 , Mt Dromedary, 23.xi.1965, Britton & Upton (ANIC); 2 , 3 , Mt.
Kaputar, 3000 ft., 30.x.1976, at light, C. W. Frazier (ANIC); 1 , Faulconbridge Ridge, 12.xii.1980, S. G. Wat-
kins Collection (ANIC); 1 , N. Wendilla, vi.1908, out of Correa, Dixon (NMV). VIC: 1 , Grampians,
6.ii.1956, Delly's Dell (SAM); 1 , Grampians, 27.xii.1979, ex light trap, G. S. Taylor (WINC); 1 , Vic
Coast, Out of Banksia (WINC); 1 , 3 , Mordiallac, F. E. Wilson Coll (NMV); 1 , 2 , same locality as
above but, 15.x.1950, A. L. B. (NMV); 1 , same data as above but 12.x.1950 (NMV); 1 , Ferntree Gully,
8.xii.1927, A. H. Westly (UQIC); 1 , same locality as above but C. Oke (NMV); 1 , Moe, 27.x.1945, C. G.
L. Gooding (ANIC); 1 , 13 miles S of Kiata, Little Desert, 7.xi.1966, I. F. B. Common & M. S. Upton
(ANIC); 1 , Springvale, Vic, 1945, E. J. Harris (ANIC); 1 , 1 , 27 miles N of Yannac, Big Desert, Vic,
27.x.1973, H. Morton, C. F. Crasby (ANIC); 1 , Kiata, Vic, 1.xi.1945, M. W. Mules (NMV); 2 , Noble Park
, Vic, C. Oke (NMV); 1 , Cheltenham, C. Oke (NMV); 1 , Deep Creek, Braybrook District, J. E. Dixon
(NMV); 1 , Braybrook, 28.iix.1904, in Correa eranh, J. E. Dixon (NMV); 1 , Kororoit Creek (NMV); 2 ,
Victoria (NMV); 1 , Aspendale District, out of Olearia (Aster) ramculoba (NMV); 1 , 1 , Mel-
bourne,v.1901, J. J. Walker (HMO); 1 , Mulwala, D. ii.90 (NMV); 1 , VIC (UQIC). SA: 1 , Lake Surprise
Area, Simson Desert, 13.ix.1971, T. F. Houston (SAM); 1 , 1 , Black Swamp, 8.vii.1963, on flower of
grape tree palm, S. W. White (SAM); 1 , 1 , Grampian, Mt. Difficult Range, 2600 f, 7.ii.1956, N. B. Tindale
(SAM); 1 , 22 miles W by SW of Mundulla, 8.xi,1966, I. F. B. Common & M. S. Upton (ANIC); 1 , SA, F.
W. Ferguson Collection (ANIC). WA: 1 , Nedlands, 4.x.1955, M. M. H. Wallace (ANIC); 1 , 63 miles E of
Esperance (33º51'S, 121º53'E), Thomas River, 20.xi.1969, Taylor & M. S. Upton (ANIC).
Other material examined. 6 , 18 . Locality unknown: 1 , Blackburn (SAM); 1 , Coll. no.
C446 (QDPI); 2 , no data (NMV); 2 , no data (SAM); 1 , Milis Geppart, 26.xii.1926, J. Mas-
queem (AM); 1 , 6.xii.1966, D. A. Dooland Collection, NRPC (AM); 6 , no data (NMV); 3 , 4 ,
no locality data, 27.viii.1905, ex. Correa speciosa (NMV);1 ?, no abdomen, Correa ahecesa (NMV);
2 , N Mall, 20.x.1910, Bred by Dixon (NMV); 1 , bred from Helichrysum, Eltham District
(NMV).
Uracanthus maculatus, sp. nov.
(Figs 20, 59, 94)
Description
Male. Body length, 18.28–30.36 mm; width, 3.14–6.23 mm.
Colour (Fig. 20). Head, thorax, basal 2/5 and apical 1/10 of elytra dark reddish brown to blackish brown;
antennae, legs and the remaining part of elytra reddish brown. Frons and clypeus with mixed dense golden
and white pubescence; vertex with dense white pubescence except a V-shaped glabrous area extending from

Zootaxa 1569 © 2007 Magnolia Press · 13
REVISION OF URACANTHUS
medial frontal groove to posterior edge of vertex. Pronotum with fairly dense pale yellow pubescence; 2 lon-
gitudinal stripes of dense white pubescence on each side: 1 on disc and 1 near ventral side. Each elytron with
a subtriangular glabrous mark extending from shoulder to basal 1/3, and margined with a line of dense white
pubescence; some areas of the mark sometimes covered with sparse white pubescence; apical area with sparse
white pubescence margined with a narrow line of dense white pubescence; the remaining part of elytral disc
with pubescence of different density, giving an appearance of clustered denser pubescence among less dense
pubescence. The subtriangular mark on elytra may be reduced or expanded. The body may vary from dark
brown to brown. Longitudinal pubescent stripes on pronotal disc are sometimes absent from the posterior
margin.
Head. Postclypeus semicircular and slightly convex, with dense large coarse punctures; frontoclypeal
suture deep and narrow in middle; median frontal groove narrow and deep, terminating near posterior edge of
vertex; distance between lower lobes of eyes 1.7–1.75 × as long as distance between upper lobes of eyes; dis-
tance between upper lobes of eyes 2.3–2.38 × as long as distance between eyes on ventral side; genal length
0.34–0.36 × as long as head width immediately below eyes. Antennae slightly shorter than body; segments 5–
10 flattened and produced on one side at apex.
Thorax and abdomen. Pronotum 1.15–1.55 × as long as width, rounded with a very feeble process at each
side; posterior margin 1.10–1.33 × as wide as anterior margin; pronotal disc binodulose in middle, and rugose
transversely near anterior and posterior margins. Scutellum semicircular, with dense pubescence. Elytra 3.95–
4.48 × as long as prothorax and 3.38–4.07 × as long as shoulder width; each elytron with 3 very feeble longi-
tudinal carinae; basal elytra finely punctate; apex widely emarginate and bispinose. Abdomen slender, apex of
terminal sternite notched.
Male terminalia. Apex of ventral median lobe more or less rounded with a very shallow notch and apex of
dorsal lobe rounded; ventral lobe longer than dorsal lobe; spined region of internal sac 4.9–5.2 × as long as
basal unspined region; spined region divided into 2 sections with an unspined gap between sections: first sec-
tion bearing dense long simple spines with fairly dense multi-branched spines near sides; second section with
sparse simple spines (Fig. 59a). Eighth sternite (Fig. 59b) strongly obliquely truncate at sides, shallowly emar-
ginate at apex, with long and fairly long setae arising terminally; ventral surface with multi-branched spines
and cloud-like processes. Eighth tergite (Fig. 59c) emarginate at apex, with dense short simple spines and
multi- branched spines on dorsal surface. Paramere 1.75–1.84 × as long as wide, cylindrical in shape; apex
rounded with long and short setae (Fig. 59d).
Female. Body length, 15.99–29.39 mm; width, 2.93–6.03 mm.
Similar to male but differs in having shorter antennae and legs and more robust abdomen; apex of terminal
sternite rounded; elytra 3.16–3.95 × as long as shoulder width and 4.17–4.90 × as long as prothorax.
Ovipositor and spermatheca. Ovipositor short; styli arising terminally with short hairs (Fig. 59e). Sper-
matheca heavily curved; spermathecal gland long and arising near base (Fig. 59f).
Distribution
Southern, southeastern and western coastal regions of Australia and central Australia (Fig. 94).
Biology
Recorded host plants for this species are Acacia coriacea, Aster ramulosus, Cassinia sp., Citrus sp., Cor-
rea speciosa var mormalis, Eriostemon lanceolatus, Helichrysum ferrugineum and Olearia ramulosa. Adults
were collected from June to February.
Comments
This new species was described as a variant (Var. B) of U. triangularis by Lea (1916). However, after hav-
ing carefully examined all specimens available, we believe that it should be a new species. This new species

THONGPHAK & WANG
14 · Zootaxa 1569 © 2007 Magnolia Press
resembles U. triangularis but differs in having more uneven pubescence; the subtriangular mark and apical
areas on the elytra covered with pubescence; the apex of ventral lobe of aedeagus much longer than dorsal
lobe, apex of ventral lobe more or less pointed; apex of eighth tergite emarginate and ventral surface of eighth
sternite with simple spines.
Material examined
Holotype. . NSW: Galore, 26.xii.1893; bearing a red holotype label (VAIC).
Paratypes. 109 , 42 . All paratypes bear blue paratype labels. QLD: 1 , Stanthorpe, xii.1964, E. Sut-
ton, E. Sutton Coll. (AM); 1 , same locality as above but 18.xi.1925 (QDPI); 1 , same locality as above but
15.x.1927, J. Henderson (QDPI); 1 , Bunya Mt, 20.xi.1930, E. Sutton, E. Sutton Coll. (QM); 1 , 320m Isla
Gorge National Park (25°11’S, 149°58’E), 2.x.1992, MV lamp, G. Daniels (NMV); 1 , Toowoonan Mt,
viii.1963, R. Ridge (UQIC). 1 , Rokehurst, ID by McKeown (WAM); 1 , Toowoomba, QLD, 10.ii.1975, J.
Macqueen (ANIC); 1 , same data as above but 12.xi.1974 (ANIC); 1 , same data as above but 26.x.1974
(ANIC). NT: 1 , Plenty River Salt Lakes, Simpson Desert, 18.ix.1992, MV light, L. Archibald (NTM); 1 ,
Tennants Creek, xi.1949, F. A. (ANIC); 1 , nr Reedy Rock hole, Amdeus Basin (32º36'S, 149º35'E),
17.viii.1962, P. Ranford (ANIC). NSW: 1 , Mt Kaputar, Bull Creek, 27.xi.1984, G. Hangay (MAM); 2 ,
Rope's Creek (MAM); 1 , Kangaroo (SAM); 1 , Broken Hill, 22.x.1944, C. E. Chadwick (VAIC); 1 , Wil-
ton, 12.i.1966, V. J. Robinson (VAIC); 1 , Crackenback Range, 9.i.1968, Q. M. Williams (VAIC); 1 , Ash-
field, 6.xi.1980, D. A. Doolan, D. A. Doolan Collection (AM); 1 , Pilliga scrub (30º56'S, 149º23'E),
10.ii.1997, M. S & B. J. Moulds (AM); 1 , West Pymble, near Sydney, 26.xii.1986, at black light, D. J.
Scambler (AM); 1 , Murrumbidgee R., Angle Crossing, 22.xii.1966 (ANIC); 1 , 4 mi SW of Mudgee
(32º36'S, 149º35'E), 18.xi.1968, Britton & Misko (ANIC); 1 , Eyres, Coll. no. 1828, ID by Lea (AM); 1 ,
same data as above but Lea (MAM). ACT: 1 , Canberra, 1–5. xii.1999, at light, A. Szito (WADA); 1 ,
Kambah, Canberra, ACT, 30.xii.1978 (ANIC). VIC: 1 , 1 , Bendigo, 18.ix, 1917, ex Cassinia (NMV); 1 ,
Donvale, xii.1951, C. I. R. J. (ASCU); 1 , Frankston, 10.xi.1927 (NMV); 1 , Ferntree Gully, 18.xii.1931,
A. W. Westley (NMV); 2 , Gyyos Lands (WADA); 1 , Victoria (MAM); 1 , Blackwaterhole, 25.x.1971, J.
C. Le Sovef (ANIC); 1 , Dromana, 16.x.1971, J. C. Le Sovef (ANIC); 1 , 17 miles Orbost, 8.xii.1956, E. F.
Riek (ANIC); 1 , Maldon, xii.1921, J. C. Gondic (NMV); 1 , Mordialloc, 4.xii.1950, A. L. Brown (NMV);
1 , same locality as above but (NMV); 1 , same data as above but iv.1950 (NMV); 1 , 1 , Yinnar, Victo-
ria, i.1956, A. L. Brown (NMV); 1 , same data as above but 10.xi.1951 (NMV); 1 , same data as above but
13.xi.1955 (NMV); 1 , same data as above but iii.1955 (NMV); 1 , same data as above xii.1952 (NMV); 1
, Inglewood, 7.xi.1933, A. L. Brown (NMV); 1 , Mt. Evelyn, 7.xi.1933, A. L. Brown (NMV); 1 , Seaford,
ex Acacia (NMV); 1 , 4 miles N by NE of Nelson, Glenelg River, 25.xi.1966, Neboise (NMV); 1 , Launch-
ing Place, C. Oke (NMV); 1 , 1 , Frankston, C. Oke (NMV); 2 , Carrum, C. Oke (NMV); 1 , Healesville,
C. Oke (NMV); 1 , Beechworth, i.1914, C. Oke (NMV); 1 , Aspendale, larva feed in Olearia ramulosa
(NMV); 1 , Youyang, ex Aster, Coll. no. 250, dept by Lea (NMV); 1 , Koroite Creek, bred in Correa spe-
ciosa var mormalis (NMV); 1 , Tarago River, ii.1930, larva in wood, J. C. Gondic, bred from Helichrysum
ferrugineum (NMV); 2 , 2 ,Victoria (NMV); 1 , Murchison R., ix.1956, at light, A. Douglas (WAM).
TAS : 1 , LiffeyValley, 16.i.1985, S. Feari (ANIC); 1 , Erith Islands, Bass strait, i.1963, J. Murray & Smith
(NMV). SA: 1 , Murray R, H. S. Cope (SAM); 1 , Purmons near Mesrong R, S. W. Falton, 20.vi.1911
(NMV); 1 , 1 , Eyres, Petch (SAM); 1 , Yalana (SAM); 1 , 1 , Franklin Island, 20.iv.1980, C. Kalf
(SAM); 4 , same locality as above but 27.x.1983, C. Watts (SAM); 2 , Athelstone, 12.xii.1979, at light, J. J.
H. Szent & Ivancy (SAM); 1 , Lyntton, 3.xi.1970, UV light trap, R. Fisher (SAM); 1 , as above but xi.1970
(SAM); 1 , as above but xii.1970 (SAM); 1 , Murray Bridge, G. H. Dutton (SAM); 1 , 850 ft, Blackwood,
30.i.1965, MV light, N. McFarland (SAM); 1 , as above but 5.xi.1957 (SAM); 1 , 1 , Yorktown (SAM); 1
, Flinders Island, T. D. Campbell (AM); 1 , Wardang Island, 9.viii.1910, M. W. Mules, F. E. Wilson Collec-
tion (NMV); 1 , SA, by inference ex. Capt. White Collection (SAM); 1 , SA (SAM); 1 , Yeelanna det. by

Zootaxa 1569 © 2007 Magnolia Press · 15
REVISION OF URACANTHUS
Lea (ANIC); 6 ,1 , Reevesby Island, SA, xii.1936, J. Clark, McCoy Soc. Expect. Sir Jos. Banks Group. SA,
xii.1936–i.1937 (NMV); 1 , Blairgowire Park, 23.xi.1970, J. C. Le Sovef (ANIC); 1 , 1 , Giffordlands
(NMV); 1 , same locality as above but out of Helichrysum ferrugineum (NMV). WA: 1 , Dorre Island,
1959, A. Douglas (WAM); 1 , Kalbarri, 12.v.1965, R. Humphries (WAM); 1 , Dumbleyung, 27.ii.1962, H.
Udell (WAM); 1 , same locality as above but xii.1966 (WAM); 1 , 400 m, Lillian Stake Rock Frank Hann
National Park (33º04'064″S 120º05'827″E), 5.xi.1996, at light, Schun & Classic, Coll. no. 196–661 (AM); 1
, Pilachilpna Well, Pilbara dist., 7.xi.1953, N. B. Tindale (SAM); 5 , Barrow Island, 1.vi.1964, W. H. Butter
(WAM); 1 , 20 km N of Leeman (29°45'S, 114°58'E), Gum Tree Bay Ca., 7–12.x.1994, R. P. McMillan
(WAM); 1 , 1 , Theveanard Island, 8.iv.1966, G. Bostack (WAM); 1 , TVL, Sand area adjacent to main
camp, 25.v. 1990, 4.00 pm, Acacia coriacea, Dominant, M. R. White (WAM); 1 , Simaon Gums, Coll. no.
41–1036 (WAM); 1 , 1 , Carnac Island of perth, (32º07'S 115º39'E), 15.i.1983, T. F. Houson (WAM); 1 ,
Dudinin, Coll. no. 30–731 (WAM); 1 , Roy Hill, Coll. no. 53–1493 (WAM); 1 , WA, 5xi.1908, C. French’s
Coll. no. 265 (NMV); 2 , 11.xi.1918, C. French's Coll. (NMV). 1 , 101 km E of Esperance (33º51'S,
121º53'E), Thomas River, 20.xi,1969, Taylor & M. S. Upton (ANIC); 2 , 63 miles E of Esperance (33º51'S,
121º53'E), Thomas River, 20.xi,1969, Taylor & M. S. Upton (ANIC); 9 , 23 km NW by W of Mt. Arid, Tho-
mas River, 4–7.xi.1977, J. F. Lawrence (ANIC); 1 , 23 miles W of Fraser Range HS (32º04'S, 122º24'E),
7.xi,1969, M. S. Upton (ANIC); 1 , 7 miles E by N of Balladonia HS, 13.x,1968, Britton, Upton & Balderson
(ANIC); 1 , Deepdene, Karridale, 17.i,1967, M. S. Upton (ANIC); 1 , 13 miles NE by E of Caiguna,
14.x.1968, no abdomen, Britton, Upton & Balderson (ANIC); 2 , 6 miles N of Mandurah, 24.i.1967, M. S.
Upton (ANIC); 1 , 11 km N of Geraldton, 10.xii.1972, at MV light, N. McFarland (ANIC); 1 , Montebello
Island, 14.xi.1953, T. G. Campbell (ANIC).
Other material examined. 11 , 18 . Locality unknown: 1 , Aust old Coll., Coll. no. k40693 (AM); 1
, same data as above but (AM); 1 , C. French (QDPI); 1 , 20.iv.1922, H. J. Carter collection (NMV); 1 ,
no data; 1 , Coll. no. K10943 (AM); 1 , Longicorn of mandarin tree, L. Gallard Collection (AM); 1 , L. C.
Haines Collection (AM); 1 , SA & WA (SAM); 1 , no locality label, 12.v.1982 (SAM); 1 , Victorian Mar-
gas (French) (AM); 1 , Coll. no. 34667, Roltnest (WAM); 1 , Colleslac, Coll. no. 45-872 (WAM); 1 , Coll.
no. 48-919, W. Henoom (WAM); 1 , Rose Beev(?), 3.vi.1942, found in Eriostemon lanceolatus roots, Frog-
gatt (ANIC); 1 , no data, ex C. French Coll. (HMO); 1 ,1 , no data under Hope/Westwood Coll. (HMO); 1
, no locality data, 5.xi.1908, C. French Collection (NMV); 1 , no locality data, x.907, ex Aster ramulosus
(NMV); 1 , no locality data, xi, 1908, ex Aster ramulosus (NMV); 2 , no locality data, xii, 1908, ex Aster
ramulosus (NMV); 1 , xii, 1907, ex Aster ramulosus (NMV); 1 , no data (NMV); 1 , no locality, ix.1918,
Coll. no. 597 (NMV); 1 , no data Coll no. 40 (NMV); 1 , no data, Coll. no. 07 (NMV); 1 , 18.ix.1917, ex
Aster sp. (NMV); 1 ?, no data, Coll. no. 350, no abdomen (NMV).
Uracanthus perthensis, sp. nov.
(Figs 21, 60, 94)
Description
Male. Body length, 17.57–21.61 mm; width, 3.23–4.84 mm.
Colour (Fig. 21). Head, thorax, basal elytra dark reddish brown; antennae, legs and the remaining part of
elytra reddish brown. Head (except a longitudinal glabrous area on vertex), legs, and abdomen with fairly
dense white pubescence. Prothorax covered with short white pubescence except glabrous median longitudinal
line and 2 raised nodules on disc; pronotum with 2 vague longitudinal stripes of dense white pubescence on
each side: 1 on disc and 1 near ventral side. Elytra covered with fairly dense short white pubescence; each
elytron with a large subtriangular glabrous mark, extending from shoulder to basal 1/3, margined with a very
vague line of dense white pubescence; apical areas with relatively sparse short white pubescence.

THONGPHAK & WANG
16 · Zootaxa 1569 © 2007 Magnolia Press
Head. Postclypeus semicircular and slightly convex, with dense coarse punctures; frontoclypeal suture
deep and narrow in middle; distance between lower lobes of eyes 1.82–1.83 × as long as distance between
upper lobes of eyes; distance between upper lobes of eyes 1.1–1.2 × as long as distance between eyes on ven-
tral side; genal length 0.48–0.55 × as long as head width immediately below eyes.
Thorax and abdomen. Pronotum 1.11–1.20 × as long as width; rounded and with a feeble process at each
side; posterior margin 1.25–1.36 × as wide as anterior margin; pronotal disc binodulose in middle, and rugose
transversely near anterior and posterior margins. Scutellum semicircular, with dense pubescence. Elytra 4.71–
4.72 × as long as prothorax and 3.41–3.69 × as long as shoulder width; each elytron with 3 feeble longitudinal
carinae; basal elytra finely punctate and smooth; apex widely emarginate and bispinose, sutural spine longer
and more acute than marginal one. Apex of terminal sternite truncate.
Male terminalia. Apex of ventral median lobe more or less truncate with a very shallow notch and apex of
dorsal lobe rounded; ventral lobe much longer than dorsal lobe; spined region of internal sac 4.98–5.12 × as
long as basal unspined region; spined region divided into 2 sections with an unspined gap between sections:
first section much longer than unspined gap and second section, with dense long simple spines; second sec-
tion with sparse simple spines (Fig. 60a). Eighth sternite obliquely truncate at sides, shallowly and widely
emarginate at apex, with long and fairly long setae arising terminally (Fig. 60b); ventral surface with very
sparse multi-branched spines and fairly dense cloud-like processes. Eighth tergite slightly emarginate (almost
truncate) at apex (Fig. 60c), with fairly dense short simple spines and multi-branched spines on dorsal surface.
Paramere long, 1.86–2.04 × as long as wide, cylindrical in shape, apex rounded with long and short setae (Fig.
60d).
Female. Unknown.
Distribution
Western Western Australia (Fig. 94)
Biology
Hosts are unknown. Adults were collected in March.
Comments
This new species was treated as Var. C of U. triangularis by Lea (1916). It resembles U. maculatus, sp.
nov., but differs in having the pubescence on the elytral disc more or less even; elytral apex more deeply emar-
ginate, and sutural apical spine longer.
Material examined
Holotype. . WA: Perth; bearing a red holotype label (NMV).
Paratype. 1 . WA: Perth, iii.1916; bearing a blue paratype label (WAM).
Uracanthus griseus, sp. nov.
(Figs 22, 61, 94)
Description
Male. Body length, 23.51–27.95 mm; width, 4.69–5.86 mm.
Colour (Fig. 22). Head, thorax, legs, antennae, and base of elytra blackish brown; abdomen and remaining
part of elytra reddish brown. Head with dense white pubescence except median frontal groove and a longitudi-
nal glabrous area on vertex. Pronotum with 2 longitudinal stripes of dense white pubescence on each side: 1
narrow on disc and 1 wide near ventral side; remaining part of pronotum covered with short white pubes-

Zootaxa 1569 © 2007 Magnolia Press · 17
REVISION OF URACANTHUS
cence, pubescence on middle disc sparser. Each elytron with a blackish, subtriangular, more or less glabous
mark, extending from shoulder to basal 1/3, margined with a very vague line of white pubescence; anterior
region of mark usually covered with sparse short white pubescence; remaining part of elytral disc with fairly
dense mixture of short white and pale golden pubescence; apical area with relatively sparse white pubescence.
Meso- and metasterna, abdomen, and legs covered with dense white pubescence.
Head. Postclypeus semicircular and slightly convex, with dense coarse punctures; frontoclypeal suture
deep and wide in middle; distance between lower lobes of eyes 1.52–1.82 × as long as distance between upper
lobes of eyes; distance between upper lobes of eyes 1.06–1.10 × as long as distance between eyes on ventral
side; genal length 0.43–0.55 × as long as head width immediately below eyes. Antennae slightly longer than
body; segments 4–10 slightly flattened and produced on one side at apex; apical ¼ of segment 11 distinctly
thinner than basal ¾.
Thorax and abdomen. Pronotum 1.15–1.29 × as long as width, with a distinct process at each side; poste-
rior margin 1.11–1.19 × as wide as anterior margin; pronotal disc binodulose in middle and rugose trans-
versely near anterior and posterior margin. Scutellum semicircular, with dense pubescence. Elytra 4.09–4.63 ×
as long as prothorax and 3.35–3.71 × as long as shoulder width; each elytron with 3 feeble longitudinal cari-
nae; basal elytra finely punctate and smooth; apex widely emarginated and bispinose. Apex of terminal stern-
ite truncate.
Male terminalia. Apex of ventral median lobe distinctly emarginate and apex of dorsal lobe pointed; ven-
tral lobe longer than dorsal lobe; spined region of internal sac 5.12–5.23 × as long as basal unspined region;
spined region divided into 2 sections with an unspined gap between sections: first section with dense long
simple spines; second section with sparse simple spines (Fig. 61a). Eighth sternite strongly obliquely truncate
at sides, strongly emarginate at apex, with long and fairly long setae arising terminally; ventral surface with
cloud-like processes (Fig. 61b). Eighth tergite rounded to slightly pointed at apex, with fairly dense short sim-
ple spines and multi-branched spines on dorsal surface (Fig. 61c). Paramere long, 1.86–2.04 × as long as
wide, cylindrical in shape, apex rounded with long and short setae (Fig. 61d).
Female. Unknown.
Distribution
Northern Victoria and southern South Australia (Fig. 94).
Biology
Hosts are unknown. Adults were collected in October and November.
Comments
This new species closely resembles U. perthensis sp. nov. but differs in having darker body colour; apical
spines of elytra shorter and thicker.
Material examined
Holotype. . SA: Marray River, H. S. Cape; bearing a red holotype label (SAM).
Paratypes. 6 . All paratypes bear blue paratype labels. VIC: 5 , 7.3 km SW of Wemen, 25.x–3.xi.1988,
T. Weir, J. Lawrence & M. Hansen (ANIC); 1 , Yanacy, 5xi.1978, J. C. Le Sovef (ANIC).
Uracanthus glabrilineatus Lea, 1917a
(Figs 23, 94)
Uracanthus glabrilineatus Lea, 1917a: 737.—McKeown, 1947: 64.

THONGPHAK & WANG
18 · Zootaxa 1569 © 2007 Magnolia Press
Description
Male. Body length, 25.35 mm; width, 5.48 mm.
Colour (Fig. 23). Head, thorax, femora, and basal elytra dark reddish brown; rest of elytra reddish brown.
Head (particularly frons and vertex) with dense pale yellowish pubescence except median frontal groove and
a longitudinal glabrous area on vertex. Pronotal disc with fairly dense pale yellowish pubescence without dis-
tinct longitudinal pubescent stripes; ventral side of prothorax with very dense pale yellowish pubescence.
Each elytron with a vague subtriangular glabrous mark, extending from shoulder to basal 1/3, without a dis-
tinct margin; some part of mark covered with sparse pubescence; disc of each elytron with 4 longitudinal lines
of dense pale yellowish pubescence interspaced with glabrous longitudinal carinae. Meso- and metasterna,
abdomen, and legs covered with fairly dense white pubescence.
Head. Postclypeus semicircular and slightly convex, with dense coarse punctures; frontoclypeal suture
deep and narrow in middle; distance between lower lobes of eyes 1.55 × as long as distance between upper
lobes of eyes; distance between upper lobes of eyes 1.42 × as long as distance between eyes on ventral side;
genal length 0.40 × as long as head width immediately below eyes. Antennae slightly shorter than or as long
as body; segments 4–10 flattened and produced on one side at apex; apical ¼ of segment 11 suddenly becom-
ing thinner.
Thorax and abdomen. Pronotum 1.35 × as long as width, with an angular process at each side; posterior
margin 1.16 × as wide as anterior margin; pronotal disc binodulose in middle and rugose transversely near
anterior and posterior margins. Scutellum semicircular, with dense pubescence. Elytra 3.53 × as long as pro-
thorax and 3.32 × as long as shoulder width; each elytron with 3 feeble longitudinal carinae; basal elytra mod-
erately punctate; apex narrowly emarginate and bispinose, sutural spine longer than marginal one. Abdomen
slender; apex of terminal sternite truncate.
Male terminalia. Unknown. The unique male specimen (holotype) has not been dissected.
Female. Unknown
Distribution
Southwestern Western Australia (Fig. 94).
Biology
Unknown
Comments
This species resembles U. perthensis sp.nov., but differs in having elytra with glabrous longitudinal lines;
pronotal disc without distinct longitudinal stripes of dense pubescence.
Material examined
Holotype. . WA: Mullewa, 18.v.1823, Miss J. F. May, bearing a name label on which ‘TYPE’ was writ-
ten in red (SAM).
Uracanthus triangularis Hope, 1833
(Figs 24, 62, 95)
Uracanthus triangularis Hope, 1833: 64.—Hope, 1834: 108; Lacordaire, 1869: 391; Best, 1882: 35; Tepper, 1887: 38;
Froggatt, 1893: 29, 1902: 710, 1923: 112; Gahan, 1893: 182; French, 1900: 135; Gurney, 1911: 58; Aurivillius,
1912: 147; Gallard, 1916: 113; Tillyard, 1926: 223; McKeown, 1947: 66; Duffy, 1963:112.
Stenochorus angustatus Boisduval, 1835: 475.—Gahan, 1893: 182 (synonymy).
Mallocera angustata.–Laporte, 1840: 425.

Zootaxa 1569 © 2007 Magnolia Press · 19
REVISION OF URACANTHUS
Description
Male. Body length, 21.68–33.13 mm; width, 4.24–6.47 mm.
Colour (Fig. 24). Body reddish brown to blackish brown with head and thorax darker. Head with dense
white or yellowish pubescence except median frontal groove and a longitudinal glabrous area on vertex.
Pronotum with 2 longitudinal stripes of white pubescence on each side: 1 on disc, vague, and 1 near ventral
side, distinct. Each elytron with a large clearly defined subtriangular glabrous mark, extending from shoulder
to basal 1/3, clearly margined with dense white pubescence; apical area glabrous, margined with a distinct line
of dense white pubescence; remaining part of elytral disc with dense and fairly long pubescence. Body colour
varies from reddish brown to blackish brown. The glabrous mark on the elytra may be reduced or expanded
from the basic pattern.
Head. Postclypeus semicircular and slightly convex, with dense coarse punctures; distance between lower
lobes of eyes 1.5–1.85 × as long as distance between upper lobes of eyes; distance between upper lobes of
eyes 1.0–1.25 × as long as distance between eyes on ventral side; genal length 0.35–0. 44 × as long as head
width immediately below eyes. Antennae about as long as body; segments 5–10 flattened and produced on
one side at apex; apical ¼ of segment 11 slightly thinner than basal ¾.
Thorax and abdomen. Pronotum 1.14–1.46 × as long as width, rounded at side; posterior margin 1.07–
1.36 × as wide as anterior margin; pronotal disc binodulose in middle and rugose transversely throughout.
Scutellum semicircular, with sparse pubescence. Elytra 4.1–5.07 × as long as prothorax and 3.7–4.15 × as
long as shoulder width; each elytron with 3 very feeble carinae; basal half of elytra with sparse coarse punc-
tures; apex widely emarginate and bispinose. Abdomen slender; apex of terminal sternite truncate or slightly
rounded.
Male terminalia. Apex of ventral median lobe narrowly emarginate, and apex of dorsal lobe rounded; ven-
tral lobe slightly longer than or as long as dorsal lobe. spined region of internal sac about 2.1–2.5 × as long as
basal unspined regions; spined region divided into 2 sections with an unspined gap between sections; first sec-
tion longer than unspined gap with dense long simple spines; second section with sparse short simple spines
(Fig. 62a). Eighth sternite obliquely truncate at sides, shallowly emarginate at apex, with long and fairly long
setae arising terminally; ventral surface with cloud-like processes (Fig. 62b). Eighth tergite truncate or shal-
lowly emarginate at apex, with fairly dense short simple spines and multi-branched spines on dorsal surface
(Fig. 62c). Paramere 1.5–1.67 × as long as wide, cylindrical in shape, apex rounded with long and short setae
(Fig. 62d).
Female. Body length, 17.17–34.19 mm; width, 3.05–6.97 mm.
Similar to males, but differs in having antennae and legs distinctly shorter, more robust abdomen; elytra
slightly longer and wider, 3.75–4.4 × as long as shoulder width and 4.09–5.13 × as long as prothorax. Apex of
terminal sternite rounded.
Ovipositor and spermatheca. Ovipositor short; styli arising terminally with short hairs (Fig. 62e). Sper-
matheca clearly curved; spermathecal gland long and arising at base (Fig. 62f).
Distribution
Northern, eastern, and southwestern coasts of Australia (Fig. 95).
Biology
The following were recorded as the host plants of this species: Acacia decurrens, A. mearnsii [=A. mollis-
sima], A. penninervis, A. pycnantha, A. sophorae [= A. longifolia var. sophorae], Acacia spp., Boronia pin-
nata, Eriostemon australasius [=E. lanceolatus], Banksia ericifolia; B. integrifolia, Banksia spp., Hakea
gibbosa, H. nodosa, H. serice and Lomatia longifolia. Adults were found on Leptospermum flavescens and
Angophora sp. flowers and Kunzea ambigua and Xanthorrhoea sp. foliage. Eggs are deposited in small
branches of the host trees. Adults emerge from September to December, although eclosion occurs as early as
June. They were collected from October to April. Adults are attracted to MV light.

THONGPHAK & WANG
20 · Zootaxa 1569 © 2007 Magnolia Press
Comments
This species closely resembles U. pallens but differs in having the pronotal disc rugose transversely
throughout; elytral apex more widely emarginate and more strongly and acutely spinose, and the subtriangular
marks on elytra broader.
Material examined
Holotype. . New Holland (= Australia); no data; type coll. 1767; bearing a red type label with Hope’s
hand writing (HMO).
Other material examined. 114 , 118 . QLD: 1 , 1 , Stanthorpe (28°39'S, 151°56'E), i.1961, C. Watts
(SAM); 1 , same locality but Sivian (AM); 1 , same locality but Von Weildt (QM); 1 , same locality but
Dr. K. K. Spence (AM); 2 , same locality but 30.iii.1963, P. Kerridge (UQIC); 1 , same locality but
2.xi.1927, S. M. Watson (QDPI); 3 , same locality but xii.1964, E. Sutton Coll. (QM); 1 , as above but
2.iv.1926 (QM); 1 , 70 km SW of Greenvale (19°00'S, 144°58'E), 16–28.i.1995, A. J. Watts (SAM); 1 ,
Bowarrady Creek (25°07'S, 153°08'E), Fraser Island, 5.x.1989, L. D. Buddle (AM); 2 , Kuranda (16°49'S,
145°38'E), F. P. Dodd (SAM); 2 , Moreton Island (27°11'S, 153°24'E), Ben Ewa Campground, 22.iii.1998, at
light, J. & A. Skevington (UQIC); 1 , Oranmore Range, 5.iii.1958, A. R. Briay (QDPI); 1 , Archer River X-
ring (13°25'S, 142°56'E), 11.v.1989, MV lamp, G. & A. Daniels (UQIC); 1 , 320 m of Isla Gorge National
Park (25°11'S, 149°58'E), 3.x.1992, MV lamp, G. Daniels (UQIC); 1 , 26 km W of Fairview (15°35'S,
144°04'E), 24.v.1989, MV lamp, G. & A. Daniels (UQIC); 1 , 1 , 10 km N of Caboolture (27°04'S,
152°58'E), Elimbah, 11.iv.1984, C. E. Hagan (UQIC); 1 , Eclipse Island, Coll. no. 39-1785 (WAM); 1 ,
QLD, 1.viii.1964, A. W. Asjary (UQIC); 1 , Wyberba, 5–7.xi.1969, at light, E. C. Dahms (QM); 3 , 1 ,
Cooloola Forestry Area, camp milo, iv.1978, at light, I. D. Galloway & E. Gympie (QDPI); 1 , 1 , same
locality but Banksia dom, 3.iii.1970, open forest to light, E. Dahms (QM); 2 , Glen Aplin, SQ,1946, S. R. E.
Brock Collection (ANIC); 3 , same locality as above but 1957, A. Gemmell, S. R. E. Brock Collection
(ANIC); 2 , same locality as above but 1912–1954, A. Gemmell, G. G. L. Gooding Collection (ANIC); 2 , 2
km N by NW Jawalbinna (15º45'S, 144º15'E), 17.i.1994, at light, P. Zborowski & E. D. Edwards (ANIC); 1 ,
4 km W by S of Cooktown, 21.v.1977, I. F. B. Common & E. D. Edwards (ANIC); 1 , 24 km E by N of
Ravenshoe, Palmerston Nation Park (17º35'S, 145º43'E), 17.xi.1981, J. Balderson (ANIC); 1 , South Dun-
more SF (27º38'S, 150º58'E), 26.xii.1998, at MV light, S. G. Watkins Collection (ANIC); 1 , 13 km N by
NW of Longreach (23º135'S, 144º04'E), Darr River, 7.iv.1976, J. F. Feehan (ANIC); 1 , Toowoomba, i.1980,
C. Brook (NMV). NT: 2 , Daly Waters (16°16'S, 133°22'E), 20.i.1971, T. Weir & A. Allwood (NTM);1 ,
October Creek (16°28'S, 135°09'E), 100 m E of Dally Water, 21.iii.1972, on Borroloola Rd, N. W. Forester
(NTM); 1 , NT, Cattle Creek (16º32' S, 136º10'E), 54 km S byW of Borroloola, 27.x.1975, M. S. Upton
(ANIC). NSW: 1 ,Engadine (34º 03' S, 151º 01' E ), 10.i.1974, R. H. Mulder Collection (AM); 1 , as above
but 14.xii.1974 (AM); 1 , as above but 17.xii.1958, H. Hughes (AM); 1 , as above but 10.i.1958 (AM); 1 ,
as above but 14.xii.1974 (AM); 1 , as above but 28.xi.1968 (AM); 1 , as above but 14.xii.1974 (AM); 1 ,
as above but 10.i.1970 (AM); 1 , as above but 6.iii.1955 (AM); 1 , as above but 12.x.1960, (AM); 2 , 1 ,
5mi, S. Mendooran (31º 47'S, 149º 17'E), 19.ii.1972, MV lamp, G. Daniels (AM); 2 , same locality but
17.ii.1972 (AM); 2 , 1 , Helensburgh (34º11'S, 150º58'E), xi.1961, H. Osborne (ASCU); 1 , same data but
xii.1961 (ASCU); 1 , same data but xii.1962 (ASCU); 1 , same data but xii.1963 (ASCU); 1 , West Pym-
ble (33º45'S, 151º07'E), near Sydney, 6.iii.1986, D. J. Scambler (AM); 1 , Goonoo State Forest (32º04' S,
148º54'E), 5 mi S. Mendooran, 24.iv.1974, D. K. McAlpine (AM); 1 , same data but 23.iv.1974 (AM); 1 ,
Budthingeroo Ceek (33º53'S, 150º00'E), Kanangra Boyd National Park, 24.xii.1977, G. Daniels (AM); 1 ,
Kuringgai Chase National Park (33º36'S, 151º12'E), 16.x.1971, MV lamp, G. Daniels (AM); 1 , same local-
ity, N. C. Llayd (ASCU); 1 , same locality but 5.iv.1973, A. B. Rose (MAM); 2 , Newcastle (33º55'S,
151º00'E), i.1988, Coll.no. K36095 (AM); 1 , Round Hill Reserve (33º00'S, 146º10'E), 23–29.iv.1968, R.
Lossin (AM); 2 , 2 , Sydney (33º51'S, 151º12'E ) (MAM); 1 , 1 , same locality but (SAM); 1 , same

Zootaxa 1569 © 2007 Magnolia Press · 21
REVISION OF URACANTHUS
locality but xii.1938 (no abdomen) (MAM); 1 , same locality (SAM); 1 , same locality but W. du Boulay,
E. F. du Boulay Collection (SAM); 1 , same locality but 9.iii.1907, W. B. G (ASCU); 1 , same locality but
Dean (UQIC); 1 , same locality but Coll. no. K36095 (AM); 1 , same locality but xi.1911, C. Gibbons
(AM); 1 , 1 , Blue Mountains National Park (33º36'S, 150º25'E), 15.xii.1984, on Leptospermum, G. R.
Brown (ASCU); 1 , Blue Mountains National Park (33º44'S, 150º30'E), 1.xii.1947 (SAM); 1 , Billy Blue
Mountain Nation Park (34º02'S, 149º27'E), 30.i.1980, N. W. Rodd (AM); 1 , Mt. Tomah (33º32' S,
150º25'E), 22.xi.1980, N. W. Rodd (AM); 1 , 1 , Mt. Wilson (33º59'S, 149º21'E), 27.xi.1980, D. A. Doolan,
D. A. Doolan Collection (AM); 1 , same locality but 6.xi.1923, Carno rear, A. Musgrave & T. Carepbull
(AM); 1 , National Park, 16.x.1921, A. Musgrave, Coll. no. K 44380 (AM); 1 , 1.5 mile E. of Freshwater
River, Iluka district, 17.i.1971, D. K. McAlpine & A. Hughes (AM); 1 , Mosman (33º13'S, 150º50'E ),1923,
E. Balton, Coll. no. K47516 (AM); 1 , Lane Cove (33°48'S, 151°10'E), 12.xii.1943, N. W. Rodd (AM); 1 ,
same as above but 7.iv.1973, G. R. Brown (ASCU); 1 , Dorrigo (30°19'S, 152°43'E), 7.x.1973, D. A. Doolan,
D. A. Doolan Collection (AM); 1 , North Ryde (33°47'S, 151°07'E), 17.i.1967, D. A. Doolan, D. A. Doolan
Collection (AM); 1 , Bobbin Head, 23.iv.1969, K. Ayers (MAM); 1 , Wahroonga, 8.xii.1969, A. B. Rose
(MAM); 1 , Oranmore, Hokas, 9.xii.1968, S. Barker (SAM); 1 , near Rivertree (28°37'S, 152°18'E), 15–
19.xii.1969, E. Dahms (QM); 1 , Bardwell Park (33°56'S, 151°08'E), 21.i.1943, C. E. Chadwick (ASCU); 1
, Wahroonga, 26.xi.1952, R. Dobson (NMV); 1 , same data as above (NMV); 1 , Sublime land,
15.xi.1902 (ASCU); 1 , Hornsby, 3.iii.1978, A. Beattic (ASCU); 1 , Wilton, 12.i.1966, V. J. Robinson
(ASCU);1 , as above (ASCU); 1 , Good Friday Mountain, 7.i.1968, on Fire trail, V. J. Robinson (ASCU); 1
, London Falls, 12.xii.1948, on Leptospermum flavescens flowers, C. E. Chadwick (ASCU); 1 , Botany,
13.xi.1994, Damagein (ASCU); 2 , 1 , Gibraltar Range (29°33'S, 152°17'E), 3,000 ft. via Glen Innes, 27–
29.xii.1972, open forest, G. B. Monteith (UQIC); 1 , Bundjalung, National Park near Iluka, 12.iv.1981, M. J.
Fletcher & G. R. Brown (ASCU); 1 , Canberra, 29.ii.1967, light trap, BCP (WINC); 1 , 8 km S of Mendoo-
ran (31°50'S, 149°06'E), 28.i.1987, G. & A. Daniels (UQIC); 1 , NSW (SAM); 1 , as above (WADA); 1 ,
same locality but I. D by lea (WADA); 1 , same locality (MAM); 1 , same locality (MAM); 1 , same
locality but 26.xi.1952, Nat Mus, R. Dobson (NMV); 1 , same locality but 15.xii.1977, Rear Beach, A. D.
Austin (WINC); 1 , Hornby, xi.1910, G. Gibbons (AM); 1 , Vaucluse, 19.ii.1920, J.M. , Coll. no. K63681
(MAM); 1 , North Sydney, 11.viii.1946, P. Hadlington (FCNI); 1 , Lucas Heights, Sydney, NSW,
23.xi.1979, on Angophora bloom, D. C. Carne (ANIC); 1 , Sydney, NSW, R. W. Ferguson Collection
(ANIC); 1 , Ebenezer, Sydney, 5.iii.1987, MV light, J. C. Keast (FCNI); 1 , Bonnet Bay, Sydney,
28.ii.1987, at light, L. S. William (FCNI); 1 , Bargo, 8.xii.1987, adult on Kunzea ambigua foliage, G. A.
Webb (FCNI); 1 , Tubbamurra Creek, Barrington Tops, 4100 ft, 11.i.1956, K. D. Fairrey (FCNI); 1 , Heath-
cote, 10.xii.1979, S. Watkins, S. G. Watkins Collection (ANIC); 1 , same data as above but 2.xii.1978
(ANIC); 1 , same data as above but 4.xii.1977 (ANIC); 2 , 3 km N of Mt. Victoria,, 22.xii.1978, on Xanth-
orrhoea, S. G. Watkins Collection (ANIC); 1 , Mt. York, NSW, 30.xi.1977, S. Watkins, S. G. Watkins Collec-
tion (ANIC); 1 , same data as above but 28.xi.1979 (ANIC); 1 , 50 miles S of Singleton, 5.i.1956, I. F. B.
Common (ANIC); 2 , 20 miles NW of Upper Colo, 8.xi.1955, T. G. Campbell (ANIC); 1 , Mt Boyce,
16.xii.1972, S. Watkins, S. G. Watkins Collection (ANIC); 1 , 9 km N by NE of Coonabarabran, 24.x.1980,
E. Britton (ANIC); 1 , Berkshire Park, 19.xi.1972, S. Watkins, S. G. Watkins Collection (ANIC); 1 , Hart-
ley Vale, 27.xi.1977, S. Watkins, S. G. Watkins Collection (ANIC); 1 , Waterfall, NSW, 26.xii.1974, S. Wat-
kins, S. G. Watkins Collection (ANIC); 1 , Kiata, 8.xii.1964, N. Dobrotworsky (ANIC); 1 , Wahroonga, H.
J. Carter (ANIC); 1 , Toganmain, 4.x.1966, I. Rawley (ANIC); 1 , 1 , National Park, xi.1952, J. G. Brook
(ANIC); 1 , NSW, H. E. Cox Collection, present 1916 by Mrs Cox (HMO);1 , Richmond R., New Holland,
Miers Collection, presented 1880 by W. Miers (HMO); 1 , Glebe Park, 11.iv.1936, E. Sutton (QM); 1 , Mit-
tagong, 20.iv.1922, H. J. Carter Coll. (NMV); 1 , Deewhy, ii.1928 (ANIC). ACT: 2 , 3 , Black Mountain,
14.ii.1966, at light trap, P. B. Carne (ANIC); 2 , 2 , same locality as above but xi.1965–ii.1966, E. B. Brit-
ton (ANIC); 1 , same locality as above but 28.ii.1969, at light trap (ANIC); 1 , same locality as above but

THONGPHAK & WANG
22 · Zootaxa 1569 © 2007 Magnolia Press
25–26.xii.1964, I. F. B. Common (ANIC); 1 , same locality as above but 16.ii.1968, M. S. Upton (ANIC); 1
, same locality as above but 15.ii.1936, A. D. Butcher (NMV); 1 , same data as above but 27.i.1936
(NMV); 2 , Lees Creek near Picadilly circus, 8.ii.1984, Calder & Stevens (ANIC). VIC: 1 , 3 km S.
Murchison, 8.i.1979, N. W. Rodd (AM); 1 : Mt. Difficult (37°00'S, 142°26'E), Grampians, 2600 ft.,
7.ii.1956, N. B. Tindale (SAM); 1 , Melbourne, 9.xi.1932, F. E. Wilson Collection (NMV); 2 , 10 miles N
of Wingan Inlet, 3.xi.1969, I. F. B. Common (ANIC); 1 , 17 miles NW Orbost , 8.xii.1956, E. F. Riek
(ANIC); 1 , Beechworth, C. Oke (NMV); 1 , Abbeyard, 27.ix.1960 (NMV); 1 , Mordialloc (no abdomen)
(NMV); 1 , Victoria (WADA); 1 , Victoria (AM). WA: 1 , Denmark (34°58'S, 117°21'E), i.1997, M. & L.
Jone (WAM); 1 , same data but i.1998 (WAM); 1 , same locality but 24.xi.1999, R. P. McMillan (WAM); 1
, Haker sterm, Turle Rock, Muckenburra, 16.x.1979, R. P. McMillan (WAM); 1 , Esperance (33°51'S,
121°53'E), 26.ix.1970, A. V. Thomas (WAM); 1 , Wurarga (28°24'S, 116°17'E), Coll. no. 37-4439 (WAM); 1
, Karridale (34°12'S, 115°06'E), 18.xii.1964, L. M. Olkalloran (QDPI); 1 , WA (WADA). Locality
Unknown: 1 , E. C. Vallis Bequest, Don. vii.1965 (QM); 1 , National Park, T. Mec. 13.x.1911 (ASCU); 1
, Nation Pk., T. Mec. 13.x.1911 (ASCU); 1 , R. R. McMillan Collection (WAM); 1 , Coll. no. 2926
(SAM); 1 , 10.ii.1920, Coll. no. K63681 (AM); 1 , no data (NMV); 1 , Relton Bequest, Coll. no. 3455
(QM); 1 , no data ( ASCTO); 1 , no data (WADA); 1 , 11.x.1915, bred from Lomatia longifolia, J. C.
Dixon (NMV); 1 , A. H. Elston collection (AM); 1 , 29.iii.1967, D. A. Doolan, D. A. Doolan Collection
(AM); 1 , N. RPC, 17.iv.1967, D. A. Doolan Collection (AM); 1 , 1 , K. G. Sound (MAM); 1 ?, no data
(no abdomen) (SAM); 1 , Deewhy, ii.1928 (ANIC); 1 , ? Park Coll, 1924, WWF (ANIC); 1 , no locality
label, S. R. E. Brock Collection (ANIC); 1 , ix.1907, ex Acacia mollissima (NMV); 1 , Slud Park (?), Aca-
cia mollissima, x.1907 (NMV); 1 , i.1908, Acacia (NMV); 2 , ix.1913, Acacia (NMV); 1 , xii., Acacia
(NMV); 1 , out of black wattle (Acacia mearnsii), S. Park (WINC); 1 , same as above but (NMV); 1 , 2 ,
no label (NMV).
Uracanthus pallens Hope, 1841
(Figs 25, 63, 96)
Uracanthus pallens Hope, 1841a: 53.—Hope, 1841b: 65, 1844: 198; Aurivillius, 1912:147;
Lea, 1916: 385; McKeown, 1947: 65; Duffy, 1963: 115.
Description
Male. Body length, 19.21–26.27 mm; width, 3.73–4.55 mm.
Colour (Fig. 25). Body, reddish brown with head, thorax, femora, and basal elytra darker. Head with
dense golden pubescence, except median frontal groove and a glabrous area on vertex. Pronotum with 2 dis-
tinct longitudinal stripes of white pubescence on each side: 1 on disc and 1 near ventral side; the remaining
pronotal disc with golden pubescence. Each elytron with a narrow, long, subtriangular glabrous mark, starting
from shoulder and ending at almost ½ of elytra, margined with a distinct line of dense white pubescence; api-
cal area glabrous, margined with a distinct line of dense white pubescence; remaining elytral disc with fairly
dense golden pubescence. The glabrous marks on elytra may be reduced or expanded to some extent.
Head. Postclypeus triangular and convex, with dense coarse punctures; frontoclypeal suture deep and
wide in middle; distance between lower lobes of eyes 1.69–1.88 × as long as distance between upper lobes of
eyes; distance between upper lobes of eyes 1.07–1.28 × as long as distance between eyes on ventral side;
genal length 0.31–0.48 × as long as head width immediately below eyes. Antennae usually slightly shorter
than body; segments 4–10 flattened and produced on one side at apex; apical ¼ of segment 11 slightly thinner
than basal ¾.
Thorax and abdomen. Pronotum 1.15–1.42 × as long as width, rounded at sides; posterior margin 1.09–
1.15 × as wide as anterior margin; disc binodulose in middle and rugose transversely anteriorly and and 1/3

Zootaxa 1569 © 2007 Magnolia Press · 23
REVISION OF URACANTHUS
posteriorly. Scutellum semicircular, with dense pubescence. Elytra 3.88–4.80 × as long as prothorax and 3.65–
4.19 × as long as shoulder width; each elytron with 3 feeble longitudinal carinae; base of elytra with coarse
punctures, decreasing in size towards apex; apex narrowly emarginate and bispinose. Abdomen slender, ven-
tral surface covered with dense pubescence; apex of terminal sternite truncate or slightly emarginate.
Male terminalia. Apex of ventral median lobe slightly emarginate and apex of dorsal lobe pointed; ventral
lobe about as long as dorsal lobe; spined region of internal sac about 5.2–5.4 × as long as basal unspined
region; spined region divided into 2 sections with an unspined gap between sections; first section longer than
unspined gap with dense long simple spines; second section with sparse short simple spines (Fig. 63a). Eighth
sternite strongly obliquely truncate at side, shallowly emarginate at apex, with long and fairly long setae aris-
ing terminally; ventral surface with cloud-like processes (Fig. 63b). Eighth tergite rounded at apex, with fairly
dense simple spines on dorsal surface (Fig. 63c). Paramere 1.5–1.67 × as long as wide, cylindrical in shape,
apex rounded with long and short setae (Fig. 63d).
Female. Body length, 15.47–26.63 mm; width, 2.65–4.85 mm.
Similar to male but differs in having antennae and legs shorter, abdomen more robust, elytra 3.96–4.09 ×
as long as shoulder width and 4.36–4.49 × as long as prothorax.
Ovipositor and spermatheca. Ovipositor short; styli arising terminally with short hairs (Fig. 63e). Sper-
matheca heavily curved; spermathecal gland arising near base (Fig. 63 f).
Distribution
Northern Queensland, northeastern and eastern New South Wales, northern Australian Capital Territory,
central Victoria and eastern & southern Tasmania (Fig. 96).
Biology
Known host is Callitris hugelii. Larvae attack and kill regenerations of the host plant. Attack appears to be
intiated near the tip of the stem, and then the larva works down below the bark in a single longitudinal chan-
nel. At about half way down the stem from the ground level, it girdles the stem. Tops of plants break off
readily at this point, and sometime fall to the ground. The larva continues to feed in the dead tops. Adults were
collected in October and March to April, and attracted to MV light trap.
Comments
This species closely resembles U. triangularis but differs in having the elytral apex relatively narrowly emar-
ginate; the glabrous marks on the elytra narrower and longer, and the pronotal disc less strongly rugose trans-
versely.
Material examined
Holotype. . Van Diemen’s Land ( = TAS): no data; type coll. 1768; bearing a white label with Hope’s
hand writing of the species name, and a red label with hand written sericus olim Van Diemans Land (HMO).
Other material examined. 10 , 7 . QLD: 1 , Archer River X-ring (13°25’S, 142°56’E), 11.iv.1989,
MV lamp, G. A. Daniels (UQIC). NSW: 1 , Crowdy Bay National Park, 15.x.1990, S. G. Watkins Collection
(ANIC); 1 , 3 miles S of Cresent Head, 27.iii.1965, I. F. B. Common & M. S. Upton (ANIC); 1 , Helens-
burgh, x.1962, H. E. Osborne (ANIC). 1 , Sydney, Ftio, W. Ferguson Coll. (ANIC); 2 , Mt Kaputar, 3000ft,
30.x.1967, at light, C. W. Frazier (ANIC); 1 , Engadine, 10.i.1970, R. H. Mulder Collection (AM); 1 , same
data as above but 12.i.1960 (AM); 1 ?, Wahroonga, no abdomen, H. J. Carter (ANIC); 1 , Vaucluse,
9.ii.1920, Coll. no.K44380 (AM). ACT: 1 , Black Mt., Canberra, 22.i.1952, L. J. Chinnick (ANIC). VIC: 1
, Delleys Dell, Grampians, 6.ii.1956, N. B. Tindale (SAM). TAS: 2 , 1 , Freycinet National Park,
28.i.1963, I. F. B. Common & M. S. Upton (ANIC); 1 , Melaleuca (43º25'S, 146º09'E), 19.ii.1990, E. S.
Nielson & P. C. McQuillan (ANIC). Locality unknown: 1 , no data, E. F. du Boulay Collection (WAM).

THONGPHAK & WANG
24 · Zootaxa 1569 © 2007 Magnolia Press
Uracanthus regalis McKeown, 1948
(Figs 26, 64, 97)
Uracanthus regalis McKeown, 1948: 54.
Description
Male. Body length, 17.19–35.69 mm; width, 3.28–7.6 mm.
Colour (Fig. 26). Body blackish brown with head, thorax, femora, and basal elytra darker. Head with
dense white pubescence except median frontal groove and a glabrous area on vertex. Pronotum with 2 wide
longitudinal stripes of white pubescence on each side: 1 on disc and 1 near ventral side; remaining part of
pronotum with golden pubescence. Each elytron with a large subtriangular glabrous mark starting from shoul-
der and ending at about basal 1/3 of elytra, margined with a vague line of dense white pubescence; disc of
each elytron with 4 vague, longitudinal lines of dense white pubescence interspaced with lines of sparse
pubescence. Body colour varies from blackish brown to reddish brown. The subtriangular glabrous marks on
elytra may be slightly reduced or expanded.
Head. Postclypeus triangular, convex and finely dense punctate; frontoclypeal suture wide and deep in
middle; distance between lower lobes of eyes 1.75–2.0 × as long as distance between upper lobes of eyes; dis-
tance between upper lobes of eyes 1.20–1.30 × as long as distance between eyes on ventral side; genal length
0.48–0.62 × as long as head width below eyes. Antennae slightly shorter than body; segments 5–10 flattened
and slightly produced on one side at apex; apical ¼ of segment distinctly thinner than basal ¾.
Thorax and abdomen. Pronotum 1.11–1.39 × as long as width, rounded at side; posterior margin 1.33–
1.43 × as wide as anterior margin; disc smooth and finely punctate with 2 small nodules in middle; disc and
side strongly rugose transversely. Scutellum semicircular, with dense pubescence. Elytra 4.37–4.92 × as long
as prothorax and 3.27–3.79 × as long as shoulder width; each elytron with 3 feeble longitudinal carinae; apex
narrowly emarginate and bispinose, sutural spine larger than marginal one. Apex of terminal sternite truncate.
Male terminalia. Apex of ventral median lobe narrowly emarginate and apex of dorsal lobe slightly
pointed; ventral lobe longer than dorsal lobe; spined region of internal sac 7.0–7.2 × as long as basal unspined
regions; spined region divided into three sections: first section with dense clustered long simple spines; no
unspined gap between first and second sections; second section with sparse short simple spines; a wide
unspined gap between second and third sections; third section with very sparse short simple spines (Fig. 64a).
Eighth sternite strongly obliquely truncate at sides, apex strongly emarginate and V- shaped, with fairly dense
setae arising terminally; ventral surface with dense cloud-like processes present and fairly dense short simple
spines (Fig. 64b). Apex of eighth tergite more or less rounded and sometimes slightly emarginated; dorsal sur-
face with dense simple spines and multi-branched spines (Fig. 64c). Paramere 1.7–1.98 × as long as wide,
cylindrical in shape; apex rounded with long and short setae (Fig. 64d).
Female. Body length, 18.21–35.69 mm; width, 3.28–7.6 mm.
Similar to males but differs in having more robust body and shorter antennae; elytra slightly longer and
wider, 3.36–3.82 × as long as shoulder with and 4.12–4.84 × as long as prothorax; apical ¼ of antennal seg-
ment 11 slightly thinner than basal ¾; apex of terminal sternite rounded.
Ovipositor and spermatheca. Ovipositor short; styli arising terminally with short hairs (Fig. 64e). Sper-
matheca clearly curved; spermathecal gland long and arising at base (Fig. 64f).
Distribution
Western and southwestern Western Australia (Fig. 97).
Biology
The only known host is Banksia sp. Adults were collected from August to February and attracted to light.

Zootaxa 1569 © 2007 Magnolia Press · 25
REVISION OF URACANTHUS
Comments
This species resembles U. dubius but differs in having the body more robust and the subtriangular mark
on the elytra broader and more distinct; the elytra with fewer and wider longitudinal lines of white pubes-
cence, and the elytral apex very narrowly emarginate.
Material examined
Holotype. . WA : Denmark (34º58'S, 117º21'E), Coll. no. K67661, bearing a red holotype label; 2 seg-
ments of left antenna missing (AM).
Other material examined. 15 , 18 . WA: 1 , South Perth (31º58'S, 115º51'E), 3.ii.1975, K. T. Richard
(WADA); 1 , same locality but 12.xii.1904, H. M. Giles (NMV); 1 , same locality as above but H. M. Giles,
F. E. Wilson Collection (NMV); 1 , Yanchep, 17.ix.1970, at light, S. J. Curry (WADA); 1 , same data as
above but 15.iii.1970 (WADA); 1 , Denmark (34º58'S, 117º21'E), ii.2002, M. L. & S. Jone (WAM); 1 ,
Lockwood Spring (27º46'S, 114º28' E), Kalbaroi, 6.vii.1940, K. Youngson & R. Johnane (WAM); 1 , Kala-
munda (31º58'S, 116º03'E), 24.x.1962, J. Dell (WAM); 1 , Bateman, 11.x.1998, hand collecting, P. R. Davis
(WADA); 1 , Kukerin, 8.xii.1993, UV light trap, A. Szito (WADA); 2 , Woodridge, intercept trap, 20–
30.x.1996, H. Demarz (WADA); 1 , same data as above but (ANIC); 1 , Esperance, 31.x.1951, E. F Wilson
Collection, W. L. Brown (NMV); 1 , 16 km NW of Eneabba, 9–12.ix.1987, at gas light, at night, T. F. Huston
(WAM); 1 , Mukcain (WAM); 1 , Dianella, in Shopping Center, 25.ix.1995, A. Szito (WADA); 1 , Moore
River National Park (31º10'S, 115º40'E), 3–4.iv.1991, at gas light, at night, T. F. Huston (WAM); 1 , 40 km N
of Badgingarra, 15.iii.1995, UV light, A. Szito (WADA); 1 , in Banksia, C446 (QDPI); 1 , Maylands,
28.iii.1974, on door, J. T. Curry (WADA); 1 , North Tarin Rock Nature Reserve (32º59'S, 118º14'E), 16–
18.x.1985, T. F. Houston, Coll. no.618-1 (WAM); 1 , 2 , Swan River, J. Clark (WADA); 1 ,
Onslow,14.iii.1965, B. G. Muir (WAM). 1 , Albany (NMV); 1 , 2 miles S by SW of Mt Ragged, 13.xi.1969,
M. S. Upton (ANIC); 1 , 25 km N Eneabba (29º36'S, 115º15'E), 24–25.x.1984, A. A. Calder (ANIC); 1 , 5
km W of Quairading, 1.x.1992, E. D. Edward & E. S. Nieson (ANIC). Locality unknown: 1 , E. F. du Bou-
lay Collection, det by K. C. McKeown as U. sp. (WAM); 2 , no data, H. J. Carter Collection (NMV); 1 , no
data, Hope/Westwood Collection (HMO).
Uracanthus dubius Lea, 1916
(Figs 27, 65, 97)
Uracanthus dubius Lea, 1916: 371.—McKeown, 1947: 64.
Description
Male. Body length, 25.08–36.08 mm; width, 4.91–7.19 mm.
Colour (Fig. 27). Body reddish brown to blackish with head, thorax, femora, and basal elytra darker. Head
covered with dense white pubescence except median frontal groove and a longitudinal glabrous area on ver-
tex. Pronotum with 2 longitudinal stripes of dense white pubescence on each side: 1 on disc, narrow and
vague, and 1 near ventral side, wide and distinct; remaining prothorax with fairly dense white pubescence.
Each elytron with an indistinct subtriangular mark covered with sparse, short, white pubescence, starting from
the shoulder and extending to basal 1/3, and the mark margined with dense white pubescence; remaining part
of elytra covered with dense white pubescence arranged like several longitudinal lines. Body colour varies
from blackish brown to reddish brown.
Head. Postclypeus triangular, convex and finely dense punctate; frontoclypeal suture wide and deep in
middle; distance between lower lobes of eyes 1.77–1.9 × as long as distance between upper lobes of eyes; dis-
tance between upper lobes of eyes 1.05–1.09 × as long as distance between eyes on ventral side; genal length

THONGPHAK & WANG
26 · Zootaxa 1569 © 2007 Magnolia Press
0.38–0.46 × as long as head width below eyes. Antennae slightly shorter than body, segments 5–10 flattened
and distinctly produced on one side at apex; apical ¼ of segment 11 clearly thinner than basal ¾.
Thorax and abdomen. Pronotum 1.17–1.34 × as long as wide, with a very small angular process at each
side; posterior margin 1.13–1.33 × as wide as anterior margin; disc finely punctate with 2 small nodules on
disc; disc and sides strongly rugose transversely. Scutellum semicircular, with dense pubescence. Elytra 4.05–
4.55× as long as prothorax and 3.38–3.60 × as long as shoulder width; each elytron with 3 feeble longitudinal
carinae; apex widely emarginate and strongly bispinose. Elytral apex may vary from widely to narrowly emar-
ginate. Apex of terminal sternite truncate.
Male terminalia. Apex of ventral lobe strongly and widely emarginate and apex of dorsal lobe rounded or
weakly pointed; ventral lobe much longer than dorsal lobe; spined region of internal sac 5.3–5.42 × as long as
basal unspined regions; spined region divided into two sections without unspined gap between sections: first
section with dense clustered long simple spines and fairly dense multi-branched spines; second section with
sparse short simple spines (Fig. 65a). Eighth sternite rounded or slightly obliquely truncate at sides, very shal-
lowly emarginate at apex, with long and fairly long setae arising terminally; ventral surface with cloud-like
processes (Fig. 65b). Eighth tergite with a small notch at apex; dorsal surface with dense simple spines and
multi-branched spines (Fig. 65c). Paramere 2.0–2.3 × as long as wide, cylindrical in shape, apex rounded with
long and short setae (Fig. 65d).
Female. Body length, 25.27–38.04 mm; width, 5.07–8.17 mm.
Similar to males but differs in having more robust body and shorter antennae; elytra 3.46–3.69 × as long as
shoulder width and 4.03–4.75 × as long as prothorax; apical ¼ of antennal segment 11 slightly thinner than
basal ¾.
Ovipositor and spermatheca. Ovipositor short; styli arising terminally with short hairs (Fig. 65e). Sper-
matheca heavily curved; spermathecal gland very long and arising near base (Fig. 65f).
Distribution
Southeastern Western Australia, southwestern South Australia and central New South Wales (Fig. 97).
Biology
Hosts are unknown. Adults were collected by light trap during September–January and a pitfall trap in
Febuary.
Comments
This species resembles U. regalis but differs in having the subtriangular mark on the elytral disc almost
completely covered with sparse short pubescence; elytra with more longitudinal lines of dense white pubes-
cences.
Material examined
Holotype. . SA: Fowlers Bay (31º58'S, 132º28'E), Coll. no. 5482, bearing a name label on which
‘TYPE’ was written in red (SAM).
Paratypes. 3 , 1 . SA: 2 , 1 , same data as above (SAM); 1 , no data (SAM).
Other material examined. 19 , 8 . NSW: 1 , 16 km W of Euabalong West, 27.x.1992, M. S. & B. J.
Moulds (AM). SA: 1 , Ceduna (32°07'S, 133°40'E), 29.xii.1957, P. Aitken (SAM); 1 , 5 km S by E of
Ceduna (32°10'S, 133°41'E), 10.x.1968, M. S. Upton & J. E. Feehan (ANIC); 5 , Yumbarra National Park,
11.xi.1975, at light, J. A. Herridge (SAM); 1 , same data as above but Inila Rock Waters (13°46'S,
133°25'E), 20–30.ii.1995, pitfall, H. Owen (SAM); 2 , Yalata, 4.x.1989, P. R. B. Det. 1971 by E. G Matthews
(WINC); 2 , Wittelee Point, 10 km SE Ceduna, 1.iii.1975, at light, J. A. Herridge (SAM); 1 , Calpatanna,
W. H. Cons Pk. Wedina Well, Eyre Pen., 30.xi.1986, at light, J. A. Forrest (SAM); 2 , 10 mi E by SE of

Zootaxa 1569 © 2007 Magnolia Press · 27
REVISION OF URACANTHUS
Koonalda HS, SA, 19.x.1968, no abdomen, Britton, Upton & Balderson (ANIC). WA: 2 , Eucla, 5.xi.08, C.
French’s Collection (NMV); 1 , Officer Basin, N. E. of Stretch Mound Great Vict Desert, 24–28.ix.1991, R.
P. McMillan (SAM); 2 , 1 , Warbla, 9.i.1960, at light, P. Aitken (SAM); 1 , 7 mi E by N of Balladonia HS,
13.x.1968, Britton, Upton & Balderson (ANIC); 2 , WA (ANIC). Locality unknown: 1 , Australia (SAM);
1 , S & W Australia (QM).
Uracanthus simulans Pascoe, 1866
(Figs 28, 66, 98)
Uracanthus simulans Pascoe, 1866: 92.—Best, 1882: 35; Tepper, 1887: 38; French, 1911: 67; Aurivillius, 1912: 147;
Lea, 1916: 383; McKeown, 1947: 66; Duffy, 1963: 117.
Description
Male. Body length, 15.89–29.26 mm; width, 3.00–5.85 mm.
Colour (Fig. 28). Body reddish to blackish brown with head, thorax, femora, and basal elytra darker. Head
with dense pale yellowish pubescence. Pronotum with 2 wide longitudinal stripes of very dense pale yellow-
ish pubescence on each side: 1 on disc and 1 near ventral side; remaining part of pronotum with less dense
pale yellowish pubescence; median line on pronotal disc more or less glabrous. Each elytron with a subtrian-
gular glabrous mark, extending from shoulder to basal 1/3, without distinct margin; some areas of the mark
sometimes covered with sparse short pale yellowish pubescence; remaining part of elytron with short dense
yellowish or pale pubescence. The glabrous mark on the elytron may be longer or shorter; body colour may be
lighter than the above described.
Head. Postclypeus semicircular, flattened, and coarsely punctate; frontoclypeal suture wide and deep in
middle; distance between lower lobes of eyes 1.42–1.53 × as long as distance between upper lobes of eyes;
distance between upper lobes of eyes 1.67–1.72 × as long as distance between eyes on ventral side; genal
length 0.33–0.41 × as long as head width immediately below eyes. Antennae slightly shorter than body; seg-
ments 5–10 flattened and distinctly produced on one side at apex.
Thorax and abdomen. Pronotum 1.26–1.47 × as long as wide, with a small rounded process at each side;
posterior margin 1.07–1.36 × as wide as anterior margin; pronotal disc finely punctate with 2 very weakly
raised nodules in middle area; disc and side strongly rugose transversely. Scutellum semicircular, with dense
pubescence. Elytra 3.54–4.15 × as long as prothorax and 3.40–3.77 × as long as shoulder width; basal elytra
coarsely punctate; apex slightly emarginate or truncate, with a small spine at suture and small process or
rounded at margin. Elytral apex may be almost rounded at margin in some specimens. Apex of terminal ster-
nite truncate or slightly emarginate.
Male terminalia. Apex of ventral lobe emarginate and apex of dorsal lobe rounded or slightly pointed;
ventral lobe about as long as dorsal lobe; spined region of internal sac 2 × as long as basal unspined region;
spined region divided into two sections with an unspined gap between sections: first section with dense clus-
tered long simple spines on sides with an unspined area in middle; second section with sparse short simple
spines (Fig. 66a). Eighth sternite obliquely truncate or slightly at side, shallowly and widely emarginate at
apex, with long and fairly long setae arising terminally; ventral surface with simple spines (Fig. 66b). Eighth
tergite rounded or slightly emarginate at apex with long and short setae; dorsal surface with fairly dense sim-
ple spines and multi-branched spines (Fig. 66c). Paramere 1.78–1.8 × as long as wide, cylindrical in shape;
apex rounded with long and short setae (Fig. 66d)
Female. Body length, 18.87–31.48 mm; width, 3.64–6.99 mm.
Similar to male but differs in having broader body and shorter antennae; elytra slightly longer and wider,
3.19–3.66 × as long as shoulder width and 3.08–4.53 × as long as prothorax.
Ovipositor and spermatheca. Ovipositor short; styli arising terminally with short hairs (Fig. 66e). Sper-
matheca slightly curved; spermathecal gland arising near base (Fig. 66f).

THONGPHAK & WANG
28 · Zootaxa 1569 © 2007 Magnolia Press
Distribution
Northern and southern Northern Territory, southern Queensland, western & northern New South Wales,
northwestern Victoria, southern and southwestern South Australia, and western and southern Western Austra-
lia (Fig. 98).
Biology
Hosts are Banksia australis (Best, 1882), Helichrysum ferrugineum and Acacia sp. (French, 1911) and
Acacia longifolia (Pierce, 1917) and Acacia sowdenii. Adults emerge during December and April (French,
1911). Adults were collected from October to March, and attracted to light.
Comments
This species resembles U. triangularis Hope but differs in having the subtriangular mark on the elytra
smaller and narrower, elytra without glabrous areas on apical areas, and the elytral apex usually with only 1
spine at the suture.
Material examined
Holotype. . SA: no data; type Pascoe Coll. 93-60; bearing a circular type label, and with Pascoe’s hand-
writing of the species name (BMNH).
Other material examined. 47 , 20 . QLD: 1 , Eulo (28°09'S, 145°02'E), 24.ix.1991, at light, G.
Daniels (UQIC). NT: 1 , Plenty River, Salt Lake, Simpson Desert (24°09'S, 136°41'E), 14.ix.1992, L. Arch-
hold (NTM); 1 , Daly Waters (13°18'S, 130°15'E), 18–21.i.1972, T. Weir & A. Allwood (NTM). 1 , 2 , Nr
Reedy Rockhole, Amadeus Basin (24°20'S, 131°35'E), 16.ix.1962, P. Ranford (ANIC); 1 , same data as
above but 1.x.1962 (ANIC); 1 , 1 , 9 km of Kulgera (25°46'S, 133°18'E), 1.x.1972, J. Upton (ANIC).
NSW: 1 , Narwarre sta., 33 miles SE of Louth, 1949, E. E. Mitchell (ANIC); 1 , Broken Hill, 22.x.1944, C.
E. Chadwick (ASCU); 1 , 1 , near Visitors Center, Mootwingee Historic Site (30°58'S, 142°04'E),
7.xi.1984, at mercur vapour lamp, G. R. Brewn & H. M. Holmes (ASCU); 1 , NSW (NMV). VIC: 1 , Mil-
dura (NMV); 1 , Yanac, 5.xi.1978, J. C. Le Sovef (ANIC). SA: 1 , Wharminda Wells, E Side Hinks Conser-
vation Park, Eyre Peninsular, 5.xii.1986, at light, J. A Forrest (SAM); 1 , 8 km N of SW Carrllin HS, Sand
End of Formby Bay, York Peninsular, 4.x.1965, N. McFarland (SAM); 1 , Clere (UQIC); 1 , no data, SA
(MAM); 1 , SA, Coll. no. K36089 (AM); 1 , Parachilna (31°07'S, 138°23'E), Flinders Range, Nat. Hist.
Exp. (SAM); 1 , Ouldea, SA, R. T. Muaurice (SAM); 1 , Hiltaba (32°07'S, 135°03'E), Gawler Ranges,
ix.1972, Field Nat. Soc (SAM); 1 , 132 km N. of Cook (30°36'S, 130°24'E), 18.iix.1980, at light, J. A. Forret
(SAM); 1 , Winbring, det by Lea (WADA); 3 , 10km N by NE Mt. Woodroofe (26°14'55″S, 131°47'36″E),
Musgrave Ra, NGO1, 13.x.1994, at light, Pitjantjatjaral Lands survey (SAM); 1 , 24 km N Mt Serle inbed of
Frome Creek, 9.xi.1976, at light, M. Minchino (SAM); 1 , N of Roxby Downs, Woodland, SA, 25.x.1976, at
light in camp, Acacia sowdenii, L. D. Williams (SAM); 1 , 17 miles S of Mt Finke, 29.x.1974, P. Aiken
(SAM); 1 , SA, Port. Lincoln, Seaford Bay area, iv –v.1964, J. G. Casavara (SAM); 1 , SA, Blackburn Col-
lection, Coll. no. 5479 (SAM).1 ?, SA, no abdomen (ANIC). WA: 1 , South Perth, 7.iii.1904, H. M. Giks,
dept by Lea (NMV); 1 , Kalgoorlie, det by K. C. McKeown (ASCU); 1 , Binnu (28°02'S, 114°40'E),
8.iii.1965, H. de Graaf (WAM); 3 , same data as above but 26.iii.1965 (WAM); 2 , Laverton (28°14'S,
122°33'E), 25.iii.1924, det by McKeown (WAM); 2 , 1 , Mullewa (28°32'S, 115°30'E), WA, Miss F. May
(SAM); 2 , same data but det by Lea, F. E. Wilson Collection (NMV); 1 , same locality but 23.iii.1939, det
by McKeown (WAM); 1 , Kalgoorlie (30°45'S, 121°27'E), 20.iv.1922, J. H. Welez and J. Carter Coll. det by
Lea (NMV); 1 , same locality as above but 10.iii.1911, W. du Boulay, det by Lea (NMV); 1 , same locality
as above but det by Lea (NMV); 1 , Thevenard Island, 8.x.1966, G. Bostock (WAM); 1 , Beverley, WA, F.
H. du Boulay (SAM); 3 , Norseman, WA, 23.iii.1971, M. S. Upton (ANIC); 1 , Ravensthorpe, 25.ii.1971,
K. T. Richards (WADA); 2 , 15 km N by NE of Ajana, Murchison (27°49'S, 114°41'E), 27.iii.1971, M. S.

Zootaxa 1569 © 2007 Magnolia Press · 29
REVISION OF URACANTHUS
Upton (ANIC); 1 , 42 km S of Moorine Rock, WA, 13.i.1980, K. & S. Carnaby (ANIC); 1 , Coolgardie,
i.1948, W. du Boulay (WAM); 1 , Geraldton (SAM). Locality unknown: 1 , Australia Old Collection det
by Lea, F. E. Wiston Collection (NMV); 1 , Australia Old Collection det by Lea (QM); 1 , JR. (?), Hope/
Westwood Coll. Cab2, d.27 (HMO); 1 , Australia Old Coll. from SAM (NMV); 1 , Coll. no. K39083 (AM).
Uracanthus fuscus Lea, 1916
(Figs 29, 67, 98)
Uracanthus fuscus Lea, 1916: 372.—McKeown, 1947: 64.
Description
Male. Body length, 14.45–23.70 mm; width, 2.95–4.98 mm.
Colour (Fig. 29). Body blackish brown with head, thorax and basal elytra darker. Head (particularly frons
and vertex) covered with dense white pubescence except a narrow longitudinal glabrous area on vertex.
Pronotal disc with a wide longitudinal stripe of very dense white pubescence on each side; remaining protho-
rax covered with dense white pubescence. Each elytron with a very vague subtriangular mark covered with
relatively sparse hairs, starting from shoulder and extending to about basal 1/3, without any margin; the
remaining of elytra with fairly dense white pubescence. Meso- and meta-sterna, abdomen, and legs covered
with dense white pubescence. Body colour may vary from blackish brown to blackish.
Head. Postclypeus semicircular and slightly convex, with dense coarse punctures; frontoclypeal suture
deep and narrow in middle; distance between lower lobes of eyes 1.82–2.0 × as long as distance between
upper lobes of eyes; distance between upper lobes of eyes 1.3–1.67 × as long as distance between eyes on ven-
tral side; genal length 0.25–0.32 × as long as head width immediately below eyes. Antennae shorter than
body; segments 5–10 strongly flattened and produced on one side at apex.
Thorax and abdomen. Pronotum 1.05–1.54 × as long as width, with a weak and rounded process at each
side; posterior margin 1.16–1.69 × as wide as anterior margin; pronotal disc binodulose in middle area; disc
and sides strongly rugose transversely near anterior and posterior margins. Scutellum semicircular, with dense
pubescence. Elytra 2.97–3.86 × as long as prothorax and 3.25–3.44 × as long as shoulder width; each elytron
with 3 feeble longitudinal carinae; basal elytra finely punctate; apex more or less truncate with a small process
or spine at suture. Segment 1 of hind tarsus 0.66–0.68 × as long as sum of segments 2+3. Apex of terminal
sternite truncate.
Male terminalia. Apex of ventral lobe strongly emarginate and apex of dorsal lobe rounded or slightly
pointed; ventral lobe as long as dorsal lobe; spined region of internal sac 4.89–5.12 × as long as basal
unspined regions; spined region divided into two sections without unspined gap between sections; first section
with dense clusters of long simple spines and fairly dense multi-branched spines at lateral sides; second sec-
tion with sparse multi-branched spines and fairly dense simple spines (Fig. 67a). Eighth sternite obliquely
truncate or nearly rounded at sides, shallowly emarginate at apex, with long and fairly long setae arising ter-
minally; ventral surface with fairly dense cloud-like processes (Fig. 67b). Eighth tergite rounded at apex, with
fairly dense simple spines and multi-branched spines near base (Fig. 67c). Paramere robust, 1.98–2.1 × as
long as wide, cylindrical in shape, apex rounded with long and short setae (Fig. 67d).
Female. Body length, 18.48–26.43 mm; width, 3.69–5.46 mm.
Ovipositor and spermatheca. Ovipositor very short; styli arising terminally with short hairs (Fig. 67e).
Spermatheca distinctly curved; spermathecal gland arising near middle (Fig. 67f).
Distribution
Southern and southeastern South Australia (Fig. 98). One specimen was collected in Queensland but exact
locality is unknown.

THONGPHAK & WANG
30 · Zootaxa 1569 © 2007 Magnolia Press
Biology
Known host is Acacia sp. Adults were collected in October to November by light trap.
Comments
This species resembles U. simulans but differs in having body more slender in shape; the triangular mark
on basal elytron vague.
Material examined
Holotype. . SA: Coll. no. 5698, bearing a name label on which ‘TYPE’ was written in red (SAM).
Paratypes. 4 . SA: 3 , South Australia, Australia old collection (SAM); 1 , no locality data, Australia
old collection (SAM).
Other material examined. 38 , 3 . QLD: 1 , Queensland (NMV). SA: 5 , Uro Bluff, W. of Lake Tor-
rens, 2.x.1971, H. Mincham (SAM); 2 , 20 km Wirraminma Stn., 2.xi.1975, at light, J. A. Herridge (SAM); 1
, same locality as above but 27.x.1953, N. B. Tindale (SAM); 1 , Kingoonya, 12.x.1977, at light, G. F.
Gross (SAM); 1 ?, no abdomen, same locality as above but R. Harvey (SAM); 2 , SA, Rev. A. E. Burgess
(SAM); 2 , Middleback Station (32°57'S, 137°24'E), at light, 26.x.1983, J. Woddell (SAM); 1 , Yalpara Sta-
tion, 27.x.1970, at light, P. R. B. (WINC); 1 , 18 mile NE of Derraroo (32°33'S, 138°49'E), Yalpara Station,
22.xi.1969, at light, P. R. B. (WINC); 1 , Penong, 18.x.1996, J. W. G. (WINC); 1 , Tarcoola (NMV); 1 ,
SA (MAM); 10 , 2 , Reevesby Island, SA, xii.1956, D. J. Mahony, McCoy Soc. Exped Sir Jos. Banks
Groups, xii 1963–i. 1937 (NMV); 1 , Port Lincoln, SA, i.1951, ex Acacia, C. Oke (NMV); 1 , Hammond
SA, i.1948, V. H. Mincham (AM); 1 , North Flinders Range, SA, A. H. Collection (AM); 1 , Kingoonya,
SA, R. Harvey (AM); 1 , Beverley, W. D. V. Firdub (AM); 1 , SA, Koonalda, 5.i.1986, P. Aitken (SAM).
WA : 2 , WA, C. French Collection (NMV). Locality unkown. 1 , no data, I.d by John Davidson (WINC);
1 , 20 miles W of ? Uenluing Rocks, i.1909, Mu Chandu Rec 26.v.1909 (NMV); 1 , no data (WINC).
Uracanthus cryptophagus Olliff, 1892
(Figs 30, 68, 99)
Uracanthus cryptophagus Olliff, 1892: 896.—Froggatt, 1907:193; Aurivillius, 1912: 147; Tillyard, 1926: 233; Brimble-
combe, 1943: 37; McKeown, 1947: 63; Duffy, 1963:114.
Uracanthus cryptophaga.—Allen, Blunno, Froggatt & Guthrie, 1898b: 1216; Froggatt, 1919: 262; 1923:96.
Description
Male. Body length, 24.17–38.73 mm; width, 4.15–7.47 mm.
Colour (Fig. 30). Body reddish brown to blackish brown with head, prothorax, basal and marginal elytra,
and legs often darker. Body covered with dense golden pubescence. Pronotal disc with 4 narrow longitudinal
stripes of dense golden pubescence; each side with a wide longitudinal stripe of dense pubescence near ventral
side. Each elytron with a very vague subtriangular basal mark covered with short golden pubescence and
obvious darker than other part, without any margin; remaining elytron covered with very dense golden pubes-
cence, arranged in 4 longitudinal lines. Body colour may vary from reddish brown to blackish brown; pubes-
cence on the basal mark of the elytron varies from very sparse to dense.
Head. Postclypeus semicircular, slightly convex, finely punctate; frontoclypeal suture wide and deep in
middle; distance between lower lobes of eyes 1.35–1.55 × as long as distance between upper lobes of eyes;
distance between upper lobes of eyes 1.17–1.36 × as long as distance between eyes on ventral side; genal
length 0.55–0.63 × as long as head width immediately below eyes. Antennae slightly shorter than body, with
dense short pubescence; segments 5–10 flattened, and produced on one side at apex.
Thorax and abdomen. Pronotum 1.27–1.45 × as long as wide, rounded at side; pronotal disc binodulose

Zootaxa 1569 © 2007 Magnolia Press · 31
REVISION OF URACANTHUS
in middle area and finely punctate; disc and side distinctly rugose transversely. Scutellum semicircular, with
dense pubescence. Elytra 4.01–4.70 × as long as prothorax and 3.79–4.11 × as long as shoulder width; each
elytron with 3 feeble longitudinal carinae; basal elytra with fairly dense punctures; apex bispinose. Abdomen
with dense pubescence; apex of terminal sternite truncate.
Male terminalia. Apex of ventral median lobe widely emarginate, dorsal median lobe rounded; dorsal
lobe shorter than ventral lobe; spined region of internal sac about 3 × as long as basal unspined regions; spined
region divided into 2 sections; first section with mixture of dense large and long simple spines and fairly dense
multi-branched spines; second section with fairly dense simple spines (Fig. 68a). Eighth sternite obliquely
truncate at terminal sides, strongly emarginate at middle area with long and fairly long setae arising termi-
nally; ventral surface with sparse basally-forked spines in middle and sparse simple spines near sides (Fig.
68b). Eighth tergite almost truncate at apex, basal area with simple spines (Fig. 68c). Paramere 1.82–1.85 × as
long as wide, cylindrical in shape, apex rounded with long and short setae (Fig. 68d).
Female. Body length, 32.33–43.79 mm; width, 5.72–8.56 mm.
Antennae and legs distinctly shorter and body more robust; elytra 3.83–4.15 × as long as shoulder width
and 4.00–4.87 × as long as prothorax; apex of terminal sternite truncate or slightly emarginate.
Ovipositor and spermatheca. Ovipositor short; styli arising laterally with both long and short hairs (Fig.
68e). Spermatheca heavily curved and C-shaped; spermathecal gland very long and arising at basal 1/3 (Fig.
68f).
Distribution
Eastern and northeastern New South Wales and southeastern Queensland (Fig. 99). One specimen was
collected in Western Australia but exact locality is unknown.
Biology
Host plants are orange, native finger lime (Citrus australasica), mandarin orange, and lychee. Eggs and
larvae were found in the smaller branches of citrus trees. Adults were collected in January, July, August, and
during September to December and attracted to MV light.
Comments
This species differs from other species of Uracanthus in having the pronotal disc and each elytron covered
with very dense long pubescence, arranged in four longitudinal lines.
Material examined
Holotype. . NSW: Richmond R. (28º47'S, 153º16'E), 5.vii.1892, Coll. no. k36086, bearing a red holo-
type label; 2 segments of right antenna missing (AM).
Paratypes. 1 , 1 . NSW: 1 , same locality as above but Coll. no. 5480 (SAM); 1 , same locality as
above but Coll. no. 630 (SAM).
Other material examined. 25 , 30 . QLD: 1 , Brisbane, R. IIIidge (SAM); 1 , same as above but
(UQIC); 1 , same data as above but (ANIC); 1 , Bucasia, Nth QLD, 9.xii.1990, K. J. Sandery (SAM); 1 ,
same data as above but 17.xi.1990 (ANIC); 1 , same data as above but 9.xii.1990 (SAM); 1 , Oakview
(26º07'S, 152º20'E) SE Barracks 15.i.2002, MV light, Monteith & Thompson, Coll. no. 10349 (QM); 1 ,
Mackay, 5.xi.1908, C. French (NMV); 1 , Montville, L. Smith (UQIC); 1 , National Park, R. Illidge
(UQIC); 1 , K'roy, 8.xii.1985, larval tunneling in citrus trunk, J. Wessels (QDPI); 1 , Macpherson's Range
(22º17'S, 146º18'E), Deane (UQIC); 1 , Imbil (26º27'S, 152º40'E), 1931, host Scrub, RB (QDPI); 1 ,
Gayndah (25º37'S, 151º36'E), 5.x.1941, A. May (QDPI); 1 , 2 , Yeppoon, QLD, 26.xi.1965, J. C. Le Sovef
(ANIC); 1 , 2 m Elimbah, 6.ix.1966, x Citrus, J. H. B (QDPI); 1 , 3 km N of Mourangee, at light, Scrub
(ANIC); 1 , Homebush, xii.1940, S. R. E. Brook Collection (ANIC); 1 , Tambourine Mt (33º49'S,

THONGPHAK & WANG
32 · Zootaxa 1569 © 2007 Magnolia Press
151º09'E), 8.i.1912, E. Sutton Collection (QM). NSW: 1 , Tweed R. (28º13'S, 153º33'E), xii.1909, E. C.
Bons (UQIC); 1 , same locality but (MAM); 2 , same locality but (WADA); 1 , 1 , same locality but
(QM); 1 , same locality but (AM); 1 , Richmond R. (28º47'S, 153º16'E), 5.vii.1892, H. J. Carter Collection
(NMV); 2 , same locality but (AM); 1 , same locality but (SAM); 1 , same locality but (VAIC); 1 , same
locality but (ASCU); 2 , same locality but 5.i.1892 (ANIC); 1 , same locality but 5.vii.1892, Froggatt Col-
lection (HMO); 1 , 1 , Alstonville (28º49'S, 153º26'E), 5.xii.1978, W. E. Wright (AM); 1 , Don (23º57'S,
149º51'E), xii.1964, E. Sutton Collection (QM); 1 , Green Pigeon via Kyogle (28º45'S, 153º00'E), iix.1940,
boring in orange, E. B. H (ASCU); 1 , Grafton (30º05'S, 152º24'E), x.1909 (ASCU); 1 , 1 , same locality
but xii.1909, Citrus tree (NMV); 1 , Lismore, xi.1953, A. L. Brown (NMV); 1 , same data as above but
20.xi.1954 (NMV); 1 , Dorrigo National Park, 3.i. 1968, at light, C. W. Frazier (ANIC); 1 , NSW,
5.xi.1908, C. French Collection (NMV); 1 , NSW, F. E. Collection (NMV). 1 , NSW (WADA). Locality
unknown: 1 , N Coast, i.1936, Citrus (ASCU); 1 , H. Binore, H. J. Carter Collection (NMV); 1 , Losinore
5–33 wells (ASCU); 1 , WA, 6.iix.1956, in Mandarin branch (QDPI); 1 , A. S. B Coll. no. 544 (QDPI).
Uracanthus fuscocinereus White, 1855
(Figs 31, 69, 100)
Uracanthus fuscocinereus White, 1855: 330.–Aurivillius, 1912: 147; Lea, 1916: 384;
McKeown, 1940: 309; McKeown, 1947: 64.
Description
Male. Body length 26.94–31.01 mm; width, 2.85–5.90 mm.
Colour (Fig. 31). Head, thorax, and basal elytra blackish brown; remaining elytra, legs, and antennal seg-
ments reddish brown; palpi and anteclypeus yellowish. Head with dense pale yellow pubescence except a gla-
brous subtriangular area on vertex. Pronotum with 2 wide longitudinal stripes of dense golden pubescence on
each side: 1 on disc and 1 near ventral side; remaining pronotum with sparse pale pubescence. Each elytron
with 4 basal glabrous marks: 1, long and narrow at margin; 1, robust at shoulder, 1, smallest, short and narrow
between suture and shoulder, and 1, longest at suture starting at base and extending towards basal 1/5 of
elytra; basal 1/3 of elytra with dense white pubescence; remaining elytra with golden pubescence. Meso- and
metasterna with white pubescence; femora and abdomen with golden pubescence.
Head. Postclypeus triangular and convex, finely punctate; frontoclypeal suture wide and deep in middle;
distance between lower lobes of eyes 1.65–1.77 × as long as distance between upper lobe of eyes; distance
between upper lobes of eyes 1.05–1.10 × as long as distance between eyes on ventral side; genal length 0.50–
0.62 × as long as head width immediately below eyes. Antennae slightly shorter than body, with uniform and
short pubescence; antennal segment 3 cylindrical, and segments 4–10 flattened, slightly produced on one side
at apex.
Thorax and abdomen. Pronotum 1.27–1.35 × as long as width, rounded or slightly produced at side; pos-
terior margin slightly wider than anterior margin; disc binodulose in middle area and finely punctate; disc and
side strongly rugose transversely throughout. Scutellum semicircular, with dense pubescence. Elytra 4.28–
4.92 × as long as prothorax and 3.60–4.26 × as long as shoulder width; with fairly dense punctures in basal
part; each elytron with 3 feeble longitudinal carinae on disc; apex widely emarginate and bispinose. Abdomen
with short dense pubescence; apex of terminal sternite truncate or slightly emarginate.
Male terminalia. Apex of ventral median lobe pointed and apex of dorsal median lobe rounded; ventral
lobe shorter than dorsal lobe; spined region of internal sac much longer than basal unspined region; spined
region divided into two sections without unspined gap between sections: first section with mixture of dense
large and long simple spines, fairly dense multi-branched spines and dense scale-like processes; second sec-
tion with dense long simple spine and fairly dense short simple spines (Fig. 69a). Eighth sternite obliquely

Zootaxa 1569 © 2007 Magnolia Press · 33
REVISION OF URACANTHUS
truncate or slightly rounded at sides; apex emarginate, with fairly dense setae arising terminally; ventral sur-
face with sparse short simple spines and cloud-like processes (Fig. 69b). Eighth tergite almost truncate at
sides, slightly emarginate at apex with long and short setae; mid area with dense simple spines and multi-
branched spine (Fig. 69c). Paramere 2 × as long as wide, cylindrical in shape, apex rounded with long dense
setae (Fig. 69d).
Female. Body length, 21.32–32.13 mm; width, 3.92–6.94 mm.
Antennae and legs distinctly shorter, body broader with robust abdomen. Elytra slightly longer and wider,
3.36–3.97 × as long as shoulder width and 4.17–4.48 × as long as prothorax; apex of terminal sternite more or
less truncate or slightly emarginate.
Ovipositor and spermatheca. Ovipositor relatively long; styli relatively large, arising terminally with
short hairs (Fig. 69e). Spermatheca slightly curved; spermathecal gland arising at basal 1/3 (Fig. 69f).
Distribution
Southeastern Queensland, northeastern New South Wales and southern Victoria (Fig. 100).
Biology
Hosts are unknown. Adults were collected in August and from October to December.
Comments
This species closely resembles U. quadristriolatus sp. nov. but differs in having middle area of pronotal
disc always covered with some pubescence; each elytron with 4 glabrous marks at base.
Material examined
Holotype. . New Holland (= Australia): no data; specimen no. 52–32; bearing a White’s hand written
label of the species name, and a circular type label of BMNH; right legs and terminal 6–7 segments of anten-
nae missing (BMNH).
Other material examined. 17 , 21 . QLD: 8 , 2 , Stanthorpe (28º39'S, 151º56'E), xii.1964 (QM); 1 ,
same as above but (ANIC); 1 , same locality as above but xi.1925 (QDPI); 1 , same locality as above but
4.xii.1922 (QDPI); 1 , same locality as above but 22.xi.1923 (UQIC); 1 , same locality as above but H.
Jarvis and F. E. Wilson (NMV); 1 , same data as above but S. Watkins Collection (ANIC); 1 , Fletcher,
29.xii.1932, E. Sutton (QM). NSW: 2 , 2 , Sydney (MAM); 1 , same locality as above but xi.1913, C.
Gibbons (AM); 1 , North Sydney, 12.iix.1976, H. J. Carter (NMV); 1 , Collaroy, x.1947, T. O. B (WINC);
1 , 1 , Fairlight near Penrith, 12.xi.1979 (AM); 1 , Iwanly, E. P. du Boulay Coll. (WAM); 1 , Sackville,
22.xi.1983, S. Watkins (ANIC); 1 , Waterfall, 11.xi.1981, S. Watkins (ANIC); 1 , 1 , Narabeen, C. Oke
(NMV). VIC: 1 , Heathcote, 9.xii.1923, Nicholson (SAM); 1 , same locality as above but, 20.xii.1978, S.
Watkins (ANIC). Locality unknown: 1 , 5.xi.1908, C. French Coll. (NMV); 1 , Coll. no. K36087 (AM); 1
, Coll. no. K36087 (AM); 1 , no data (QDPI); 1 , no locality label, S. R. E. Brock Collection (ANIC).
Uracanthus quadristriolatus, sp. nov.
(Figs 32, 70, 100)
Description
Male. Body length, 30.00 mm; width, 5.49 mm.
Colour (Fig. 32). Body dark reddish brown, with head, thorax and basal elytra darker. Head with dense,
long, pale orange pubescence; vertex with a large and long longitudinal triangular hairless mark. Pronotum
with 2 wide, clearly defined stripes of very dense pale orange pubescence on each side: 1 on disc and 1 near

THONGPHAK & WANG
34 · Zootaxa 1569 © 2007 Magnolia Press
ventral side; the remaining pronotum hairless. Each elytron with 3 small basal glabrous marks: 1 on shoulder,
largest, 1 between shoulder and suture, smallest and narrowest, and 1 at suture, rest of elytra covered with uni-
formly dense short pale yellow pubescence.
Head. Postclypeus triangular and convex, finely punctate, and sparsely pubescence; frontoclypeal suture
wide and deep in middle; distance between lower lobes of eyes 1.75 × as long as distance between upper lobe
of eyes; distance between upper lobes of eyes 1.0 × as long as distance between eyes on ventral side; genal
length 0.53 × as long as head width immediately below eyes. Antennae shorter than body, covered with uni-
form short pubescence; antennal segment 3 cylindrical, segments 4 to 10 flattened, slightly produced on one
side at apex.
Thorax and abdomen. Pronotum 1.42 × as long as width, with a small rounded process at each side; pos-
terior margin wider than anterior margin; disc binodulose near middle area; disc and sides heavily rugose
transversely throughout. Scutellum semicircular, with sparse pubescence. Elytra 4.27 × as long as prothorax
and 3.94 × as long as shoulder; basal elytra sparsely punctate; apex bispinose with spine at suture shorter than
that at margin. Abdomen with dense short pubescence, apex of terminal sternite truncate with a shallow notch.
Male terminalia. Apex of ventral median lobe pointed; dorsal median lobe rounded, ventral lobe longer
than dorsal lobe. Spined region of internal sac more than 4 × as long as basal unspined regions; spined region
divided into two sections: first section with mixture of dense large and long simple spines, fairly dense multi-
branched spines and dense scale-like processes; second section with dense long simple spines, becoming
sparser towards the end of this section (Fig. 70a). Eighth sternite obliquely truncate at sides, apex widely
emarginate with fairly dense setae; ventral surface with sparse cloud-like processes (Fig. 70b). Eighth tergite
almost truncate at sides, slightly emarginate at apex with long and short setae, with dense simple spines and
multi-brancheds spine in mid and basal area (Fig. 70c). Paramere 2.2 × as long as wide, cylindrical in shape,
apex rounded with long dense setae (Fig. 70d).
Female. Body length, 23.98–27.09 mm; width, 4.34–4.69 mm.
Similar to males but antennae and legs distinctly shorter; body broader with robust abdomen; elytra slightly
longer and wider, 3.97–4.19 × as long as shoulder width and 4.02–4.28 × as long as prothorax.
Ovipositor and spermatheca. Ovipositor relatively long; styli arising terminally with short hairs (Fig.
70e). Spermatheca curved; spermathecal gland arising at base (Fig. 70f).
Distribution
Northeastern New South Wales and northern Queensland (Fig. 100).
Biology
Hosts are unknown. Specimens were collected November.
Comments
This species resembles to U. fuscocinereus but differs in having the elytra covered with a uniform dense
pale yellowish pubescence; middle area of pronotal disc without hair; the base of each elytron with only 3 gla-
brous marks.
Material examined
Holotype. . NSW: Dorrigo Natural Park, xi.1971, MV light, R. Hardie; bearing a red holotype label
(ANIC).
Paratypes. 2 . NSW: 1 , Armidale, C. F. Deuquet (AM). QLD: 1 , Black Mt., 2m N Kuranoa, N QLD,
4.xi.1976, at light, J. G. Brooks (ANIC).

Zootaxa 1569 © 2007 Magnolia Press · 35
REVISION OF URACANTHUS
Uracanthus insignis Lea, 1916
(Figs 33, 71, 101)
Uracanthus insignis Lea, 1916: 370.—McKeown, 1942: 87; 1947: 64.
Description
Male. Body length, 23.58–28.1 mm; width, 4.27–4.99 mm.
Colour (Fig. 33). Head, thorax, basal 1/3 and apical 1/10 of elytra, and elytral suture blackish brown;
remaining body yellowish to reddish brown. Frons, vertex, antennae and legs covered with fairly dense golden
pubescence. Pronotum with a large longitudinal glabrous area on disc, wider at anterior edge and becoming
narrower at posterior edge, and margined by a narrow line of dense white pubescence at each side of glabrous
area; each side of pronotum with a wide longitudinal stripe of dense white pubescence near ventral side; area
between narrow pubescent line on disc and wide pubescent stripe near ventral side, with dense yellowish
brown pubescence; ventral surface of prothorax sparsely pubescent or almost glabrous. Basal 1/3 suture
behind scutellum and margin of elytron with white pubescence; elytral apex and suture from basal 1/3 to apex
sparsely pubescent or glabrous, and blackish brown; remaining part of elytra with dense short yellowish
pubescence. Meso- and metasterna, and sternites with short yellowish pubescence. Body and apical elytra
colour varies from blackish brown to reddish brown.
Head. Postclypeus triangular, convex, sparsely and coarsely punctate; frontoclypeal suture wide and deep
in middle; frons and vertex finely punctate; distance between lower lobes of eyes 2.27–2.4 × as long as dis-
tance between upper lobes of eyes; distance between upper lobes of eyes 1.0–1.1 × as long as distance
between eyes on ventral side; genal length 0.63–0.79 × as long as head width immediately below eyes. Anten-
nae shorter than body, thin with dense short pubescence and fine punctures; segments 5–10 flattened and pro-
duced on one side at apex.
Thorax and abdomen. Pronotum 1.19–1.29 × as long as width, rounded at side; posterior margin 1.16–
1.34 × as wide as anterior margin; pronotal disc binodulose in middle; disc and side with distinct transverse
rugae. Scutellum semicircular, with dense pubescence. Elytra 4.12–4.66 × as long as prothorax and 3.9–4.04 ×
as long as shoulder; apex bispinose. Apex of terminal sternite truncate.
Male terminalia. Apex of ventral lobe rounded with a very shallow notch; apex of dorsal lobe pointed;
ventral lobe distinctly shorter than dorsal lobe; spined region of internal sac much longer than basal unspined
region, divided into 2 sections: first section with mixture of dense simple spines and multi-branched spines;
second section with fairly dense multi-branched spines (Fig. 71a). Eighth sternite strongly obliquely truncate
at sides, widely emarginate at apex, with fairly dense long setae arising terminally; ventral surface with cloud-
like processes (Fig. 71b). Eighth tergite parallel at sides and truncate at apex with long and short setae, mainly
arising terminally, and dorsal surface with dense simple spines, and multi-branched spines (Fig. 71c).
Paramere short and robust, 1.80–1.83 × as long as wide, cylindrical in shape, apex rounded with long and
short setae (Fig. 71d).
Female. Body length, 23.77–39.14 mm; width, 4.11–7.40 mm
Body more robust and broader, antennae and legs shorter. Elytra slightly longer and wider, 3.83–4.09 × as
long as shoulder width and 4.38–5.29 × as long as prothorax.
Ovipositor and spermatheca. Ovipositor relatively long; styli arising terminally with short hairs (Fig.
71e). Spermatheca slightly curved; spermathecal gland short and arising at base (Fig. 71f).
Distribution
Northern andn eastern Queensland and northeastern New South Wales (Fig. 101).
Biology
Hosts are unknown. Adults were collected from January to July by MV light trap.

THONGPHAK & WANG
36 · Zootaxa 1569 © 2007 Magnolia Press
Comments
This species closely resembles U. punctulatus, sp. nov. but differs in having white pubescent lines on the
pronotal disc, and the elytral suture from basal 1/3 to apex and apical area sparsely pubescent or glabrous, and
blackish brown.
Material examined
Holotype. . NSW: Narara (33º20'S, 151º22'E), Coll. no. I. 5691, W. du Bouday, bearing a name label on
which ‘TYPE’ was written in red (SAM).
Paratype. 1 . NSW: same data as above but bearing a blue paratype label; last segment of abdomen
missing (AM).
Other material examined. 8 , 6 . QLD: 4 , 30 km NE of Coen (13º32'S, 143º29'E), 500 m McIlwraith
Range, Leo Creek Road., 29.vi–4.vii.1976, G. B & S. R. Monteith (QM); 1 , 15 km W Captain Billy Creek
(11º40'S, 142º45'E), Dividing Range, Cape York Penisular, 4–9.vii.1975, G. B. Monteith (QM); 1 , Lam
National Park, 4.ii.1966 (UQIC); 1 , Paluma (15º00'S, 146º12'E), 900m, 10.ii.1980, D. W. Frith (ANIC); 2 ,
8 km W by N of Bald Hill (13º45'S, 143º22'E), Mcllwraith Range, 500 m upper Leo Creek site, 27.v–12.1989,
at light, T. A. Weir (ANIC); 1 , Eungella, C of QLD, i.1976, G. B (ANIC); 1 , Mt. Spec (18º57'S, 146º11'E),
i.1964, G. B (ANIC). NSW: 1 , Comboyne, 1973 (AM); 1 , Newport, 5.ii.1964, R. H. Mulder Collection
(AM); 1 , NSW, Caparra, 1.iv.1992, S. G. Watkins Collection (ANIC).
Uracanthus punctulatus, sp. nov.
(Figs 34, 72, 101)
Description
Male. Body length, 28.03–39.96 mm; width, 5.19–7.83 mm.
Colour (Fig. 34). Body blackish brown, covered with dense and thick pale or pale yellow pubescence;
pronotal disc with a large longitudinal glabrous area, wider at posterior edge and becoming slightly narrower
at anterior edge; punctures on basal half of elytra more or less exposed, giving an appearance of a number of
small blackish dots in pubescence on basal elytra.
Head. Postclypeus subtriangular, slightly convex, sparsely and coarsely punctate; frontoclypeal suture
wide and deep in middle; distance between lower lobes of eyes 1.75–2 × as long as distance between upper
lobes of eyes; distance between upper lobes of eyes 0.7–0.85 × as long as distance between eyes on ventral
side; genal length 0.8–0.85 × as long as head width immediately below eyes. Antennae shorter than body, seg-
ments 4–10 flattened and produced on one side at apex.
Thorax and abdomen. Pronotum 1.18–1.39 × as long as width, side rounded with a feeble process; posterior
margin 1.24–1.46 × as wide as anterior margin; pronotal disc binodulose in middle area, with distinct trans-
verse rugae throughout. Scutellum semicircular, with sparse pubescence. Elytra 4.45–4.83 x as long as protho-
rax and 3.67–4.01 × as long as shoulder width; basal 2/3 of elytra with coarse punctures; apex more or less
truncate with 2 sharp spines. Apex of terminal sternite truncate or slightly emarginate.
Male terminalia. Apex of ventral and dorsal median lobes rounded; ventral lobe shorter than dorsal lobe;
unspined region of internal sac very short: spined region divided into 2 sections: first section with dense scale-
like processes near base and dense large and long simple spines in most part; second section with fairly dense
multi-branched spines (Fig. 72a). Eighth sternite obliquely truncate at sides, strongly emarginate at apex, with
long and fairly long setae arising terminally (Fig. 72b). Eighth tergite more or less truncate and slightly emar-
ginate at apex (Fig. 72c). Paramere short, 2.0–2.3 × as long as wide, cylindrical in shape, apex rounded with
long and short setae (Fig. 72d).
Female. Body length, 35.41–37.56 mm; width, 7.14–7.32 mm

Zootaxa 1569 © 2007 Magnolia Press · 37
REVISION OF URACANTHUS
Body more robust and broader; antennae and legs shorter; elytra 3.61–3.82 × as long as shoulder width
and 4.23–4.79 × as long as prothorax; apex of terminal sternite truncate.
Ovipositor and spermatheca. Ovipositor short; styli arising terminally with sparse long and short hairs
(Fig. 72e). Spermatheca slightly curved; spermathecal gland arising at base (Fig. 72f).
Distribution
Northeastern New South Wales and southeastern Queensland (Fig. 101).
Biology
Hosts are unknown. Adults were collected during October–December.
Comments
This species was wrongly identified as females of U. insignis by previous workers. It resembles U. insig-
nis but differs in having a body covered with uniform pubescence without white pubescent lines on the prono-
tal disc and the elytral suture not glabrous.
Material examined
Holotype. . NSW: Port Macquarie, vi.1982, K. R. Pullen; bearing a red holotype label (ANIC).
Paratypes. 4 , 2 . All specimens bear blue paratype labels. QLD: 1 , Mt Glorious (27º20'S, 152º46'E),
23.x.1971, B. Camtrell (UQIC); 1 , QLD, Relton Bequest, Coll.no. 3455 (QM); 1 , Imbil (26º27'S,
152º40'E) (AM). NSW: 1 , Tuncurry (32º05'S, 152Sº30'E), i.1930 (AM); 1 , Bruxner Park (29º02'S,
152º01'E) near Coffs Harbour, 22.i.1971, D. K. McAlpine (AM). Locality unknown: 1 , no data (ANIC).
Uracanthus maleficus Lea, 1917a
(Figs 35, 73, 102)
Uracanthus maleficus Lea, 1917a: 739.—McKeown, 1947: 65; Duffy, 1963: 117.
Description
Male. Body length, 22.12–26.38 mm; width, 4.92–6.26 mm.
Colour (Fig. 35). Body light reddish to blackish brown. Head with dense golden pubescence. Pronotum
with 2 wide longitudinal stripes of dense golden pubescence on each side: 1 on disc and 1 near ventral side;
remaining pronotum almost hairless; ventral surface of prothorax with fairly dense golden pubescence. Elytra
covered with dense, short golden pubescence.
Head. Postclypeus semicircular, with dense large coarse punctures; frontoclypeal suture deep and narrow
in middle; distance between lower lobes of eyes 2.20–2.36 × as long as distance between upper lobes of eyes;
distance between upper lobes of eyes 0.75–0.78 × as long as distance between eyes on ventral side; genal
length 0.25–0.30 × as long as head width immediately below eyes. Antennae much longer than body; seg-
ments 5–10 flattened and produced on one side at apex.
Thorax and abdomen. Pronotum 0.83–0.95 × as long as width, rounded at side; posterior margin 1.27–
1.62 × as wide as anterior margin; pronotal disc smooth and weakly binodulose. Scutellum semicircular, with
dense pubescence. Elytra 4.62–5.61 × as long as prothorax and 3.23–3.47 × as long as shoulder width; each
elytron with 2 slightly raised longitudinal carinae on disc; basal 1/3 of elytra coarsely punctuate; apex of
elytron bispinose with sutural spine significantly larger. Abdomen paralell sided; apex of terminal sternite
truncate.
Male terminalia. Apex of ventral median lobe rounded; ventral lobe slightly longer than or as long as dor-
sal lobe; spined region of internal sac much longer than basal unspined region; spined region with dense long

THONGPHAK & WANG
38 · Zootaxa 1569 © 2007 Magnolia Press
simple spines throughout except a short unspined terminal area (Fig. 73a). Eighth sternite rounded at side,
shallowly emarginate at apex, with long and fairly long setae arising terminally; ventral surface with simple
spines and cloud-like processes (Fig. 73b). Eighth tergite emarginate at apex, with fairly dense basally-forked
spines on dorsal surface (Fig. 73c). Paramere long, 2.8–2.9 × as long as wide, cylindrical in shape, apex
rounded with long and short setae (Fig. 73d).
Female. Body length, 29.07–32.20 mm; width, 6.66–7.23 mm.
Abdomen more robust and broader, antennae and legs shorter. Elytra longer and wider, 3.32–3.60 × as
long as shoulder width and 6.03–6.55 × as long as prothorax.
Ovipositor and spermatheca. Ovipositor short; styli arising terminally with short hairs (Fig. 73e). Sper-
matheca C-shaped; spermathecal gland short and arising at base (Fig. 73f).
Distribution
This species is only found in Tasmania (Fig. 102).
Biology
Known host is hazelnut (Corylus spp.). Adults were collected from December to April and attracted to
light.
Comments
This species resembles U. pseudogigas sp. nov. but differs in being a smaller size, the pronotal disc having
2 longitudinal stripes of dense pubescence and the surface of the disc being smooth; elytral apex bispinose.
Material examined
Holotype. . TAS: Sprent (41º16'S, 146º09'E), Coll. no. 12588, bearing a name label on which ‘TYPE’
was written in red (SAM).
Paratypes. 1, 2 . TAS: 1 , same data as above but reared from hazelnut tree, Coll. no. 5705 (SAM); 1
, Ulverstone (41º09'S, 146º10'E), reared from hazelnut tree (SAM); 1 , same data as above but (AM).
Other material examined. 12 , 5 . TA S: 1 , Tasmania, A. Simson, Coll. no.3648 (UQIC); 1 , Strahan
(13º32'S, 143 º19'E), ii.1924, G. H. Hardy, F. E. Wilson Collection (UQIC); 1 , Ulverstone (41º09'S,
146º10'E), reared from Hazelnut tree, 11.xii.1918, C. French (NMV); 2 , Melaleuca (43º25'S, 146º09'E),
17.ii.1990, E. S. Nielsen & P. B. McQuillan (ANIC); 3 , same locality as above but nr Bathurst Harbour, 18–
21.ii.1991, at light, A. Calder & W. Dressler (ANIC); 1 , Claytons (43º23'S, 146º08'E), Bathurst Harbour,
Tas, 14.ii.1990, E. S. Nielsen & P. B. McQuillan (ANIC); 1 , same locality as above but 16.iii.1991, M.
Horak (ANIC); 1 , same locality as above but 12.iv.1991, E. D. Edwards (ANIC); 1 , 4 km W Orford,
(42º33'S, 147º52'E), 27.i.1983, at light, J. C. Cardale Collection (ANIC); 1 , Commandant Creek (42º28'S,
145º25'E), 15.ii.1971, A. Neboiss (NMV); 1 , Arve River (43º08'S, 146º48'E), 23.ii.1963, I. F. B. Common
& M. S. Upton (ANIC). 1 , no locality label, TAS (NMV). Locality unknown: 1 , 11.xii.1918, C. French
(NMV); 1 , no data (SAM).
Uracanthus gigas Lea, 1916
(Figs 36, 74, 102)
Uracanthus gigas Lea, 1916: 372.—McKeown, 1947: 64.
Description
Male. Body length, 40.61–53.78 mm; width, 8.87–10.25 mm.

Zootaxa 1569 © 2007 Magnolia Press · 39
REVISION OF URACANTHUS
Colour (Fig. 36). Head, prothorax, legs and elytra blackish brown to black; head and thorax with fairly
long golden or pale hairs and the remaining body with fairly dense short white pubescence.
Head. Postclypeus semicircular or subtriangular, with fairly dense punctures; frontoclypeal suture deep
and narrow in middle; frons and vertex with fairly coarse punctures; distance between lower lobes of eyes
2.0–2.09 × as long as distance between upper lobes of eyes; distance between upper lobes of eyes 2.09–2.1 ×
as long as distance between eyes on ventral side; genal length 0.31–0.36 × as long as head width immediately
below eyes. Antennae longer than body; segments 4–10 slightly produced on one side at apex.
Thorax and abdomen. Pronotum 0.83–0.89 × as long as width, rounded at side; posterior margin 1.34–
1.52 × as wide as anterior margin; pronotal disc and sides uneven but no obvious nodules on disc. Scutellum
semicircular, with dense pubescence. Elytra 5.16–6.74 × as long as prothorax and 3.11–4.01 × as long as
shoulder width; each elytron with 3–4 raised longitudinal carinae; elytra with fairly dense, fine punctures
throughout; elytral apex bispinose. Apex of terminal sternite truncate or slightly emarginate.
Male terminalia. Apex of ventral median lobe rounded; ventral lobe slightly longer than or as long as dor-
sal lobe; internal sac with dense long simple spines almost throughout except short basal and terminal areas
(Fig. 74a). Eighth sternite obliquely truncate, side sharply pointed and apex heavily emarginated, with dense
long setae; ventral surface with sparse simple spines and cloud-like processes (Fig. 74b). Eighth tergite round
at sides and rounded or slightly truncate at apex, with fairly dense basally-forked spines on dorsal surface
(Fig. 74c). Paramere long, 2.8–2.9 × as long as wide, cylindrical in shape, apex rounded with long and short
setae (Fig. 74d).
Female. Body length, 31.65–60.56 mm; width, 11.10–12.30 mm
Body broader with robust abdomen; antennae and legs shorter. Pronotum 0.78–0.91 × as long as width.
Elytra 3.28–4.04 × as long as shoulder width and 6.45–7.06 × as long as prothorax. Body colour may be
brighter in females of some specimens
Ovipositor and spermatheca. Ovipositor relatively long; styli arising terminally with short hairs (Fig.
74e). Spermatheca relatively short and robust, moderately curved; spermathecal gland arising at base (Fig.
74f).
Distribution
Western Western Australia and eastern Queensland (Fig. 102).
Biology
Known hosts are Brigalow (Acacia harpophylla). Adults were attracted to MV light and were collected
during May to December, and February to March.
Comments
This species resembles U. pseudogigas sp. nov. but differs in having the elytral apex bispinose and the
ventral median lobe of the aedeagus rounded.
Material examined
Holotype. . QLD: Kingaroy (26º32'S, 151º50'E), French Collection (NMV)
Paratypes. 1 , 1 . QLD: 1 , same data as above but Coll. no. 18529 (SAM). WA: 1 , locality
unknown (NMV).
Other material examined. 31 , 6 . QLD: 1 , Brialow Res. Site 1 (28º48'S, 149º45'E), softwood scrub,
160m, 18.xi.2000, at MV light, A. Ewart, Coll. no. 9595 (QM); 1 , East of Wonga Hill, 20 m, site 3 (26º04'S,
150º49'E), 11.xii.2001, vine scrub, at MV light, Cook & Wright (QM); 1 , Coomooboolaroo (23º54'S,
149º30'E), C. Bainara (SAM); 1 , Edungalba (23º43'S, 149º50'E), 1939, E. E. Adams, terminalia slide no. f-
030414-3 (QM); 1 , N Taroom (25º36'S, 149º46'E), 2.x.1991, MV light, G. Danniels (UQIC); 1 , Mut-

THONGPHAK & WANG
40 · Zootaxa 1569 © 2007 Magnolia Press
taburra, E. Monteith, terminalia slide no. m-030414-5 (UQIC); 1 , Hughenden RW, Bommbrick snap stack,
19.v.1934, terminalia slide no. f-030414-1 (UQIC); 1 , Coroorok R (UQIC); 2 , SE by E of Lifracombe,
17.iii.1971, R. A. H. Davis (ANIC); 1 , same data as above but 19.iii.1971 (ANIC); 1 , Edungalba, hatched
from brigalow, i.1946, A. & B. Smith (ANIC). 1 , same locality as above but 1944 (ANIC). WA: 1 , Wadg-
ingarra (WAM); 1 , Gullewa (WAM); 1 , same data as above but (QM); 1 , Boologooro HS (24º39'S,
113º42'E), 27–29.iix.1980, C. A. Howard & T. F. Houston (WAM); 1 , same data as above but terminalia
slide no. m-030414-7 (WAM); 1 , Mooka stn upper Gasecoyne, 10.x.1971; P. Bridge & M. Thomas, termina-
lia slide no. m-030414-4 (WAM); 1 , Forrest, iix.1873 (WAM); 1 , Middalya, 4.ii.1904, T. Weir (NMV); 1
, Carnarvon, Yalbalgo stn., 12.ix.1950, A. Snell (WADA); 1 , same data as above but terminalia slide no.
m-030414-6 (WADA); 1 , same data as above but 21.iix.1950, terminalia slide no. f-030414-2 (WADA); 1 ,
Carnarvon, Jimba Jimba, ix.1949, A. Snell (WADA); 4 , N by E Singlatan Mt. (29º21'S, 117º20'E),
28.ix.1981, J. C. Cardale (ANIC); 8 , NE Ninghan HS, 26 m, nr Lake Moore, 23.x.1963, at light, A. V. Will-
iamson (ANIC).
Uracanthus pseudogigas, sp. nov.
(Figs 37, 75, 102)
Description
Male. Body length, 35.86–49.88 mm; width, 8.27–10.83 mm.
Colour (Fig. 37). Head, prothorax, legs and basal 1/10 elytra dark reddish brown to blackish brown;
remaining elytra pale yellowish brown; antennae reddish brown. Head and thorax with dense, long, pale yel-
lowish pubescence except a glabrous ψ-shaped or ‡-shaped mark on pronotal disc; elytra and abdomen with
dense, short, pale yellowish pubescence. The ψ-shaped glabrous mark on pronotal disc may be complete or
incomplete.
Head. Postclypeus semicircular, flattened, with fairly dense punctures; frontoclypeal suture deep and nar-
row in middle; distance between lower lobes of eyes 1.48–1.69 × as long as distance between upper lobes of
eyes; distance between upper lobes of eyes 1.04–1.17 × as long as distance between eyes on ventral side;
genal length 0.40–0.42 × as long as head width immediately below eyes; Antennae longer than body; seg-
ments 4 to10 flattened , produced on one side at apex.
Thorax and abdomen. Pronotum 1.02–1.53 × as long as width, with a small rounded process at each side;
posterior margin 1.19–1.29 × as wide as anterior margin; pronotal disc finely punctate with two slightly raised
nodules in middle. Scutellum semicircular and glabrous. Elytra 4.15–5.44 × as long as prothorax and 3.16–
3.46 × as long as shoulder width; each elytron with distinctly raised 4 longitudinal carinae; apex with an acute
spine at suture. Apex of terminal sternite truncate.
Male terminalia. Apex of ventral median lobe widely emarginate and apex of dorsal lobe pointed; ventral
lobe slightly shorter than dorsal lobe; internal sac with long simple spines throughout except short areas at
base and terminal (Fig. 75a); Eighth sternite rounded at side, emarginate at apex, with long and fairly long
setae arising terminally; ventral surface with fairly dense short setae and dense cloud-like processes (Fig.
75b). Eighth tergite emarginate at apex; basal surface with fairly dense simple and multi-branched spines (Fig.
75c). Paramere 3.8–4.0 × as long as wide, cylindrical in shape, apex rounded with long and short setae (Fig.
75d).
Female. Unknown.
Distribution
Southeastern Western Australia, southern Northern Territory and southern and eastern South Australia
(Fig. 102).

Zootaxa 1569 © 2007 Magnolia Press · 41
REVISION OF URACANTHUS
Biology
Known host is Acacia sowdenii. Adults were collected by pitfall and light traps during September–
November.
Comments
This new species resembles U. gigas but differs in having the elytral apex with only one spine at the
suture; ventral median lobe of aedeagus emarginate. This species was considered a variant of U. gigas by
Rondonuwu and Austin (1988).
Material examined
Holotype. . SA: Mulgathing HS, on bushes forming a hedge, F. & S. Aslin, terminalia slide no. m-
040529-1 (SAM).
Paratypes. 24 . SA: 1 , Roxby Downs, from camp site, 26.x.1976, L. Williams (SAM); 1 , Woodland,
H of Roxby, 25.x.1978, at light, Acacia sowdenii, L. D. Williams (SAM); 1 , S Moonaree HS , 32 km,
Gawler Ra, 4.x.1988, at light, J. A. Forrest (SAM); 1 , same data as above but terminalia slide no. m-
040529-2 (SAM); 1 , S Pimba, 106 km, 22.ix.1960, Austa (SAM); 1 , Arcoona W May Hill, 4–9.xi.1996,
pitfall (SAM); 1 , Wirraminna, 27.x.1953, W. B. Tindale, E. S. J. (SAM); 1 , HW of SA, H. Basedow & M.
W. Grace (SAM), terminalia slide no. m-040529-3 (SAM); 1 , SW Table Hill, 7.2 km, Innamincka, 4–9
.xi.1996, Slony D. S (SAM); 1 , Hast Bluff, C. A, 5.ix.1957, N. B. Tindale (SAM). WA: 1 , Gill Pinnacle,
Mural cres, 10.xi.1963, at light, P. Aitken & N. B. Tindale (SAM); 1 , Lauviam, Kalgoorlie, 1898 (SAM); 1
, E of Terhan RH, 11 mile, 15.x.1960, Chinnick, McCade, Corby (ANIC). NT: 2 , 4 mile S of Levin Mt.,
20.ix.1963, P. Ranford (ANIC); 1 , S by 41 km E of Alice Spring area , 4.x.1978, Upton & Barret (ANIC); 6
, 60 km Hamilton Downs Sin. NW Alice Spring, 11.x.1976, at light, G. Griffin (ANIC); 1 , Alice Spring,
vii–ix.1957, N. Mollett, terminalia slide no. m-040529-4 (SAM). Locality unknown: 1 , no data (NMV).
Uracanthus acutus Blackburn, 1889
(Figs 38, 76, 103)
Uracanthus acutus Blackburn, 1889: 451.—Froggatt, 1898: 46; Allen, Blunno, Froggatt and Guthrie, 1898a: 1038;
Dixon, 1908:77; Aurivillius, 1912: 147; McKeown, 1947: 62; Duffy, 1963:117.
Description
Male. Body length, 11.81–20.45 mm; width, 2.13–3.10 mm.
Colour (Fig. 38). Body reddish brown with head, thorax, antennae segments, scutellum, and basal 1/5 to
1/3 of elytra often darker. Head with dense golden pubescence. Pronotal disc with a broad longitudinal stripe
of very dense golden pubescence on each side and two longitudinal stripes of relatively sparse pubescence in
middle; side with a narrow longitudinal stripe of very dense golden pubescence near ventral side. Elytron with
dense short golden pubescence, particularly on apical ½. The longitudinal stripes on middle pronotal disc may
vary from very distinct to vague.
Head. Postclypeus semicircular, slightly convex, with fairly dense coarse punctures; frontoclypeal suture
deep and narrow in middle; distance between lower lobes of eyes 2.0 –2.33 × as long as distance between
upper lobes of eyes; distance between upper lobes of eyes 0.80–0.84 × as long as distance between eyes on
ventral side; genal length 0.26–0.31 × as long as head width immediately below eyes; Antennae shorter than
body; segments 5–10 subcylindrical and slightly produced on one side at apex.
Thorax and abdomen. Pronotum 1.42–1.57 × as long as width, rounded at side; posterior margin 1.24–
1.53 × as wide as anterior margin; pronotal disc finely punctate with two very feebly raised nodules in middle
area; posterior ¼ of disc weakly, transversely rugose. Scutellum semicircular, glabrous. Elytra 3.70–3.87 × as

THONGPHAK & WANG
42 · Zootaxa 1569 © 2007 Magnolia Press
long as prothorax and 3.81–4.15 × as long as shoulder width; each elytron with 3 longitudinal carinae; basal 1/
3 of elytra with dense coarse punctures and 2/3 of elytra with fine punctures; apex sharply pointed. Apex of
terminal sternite truncate or truncate with a small notch.
Male terminalia. Apex of ventral median lobe slightly pointed and apex of dorsal lobe rounded; ventral
lobe slightly longer than dorsal lobe; internal sac with dense simple and long spines except short unspined
basal and terminal areas (Fig. 76a). Eighth sternite obliquely truncate at side, widely emarginate at apex, with
long setae arising terminally; ventral surface with fairly dense cloud-like processes (Fig. 76b). Eighth tergite
rounded at sides with a small notch at apex; surface with fairly dense, simple and short spines in middle area
(Fig. 76c). Paramere long and thin, 2.5–2.6 × as long as wide, cylindrical in shape, apex rounded with long
and short setae (Fig. 76d).
Female. Body length, 18.60–23.87 mm; width, 3.8–4.49 mm.
Abdomen more robust, antennae and legs shorter; distance between lower lobes of eyes less than 2 × as
long as distance between upper lobes of eyes; elytra 4.33–4.95 × as long as prothorax and 3.80–4.06 x as long
as shoulder width.
Ovipositor and spermatheca. Ovipositor very short; styli arising terminally with short hairs (Fig. 76e).
Spermatheca slightly curved; spermathecal gland arising near base (Fig. 76f).
Distribution
Western, southern and eastern Victoria, eastern New South Wales, southern and southeastern South Aus-
tralia, and northeastern and southeastern Queensland (Fig. 103).
Biology
Hosts are peach, plum and apricot, Amygdalus persica, Armeniaca vulgaris, Acacia dealbata and A. mol-
lissima, Pomaderris spp. and Loranthus sp. Adults are found on Eucalyptus foliage. Larvae at first bore
towards and then into the heartwood, but later tunnel right round beneath the bark, causing the latter to wilt
and die, and finally break off. Adults were collected from December to February. Adults are attracted to light.
Comments
This species resembles U. loranthi but differs in having the body more slender, the pronotal disc with
transverse rugae only in the area near the posterior margin, and the apex of the elytra sharply pointed.
Material examined
Paratypes. 1 , 1 . VIC: 1 , Victoria, Coll. no. I. 5488 (SAM); 1 , locality unknown, abdomen and left
antennae missing, Coll. no. 2909 (SAM). The holotype is supposed to be in BMNH according to McKeown
(1947) but could not be located.
Other material examined. 84 , 19 . QLD: 1 , Gap Creek, 6 mile N of Bloomfield R., N of QLD, 13–
14.xi.1965, G. Monteith (QM); 1 , Bunya Mt (26°50’S, 151°33’E), 5 km NW of Mt Mowbullan, 3350 ft.
8.i.1970, at light, Britton, Holloway & Misko (ANIC); 1 , Brisbane, 3.ii.1962, B. F. Stone (UQIC); 1 , QLD
(NMV). NSW: 2 , Pt. Lookout via Ebor, Nothofagus Forest, 22.i.1967, B. Cantrell (UQIC); 1 , Jenolan
Caves, 18.i.1985, George Hangay (ANIC); 1 , Wiangaree state Forest, via Kyogle, 28.xi.1970, G. Monteith
(UQIC); 2 , West Pymble, near Sydney, 5.xi.1985, at black light, D. J. Scambler, (AM); 1 , same as above
but 10.xi.1987 (AM); 1 , Willougby, Sydney, 10.xi.1986, D. K. McAlpine (AM); 2 , Ashfield, 26.x.1980,
D.A. Doolan (AM); 1 , National Park, i.1965, MV light, D. K. McAlpine (AM); 1 , Rydalmere, 16.xi.1976,
at light trap, G. R. Brown (ASCU); 1 , same locality as above but 27.x.1971, at mercury vapour light
(ASCU); 1 , Prospect, 11.xii.1911 (AM); 1 , Arrawarra, H. J. Carter (NMV); 1 , Beecroft (33°44’S,
151°03’E), 12.xii.1967, at mercury vapour light, O. M. Williams (ASCU); 1 , Clovelly, 2.xi.1956, on Euca-
lyptus foliage, W. E. Wright (ASCU); 1 , same locality as above but 15.xi.1953 (ASCU); 6 , Kiama,

Zootaxa 1569 © 2007 Magnolia Press · 43
REVISION OF URACANTHUS
xi.1958, L. Cady, (ASCU); 2 , same locality as above but xi.1959 (ASCU); 1 , same data as above as above
but xi.1960 (ASCU); 1 , same locality as above but 5.xi.1921, Allowire (MAM); 2 , Minnamurra Falls,
xii.1968, C. E. Chadwick (ASCU); 1 , Mt. Keira, 23.ii.1966, at mercury vapour light, V. J. Robinson
(ASCU); 1 , Sydney, xi.1937, A. Musgrave (AM); 1 , Sydney (MAM); 1 , same locality as above but
(NMV); 1 , same locality as above but 10.xi.1968, C. E. Chadwick (ASCU); 1 , no data, NSW (MAM); 2
, same as above but (NMV). ACT: 2 , Black Mt., Canberra, ACT, 25.i.1968, M. S. Upton (ANIC); 1 ,
same as above but 11.i.1972, K. R. Pullen (ANIC); 1 , same as above but 16.i.1952, no abdomen, L. J.
Chimick (ANIC). VIC: 4 , Red Hill, 10.ii.1970, Dr. Holmes (ANIC); 1 , Kiata, x.1938, F. E. Wilson
(NMV); 1 , Melton, xii.1937, bred ex Loranthus, F. E. Wilson (ANIC); 1 , Hayfield (36°56'S, 145°45'E),
16.xi.1921, ex Acacia (NMV); 1 , Melbourne, French (SAM); 1 , Mordialloc, 1.i.1931, A. L. Brown
(NMV); 1 , Cheltenham, C. Oke (NMV); 2 , Hazelwood, 29.xii.1948, A. L. Brown (NMV); 1 , Saint
Kilda, C. Oke (NMV); 1 , Yinnar, xii.1948, A. L. Brown (NMV); 1 , 3km SE Taggerty, Little River,
6.i.1972, Neboiss (NMV); 1 , Mitta Mitta River, Neboiss & Malcolm (NMV); 1 , Mitcham, xi.1950, A. L.
Brown (NMV); 1 , Thompson Reserve, 12.xii.1973, La Trobe R. Survey (NMV); 1 , Rainbow Creek, 4 km
NNW of Cowarr, 4.xii.1976, A. A. Calder (NMV); 1 , N. Mallee, Dixon (NMV); 1 , Moe, 5.i.1949, C. G. L.
Gooding (NMV); 2 , Studley Park, 12.vii.1916, bred from Pomaderris, Dixon (NMV); 1 , Sea Lake,
xi.1912, G. C. Goodie (NMV); 1 , Gippsland Lake Park, (NMV); 1 , Richmond, 26.xii.1904 (NMV); 1 ,
Victoria (MAM); 4 , 4 , same as above but (NMV); 4 , Victoria, ex Acacia, xi.1919 (NMV); 2 , same
data as above but xi.1905 (NMV); 3 , same data as above but xi.1912 (NMV); 1 , same data as above but
xi.1916 (NMV); 2 , Vic, 6.xi.1909, ex Pomaderris, (NMV); 1 , same locality as above but x.1907, ex Aca-
cia mollissima (NMV). SA: 1 , Kurlge Blackwood, 850 ft. MV light, 5.xi.1960, N. B. Tindale (SAM); 1 ,
20 km S of Wirraminna HS, at light, 2.xi.1975, J. A. Herridge (SAM); 1 , Burrawing, 10.i.1933 (AM).
Locality unknown: 1 , 1 , Jemey Moryma, 19.x.1896 (NMV); 1 , 11.xi.1975, light trap, V. P. R. I.
(VAIC); 1 , no data, no abdomen (WINC). 1 , Australia, H. E. Cox Collection 1916 (HMO); 1 , no data
(UQIC).
Uracanthus loranthi Lea, 1916
(Figs 39, 77, 104)
Uracanthus loranthi Lea, 1916: 380.—McKeown, 1947: 65; Duffy, 1963: 117.
Description
Male. Body length, 12.58–15.70 mm; width, 2.13–2.87 mm.
Colour (Fig. 39). Body reddish brown with head, thorax, antennae, scutellum, and basal elytra often
darker. Head with fairly dense golden pubescence. Pronotum with a wide longitudinal stripe of dense golden
pubescence on each side of disc, a median wide longitudinal stripe of sparse pubescence (may be divided into
two by a narrow median glabrous line), and a narrow longitudinal stripe of dense golden pubescence on each
side near sternum. Elytral disc covered with fairly dense short golden pubescence particularly on apical ½.
Body colour varies from reddish brown to yellowish brown.
Head. Postclypeus semicircular, flattened, with fine punctures; frontoclypeal suture deep and narrow in
middle; distance between lower lobes of eyes 1.92–2.0 × as long as distance between upper lobes of eyes; dis-
tance between upper lobes of eyes 0.68–0.70 × as long as distance between eyes on ventral side; genal length
0.27–0.30 × as long as head width immediately below eyes; Antennae slightly shorter than body; segments 5–
10 subcylindrical and slightly produced on one side at apex.
Thorax and abdomen. Pronotum 1.22–1.46 × as long as width, strong rounded at side; posterior margin
1.05–1.06 × as wide as anterior margin; pronotal disc feebly binodulose in middle area and finely punctate;

THONGPHAK & WANG
44 · Zootaxa 1569 © 2007 Magnolia Press
disc and side strongly rugose transversely. Scutellum semicircular, glabrous. Elytra 3.72–4.36 × as long as
prothorax and 3.67–4.35 × as long as shoulder width; each elytron with 3 feeble longitudinal carinae; basal
half of elytra with dense coarse punctures and apical half with fine punctures; apex truncate or slightly emar-
ginate with a small process at margin and suture, respectively. However, the elytral apex may be clearly bispi-
nose in some specimens. Apex of terminal sternite truncate.
Male terminalia. Apex of ventral median lobe rounded to slightly pointed and and apex of dorsal lobe
rounded; dorsal lobe almost as long as ventral lobe; internal sac with dense simple and long spines except
short unspined basal and terminal areas (Fig. 77a). Eighth sternite obliquely truncate at sides, truncate and
slightly emarginate at apex, with long and fairly long setae arising terminally; ventral surface with sparse
cloud-like processes (Fig. 77b). Eighth tergite shallowly emarginate at apex, with dense setae arising from
apical ½; surface with fairly dense multi-branched spines near base and fairly dense simple spines in mid area
(Fig. 77c). Paramere 2.0–2.12 × as long as wide, cylindrical in shape; apex rounded with long and short setae
(Fig. 77d).
Female. Body length, 12.86–17.41 mm; width, 2.48–3.12 mm.
Abdomen more robust, antennae and legs shorter; the median longitudinal glabrous line on pronotal disc
more obvious and broader; elytra 3.44–4.72 × as long as prothorax and 3.68–4.21 × as long as shoulder width.
Ovipositor and spermatheca. Ovipositor short; styli arising terminally with short hairs (Fig. 77e). Sper-
matheca short and robust, slightly curved; spermathecal gland short, arising from basal 1/3 (Fig. 77f).
Distribution
Eastern, southeastern and southern Queensland, eastern New South Wales, Southern and central Victoria,
southeastern South Australia, southern Western Australia, northern & central Australian Capital Territory,
and southern Northern Territory (Fig. 104).
Biology
Known hosts are Loranthus sp., Acacia sp. and mistletoe on Casuarina sp. Adults were collected between
September and April by light trap and beating from Loranthus sp.
Comments
This species closely resembles U. acutus but differs in having the elytral apex truncate or bispinose and
the male eighth tergite truncate and slightly emarginate.
Material examined
Holotype. . VIC: Melbourne, 15.v.1916, Coll. no. T-8880; bearing a red holotype label; left antenna
damaged; right antenna and left mid tibia and tarsus missing (NMV).
Paratypes. 2 . VIC: 1 , Victoria, Coll. no. 9084, 3 segments of left antenna and 2 segment of right
antenna, and right elytron missing (SAM); 1 , locality unknown, i.1908, Coll. no. J. 5693 (SAM).
Other material examined. 33 , 27 . QLD: 1 , Toowoomba, 10.xi.1974, Macqueen (ANIC); 1 , Vallis,
v.1984 (ANIC); 1 , W of Shore, Tinaroo, 7.xi.1966, at light, E. Britton (ANIC); 1 , 26 km N of Mazeppa,
National Park, 24.iii.1993, G. Monteith (QM); 1 , Booloumba Creek, Conondale Ra., 29–30.x.1998, G. B.
Monteith (QM); 1 , Nocundra Waterhole, Wilson River, 20.x.1968, J. A. L. Watson (ANIC); 1 , Lake
Broadwater via Dalby, 26–28.i.1985, G. Monteith & G. Thompson (QM); 1 , Mt. Mistake Plateau via Goom-
burra, 21–22.xi.1987, G. B. Monteith (QM); 1 , 350 m, Mt. Hayward (20°20'S, 148°45'E), 19–20.xi.1992,
Monteith, Thompson & Janetzki (QM); 1 , Brisbane, R. Illidge (UQIC). NT: 1 , 39 km E of Alice Springs,
26.xi.1978, Upton & Barret (ANIC). NSW: 1 , 65 NW Nyngan, 21.x.1949, E. F. Riek (ANIC); 1 , Jenolan
Caves, 18.i.1985, George Hangay (ANIC); 1 , Burrapine (30°42'S, 152°38'E), 10.i.1933 (AM). ACT: 1 ,
Black Mtn, Canberra, ACT, 11.i.1972, K. R. Pullen (ANIC); 2 , same as above but 25.i.1968, M. S. Upton

Zootaxa 1569 © 2007 Magnolia Press · 45
REVISION OF URACANTHUS
(ANIC); 1 , same as above but 16.i.1952 (no abdomen), L. J. Chimick (ANIC). VIC: 1 , 2 , Melton,
i.1973, bred from Loranthus sp., F. E. Wilson (NMV); 1 , Burnley, 28.i.1975, at light trap, V. P. R. I. (VAIC);
1 , same as above but xii.1975 (VAIC); 1 , same as above but 24.i.1974 (VAIC); 1 , South Morang
(37°39'S, 145°05'E) (NMV); 1 , Melbourne, French (SAM); 1 , 1 , Kew (37°48’S, 145°02’E), ex. Loran-
thus, 11.i.1920 (NMV); 1 , Kiata, Victoria, x.1928, F. E. Wilson (NMV); 4 , Red Hill, 10.ii.1970, Dr.
Holmes (ANIC); 1 , Moorooduc (38°14'S, 145°06'E), x.1914, J. E. D. (NMV); 1 , Mordialloc, 19.xi.1930,
A. L. Brown (NMV); 1 , same data as above but 1.ii.1931 (NMV); 1 , Parwan, ex. Loranthus, xi.1911
(NMV); 1 , same locality as above but ex mistletoe on Casuarina sp., 5.ix.1915 (NMV); 1 , Birchip, J. C.
Goudie (NMV); 1 , Studley Park (NMV); 1 , 1 , Victoria (NMV). SA: 2 , 850 ft. Kurge, Blackwood,
17.xii.1968, at MV light, N. E. Tindale (SAM); 1 , 20 km, S. Wirraminna, 2.xi.1975, at light, J. A. Herridge
(SAM). WA: 1 , Mcdermid Rock (32°01'S, 120°44'E), 27.xi–x.1978, T. F. Houston et al. (WAM). Locality
unknown: 2 , 4 , no locality data, ex. Loranthus, i.1905 (NMV); 1 , no locality data, 24.x.1920, bred out
mistletoe, D. Millian (NMV); 1 , no locality data, ex Acacia, i.1910 (NMV); 1 , 1 , no data (NMV); 1 ,
no data (AM); 3 , no locality data, ex mistletoe, xii.1908 (NMV).
Uracanthus cupressianus Rondonuwu and Austin, 1988
(Figs 40, 78, 105)
Uracanthus cupressianus Rondunuwu and Austin, 1988: 110.
Description
Male. Body length, 12.76–18.06 mm; width, 2.11–3.20 mm.
Colour (Fig. 40). Body light reddish brown with head, thorax, antennae, scutellum, femora, and basal
elytra often darker. Head with dense, long, golden pubescence. Pronotal disc with 2 broad longitudinal stripes
of dense, long, golden pubescence, a narrow, longitudinal, glabrous median line between stripes, and 4 vague
sparsely haired or glabrous spots near the median line; pronotum with a narrow longitudinal stripe of dense,
long, golden pubescence on each side near sternum; ventral surface of prothorax with sparse golden pubes-
cence. Elytra covered with dense, short, golden pubescence but pubescence near prothorax much longer. Body
colour may vary from yellowish brown to reddish brown. The glabrous median longitudinal line on the prono-
tal disc may vary from narrow to wide.
Head. Postclypeus subtriangular and flattened, with sparse punctures; frontoclypeal suture deep and nar-
row in middle; distance between lower lobes of eyes 1.72–1.82 × as long as distance between upper lobes of
eyes; distance between upper lobes of eyes 0.76–0.94 × as long as distance between eyes on ventral side;
genal length 0.26–0.32 × as long as head width immediately below eyes. Antennae shorter than body; seg-
ments 5–10 subcylindrical.
Thorax and abdomen. Pronotum 1.26–1.54 × as long as width, rounded at side; posterior margin 1.14–
1.32 × as wide as anterior margin; pronotal disc finely punctate; disc and sides strongly rugose transversely.
Scutellum semicircular, glabrous. Elytra 4.28–5.47 × as long as prothorax and 4.13–4.44 × as long as shoulder
width; each elytron with 3 longitudinal carinae; elytra with coarse punctures on basal 1/3 and fine punctures
on the rest 2/3; elytral apex rounded or slightly pointed. Apex of terminal sternite slightly emarginate.
Male terminalia. Apex of ventral median lobe pointed and apex of dorsal lobe shallowly emarginate; dor-
sal lobe slightly longer than ventral lobe; Spined region of internal sac divided into two sections; first section
longer than basal unspined region, with sparse multi-branched spines at base and very dense long simple
spines on the remaining part; second section about 3 × shorter than first section, with sparsely simple spines; a
short unspined gap between sections (Fig. 78a). Eighth sternite, rounded at sides, widely emarginate at apex,
long and fairly long setae arising terminally; ventral surface with sparse cloud-like processes (Fig. 78b).

THONGPHAK & WANG
46 · Zootaxa 1569 © 2007 Magnolia Press
Eighth tergite long and narrow, emarginate at apex, with fairly dense simple spines in middle area (Fig. 78c).
Paramere 3.0–3.1 × as long as wide, cylindrical in shape, apex rounded with sparse long and short setae (Fig.
78d).
Female. Body length, 15.29–19.67 mm; width, 2.68–3.87 mm.
Abdomen more robust, antennae and legs shorter; the median glabrous longitudinal line at pronotal disc
broader; elytra 4.51–5.13 × as long as prothorax and 3.92–4.12 × as long as shoulder width.
Ovipositor and spermatheca. Ovipositor relatively long in the genus; styli arising terminally with short
hairs (Fig. 78e). Spermatheca heavily curved; spermathecal gland arising near base (Fig. 78f).
Distribution
Southeastern South Australia (Fig. 105).
Biology
Known hosts are Cupressus sempervirens, C. macrocarpa, and C. arizonica. Adults were collected from
December to February.
Comments
This species resembles U. acutus but differs in having the pronotal disc with denser pubescence and more
transverse rugae on disc, and the apex of the elytra not strongly pointed.
Material examined
Holotype. . SA: Glenelg, reared ex Cupressus sempervirens, 5.x.1986, S. Rondunuwu, bearing a circular
type label (SAM).
Paratypes. 15 , 10 . SA: 12 , 9 , North of Glenelg, reared ex Cupressus sempervirens, 5.x.1986, S.
Rondunuwu (WINC); 3 , 1 , same as above but (SAM).
Other material examined. 4 , 5 . SA: 1 , 1 , Adelaide, adult 1960 (larvae iii.1959), reared ex Cupres-
sus macrocarpa, F. D. Morgan Coll. (WINC); 1 , 1 , same locality as above but R. Brown (SAM); 1 ,
same locality as above but ex limb of cypress pine, 17.x.1984 (WINC); 1 , Wood Ville West, xi.1963, reared
ex Cupressus arizonica, B. Rankine Coll. (WINC); 1 , near Torreng Bridge 26.x.1973, H. Mincham (SAM);
1 , Warradale, 16.xi.1973, MV light, B. P. McQuillan(SAM); 1 ?, Plympton Park, 8.xi.1961, boring in
Cupressus sp.,(no abdomen) (SAM). Locality unknown: 1 (WINC).
Uracanthus minatus Pascoe, 1866
(Figs 41, 79,105)
Uracanthus minatus Pascoe, 1866: 93.–Aurivillius, 1912: 147; McKeown, 1947: 65.
Description
Male. Body length, 16.36 mm; width, 2.58 mm.
Colour (Fig. 41). Antennae, legs, metasternite, and abdomen blackish brown; remaining body yellowish
brown with head, prothorax and basal 1/3 elytra darker. Head with dense, long, yellowish pubescence. Prono-
tal disc with 2 broad longitudinal stripes of dense, long, long yellowish pubescence, a narrow, longitudinal,
glabrous median line between stripes; side with a narrow longitudinal stripe of very dense golden pubescence
near ventral side; Elytron with very dense short yellowish pubescence.
Head. Postclypeus triangular, convex, with fairly dense punctures; frontoclypeal suture deep and wide in
middle; distance between lower lobes of eyes 1.94 × as long as distance between upper lobes of eyes; distance

Zootaxa 1569 © 2007 Magnolia Press · 47
REVISION OF URACANTHUS
between upper lobes of eyes 0.95 × as long as distance between eyes on ventral side; genal length 0.45 × as
long as head width immediately below eyes. Antennae longer than body; segments 5–10 subcylindrical and
slightly produced on one side at apex.
Thorax and abdomen. Pronotum 1.38 × as long as width, rounded at side with a very small process on
each side; posterior margin 1.33 × as wide as anterior margin; pronotal disc and side strongly rugose trans-
versely. Scutellum semicircular, with dense pubescence. Elytra 4.31 × as long as prothorax and 4.43 × as long
as shoulder width; basal 1/3 of elytra with dense coarse punctures and the rest with fine punctures; apex
rounded.
Male terminalia. Apex of ventral median lobe pointed and apex of dorsal lobe rounded; ventral lobe
shorter than dorsal lobe; spined region of internal sac divided into two sections: first section with dense long
simple spines and sparse multi-branched spines; second section about as long as first section, with sparse sim-
ple spines; no distinct unspined gap between sections (Fig. 79a). Eighth sternite obliquely truncate at sides;
widely emarginate at apex, with long and fairly long setae arising terminally; ventral surface with short simple
spines and fairly dense cloud-like processes (Fig. 79b). Eighth tergite truncate or slightly rounded at apex,
with fairly dense simple spines and multi-branched spines on surface (Fig. 79c). Paramere short, cylindrical in
shape, apex rounded with long and short setae (Fig. 79d).
Female. Body length, 12.73–17.46 mm; width, 2.26–3.52 mm.
Body more robust, antennae shorter than body; distance between lower lobes of eyes less than 2 × as long
as distance between upper lobes of eyes; elytra 3.70–3.75 ×as long as prothorax and 3.96–4.16 ×as long as
shoulder width.
Ovipositor and spermatheca. Ovipositor long; styli short, arising terminally with short hairs (Fig. 79e).
Spermatheca heavily curved; spermathecal gland arising near base (Fig. 79f).
Distribution
Southwestern Western Australia (Fig. 105).
Biology
Hosts unknown. Adults were collected in October and November.
Comment
This species resembles U. corrugicollis but differs in having the antennae, legs and abdomen black; the
elytral apex rounded without any spine.
Material examined
Holotype. . WA: no data, bearing a circular type label, and Pascoe’s hand writing (BMNH).
Other material examined. 1 , 3 . WA: 1 , Lake Grace (33º05'S, 118º24'E), 17.xi.1928, K. & E. Carn-
aby (ANIC); 1 , Thomas River (33º51'S, 123º05'E), 1.xi.1983, K. & E. Carnaby (ANIC); 1 , Marloo Stn.,
Wurarga, 1931–1941, A. Goerling (ANIC); 1 , 18 km E by S of Kalbarri (27º44'S, 114º21'E), Kalbarri
National Park, 19.x.1984, D. C. Y. Rentz (ANIC).
Uracanthus longicornis Lea, 1916
(Figs 42, 80, 105)
Uracanthus longicornis Lea, 1916: 381.—McKeown, 1947: 65.
Uracanthus daviumbus Gressitt, 1951: 24.–Gressitt, 1959: 147, 148. syn. nov.

THONGPHAK & WANG
48 · Zootaxa 1569 © 2007 Magnolia Press
Description
Male. Body length, 13.49–21.76 mm; width, 2.41–4.18 mm.
Colour (Fig. 42). Head and thorax dark reddish to blackish brown; antennae, legs, and basal elytra reddish
brown; remaining parts reddish to yellowish brown. Head and pronotal disc with dense white pubescence;
pronotum with a wide longitudinal stripe of very dense white pubescence on each side near ventral side;
sterna and pleura with fairly dense white pubescence. Elytra with dense white pubescence on disc and a stripe
of very dense white pubescence on margins, gradually diminishing towards apex. Legs and abdomen with
dense white pubescence. Pubescence on head, thorax and elytral margins longer than that on the remaining
parts. The glabrous longitudinal mark on pronotal disc may vary from very narrow to narrow. Body colour
may vary from reddish brown to yellowish brown.
Head. Postclypeus semicircular, flattened and depressed toward to anteclypeus, with fairly dense coarse
punctuate; frontoclypeal suture deep and narrow in middle; distance between lower lobes of eyes 2.09–2.27 ×
as long as distance between upper lobes of eyes; distance between upper lobes of eyes 1.37 –1.57 × as long as
distance between eyes on ventral side; genal length 0.18–0.21 × as long as head width immediately below
eyes. Antennae longer than body; segments 4–10 flattened and strongly produced on one side at apex.
Thorax and abdomen. Pronotum 1.02–1.38 × as long as width, rounded at side; posterior margin 1.11–
1.34 × as wide as anterior margin; pronotal disc and side strongly rugose transversely. Scutellum semicircular,
with dense pubescence. Elytra 3.60–4.40 × as long as prothorax and 3.56–3.90 × as long as shoulder width;
each elytron with 3 feeble longitudinal carinae; basal 1/3 of elytra coarsely punctate and the rest finely punc-
tate diminishing toward apex; apex narrowly emarginate with a sharp spine at suture. However, in some spec-
imens, the elytral apex varies from rounded to narrowly emarginate; the apical spine of the elytron varies from
sharp to none. Apex of terminal sternite truncate.
Male terminalia. Apex of ventral median lobe slightly pointed and apex of dorsal lobe rounded; ventral
lobe distinctly longer than dorsal lobe; spined region of internal sac almost occupying the whole internal sac,
only leaving a very short basal unspined region; spined region divided into 3 sections: first section with fairly
dense short and simple spines and some multi-branch spines; second section longer than first section, with
dense long and simple spines on each side; third section with fairly dense simple spines and multi-branched
spines (Fig. 80a). Eighth sternite rounded or slightly obliquely truncate at sides, strongly emarginate at apex,
with long setae arising terminally; ventral surface with fairly dense cloud-like processes (Fig. 80b). Eighth
tergite rounded to truncate and shallowly emarginate at apex; surface with sparse multi-branched spines near
based and dense short simple spines basal-mid area (Fig. 80c). Paramere robust, 2.0–2.1 × as long as wide,
tapering toward to apex; apex rounded with long and short setae (Fig. 80d).
Female. Body length, 19.78–24.78 mm; width, 3.65–4.63 mm.
Abdomen more robust, antennae and legs shorter; distance between lower lobes of eyes 1.69–2.22 × as
long as distance between upper lobes of eyes; elytra 3.87–4.72 × as long as prothorax and 3.79–3.92 × as long
as shoulder width.
Ovipositor and spermatheca. Ovipositor short; styli arising terminally with short hairs (Fig. 80e). Sper-
matheca robust, heavily curved; spermathecal gland arising near base (Fig. 80f).
Distribution
Northern and southeastern Queensland (Fig. 105) and souther New Guinea.
Biology
Hosts are unknown. Adults were collected by light trap during mid April to July and August to mid Octo-
ber.

Zootaxa 1569 © 2007 Magnolia Press · 49
REVISION OF URACANTHUS
Comments
This species resembles U. cupressianus and U. acutus but differs from those species in having the dis-
tance between the eyes on the ventral side closer, stronger transverse rugae on the pronotal disc, antennae
more robust and longer and the apex of the elytra rounded and acutely spined at the suture.
Material examined
Uracanthus longicornis. Holotype. . QLD: Coll. no. I. 5704; bearing a name label on which ‘TYPE’ was
written in red (SAM). Paratypes. 3 , QLD: 2 , Coll. no. 18539 (SAM); 1 , Endeavour River (SAM).
Uracanthus daviumbus. Holotype. . New Guinea: Lake Daviumbu, Fly River, Papua New Guinea, 19–
30.viii.1936, Archbold Expedition; bearing a red Holotype label (AMNH). Allotype. . New Guinea: same
locality as above; bearing a red allotype label (AMNH).
Other material examined. 72 , 8 . QLD: 9 , Cooktown, 2.viii.1978, J. C. Le Sovef (ANIC); 1 , same
locality as above but (NMV); 2 , Cairns, vi.1969, at MV light, R. Hardicb (ANIC); 1 , same locality as
above but 28.vii.1941 (ANIC); 1 , same locality as above but v.1956, G. B. (ANIC); 1 , same locality as
above but viii.1949, J. G. Brooks (AM); 1 , same locality as above but 1.iv.1935 (QM); 1 , Keating Gap
(15° 30'S, 145°15'E), 3 km SW by S of Cooktown, 16.v.1977, Common & Edward (ANIC); 7 , Dividing
Range, 15 km W. of Catain Billy Creek (15°18'S, 144° 51'E), Cape York Pen., 5–9.vii.1975, J. F. R. Kerr
(ANIC); 1 , Battle Camp Range, 26.VI.1993, at light, P. Zborowski & I. Naumann (ANIC); 7 , 7 km N of
Hope Vale Mission (15°14'S, 145°07'E), Station Creek, 10.v.1981, A. Calder (ANIC); 1 , same locality as
above but 4.x.1980, Tom Weir (ANIC); 2 , 14 km WN of Hope vale Mission (15°16'S, 144°59'E), 8–
10.x.1980, Tom Weir (ANIC); 2 , same locality as above but 7–10.v.1981, A. Calder (ANIC); 1 , same
locality as above but 9.vii.1976, G. B. and S. R. Monteith (QM); 1 , 4 Km of WS Cooktown, 12.v.1980, I. F.
B. Common & E. D. Edwards (ANIC); 1 , 1 Km W. of Cooktown (15°28'S, 145°15'E), 12–13.v.1981, A.
Calder (ANIC); 4 , 15 km WNW of Bald Hill (13°43'S, 143°19'E), Mcllwralth Range, 27.vii.1989, weather
station site, at night, Tom Weir (ANIC); 1 , Batavia Downs (12°40'S, 142°40'E), 22.vii.1992, at light, P.
Zborowski & E. S. Nielsen (ANIC); 4 , Monton (12°27'S, 142°38'E), 27.vi.1992, at light, P. Zborowski & E.
S. Nielsen (ANIC); 1 , Morton, Wenlock R., Cape York Pen., 14.ix.1974, G. B. Monteith (QM); 4 , Turn off
to Captain Billy Landing (11°41'S, 142°42'E), 20.viii.1992, at light, J. Cardale & P. Zboroski (ANIC); 1 ,
Heartland (11°45'S, 142°35'E), 26.vii.1992, P. Zborowski & E. S. Nielsen (ANIC); 2 , Archer River
(13°33'S, 143°03'E), 28.vii.1992, at light, P. Zborowski and E.S. Nielsen (ANIC); 1 , 1 km WN of Rounded
Hill (15°17'S, 145°13'E), 8–10.x.1980, Tom Weir (ANIC); 1 , NE of Mt. Webb (15°03'S, 145°09'E), 1–
3.x.1980, Tom Weir (ANIC); 3 , 2 km S of Horseshae lookout (23°46'S, 149°06'E), Black Down Tab, 23–
24.iv.1981, A. Clader (ANIC); 1 , Mt. Cook National Park (15°29'S, 145°16'E), 11–12.x.1980, T. Weir
(ANIC); 1 , Edae Hill, x.1956, G.B. (ANIC); 1 , 8 km WN of Mt. Molloy, 21–22.v.1980, I. D. Naumann &
T.C. Cardale (ANIC); 1 , Torrus Station, C. T. McNamara (AM); 1 , Brisbane, 25.x.1964, L. Grossett
(UQIC); 1 , Iron Range, Cape York Penn., 30.vi.–4.vii.1977, G. B. Monteith (QM); 1 , Edmonton, 5–
12.ix.1971, R. E. Mascord (ANIC); 1 , 3 km WS of Black Mt. Annan R., 27.vi.1980, T. Weir (ANIC); 1 , 2
, Hann R., 2.vi.1970, J. C. Le Sovef (ANIC). 1 , Monto Q, 17.vi.1978, J. C. Le Sovef (ANIC). Locality
unknown: 1 , N of Queensland (SAM); 1 , 4 , Queensland (NMV).
Uracanthus albatus Lea, 1916
(Figs 43, 81, 106)
Uracanthus albatus Lea, 1916: 378.—Lea, 1917b: 619; McKeown, 1947: 63; Duffy, 1963: 116.
Description
Male. Body length, 13.37–24.56 mm; width, 2.38–5.23 mm.

THONGPHAK & WANG
50 · Zootaxa 1569 © 2007 Magnolia Press
Colour (Fig. 43). Body reddish brown with head, thorax and basal 1/5 of elytra often darker. Head, prono-
tal disc and basal 1/10 of elytra with very dense white and long pubescence; a longitudinal line, more or less
glabrous, starting from frontoclypeal suture and ending near posterior margin of pronotal disc; pronotal disc
with a small glabrous spot on each side of median longitudinal line; each side of pronotum with a wide longi-
tudinal stripe of dense white pubescence near ventral side; sterna and pleura with dense yellowish white
pubescence. Each elytron covered with dense short white pubescence, and 3 narrow stripes of denser pubes-
cence on disc excluding 1 at suture and 1 at margin. Legs covered with fairy dense pubescence; mid and hind
femora with very dense, brush-like brownish hairs on ventral side. Abdomen with fairly dense yellowish
pubescence; each of visible sternites 1–3 with a medial tuft of very dense brownish hairs. Body colour varies
from reddish brown to yellowish brown. Specimens from Victoria differ from other specimens in having
sparser pubescence.
Head. Postclypeus subtriangular and flattened, with dense coarse punctures but mostly hidden in dense
pubescence; frontoclypeal suture deep and narrow in middle; distance between lower lobes of eyes 1.82–2.00
× as long as distance between upper lobes of eyes; distance between upper lobes of eyes 0.86–0.94 × as long
as distance between eyes on ventral side; genal length 0.24–0.37 × as long as head width immediately below
eyes. Antennae slightly longer than body; segments 4–10 flattened and produced on one side at apex.
Thorax and abdomen. Pronotum 1.14–1.25 × as long as width, with a small conical process at each side;
posterior margin 1.25–1.43 × as wide as anterior margin; pronotal disc weakly binodulose in middle area; disc
and side strongly rugose transversely throughout. Scutellum semicircular, with dense pubescence. Elytra
4.07–4.56 × as long as prothorax and 3.34–4.07 × as long as shoulder width; each elytron with 3 longitudinal
carinae on disc; apex distinctly bispinose, with sutural spine usually longer. Apex of terminal sternite more or
less truncate.
Male terminalia. Apices of ventral and dorsal median lobes rounded; ventral lobe slightly shorter than or
as long as dorsal lobe; spined region of internal sac divided into two sections: first section with mixture of
dense simple spines and multi-branched spines along side areas; second section with sparse short simple
spines (Fig. 81a). Eighth sternite strongly obliquely truncate at sides, widely and deeply emarginate at apex,
with long setae arising terminally; ventral surface with dense cloud-like processes (Fig. 81b). Eighth tergite
truncate or slightly emarginate at apex, with some short simple spines and multi-branched spines on mid area
(Fig. 81c). Paramere 1.46–1.48 × as long as wide, cylindrical in shape; apex rounded with long and short setae
(Fig. 81d).
Female. Body length, 17.21–26.55 mm; width, 4.27–5.43 mm.
Antennae and legs shorter; abdomen more robust; femora without brownish brush-like hairs and visible
sternites without medial tufts of very dense brownish hairs.
Ovipositor and spermatheca. Ovipositor short; styli arising terminally with short hairs (Fig. 81e). Sper-
matheca slightly curved; spermathecal gland arising at base (Fig. 81f).
Distribution
Widely distributed in southern and southeastern Queensland, northeastern New South Wales, Victoria,
northern, eastern and southeastern South Australia, and southern Western Australia (Fig. 106).
Biology
Known hosts are Grevillea huegelii and Acacia sp. and A. sowdenii. Adults were collected from July to
December and February to March, by MV light trap.
Comments
This species resembles U. ventralis but differs in having the apex of the elytra distinctly bispinose; head,
pronotal disc and basal 1/10 of the elytra with pubescence distinctly denser, whiter and longer than that on the
remaining parts, and each elytron with only 3 longitudinal stripes of dense pubescence on the disc.

Zootaxa 1569 © 2007 Magnolia Press · 51
REVISION OF URACANTHUS
Material examined
Holotype. . VIC: Dimboola, Coll. no. I. 5492, bearing a name label on which ‘TYPE’ was written in red;
6 segments of left antenna missing (SAM).
Paratypes. 2 , 3 . SA: 2 , Parachilna, Flinder Range (SAM). VIC: 1 , Birchip, 18.v.1937 (no abdo-
men) (SAM); 1 , Leigh Creek, Creewood (SAM); 1 , 5.xi.1908, C. French Collection (NMV).
Other material examined. 70 , 20 . QLD: 1 , 160 m. Brigalow Res. Stn. Site (22°48'S, 149°45'E), 27–
28.x.2000, at MV light, vine scrub, Coll. no. 9804, QM party (QM); 1 , 560m Amphitheare Camp, Expedi-
tion Range National Park (25º12'S,148º59'E), 17.xii.1997, Monteith, Cook, & Thompson (QM); 1 , S Cun-
namulla (28º04'S,145º41'E), 15.x.1968, at light, J. A. Watson (ANIC). NSW: 1 , 20 m Coolabah (31°01'S,
146°41'E), 11.xii.1963, R. Lossin (AM); 3 , Round Hill (31°45'S, 149°40'E), Fauna Reserve, 27.x.1977, MV
lamp, G. Daniels (AM); 1 , Clements (29°49'S, 149°07'E), xii.1928, K. K. S (AM); 2 , 65 NW Nyngan
(31°32'S, 147°10'E), 21.x.1949, E. F. Riek (ANIC); 1 , Nyngan-Bourke, 22.x.1949, S. J. Paramonov
(ANIC); 1 , Bourke (30°04'S, 145°57'E), 24.x.1949, S. J. Paramonov (ANIC); 3 , 4 miles, Sand hills
(35°12'S, 149°33'E), Pt. Parie, 23.xi.1958, T. Greaves (ANIC). VIC: 2 , Kiata (36°21'S, 141°47'E), F. E.
Wilson, F. E. Wilson Collection (NMV); 1 , 2 , same as above but ex Acacia, iv.1916 (NMV); 1 ,
Myrniong (37°37'S, 144°20'E), 528A (AM); 1 , Mallee District (36°21'S, 141°47'E), bred out of Grevillea
huegelii, 26.xi.1915 (NMV); 1 , same as above but xi.1915 (no abdomen), T. S. Dixon (NMV); 1 , same
locality but (WADA); 1 , Victoria (AM); 1 , Victoria (NMV); 1 , Gypsum (35°16'S, 142°23'E), C. Oke
(NMV); 1 , Victoria, no abdomen (NMV); 1 , Birchip (35°58'S, 142°54'E) (no abdomen) (NMV). SA: 1 ,
just west of Lake Eyre (28°22'S, 137°22'E), 17.viii.1985, T. D. Schwaner (SAM); 1 , Port Lincoln (34°43'S,
135°51'E), xi.(no year), S. H. Curnow Collection (SAM); 1 , same as above but 29.ii.1936 (SAM); 3 , 1 ,
Myrtle Spring (30°43'S, 137°09'E), Pine Dam, 13.ix.1964, N. B. Tindale (SAM); 1 , Iron Knob (32°43'S,
137°09'E), S. H. Curnow Collection (SAM); 1 , Baroota (32°55'S, 137°58'E), xi, S. H. Curnow Collection
(SAM); 1 , 8 km NE Mt. Woodroofe (26°17'10″S, 131°48'20″E), Musgrave Range, at light, 14.x.1994, Pit-
jantjatjara Land Survey (SAM); 1 , 20 m S. Cradock (32°04'S, 138°29'E), 20.x.1972, H. E. Evans & T.
Houston (SAM); 3 , Middleback stn. (32°57'S, 137°24'E), 26.x.1983, at light, T. Woodell (SAM); 1 , Great
Victoria Desert (27°52'S, 130°22'E), 100 km N of Cook, leg. CRA team (SAM); 1 , 51 km N Morgan
(34°02'S, 139°40'E), on Canegrass stn. Rd, 1.xi.1995, at light, G. F. Gross (SAM); 2 , Frome River (29°04'S,
137°54'E), Crossing of Birdsville Track nr Marree, 25.x.1966, G. F. Cross (SAM); 1 , 30 km SW Mabel
Creek. HS Sand plain (29º09'40″S, 134º06'30″), at ligh, 25.x.1984, CRA Suvey (SAM); 3 , Simpson Desert
(26°07'S, 138°37'E), vii–viii.1972, C. R. Harris (SAM); 1 , same locality but Poeppel Corner and Birdsville,
6.ix.1986, at night on sand, J. A. Forrest (SAM); 1 , N of Roxby Downs (30°33'S, 136°54'E), Woodland,
25.x.1976, around light in camp in Acacia sowdenii , L. D. Williums (SAM); 2 , Dalhousie Springs (26°27'S,
135°31'E), 6.x.1987, at light, J. A. Forrest (SAM); 2 , Gum Creek (33°35'S, 138°40'E), 6m. N of Mulligan
Springs, 24.ix.1967, to light (SAM); 1 , Farina(30°04'S, 138°16'E), Creek bed , 27.x.1970, G. Gross & E.
Matthews (SAM); 1 , Yorke Peninsula (34°21'S, 137°37'E), Marion Bay, 8.xi.1976, at light, I. Kowanko
(SAM); 1 , Minnie Downs (27°49'S, 134°40'E), NE Corner of SA, L. Reese (SAM); 1 , Mernmerna
(31°39'S, 138°22'E), 1948 (SAM); 1 , Parachilna (31°07'S, 138°23'E), Flinders Range, H. Hule (AM); 2 ,
Flinders Ranges National Park, Dingleys Dell Camp, Oraparinna Creek (32º21'17″S, 138 º42'16″E),
9.x.1997, at light, S. & A. Skevington, C. Lambkin & S. Winterton (UQIC); 1 , Algebuckina (27°54'S,
135°48'E), 50 km SE of Oodmadatta, 8.ix.1988, G. B. Monteith (QM); 2 , Poochera (32°43'S, 134°50'E),
Mallee Woodland, x–xi.1984, R. W. Taylor & R. J. Bartell (ANIC); 2 , 11 km E by SE of Poochera (32°46'S,
134°56'E), SA, 25–26.ix.1981, at MV light, A. A. Calder (ANIC);1 , Birkenshaw Well (28°57'S, 136°07'E),
nr William Creek, x.1992, I. Gee (ANIC); 1 , Lake George (27°20'S, 140°10'E), SA, 13.x.1972, Roffey &
Mitchell (ANIC); 2 , Leigh Creek (30°32'S, 138°28'E), SA, 29.ix.1965, G. C. Gregory (ANIC); 1 , Ang-
orichina Creek Below Hostel (31°07'S, 138°33'E), 22.x.1978, E. B. Britton (ANIC); 1 , 2 , William Creek
(28°54'S, 136°21'E), 6.ix.1990, attracted to light, I. Bunic (ANIC). WA : 1 , 130 m NE Carnegie (26°35'S,

THONGPHAK & WANG
52 · Zootaxa 1569 © 2007 Magnolia Press
120°13'E), 3.xi.1971 (WAM); 1 , 2 mile west of Warburton (26°13'S, 126°39'E), 23.vii.1962, W. H. Buther
(WAM); 1 , Merredin (31°29'S, 118°17'E), 25.ix.1938, A. & T. Winere (QM); 1 , 24–25 km End of Beyon-
die HS (24°47’S, 120°02’E), 17–20.iix.1964, T. F. Houston & B. Hanich, Coll no. 537-18 (WAM); 2 , 2 km
W by SW Muggan RH, 156 km SW of Warburton (27°00’S, 129°19’E), 13–14.ix.1982, T. F. Houston & B.
Hanich (WAM); 1 , Giles (29°24'S, 125°20'E), iix.1965, D. G. Matthews (WAM); 1 , same as above but
1960, B. Gilmour (ANIC); 1 , Cullculli (27°05'S, 118°14'E) station, iix.1972, J. Cark (ANIC); 1 , Cooper
Hills (27°22'S, 126°22'E), 5.viii.1967, W. D. L. Ride (WAM); 1 , Koondra (23°34'S, 120°05'E), Goyder
Lagoon, WA, 30.iix.1975, J. Blyth (NMV); 1 , NE of Streich Mound (30°27'S, 123°41'E), Great Victoria
Desert, Officer Basin, 24–28, ix.1991, R. P. McMillan (WAM). Locality unknown: 1 , John Davidson
(WINC).
Uracanthus ventralis Lea, 1917a
(Figs 44, 82, 106)
Uracanthus ventralis Lea, 1917a: 741.—McKeown, 1947:67.
Uracanthus multilineatus McKeown, 1948: 54. syn. nov.
Description
Male. Body length, 13.53–20.79 mm; width, 2.72–4.27 mm.
Colour (Fig. 44). Body reddish brown with head, thorax and basal 1/5–1/3 of elytra often darker. Head
and thorax with dense white pubescence; pronotal disc with a glabrous median longitudinal line and a glabous
spot on each side of the median line. Elytra covered with whitish pubescence; each elytron with 4 longitudinal
stripes of very dense white pubescence on disc excluding 1 at margin and 1 at suture. Legs covered with fairy
dense pubescence; mid and hind femora with very dense, brush-like brownish hairs on ventral side. Abdomen
with fairly dense yellowish pubescence; each of visible sternites 1–3 with a medial tuft of very dense brown-
ish hairs.
Head. Postclypeus semicircular and flattened, with dense large coarse punctures; distance between lower
lobes of eyes 1.62–1.82 × as long as distance between upper lobes of eyes; distance between upper lobes of
eyes 1.0–1.15 × as long as distance between eyes on ventral side; genal length 0.23–0.25 × as long as head
width immediately below eyes. Antennae slightly longer than body; segments 4–10 flattened and produced on
one side at apex.
Thorax and abdomen. Pronotum 1.14–1.43 × as long as width, with a small conical process at each side;
posterior margin 1.08–1.30 × as wide as anterior margin; pronotal disc binodulose in middle area; disc and
side strongly rugose transversely. Scutellum semicircular, with dense pubescence. Elytra 3.45–4.15 × as long
as prothorax and 3.29–3.57 x as long as shoulder width; apex rounded or truncate, with an acute spine at
suture. Apex of terminal sternite emarginate.
Male terminalia. Apex of ventral median lobe narrowly rounded and apex of dorsal lobe rounded; ventral
lobe slightly shorter than or as long as dorsal lobe; spined region of internal sac divided into two sections: first
section located near base with a mixture of dense simple spines and fairly dense multi-branched spines; sec-
ond section with sparse short simple spines; a long unspined gap between the two sections (Fig. 82a). Eighth
sternite obliquely truncate at side, shallowly emarginate at apex, with sparse long setae arising terminally;
ventral surface with cloud-like processes (Fig. 82b). Eighth tergite truncate or shallowly emarginate at apex;
dorsal surface with fairly dense short simple spines and multi-branched spines (Fig. 82c). Paramere 1.96–1.98
× as long as wide, cylindrical in shape, apex rounded (Fig. 82d).
Female. Body length,12.81–19.72 mm; width, 2.38–4.04 mm.
Antennae distinctly shorter than body; femora without brownish brush-like hairs and visible sternites
without medial tufts of very dense brownish hairs.

Zootaxa 1569 © 2007 Magnolia Press · 53
REVISION OF URACANTHUS
Ovipositor and spermatheca. Ovipositor relatively long; styli arising terminally with short hairs (Fig.
82e). Spermatheca slightly curved; spermathecal gland arising at base (Fig. 82f).
Distribution
Southwestern Queenland, central Northern Territory, northern and sourthern South Australia, and central
and southwestern Western Australia (Fig. 106).
Biology
Hosts are unknown. Adults were collected by light in November and January to March.
Comments
This species resembles U. albatus but differs in having the elytral apex unispinose and each elytron with 4
longitudinal stripes of dense pubescence on the disc.
Material examined
U. ventralis. Holotype. . WA: Mullewa, J. F. May, bearing a name label on which ‘TYPE’ was written in
red; 1 segment of right antenna missing (SAM).
U. multilineatus. Holotype. . WA : Lake Violet, Coll. no. 27–1497, Reg. no. 33556, antennae damaged
(WAM).
Other material examined. 14 , 5 . QLD: 1 , 24 ml. NE of Kihee H.S (27°23'S, 142°37'E), QLD,
14.i.1965, at MV light, L. J. Chinnick (ANIC). NT: 1 , Stuart H’way, 26 km. S of Tennant Creek (19°33'S,
134°14'E), 29.xi.1972, D. H. Colless (ANIC); 1 , 30 km NW by W of Alice Springs (23°32'S, 133°38'E),
7.x.1978, Upton & Barrett (ANIC). SA: 1 , Musgrave Ranges, 12.ii.1966, at light, P. Aitken & N. B. Tindale
(SAM); 1 , Mulgathing HS (30°14'S, 133°59'E), 8.xi.1977, on brushes forming a hedge around the SH, F. &
J. Austin (SAM). WA: 1 , Sandstone, det by Lea, H. J. Carter Collection (NMV); 1 , same as above but
(WAM); 2 , 37 km SW of Youanmi (28°45'S, 118°31'E), 13–14.iii.1982, at light, Coll. no. 437-8, T. F. Hous-
ton & B. Hanich (WAM); 1 , 12 km NE of Warriedar (29°04'S, 117°18'E), 18.iii.1982, at light, Coll. no. 442-
1, T. F. Houston & B. Hanich (WAM); 1 , Wiluna (26°35'S, 120°13'E), ii.1961, A. Douglas leg., G. F. Mus
(WAM); 1 , Ellavalla stn., Carnarvon, ii.1955, A. Snell (NMV); 1 , 3 , Marloo Stn.(28°22'S, 122°37'E),
Wurarga, 1931–1941, A. Goerling (ANIC); 2 , Deeba Rockhole (28°19'S, 116°10'E), 34 km NE by N of
Laverton, 12.xi.1977, T. A. Weir (ANIC); 1 , Binnu (28°02'S, 114°40'E), 8.ii.1965, M. de. Graaf (WAM).
Uracanthus strigosus Pascoe, 1875
(Figs 45, 83, 107)
Uracanthus strigosus Pascoe, 1875: 62.—French, 1911: 67, pl. cxi, Fig.6; Aurivilius, 1912:
147; Lea, 1916: 384; McKeown, 1947: 66; Duffy, 1963:117; Monné, 2005: 660.
Description
Male. Body length, 15.26–22.93 mm; width, 2.98–4.5 mm.
Colour (Fig. 45). Body reddish brown with head, prothorax and basal 1/3 of elytra often darker. Head with
fairly dense yellowish pubescence. Pronotal disc with 4 longitudinal stripes of yellowish pubescence: 2 wider
and denser ones on sides and 2 narrower and sparser ones between the former; each side of pronotum with a
longitudinal stripe of dense pubescence near ventral side; ventral surface of prothorax with fairly dense white
pubescence. Each elytron with 4 longitudinal stripes of dense yellowish white pubescence; the two closer to
suture appearing to merge into one from basal 1/3 or middle to apex. Mid and hind femora with very dense,
brush-like brownish hairs on ventral side. Abdomen with fairly dense yellowish pubescence; each of visible

THONGPHAK & WANG
54 · Zootaxa 1569 © 2007 Magnolia Press
sternites 1–3 with a medial tuft of very dense brownish hairs. Body colour may vary from reddish brown to
blackish. Pubescence colour may vary from whitish to greyish. Specimens from Queensland are darker than
those from New South Wales.
Head. Postclypeus semicircular to subtriangular and flattened, with fine punctures; frontoclypeal suture
deep and narrow in middle; distance between lower lobes of eyes 2.06–2.21 × as long as distance between
upper lobes of eyes; distance between upper lobes of eyes 0.77–0.85 × as long as distance between eyes on
ventral side; genal length 0.31–0.41 × as long as head width immediately below eyes. Antennae slightly
shorter than body; segments 4–10 slightly produced on one side at apex.
Thorax and abdomen. Pronotum 1.12–1.30 × as long as width, with a strongly raised conical process at
each side; posterior margin 1.17–1.44 × as wide as anterior margin; pronotal disc weakly binodulose in middle
area; disc and side strongly rugose transversely. Scutellum semicircular, with dense pubescence. Elytra 3.87–
4.61 × as long as prothorax and 3.64–4.05 × as long as shoulder width, with coarse punctures in basal 1/3,
gradually diminishing toward apex; apex bispinose with marginal spine much longer. The apical spines of
elytra may vary from long to short. Apex of terminal sternite slightly emarginate.
Male terminalia. Apex of ventral median lobe emarginate and apex of dorsal lobe narrowly rounded or
slightly pointed; ventral lobe longer than dorsal lobe; spined region of internal sac divided into two sections:
first section longer than unspined gap between first and second sections, with fairly dense short simple spines;
second section about as long as first section, with sparse short simple spines (Fig. 83a). Eighth sternite
obliquely truncate or more or less rounded at side, clearly emarginate at apex, with sparse long setae arising
terminally; ventral surface with cloud-like processes (Fig. 83b). Eighth tergite shallowly emarginate at apex;
dorsal surface with fairly dense short simple spines and multi-branched spines in basal area (Fig. 83c).
Paramere 1.96–1.98 × as long as wide, cylindrical in shape, apex rounded with long and short setae (Fig. 83d).
Female. Body length, 19.82–22.19 mm; width, 3.56–4.63 mm.
Antennae distinctly shorter than body; femora without brownish brush-like hairs and visible sternites
without medial tufts of very dense brownish hairs.
Ovipositor and spermatheca. Ovipositor relatively long; styli arising terminally with short and long hairs
(Fig. 83e). Spermatheca curved, particularly near apex; spermathecal gland short and arising at basal 1/3 (Fig.
83f)
Distribution
Northeastern and southeastern Queensland, eastern New South Wales, southern and central Victoria,
northern South Australia, and southwestern Western Australia (Fig. 107). This species was accidentally intro-
duced to Ecuador (Monné 2005).
Biology
Known hosts are Acacia spp, Helichrysum ferrugineum, Melaleuca uncinata and Casuarina sp. Adults
were collected from October to April, and attracted to light.
Comments
This species resembles U. albatus but differs in having the pronotal disc with 4 longitudinal stripes of
pubescence on the disc; pubescent stripes close to the elytral suture tending to merge from middle towards the
apex.
Material examined
Holotype. . NSW: Rope’s Creek, Pascoe Coll. 93-60, bearing circular type label, and Pascoe’s handwrit-
ing of the species names (BMNH).
Other material examined. 59 , 41 . QLD: 1 , Toowoomba, 1.x.1928, W. B. Branard (QDPI); 1 , 5 ,

Zootaxa 1569 © 2007 Magnolia Press · 55
REVISION OF URACANTHUS
Yuleba SF Site 1 (26°58'S, 149°44'E), 270 m, Cypress/ Box, 8–10.iii.2002, MV light, Monteith & Wright
(QM); 1 , 330 m, Barakula (26°26'S, 150°31'E), 17–18.xii.2001, MV light, Monteith & Wright (QM); 1 ,
Lake Broadwater (27°21'S, 151°07'E) via Dalby, Gravel Ridge, 12.ii.1985, to light, M. Benie (QM); 1 , same
locality but 23.ii.1986, MV light, G. B. Monteith & G. Thompson (QM); 1 , Mt. Moffat National Park
(25°01'S, 147°57'E), Top Moffat Camp, 13–15.xii.1987, Monteith, Thompson & Yeates (QM); 1 , 1 , 520
m, Wonga Hills site 3 (26°04'S, 150°49'E), 11.xii.2001, MV light, vine scrub, Monteith, Cook & Wright
(QM); 1 , Blackdown Tablelands and Staney Ck. Via Dinge, 17–19.xii.1985, at light, S. Hamlet (QM); 1 ,
24 km N. Tinaroo Dam (17°09'S, 145°33'E), 10–11.xii.1999, at MV light, A. E. Wart (QM); 1 , Bulimba
(16°52'S, 143°29'E), 1.vii.1923 (UQIC); 1 , Miles (26°39'S, 150°11'E), 12.i.1939, N. Geary (AM); 1 , Con-
damine (26°55'S, 150°08'E), 6.ii.1939, N. Geary (AM); 1 , Roma (26°34'S, 148°47'E), 28.iii.1957, E. F.
Riek (ANIC); 1 , Tambourine (27°52'S, 153°07'E) (UQIC). NSW: 1 , Rope’s Creek (QM); 1 , same local-
ity but 29.ix.1913, in wattle nut, W. du B. (SAM); 1 , same locality but (MAM); 1 , Sydney, H. W. Cox
(WADA); 2 , same as above but (NMV); 1 , Grose Vale, 27.ix.1985, J. A. Humphreys (ASCU); 1 , Man-
grove Mt., 6.xi.1960, Mosse and Robinson (ASCU); 1 , Clarence, Blue Mt, 12.ii.1979, N.W. Rodd (AM); 1
, Bombala (AM); 1 , Mogriguy (32°03'S, 148°39'E), 9.ii.1981, R. H. Mulder Collection (AM); 1 , Mt.
Kaputar (30°16'S, 150°10'E), Bullawa Creek, 28.xi.1984, George Hangay (MAM); 1 , NSW (WADA); 1 ,
20 miles SW of Singleton, NSW, 12.xi.1968, no abdomen, I. F. B. Common (ANIC); 1 , Yarras, 27.xi.1964
(ANIC); 1 , Colo Vale, 21.v.1900 (AM). VIC: 1 , Bright (36°43'S, 146°57'E), Coll. no. I. 5485 (SAM); 3 ,
1 , Melton (37°40'S, 144°34'E), 28.iii.1920, on Casuarina, F. E. Wilson, F. E. Wilson Collection (NMV); 1
, 1 , same locality but (VAIC); 1 , same locality but (AM); 1 , Kiata (36°21'S, 141°47'E), i.1928, bred
from Melaleuca uncinata, F. E. Wilson Collection (NMV); 1 , Walpeup (35°08'S, 142°20'E), 25.x.1984,
light trap (VAIC); 1 , Birchip (35°58'S, 142°54'E), x.1902, det. Lea (NMV); 1 , Victoria (AM); 1 ,
7.vi.1937, Coll. no. I 5485 (SAM); 1 , Victoria, 15.v.1916, D. Bcatly (NMV); 1 , You Yangs (37°56'S,
144°26'E), C. Oke (NMV); 1 , Grampians (37°08'S, 142°21'E), 1400ft., Wannon Dividing, Vic., 8.ii.1956,
no abdomen, I. F. B. Common (ANIC); 1 , 17 NW Orbost (37°42'S, 148°27'E),Vic, 8.xii.1955, no abdomen,
E. F. Riek (ANIC); 1 , Wyperfeld (35°33'S, 142°07'E), Vic, i.1974, GWA (ANIC); 1 , 4 , N Mallee,
Dixon, 1917 (NMV); 1 , 5 miles W of Jeparit (36°08'S, 141°59'E), VIC, 4.ii.1956, I. F. B. Common (ANIC);
1 , Big Desert, Broken Bucket Crown Land Res. 7.iii.1975, UV light, P. B. McQuillan (SAM); 1 , Vic,
4.vi.1908 (ANIC). SA: 3 , Glenmanyie Bore (30°50'S, 140°34'E), E of Lake Frome, 28.ix.1975, at light,
sand ridges with black oak Acacia, P. Aitken & T. F. Houston (SAM); 2 , 15 km W Sturt Vale Stn. (35°29'S,
138°59'E), 17.xi.1975, at light, G. F. Gross (SAM); 1 , 185 km S Radium Hill, 3.x.1926, on powerline track,
at light, P. Aitken (SAM); 1 , 10 km NW Emu Junc. (28°38'S, 132°12'E), Great Victoria Desert., 10.x.1976,
at light, J. A. Herridge & G. F. Gross (SAM); 1 , ‘Kurlge’ Blackwood, 860ft, 1.x.1956, at MV light, N. B.
Tindale (SAM); 1 , nr Morgan, xi.1954, F. J. Mitchell (SAM); 1 , Wirrabara, xi.(no year) (SAM); 1 , no
data, SA, no abdomen (SAM). WA: 1 , 4 km W of Hyden, 29.i.1993, E. D. Edward & E. S. Nielsen (ANIC);
1 , no data, det by Lea, no abdomen (ANIC); 1 , Wialki, 6.x.1983, R. P. McMillan (WAM); 5 , 7.5 km SE
of Banjiwarn HS (27°42'S, 121°37'E), 24.iii. 1979, T. F. Houston et. al. (WAM); 1 , Binnu, 2.ii.1965, M. de.
Graaf (WAM); 1 , Condingup, E of Esperance, 27.i.1977, A. M. & M. J. Douglas (WAM); 1 , 10 km W by
SW of Point Malcolm (33°48'S, 123°48'E), 15–18.i.1982, B. Hanich & T. F. Houston (WAM); 1 , Fitzgerald
River Nation Park (33°47'S, 119°45'E), 5.i.1986, MV lamp, no abdomen, G. & A. Daniels (UQIC). Locality
unknown: 3 , Coll. no. 3455, Relton Bequest (QM); 2 , ex Casuarina, 1.ii.1911 (NMV); 1 , same as
above but ix.1911 (AM); 2 , same as above but xi.1912 (NMV); 1 , same as above but (NMV); 1 , same as
above but (NMV); 1 , same as above but xii.1908 (NMV); 1 , same as above but xi.1908 (NMV); 1 , same
as above but i.1907 (NMV); 1 , same as above but ii.1914 (NMV); 1 , same as above but iii.1918 (NMV);
1 , no data (NMV).

THONGPHAK & WANG
56 · Zootaxa 1569 © 2007 Magnolia Press
Uracanthus parallelus Lea, 1916
(Figs 46, 84, 108)
Uracanthus parallelus Lea, 1916: 377.—McKeown, 1947: 65.
Description
Male. Body length, 14.22–17.94 mm; width, 2.08–3.28 mm.
Colour (Fig. 46). Body reddish brown with head thorax, femora and basal elytra often darker. Head, tho-
rax, abdomen and elytra with dense, long, pale yellow pubescence; pronotum with a large more or less gla-
brous spot in the middle; elytral shoulder with a small longitudinal glabrous mark.
Head. Postclypeus subtriangular, slightly convex, with fairly dense coarse punctures; distance between
lower lobes of eyes 1.66–1.80 × as long as distance between upper lobes of eyes; distance between upper
lobes of eyes 1.00–1.09 ×as long as distance between eyes on ventral side; genal length 0.40–0.48 × as long as
head width immediately below eyes. Antennae slightly shorter than body; segments 5–10 subcylindrical and
slightly produced on one side at apex.
Thorax and abdomen. Pronotum 1.18–1.20 × as long as width, rounded or sligthyl produced at side; pos-
terior margin 1.08–1.25 × as wide as anterior margin; pronotal disc without obvious nodules in middle area;
disc and side smooth with transverse rugae near anterior and posterior margins. Scutellum semicircular, with
dense pubescence. Elytra 4.74–4.83 ×as long as prothorax and 4.50–5.04 × as long as shoulder width; longitu-
dinal carinae on disc not obvious; basal 1/3 of elytra with coarse punctures and the remaining 2/3 with fine
punctures gradually diminishing toward apex; apex emarginate and bispinose but spines small. Apex of termi-
nal sternite emarginate.
Male terminalia. Apex of ventral median lobe pointed and apex of dorsal lobe truncate; ventral lobe
shorter than dorsal lobe; spined region of internal sac divided into two sections: first section longer than sec-
ond section, with dense long simple spines in basal half and fairly dense muti-branched spines on sides in api-
cal half; second section about as long as unspined gap between two sections, with sparse simple spines (Fig.
84a). Eighth sternite rounded or slightly obliquely truncate at sides, shallowly emarginate at apex, with long
and fairly long setae arising terminally; ventral surface with fairly dense cloud-like processes (Fig. 84b).
Eighth tergite more or less rounded at apex with a small pointed process in middle of apex; dorsal surface with
very sparse multi-branched spines near base and sparse short simple spines in middle area (Fig. 84c).
Paramere 1.38–1.4 × as long as wide, subcylindrical in shape, apex rounded with sparse long and short setae
(Fig. 84d).
Female. Unknown.
Distribution
Eastern Queensland (Fig. 108).
Biology
Hosts are unknown. One specimen was found on vine scrub. Adults were collected from September to
December.
Comments
This species resembles U. froggatti but differs in having the pubescence on the elytral disc more or less
evenly distributed, and the elytral apex bispinose.
Material examined
Holotype. . QLD: Cooktown, Coll. no. T-8879, bearing a red holotype label; left antenna missing
(NMV).

Zootaxa 1569 © 2007 Magnolia Press · 57
REVISION OF URACANTHUS
Other material examined. 2 . QLD: 1 , Carnarvon Range, 14.xii.1938, N. Geary (AM); 1 , Boomer
Range (23°12'S, 149°44'E), 240 m Pyton Scrub site 5, 28–29.ix.1999, Vine scrub, Monteith, Cook, Burwell &
Evan (QM); 1 ?, N Qld, no abdomen (SAM).
Uracanthus froggatti Blackburn, 1894
(Figs 47, 85, 108)
Uracanthus froggatti Blackburn, 1894: 106.—Froggatt, 1894: 116; Aurivillius, 1912: 147; Lea, 1916: 385; McKeown,
1947: 64; Duffy, 1963: 116.
Description
Male. Body length, 15.48–18.99 mm; width, 2.23–3.26 mm.
Colour (Fig. 47). Body dark reddish brown with head, thorax and basal elytra darker. Head with fairly
dense white pubescence; thorax and basal elytra with dense white pubescence; pronotal disc with 3 vague and
not well-defined, longitudinal, glabrous lines. Elytra with fairly dense white pubescence of uneven distribu-
tion, giving an appearance of irregular pubescent patterns; elytra apex with very dense white pubescences.
Head. Postclypeus triangular, convex, with shiny and smooth surface; frontoclypeal suture deep and wide
in middle; distance between lower lobes of eyes 1.63–1.71 × as long as distance between upper lobes of eyes;
distance between upper lobes of eyes 1.0–1.2 × as long as distance between eyes on ventral side; genal length
0.56–0.57 × as long as head width immediately below eyes. Antennae slightly shorter than body; segments 4–
10 thin and subcylindrical, and slightly produced on one side at apex.
Thorax and abdomen. Pronotum 1.26–1.34 × as long as width, rounded at side; posterior margin 1.15–
1.30 × as wide as anterior margin; pronotal disc weakly binodulose in middle area; disc and side rugose trans-
versely. Rugae on pronotal disc may vary from very distinct to vague. Scutellum semicircular, with dense
pubescence. Elytra 4.60–4.97 ×as long as prothorax and 4.20–4.77 × as long as shoulder width; longitudinal
carinae on disc not obvious; disc with fine punctures; apex more or less truncate. Apex of terminal sternite
rounded with a small notch.
Male terminalia. Apex of ventral and dorsal median lobes narrowly rounded or slightly pointed; ventral
lobe shorter than dorsal lobe; basal unspined region of internal sac with wave-like patterns throughout, about
as long as spined region; spined region with fairly dense long simple spines and sparse multi-branched (Fig.
85a). Eighth sternite obliquely truncate at side, widely emarginate at apex, with long and fairly long setae aris-
ing terminally, ventral surface with sparse cloud-like processes (Fig. 85b). Eighth tergite semicircular in shape
and rounded at apex, with fairly dense basally-forked spines on dorsal surface (Fig. 85c). Paramere short, as
long as wide, cylindrical in shape, apex rounded with long and short setae (Fig. 85d).
Female. Body length, 14.46–15.82 mm; width, 2.17–2.40 mm
Body more robust; antennae and legs shorter. Elytra 5.30–5.27 × as long as prothorax and 4.64–5.15 × as
long as shoulder width.
Ovipositor and spermatheca. Ovipositor very short; styli arising terminally with short and long hairs (Fig.
85e). Spermatheca curved; spermathecal gland long and arising near base (Fig. 85f).
Distribution
Southeastern Queensland and eastern New South Wales (Fig. 108).
Biology
The known host is Lasiopetalum ferrugineum. Adults emerge in April. Larvae feed in the stems of the
host, completely hollowing them for a considerable length, and usually cutting the branch off before pupating,
and forming a cell at the end of the gallery. Adults were collected in November by light trap.

THONGPHAK & WANG
58 · Zootaxa 1569 © 2007 Magnolia Press
Comments
This species resembles U. parallelus but differs in having pubescence on the elytral disc unevenly distrib-
uted and the elytral apex truncate.
Material examined
Paratype. 2 . NSW: 1 , Coll. no. I. 5489, no abdomen (SAM); 1 , Coll. no. 5170, bearing a blue
paratype label; left antenna missing (NMV). The holotype is supposed to be in BMNH according to McKe-
own (1947) but could not be located.
Other material examined. 3 , 2 . QLD: 1 , Toowoomba, R. Illidge (UQIC). NSW: 1 , Coogee,
8.x.1925, F. A. McNell (AM); 1 , Sydney (ASCU); 1 , no locality, 20.iv.1893, bred from Lasiopetalum fer-
rugineum, Froggatt (ANIC); 1 , Thomas River, 23 km NW by W of Mt. Arid, 4–7.xi.1977, at light, J. F.
Lawrence (ANIC).
Uracanthus tropicus Lea, 1916
(Figs 48, 86, 108)
Uracanthus tropicus Lea, 1916: 376.—McKeown, 1947: 67.
Description
Male. Body length, 13.99–15.58 mm; width, 2.21–2.58 mm.
Colour (Fig. 48). Head, thorax, antennae, and basal 1/3 elytra reddish brown to blackish brown, and
remaining elytra yellowish brown. Body covered with fairly dense pale yellow pubescence. Each elytron usu-
ally with a wide submarginal longitudinal glabrous mark starting from shoulder extending to basal 1/3–apical
1/3 of elytra. The submarginal longitudinal glabrous mark on the elytra varies from very clear to almost com-
pletely absent.
Head. Postclypeus triangular, convex, smooth, shiny and glabrous; frontoclypeal suture deep and narrow
in middle; distance between lower lobes of eyes 1.58–1.76 × as long as distance between upper lobes of eyes;
distance between upper lobes of eyes 1.44–1.50 × as long as distance between eyes on ventral side; genal
length 0.26–0.34 × as long as head width immediately below eyes. Antennae as long as or slightly longer than
body; segments 4–10 thin, subcylindrical in shape, and slightly produced on one side at apex.
Thorax and abdomen. Pronotum 1.25–1.48 × as long as width, rounded at sides; posterior margin 1.10–
1.21 × as wide as anterior margin; pronotal disc without obvious nodulose area in middle; disc and side
smooth with feeble transverse rugae near posterior margin. Scutellum semicircular, with dense pubescence.
Elytra 3.70–3.89 × as long as prothorax and 4.19–4.53 × as long as shoulder width; basal 1/3 of elytra with
fine puncture, diminishing toward apex; apex truncate or slightly marginate with a small tooth at suture and
margin, respectively. Apex of terminal sternite rounded or widely truncate.
Male terminalia. Apex of ventral median lobe and apex of dorsal lobe slightly pointed or widely rounded;
ventral lobe shorter than dorsal lobe; spined region of internal sac about 3 × as long as terminal unspined area,
with dense long simple spines (Fig. 86a). Eighth sternite rounded or obliquely truncate at sides, shallowly
emarginate at apex, with some setae arising terminally; ventral surface with simple spines (Fig. 86b). Eighth
tergite rounded or very weakly emarginate at apex; dorsal surface with fairly dense basally-forked spines on
basal-mid area (Fig. 86c). Paramere 2.8–2.9 × as long as wide, cylindrical in shape, apex rounded with long
and short setae (Fig. 86d).
Female. Body length, 11.11–17.50 mm; width, 2.0–2.90 mm
Body more robust; antennae shorter than body; elytra 4.23–4.53 × as long as prothorax and 4.06–4.39 ×as
long as shoulder width.

Zootaxa 1569 © 2007 Magnolia Press · 59
REVISION OF URACANTHUS
Ovipositor and spermatheca. Ovipositor relatively long; styli arising terminally with short hairs (Fig.
86e). Spermatheca heavily curved, particularly near base; spermathecal gland arising at base (Fig. 86f).
Distribution
Northern and eastern Queensland (Fig. 108).
Biology
Hosts are unknown. Adults were collected during October to December.
Comments
This species resembles U. parallelus but differs in having each elytron with a wide submarginal longitudi-
nal glabrous mark starting from the shoulder and extending to 1/3–2/3 of elytral length; the elytral apex more
or less truncate. The label of the holotype states that the specimen is a male but it is in fact a female.
Material examined
Holotype. 1 . Northern Queensland, Coll no. I. 5697, bearing a name label on which ‘TYPE’ was written
in red (SAM).
Other material examined. 4 , 7 . QLD: 2 , 2 , 15 km W of Irvinebank (17°25'S, 145°04'E), 27–
28.xi.1981, J. Balderson (ANIC); 1 , 24 km E by N of Ravenshoe (17°35'S, 145°43'E), 14.xi.1981, J. Balder-
son (ANIC); 1 , Mossman Gorge National Park (16°28'S, 45°20'E), nr Mossman, 23.xi.1981, J. Balderson
(ANIC); 1 , SW of Mount Garnet (18°05'S, 144°40'E), 4.x.1989, Forty Mile Scrub, L. Ring (UQIC); 1 ,
Brisbane, 14.x.1956, Rirhpateacts (UQIC); 1 , Mt. Abbott (20°06'S, 147°45'E), summit area, 8–10.xii.1996,
750–1000 m, G. Monteith, D. & I. Cook (QM); 1 , Flinder River, 59 km SW of Normanton, QLD, 1.xii.1981,
J. Balderson (ANIC); 1 , 6 km SE of Chillagoe, QLD, 26.xi.1981, J. Balderson (ANIC).
Uracanthus parvus Lea, 1916
(Figs 49, 87, 109)
Uracanthus parvus Lea, 1916: 375.–McKeown, 1947: 65.
Uracanthus dentiapicalis McKeown, 1948:54. syn. nov.
Description
Male. Body length, 11.9–20.4 mm; width, 1.71–2.93 mm.
Colour (Fig. 49). Head, thorax, antennae, and basal elytra blackish brown; the remaining part reddish
brown. Body covered with sparse to fairly dense white or yellowish pubescence; elytra with denser and longer
whitish pubescence on base along suture and from basal 1/10 to 1/3 near margin. Antennal segments 4 to 10
with dense fringe of yellowish pubescence on one side. Body colour varies from blackish brown to reddish
brown.
Head. Postclypeus semicircular, convex, smooth and shiny; frontoclypeal suture deep and narrow in mid-
dle; distance between lower lobes of eyes 1.37–1.75 × as long as distance between upper lobes of eyes; dis-
tance between upper lobes of eyes 1.5–1.67 × as long as distance between eyes on ventral side; genal length
0.35–0.38 × as long as head width immediately below eyes. Antennae slightly longer than body; segments 4–
10 slightly flattened and strongly produced on one side at apex.
Thorax and abdomen. Pronotum 1.25–1.41 × as long as width, rounded at side; posterior margin 1.23–
1.44 × as wide as anterior margin; pronotal disc very weakly binodulose in middle area; disc and side strongly
rugose transversely. Scutellum semicircular, with sparse pubescence. Elytra 4.94–5.30 × as long as prothorax
and 4.2–4.65 × as long as shoulder width; basal 1/3 of elytra coarsely punctate and then 2/3 of elytra finely

THONGPHAK & WANG
60 · Zootaxa 1569 © 2007 Magnolia Press
punctate, gradually diminishing toward to apex; apex bispinose. The apical spines of the elytra may vary from
highly developed to small processes. Apex of terminal sternite truncate or slightly emarginate.
Male terminalia. Apex of ventral median lobe rounded and apex of dorsal lobe narrowly rounded or
slightly pointed; dorsal lobe shorter than ventral lobe. Spined region of internal sac divided into two sections:
first section longer than unspined gap and second section combined, with fairly dense multi-branched spines
on basal 2/3 and some scale-like spines on apical 1/3; second section with sparse short simple spines (Fig.
87a). Eighth sternite rounded or slightly obliquely truncate at side, almost truncate at apex, with fairly long
setae arising terminally; ventral surface with cloud-like processes (Fig. 87b). Eighth tergite truncate and shal-
lowly emarginate at apex; surface with fairly dense basally-forked spines and simple spines (Fig. 87c).
Paramere robust, 2.0–2.09 × as long as wide, apex obliquely truncate with fairly dense long and short setae
(Fig. 87d).
Female. Body length, 11.21–18.33 mm; width, 1.87–2.80 mm.
Body more robust; pubescence on elytra more or less evenly distributed; antennae shorter than body, without
fringe.
Ovipositor and spermatheca. Ovipositor relatively long; styli long, arising terminally with short and long
hairs (Fig. 87e). Spermatheca curved, particularly near apex; spermathecal gland relatively long, arising at
base (Fig. 87f).
Distribution
Southern, southwestern and northeastern Western Australia, southern and northern Northern Territory,
northern and southern South Australia, and southwestern New South Wales (Fig. 109).
Biology
Hosts are unknown. Adults were collected from October to March, and in May and August by UV light
and pitfall.
Comments
This species closely resembles U. pertenuis but differs in having the body more slender, male elytra with
denser and longer whitish pubescence along the suture and margin, while female elytra have pubescence more
or less evenly distributed, and the elytral apex with two small spines of similar size or length.
Material examined
Uracanthus parvus. Holotype. . WA : no locality, Coll. no. I. 5703, W. du Boulay (SAM).
Uracanthus dentiapicalis. Holotype. . WA : Wandagee Station (23°50'S, 114°30'E) , Coll. no. 41-740
(WAM).
Other material examined: 31 , 6 . NT: 1 , 26 km WSW of Mulga Park (26º00'S, 131º25'E) HS,
18.i.1982, D. C. F. & B. G. F. Rentz, & R. Honeycutt (ANIC); 1 , Rimbija (11º01'S, 136º45'E), Wessel
Island, 21.i–3.ii.1977, Farrow & Dowse (ANIC); 1 , same data as above but no abdomen (ANIC); 1 , Finke
Gorge National Park, 4 km N Finke Ellery Ck, JCL, 26.iii.1993, J. A. Forrest & D. Hirst (SAM); 1 , Yuen-
dumu, Central Australia (QM). NSW: 5 , 11 km N of Mossgiel (33º16'S, 144º34'E), Willandra Bridge, dry
swamp, 21.xii.1970, at light, Britton, Misko & Pullen (ANIC). SA: 1 , Mt. Davies (26º12'S, 129º15'E),
11.xi.1963, at light, P. Aitken & N. B. Tindale (SAM); 1 , Minnie Downs, N.E. Corner of SA, L. Reese
(SAM); 1 , Everard Park (27º00'S, 132º42'E), 30.x.1970, at light, E. Matthews (SAM); 1 , same data as
above but 2.xi.1970 (SAM); 2 , 48.9 km SE Mt. Lindsay (27º18'50″S, 130º15'54″E), Sand plain PIL camp,
22–25.x.1996, pitfall, Pitjantjatjara Lands (SAM); 1 , 6 mile W of Iron Knob (32º43'S 137º49'E), 16.iii.1968,
I. F. B. Common & M. S. Upton (ANIC); 1 , 40 km W Vokes Hill Junct, (28º32'S, 130º16'E), 14–15.iv.1994,
Malaise trap, J. A. Forrest (SAM); 1 , Margaret Rv. 9.5 km SE Coward Spring. 2.xii.1974, at light, J. A. Her-

Zootaxa 1569 © 2007 Magnolia Press · 61
REVISION OF URACANTHUS
ridge (SAM); 1 , Frome River crossing of Birdsville Track nr Marree, 25.x.1966, G. F. Gross (ANIC). WA: 2
, Milly Milly (26º04'S, 116º41'E), 27.v.1922 (WAM); 1 , 3.75 km NE of Comet Vale Siding (29º57'S,
121º07'E), 7–15.iii.1979, T. F. Houston et. al. (WAM); 1 , 29 km SE by E of Coolgardie (31º07'S, 121º24'E),
5.v.1983, E. S. Nielson & E. D. Edwards (ANIC); 1 , Carson Escarpment (14º49'S, 126º49'E), 9–
15.viii.1975, no abdomen, I. F. B. Common & M. S. Upton (ANIC); 1 , Yanchep NP (31º32'S, 115º41'E),
28.iii.1990, Horak Wooks, Walsh & Yeates (ANIC); 1 , 10 km W by SW of point Malcolm (33º48'S,
123º46'E), 15–18.i.1982, at light, B. Hanich & T. F. Houston (WAM); 1 , 1 , Bernier Island, 18.v.1963, R.
D. Hughes (ANIC); 1 , Drysdale River (15°02'S, 126°55'E), 3–8.viii.1975, I. F. B. Common & M. S. Upton
(ANIC); 1 , Kalgoorlie, iii.1845, Mathews (WADA); 1 , 16km W of Lyons River HS (24º38'S, 115º20'E),
13–15.v.1981, B. Hanich & T. F. Houston (WAM); 1 , 11 km N of Geraldton, at Drummond Cove, 2–
11.v.1973, UV light, N. McFarland (ANIC); 1 , 9 km NW of Mt Ragged, Junana Rock (33º23'S, 123º24'E),,
26.x.1977, J. F. Lawrence (ANIC); 1 , 1.7–2.7 Km NW of Toolinna Rockhole (32º45'S, 124º59'E), 24–
28.ii.1984, on Colluvium swale-stunted grass, T. F. Houston (ANIC); 1 , 1 , Wittenoom (22º14'S, 118º20'E)
(WAM).
Uracanthus pertenuis Lea, 1916
(Figs 50, 88, 110)
Uracanthus pertenuis Lea, 1916: 374.—McKeown, 1947: 66; Duffy, 1963:117.
Description
Male. Body length, 12.62–21.6 mm; width, 1.8–3.45 mm.
Colour (Fig. 50). Body reddish brown with head, thorax and basal elytra darker. Frons with sparse white
pubescence, clypeus glabrous or with very sparse white pubescence. Antennal segments 3 to 10 with yellow-
ish fringe. Pronotal disc with 2 broad longitudinal stripes of fairly dense white pubescence; prothorax with a
longitudinal stripe of very dense white pubescence on each side near ventral side. Each elytron with 2 longitu-
dinal stripes of dense white pubescence: 1 along suture starting from base, widest at base, and becoming nar-
rower toward apex, and 1 along margin, starting from or behind shoulder, widest at base tapering toward apex;
the rest of elytra with sparse white pubescence. Body colour varies from blackish brown to reddish brown.
Head. Postclypeus subtriangular, convex, smooth and shiny; frontoclypeal suture deep and wide in mid-
dle; distance between lower lobes of eyes 1.67–1.9 × as long as distance between upper lobes of eyes; distance
between upper lobes of eyes 1.23–1.31 × as long as distance between eyes on ventral side; genal length 0.35–
0.39 × as long as head width immediately below eyes. Antennae as long as or slightly longer than body; seg-
ments 5–10 produced on one side at apex.
Thorax and abdomen. Pronotum 1.25–1.47 × as long as width, rounded at side; posterior margin 1.15–
1.35 × as wide as anterior margin; pronotal disc weakly binodulose in middle area; disc and side strongly rug-
ose transversely. Scutellum semicircular, with sparse pubescence. Elytra 4.25–4.78 × as long as prothorax and
4.50–4.87 × as long as shoulder width; elytra finely punctate; elytral apex strongly bispinose with spine at
margin distinctly longer than that at suture. The sutural spine at the elytral apex varies from a small to long but
never longer than marginal spine. Apex of terminal sternite truncate or slightly emarginate.
Male terminalia. Apices of both ventral and dorsal median lobes rounded; dorsal lobe slightly longer than
ventral lobe; spined region of internal sac divided into 2 sections: first section about as long as unspined gap
between first and second sections, with fairly dense multi-branched spines and scale like spines; second sec-
tion shorter than first section, with sparse short simple spines (Fig. 88a). Eighth sternite obliquely truncate at
sides, shallowly emarginate at apex, with fairly long setae arising terminally; ventral surface with cloud-like
processes (Fig. 88b). Eighth tergite shallowly emarginate at apex; surface with fairly dense basally-forked

THONGPHAK & WANG
62 · Zootaxa 1569 © 2007 Magnolia Press
spines and simple spines (Fig. 88c). Paramere robust, 2.0–2.09 × as long as wide; apex obliquely truncate with
fairly dense long and short setae (Fig. 88d).
Female. Body length, 15.57–22.69 mm; width, 2.34–3.50 mm.
Body more robust and broader; pubescent patterns on elytra similar to those in males; antennae shorter
than body, without the fringe;
Ovipositor and spermatheca. Ovipositor relatively long; styli arising terminally with short and long hairs
(Fig. 88e). Spermatheca heavily curved; spermathecal gland short, arising near base (Fig. 88f).
Distribution
Northeastern, southern and southeastern Queensland, northern, northeastern and southeastern New South
Wales, eastern and southern Victoria, southern Tasmania, southeastern South Australia, and southwestern and
southeastern Western Australia (Fig. 110).
Biology
Known hosts are Acacia armata, and Loranthus sp. and A. oxycedrus. Adults were collected by light trap
between September and April.
Comments
This species closely resembles U. parvus but differs in having the body more robust and longer; elytra of
both sexes with denser and longer whitish pubescence along the suture and margin, and the elytral apex with
the marginal spine always distinctly longer than the sutural spine.
Material examined
Holotype. . VIC: no locality, Coll. no. I. 5488; bearing a name label on which ‘TYPE’ was written in
red; abdomen damaged (SAM).
Paratypes. 1 , 1 . TAS: 1 , Hobart, Coll. no. I. 4504, A. M. Lea (SAM). SA: 1 , Goolwa, Old collec-
tion (SAM).
Other material examined: 37 , 23 . QLD: 1 , 1 , Brisbane (27º28'S, 153º01'E), 4.xi.1960, K. Roarao
(UQIC); 1 , same locality as above but 15.iv.1949, W. Wetherall (UQIC); 1 , 16 km N of Boonah (27º60'S,
152º01'E), 27.ix.1986, C. Burwell (UQIC); 1 , Carnarvon National Park, Mount Moffatt section; Ranger
Headquarters (25º00'122″S, 147º56'59″E), 1.xii.1987, MV light, J. Skevington, C. Lambkin & S. Evans
(UQIC); 1 , Cunnamulla (28º04'S, 145º41'E), 22.xi.1938, N. Geary (AM); 1 , Rockhampton (23º22'S,
150º32'E) (MAM); 1 , Watalgan Range (24º40'S, 128º53'E), N of Bundaberg, 7.x.1972, H. Frauca (ANIC).
NSW: 1 , 5 mi S Mendooran (31º47'S, 149º17'E), Goonoo State Forest, 25.iv.1971, D. K. McAlpine (AM); 1
, Killara (33º45'S, 151º09'E), 20.xii.1924, Allowrie (MAM); 2 , Ropes Creek (29º55'S, 146º16'E) (MAM);
1 , Sydney, 24.i.1904 (ASCU); 1 , 15 km NE Ulan, 4.xii.1982, L. O'Donnell (ANIC). ACT: 1 , Canberra
(35º18'S, 149º08'E), at MV light, N. B. Tindale (SAM); 1 , same locality as above but 28.i.1988, B. J. Leps-
chi (ANIC); 1 , Black Mountain (35º18'S 149º08'E), 29–31.xii.1985, K. R. Pullen (ANIC); 1 , same local-
ity as above but 31.xii.1951, L. Obinaick (ANIC); 1 , same locality as above but ii.1986, light trap, Z. Liepa
(ANIC); 1 , same locality as above but 18.i.1956, light trap, P. B. Carne (ANIC); 1 , same locality as above
but 28.i.1988, L. Obinaick (ANIC); 1 , Guabgahlin, 21.xii.1962, S. Baker (SAM). VIC: 3 , 3 , Baxter
(38º11'S, 145º09'E), C. Oke (NMV); 2 , 1 , Langwarrin (38º09'S 145º10'E), J. E. Dixon (NMV); 1 , same
data as above but (AM); 1 , same locality as above but i.1914, C. French Collection (NMV); 1 , 1 , same
locality as above but 4.ii.1922, ex Acacia oxycedrus (NMV); 3 , same locality as above but 5.ii.1922
(NMV); 3 , same locality as above but 12.ii.1922 (NMV); 2 , same data as above but (VAIC); 1 , Mallee
(36º10'S, 146º54'E) (VAIC); 1 , Clayton (37º55'S, 145º07'E) (NMV); 1 , Victoria (NMV). SA: 1 , Ade-
laide (34º55'S, 138º36'E), C. Watts (SAM); 1 , Wirrabara (33º01'S, 138º16'E), xi (no year), S. H. Curnow

Zootaxa 1569 © 2007 Magnolia Press · 63
REVISION OF URACANTHUS
Collection (SAM); 1 , Gordon (32º07'S, 138º16'E), 5.v.1924, Hanson (MAM). TAS : 1 , Tasmania, A.
Simon (SAM). WA: 1 , Beverley (32º06'S, 116º55'E), E. F. du Boulay (SAM); 1 , Marloo Stn, Wurarga,
1931–1941, A. Goerling (ANIC); 1 , Eucla (31º40'S, 128º53'E), 30.iv.1995 (SAM); 1 , Duke of Orleans
(33º55'S, 122º35'E), Bay Caravan Park, Ablution block, 21.i.1995, A. F. Longbotton (WAM). Locality
unknown: 1 , no locality, S. R. E. Brock Collection donate (ANIC); 1 , no data (QM); 1 , G. 355 (QDPI);
1 , no data (QDPI); 1 , 1 , no data (NMV).
Uracanthus bivitta Newman, 1838
(Figs 51, 89, 111)
Uracanthus bivitta Newman, 1838: 172.—Best, 1882: 35; Aurivillius, 1912: 147; 1917: 16; McKeown, 1947: 63; Duffy,
1963: 116.
Uracanthus bivittata.—French, 1911: 67.
Uracanthus bivittatus.—Lea, 1916: 385.
Uracanthus marginellus Hope, 1841a: 54.—Hope, 1841b: 65; 1844: 198; Aurivillius, 1912: 147; Lea, 1916: 384; McKe-
own, 1947: 65. syn. nov.
Uracanthus inermis Lea (not Aurivillius), 1917a: 738. syn. nov.
Uracanthus leai McKeown, 1938: 214 (nom. nov. for U. inermis Lea).–McKeown, 1947: 64.
Description
Male. Body length, 17.17–32.12 mm; width, 2.68–5.59 mm.
Colour (Fig. 51). Body reddish brown with head, thorax and basal elytra darker. Frons and vertex with
mixture of fairly dense white and yellowish pubescence. Pronotum with 2 longitudinal stripes of very dense
yellowish pubescence at each side: 1 on disc and 1 near ventral side; each side with 2 spots of very dense yel-
lowish pubescence between above longitudinal stripes: anterior one larger than posterior one; rest of prono-
tum with fairly sparse to very sparse pubescence. Each elytron with two longitudinal stripes of very dense
yellowish pubescence: 1 wide on disc and 1 narrow at margin; the area between the two stripes glabrous or
sparsely pubescent. Body colour may be darker or paler. Specimens from Western Australia are more hairy,
for example, pronotal disc between two longitudinal stripes can have fairly dense pubescence, and the area
between the two longitudinal stripes of very dense pubescence on the elytron, can be densely pubescent.
Head. Postclypeus triangular, convex, with fairly dense fine punctures; frontoclypeal suture deep and
wide in middle; distance between lower lobes of eyes 2.12–2.37 × as long as distance between upper lobes of
eyes; distance between upper lobes of eyes 1.14–1.23 × as long as distance between eyes on ventral side;
genal length 0.20–0.38 × as long as head width immediately below eyes. Antennae slightly shorter than body;
segments 4–10 thin and slightly produced on one side at apex.
Thorax and abdomen. Pronotum 1.17–1.44 × as long as width, rounded at side; posterior margin 1.02–
1.33 × as wide as anterior margin; pronotal disc weakly binodulose in middle area; disc and side strongly rug-
ose transversely. The transverse rugae on the pronotal disc may vary from very strong to weak. Scutellum
semicircular, with sparse to faily dense pubescence. Elytra 4.20–4.89 × as long as prothorax and 4.03–4.51 ×
as long as shoulder width; apex emarginate or weakly bispinose. The elytral apex varies from widely emargin-
ate to narrowly emarginate or even more or less truncate; apical spines may be well developed or reduced to
the minimum. Apex of terminal sternite emarginate.
Male terminalia. Apices of both ventral and dorsal median lobes slightly pointed; dorsal lobe about as
long as or slightly longer than ventral lobe; spined region of internal sac occupying most of internal sac; basal
unspined region very short with 1–2 lines of large spines near base, sometimes forming an arch in shape;
spined region divided into two sections; first section much longer than second section, with sparse scale-like
spines, dense long simple spines; second section with sparse simple spines and multi-branched spines near the
end of this section; a short unspined gap between sections (Fig. 89a). Eighth sternite obliquely truncate at side,

THONGPHAK & WANG
64 · Zootaxa 1569 © 2007 Magnolia Press
narrowly emarginate at apex, with fairly long setae arising terminally; ventral surface with cloud-like pro-
cesses (Fig. 89b). Eighth tergite rounded to truncate or slightly emarginate at apex, with dense simple spines
or basally-forked spines on ventral surface (Fig. 89c). Paramere 2.53–3.0 × as long as wide, subcylindrical in
shape; apex rounded with fairly dense long and short setae (Fig. 89d).
Female. Body length, 18.14–29.59 mm; width, 2.92–5.23 mm.
Body more robust, antennae distinctly shorter than body.
Ovipositor and spermatheca. Ovipositor relatively long; styli long, arising terminally with short hairs
(Fig. 89e). Spermatheca short, slightly curved; spermathecal gland arising near base (Fig. 89f).
Distribution
Southeastern Queensland, eastern New South Wales, Australian Capital Territory, southern and western
Victoria, southeastern South Australia, eastern Tasmania, and southwestern Western Australia (Fig. 111).
Biology
Known hosts are Ulex sp., Acacia longifolia, Pultenaea stipularis, Helichrysum ferrugineum, Cytisus sco-
parius, Jacksonia sp., J. sternbergiana, Pultanea sp., and Eucalyptus sp. Adults were found feeding on Lep-
tospermum scoparium in November and visiting Eucalyptus and Xanthorrhoea flowers.
Comments
This species is similar to U. discicollis but differs in having the pronotal disc strongly rugose transversely;
the elytral apex widely emarginate to narrowly emarginate or even more or less truncate; each side of the
pronotum with 2 spots of very dense yellowish pubescence.
Material examined
U. bivitta. Holotype. . New Holland (Australia): no locality, Ent. Club 44–12; bearing a red circular
type label; 4 segments of left antenna and 3 segments of right antenna missing (BMNH).
U. marginellus. Holotype. . No locality, Type Coll. no.1769 in Hope Coll.; bearing a type label; right
antenna missing (HMO).
U. inermis. Holotype. . QLD: Cairns, type no 12587; bearing a name label with Lea’s handwriting; 3
segments of left antenna missing (SAM). Allotype. . QLD: same locality as above; bearing an allotype label
(SAM).
Other material examined: 115 , 102 . QLD: 2 , Mt. Lofty Ranges, S. H. Curnow (SAM); 1 , same as
above but (AM); 1 , same as above but (QM); 1 , Bowen, A. Simson (SAM); 1 , 3 km N of Lake Bowar-
rady, Fraser Island, 12.x.1978, G. B. Monteith (QM); 1 , Inglewood Ranges (NMV); 1 , same locality as
above but 4.xi.1925 (UQIC); 1 , Charteville-Bollon Rd, 70 km SE Charteville (26°52'S, 146°35'E),
28.x.1991, MV lamp, G. Daniels (UQIC); 1 , Brisbane, 30.xii.1970, Acacia ridge, E. C. Dahms (QM); 1 ,
same locality as above but 8.xii.1914, H. Hacker (QM); 1 , same locality as above but Illidge (UQIC); 1 , 3
km S of Stanthorpe, 21.xi.1980, MV lamp, M. A. Schneider & G. Daniels (UQIC); 1 , Stanthorpe, 29.x.1926
(NMV); 1 , same locality as above but 19.x.1926 (QDPI); 1 , same locality but E. Sutton (QM); 1 , Colton
Vale, E. F. du Boulay Collection (WAM); 1 , Fletcher, 3.xii.1938, E. Sutton (QM); 2 , 2km NbyNW Jowal-
bina (15°45’S, 144°15’E), 17.i.1994, at light, P. Zborowski & E. D. Edwards (ANIC); 1 , Cooktown (QM); 1
,1 , 560 m Amphitheare Camp, Expedition Range National Park (25º12'S, 148º59'E), 17.xii.1997, MV
light, Monteith, Cook & Thompson (QM); 1 , Blackdown Tableland, Stoney Creek via Dingo, NE QLD, 17–
19. xii.1985, at light, S. Hamlet (SAM); 1 , Glen Aplin, 1.i.1965, P. Kerridge (UQIC). NSW: 1 , 1 , Dar-
key Creek near Mt. Papong, 1.i.1979, on Eucalyptus bloom (ANIC); 1 , Glen Apline, 27.x.1929 (ANIC); 1
, same as above but 27.x.1941, S. R. E. Brock Collection (ANIC); 1 , same as above but 1.i.1965 (NMV); 1
, Shoalhaven, 1895, G. W. F. (ANIC); 2 , Galston, Dumbrell (SAM); 1 , Cumberland Park, 9.i.1958, G.

Zootaxa 1569 © 2007 Magnolia Press · 65
REVISION OF URACANTHUS
Dolazel (AM); 1 , Pt. Denison (MAM); 1 , Nelson Bay, 12.xi.1960, I. F. B. Common & M. S. Upton
(ANIC); 1 , 50 miles S of Singleton, NSW, 5.i.1956, I. F. B. Common (ANIC); 1 , Mt. Kaputar, 3000 ft,
30.xi.1968, at light, C. W. Frazier (ANIC); 1 , same data as above but 18.xi.1968 (ANIC); 1 , same locality
as above but Bullewa Creek, 29.xi.1984, George & Hangay (ANIC); 1 , Lane Cove, 16.x.1943, N. W. Rodd
(AM); 1 , Roseville, 28.i.1961, C. E. Chadwick (ASCU); 1 , Black Hill, Athelstone, 22.x.1966, on scrub
Cryptandra tomentosa, N. McFarland (SAM); 1 , Hornsby, xi.1911, C. Gibbons (AM); 1 , Helensburgh,
vii.1996, H. E. Osburne (ASCU); 1 , Frenchs Forest (33°44'S, 151°12'E), 11.xii.1937, N. C. Lioyel (ASCU);
1 , Springwood, 30.x.1961, A. E. Searle (ASCU); 1 , Ropes Creek (MAM); 1 , 1 , 14 km W of Braid-
wood, emerged 2.xi.1992, ex thick stems of Cytisus scoparius, P. Hodge Coll. (ANIC); 1 , Allowrie,
22.x.1923, Killarn (AM); 2 , 3 , NSW (AM); 2 , 2 , same data as above but (SAM); 1 , same data as
above (NMV); 1 ? (no abdomen) (AM); 1 ? (no abdomen) (WADA); 1 , National Park, x.1920 (NMV); 1 ,
1 , Sydney, 1.xi.1902, W. B. G. Warerley (ASCU); 1 , 1 , Sydney (MAM); 1 , 1 , same locality as
above but (HMO); 2 , same locality as above but (SAM); 1 , same locality as above but Deane (UQIC); 1
, 1 , same locality as above but H. J. Carter (NMV); 1 , same locality as above but xi.1911, C. Gibbons
(AM); 1 , same locality as above but 1924, W. W. Froggatt (ANIC); 1 , Cessnock, NSW (WAM); 1 , New-
ell H’ Way, 9 km N by NE Coonabarabran, 9.xii.1980, at light, E. Britton (ANIC); 1 , 23 km N by NE Coon-
abarabran (30°05'S, 149°24'E), 488 m, 7.xii.1974, at light, I. F. B. Common & E. D. Edwards (ANIC); 1 ,
Merimbula (36°54'S, 149°56'E), 23.ix.1999, E. C. Zimmerman (ANIC); 1 , 3ml NW. of Evans Head,
13.xii.1955, M. J. D. White (ANIC); 1 , Kurina, xi.1973, A. Werl (ANIC); 2 , 1 , Wahroonga, H. J. Carter
(ANIC); 2 , same locality as above but x.1952, R. D. (ANIC) 1 , Thredbo River, Mt. Kosciusko, 3000ft,
15.xii.1931, L. Graham (ANIC); 1 , La Perouse (33°58'S, 151°14'E), 8.xi.1978, E. E. Taylor (FCNI); 1 ,
Narrabeen (33°42'S, 151°17'E), 24.x.1956, H. Booth (FCNI). ACT: 3 , 2 , Black Mountain (35°17'S,
149°07'E), at light, xi.1965–ii.1966 (ANIC); 1 , same locality as above but 16.xii.1952, L. J. Chinnick
(ANIC); 1 , same locality as above but 19.i.1956, light trap, I. F. B. Common (ANIC); 1 , same data as
above but 24.xii.1964 (ANIC) 1 , same data as above but 12.xi.1964 (ANIC); 1 , same data as above but
27.x.1964(ANIC). VIC: 1 , Donvale, x.1952, C. J. R. J (VAIC); 1 , N of Melbourne, no abdomen (NMV);
1 , Melbourne (NMV); 1 , same locality as above but (VAIC); 2 , 1 , Ringwood, C. Oke (NMV); 1 ,
Ferntree Gully, xi.1936, C. Oke (NMV); 1 , Boronia, xii.1955, Fleet (NMV); 1 , Cheltenham, x.1930, J. C.
Goudie (NMV); 1 , N. Mallee, Dixon (NMV); 1 , Somerville, 12.iii.1902 (NMV); 1 , Wellington, Carey
River, 15.ii.1977, A. A. Calder (NMV); 1 , no locality (NMV); 1 , no locality, Coll. no. 3455, Relton
Bequest (QM); 1 , Melbourne, H.W. Davey (VAIC); 1 , no locality, Vic (NMV); 1 , 1 , Little Desert, 13
miles S of Kiata, Vic, 7.xi.1966, I. F. B. Common & M. S. Upton (ANIC); 1 ? (no abdomen), same as above
but 6.xi.1966 (ANIC); 1 , same locality as above but 14.xi.1945, F. E. Wilson (NMV); 1 , same as above
but 23.x.1946 (NMV); 1 , Kiata, 26.x.1961, J. C. Le Sovef (NMV); 1 , same locality as above but x.1939,
R. Oldneld (NMV); 1 , 1 , Lake Hattah, J. E. Dixon (NMV); 1 , 90 miles Desert, N of Yanac, 6.xi.1958, F.
E. Wilson (NMV); 1 , Grampians, xi.1950 (ANIC); 1 , Melboune, 1901 (NMV); 2 , 4 , Victoria (NMV);
1? (no abdomen), same locality as above (NMV). SA: 1 , Murray River, H. S. Cope (SAM); 1 , Port Lin-
coln, Blackburn (SAM); 1 , Cape Donnington Lincoln, 19.i.1982, at light, E.G. Matthews (SAM); 1 ,
Karoonda to Peebinga, G. E. H. Wright (SAM); 1 , Adelaide, R. F. Kemp (AM); 1 , 1 , same locality as
above but Wilson (HMO); 2 , 1 , no locality, SA (NMV); 1 , 1 , no locality ( in AM, in SAM); 1 ,
no locality, A. P. Burgess (AM); 1 , same as above but (SAM); 1 ? (no abdomen), no locality (QM); 1 ,
‘Kurlge’ Blackwood, SA, 850ft, xi.1965, N. B. Tindale (SAM); 1? (no abdomen), Yorktown (SAM). WA: 1
, Geraldton, J. Clark (AM); 1 , nr Howathara Hill, 25 km E by NE of Geraldton, 28.x.1992, E. D. Edwards
& E. S. Nielsen (ANIC); 1 , Kellerberrin, 2.xii.1926, K. & E. Carnaby (ANIC); 1 , Murchison, WA,
x.1935, H. Brown (ANIC); 1 , Denmark, larva in Pultanea stem: 14.i.1975; emerged 18.iv.1975, R. P.
McMillan (WAM); 1 , Cottesloe, cut from stem of Jacksonia, R. P. McMillan (WAM); 1 , Yallingup,
22.xii.1979, ex Eucalyptus, R. M. Bohart (ANIC); 1 , near Howatharra Hill, 25 km E by NE of Geraldton,

THONGPHAK & WANG
66 · Zootaxa 1569 © 2007 Magnolia Press
28.x.1992, E. D. Edwards & E. S. Nielsen (ANIC); 1 , Bruce Rock, i.1951, A. Douglas Collection (WAM); 1
, Aquinas (32º01'S, 115º51'E), 30.v.1967, Murchison (WAM); 1 , Star swamp, North Beach (Perth),
3.xi.1983, on flower of Xanthorrhoea, R. P. McMillan (WAM); 1 , Lake Esperance, E. Sutton Collection
(QM); 1 , 2 km N of Mundaring, 19.xii.1962, J. Dell (WAM); 1 , Drummond Cove (28º40'S, 114º36'E),
Geraldton, 24.ix.1972, light trap, N. MaCfresh (WADA); 1 , Swan River (32º03'S, 115º44'E), L. J. Newman
(WADA); 1 , same locality as above but xi (no year), Little (WADA); 1 , same locality as above but 1869,
de Boulay (HMO); 1 , Kings Park (31º57'S, 115º49'E), 23.ix.1963 (WADA); 1 , Cottesloe (31º59'S,
115º45'E), larva 20.xi.1975, adult 4.iii.1976, as larva in stem Jacksonia sternbergiana, R. P. McMillan
(WAM); 3 , same locality and host as above but v.1987 (WAM); 1 , same locality as above but larva
16.viii.1975, adult 6.ix.1975, cut from stem of Jacksonia sternbergiana, R. P. McMillan (WAM); 1 , Ene-
abba (29º49'S, 115º16'E), 26.x.1993, R. P. McMillan (WAM); 1 , 1 , Jurien Bay (30º17'S, 115º02'E),
30.i.1976, A. M & M. J. Douglas (WAM); 3 , Talbot Road Res. (31º52'S, 116º02'E), 17.iv.1967, in Jacksonia
sternbergiana, R. P. McMillan (WAM); 1 , South Perth (31º58'S, 115º51'E), 19.x.1971, S. J. Curay (WADA);
1 , Maylands (31º55'S, 115º53'E), 5.xi.1973, Exlucernce tree, P. S. Lawrence (WADA); 1 , Coorow
(29º52'S, 116º01'E), 15.x.1990, UV light, A. Szito (WADA); 1 , Jandakot (32º07'S, 115º50'E), S Perth,
22.xi.1977, dead on ground, A. Page (WAM); 1 , Bayswater (31º55'S, 115º55'E), 19.i.1966, B. G. Muir
(WAM); 1 , Salter Point (Perth) (32º01'S, 115º52'E), 28.x.1983, L. E. Koch (WAM); 1 , Perth (NMV); 2 ,
Thomas River, 63 mi E of Esperance (33?51 S, 121?53 E), 20.xi.1969, UV light, E. B. Briton (ANIC); 1 ,
Mt Yorkrakine (32º22'S, 117º35'E), 27.x.1957, I. M. (ANIC); 1 , 9km W of Hopetoun (33º57'S, 120º07'E),
23.x.1971, UV light, D. &N. McFarland (ANIC); 1 , South Perth, 5.x.1905, H. M. Giles (NMV); 1 , Glen
Forrest (33º14'S, 115º55'E), WA, xi.1950, A. B. (ANIC); 1 , 11 km N Geraldton (28º46'S, 114º36'E) WA,
25.x.1972, UV light (ANIC); 1 , no data ( HMO); 2 , Pt D’Entrecasteaux (34°50'S, 115°57'E), 6.xi.1926, K.
R. Norris (ANIC); 1 , 11 miles E of Pingrup, 21–27.xi.1958, Greave & Dowse (ANIC). TAS: 1 , Kingston,
TAS, 2.i.1947, J. R. Cunningham (AM); 1 , same data as above but 23.ii.1946 (AM); 1 ? (no abdomen),
Freycinet Nation Park, 28.ii.1963, I. F. B. & M. S. Upton (ANIC); Locality unknown: 2 , K 36096 (AM); 1
, 1 , K10942 (AM); 1 , K22588 (AM); 1 , 3455, Relton Bequest (QM); 1 , C. French Collection
(NMV); 1 , same data as above but (QDPI); 1 , F. E. Wilson Coll. (NMV); 1 , L. C. Haines Collection
(AM); 1 , no data (ANIC); 2 , 6 , no data (NMV); 1 , no data but (SAM).
Uracanthus discicollis Lea, 1916
(Figs 52, 90, 112)
Uracanthus discicollis Lea, 1916: 373.—McKeown, 1947: 63.
Description
Male. Body length, 13.58–25.34 mm; width, 2.52–4.93 mm.
Colour (Fig. 52). Body reddish brown to brown with head, prothorax and basal elytra darker. Head with
sparse to fairly dense white or pale yellow pubescence. Pronotal disc with 2 longitudinal stripes of very dense
white pubescence; each side of pronotum with a longitudinal stripe of very dense white pubescence near ven-
tral side; the remaining pronotum glabrous or very sparsely pubescent; prosternum with fairly dense white
pubescence. Each elytron with 2 longitudinal stripes of very dense whitish pubescence: 1 wide on disc near
suture and 1 narrow near margin; the remaining elytra glabrous or very sparsely pubescent. Body colour var-
ies from blackish brown to reddish brown or brown.
Head. Postclypeus triangular or subtriangular, shiny and convex, with fairly dense fine punctures; fronto-
clypeal suture deep and wide in middle; distance between lower lobes of eyes 1.76–2.12 × as long as distance
between upper lobes of eyes; distance between upper lobes of eyes 1.05–1.13 × as long as distance between

Zootaxa 1569 © 2007 Magnolia Press · 67
REVISION OF URACANTHUS
eyes on ventral side; genal length 0.36–0.51 × as long as head width immediately below eyes. Antennae
slightly longer than body; segments 5–10 thin, subcylindrical in shape, and slightly produced on one side at
apex.
Thorax and abdomen. Pronotum 1.10–1.26 × as long as width, with a conical process at each side; poste-
rior margin 1.05–1.34 × as wide as anterior margin; pronotal disc very weakly binodulose or not nodulose in
middle area; disc and side smooth and shiny, with very feeble transverse rugae near posterior margin. Scutel-
lum semicircular, with fairly dense pubescence. Elytra 3.34–4.28 × as long as prothorax and 3.6–4.01 × as
long as shoulder width; basal 1/3 of elytra coarsely punctate and the remaining 2/3 finely punctuate toward
apex; elytral apex rounded at margin and spinose or processed at suture. The apical spine of the elytron varies
from sharp to none. Apex of terminal sternite distinctly emarginate.
Male terminalia. Apex of ventral median lobe strongly pointed and apex of dorsal lobe rounded or
broadly projected; ventral lobe shorter than dorsal lobe; spined region of internal sac not clearly divided, with
dense long simple spines (Fig. 90a). Eighth sternite obliquely truncate or even rounded at sides, very shal-
lowly emarginate at apex, with sparse long and short setae arising terminally; ventral surface with sparse
cloud-like processes (Fig. 90b). Eighth tergite slightly point at apex, with fairly dense basally-forked spines
and simple spines on ventral surface (Fig. 90c). Paramere long, 2.0–2.09 × as long as wide, gradually tapering
toward apex; apex with sparse to dense long and short setae (Fig. 90d).
Female. Body length, 13.71–24.55 mm; width, 2.54–4.83 mm.
Body more robust, antennae shorter than body; elytra 3.81–4.42 × as long as prothorax and 3.59–3.85 × as
long as shoulder width.
Ovipositor and spermatheca. Ovipositor relatively long; styli long, arising almost terminally with short
hairs (Fig. 90e). Spermatheca slightly curved; spermathecal gland arising near base (Fig. 90f).
Distribution
Northeastern New South Wales, Australian Capital Territory, southern and central Victoria, southeastern
South Australia, and southern and southwestern Western Australia (Fig. 112).
Biology
Hosts are unknown. Adults visit flowers of Melaleuca sp. Adults were collected by light trap between
October and January.
Comments
This species resembles U. bivitta but differs in having the pronotal disc mostly smooth and shiny; the ely-
tral apex rounded or with a sharp spine or process at the suture; each side of the pronotum without 2 spots of
very dense yellowish pubescence.
Material examined
Holotype. . SA: Mount Lofty Rgs (34º07'S, 139º02'E), Coll. no. I.5696 (SAM).
Cotypes. 7 , 1 . SA: 2 , Kangaroo Island, J. G. O. Tepper (SAM); 1 , 1 , Karoonda to Peebinga, G.
E. H. Wright (SAM); 1 , Mount Lofty Rgs, Coll. no. 12586, S. H. Curnow (SAM); 1 , Marree (29º38'S,
138º03'E) (SAM); 1 , no locality, SA (SAM). WA : 1 , Albany, Helms (SAM).
Other material examined: 125 , 52 . NSW: 1 , 1 , 40 mile S of Narrabri (30º19'S, 149º46'E), 19–
20.x.1973, Pilliga scrub, G. B. Monteith (UQIC); 1 , 1 , W. Wyalong (33º54'S, 147º14'E), 24.xi.1963, J. C.
Le Sovef (ANIC); 1 , 23 km W by SW of Ranklin Spring (33º43'S, 146º03'E), Gap Dam SF., 3.xii.1992, at
light, McEvoy Moulds & McAlpine (AM); 1 , Jibbon beach (34º04'S, 151º10'E), Port Macquarie, x.1928, F.
H. Rodda (AM); 1 , West Pymble (33º45'S, 151º07'E), nr Sydney, 27.x.1983, at UV light, D. J. Scambler
(AM); 2 , 23 km N by NE of Coonabarabran (30º09'S,149º24'E), 7.xii.1974, I. F. B. Common & E. D.

THONGPHAK & WANG
68 · Zootaxa 1569 © 2007 Magnolia Press
Edwards (ANIC); 1 , 9 km W of Coonabarabran, 2.xii.1974, I. F. B. Common & E. D. Edwards (ANIC); 1 ,
1 , Sydney, 9.xi.1930, K. K. Spence (AM); 1 , NSW (AM); 1 , same data as above but (NMV); 3 , Mt.
Kaputar (30º16'S,150º10'E), NSW, 300 ft., 30.x.1967, at light, C. W. Frazier (ANIC); 1 , same data as above
but 18.xi.1968 (ANIC);1 , 2 , Mulla (31º49'S, 147º19'E), xi.1930, H. J. Carter (ANIC). ACT: 1 , Black
Mt. (35º16'S, 149º06'E), 28.xi.1977, E. C. Zimmerman Collection (ANIC). VIC: 3 , 1 , 30 km NW of
Rainbow, Big Desert National Park (35º43'S, 141º46'E), Milmed Rock Track, 1.xii.1992, at light, McAlpine,
McEvoy & Moulds (AM); 1 , 14 km S. of Murrayville (35º15'S, 141º11'E), Big Desert Wilderness,
31.x.1982, J. & D. Gardner (SAM); 1 , 44 km E by SE of Murrayville (35º38'S, 141º98'E), 2.xi.1985, The
Springs, Big Desert, 2.xi.1985 (NMV); 1 , Wyperfeld National Park (35º37'S, 142º01'E), Ranger’s House,
light trap, 17.xi.1973, Misko & Anderson (ANIC); 1 , Glenaladale (37º45'S, 147º19'E), 1.xii.1978, J. C. Le
Sovef (ANIC); 1 , 13 mile S. of Kiata, Little Desert (36º33'S, 141º38'E), 7.xi.1966, I. F. B. Common & M.S.
Upton (ANIC); 1 , Little Desert (36º33'S, 141º38'E), 13.xi.1942, A. D. Bishop (NMV); 1 , same locality as
above but 12.xi.1945, F. E. Wilson (NMV); 1 , Inglewood (36º34'S, 143º52'E), 24.xi.1952, J. C. Le Sovef
(ANIC); 1 , 1 , same locality as above but 21.xi.1945, I. Smith (NMV); 1 , Kiata (36º21'S, 141º47'E),
12.xi.1945, A. B. (ANIC); 1 , same data as above but (NMV); 3 , same data as above but 7.xi.1945, A. B.
(ANIC ); 3 , same locality as above but 5.xi.1949, J. C. Le Sovef (ANIC); 1 , same data as above but
26.x.1967, J. C. Le Sovef (ANIC); 1 , Birchip (35º58'S, 142º54'E), xii.1900 (NMV); 1 , Bacchus Marsh
(37º40'S, 144º26'E), xi.1908, C. Oke (NMV); 4 , Gypsum (35º16'S, 142º23'E), xi.1924, C. Oke (NMV); 1 ,
Sea Lake (35º29'S, 142º51'E), ix.1917, Goudie (NMV). SA: 5 , 3 , Calpatanna Waterhole Conservation
Park, Wedina Well, Eyre Peninsular, 30.xi.1986, at light, J. A. Forrest (SAM); 4 , 4 , Yumbarra National
Park (31º39'S, 133º22'E), 11.xi.1975, J. A. Herridge (SAM); 4 , 2 , Cold & Wet station (35º44'S, 139º45'E),
19.xi.1962, at light, P. Aitken, G. Pretty & N. B. Tindale (SAM); 4 , 1 , Ngarkat Conservation Park
(35º50'S, 140º37'E), 18.xi.1991, Box Flat at light, J. A. Forrest (SAM); 1 , 5 km Scorpion Spring Conserva-
tion Park (35º27'S, 140º53'E), S. W. Namam’s Well, 16.xii.1983, at light, Museum Party (SAM); 1 , same
data as above but 17.xii.1983 (SAM); 1 , 1 , Athelstone (34º52'S, 138º42'E), 24.x.1967, UV light 7 pm–2
am, J. Szent & lvany (SAM); 1 , same data as above but 5.xi.1967 (SAM); 1 , same data as above but
19.xi.1967 (SAM); 1 , Blackwood (35º01'S, 138º36'E), 850 ft, 17.xi.1961, N. B. Tindale (SAM); 1 , 2 km
W of Blewitt Spring (35º10'S, 138º35'E), 9.xi.1976, 18 ºC, UV light, P. B. McMillan (SAM); 1 , Rupara
(33º26'S, 138º57'E), 2.xi.1922, J. D. Wilson (SAM); 1 , 1 , 2mi W Darke Peak (33º28'S, 136º12'E),
8.xii.1968, N. McFarland & N. B. Tindale (SAM); 1 , Ferries-McDonald Conservation Park (35º13'S,
139º08'E), 19.x.1988, A. Stolarski (SAM); 1 , Karkoo (34º01'S, 135º43'E) nr Pt. Linncoln, C. Kimber
(SAM); 1 , Coonalpyn (35º41'S, 139º51'E) (SAM); 1 , 1.5 km NNW Horne Lookout, E. Bascombe Wells
Conservation Park (33º39'S, 135º31'E), Kappawanta Basin Eyre Pen, at light, 2–3.xii.1986, J. A. Forrest
(SAM); 1 , 22 mile W by SW of Mundulla (36º21'S, 140º41'E), SA, 8.xi.1966, I. F. B. Common & M. S.
Upton (ANIC); 1 , Kuitpo Forrest (35º13'S, 138º42'E), 30.vii.1979, on Melaleuca flower, F. D. M (WINC);
2 , Yeelanna (SAM); 1 , Waite Institute, 16.xi.1955, light trap (WINC); 1 , no locality label, SA, A. H. C.
Zietz (SAM); 1 , SA, A. H. Elston, A. H. Elston Collection(AM); 1 , SA (AM);1 , SA (MAM). WA: 1 ,
20 km N of Leeman (29º45'S, 114º58'E), Gum Tree Bay, 6.x.1993, R. P. McMillan (WAM); 1 , same data as
above but 7.x.1993 (WAM); 2 , 1 , 1 km S of Leeman (29º58'S, 114º59'E), 30.x.1992, E. D. Edwards & E.
S. Nielsen (ANIC); 1 , 6 mile West of Hopetoun (33º51'S, 120º07'E), 25.xi.1968, N. McFarland (SAM); 2 ,
Salmon Gums (32º58'S, 121º38'E), Coll. no. 42-389 (WAM); 1 , Scaddan (32º27'S, 121º44'E), 25.i.1977, A.
& M. J. Douglas (WAM); 1 , Watheroo National Park (30º12'S, 115º50'E), 3–5.x.1980, T. F. Houston
(WAM);1 , Eneabba (29º49'S, 115º16'E), 25.x.1993, R. P. McMillan (WAM); 1 , Western Flora Caravan
Park, 22 km NNW Eneabba (29º37'39″S, 115º13'26″E), 4.xi.1999, T. F. Houston (WAM); 1 , Robb Road
Bushland, 26 km N of Eneabba (29º35'3″S, 115º17'55″E), 18.xi.2000, T. F. Houston (WAM); 1 , 25 km N of
Eneabba (29º36'S, 115º15'E), 24–25.x.1984, A. A. Clader Coll.(ANIC); 1 , 2 , Coonadgee (31º22'S,
115º50'E), WA, (WAM); 1 , Claremont (31º58'S, 115º46'E), WA, 16.x.1978, R. P. McMillan (WAM); 1 , 5

Zootaxa 1569 © 2007 Magnolia Press · 69
REVISION OF URACANTHUS
mi W Denmark (34º58'S, 117º21'E), iii.1965, W. H. Butter (WAM); 1 , Perth (31º57'S, 115º51'E) (WAM); 1
, 1 , Wembley (31º56'S, 115º48'E), 6.xi. 1953 (WAM); 1 , Jurien Bay (30º17'S, 115º02'E), WA, 30.i. 1976,
A. M & M. J. Douglas (WAM); 1 , Mt Toolbrunup (34º23'S, 118º02'E), Stirling Ranges, WA, 7.xii.1970, MV
lamp, G. A. Holloway (AM); 1 , Moingup Spring (34º24'S, 118º26'E), Stirling Ranges, 7.xii.1970, MV lamp,
G. A. Holloway (AM); 1 , Sheoaks Hill (33º36'S, 123º39'E), W Israelite Bay, WA, 8.i.1993, E. D. Edwards &
E. S. Nielsen (AM); 1 , Geraldton (28º46'S, 114º36'E), WA, J. Clark (AM); 1 , same data as above but
(WADA); 1 , same locality as above but 12.xi.1981, K. &. E. Carnaby (ANIC); 1 , 50 km NW Yuna
(33º38'S, 123º40'E), 6.ix.1981, G. A. Holloway (AM); 3 , 15 mi NW of Mt Arid (33º49'S, 123º02'E),
13.xi.1969, M. S. Upton (ANIC); 3 , Boxwood Hill (34º21'S, 118º44'E), WA, 6–7.xi.1980, K. & E. Carnaby
(ANIC); 4 , 23 km NW by W of Mt. Arid (33º51'S, 123º00'E), Thomas River, WA, 4–7.xi.1977, at light, J. F.
Lawrence (ANIC); 1 , 1 , 101 km. E of Esperance (33º51'S, 121º53'E), Thomas River, WA, 20.xi.1962, at
light, beach dumes, Britton, Taylor & Upton (ANIC); 1 , same locality as above but E. of Esperance
(33º51'S, 121º53'E), WA, 20.xi.1962, UV light, E. B. Britton (ANIC); 1 , same data as above but 19.xi.1969
(ANIC); 2 , 11 mi. S by W of Cocklebiddy (32º12'S, 126º03'E), 2.xi.1969, Key & Upton (ANIC); 2 , 2mi S
by SW of Mt Ragged (33º27'S, 123º28'E), WA, 13.xi.1969, M. S. Upton (ANIC); 1 , Swan River (32º03'S,
155º44'E), WA, J. Clark (WADA); 1, same locality as above but 30.xi.(no year) (WADA); 1 , Yanchep
(31º33'S, 115º41'E), 25.x.1969, at light trap, S. J. Curry (WADA); 1 , same data as above but 26.x.1969
(WADA); 1 , 23 mi W of Fraser Range (32º03'S, 122º47'E), 7.xi.1969, M. S. Upton (ANIC); 1 , Carnarvon
(24º53'S, 113º39'E), Blow Holes Rd., 3.ix.1978, K. & E. Carnaby (ANIC); 1 , 12 km S by E Kalbarri
National Park (27º49'S, 114º11'E), 19.x.1984, at MV light, A. A. Calder Collection (ANIC); 1 , 34 Km NE
by N of Laverton (28º22'S, 122º37'E), Deeba Rock Hole, 12.xi.1977, T. A. Weir (ANIC); 2 , 2 , 11 mi. E. of
Pingrup (33º36'S, 118º36'E), WA, 21–27.xi.1958, at light, Greaves & Dowse (ANIC); 1 , Junana Rock, 9 km
NW of Mt Ragged (33º23'S, 123º24'E), 26.x.1977, J. F. Lawrence (ANIC); 1 , Dongara, 5.xi.1958, at light,
Greaves & Dowse (ANIC); 1 , Bremer Bay (34º24'S, 119º26'E), 15.xi.1941, K. K. Norris (ANIC); 3 , Woo-
dridge (31º20'S, 115º35'E), 20–30.x.1996, intercept trap, H. Demarz (WADA); 3 , same data as above but
x.1997 (WADA); 1 , Perth, 1913, H. M. Giles (HMO); 1 , Tachalarup, 25.x.1938, at light trap, D. Hardy
(WADA); Locality unknown: 1 , K 58326 (AM); 1 , Australia old Collection (ANIC). 1 , no data
(VAIC).
Uracanthus corrugicollis Lea, 1917a
(Figs 53, 91, 113)
Uracanthus corrugicollis Lea, 1917a: 742.—McKeown, 1947: 63.
Description
Male. Body length, 25.26 mm; width, 5.41 mm.
Colour (Fig. 53). Body reddish brown with head, thorax, and basal and apical elytra darker. Head, prono-
tum and elytra covered with fairly dense, more or less evenly distributed, whitish pubescence; pubescence on
elytra generally shorter.
Head. Postclypeus subtriangular and flattened, with fairly dense coarse punctures; frontoclypeal suture
deep and wide in middle; distance between lower lobes of eyes 1.8 × as long as distance between upper lobes
of eyes; distance between upper lobes of eyes 1.25 × as long as distance between eyes on ventral side; genal
length 0.19 × as long as head width immediately below eyes. Antennae largely damaged.
Thorax and abdomen. Pronotum 0.94 × as long as width, rounded with a small process at each side; pos-
terior margin 1.22 × as wide as anterior margin; pronotal disc without distinct nodules in middle area; disc and
side strongly rugose transversely. Scutellum semicircular, with sparse pubescence. Elytra 5.26 × as long as

THONGPHAK & WANG
70 · Zootaxa 1569 © 2007 Magnolia Press
prothorax and 3.44 × as long as shoulder width; basal 1/3 with coarsely punctures; apex more or less rounded
at margin and with a small spine at suture. Apex of terminal sternite rounded or slightly truncate.
Male terminalia. Apex of ventral median lobe emarginate and apex of dorsal lobe with rounded; ventral
lobe distinctly longer than dorsal lobe; spined region of internal sac divided into two sections; first section
with sparse short simple spines on sides near base; second section with fairly dense long simple spines;
unspined gap between first and second section above as long as second section (Fig. 91a). Eighth sternite
rounded at side, shallowly emarginate at apex, with long and fairly long setae arising terminally; ventral sur-
face with sparse cloud-like processes (Fig. 91b). Eighth tergite rounded at apex, with fairly dense simple
spines and multi- branched spine on surface (Fig. 91c). Paramere long, 2.5 × as long as wide, cylindrical in
shape, apex rounded with long and short setae (Fig. 91d).
Female. Unknown.
Distribution
Southwestern Western Australia (Fig. 113).
Comments
This species differ from others species of Uracanthus in having very short genal length; pubescence on
the elytra uniform without any line or stripe of denser pubescence. No other specimen matches the type.
Material examined
Holotype. . WA: Mullewa (28º26'S, 115º35'E), Miss J. F. May; bearing a name label on which ‘TYPE’
was written in red; 8 segments of left antenna and 8 segment of right antenna damaged (SAM).
Uracanthus lateroalbus Lea 1916
(Figs 54, 92, 113)
U. lateroalbus Lea, 1916: 378.—McKeown, 1947: 65.
U. fuscostriatus McKeown, 1948: 55. syn. nov.
Description
Male. Body length, 26.63–28.93 mm; width, 5.25–5.73 mm.
Colour (Fig. 54). Body reddish brown with head, thorax, legs and basal elytra darker. Head with fairly
dense pale yellow and white pubescence. Pronotum with mixture of fairly dense whitish and yellowish pubes-
cence; each side with a longitudinal stripe of very dense white pubescence near ventral side. Elytra covered
with dense yellowish pubescence on disc; pubescence near suture and margin (basal elytra in particular)
almost white; each elytron with a narrow glabrous longitudinal stripe starting from shoulder and extending to
about basal 1/3 of elytra. Colour varies from reddish brown to blackish.
Head. Postclypeus triangular, convex, coarsely punctuate; frontoclypeal suture deep and wide in middle;
distance between lower lobes of eyes 2.08–2.3 × as long as distance between upper lobes of eyes; distance
between upper lobes of eyes 1.0–1.04 × as long as distance between eyes on ventral side; genal length 0.32–
0.39 ×as long as head width immediately below eyes. Antennae slightly shorter than body; segments 4–10
flattened and produced on one side at apex.
Thorax and abdomen. Pronotum 1.15–1.30 × as long as width, sides rounded with a small rounded pro-
cess at each side; posterior margin 1.09–1.18 × as wide as anterior margin; pronotal disc weakly binodulose in
middle area; disc and sides strongly rugose transversely, particularly near anterior and posterior margins.
Scutellum semicircular, with dense pubescence. Elytra 4.15–5.04 × as long as prothorax and 3.60–3.90 ×as
long as shoulder width; elytron finely puncture; apex slightly emarginated with a small process at margin and

Zootaxa 1569 © 2007 Magnolia Press · 71
REVISION OF URACANTHUS
a small spine at suture. The apical spine of the elytron at the suture may vary from very small to relatively
long. Apex of terminal sternite truncate with a small notch.
Male terminalia. Apex of ventral median lobe pointed and apex of dorsal lobe rounded; dorsal lobe about
as long as ventral lobe; spined region of internal sac almost occupying the entire internal sac, divided into two
sections: first section about as long as second section, with very dense long simple spines; second section with
fairly dense multi-branched spines at base near first section and sparse simple spines on the remaining part
(Fig. 92a). Eighth sternite rounded or slightly obliquely truncate at sides; emarginate at apex, with long and
fairly long setae arising terminally; ventral surface with short simple spines (in mid area) and fairly dense
cloud-like processes (Fig. 92b). Eighth tergite truncate at apex, with fairly dense simple spines and multi-
branched spines on surface (Fig. 92c). Paramere 2.3–2.38 × as long as wide, cylindrical in shape; apex
rounded with long and short setae (Fig. 92d).
Female. Body length, 21.39–28.67 mm; width, 4.96–5.51 mm.
Antennae and legs distinctly shorter, body broader. Elytra 3.10–3.64 × as long as shoulder width and
4.31–4.91 × as long as prothorax.
Ovipositor and spermatheca. Ovipositor short; styli arising terminally with short hairs (Fig. 92e). Sper-
matheca C-shaped; spermathecal gland arising at base (Fig. 92f).
Distribution
Southwestern Western Australia (Fig. 113).
Biology
Hosts are unknown. Adults were collected in February and December.
Comments
This species resembles U. discicollis but differs in having pubescence on the pronotal disc more or less
evenly distributed; the pronotal disc strongly rugose transversely, and the glabrous areas on the elytra only
occurring laterally between the shoulder and basal 1/3 of the elytra.
Material examined
U. lateroalbus: Holotype. . WA: Swan River (32º03'S, 115º44'E), Coll. no. I. 5695, bearing a name label
on which ‘TYPE’ was written in red; 8 segments of left antenna and 1 segment of right antenna missing
(SAM).
U. fuscostriatus: Holotype. . WA: Maylands (31º55'S, 155º28'E), Coll. no. 32-242; bearing a red holo-
type label; 4 segments of left antenna missing (WAM).
Other material examined. 4 , 2 .WA: 1 , Swan River, J. Clark (SAM); 1 , Jandakot (south of Perth),
12.ii.1950, F. H. Uther Baker (WADA); 1 , Dumbleyung (33º15'S, 117º14'E), 26.xii.1962 (WAM); 1 , Sor-
rento (31º49'S, 115º44'E), 6.xii.1972, P. G. & A. J. Kendrick (WAM); 1 , 1 , no locality, WA (WADA).
Uracanthus bistriolatus, sp.nov.
(Figs 55, 93, 113)
Description
Male. Body length, 24.82–29.29 mm; width, 4.49–5.36 mm.
Colour (Fig. 55). Body dark reddish brown with head, thorax, and basal elytra darker. Head with fairly
dense white pubescence. Antennal segments 3–11 with very short white fringe on one side. Pronotal disc with
fairly dense white pubescence divided by a very narrow glabrous median line; each side of pronotum with a

THONGPHAK & WANG
72 · Zootaxa 1569 © 2007 Magnolia Press
longitudinal stripe of very dense white pubescence near ventral side, and a wide glabrous longitudinal stripe
between pubescent area on disc and pubescent stripe near ventral side. Elytra covered with more or less
evenly distributed dense short white pubescence, with pubescence denser and longer on base and margins;
each elytron with a narrow glabrous longitudinal stripe ranging from shoulder to basal 1/4 of elytra. Body
colour varies from dark reddish to blackish brown.
Head. Postclypeus triangular, convex, finely punctate; frontoclypeal suture deep and wide in middle; dis-
tance between lower lobes of eyes 2.06–2.46 × as long as distance between upper lobes of eyes; distance
between upper lobes of eyes 0.78–0.94 × as long as distance between eyes on ventral side; genal length 0.28–
0.36 × as long as head width immediately below eyes. Antennae longer than body; segments 4–10 flattened
and produced on one side at apex.
Thorax and abdomen. Pronotum 1.09–1.25 × as long as width, with a small rounded process at each side;
posterior margin 1.08–1.35 × as wide as anterior margin; pronotal disc weakly binodulose in middle area; disc
and side strongly rugose transversely. Scutellum semicircular, with dense pubescence. Elytra 4.6–4.94 × as
long as prothorax and 3.67–3.95 × as long as shoulder width; elytron disc with fine punctures; apex bispinose.
The apical spine at the suture may be reduced to a small process. Apex of terminal sternite truncate or slightly
emarginate.
Male terminalia. Apex of ventral median lobe with a small notch and apex of dorsal lobe with rounded;
dorsal lobe as long as or slightly longer than ventral lobe; spined region occupying almost all internal sac,
divided into two sections: first section about as long as second section, with dense long simple spines; second
section with sparse short simple spines (Fig. 93a). Eighth sternite rounded at side, shallowly emarginate at
apex, with long and fairly long setae arising terminally; ventral surface with simple spines and cloud-like pro-
cesses (Fig. 93b). Eighth tergite truncate, shallowly emarginate at apex, with fairly dense basally-forked
spines on surface of basal ½ of tergite (Fig. 93c). Paramere long, 2.22–2.27 × as long as wide, cylindrical in
shape, apex rounded with long and short setae (Fig. 93d).
Female. Body length, 29.81 mm; width, 5.47 mm.
Antennae as long as or slightly shorter than body; antennal segments without white fringe; elytra 3.89 × as
long as shoulder width and 4.71 × as long as prothorax.
Ovipositor and spermatheca. Ovipositor long; styli short, arising terminally with short and long hairs
(Fig. 93e). Spermatheca curved; spermathecal gland arising at base (Fig. 93f).
Distribution
This species is found only in southeastern Queensland (Fig. 113).
Biology
Hosts are unknown. Adults were collected at light in December and February.
Comments
This new species resembles U. lateroalbus but differs in having the pronotum with a glabrous stripe on
each side; the glabrous stripe on the lateral elytron shorter; the elytra usually bispinose, and male antennae
usually longer than body, with a short white fringe on one side.
Material examined
Holotype. . QLD: Carnarvon Range (25º17'S, 148º44'E), 16.i.1940, N. Geary; bearing a red holotype
label (AM).
Paratypes. 7 , 1 . QLD: 2 , same data as holotype (AM); 1 , same data as above but 11.i.1940 (AM);
3 , 1 , 560m Amphitheare Camp, Expedition Range National Park (25º12'S, 148º59'E), 17.xii.1997, MV
light in open forest, Monteith, Cook and Thompson (QM); 1 , same data as above but Evens, Burwell and
Ewart (QM).

Zootaxa 1569 © 2007 Magnolia Press · 73
REVISION OF URACANTHUS
Uracanthus suturalis Lea, 1916
(Figs 56, 113)
Uracanthus suturalis Lea, 1916: 380.—McKeown, 1947: 66.
Description
Male. Body length 19.14–21.60 mm; width, 3.83–4.34 mm.
Colour (Fig. 56). Body blackish brown, with head, thorax and basal elytra almost back. Head with fairly
dense white pubescence. Pronotal disc covered with dense white pubescence, with 5 glabrous areas: 4 more or
less round spots (2 on each side) and 1 longitudinal stripe (on middle line) starting from posterior margin and
extending to posterior 1/3; each side of pronotum with a wide longitudinal stripe of very dense white pubes-
cence near ventral side, and a wide longitudinal glabrous area between pubescent disc and pubescent stripe
near ventral side. each elytron with 6–7 very narrow longitudinal lines of white pubescence interspaced with
glabrous longitudinal lines on disc, and a wide longitudinal stripe of dense white pubescence on margin.
Head. Postclypeus semicircular or subtriangular, slightly convex, with densely fine punctures; frontocly-
peal suture deep and narrow in middle; distance between lower lobes of eyes 1.60–1.87 × as long as distance
between upper lobes of eyes; distance between upper lobes of eyes 1.45–1.53 × as long as distance between
eyes on ventral side; genal length 0.31–0.33 × as long as head width immediately below eyes; ventral side of
head with transverse wrinkles. Antennae slightly longer than body; segments 4–10 flattened and strongly pro-
duced on one side at apex.
Thorax and abdomen. Pronotum 1.02–1.06 × as long as width, conical process at side; posterior margin
1.25–1.32 × as wide as anterior margin; pronotal disc with 4 weakly raised and glabrous nodules; pronotal
disc smooth with transverse rugae near posterior margin; side strongly rugose transversely. Scutellum semicir-
cular, with dense pubescence. Elytra 4.45–4.88 × as long as prothorax and 3.63–3.67 × as long as shoulder
width; elytron finely punctate toward apex; apex with an acute spine at suture.
Male terminalia. Unknown. The unique type specimen was not dissected.
Female. Unknown
Distribution
Southern South Australia, and southern Western Australia (Fig. 113).
Biology
Hosts are unknown. Adults were collected in October.
Comments
This species resembles U. ater but differs in having the elytral apex with an acute spine at the suture and
the elytra with clearly defined longitudinal lines of white pubescence.
Material examined
Holotype. . SA: Murat Bay (32º07'S, 133º37'E), E. A. King, Coll. no. I. 5692, bearing a type label and a
species name label on which ‘TYPE’ was written in red (SAM).
Other material examined. WA: 1 , 9 km NW of Mt. Ragged (33º27'S, 123º28'E), Junana Rock,
26.x.1977, no abdomen, J. F. Lawrence (ANIC).
Uracanthus ater Lea, 1917a
(Fig. 57)
Uracanthus ater Lea, 1917a: 742.–Lea, 1917b: 619; McKeown, 1947: 63.

THONGPHAK & WANG
74 · Zootaxa 1569 © 2007 Magnolia Press
Description
Male. Body length, 21.50 mm; width, 4.39 mm.
Colour (Fig. 57). Body blackish brown except middle area of elytra reddish brown. Head, pronotum and
elytra with more or less evenly distributed white pubescence.
Head. Postclypeus semicircular and flattened, with sparse coarse punctures; frontoclypeal suture deep and
narrow in middle; distance between lower lobes of eyes 1.53 × as long as distance between upper lobes of
eyes; distance between upper lobes of eyes 1.08 × as long as distance between eyes on ventral side; genal
length 0.24 × as long as head width immediately below eyes. Antennae about as long as body; segments 4–10
flattened and produced on one side at apex.
Thorax and abdomen. Pronotum 1.22 × as long as width, with a small conical process at each side; poste-
rior margin 1.46 × as wide as anterior margin; pronotal disc binodulose in middle area; disc and side strongly
rugose transversely. Scutellum semicircular, with dense pubescence. Elytra 4.03 × as long as prothorax and
3.60 × as long as shoulder width; elytra coarsely and shallowly punctate from base to apex; apex slightly
emarginate, with a small spine at suture.
Male terminalia. Unknown.
Female. Unknown.
Distribution
Central Australia.
Comments
This species resembles U. suturalis but differs in having the elytral apex slightly emarginate with a very
small spine at the suture; pubescence on elytra more or less evenly distributed.
Material examined
Holotype. , Central Australia, type no. 18528, abdomen and 6 segments of right antenna missing, H.
Hacker; bearing a type label, and a name label on which ‘TYPE’ was written in red (SAM).
Acknowledgements
We would like to thank M. Moulds (AM), T. Weir (ANIC), M. Fletcher (ASCU), S. Shute (BMNH), D. Kent
(FCNI), J. Hogan (HMO), M. Humphrey (MAM), G. Dally (NTM), P. Lillywhite (NMV), G. B. Monteith
(QM), J. Donaldson (QDPI), J. Forrest (SAM), G. Daniel (UQIC), A. Szito and N. Zlim (WADA), T. Houston
and B. Hanich (WAM), J. Jennings (WINC), and M. Malipatil (VAIC) for making available the material
included in this study. R. de Keyzer and an anonimous referee made thorough corrections and constructive
comments on the manuscript, for which we are greatly grateful. This study was partially supported by a schol-
arship from the Asian Development Bank (ADB) loan to CAB (loan no. 1699-THA), a Royal Thai Govern-
ment Scholarship, and financial assistance from Massey University.
References
Allen, W.J., Blunno, M., Froggatt, W.W. & Guthrie, F.B. (1898a) Insect and fungus diseases of fruit-tree and their reme-
dies. Agricultural Gazette of New South Wales, 9, 1028–1043.
Allen, W.J., Blunno, M., Froggatt, W.W. & Guthrie, F.B. (1898b) Insect and fungus diseases of fruit-tree and their reme-
dies. Agricultural Gazette of New South Wales, 9, 1216–1221.
Aurivillius, C. (1912) Coleopterorum Catalogus. Pars 39, Cerambycidae: Cerambycinae. W. Junk, Berlin, 574 pp.

Zootaxa 1569 © 2007 Magnolia Press · 75
REVISION OF URACANTHUS
Aurivillius, C. (1917) Results of Dr. E. Mjöberg’s Swedish scientific expeditions to Australia 1910–1913, Part XII. Cer-
ambycidae. Arkiv för Zoologi, 10, 1–50.
Bense, U. (1995) Longhorn Beetles: Illustrated Key to the Cerambycidae and Vesperidae of Europe, Margraf, Germany,
512 pp.
Best, D. (1882) Longicorn beetles of Victoria, Part V. The Southern Science Record, 2, 35–38.
Blackburn, T. (1889) Notes on Australian Coleoptera, with descriptions of new species, Part III. Proceedings of the Lin-
nean Society of New South Wales, 4, 445–482.
Blackburn, T. (1894) Notes on Australian Coleoptera, with descriptions of new species, Part XV. Proceedings of the Lin-
nean Society of New South Wales, 9, 85–108.
Boisduval, J. B. A. (1835) Faune entomologique de l’ ocean pacifique, deuxième partie. Coleopteres et autres orders.
Voyage de Découvertes de l’Astrolabe, execute par order du Roi, pendant des Années 1826–29, 2, 1–717, Tastu,
Paris.
Brimblecombe, A. R. (1943) Citrus branch borer. Queensland Agricultural Journal, 57, 37–39.
Dixon, J. E. (1908) Food plants of Victorian longicorn beetles. Victorian Nature, 25, 77.
Duffy, E. A. J. (1963) A Monograph of Immature Stages of Australian Timber Beetles (Cerambycidae), Bristish Museum
(Natural History), London, 235 pp.
Ehara, S. (1954) Comparative anatomy of male genitalia in some cerambycid beetles (Coleoptera: Cerambycidae). Jour-
nal of Faculty of Science, Hokkaido University, Series VI, Zoology, 12, 61–114.
French, C. (1900) A Handbook of Destructive Insects of Victoria, with Notes on the Methods to Be Adopted to Check and
Extirpate Them, Part III. Government Printer, Melbourne, 229 pp.
French, C. (1911) A Handbook of Destructive Insects of Victoria, with Notes on the Methods of Prevention and Extirpa-
tation, Part V. Government Printer, Melbourne, 169 pp.
Froggatt, W.W. (1893) On the life-histories of Australian Coleoptera, Part I. Proceedings of the Linnean Society of New
South Wales, 8, 27–42.
Froggatt, W.W. (1894) On the life-histories of Australian Coleoptera. Part II. Proceedings of the Linnean Society of New
South Wales, 9, 113–125.
Froggatt, W.W. (1898) Fruit–tree and vine pests. Agricultural Gazette of New South Wales, 9, 41–47.
Froggatt, W. W. (1902) Insects of wattle trees. Agricultural Gazette of New South Wales, 13, 701–720.
Froggatt, W. W. (1907) Australian Insects. W. Brooks & Co., Sydney, 449 pp.
Froggatt, W. W. (1919) The native lime-tree borer. Agricultural Gazette of New South Wales, 30, 261–267.
Froggatt, W. W. (1923) Forest Insects of Australia. Forestry Commission of New South Wales, Sydney, 171 pp.
Gahan, C. J. (1893) Notes on the longiconia of Australia and Tasmania. Part I, with a list of the species collected by Mr.
J. J. Walker, and descriptions of new forms. Transactions of the Entomological Society of London, 2, 165–197.
Gallard L. (1916) List of insects associated with Acacia decurrens. Australian Nature, 3, 112–114.
Gressitt, J. L. (1951) Longicorn beetles from New Guinea and the South Pacific (Coleoptera: Cerambycidae). Annals of
the Entomological Society of America, 44, 1–30.
Gressitt, J. L. (1959) Longicorn beetles from New Guinea, I (Cerambycidae). Pacific Insects, 1, 59–171.
Gurney, W. B. (1911) A study of wattle trees (Acacia) and a list of insects of wattle trees. Australian Nature, 2, 56–59.
Hawkeswood, T. J. (2002) Review of the biology and host plants of the Australian longicorn beetle Uracanthus triangu-
laris (Hope, 1833) (Coleoptera: Cerambycidae). Entomologische Zeitschrift mit Insekten-Borse, 112, 59–62.
Hope, F.W. (1833) On the characters of several new genera and species of coleopterous insects. Proceedings of the Zoo-
logical Society of London, 1, 60–64.
Hope, F.W. (1834) Characters and descriptions of several new genera and species of coleopterous insects. Transactions
of the Zoological Society of London, 1, 91–112.
Hope, F.W. (1841a) An extract of observations on the Stenochoridae of New Holland, with description of new genera and
species of that family. Proceeding of the Zoological Society of London, 8, 46–55.
Hope, F.W. (1841b) Observations on the Stenochoridae of New Holland with descriptions of new species. The Annals
and Magazine of Natural History, 7, 58–67.
Hope, F.W. (1844) Observations on the Stenochoridae of New Holland with description of new genera and species of that
family. Transactions of the Zoological Society of London, 3, 187–201.
Lacordaire, J. T. (1869) Histoire Naturelle des Insectes. Genera des Coléoptères ou exposé Methodique et Critique de
tous les Genres Proposés jusqu’ici dans cet Ordre d’Insectes. VIII. Tricténotomides et Logicornes. Roret, Paris, 552
pp.
Laporte, F. L. (1840) Histoire Naturelle des Insectes. Tom. I–II. Coléoptères. P. Duménil, Paris, 454 pp.
Lawrence, J. F. & Britton, E. B. (1991) Coleoptera. In The Insects of Australia: A Textbook for Students and Research
Workers. (ed. CSIRO), Melbourne University Press, Melbourne, pp 543–683.
Lea, A. M. (1916) Notes on some miscellaneous Coleoptera with descriptions of new species, Part II. Transactions of the
Royal Society of South Australia, 40, 272–436.
Lea, A. M. (1917a) Descriptions of new species of Australian Coleoptera, Part XII. Proceedings of the Linnean Society

THONGPHAK & WANG
76 · Zootaxa 1569 © 2007 Magnolia Press
of New South Wales, 41, 720–745.
Lea, A. M. (1917b) Results of the South Australian Museum expedition to Strzelecki and Cooper Creeks, September and
October, 1916, Insecta. Transactions of the Royal Society of South Australia, 41, 489–630.
Lu, W. & Wang, Q. (2005a) Systematics of the New Zealand longicorn beetle genus Oemona Newman with discussion of
the taxonomic position of the Australian species, O. simplex White (Coleoptera: Cerambycidae: Cerambycinae).
Zootaxa, 971, 1–31.
Lu, W. & Wang, Q. (2005b) Systematics of the longicorn beetle genus Ophryops (Coleoptera: Cerambycidae). Inverte-
brate Systematics, 19, 169–188.
Matthew, E. G. (1997) A Guide to the Genera of Beetles of South Australia Part 7. Adelaide, South Australia, 61 pp.
McKeown, K. C. (1938) Notes on Australian Cerambycidae IV. Records of the Australian Museum, 20, 200–216.
McKeown, K. C. (1940) Notes on Australian Cerambycidae V. Records of the Australian Museum, 20, 293–312.
McKeown, K. C. (1942) Australian Cerambycidae VI: Descriptions of new species mostly from Queensland. Records of
the Australian Museum, 21, 86–88.
McKeown, K. C. (1947) Catalogue of the Cerambycidae (Coleoptera) of Australia. Memoirs of the Australia Museum,
10, 1–190.
McKeown, K. C. (1948) Australian Cerambycidae VIII. Notes on collection from the Western Australia Museum with
descriptions of new species. Records of the Australian Museum, 22, 49–63.
Monné, M. A. (2005) Catalogue of the Cerambycidae (Coleoptera) of the Neotropical Region. Part I. Subfamily Ceram-
bycinae. Zootaxa, 946, 1–765.
Moore, K. M. (1972) Observations on some Australian forest insects. 27. Some insects attacking Callitris hugelii (white
cypress pine). Australian Zoologist, 17, 40–46.
Newman, E. (1838) Entomological notes. Entomological Magazine, 5, 168–181.
Olliff, A. S. (1892) Entomological notes. Agricultural Gazette of New South Wales, 3, 895–899.
Pascoe, F. P. (1866) On the Longicornia of Australia, with a list of all the described species. Journal of the Linnean Soci-
ety of London, Zoology, 9, 80–112.
Pascoe, F. P. (1875) Notes on Coleoptera, with descriptions of new genera and species. Part III. The Annals and Maga-
zine of Natural History, 15, 59–73.
Pierce, W. D. (1917) A Manual of Dangerous Insects Likely to Be Introduced in the United States through Importation.
U.S. Government Printing Office, Washington, DC, 256pp.
Rondonuwu, S. A. & Austin, A. D. (1988) A new species of Uracanthus (Coleoptera: Cerambycidae): a pest on orna-
mental cypresses in the Adelaide region. Transactions of the Royal Society of South Australia, 112, 109–117.
Sharp, D., & Muir, F. (1912) The comparative anatomy of male genitalia tube in Coleoptera. Transactions of the Entomo-
logical Society of London, 1912, 477–642.
Tepper, J. G. O. (1887) Common Native Insects of South Australia: A Popular Guide to South Australian Entomology,
Part I. Coleoptera, or Beetles. E. S. Wigg & Son, Adelaide, 46pp.
Thomson, J. (1860) Essai d’une Classification de la Famille des Cérambycides, et Matériaux pour Servir a une Monog-
raphie de cette Famille. 8vo. i–xvi, Liége, Paris, 396 pp.
Thomson, J. (1864) Systema Cerambycidarum ou exposé de tous les Genres Compris dans la Famille des Cérambycides
et Familles Limitrophes. 8vo. livr. 1–4, Liege, Paris, 578 pp.
Tillyard, R. J. (1926) Insects of Australia and New Zealand. Augus & Robertson, Sydney, 560 pp.
Wang, Q. (1993) A revision of Atesta Pascoe (Coleoptera: Cerambycidae, Cerambycinae: Phoracanthini) from Australia,
with descriptions of 18 new species. Invertebrate Taxonomy, 7, 961–1024.
Wang, Q. (1998) The phoracanthine beetles (Coleoptera: Cerambycidae): Taxonomic overview and generic relation-
ships. Invertebrate Taxonomy, 12, 667–684.
Wang, Q. & Lu, W. (2004) A systematic revison of New Zeand longicorn beetle genus Drototelus Broun (Coleoptera:
Cerambycidae). Invertebrate Systematics, 18, 649–659
White, A. (1855) Catalogue of Coleopterous Insects in the Collection of the British Museum, part 8 Longicornia 2, Brit-
ish Museum, London, pp. 175–412.

Zootaxa 1569 © 2007 Magnolia Press · 77
REVISION OF URACANTHUS
FIGURE 1. Uracanthus gigas, dorsal view: BL, body length from tip of mandibles to apex of elytra; SW, shoulder
width; PL, pronotal length; PW, pronotal width; EL, length of elytron along suture.

THONGPHAK & WANG
78 · Zootaxa 1569 © 2007 Magnolia Press
FIGURE 2. Head of Uracanthus. A: c , compound eye; a, antennae; an, antennal socket; pc, postcypleus; ac, ante-
clypeus; fr, frons; frs, frontoclypeal suture; aa, tentorial pit or anterior articulation; g, gena; l, labrum; md, mandible; mf,
median frontal groove; v, vertex. B: DUE, distance between upper lobes of eyes; DLE, distance between lower lobes of
eyes; HWBE, head width immediately below eyes; GL, genal length. C: DVE, distance between lobes of eyes on ventral
side. D: GL, genal length.

Zootaxa 1569 © 2007 Magnolia Press · 79
REVISION OF URACANTHUS
FIGURE 3. Antennae of Uracanthus: a, U. dubius; b, U. fuscus; c, U. cupressianus.

THONGPHAK & WANG
80 · Zootaxa 1569 © 2007 Magnolia Press
FIGURE 4. Antennal segments of Uracanthus: a, U. gigas, produced at apex; b, U. pertenuis male, fringed; c, U. gigas,
antennal segment 11 of male.

Zootaxa 1569 © 2007 Magnolia Press · 81
REVISION OF URACANTHUS
FIGURE 5. Pronotum of U. fuscocinereus, dorsal view: PL, pronotal length; AW, pronotal width at anterior margin; PW,
pronotal width at posterior margin.

THONGPHAK & WANG
82 · Zootaxa 1569 © 2007 Magnolia Press
FIGURE 6. Pronotum of U. insignis and U. albatus, dorsal view.

Zootaxa 1569 © 2007 Magnolia Press · 83
REVISION OF URACANTHUS
FIGURE 7. Surface of pronotal disc of Uracanthus: 1, U. tropicus; 2, U. parallelus; 3, U. froggatti; 4, U. loranthi; 5, U.
cupressianus; 6, U. maleficus; 7, U. longicornis; 8, U. glabrilineatus; 9, U. albatus; 10, U. pallens; 11, U. corrugicollis;
12, U. pertenuis; 13, U. suturalis; 14, U. fuscocinereus; 15, U. dubius; 16, U. ventralis; 17, U. regalis; 18, U. albatus; 19,
U. lateroalbus; 20, U. cryptophagus; 21, U. discicollis ; 22, U. insignis; 23, U. bivitta; 24, U. parvus; 25, U. fuscus.

THONGPHAK & WANG
84 · Zootaxa 1569 © 2007 Magnolia Press
FIGURE 8. Pronotal pubescence pattern of Uracanthus, dorsal view: a, U. pallens; b, U. quadristriolatus; c, U. bicolor-
atus; d, U. bivitta; e, U. strigosus; f, U. cryptophagus; g, U. maleficus; h, U. insignis; i, U. albatus; j, U. suturalis; k, U.
longicornis; l, U. lateroalbus; m, U. parallelus; n, U. pseudogigas; o, U. ater; p, U. ventralis.

Zootaxa 1569 © 2007 Magnolia Press · 85
REVISION OF URACANTHUS
FIGURE 9. Right elytra of Uracanthus, dorsal view: 1, U. bicoloratus; 2, U. maculatus; 3, U. perthensis; 4, U. griseus;
5, U. glabrilineatus; 6, U. triangularis; 7, U. pallens; 8, U. regalis; 9, U. dubius; 10, U. simulans; 11, U. fuscus; 12, U.
cryptophagus; 13, U. fuscocinerieus; 14, U. quadristriolatus; 15, U. insignis; 16, U. punctulatus; 17, U. maleficus; 18, U.
gigas; 19. U. pseudogigas; 20; U. acutus; 21, U. loranthi; 22, U. cupressianus; 23, U. minatus; 24, U. longicornis; 25, U.
albatus; 26, U. ventralis; 27, U. strigosus; 28, U. parallelus; 29, U. froggatti; 30, U. tropicus; 31, U. parvus; 32, U.
pertenuis; 33, U. bivitta; 34, U. discicollis; 35, U. corrugicollis; 36, U. lateroalbus; 37, U. bistriolatus; 38, U. suturalis;
39, U. ater.

THONGPHAK & WANG
86 · Zootaxa 1569 © 2007 Magnolia Press
FIGURE 10. Legs of Uracanthus: a, U. cryptophagus; b, U. gigas.

Zootaxa 1569 © 2007 Magnolia Press · 87
REVISION OF URACANTHUS
FIGURE 11. Visible sternites of male: a, U. gigas; b, U. albatus.

THONGPHAK & WANG
88 · Zootaxa 1569 © 2007 Magnolia Press
FIGURE 12. Aedeagus of Uracanthus, ventral view: a, complete structure: vl, ventral lobe; dl,dorsal lobe; ms, median
strut; sr, spined region; ur, unspined region; te, terminal region; b, ventral lobe pointed; c, ventral lobe rounded; d, ventral
lobe truncate; e, ventral lobe emarginate. Scale bars = 0.5 mm.

Zootaxa 1569 © 2007 Magnolia Press · 89
REVISION OF URACANTHUS
FIGURE 13. Forms of spines or processes on internal sac of aedeagus: a, simple spines; b, basally-forked spines; c,
multi-branched spines; d, scale-like processes.

THONGPHAK & WANG
90 · Zootaxa 1569 © 2007 Magnolia Press
FIGURE 14. Tegmens of Uracanthus, dorsal view: a, U. pertenuis; b, U. pallens; c, U. froggatti; d, U. gigas. Scale bars
= 0.5 mm.