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Butterflies and Moths (Insecta: Lepidoptera) of the Bulgarian part of Eastern Rhodopes

Authors:
  • National Museum of Natural History, Bulgarian Academy of Sciences
Insecta: Lepidoptera 525
Beron page 525
Beron P., Popov A. (eds). Biodiversity of Bulgaria. 2. Biodiversity of
Eastern Rhodopes (Bulgaria and Greece). Pensoft & Nat. Mus. Natur. Hist., Sofia
Butterflies and Moths (Insecta: Lepidoptera) of the
Bulgarian part of Eastern Rhodopes
Stoyan BESHKOV, Mario LANGOUROV
Beshkov S., Langourov M. 2004. Butterflies and Moths (Insecta: Lepidoptera) of the Bulgarian
part of Eastern Rhodopes. - In: Beron P., Popov A. (eds). Biodiversity of Bulgaria. 2. Biodiversity
of Eastern Rhodopes (Bulgaria and Greece). Pensoft & Nat. Mus. Natur. Hist., Sofia, 525-676.
Abstract. This article gives faunistical data for 1054 Lepidoptera species, of which 179 Microl-
epidoptera and 875 Macrolepidoptera, collected in 127 localities in the area explored. The
group Macrolepidoptera is relatively well explored. The Microlepidoptera group from the
Bulgarian part of the Eastern Rhodopes is however poorly known. For more than 15 years
data, which covers the biggest part of the Eastern Rhodopes, has been collected from January
to the end of December in different parts of the region and with different intensity over the
years. Butterf lies and skippers are collected with the classical methods. Moths are collected on
different kinds of lamps, light trap and sugaring. Clearwings moths (Sesiidae) are collected by
net and on pheromones as well. Some materials are collected using a Malaise trap. Several
problematic species are determined by taking into account the differences in the genitalia,
including the everted vesica, some of which are illustrated here. All literature sources and data
from faunistical records are also included here. As a result of our research about 670 species
are reported for the first time in the scientific literature as new for the Eastern Rhodopes. One
subspecies was described as new for the science, one species was found to be new for Europe,
seven species were new for Bulgaria, two species were new for the cave fauna of Bulgaria, one
more (Xanthorhoe oxybiata) is reported here as new for the cave fauna in general. One genus -
Stygoides Bruand, 1853 is new for the Bulgarian fauna. 13 species (Melasina christenseni, Stygoides
colchica, Depressaria badiella, Helcystogramma lineolella, Epinotia caprana, Epinotia nisella, Aethes
margarotana, Dioryctria simplicella, Trachycera advenella, Udea numeralis, Oulobophora externata,
Griposia wegneri and Hypena lividalis) are reported here as new for Bulgaria. Five more species
(Zebeeba falsalis, Dasypolia ferdinandi petrovi, Leucania herrichi, Leucania punctosa and Lycophotia
molothina) in Bulgaria are known from the Eastern Rhodopes only. Many species, which till
our investigation have been known only from SW Bulgaria and from singles localities in the
country, were found in the Eastern Rhodopes as well. The overall number of the Lepidoptera
species from the Bulgarian and the Greek parts of the Eastern Rhodopes published in this
volume 1186 species, 918 of which are Macrolepidoptera. From the group Macrolepidoptera
43 species are found only in the Greek part of the mountain and 554 Macrolepidoptera species
are found only in the Bulgarian part of the mountain. The number of the species common for
the Bulgarian and the Greek parts of the region is 321, as all of them are Macrolepidoptera. The
overall number of the Lepidoptera species (Macrolepidoptera and Microlepidoptera) found
only in the Bulgarian part of the mountain and published in this volume is 1143.
Key words: Lepidoptera, Macrolepidoptera, Microlepidoptera, Eastern Rhodopes, Bulgaria,
distribution, cave fauna, genitalia illustrations
526 S. BESHKOV, M. LANGOUROV
Beron page 526
Introduction
In Bulgaria the order Lepidoptera is one of the best-explored groups of invertebrates
although there are still some assumptions and errors about the knowledge of the group.
Concerning Macrolepidoptera, this is almost the only group of insects, which is so well
explored as are the vertebrates, although it is considerably larger. In Bulgaria about 2% are
endemic taxa as most of them are Balkan endemic on a subspecific level. One of the
reasons for the low number of endemic taxa is the well-elaborated taxonomy and faunis-
tics of the group. In Bulgaria the order Lepidoptera is extremely abundant, which is a
reason for its high diversity in terms of living forms. This requires different and special-
ized collecting methods in the field. In Bulgaria, respectively in the Eastern Rhodopes the
adult representatives of the group can be collected during the whole year and in all possible
terrestrial habitats.
Concerning the Macrolepidoptera group, from the approximately 1600 species, reg-
istered in the country, about 55% of the species, which constitute Bulgarian diversity, are
reported in this article as inhabiting the Bulgarian part of the Eastern Rhodopes. Com-
pared to the other parts of Bulgaria, at present the Eastern Rhodopes although not studied
in detail yet, can be pointed as one of the richest regions, in terms of species diversity. For
comparison here are presented some data for other parts of the country:
Kresna Gorge - 946
Western Rhodopes - 822
Vitosha Mountain – 888
Sinite Kamani Park near Sliven – 820
Western Stara Planina Mts. – 727
Belassitsa Mountain – 720
Zemen Gorge - 692
Volcanic Hill of Kozhuh with Rupite Place near Petrich – 692
Iskar Gorge - 550
Kastrakli Reserve in Western Rhodopes - 471
Upper Thracian Lowland – 440
In a more intensive research the number of the Macrolepidoptera species in the Eastern
Rhodopes is expected to increase to about 1000. The rate of the survey of the Macrolepi-
doptera group in the area is about 83%, but the different parts of the region are investigated
inadequately. At the same time, faunistically the Eastern Rhodopes are one of the most
interesting areas in Bulgaria and Europe. Many species, previously known in Bulgaria only
from the southwestern part of the country (e.g. Rethera komarovi, Agrochola osthelderi, Xanthor-
hoe oxybiata), were found in the Eastern Rhodopes as well. At present in Bulgaria two genera
Stygoides and Zebeeba and fifteen moth species – Melasina christenseni, Stygoides colchica, Depres-
saria badiella, Helcystogramma lineolella, Aethes margarotana, Dioryctria simplicella, Trachycera ad-
venella, Udea numeralis, Oulobophora externata, Hypena lividalis, Zebeeba falsalis, Dasypolia ferdi-
nandi petrovi, Leucania herrichi, Leucania punctosa and Lycophotia molothina are known from
the Eastern Rhodopes only. For the last one the Eastern Rhodopes is the only locality on the
Balkan Peninsula. For many other species it is the most distant point of the species range.
Insecta: Lepidoptera 527
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Concerning Microlepidoptera, the data from the Eastern Rhodopes are very poor. As
the present authors are not well familiar with this group, almost all the data are taken
from literature sources, quoted below. This is the reason for some records (TULESCHK-
OV & SLIVOV, 1975b) to be quoted exactly as they were initially published. The records
of Ganev for Pyralids however are accepted without any doubt, because he was a leading
specialist on this family. His original data however are from one spot only: Belite Brezi
Hut near Ardino. The present authors’ original data on Microlepidoptera concern mostly
caves and galleries and migrants.
Taking into account the present data for the Eastern Rhodopes, the original records
and the data from literature sources the region, which includes the Arda Valley together
with the dams Studen Kladenets and Ivaylovgrad from Studen Kladenets Dam, Sredna
Arda Railway Station from West to Ivaylovgrad Dam, Arda Hut near Dabovets Village to
the East, can be pointed as one with higher biodiversity. In this area, which is also the
best-studied one, the biodiversity is higher – about 430 species near Studen Kladenets
Village. At the second place in terms of biodiversity is the area around Madjarovo, togeth-
er with Momina Skala Hut – about 428 species. Ivaylovgrad Dam, Arda Hut near Dabovets
Village and Studen Kladenets Dam, Sredna Arda Railway Station follow with about 260
species registered. From Studen Kladenets Dam, Kroyatsi Hunting Farm near Nanovitsa
Village about 180 species are registered. About 150 species are known from Dishlik-Dere
Valley and the Iron Bridge on Arda River near Dolno Cherkovishte Village. The low
number of the species recorded is a result of the irregular research there. The same is the
situation with Studen Kladenets Dam, OSO Sport Center between Kaloyantsi and Gn-
yazdovo villages, where about 100 species only are known.
Another region with high biodiversity is the valley of Byala Reka River from Be-
lopolyane Village (=Byala Polyana) to Meden Buk Village and the valley of Luda Reka
River near Siv Kladenets Village. The best explored in that area is Siv Kladenets Village
with more than 270 recorded species. Second comes Meden Buk Village with more than
140 species. Next is Belopolyane Village with 130 species, all of them moths and recorded
in 3 collecting nights only – one in April, one in September and one in October. From
Dolno Lukovo Village about 90 species are known, from Mandritsa Village - 56 and from
Odrintsi Village - 40 species. The low number of the species recorded is also a result of the
irregular collecting there.
In both large areas pointed above, there is a significant overlap of the species compo-
sition and there are no vertical zones because of the minor differences in the altitude of the
different localities. Significant differences can be foundnd however in the higher western
parts of the Eastern Rhodopes. The data on them come from literature sources only (Belite
Brezi Hut near Ardino – 52 species), concerning only the tribe Eupitheciini and some
“rare” species, which were later found nowhere in the Eastern Rhodopes. These data can
not be used to make conclusions about the biodiversity. In the relatively high region near
Lebed Village, Djebel District material was collected two times only. From the 22 species
recorded there, 3 are found nowhere in the Eastern Rhodopes. From the higher parts, the
list here includes some data from the villages Egrek, Kremen (=Akcha) and Kirkovo as
well. The data from there concern only late-autumn fauna.
Faunistically, most interesting are the areas around Madjarovo, Studen Kladenets
Village, Siv Kladenets Village, Meden Buk Village, Belopolyane Village, the Iron Bridge on
528 S. BESHKOV, M. LANGOUROV
Beron page 528
Arda River near Dolno Cherkovishte Village, Egrek Village and Belite Brezi Hut. In
further research some new, rare and faunistically interesting species are expected to be
found near the villages Belopolyane and Egrek, near the Iron Bridge on Arda River near
Dolno Cherkovishte Village and in some other places. These areas are so interesting
because of the limestone rocks there. Unfortunately, they are still not well explored. Such
not well explored regions are all the other regions except for Arda Valley eastward from
Sredna Arda Railway Station, Byala Reka River valley from Meden Buk Village to its exit
from Bulgaria and Luda Reka River Valley near Siv Kladenets Village.
Here it is necessary to mention that the differences in the number of the species, the
differentiation of the richest of species and the faunistically most interesting area are
subjective, due to the different collecting methods used during the years and the different
numbers of visits. It is interesting to mention that such a common species as Cucullia
umbratica, Pyrria umbra, Polia nebulosa, Eremobia ochroleuca etc. have never been found in
the Eastern Rhodopes, although according to the present authors they should occur there.
From the rare species the same is the situation with Acronicta orientalis, Calophasia barthae,
Pyrrhia treitschkei, Agrochola wolfschlaegri, Agrochola rupicapra, Dichagyris melanura, Meganola
gigantula etc., which were expected, but have never been found there.
Literature sources
First records of the Lepidoptera fauna of the Eastern Rhodopes can be found in the
article of BURESCH (1912), where Zerynthia cerisy from Haskovo is mentioned. Next one
is the article of BURESCH & ILTSCHEW (1915), where Lemonia balcanica, Perisomena
caecigena and Aporophila australis are reported as ocurring “between Lyubimets and Kardja-
li”. BURESCH & ILTSCHEW (1921) reported Heliothis peltigera from Kardjali. Later on,
MARKOWITSCH (1923) reported Iphiclides podalirius and Nymphalis polychloros (Kardja-
li), Pieris brassicae (Haskovo, Kardjali, Krumovgrad), Pieris rapae (Haskovo, Kardjali, Kru-
movgrad, Momchilgrad), Pontia edusa, Leptidea sinapis, Polygonia c-album, Melitaea trivia,
Melitaea cinxia, Argynnis paphia, Hipparchia fagi or Hipparchia syriaca (as Satyrus hermione),
Lycaena phlaeas, Aricia agestis, Polyommatus belargus, Prodotis stolida, Lythria purpuraria (Kru-
movgrad), Limenitis camilla (Haskovo, Krumovgrad), Pyronia tithonus (Krumovgrad Dis-
tr.), Maculinaea arion (Eastern Rhodopes, without locality), Cerula vinula (Eastern Rhodopes
near Arda River), Scopula ornata (southern slopes of Eastern Rhodopes). There are men-
tions of some other species, all of them common ones, but without localities. Some of the
localities in that article at present are in Greece and are not included in the present article.
Next is the article of TSCHORBADJIEV (1936), who reported Sesamia cretica Led. from
the villages Novoselishte, Goren Chitak (=Gormo Voyvodino) and Maslinovo. Sesamia
cretica is reported also by POPOV (1939) from Ivaylovgrad and Haskovo. In the article of
TSCHORBADJIEW (1938 [1939]) Thaumetopoea solitaria is questionably reported from
Alenbeyer [=Bolyartsi] Village as a harmful species on Cоtinus coggigria, Fagus, Carpinus,
Quercus and other trees. Its correct identification as Thaumetopoea solitaria is confirmed by
BURESCH (1940: 247). In his work the caterpillars are reported from Haskovo on Cot-
tinus coggigria. In the catalogues of BURESCH & TULESCHKOW (1929-1947) there
was reference of all published data for the Eastern Rhodopes at that time (only these of
Insecta: Lepidoptera 529
Beron page 529
MARKOWITSCH quoted above) and new data, originating from the collections of the
Royal Entomological Station, were included.
POPOV (1952: 68-69) reported a large-scale flight of Pyrameis cardui (about 3 km long
wall) on 03.4.1952 in the area around Madrets Village and Harmanli, coming from northeast.
Some data for the region, mainly from Haskovo and Harmanli, can be found in the
work of TULESKOV (1965). The data from Harmanli are mainly from the Harmanliys-
ka Reka valley, which indeed belongs to the Eastern Rhodopes. Other records but only for
the groups Bombyces and Sphinges, most of them original are present in the catalogue of
GANEV (1984d). There are reports on the following species: Lemonia balcanica, Perisomena
caecigena, Trichiura crataegi (Kardjali), Phalera bucephala, Stauropus fagi (last one probably
wrongly instead Cerula vinula), Spatalia argentina, Notodonta dromedarius, Drymonia melag-
ona, Drymonia velitaris, Clostera curtula, Macrothylacia rubi, Dendrolimus pini, Phillodesma
tremulifolia, Agrius convolvuli, Hyloicus pinastri, Dilina tiliae, Laothoe populi, Proserpinus pros-
erpina, Deilephila porcellus, Lamellocossus terebra, Apoda limacodes, Drepana binaria, Drepana
cultraria, Cilix glaucata, Habrosyne pyritoides, Ocneria rubea, Miltochrista miniata, Eilema soror-
cula, Lithosia quadra, Spilosoma lubricipeda, Diaphora luctuosa (Belite Brezi Hut near Ardino)
and Phragmatobia caesarea (Harmanli).
Data for the tribe Eupitheciini can be found in the catalogue of GANEV (1985a),
where from the Eastern Rhodopes the following species are published: Eupithecia distinctar-
ia (Belite Brezi Hut near Ardino and Zlatograd), Eupithecia vulgata, Eupithecia subfuscata,
Eupithecia icterata, Eupithecia orphanata, Chloroclystis chloerata and Chloroclystis rectangulata
(Belite Brezi Hut near Ardino).
GANEV (1988a) reported Hipparchia syriaca from Kardjali. Another article, containing
some data from the Eastern Rhodopes is that of TULESCHKOV & SLIVOV (1975a). In the
article, dedicated entirely to the whole Rhodopes, the following species are reported from the
Eastern Rhodopes: Idaea filicata (The border post near Avren Village), Rhodometra sacraria
(The border post near Avren Village and Kardjali), Horisme corticata (The border post near
Avren Village and Momchilgrad), Scopula immorata (The border post near Avren Village and
Krumovgrad), Phytometra viridaria (The border post near Avren Village, Dabovets Village,
Ivaylovgrad), Eucrostes indegenata (The border post near Avren Village and Ivaylovgrad),
Idaea dimidiata (reported as didimata), Idaea seriata, Idaea inquinata, Selidosema plumaria, Yi-
goga forcipula, Hadena compta, Mythimna turca, Catocala sponsa, Cucullia santonici (Kardjali),
Scoliopteryx libatrix (Kardjali, Krumovgrad), Idaea ostrinaria (Kardjali, Popsko and Mandritsa
villages), Euphyia putridaria, Epione repandaria (Kardjali, Dabovets Village), Macaria aestimaria
(Kardjali, Podrumche Village), Macaria alternaria (Kardjali, Plevun Village), Paracolax tristalis
(Kardjali, Chorbadjiysko Village), Copiphana olivina, Acronicta megacephala (Kardjali, Ivay-
lovgrad), Orthonama obstipata (Kardjali, Svirachi Village), Melipotis suava (Kardjali, Ivay-
lovgrad, Mandritsa Village), Cyclophora albiocelaria, Perconia strigillaria, Hypena munitalis (Kardjali,
Mandritsa Village), Aplasta ononaria (Momchilgrad, Dabovets Village and Mandritsa Vil-
lage), Cyclophora punctaria, Diarsia rubi, Acronicta leporina, Ipimorpha subtusa, Heliothis armigera
(Krumovgrad), Idaea pallidata (Chorbadjiysko Village), Xanthorhoe designata (Podrumche Vil-
lage), Hemithea aestivaria, Ligdia adustata, Omphalophana antirrhini (Ivaylovgrad), Glossotrophia
confinaria (Ivaylovgrad, Popsko Village), Boarmia viertlii (Dabovets Village), Lacanobia suasa,
Calophasia opalina (Plevun Village), Idaea moniliata, Opisthograptis luteolata, Catocala elocata,
Laspeyria flexula (Mandritsa Village).
530 S. BESHKOV, M. LANGOUROV
Beron page 530
The next article of the same authors (TULESCHKOV & SLIVOV, 1975b) is dealing
with Microlepidoptera, also in the entire Rhodopes. There the following Sesiidae species
are reported from the Eastern Rhodopes: Synanthedon vespiformis, Chamaesphecia bibionifor-
mis (The border post near Avren Village), Sesia apiformis (The border post near Avren
Village, Mandritsa Village), Synanthedon myopaeformis (as myopiformis) (Chorbadjiysko
Village), Bembecia uroceriformis (Kardjali, Chorbadjiysko Village), Chamaesphecia annellata
(Kardjali, Mandritsa Village), Chamaesphecia astatiformis (Ivaylovgrad). In the same article
there is much data concerning some families of Microlepidoptera. All of them are quoted
and included in the present list below.
In SLIVOV (1979) two species only are reported: Chersotis rectangula (Belite Brezi Hut
near Ardino and Studen Kladenets) and Mesoligia literosa (Studen Kladenets). Another
article with data, concerning Lepidoptera in the area studied, is that of SLIVOV (1972),
where Eucrostes indigenata (Ivaylovgrad), Semiothisa aestimaria (Podrumiche Village) and
Oria musculosa (Ivaylovgrad) are reported. SLIVOV (1973) reported a single species: Hippar-
chia syriaca from Mandritsa Village. In GOGOV & LOUKOV (1964) Erannis declinans is
reported from Kardjali. In SLIVOV & LUKOV (1976 [1977]) data for two species only can
be found, one of them (Boarmia viertlii from Dabovets Village) already published from the
same locality. SLIVOV (1984) reported Janthinea frivaldszkyi from Ivaylovgrad.
GANEV (1980) reported Lamelocossus terebrus, Boarmia maeoticaria, Ochrostigma velital-
is, Trisatelis emortualis and Ocneria ruibea, all from Belite Brezi Hut near Ardino. Later on,
GANEV (1982a) reported the following species, again from Belite Brezi Hut: Scopula flo-
slactata, Scotopteryx mucronata, Chesias rufata, Plagodis dolabraria, Ennomos quercinaria, Deu-
teronomos fuscantaria, Fagivorina arenaria, Aethalura punctulata, Bapta bimaculata, Gnophos
furvata, Ochrostigma melagona, Diaphora luctuosa, Cucullia lactucae, Euchalcia modesta, Proserpi-
nus proserpina. In GANEV (1984a) and GANEV (1987) Eupithecia orphnata is reported
again from Belite Brezi Hut near Ardino and in addition Photedes captiuncula is reported
from the same locality. GANEV (1982b, 1983, 1984c, 1985c, 1986a, 1986b) reported
several Pyraloidea species from the Eastern Rhodopes, mostly from Belite Brezi Hut near
Ardino. In the article of NESTOROVA (1980) Chesias rufata and Eupithecia abbreviata are
reported from Shadiytsa Place northweste of Zlatograd. In another article of NESTORO-
VA (1984) Perizoma affinitata is reported from the same place.
One butterfly species - Pontia chloridice is reported by ABADJIEV (1992) from Studen
Kladenets Dam, Sredna Arda Railway Station and from Studen Kladenets Village. ABAD-
JIEV (1993) reported Kirinia roxelana (Arda Valley), Pieris krueperi, Pieris rapae, Pieris mannii
and Pontia chloridice (Studen Kladenets Dam: Sredna Arda Railway Station). ABADJIEV
(1995) reported Polygonia egea (Oreshari Village), Euphydryas aurinia and Hipparchia syriaca
(Ivaylovgrad Dam: Arda Hut near Dabovets Village), Hipparchia syriaca and Melitaea didyma
meridionalis (Studen Kladenets Dam: Sredna Arda Railway Station), Hipparchia fatua (Studen
Kladenets Village) and Limenitis reducta (Byala Reka Valley: Dolno Lukovo Village). Another
three articles containing data from the Eastern Rhodopes are: ABADJIEV (1993): (Kirinia
roxelana and Quercusia quercus from Momina Skala Hut near Madjarovo), ABADJIEV (1994а):
(Pontia chloridice from Studen Kladenets Dam: Sredna Arda Railway Station) and ABADJIEV
(1994b): (Pontia chloridice from Studen Kladenets Dam: Sredna Arda Railway Station).
GANEV & BESCHKOV (1987) reported Pyrgus sidae, Satyrium w-album and Sphinx
ligustri from Studen Kladenets Dam [Kroyatsi Hunting Farm near Nanovitsa Village].
Insecta: Lepidoptera 531
Beron page 531
BESHKOV (1992) reported Colias erate f. chrysodona, Pieris pseudorapae balcana, Tarucus
balcanicus (Garvanovo Village), Phalera bucephaloides, Cryphia ochsi (Ayda Hut near Hasko-
vo), Orthosia rorida, Craniophora pontica (Studen Kladenets Village), Cryphia seladona burgeffi
(Sussam Village near Haskovo). BESHKOV (1994) reported Colias crocea and Pieris rapae
(Studen Kladenets Dam: Sredna Arda Railway Station and Momina Skala Hut near Mad-
jarovo), Macroglossum stellatarum (Studen Kladenets Dam: Sredna Arda Railway Station
and Ivaylovgrad Dam: Arda Hut), Issoria lathonia (Studen Kladenets Dam: Sredna Arda
Railway Station, Momina Skala Hut and Siv Kladenets Village), Lycaena phlaeas (Studen
Kladenets Dam: Sredna Arda Railway Station; Studen Kladenets Village; Momina Skala
Hut), Prodotis stolida (Studen Kladenets Dam: Sredna Arda Railway Station and Luda
Reka River Valley: Siv Kladenets Village), Autographa gamma (Studen Kladenets Dam:
Sredna Arda Railway Station; Ivaylovgrad Dam: Arda Hut; Siv Kladenets Village), Nymph-
alis polychloros (Studen Kladenets Dam: Kroyatsi Hunting Farm near Nanovitsa Village),
Aporia crataegi, Pieris brassicae (Madjarovo and Siv Kladenets Village), Pontia edusa (Momi-
na Skala Hut and Siv Kladenets Village), Pontia chloridice and Nycteola asiatica (Siv Klad-
enets Village), Libythea celtis (Madjarovo), Cynthia cardui and Hyles euphorbiae (Momina
Skala Hut near Madjarovo), Tyta luctuosa (Studen Kladenets Village, Momina Skala Hut;
Ivaylovgrad Dam, Arda Hut near Dabovets Village and Siv Kladenets Village, Ivaylovgrad
District), Acontia lucida (Ivaylovgrad Dam: Arda Hut; Siv Kladenets Village), Macdun-
noughia confusa and Spodoptera exigua (Ivaylovgrad Dam: Arda Hut), Hoplodrina blanda
(Studen Kladenets Dam: Studen Kladenets Village; Arda Hut near Dabovets Village), My-
thimna albipuncta (Studen Kladenets Village; Ivaylovgrad Dam: Arda Hut near Dabovets
Village), Mythimna vitellina and Agrotis ipsilon (Studen Kladenets Dam: Sredna Arda Rail-
way Station), Mythimna l-album (Ivaylovgrad Dam: Arda Hut; Momina Skala Hut and Siv
Kladenets Village), Acantholeucania lorey, Noctua pronuba and Agrotis exclamationis (Ivay-
lovgrad Dam: Arda Hut; Siv Kladenets Village).
BESHKOV (1995) reported Pieris krueperi (Studen Kladenets Dam: Sredna Arda Rail-
way Station; Studen Kladenets Village), Pontia chloridice (Studen Kladenets Dam: Shiroko
Pole Village, Sredna Arda Railway Station; Studen Kladenets Village; Siv Kladenets Vil-
lage), Pieris ergane, Hypena palpalis (Studen Kladenets Village and Madjarovo), Heodes otto-
mana (Siv Kladenets Village and Ivaylovgrad Dam: Arda Hut near Dabovets Village),
Brenthis hecate (Studen Kladenets Dam: Darets, Sredna Arda Railway Station; Studen Klad-
enets Village), Trichiura verenae (Momchilgrad; Svezhest Hut near Krumovgrad; between
Trakiets and Konush villages; Studen Kladenets Village), Saturnia spini (Studen Kladenets
Dam: Kroyatsi Hunting Farm near Nanovitsa Village, Sredna Arda Railway Station; Studen
Kladenets Village), Perisomena caecigena (Svezhest Hut near Krumovgrad, Studen Kladenets
Dam: Sredna Arda Railway Station; Studen Kladenets Village; Momina Skala Hut; Ivay-
lovgrad Dam: Arda Hut; Siv Kladenets Village), Orthistixis cribraria (Studen Kladenets
Dam: between Darets and Studen Kladenets Village; Siv Kladenets Village), Еumera regina
(Studen Kladenets Dam: Kroyatsi Hunting Farm near Nanovitsa Village; Studen Klad-
enets Village; Momina Skala Hut), Dolbina elegans and Rethera komarovi (Studen Kladenets
Village, Mandritsa Village), Thaumetopoea solitaria (Studen Kladenets Dam: Kroyatsi Hunting
Farm near Nanovitsa Village; Momina Skala Hut; Ivaylovgrad Dam: Arda Hut), Ocneria
lederi (Studen Kladenets Dam: Sredna Arda Railway Station; Momina Skala Hut; Siv
Kladenets Village), Parocneria terebinthi (Studen Kladenets Dam: Sredna Arda Railway Sta-
532 S. BESHKOV, M. LANGOUROV
Beron page 532
tion; Momina Skala Hut), Nycteola siculana (Studen Kladenets Village), Cryphia seladona
(Studen Kladenets Dam: Kroyatsi Hunting Farm near Nanovitsa Village, Sredna Arda
Railway Station; Momina Skala Hut), Nychiodes amigdalaria, Zebeeba falsalis, Agrochola dele-
ta (Siv Kladenets Village), Cucullia celsiae, Copiphana olivina (Ivaylovgrad Dam: Arda Hut),
Amephana dalmatica (Momina Skala Hut near Madjarovo; Siv Kladenets Village), Pyrois
effusa (Madjarovo), Orectis proboscidata, Amphipyra tetra (Studen Kladenets Village; Momi-
na Skala Hut near Madjarovo), Polyphaenes subsericata (Djebel District: Lebed Village; Studen
Kladenets Dam: Kroyatsi Hunting Farm near Nanovitsa Village; Studen Kladenets Vil-
lage; Momina Skala Hut), Xanthia cypreago (Momchilgrad; Svezhest Hut near Krumovgrad;
between Konush and Trakiets villages; between Kaloyantsi and Gnyazdovo villages; Studen
Kladenets Village; Momina Skala Hut near Madjarovo), Agrochola osthelderi (Studen Klad-
enets Village; Momina Skala Hut), Agrochola gratiosa (Svezhest Hut near Krumovgrad;
Momina Skala Hut), Conistra ragusae (Studen Kladenets Dam: Kroyatsi Hunting Farm
near Nanovitsa Village), Xylena lunifera (Svezhest Hut near Krumovgrad), Dryobotodes mono-
chroma, Leucania herrichi (Momina Skala Hut), Spudaea pontica, Dryobotodes tenebrosa (Studen
Kladenets Dam: Sredna Arda Railway Station), Polymixis trisignata (between Kaloyantsi
and Gnyazdovo villages; Siv Kladenets Village), Gortyna moesiaca (Momchilgrad; Momi-
na Skala Hut; Studen Kladenets Village; Sredna Arda Railway Station), Hadula mendax
(Momina Skala Hut; Siv Kladenets Village; Ivaylovgrad Dam: Arda Hut), Orhosia (Di-
oszeghyana) schmidtii (“Kenana” near Haskovo; Studen Kladenets Dam: Kroyatsi Hunting
Farm near Nanovitsa Village; Studen Kladenets Village).
Lycaena ottomana and Euphydryas aurinia from Arda Hut near Davovets Village are
reported as well as in ABADJIEV (2003). The site Arda Hut is recognised there as one of
the prime butterfly areas in Bulgaria.
BESHKOV (1996a) reported Yigoga flavina from Siv Kladenets Village and Ivay-
lovgrad Dam: Arda Hut and from Dabovets Village. In BESHKOV (1993) and BESHK-
OV (1998) a large number of species for the Eastern Rhodopes are reported, but only one
species has a locality: Cryphia seladona burgeffi (Sussam Village). Other data for the Eastern
Rhodopes are present in BESHKOV & GOATER (2000): Triodia amasinus dobrogensis,
Spodoptera exigua, Luperina dumerilii (Studen Kladenets Village, Momina Skala Hut, Meden
Buk Village); Triodia silvina, Crocallis elinguaria, Peribatodes umbraria, Scopula ochraceata,
Eilema caniola, Dysauxes famula pontica, Hoplodrina ambigua, Thalpophila matura, Episema
korsakovi, Mythimna vitellina, Noctua pronuba, Noctua comes (Studen Kladenets Village,
Momina Skala Hut, Meden Buk Village, Belopolyane Village); Iphiclides posdalirius (Hask-
ovo); Pieris rapae, Pontia edusa, Issoria lathonia, Cilix glaucata, Ennomos fuscantaria, Poly-
pogon plumigeralis, Euxoa cos, Agrotis ipsilon (Meden Buk Village); Colias crocea, Polymixis
serpentina, Tholera decimalis (Momina Skala Hut, Meden Buk Village); Pandoriana pandora,
Cymatophorima diluta, Idaea camparia, Eupithecia ericeata, Sphrageidus similis, Orectis probosci-
data, Pseudoips prasinanus, Episema glaucina, Dryobotodes monochroma, Chersotis rectangula,
Euxoa obelisca (Momina Skala Hut); Hipparchia syriaca, Drepana binaria (Studen Kladenets
Village, Momina Skala Hut, Meden Buk Village); Hipparchia fatua, Cryphia amasina, Mor-
mo maura, Agrotis crassa (Studen Kladenets Village, Meden Buk Village); Lycaena phlaeas,
Polyommatus icarus, Eucrostes indigenata, Glossotrophia confinaria, Gnophos sartatus, Scopula
imitaria, Nycteola asiatica, Cryphia ochsi, Cryphia seladona burgeffi, Mythimna l-album (Studen
Kladenets Village); Cyclophora puppillaria, Idaea degeneraria, Mesogona acetosellae, Mythimna
Insecta: Lepidoptera 533
Beron page 533
albipuncta, Xestia xanthographa, Agrotis trux (Studen Kladenets Village, Momina Skala Hut,
Belopolyane Village); Timandra griseata, Idaea dimidiata, Camptogramma bilineata, Nebula
ablutaria, Eupithecia orphnata, Artiora evonymaria, Crocallis tusciaria, Pterostoma palpinum,
Eilema sororcula, Phragmatobia fuliginosa, Schrankia costaestrigalis, Nycteola siculana, Acontia
melanura, Euchalcia consona, Trichoplusia ni, Abrostola agnorista, Amphipyra livida, Atethmia
centrago, Phlogophora meticulosa, Meganephria bimaculosa, Dryobotodes carbonis, Gortyna flava-
go, Lacanobia oleracea, Mythimna ferrago, Leucania punctosa, Agrotis puta, Agrotis segetum (Be-
lopolyane Village); Atethmia ambusta (Ivaylovgrad Dam: Borislavtsi Village); Scopula mar-
ginepunctata, Aspilates ochrearia, Gymnosceles rufifasciata, Thaumetopoea solitaria, Prodotis stol-
ida, Autographa gamma, Episema tersa, Hadula trifolii (Studen Kladenets Village, Meden Buk
Village, Belopolyane Village); Rhodometra sacraria, Xestia cohaesa (Momina Skala Hut, Meden
Buk Village, Belopolyane Village); Xanthorhoe fluctuata, Lemonia balcanica, Praestilbia arme-
niaca (Momina Skala Hut, Belopolyane Village); Eupithecia centaureata, Hyles euphorbiae,
Macroglossum stellatarum, Catocala elocata, Lygephila craccae, Acontia lucida, Amphipyra tragop-
ogonis, Amphipyra tetra, Helicoverpa armigera, Episema lederi, Aporophila canescens, Mesapamea
secalis (Studen Kladenets Village, Belopolyane Village); Eumera regina, Nychiodes waltheri,
Selidosema plumarium, Eilema pseudocomplana, Parascotia fuliginaria, Catocala nupta (Meden
Buk Village, Belopolyane Village); Peribatodes rhomboidaria, Lasiocampa trifolii (Studen Klad-
enets Village, Momina Skala Hut, Meden Buk Village, Belopolyane Village, Borislavtsi
Village); Campaea margaritata, Еuplagia quadripunctaria, Bena bicolorana, Paradrina flavirena,
Polyphaenis subsericata, Agrochola ostheldei, Leucania herrichi, Lampra fimbriata, Epilecta lino-
grisea (Studen Kladenets Village, Momina Skala Hut); Dyscia sicanaria (Studen Kladenets
Village, Momina Skala Hut, Belopolyane Village, Ivaylovgrad Dam: Barrage); Agrius con-
volvuli (Madjarovo). In the same article data for 27 Pyralidae species from the same local-
ities is given and all of them are included in the species list below. Almost the same data,
with minor omissions, are present in the article of GOATER (1996). BESHKOV (1999)
reported one species, which is new for Europe (Egira tibori), from Studen Kladenets Village
and from “Kroyatsi” Hunting farm near Nanovitsa Village. In the same article Egira
conspicillaris is reported from Arda Hut near Dabovets Village.
BESHKOV & VASSILEV (1996) reported Pieris brassicae (Studen Kladenets Dam:
Zvezdelina Village, Sredna Arda Railway Station; between Kulich Village and the Iron
Bridge on Arda River near Dolno Cherkovishte Village; Siv Kladenets Village); Pontia
edusa, Colias crocea (Zvezdelina Village, Krumovgrad, between Kulich Village and the Iron
Bridge on Arda River near Dolno Cherkovishte Village, Siv Kladenets Village); Issoria
lathonia (Zvezdelina Village, between Kulich Village and the Iron Bridge on Arda River
near Dolno Cherkovishte Village, Arda Hut near Madjarovo, Siv Kladenets Village); Pon-
tia chloridice, Hipparchia semele, Prodotis stolida, Pseudaletia unipuncta (Sredna Arda Railway
Station); Lycaena phlaeas (Sredna Arda Railway Station, between Kulich Village and the
Iron Bridge on Arda River near Dolno Cherkovishte Village); Autographa gamma (Sredna
Arda Railway Station, Studen Kladenets Village, Momina Skala Hut, Siv Kladenets Vil-
lage); Rhodometra sacraria, Mythimna albipuncta (Studen Kladenets Village); Mythimna vitell-
ina (Studen Kladenets Village, Siv Kladenets Village); Syntarucus pirithous (Krumovgrad),
Vanessa atalanta, Mythimna l-album (Sredna Arda Railway Station, Siv Kladenets Village);
Amphipyra pyramidea, Spodoptera exigua, Noctua pronuba, Peridroma saucia (Sredna Arda
Railway Station, Studen Kladenets Village, Momina Skala Hut); Helicoverpa armigera (Sred-
534 S. BESHKOV, M. LANGOUROV
Beron page 534
na Arda Railway Station, Studen Kladenets Village, between Kulich Village and the Iron
Bridge on Arda River near Dolno Cherkovishte Village, Momina Skala Hut); Agrotis
ipsilon (Sredna Arda Railway Station, Studen Kladenets Village, Momina Skala Hut, Siv
Kladenets Village); Agrotis segetum (Sredna Arda Railway Station, Momina Skala Hut,
Siv Kladenets Village); Papilio machaon (between Kulich Village and the Iron Bridge on
Arda River near Dolno Cherkovishte Village); Orthonama obstipata, Acherontia atropos,
Catocala sponsa, Chrysodeixis chalcites, Heliothis peltigera (Momina Skala Hut); Nymphalis
polychloros, Polygonia c-album (Arda Hut near Dabovets Village, Siv Kladenets Village);
Aporia crataegi, Gonepteryx rhamni, Nymphalis antiopa, Inachis io, Aglais urticae, Macroglos-
sum stellatarum, Tyta luctuosa, Paradrina clavipalpis, Hoplodrina blanda, Xestia c-nigrum
Agrotis exclamationis (Siv Kladenets Village). Some data on late autumn butterflies can be
found in the article of ABADJIEV, TSHIKOLOVETS & BESHKOV (1999). They are
all included in the species list below.
Other data for the Macrolepidoptera fauna of the Eastern Rhodopes can be found in
the article of BESHKOV (1996b), who reported: Iphiclides podalirius (Haskovo); Nymphalis
polychloros, Paradrina clavipalpis, Acantholeucania loreyi (Studen Kladenets Dam: Sredna Arda
Railway Station); Aglius convolvuli (Madjarovo); Lycaena phlaeas (Studen Kladnets Village,
Ivaylovgrad); Nycteola asiatica, Mythimna l-album (Studen Kladenets Village); Pandoriana
pandora, Chrysodeixis chalcites (Momina Skala Hut); Pieris rapae (Perperek Village, Kaloyan-
tsi Village, Ivaylovgrad, Zhaltichalskoto Dere Valley near Meden Buk Village); Pontia
edusa (Perperek Village, Zhaltichalskoto Dere Valley near Meden Buk Village); Colias crocea
(Perperek Village, Kaloyantsi Village, Sredna Arda Railway Station, Momina Skala Hut,
Ivaylovgrad, Zhaltichalskoto Dere Valley near Meden Buk Village); Issoria lathonia (Sred-
na Arda Railway Station, Zhaltichalskoto Dere Valley near Meden Buk Village); Helicover-
pa armigera (Sredna Arda Railway Station, Studen Kladenets Village, Belopolyane Village);
Spodoptera exigua (Sredna Arda Railway Station, Studen Kladenets Village, Momina Skala
Hut, Zhaltichalskoto Dere Valley near Meden Buk Village); Agrotis segetum (Sredna Arda
Railway Station, Belopolyane Village); Agrotis ipsilon (Sredna Arda Railway Station, Mo-
mina Skala Hut, Zhaltichalskoto Dere Valley near Meden Buk Village); Cyclophora pup-
pilaria, Mythimna albipuncta (Studen Kladenets Village, Momina Skala Hut, Belopolyane
Village); Prodotis stolida (Studen Kladenets Village, Zhaltichalskoto Dere Valley near Meden
Buk Village, Belopolyane Village); Macroglossum stelatarum, Hyles euphorbiae, Acontia lucida
(Studen Kladenets Village, Belopolyane Village); Lampra fimbriata, Euplagia quadripunctar-
ia (Studen Kladenets Village, Momina Skala Hut near Madjarovo); Autographa gamma
(Studen Kladenets Village, Momina Skala Hut, Ivaylovgrad, Belopolyane Village, Zhal-
tichalskoto Dere Valley near Meden Buk Village, Siv Kladenets Village); Mythimna vitelli-
na, Noctua pronuba (Studen Kladenets Village, Momina Skala Hut, Belopolyane Village,
Zhaltichalskoto Dere Valley near Meden Buk Village); Rhodometra sacraria (Momina Skala
Hut, Belopolyane Village, Zhaltichalskoto Dere Valley near Meden Buk Village); Trichop-
lusia ni, Amphipyra pyramidea, Phlogophora meticulosa (Belopolyane Village); Pseudaletia uni-
puncta (Siv Kladenets Village).
Slivov (pers. comm.) collected Theretra alecto cretica in the districts of Kardjali.
Data for some butterflies species from Komuniga Village are published by KRZY-
WICKI (1981). Certain Geometridae species from the Eastern Rhodopes are present in the
catalogue of NESTOROVA (1998). Many data for the superfamilies Papilionoidea and
Insecta: Lepidoptera 535
Beron page 535
Hesperioidea can be found referred to and mapped on a UTM 10х10 кm grid in ABAD-
JIEV (2001). A large amount of data for the family Noctuidae, published in the literature
for the region or original unpublished data, can be found in the monograph of BESHK-
OV (2000). For some Microlepidoptera species original or quoted data are present in the
articles of GANEV (1985b, 1985c, 1986a, 1986b, 1988b). In his articles the original data
come from Belite Brezi Hut near Ardino, an area from which the present authors do not
have their own data. One noctuid species (Polymixis trisignata) from Siv Kladenets Village
is illustrated in RONKAY, YELA & HREBLAY (2001).
Material collected
The data presented here results mostly from the authors’ investigations in the period
1984-2004. During 1984-1988 material was collected occasionally. Detailed study began
in the autumn of 1989. Material was collected each month, including December and
January, sometime several times monthly as usually several regions of the area were ex-
plored. Diurnal Lepidoptera species were collected using the classical methods. Nocturnal
moths were attracted at 160, 250 or 400 W MVL, using electricy of buildings as an
electrical source (chalets and single buildings with electricity). For a short period a1500
W Tg lamp was used, as well as a 400 UV lamp. After 1995 a generator was used as a
source of electricity. In the last 10 years the lamps were powered by a generator, as usually
a 160 W MVL together with a 25 W or an 18 W “black” lamp were used. In the last 9
years a standard light trap powered by an accumulator was also used. Last several years a
sugar-wine mixture 1:1 was used as well, mostly in early spring and late autumn. Only
during 1996-1997 clearwings moths (Sesiidae) were collected at different kinds of species
specific and combined pheromones. Aestivated and hibernated Lepidoptera species were
collected in several caves and galleries. Some materials from Madjarovo and its districts
were collected by the second author, who had used a Malaise trap.
Some problematic species are determined taking into account the differences in the
genitalia, including the everted vesica.
We received a large amount of data from the data-base of Mr Peter Davey. All these
data are incorporated here. Because we have not seen the specimens observed and recorded
by him, we accepted all his records without any doubt. We have commented here only on
some species, which occurence in the region is doubtful and the data provided might be
the result of a missidentification.
Before our studies, from the territory of the Eastern Rhodopes very few species were
known from single localities only. All literature sources and data from the faunistic records
are included here. As a result of our research about 670 species are new for the Eastern
Rhodopes, one subspecies (Dasypolia ferdinandi petrovi Beshkov, 2000) was new for the
science, one species (Egira tibori Hreblay, 1994) was new for Europe, five other species
(Zebeeba falsalis, Orthosia schmidti, Leucania herrichi, Leucania punctosa, Lycophotia molothina)
were new for Bulgaria, twelve more species (Melasina christenseni, Stygoides colchica, Depressar-
ia badiella, Helcystogramma lineolella, Aethes margarotana, Dioryctria simplicella, Trachycera ad-
venella, Udea numeralis, Oulobophora externata and Hypena lividalis) and one genus (Stygoides)
are reported for the first time from Bulgaria, two species were new for the cave fauna of
536 S. BESHKOV, M. LANGOUROV
Beron page 536
Bulgaria, one more (Xanthorhoe oxybiata) is reported here as new for the cavian fauna in
general. Many species, which until our study had been known only from SW Bulgaria
and from single localities in the country, were found in the Eastern Rhodopes. As a result
of our investigations it turned out that the Eastern Rhodopes are one of the faunistically
richest and one of the most interesting regions in Bulgaria, showing extremely high diver-
sity of butterflies and moths species.
In this article from the Bulgarian part of the Eastern Rhodopes 1054 Lepidoptera
species from 127 localities are known, distributed in families as follows:
Psychidae 10
Limacodidae 1
Zygaenidae 9
Sesiidae 15
Cossidae 7
Thyrididae 1
Hepialidae 2
Lasiocampidae 15
Saturniidae 4
Lemoniidae 1
Sphingidae 18
Hesperiidae 17
Papilionidae 5
Pieridae 18
Lycaenidae 35
Nymphalidae 53
Drepanidae 9
Geometridae 207
Notodontidae 29
Noctuidae 359
Pantheidae 1
Lymantriidae 12
Nolidae 15
Arctiidae 25
Total Macrolepidoptera 875
Adelidae 2
Tineidae 8
Yponeomeutidae 2
Ypsolophidae 6
Plutellidae 1
Acrolep i i d ae 1
Glyphipterygidae 1
Ethmiidae 3
Depressariidae 3
Oecophoridae 5
Insecta: Lepidoptera 537
Beron page 537
Lecithoceridae 1
Momphidae 1
Cosmopterigidae 1
Gelechiidae 9
To rt r i ci d ae 3 7
Alucitidae 2
Pterophoridae 4
Carposinidae 1
Pyralidae 89
Total Microlepidoptera 179
The number of the Macrolepidoptera species from the Eastern Rhodopes is expected
to increase to 1000 or more. This means that in the Eastern Rhodopes probably about
61% of the Bulgarian Macrolepidoptera species occur.
The most interesting areas in the Eastern Rhodopes are the Arda Valley and the
basin of Byala Reka and Luda Reka Rivers. Several species, which have disappeared (Pontia
chloridice for example) or which have low density in the other parts of the country, have
very stabile populations in these regions. Only one legally protected species in Bulgaria
(Perisomena caecigena Kupido) occurs in the Eastern Rhodopes. It is a very common species
and in most of the localities it is extremely abundant. All species included in the appendixes
of the international conventions are mentioned in the text. Many other records on phenology,
density, ecology and other features, which were collected during the investigation, are not
included here. Such data are given only for the most interesting species.
Further trends
Further field research needs to explor more in detail the area west of Varbitsa river,
the area between Arda River and the Thracian lowland, as well as the region between Arda
River and Greece. As a whole, the data available from May to August are not sufficient
enough. At the beginning of the investigation, because of the lack of own source of electricy,
some important habitats were not sufficiently studied. Collecting in such habitats (limestone
areas, habitats with Phyllirea etc.) is urgently needed. Collecting with light traps and sugaring
in all seasons and all habitats is also urgently needed in further research on nocturnal
species, as well as using pheromones for collecting Sesiidae. In Further trends it is necessary
to pay attention to the larvae, especially in searching for species whose adults are not
attracted by artificial light and sugar-wine mixture.
Recommendations for biodiversity support
1. To save the forest, it is necessary to forbid clear felling everywhere in the Eastern
Rhodopes. The main problem is that in the region mostly wood is used for heating and in
many places in the Eastern Rhodopes charcoal is produced. It is necessary to find an alterna-
tive source of heating. In felling only dead trees are to be cut and introduced foreign tree species.
538 S. BESHKOV, M. LANGOUROV
Beron page 538
2. In further afforestation only local tree species must be used.
3. Mercury-vapor lamps in settlements and on single buildings to be replaced by
halogen lamps with reddish or orange light, which are a slight attraction for insects.
Insects are strongly attracted by mercury-vapor lamps, they are lost from their habitats
and consequently get burned, squashed or fall victims to predators.
4. Control on mowing and pasture in natural habitats and establishing new protected areas.
5. Strict ban on the usage of pesticides in forests and non-cultivated lands. In the
event of usage of pesticides in agricultural lands, airplanes are not to be used.
6. Strict ban on firing of stubblefields.
Acknowledgements
We would like to express our thanks to the people who contributed and helped in the realization of this
article. We are especially grateful to Mr Peter Davey, Mrs Veronica Ferdinandova, Mrs Svetlana Hristova, Mag.
Boyan Petrov, Dr Milen Marinov, Mr Viktor Gashtarov, Dr Herbert Beck and Mr Dimitur Vassilev for the
material and data provided.
Species collected in caves and galleries
Data on cave Lepidoptera from the Eastern Rhodopes can be found only in the
articles of BESHKOV (1995) and BESHKOV & PETROV (1996). All these, together with
some original unpublished data are included in the list below. Up to now from the caves
and galleries of the Eastern Rhodopes 15 Lepidoptera species are known, recorded in 9
caves and 5 galleries.
TINEIDAE
Monopis laevigella ([Denis & Schiffermüller], 1775) =rusticella (Hübner, 1796),
=saturella (Haworth, 1828)
Subtroglophil.
Tilki Ini Cave near Ostrovitsa Village, Kardjali District.
Monopis obviella ([Denis & Schiffermüller], 1775) (=ferruginella (Hübner,
1813), nec Thunberg, 1788)
Karangil Cave above Shiroko Pole Village, Kardjali district, 280m, 21.10.2003, S.
Beshkov, P. Beron & B. Petrov leg. 4 ‡‡ and 5 †† specimens, genitalia checked.
ACROLEPIIDAE
Digitivalva pulicariae (Klimesch, 1956)
Subtroglophil?
Karagug Cave on Djebelska Reka river below Tarnovtsi Village, Djebel district, 400m,
22.10.2003, S. Beshkov, P. Beron & B. Petrov leg. single specimen, genitalia checked;
Gyaur Hambar Cave above Ridino Village, 12.12.2000, hibernating in the cave, 2 ‡‡ and
1 , genitalia checked.
Insecta: Lepidoptera 539
Beron page 539
ALUCITIDAE
Alucita cymatodactyla Zeller, 1852
Subtroglophil.
Prilepnata Peshtera Cave near Sredna Arda Railway Station, Kardjali District; Tilki
Ini Cave near Ostrovitsa Village, Kardjali District; Maarata Cave near Madrets Village,
Perperek District; Razklonenata Peshtera (Gouk-In) Cave in Dishlik-Dere Valley, between
Dolno Cherkovishte and Oreshari villages.
NYMPHALIDAE
Inachis io (Linnaeus, 1758)
Trog o xe n .
Gyaur Hambar Cave near Ridino Village; Zlatnata Yama Cave near Kremen (=Akcha)
Village, Podkova District, 520 m, hibernating.
Nymphalis polychloros (Linnaeus, 1758)
Trog o xe n .
Kodja-Kaya Hill near Odrintsi Village 320 m, hibernating in a bunker; Gyaur Ham-
bar Cave near Ridino Village.
GEOMETRIDAE
Timandra comae A. Schmidt, 1931 =griseata W. Petersen, 1902
Facultative trogoxen
Gallery near Dyadovskiya Kemer Bridge near Dyadovtsi Village, Ardino District.
Xanthorhoe oxybiata (Milliére, 1872)
Facultative trogoxen
Samara Cave near Ribino Village.
Triphosa dubitata (Linnaeus, 1758)
Trog l o x en
Razklonenata Peshtera (Gouk-In) Cave in Dishlik-Dere Valley, between Dolno Cherk-
ovishte and Oreshari villages.
NOCTUIDAE
Autophila limbata (Staudinger, 1871)
According to BERON (1972, 1975, 1995) Autophila limbata is trogloxen. Here this
species is accepted as subtroglophil.
Disused mine gallery near Madjarovo on the road to Borislavtsi Village; Samara
Cave near Ribino Village; Gyaur Hambar Cave near Ridino.
Scoliopteryx libatryx (Linnaeus, 1758)
According to GUEORGUIEV & BERON (1962), BERON & GUEORGUIEV (1967)
Scoliopteryx libatryx is trgloxen; According to BERON (1972, 1975, 1994) it is subtroglo-
phil. Here also it is accepted as subtroglophil.
540 S. BESHKOV, M. LANGOUROV
Beron page 540
Razklonenata Peshtera (Gouk-In) Cave in Dishlik-Dere Valley, between Dolno Cherk-
ovishte and Oreshari villages; Samara Cave near Ribino Village; Maarata Cave near Ma-
drets Village, Perperek District; Disused Mine Gallery near Madjarovo on the road to
Borislavtsi Village; Disused Mine Gallery in Gurgen Dere Valley near Gaberevo Village,
Madjarovo District; Gyaur Hambar Cave near Ridino Village.
Hypena rostralis (Linnaeus, 1758)
Trogloxen according to GUEORGUIEV & BERON (1962), BERON (1995).
Prilepnata Peshtera Cave near Sredna Arda Railway Station; “Maarata” cave near Madrets
Village, Perperek District; Disused Mine Gallery betwen Lozen and Cherna Mogila villages.
Hypena palpalis (Hübner, 1796)
Trog l oxen or S ub t ro g loph i l ?
Disused Mine Gallery near Madjarovo on the road to Borislavtsi Village; Samara
Cave near Ribino Village.
Pyrois effusa (Boisduval, [1828])
Trog l oxe n .
“Zlatnata Yama” Cave near Kremen (Akcha) Village, Djebel district; Karagug Cave
on Djebelska Reka river below Tarnovtsi Village, Djebel district, 22.10.2003 on copula;
Disused Mine Gallery near Madjarovo on the road to Borislavtsi Village; Gyaur Hambar
Cave near Ridino Village.
Mormo maura (Linnaeus, 1758)
Trog l oxe n .
Gallery near Dyadovskiya Kemer Bridge near Dyadovtsi Village, Ardino District.
List with UTM codes of the localities
1. Tilki Ini Cave near Ostrovitsa Village, Kardjali District - LG 60.
2. Prilepnata Peshtera Cave near Sredna Arda Railway Station, Kardjali District - LG 71.
3. Maarata Cave near Madrets Village, Perperek District - LG 71.
4. Razklonenata Peshtera (=Gouk-In) Cave in Dishlik-Dere Valley, between Dolno Cherkovishte and
Oreshari villages – LG 90.
5. Zlatnata Yama Cave near Kremen (=Akcha) Village, Podkova District – LF 67.
6. Samara Cave near Ribino Village - LF 78.
7. Gyaur Hambar Cave near Ridino Village – LF 58.
8. Karagug Cave on Djebelska Reka river below Tarnovtsi Village, Djebel district, 400m, N41°30’25.1";
E025°15’07.1" MH KN 21PM.
9. Karangil Cave above Shiroko Pole Village, Kardjali district, 280m - LG 71.
10. Gallery near Dyadovskiya Kemer Bridge near Dyadovtsi Village, Ardino District – LG 40.
11. Disused Mine Gallery between Lozen and Cherna Mogila villages – MG 12.
12. Disused Mine Gallery in Gyurgen Dere Valley near Gaberevo Village, Madjarovo District – MG 00.
13. Disused Mine Gallery near Madjarovo on the road to Borislavtsi Village - MG 01.
14. Disused Bunker on Kodja-Kaya Hill near Odrintsi Village – MF 28.
Insecta: Lepidoptera 541
Beron page 541
List of species
The systematics of the families follows KARSHOLT & RAZOWSKY (1996): The
Lepidoptera of Europe. Appolo Books, Stenstrup, 380 pp. The systematics of the families
mostly follows the different authors in KARSHOLT & RAZOWSKY (1996). Localities are
given mostly from west to east and from south to north.
HEPIALIDAE
Triodia silvina (Linnaeus, 1761)
Lebed Village, Djebel District; Sredna Arda Railway Station; Studen Kladenets Vil-
lage; The crossroad to Haskovo near Knizhovnik Village; Momina Skala Hut; Harmanli;
Meden Buk Village; Belopolyane Village.
Triodia amasinus dobrogensis (Caradja, 1932)
The crossroad to Haskovo near Knizhovnik Village; Studen Kladenets Village;
Momina Skala Hut; the bridge on Arda River near Madjarovo; Meden Buk Village.
ADELIDAE
Nemophora metallica (Poda, 1761)
Krumovgrad, 22.7.1968 (TULESCHKOV & SLIVOV, 1975b: 162).
Nematopogon adansoniella (Villers, 1789) =panzerella (Fabricius, 1794)
Krumovgrad, 3.6.1967, 1 ; Kardjali, 1.6.1967, 2 †† (TULESCHKOV & SLIVOV,
1975b: 162).
TINEIDAE
Triaxomera parasitella (Hübner, 1796)
Dabovets Village, 600 m, 18.6.1969, 1 (TULESCHKOV & SLIVOV, 1975b: 162).
Nemapogon clematella (Fabricius, 1781) =arcella auct., nec (Fabricius, 1776)
Krumovgrad, 3.6.1967, 2 †† (TULESCHKOV & SLIVOV, 1975b: 162).
Neurothaumasia ankerella (Mann, 1867)
Krumovgrad, 3.6.1967, 1 (TULESCHKOV & SLIVOV, 1975b: 162); The bor-
der post near Avren Village, 21.6.1969, 2 †† and 1 (TULESCHKOV & SLIVOV,
1975b: 162).
Tinea pellionella Linnaeus, 1758
Mandritsa Village, 19.6.1969, 2 †† (TULESCHKOV & SLIVOV, 1975b: 162).
Monopis laevigella ([Denis & Schiffermüller], 1775) (=rusticella (Hübner, 1796),
=saturella (Hawirth, 1828)
Tilki Ini Cave near Ostrovitsa Village, Kardjali District.
542 S. BESHKOV, M. LANGOUROV
Beron page 542
Monopis obviella ([Denis & Schiffermüller], 1775) (=ferruginella (Hübner,
1813), nec Thunberg, 1788)
Karangil Cave above Shiroko Pole Village, Kardjali district, 280m, 21.10.2003, S.
Beshkov, P. Beron & B. Petrov leg. 2 ‡‡ and 3 †† specimens, genitalia checked; The
border post near Avren Village, 21.6.1969, 2 ††, 1 (TULESCHKOV & SLIVOV, 1975b:
162); Dabovets Village, 18.6.1969, 2 †† (TULESCHKOV & SLIVOV, 1975b: 162).
Euplocamus anthracinalis (Scopoli, 1763)
The bridge on Arda River near Madjarovo. Flies at a day time. Collected once in the
second half of May in an oak forest.
Euplocamus ophisa (Cramer, 1779)
The bridge on Arda River near Madjarovo; Momina Skala Hut nar Madjarovo. Flies
at a day time. Collected in the second half of May in an oak forest.
PSYCHIDAE
Melasina christenseni Hattenschwiler, 1990
Sredna Arda Railway Station, single specimen (bag), P. Hattenschwiler det. New spe-
cies for the Bulgarian fauna.
Eochorica balcanica (Rebel, 1919)
The Iron Bridge on Arda River near Dolno Cherkovishte; the bridge on Arda River
near Madjarovo. Rare and local species, previously known as a Balkan endemic species,
recently found in Asia Minor. From the Eastern Rhodopes known only by two speci-
mens collected in August-September.
Rebelia perlu cidella (Bruand, 1853)
Studen Kladenets Village; The bridge on Arda River near Madjarovo; Ivaylovgrad
Dam, Arda Hut near Dabovets Village; Belopolyane Village. Local and rare species.
Rebelia macedonica Pinker, 1956
Siv Kladenets Village, Ivaylovgrad District. A local and rare species. The taxa of this
genus need revision.
Epichnopterix kova csi Sieder, 1955
Kulich Village, Potochnitsa District. A local and rare species.
Pachythelia villosella (Ochsenheimer, 1810)
Sredna Arda Railway Station.
Ptilocephala plumifera (Ochsenheimer, 1810)
Sredna Arda Railway Station; Gorna Krepost Village; Studen Kladenets Village;
Kulich Village, Potochnitsa District; Ivaylovgrad; Siv Kladenets Village, Ivaylovgrad
District.
Insecta: Lepidoptera 543
Beron page 543
Megalophanes viciella orientalis Driel
Studen Kladenets Village; Dishlik-Dere Valley near Oreshari Village; the bridge on
Arda River near Madjarovo; Ivaylovgrad Dam, Arda Hut near Dabovets Village; Mandrit-
sa Village; Siv Kladenets Village.
Apterona crenulella (Bruand, 1853) (bisex. form)
[Haskovski] Mineralni Bani, 2.7.1977, often mining leafs of Teucrium chamaedrys L.
(BEIGER, 1979: 498)
Apterona helicoidella (Vallot, 1827) parth. form
Sredna Arda Railway Station.
YPONEOMEUTIDAE
Yponomeuta plumbella ([Denis & Schiffermüller], 1775)
Chorbadjiysko Village, 22.6.1969, 2 †† (TULESCHKOV & SLIVOV, 1975b: 163).
Swammerdamia pyrella (Villers, 1789)
Dabovets Village, 20.7.1968, 1 (TULESCHKOV & SLIVOV, 1975b: 163).
YPSOLOPHIDAE
Ypsolopha asperella (Linnaeus, 1761)
Often in the oak forests in the Eastern Rhodopes (TULESCHKOV & SLIVOV,
1975b: 163), but without specified localities.
Ypsolopha lucella (Fabricius, 1775)
Djebelska Reka river below Tarnovtsi Village, Djebel district, 380m alt., 21.10.2003,
S. Beshkov, P. Beron & B. Petrov leg. 3 ‡‡ (misidentification is possible); Kardjali, 23.7.1968,
1 (TULESCHKOV & SLIVOV, 1975b: 163); Plevun Village, 22.7.1068, 1 , 2 ‡‡
(TULESCHKOV & SLIVOV, 1975b: 163); Ivaylovgrad, 20.6.1969, 2 ††, 1 (TULE-
SCHKOV & SLIVOV, 1975b: 163).
Ypsolopha sylvella (Linnaeus, 1767)
Near the Zlatnata Yama Cave near Kremen (Akcha) Village, Djebel district, 280m,
22.10.2003, S. Beshkov, P. Beron & B. Petrov leg. 1 , genitalia checked.
Ypsolopha parenthesella (Linnaeus, 1761)
Near the Zlatnata Yama Cave near Kremen (Akcha) Village, Djebel district, 280m,
22.10.2003, S. Beshkov, P. Beron & B. Petrov leg. 1 , genitalia checked.
Ypsolopha sequella (Clerck, 1759)
Dabovets Village, 18.6.1969, 1 (TULESCHKOV & SLIVOV, 1975b: 163); Plevun
Village, 22.7.1068, 2 †† (TULESCHKOV & SLIVOV, 1975b: 163); Madjarovo, 07.8.2003,
08.8.2003, 11.8.2003, at a light trap, Peter Davey leg.; The Bridge on the Arda River near
Madjarovo, 09.8.2003, at a light trap, Peter Davey leg.
544 S. BESHKOV, M. LANGOUROV
Beron page 544
Ypsolopha vitella (Linnaeus, 1758)
The Bridge of Arda Rived near Madjarovo.
PLUTELLIDAE
Plutella xylostella (Linnaeus, 1758) (=maculipennis Curtis, 1832)
Djebelska Reka river below Tarnovtsi Village, Djebel district, 380m alt., 21.10.2003,
S. Beshkov, P. Beron & B. Petrov leg. 1 (misidentification is possible); Madjarovo,
08.8.2003, 11.8.2003, at a light trap, Peter Davey leg.
ACROLEPIIDAE
Digitivalva pulicariae (Klimesch, 1956)
Karagug Cave on Djebelska Reka river below Tarnovtsi Village, Djebel district, 400m,
22.10.2003, S. Beshkov, P. Beron & B. Petrov leg. single specimen, genitalia checked;
Gyaur Hambar Cave above Ridino Village, 12.12.2000, hybernating in the cave, 2 ‡‡
and 1 , genitalia checked.
GLYPHIPTERYGIDAE
Glyphipteryx thrasonella (Scopoli, 1763)
Kardjali, 1.6.1967, 1 , 23.7.1968, 1 (TULESCHKOV & SLIVOV, 1975b: 167).
DEPRESSARIIDAE
Agonopterix alstromeriana (Clerck, 1759)
Chorbadjiysko Village, 22.6.1969, 1 (TULESCHKOV & SLIVOV, 1975b: 166).
Depressaria badiella (Hübner, 1796)
Djebelska Reka river below Tarnovtsi Village, Djebel district, 380m alt., 21.10.2003,
S. Beshkov, P. Beron & B. Petrov leg., 1 , genitalia checked; Between Shiroko Pole and
Sedlovina villages, Kardjali district, 260m alt., 20.10.2003, S. Beshkov, P. Beron & B.
Petrov leg. 2 ‡‡, genitalia checked. This species seems to be new for Bulgaria. It is not
included as present in Bulgaria in KARSHOLT & RAZOWSKI (1996: 67).
Depressaria douglasella Stainton, 1849
Djebelska Reka river below Tarnovtsi Village, Djebel district, 380m alt., 21.10.2003,
S. Beshkov, P. Beron & B. Petrov leg. 1 , genitalia checked; Between Shiroko Pole and
Sedlovina villages, Kardjali district, 260m alt., 20.10.2003, S. Beshkov, P. Beron & B.
Petrov leg. 1 , genitalia checked.
ETHMIIDAE
Ethmia pusiella (Linnaeus, 1758)
The bridge on Arda River near Madjarovo.
Ethmia aurif luella (Hübner, 1810)
Siv Kladenets Village, Ivaylovgrad district.
Insecta: Lepidoptera 545
Beron page 545
Ethmia bipunctella (Fabricius, 1775)
The bridge on Arda River near Madjarovo, 18.05.2004, 1 ; Near the Zlatnata Yama
Cave near Kremen (Akcha) Village, Djebel district, 280m, 22.10.2003, S. Beshkov, P.
Beron & B. Petrov leg. 1 and 1 . This species is not included as present in Bulgaria in
KARSHOLT & RAZOWSKI (1996), although it has many published localities in the
country.
OECOPHORIDAE
Epicallima formosella ([Denis & Schiffermüller], 1775)
Kardjali, 23.6.1969, 1 ; Ibid, 23.7.1968, 1 , 2 ‡‡ (TULESCHKOV & SLIVOV,
1975b: 166).
Esperia oliviella (Fabricius, 1794)
Kardjali, 23.6.1969, 4 ††, 23.7.1968, 1 (TULESCHKOV & SLIVOV, 1975b:
165); Mandritsa Village, 19.6.1969, 2 ††, 1 (TULESCHKOV & SLIVOV, 1975b: 165).
Oecophora bractella (Linnaeus, 1758)
Kardjali, 1.6.1967, 1 , 2 ‡‡ (TULESCHKOV & SLIVOV, 1975b: 165); Mandritsa
Village, 19.6.1969, 1 , 1 (TULESCHKOV & SLIVOV, 1975b: 165).
Cephalispheira ferrugella ([Denis & Schiffermüller], 1775)
The border post near Avren Village, 21.6.1969, 2 †† (TULESCHKOV & SLIVOV,
1975b: 165).
Telechrysis tripuncta (Haworth, 1828)
Plevun vilage, 22.7.1968, 2 ††, 2 ‡‡ (TULESCHKOV & SLIVOV, 1975b: 166);
Mandritsa Village, 19.6.1969, 2 ††, 1 (TULESCHKOV & SLIVOV, 1975b: 166).
LECITHOCERIDAE
Odites kollarella (O. Costa, 1832) (=luteella Duponchel, 1840; =lutrella, incor-
rect subsequent spelling)
Kardjali, 23.6.1969, 1 (TULESCHKOV & SLIVOV, 1975b: 167).
MOMPHIDAE
Mompha miscella ([Denis & Schiffermüller], 1775)
Kardjali, 23.6.1969, 2 †† (TULESCHKOV & SLIVOV, 1975b: 165); Popsko Vil-
lage, 22.6.1969, 2 †† (TULESCHKOV & SLIVOV, 1975b: 165).
COSMOPTERIGIDAE
Pyroderces argyrogrammos (Zeller, 1847)
Kardjali, 1.6.1967, 2 ††, 2 ‡‡ (TULESCHKOV & SLIVOV, 1975b: 165); Chorbad-
jiysko Village, 21.6.1969, 1 , 1 (TULESCHKOV & SLIVOV, 1975b: 165).
546 S. BESHKOV, M. LANGOUROV
Beron page 546
GELECHIIDAE
Metzneria aprilella (Herrich-Schäffer, 1854)
Kardjali, 1.6.1967, 1 , 23.6.1969, 1 , 2 ‡‡ (TULESCHKOV & SLIVOV, 1975b:
164); Mandritsa Village, 19.6.1969, 1 (TULESCHKOV & SLIVOV, 1975b: 164).
Teleiopsis diffinis (Haworth, 1829)
Near the Zlatnata Yama Cave near Kremen Village, Djebel District, 280m, 22.10.2003,
S. Beshkov, P. Beron & B. Petrov leg. 3 †† and 1 , genitalia of all specimens checked;
Between Shiroko Pole and Sedlovina villages, Kardjali district, 260m alt., 20.10.2003, S.
Beshkov, P. Beron & B. Petrov leg. 1 , genitalia checked; Popsko Village, 22.7.1968, 1
(TULESCHKOV & SLIVOV, 1975b: 164).
Pseudotelphusa scalella (Scopoli, 1763)
Plevun Village, 22.6.1969, 1 (TULESCHKOV & SLIVOV, 1975b: 164); Ivay-
lovgrad, 29.6.1969, 3 †† (TULESCHKOV & SLIVOV, 1975b: 164).
Scrobipalpa atriplicella (Fischer v. Röslerstamm, 1841)
The border post near Avren Village, 21.6.1969, 2 †† (TULESCHKOV & SLIVOV,
1975b: 165).
Stomopteryx detersella (Zeller, 1847)
Krumovgrad, 20.6.1969, 2 †† (TULESCHKOV & SLIVOV, 1975b: 165); Mandrit-
sa Village, 21.7.1968, 1 (TULESCHKOV & SLIVOV, 1975b: 165).
Syncopacma taeniolella (Zeller, 1839)
The border post near Avren Village, 21.6.1969, 1 , 1 (TULESCHKOV & SLIVOV,
1975b: 165); Ivaylovgrad, 20.6.1969, 3 †† (TULESCHKOV & SLIVOV, 1975b: 165).
Nothris verbascella ([Denis & Schiffermüller], 1775)
Chorbadjiysko Village, 22.6.1969, 1 and 1 (TULESCHKOV & SLIVOV, 1975b: 164).
Dichomeris marginella (Fabricius, 1781)
Mandritsa Village, 19.6.1969, 2 ††, 1 (TULESCHKOV & SLIVOV, 1975b: 165).
Helcystogramma lineolella (Zeller, 1839)?
Near the Zlatnata Yama Cave near Kremen (Akcha) Village, Djebel district, 280m,
22.10.2003, S. Beshkov, P. Beron & B. Petrov leg. 1 , genitalia checked. This species
seems to be new for Bulgaria, if it is correctly identified. It is not included as present in
Bulgaria in KARSHOLT & RAZOWSKI (1996: 121).
LIMACODIDAE
Apoda limacodes (Hufnagel, 1766)
Belite Brezi Hut near Ardino; Sredna Arda Railway Station; near Nanovitsa Village;
Krumovgrad; Dishlik-Dere Valley near Oreshari Village; the Iron Bridge on Arda River
near Dolno Cherkovishte; Momina Skala Hut near Madjarovo; 4 km from the bridge on
Insecta: Lepidoptera 547
Beron page 547
Arda River near Madjarovo to Borislavtsi; Ayda Hut near Haskovo; Ivaylovgrad Dam,
Arda Hut near Dabovets Village; Siv Kladenets Village.
ZYGAENIDAE
Theresimima amphelophaga (Bayle-Barelle, 1808)
4 km from the Bridge on Arda River near Madjarovo to Borislavsi, 1; Above
Dishlik-Dere Valley below Oreshari Village, 1†, genitalia checked.
Rhagades pruni ([Denis & Schiffermüller], 1775)
The bridge on Arda River near Madjarovo, 1 in a Malaise trap, genitalia checked.
Adscita drenowskii Alberti, 1939
Dishlik-Dere Valley between Dolno Cherkovishte and Oreshari villages; Near Ko-
vankaya, vicinity of Strandjevo Village; The Bridge on Arda River near Madjarovo. Gen-
italia checked.
Adscita sp.
Dishlik-Dere Valley between Dolno Cherkovishte and Oreshari villages, a single
specimen, sympatric and synchronic with A. drenowskii (4.5.2000). This specimen is ex-
ternally similar to the previous one, but the genitalia differ greatly (aedoeagus without
cornuti) and are unknown to the present authors.
Zygaena punctum Ochsenheimer, 1808
Kroyatsi Hunting Farm near Nanovitsa Village.
Zygaena loti ([Denis & Schiffermüller], 1775) (=achilleae Esper, 1780)
Studen Kladenets Dam, Sredna Arda Railway Station.
Zygaena ephialtes (Linnaeus, 1767)
Near Nanovitsa Village; the Iron Bridge on Arda River near Dolno Cherkovishte.
Zygaena angelicae Ochsenheimer, 1808
Near Gorno Pole Village, Madjarovo district.
Zygaena filipendulae (Linnaeus, 1758)
Between Ardino and Nedelino, 2 km before the crossroad to Nedelino; Studen Klad-
enets Dam, Sredna Arda Railway Station; Studen Kladenets Dam, Kroyatsi Hunting Farm
near Nanovitsa Village; The crossroad to Manastir Village from the road Haskovo-Kard-
jali; Southern of Madjari Village; Ivaylovgrad Dam, Arda Hut near Dabovets Village;
Kamilski Dol Village.
SESIIDAE
Tintia myrmosaeformis (Herrich-Schäffer, 1846)
Studen Kladenets Dam, Sredna Arda Railway Station. Only once 2 specimens were
collected with a feromone.
548 S. BESHKOV, M. LANGOUROV
Beron page 548
Sesia apiphormis (Clerck, 1759)
The border post near Avren Village; Siv Kladenets Village, Ivaylovgrad District;
Mandritsa Village.
Paranthrene tabaniformis (Rottemburg, 1775)
Mezek Village, Svilengrad District.
Synanthedon formicaeformis (Esper, 1783)
The bridge on Arda River near Madjarovo, a single in a Malaise trap, genitalia
checked, but incorrect identification seems possible. In this species the uncus does not go
ahead to the scopula.
Synanthedon vespiformis (Linnaeus, 1761)
The border post near Avren Village.
Synanthedon myopaeformis (Borkhausen, 1789)
Chorbadjiysko Village.
Synanthedon conopiformis (Esper, 1782)
The bridge on Arda River near Madjarovo, 22.05.2004, a single specimen at light.
Bembecia uroceriformis (Treitschke, 1834)
Chorbadjiysko Village; Kardjali.
Pyropteron minianiformis (Freyer, 1843)
Studen Kladenets Village.
Synansphecia triannuliformis (Freyer, 1845)
The bridge on Arda River near Madjarovo, a single in a Malaise trap, genitalia checked.
Synansphecia affinis (Staudinger, 1856)
Studen Kladenets Dam, Sredna Arda Railway Station. Only once 2 specimens were
collected at feromons.
Chamaesphecia annellata (Zeller, 1847)
Kardjali; Mandritsa Village.
Chamaesphecia bibioniformis (Esper, 1800)
The border post near Avren Village.
Chamaesphecia astatiformis (Herrich-Schäffer, 1846)
Ivaylovgrad.
Chamaesphecia tentrediniformis ([Denis & Schiffermüller], 1775)
The Iron Bridge on Arda River near Dolno Cherkovishte Village.
Insecta: Lepidoptera 549
Beron page 549
COSSIDAE
Cossus cossus (Linnaeus, 1758)
Ayda Hut near Haskovo; Studen Kladenets Village; the Iron Bridge on Arda River near
Dolno Cherkovishte; Momina Skala Hut near Madjarovo; Madjarovo; the bridge on Arda
River near Madjarovo; above the bridge on Arda River near Madjarovo; Ivaylovgrad Dam,
Arda Hut near Dabovets Village; Dolno Lukovo Village; Siv Kladenets Village.
Lamellocossus terebra ([Denis & Schiffermüller], 1775)
Belite Brezi Hut near Ardino.
Parahypopta caestrum (Hübner, 1808)
Dolno Lukovo Village; Siv Kladenets Village.
Dyspessa salicicola (Eversmann, 1848)
Studen Kladenets Village; Siv Kladenets Village.
Dyspessa ulula (Borkhausen, 1790)
Studen Kladenets Village; above Dishlik-Dere Valley near Oreshari Village; the bridge on Arda
River near Madjarovo; above the bridge on Arda River near Madjarovo; 4 km from the bridge on
Arda River near Madjarovo to Borislavtsi; Arda Hut near Dabovets Village; near Bivolene Village;
Dolno Lukovo Village; Mandritsa Village; Siv Kladenets Village; Belopolyane Village.
Stygoides colchica (Herrich-Schäffer, 1851) (=Psychidostygia tricolor (Lederer, 1858))
The bridge on Arda River near Madjarovo, a single specimen in a Malaise trap,
29.4.-20.5.2001, M. Langourov leg, genitalia checked. New genus and species for the Bul-
garian fauna, in Europe known from Southern Greece only and from Turkey.
Zeuzera pyrina (Linnaeus, 1761)
Sredna Arda Railway Station; Studen Kladenets Village; Madjarovo; the bridge on
Arda River near Madjarovo; Arda Hut near Dabovets Village; Dolno Lukovo Village; Siv
Kladenets Village.
TORTRICIDAE
Cochylimorpha straminea (Haworth, [1811])
Between Shiroko Pole and Sedlovina villages, Kardjali district, 260 m alt., 20.10.2003,
S. Beshkov, P. Beron & B. Petrov leg., 1 , genitalia checked.
Eugnosta lathoniana (Hübner, 1800)
The border post near Avren Village, 21.6.1969, 1 (TULESCHKOV & SLIVOV,
1975b: 173); Ivaylovgrad, 21.6.1969, 1 (TULESCHKOV & SLIVOV, 1975b: 173).
Aethes margarotana (Duponchel, 1836)
Siv Kladenets Village, Ivaylovgrad district, 04.5.2003, S. Beshkov & V. Gashtarov
leg., 1 at day time, genitalia checked. Missidentification with its closely related species
Aethes williana (Brahm, 1791) however is possible. For a certain identification genitalia
550 S. BESHKOV, M. LANGOUROV
Beron page 550
are needed. This species is not included in RAZOWSKI: Tortricidae: in KARSHOLT &
RAZOWSKI (1996) and if it is correctly identified, it will be new for the Bulgarian fauna.
Aethes bilbaensis (Rössler, 1847)
Krumovgrad, 22.7.1968, 1 (TULESCHKOV & SLIVOV, 1975b: 173).
Cochylis posterana Zeller, 1847
Mandritsa Village, 21.7.1968, 1 , 1 (TULESCHKOV & SLIVOV, 1975b: 174).
Alseuncaria ruficiliana (Haworth, 1811) (=cliella Hübner, 1796, nec [Denis &
Schiffermüler], 1775)
Kardjali, 23.6.1969, 1 , 1 (TULESCHKOV & SLIVOV, 1975b: 173); Svirachi
Village, Ivaylovgrad District, 19.6.1969, 1 , 1 (TULESCHKOV & SLIVOV, 1975b: 173).
Tortrix viridana Linnaeus, 1758
The bridge on Arda River near Madjarovo.
Acleris forsskaleana (Linnaeus, 1758)
Kardjali, 23.7.1968, 2 ††, 3 ‡‡ (TULESCHKOV & SLIVOV, 1975b: 170); the border
post near Avren Village, 21.6.1969, 2 ††, 2 ‡‡ (TULESCHKOV & SLIVOV, 1975b: 169);
Mandritsa Village, 19.6.1969, 3 ††, 1 (TULESCHKOV & SLIVOV, 1975b: 169).
Acleris rhombana ([Denis & Schiffermüller], 1775)
The Bridge of Arda River near Madjarovo.
Acleris variegana ([Denis & Schiffermüller], 1775)
Djebelska Reka river below Tarnovtsi Village, Djebel district, 380m alt., 21.10.2003, S. Beshk-
ov, P. Beron & B. Petrov leg. 1 (genitalia checked) and 3 ‡‡; Near the Zlatnata Yama Cave near
Kremen (Akcha) Village, Djebel district, 280m, 22.10.2003, S. Beshkov, P. Beron & B. Petrov leg.
single specimen; Ivaylovgrad, 22.6.1968, 2 †† (TULESCHKOV & SLIVOV, 1975b: 170).
Acleris hastiana (Linnaeus, 1758)
Mandritsa Village, 21.7.1968, 2 †† (TULESCHKOV & SLIVOV, 1975b: 170).
Acleris quercinana (Zeller, 1849)
Chorbadjiysko Village, 22.6.1969 2 †† (TULESCHKOV & SLIVOV, 1975b: 170);
Dabovets Village, 18.6.1969, 1 (TULESCHKOV & SLIVOV, 1975b: 170).
Acleris literana (Linnaeus, 1758) f. irrorana Hübner
Kardjali, 23.6.1969, 2 †† (TULESCHKOV & SLIVOV, 1975b: 170).
Cnephasia incertana (Treitschke, 1835)
Kardjali, 23.6.1969, 2 †† (TULESCHKOV & SLIVOV, 1975b: 169); Krumovgrad,
3.6.1967, 2 †† (TULESCHKOV & SLIVOV, 1975b: 169); Dabovets Village, 19.6.1969, 1
(TULESCHKOV & SLIVOV, 1975b: 169).
Insecta: Lepidoptera 551
Beron page 551
Epagoge grotiana (Fabricius, 1781)
Between Shiroko Pole and Sedlovina villages, Kardjali district, 260m alt., 20.10.2003,
S. Beshkov, P. Beron & B. Petrov leg. 3 ††, genitalia checked.
Archips podana (Scopoli, 1763)
Madjarovo, 11.8.2003, at a light trap, Peter Davey leg.
Choristoneura hebenstreitella (Müller, 1764) (=sorbiana (Hübner, 1799))
The bridge on Arda River near Madjarovo, second half of May.
Argyrotaenia ljungiana (Thunberg, 1797) (=lepidana Herrich-Schäffer, 1854;
=lepideana, incorrect subsequent spelling)
Chorbadjiysko Village, 22.6.1969, 1 (TULESCHKOV & SLIVOV, 1975b: 172).
Ptycholoma lecheana (Linnaeus, 1758)
The bridge on Arda River near Madjarovo, second half of May.
Apotomis turbidana (Hübner, 1825)
Belite Brezi Hut near Ardino, 900 m, 24.6.1969, 2 ††, 1 (TULESCHKOV & SLIVOV,
1975b: 172); Mandritsa Village, 19.6.1969, 3 †† (TULESCHKOV & SLIVOV, 1975b: 172).
Epinotia caprana (Fabricius, 1798)
Djebelska Reka river below Tarnovtsi Village, Djebel district, 380m alt., 21.10.2003,
S. Beshkov, P. Beron & B. Petrov leg. 1 , genitalia checked, but misidentification is
possible; the specimen differs from other specimens from this species from Europe. This
species is not included for Bulgarian fauna in KARSHOLT & RAZOWSKI (1996: 147),
therefore if it is correctly identified, it seems to be a new species for Bulgaria.
Epinotia abbreviana (Fabricius, 1794) (=trimaculana Donovan, 1806)
Chorbadjiysko Village, 2.6.1969, 1 (TULESCHKOV & SLIVOV, 1975b: 172);
Ivaylovgrad, 20.6.1969, 1 (TULESCHKOV & SLIVOV, 1975b: 172).
Epinotia nisella (Clerck, 1759)
Djebelska Reka river below Tarnovtsi Village, Djebel district, 380 m alt., 21.10.2003,
S. Beshkov, P. Beron & B. Petrov leg. 1 , genitalia checked. This species is not included
for Bulgarian fauna in KARSHOLT & RAZOWSKI (1996: 147), therefore it is reported
here as a new species for Bulgaria.
Zeiraphera isertana (Fabricius, 1794)
Kardjali, 23.6.1969, 2 ††, 1 (TULESCHKOV & SLIVOV, 1975b: 171); Madjaro-
vo, 07.8.2003, at a light trap, Peter Davey leg.; the Bridge on Arda River near Madjarovo,
09.8.2003, at a light trap, Peter Davey leg.
Eucosma cana (Haworth, 1811)
Ivaylovgrad, 22.7.1968, 2 †† (TULESCHKOV & SLIVOV, 1975b: 171).
552 S. BESHKOV, M. LANGOUROV
Beron page 552
Gypsonoma dealbana Frölich, 1829)
Plevun Village, 20.6.1969, 2 †† (TULESCHKOV & SLIVOV, 1975b: 171).
Notocelia uddmanniana (Linnaeus, 1758)
Krumovgrad, 22.7.1968, 1 (TULESCHKOV & SLIVOV, 1975b: 171); Popsko
Village, 22.6.1968, 1 (TULESCHKOV & SLIVOV, 1975b: 171); Mandritsa Village,
19.6.1969, 2 †† (TULESCHKOV & SLIVOV, 1975b: 171).
Notocelia trimaculana (Haworth, 1811) (=suffusana Duponchel, [1843])
Kardjali, 23.7.1968, 1 (TULESCHKOV & SLIVOV, 1975b: 171); Krumovgrad,
3.6.1967, †† (TULESCHKOV & SLIVOV, 1975b: 171).
Enarmonia formosana (Scopoli, 1763) (=formosaria, incorrect subsequent spell-
ing, =Grapholitha woeberiana [Denis & Schiffermüller], 1775)
Kardjali, 1.6.1967, 1 ; Dabovets Village, 19.6.1969, 1 (TULESCHKOV & SLIVOV,
1975b: 170).
Ancylis comptana (Frölich, 1828)
Kardjali, 23.6.1969, 1 (TULESCHKOV & SLIVOV, 1975b: 172).
Ancylis mitterbacheriana ([Denis & Sshiffermüller], 1775)
Kardjali, 1.6.1967, 2 †† (TULESCHKOV & SLIVOV, 1975b: 172); near Madjarovo.
Cydia fissana (Frölich, 1828)
The border post near Avren Village, 21.6.1969, 1 , 1 (TULESCHKOV & SLIVOV,
1975b: 170); Mandritsa Village, 19.6.1969, 2 †† (TULESCHKOV & SLIVOV, 1975b: 170).
Cydia pomonella (Linnaeus, 1758)
Madjdarovo, 07.8.2003, at light trap, Peter Davey leg.
Cydia pyrivora (Danilevsky, 1947)
Dabovets Village, 18.6.1969, 1 (TULESCHKOV & SLIVOV, 1975b: 170).
Cydia amplana (Hübner, 1800)
The Bridge on Arda River near Madjarovo, 09.8.2003, at a light trap, Peter Davey leg.
Pammene amygdalana (Duponchel, 1842)
Mandritsa Village, 19.6.1969, 1 (TULESCHKOV & SLIVOV, 1975b: 170).
Dichrorampha cinerosana (Herrich-Schäffer, 1851)
The border post near Avren Village, 21.6.1969, 2 †† (TULESCHKOV & SLIVOV,
1975b: 170).
Insecta: Lepidoptera 553
Beron page 553
ALUCITIDAE
Alucita cymatodactyla Zeller, 1852
Prilepnata Peshtera Cave near Sredna Arda Railway Station, Kardjali District; Tilki
Ini Cave near Ostrovitsa Village, Kardjali District; Maarata Cave near Madrets Village,
Perperek District; Razklonenata Peshtera (Gouk-In) Cave in Dishlik-Dere Valley, between
Dolno Cherkovishte and Oreshari villages.
Alucita huebneri Wallengren, 1859
Djebelska Reka river below Tarnovtsi Village, Djebel district, 380 m alt., 21.10.2003,
S. Beshkov, P. Beron & B. Petrov leg. 1 and 1 , genitalia checked. This species is not
included as present in Bulgaria in KARSHOLT & RAZOWSKI (1996), although it is
reported several times from Bulgaria.
PTEROPHORIDAE
Stenoptilia gratiolae Gibeaux & Nel, 1990 (=paludicola auct., nec (Wallengren, 1862)
Near the Zlatnata Yama Cave near Kremen Village, Djebel District, 280 m, 22.10.2003,
S. Beshkov, P. Beron & B. Petrov leg. 1 , genitalia checked; Djebelska Reka River below
Tarnovtsi Village, Djebel District, 380 m alt., 21.10.2003, S. Beshkov, P. Beron & B.
Petrov leg. 1 , genitalia checked.
Oxyptilus parvidactyla (Haworth, 1811)
The Bridge on Arda River near Madjarovo, 17-23.05.2004, 1 , genitalia checked.
Stangeia siceliota (Zeller, 1847)
Above the Bridge on Arda River near Madjarovo.
Emmelina monodactyla (Linnaeus, 1758)
The Bridge on Arda River near Madjarovo.
CARPOSINIDAE
Carposina scirrhosella Herrich-Sschдffer, 1854
Ivaylovgrad, 22.2.1968, 1 (TULESCHKOV & SLIVOV, 1975b: 174).
THYRIDIDAE
Thyris fenestrella (Scopoli, 1763)
Kulich Village, Potochnitsa District; the Iron Bridge on Arda River near Dolno
Cherkovishte Village.
PYRALIDAE
Aphomia sociella (Linnaeus, 1758)
Madjarovo, 11.8.2003, at a light trap, Peter Davey leg.
Aphomia zelleri Jonnis, 1932
Mandritsa Village, 19.6.1969, 1 (TULESCHKOV & SLIVOV, 1975b: 174).
554 S. BESHKOV, M. LANGOUROV
Beron page 554
Lamoria anella ([Denis & Sschiiermьller], 1775)
Popsko Village, 2.7.1968, 1 (TULESCHKOV & SLIVOV, 1975b: 174).
Galleria mellonella (Linnaeus, 1758)
Popsko Village, 20.6.1969, 1 (TULESCHKOV & SLIVOV, 1975b: 175).
?Synaphe moldavica (Esper, 1794)
2 km before Sussam Village from Gorski Izvor.
Synaphe antennalis (Fabricius, 1794) (=connectalis Hьbner, 1796)
Kardjali, 23.6.1969, 1 (TULESCHKOV & SLIVOV, 1975b: 177).
Synaphe punctalis (Fabricius, 1775)
The Bridge on Arda River near Madjarovo, 09.8.2003, at a light trap, Peter Davey leg.
Pyralis regalis ([Denis & Sschiiermьller], 1775)
Near the Zlatnata Yama Cave near Kremen Village, Djebel District, 280m, 22.10.2003,
S. Beshkov, P. Beron & B. Petrov leg. single specimen; The border post of Avren Village,
1.6.1969, 1 , 1 (TULESCHKOV & SLIVOV, 1975b: 177); The bridge on Arda River
near Madjarovo.
Pyralis farinalis (Linnaeus, 1758)
Krumovgrad; The bridge on Arda River near Madjarovo.
Aglossa pinguinalis (Linnaeus, 1758)
Djebelska Reka river below Tarnovtsi Village, Djebel District, 380 m alt., 21.10.2003,
S. Beshkov, P. Beron & B. Petrov leg. 1 , genitalia checked.
Stemmatophora combustalis (Fischer v.Röslerstamm, 1842)
Krumovgrad, 22.7.1968 1 (TULESCHKOV & SLIVOV, 1975b: 177); Ivaylovgrad,
20.6.1969, 2 ††, 1 (TULESCHKOV & SLIVOV, 1975b: 177).
Actenia brunnealis (Treitschke, 1829)
Krumovgrad, 22.7.1968, 1 , 1 (TULESCHKOV & SLIVOV, 1975b: 177); Momi-
na Skala Hut near Madjarovo.
Hypsopygia costalis (Fabricius, 1775)
Near the Zlatnata Yama Cave near Kremen (Akcha) Village, Djebel District, 280m,
22.10.2003, S. Beshkov, P. Beron & B. Petrov leg. single specimen; Madjarovo, 11.8.2003,
at a light trap, Peter Davey leg.; Belopolyane Village, Ivaylovgrad district.
Herculia rubidalis ([Denis & Schiffermüller], 1775)
The border post near Avren Village, 21.6.1969, 2 †† (TULESCHKOV & SLIVOV,
1975b: 177).
Insecta: Lepidoptera 555
Beron page 555
Orthopygia glaucinalis (Linnaeus, 1758)
Near the Zlatnata Yama Cave near Kremen Village, Djebel District, 280m, 22.10.2003,
S. Beshkov, P. Beron & B. Petrov leg. single specimen; Podrumche Village, Krumovrad
District, 21.6.1969, 2 ††, 1 (TULESCHKOV & SLIVOV, 1975b: 177).
Eudotrica f lammealis ([Denis & Schiffermüller], 1775)
Madjarovo, 07.8.2003, at a light trap, Peter Davey leg.; Zhaltichalskoto Dere Valley
near Meden Buk Village.
Pimpeliella dilutella ([Denis & Schiffermüller], 1775) (=dilutella Hübner, 1796)
Near the Zlatnata Yama Cave near Kremen (Akcha) Village, Djebel District, 280m,
22.10.2003, S. Beshkov, P. Beron & B. Petrov leg. 1 , genitalia checked.
Sciota (Denticera) divisella (Duponchel, 1842)
Near the Zlatnata Yama Cave near Kremen Village, Djebel District, 280m, 22.10.2003,
S. Beshkov, P. Beron & B. Petrov leg. 1 , genitalia checked.
Selagia spadicella (Hübner, 1796)
Krumovgrad, 22.7.1968, 2 †† (TULESCHKOV & SLIVOV, 1975b: 176).
Etiella zinckenella (Treitschke, 1832)
Studen Kladenets Dam, Sredna Arda Railway Sation; Madjarovo; Zhaltichalskoto
Dere Valley near Meden Buk Village.
Oncocera semirubella (Scopoli, 1763)
Madjarovo, 11.8.2003, at a light trap, Peter Davey leg.
Dioryctria simplicella Heinemann, 1863 (=mutatella Fuchs, 1903)
Between Shiroko Pole and Sedlovina villages, Kardjali district, 260m alt., 20.10.2003,
S. Beshkov, P. Beron & B. Petrov leg. 1 , genitalia checked. This species seems to be new
for Bulgaria, if it is correctly identified. It is not included as present in Bulgaria in KAR-
SHOLT & RAZOWSKI (1996: 172).
Phycita roborella ([Denis & Schiffermüller], 1775) (=spissicella Fabricius, 1776)
Kardjali: 23.6.1969, 1 , 23.7.1968, 1 , 2 ‡‡ (TULESCHKOV & SLIVOV, 1975b:
176); Madjarovo, 11.8.2003, at a light trap, Peter Davey leg.
Oxybia transversella (Duponchel, 1836)
Belopolyane Village, Ivaylovgrad district.
Conobathra tumidana ([Denis & Schiffermüller], 1775)
Popsko Village, 2.6.1969, 1 (TULESCHKOV & SLIVOV, 1975b: 176); Madjaro-
vo, 07.8.2003, at a light trap, Peter Davey leg.
556 S. BESHKOV, M. LANGOUROV
Beron page 556
Conobathra repandana (Fabricius, 1798)
Momina Skala Hut near Madjarovo.
Trachycera advenella (Zincken, 1818)
Madjarovo, 11.8.2003, at a light trap, Peter Davey leg. This species is not included in
SPEIDEL: Pyralidae, in KARSHOLT & RAZOWSKI (1996) and it is new for the Bulgar-
ian fauna.
Eurhodope rosella (Scopoli, 1763)
The border post near Avren Village, 21.6.1969, 1 , 1 (TULESCHKOV & SLIVOV,
1975b: 176); Mandritsa Village, 19.6.1969, 1 (TULESCHKOV & SLIVOV, 1975b: 176).
Myelois circumvoluta (Frourcroy, 1785) (=cribrella Hübner, 1796)
Above the bridge on Arda River near Madjarovo.
?Pterothrixidia rufella (Duponchel, 1836)
The bridge on Arda River near Madjarovo, 17-23.05.2004, S. Beshkov & H. Beck
leg., in coll. S. Beshkov,1 , genitalia checked.
Bradyrrhoa gilveolella (Treitschke, 1832)
Studen Kladenets Village; Zhaltichalskoto Dere Valley near Meden Buk Village; Be-
lopolyane Village.
Ancylosis cinnamomela (Duponchel, 1836)
Krumovgrad, 2.7.1968, 1 , 1 (TULESCHKOV & SLIVOV, 1975b: 176); Plevun
Village, 20.6.1969, 2 ††, 1 .
Homoeosoma sinuella (Fabricius, 1794)
Madjarovo, 06.8.2003, at a light trap, Peter Davey leg.
Scoparia luteolaris (Scopoli, 1772) (=ochrealis [Denis & Schiffermüller], 1775)
Podrumche Village, 21.6.1969, 2 ††, 1 (TULESCHKOV & SLIVOV, 1975b: 178).
Scoparia ingratella (Zeller, 1846)
Belite Brezi Hut near Ardino, 1000 m.
Dipleurina lacustrata (Panzer, 1804) = crataegella auct., nec (Linnaeus, 1758)
Belite Brezi Hut near Ardino, 1000 m.
Eudonia mercurella (Linnaeus, 1758)
Studen Kladenets Village.
Euchromius superbellus (Zeller, 1849)
The Bridge on Arda River near Madjarovo, 17-23.05.2004, 1 , genitalia checked.
Insecta: Lepidoptera 557
Beron page 557
Calamotropha paludella (Hübner, 1824)
Near Zhelezino Village.
Agriphila tristella ([Denis & Schiffermüller], 1775)
Near the Zlatnata Yama Cave near Kremen Village, Djebel District, 280m, 22.10.2003,
S. Beshkov, P. Beron & B. Petrov leg. 1 , genitalia checked.
Agriphila inquinatella ([Denis & Schiffermüller], 1775)
Studen Kladenets Village; Zhaltichalskoto Dere Valley near Meden Buk Village.
Agriphila latistria (Haworth, [1811])
Zhaltichalskoto Dere Valley near Meden Buk Village.
Agriphila dalmatinellus (Hampson, 1900)
Studen Kladenets Village.
Catoptria mytilella (Hübner, 1805)
Krumovgrad, 22.7.1968, 1 (TULESCHKOV & SLIVOV, 1975b: 175).
Catoptria pinella (Linnaeus, 1758)
Madjarovo, 06.8.2003, at a light trap, Peter Davey leg.
Catoptria falsella (([Denis & Schiffermüller], 1775)
The Bridge on Arda River near Madjarovo, 09.8.2003, at a light trap, Peter Davey leg.
Catoptria confusellus (Staudinger, 1882)
Djebelska Reka river below Tarnovtsi Village, Djebel district, 380 m alt., 21.10.2003,
S. Beshkov, P. Beron & B. Petrov leg. 1 specimen, genitalia checked; Studen Kladenets
Village; Momina Skala Hut; Zhaltichalskoto Dere Valley near Meden Buk Village; Be-
lopolyane Village.
Pediasia matricella (Treitschke, 1832)
Studen Kladenets Village; between Shiroko Pole and Sedlovina villages, Kardjali
District, 260m alt., 20.10.2003, S. Beshkov, P. Beron & B. Petrov leg. 2 ††, genitalia
checked; Zhaltichalskoto Dere Valley near Meden Buk Village.
Ancylolomia tentaculella (Hübner, 1796)
Madjarovo, 11.8.2003, at a light trap, Peter Davey leg.; Zhaltichalskoto Dere Valley
near Meden Buk Village.
Scirpophaga praelata (Scopoli, 1763)
2 km before Sussam Village from Gorski Izvor.
Cataclysta lemnata (Linnaeus, 1758)
Mandritsa Village, 19.6.1969, 1 (TULESCHKOV & SLIVOV, 1975b: 178).
558 S. BESHKOV, M. LANGOUROV
Beron page 558
Elophila nymphaeata (Linnaeus, 1758)
Kardjali, 23.6.1969, 4 ††, 1 (TULESCHKOV & SLIVOV, 1975b: 178, GANEV,
1982b: 228); Mandritsa Village, 19.6.1969, 3 †† (TULESCHKOV SLIVOV, 1975b:
178, GANEV, 1982b: 228); 2 km before Sussam Village from Gorski Izvor.
Nymphula stagnata (Donovan, 1806)
Kardjali, 23.6.1969, 3 ††, 2 ‡‡ (TULESCHKOV & SLIVOV, 1975b: 178).
Aporodes floralis (Hübner, 1809)
Krumovgrad, 23.7.1968, 1 , 3 ‡‡ (TULESCHKOV & SLIVOV, 1975b: 180); Po-
drumche Village, 21.6.1969, 1 , 2 ‡‡ (TULESCHKOV & SLIVOV, 1975b: 180); Mad-
jarovo, 06.8.2003, at a light trap, Peter Davey leg.; Siv Kladenets Village, Ivaylovgrad
District, 04.5.2003, S. Beshkov & V. Gashtarov leg., 1 at day time.
Ephelis cruentalis (Geyer, 1832)
Krumovgrad, 22.7.1968, 2 †† (TULESCHKOV & SLIVOV, 1975b: 178); Ibid,
03.07.2004, S. Beshkov leg.
Titanio normalis (Hübner, 1796)
Popsko Village, 2.7.1968, 1 (TULESCHKOV & SLIVOV, 1975b: 178).
Eurrhypis pollinalis ([Denis & Schiffermüller], 1775)
Belite Brezi Hut near Ardino, 1000 m.
Evergestis frumentalis (Linnaeus, 1761)
The bridge on Arda River near Madjarovo, 17-23.05.2004, S. Beshkov & H. Beck
leg., in coll. S. Beshkov, 1 , genitalia checked.
Evergestis segetalis (Herrich-Schäffer, 1851)
Chorbadjiysko Village, 2.6.1969, 2 †† (TULESCHKOV & SLIVOV, 1975b: 178);
Kardjali, 3.7.1968, 1 , 2 ‡‡ (TULESCHKOV & SLIVOV, 1975b: 178).
Evergestis serratalis (Staudinger, 1871)
Studen Kladenets Village; Momina Skala Village; Zhaltichalskoto Dere Valley near
Meden Buk Village; Belopolyane Village.
Hellula undalis (Fabricius, 1781)
Zhaltichalskoto Dere Valley near Meden Buk Village; Belopolyane Village.
?Udea numeralis (Hübner, 1796)
The bridge on Arda River near Madjarovo, 17-23.05.2004, S. Beshkov & H. Beck
leg., in coll. S. Beshkov, 1 , genitalia checked. This species seems to be new for Bulgaria,
if it is correctly identified. It is not included as present in Bulgaria in KARSHOLT &
RAZOWSKI (1996: 192).
Insecta: Lepidoptera 559
Beron page 559
Udea ferrugalis (Hübner, 1796)
Near the Zlatnata Yama Cave near Kremen Village, Djebel District, 280m; Between Shiroko
Pole and Sedlovina villages, Kardjali District; Studen Kladenets Dam, Sredna Arda Railway
Station; Studen Kladenetz Village; Momina Skala Hut near Madjarovo; The bridge on Arda River
near Madjarovo; Zhaltichalskoto Dere Valley near Meden Buk Village; Belopolyane Village.
Paracorsia repandalis ([Denis & Schiffermüller], 1775)
Studen Kladenets Village; Madjarovo, 11.8.2003, at a light trap, Peter Davey leg.
Loxostege aeruginalis (Hübner, 1796)
Kardjali, 23.6.1969, 2 ††, 2 ‡‡ (TULESCHKOV & SLIVOV, 1975b: 179); The
border post near Avren Village, 21.6.1969, 1 , 1 (TULESCHKOV & SLIVOV, 1975b:
179); Dabovets Village, 19.6.1969, 1 (TULESCHKOV & SLIVOV, 1975b: 179).
Margaritta sticticalis (Linnaeus, 1761)
Harmanli; Madjarovo.
Ecpyrrhorroe rubiginalis (Hübner, 1796)
Studen Kladenets Village; Belopolyane Village, Ivaylovgrad District.
Pyrausta cingulata (Linnaeus, 1758)
Above the bridge on Arda River near Madjarovo.
Pyrausta despicata (Scopoli, 1763) (=cespitalis ([Denis & Schiffermüller], 1775)
Madjarovo, 07.8.2003, at a light trap, Peter Davey leg.
Pyrausta purpuralis (Linnaeus, 1758)
Madjarovo, 06.8.2003, at a light trap, Peter Davey leg.
Pyrausta rectefascialis (Toll, 1936)
Belopolyane Village.
Pyrausta sanguinalis (Linnaeus, 1767) (=sanguinalsi, incorrect subsequent spelling)
The bridge on Arda River near Madjarovo; Zhaltichalskoto Dere Valley near Meden
Buk Village.
Pyrausta aurata (Scopoli, 1763)
The crossroad to Manastir Village from the road Haskovo-Kardjali; Belopolyane Village.
Sitochroa verticalis (Linnaeus, 1758)
Madjarovo, 07.8.2003, 08.8.2003, 11.8.2003, at a light trap, Peter Davey leg.; the
Bridge on Arda River near Madjarovo, 09.8.2003, at a light trap, Peter Davey leg.
Sclerocona acutella (Eversmann, 1842)
The Bridge on Arda River near Madjarovo.
560 S. BESHKOV, M. LANGOUROV
Beron page 560
Ostrinia nubilalis (Hübner, 1796)
Madjarovo; Belopolyane Village.
Ebulea crocealis (Hübner, 1796)
Kardjali, 23.6.1969, 1 (TULESCHKOV & SLIVOV, 1975b: 180); The border post
near Avren Village, 21.6.1969, 2 †† (TULESCHKOV & SLIVOV, 1975b: 180).
Anania verbascalis ([Denis & Schiffermüller], 1775)
Siv Kladenets Village, Ivaylovgrad district, 04.5.2003, S. Beshkov & V. Gashtarov
leg., 1 at day time.
Eurrhyparia hortulata (Linnaeus, 1758)
Podrumche Village, 21.6.1969, 2 †† (TULESCHKOV & SLIVOV, 1975b: 179);
Popsko Village, 22.7.1968, 1 , 1 (TULESCHKOV & SLIVOV, 1975b: 179).
Paratalanta pandalis (Hübner, 1825)
Belite Brezi Hut near Ardino.
Pleuroptya ruralis (Scololi, 1763)
Djebelska Reka river below Tarnovtsi Village, Djebel District, 380m alt., 21.10.2003, S.
Beshkov, P. Beron & B. Petrov leg. single specimen; Near the Zlatnata Yama Cave near Kremen
Village, Djebel district, 280m, 22.10.2003, S. Beshkov, P. Beron & B. Petrov leg. 2 ‡‡ specimens;
Kardjali, 23.7.1968, 2 †† (TULESCHKOV & SLIVOV, 1975b: 179); Studen Kladenets Village;
Zhaltichalskoto Dere Valley near Meden Buk Village; Mandritsa Village, 19.6.1969, 1 , 2 ‡‡
(TULESCHKOV & SLIVOV, 1975b: 179); Belopolyane Village, Ivaylovgrad District.
Mecina f lavalis ([Denis & Schiffermüller], 1775)
The border post near Avren Village, 21.6.1969, 1 (TULESCHKOV & SLIVOV, 1975b: 179).
Mecyna trinalis ([Denis & Schiffermüller], 1775)
Kardjali, 23.7.1968, 2 ††, 3 ‡‡ (TULESCHKOV & SLIVOV, 1975b: 179); Podru-
miche Village, 21.6.1969, 1 , 1 (TULESCHKOV & SLIVOV, 1975b: 179).
Diasemia reticularis (Linnaeus, 1761) (=litterata Scopoli, 1763)
Belite Brezi Hut near Ardino, 1000 m.
Palpita unionalis (Hübner, 1796)
Studen Kladenets Village; Momina Skala Hut; Zhaltichalskoto Dere Valley near
Meden Buk Village; Belopolyane Village.
Palpita testalis (Fabricius, 1794) (=incoloralis (Guenée, 1854)
Momina Skala Hut near Madjarovo.
Amaurophanes stigmosalis (Herrich-Schäffer, 1848)
Mandritsa Village, 19.6.1969, 3 ††, 2 ‡‡ (TULESCHKOV & SLIVOV, 1975b: 178).
Insecta: Lepidoptera 561
Beron page 561
Dolicharthria punctalis ([Denis & Schiffermüller], 1775)
Between Shiroko Pole and Sedlovina villages, Kardjali district, 260m alt., 20.10.2003,
S. Beshkov, P. Beron & B. Petrov leg. 1 .
Antigastra catalaunalis (Duponchel, 1833)
Zhaltichalskoto Dere Valley near Meden Buk Village; Belopolyane Village.
Metasia suppandalis (Hübner, 1823)
Svirachi Village, 19.6.1969, 4 ††, 1 (TULESCHKOV & SLIVOV, 1975b: 178);
Mandritsa Village, 19.6.1969, 2 ††, 1 (TULESCHKOV & SLIVOV, 1975b: 178).
Nomophila noctuella ([Denis & Schiffermüller], 1775)
Near the Zlatnata Yama Cave near Kremen Village, Djebel district, 280m; Between Shiro-
ko Pole and Sedlovina villages, Kardjali district, 20.10.2003; Krumovgrad; Studen Kladenets
Dam, Sredna Arda Railway Station; Studen Kladenetz Village, 21.9., 51 ‡‡, 17 ††, S. Beshkov
& V. Tschikolowez leg.; between Kulich Village and the bridge on Arda River near Kotlari
Village; Momina Skala Hut near Madjarovo, 22.IX. - more than 60 specimens, S. Beshkov &
V. Tschikolowez leg.; Madjarovo; the bridge on Arda River near Madjarovo; Zhaltichalskoto
Dere Valley near Meden Buk Village; Siv Kladenets Village; Belopolyane Village.
LASIOCAMPIDAE
Poecilocampa populi (Linnaeus, 1758)
Near Lebed Village, Djebel District, 11.12.2000, S. Beshkov, M. Langourov & B.
Petrov leg. on a lamp, 1 .
Trichiura crataegi (Linnaeus, 1758)
Djebelska Reka river below Tarnovtsi Village, Djebel district, 380m; Kardjali; Studen
Kladenets Dam, OSO Sport Center between Kaloyantsi and Gnyazdovo villages; Studen
Kladenets Village; the bridge on Arda River near Madjarovo; Belopolyane Village; Siv
Kladenets Village, Ivaylovgrad District.
Trichiura verenae Witt, 1981
Djebelska Reka river below Tarnovtsi Village, Djebel district, 380m; Momchilgrad;
between Momchilgrad and Raven Village; between Konush and Trakiets villages; near
Egrek Village; Studen Kladenets Dam, Sredna Arda Railway Station; Studen Kladenets
Village; Svezhest Hut near Krumovgrad; the bridge on Byala Reka River near Meden Buk
Village; Siv Kladenets Village, Ivaylovgrad District; Odrintsi Village. Late autumn species,
it flies in October-November. It is abundand in some places. In Bulgaria this species is
known from very few localities outside the Eastern Rhodopes.
Eriogaster rimicola ([Denis & Schiffermüller], 1775)
Near the Zlatnata Yama Cave near Kremen Village, Djebel District, 280m; Near
Kirkovo Village; Orlitsa Village; Kremen Village, Makaza District; Djebelska Reka river
below Tarnovtsi Village, Djebel District, 380m alt.; Sredna Arda Railway Station; OSO
Sport Center between Kaloyantsi and Gnyazdovo villages; near Egrek Village; Studen
562 S. BESHKOV, M. LANGOUROV
Beron page 562
Kladenets Village; Svezhest Hut near Krumovgrad; the Iron Bridge on Arda River near
Dolno Cherkovishte; Momina Skala Hut; Ivaylovgrad Dam, Arda Hut near Dabovets
Village; the bridge on Byala Reka River near Meden Buk Village; Odrintsi Village; Siv
Kladenets Village, Ivaylovgrad District.
Eriogaster catax (Linnaeus, 1758)
Near the Zlatnata Yama Cave near Kremen Village, Djebel District, 280m, 22.10.2003,
S. Beshkov, P. Beron & B. Petrov leg. single specimen. A strictly protected species from
appendix II. of the Bern Convention. A species from Appendix IV of the EEC Habitat
Directive (92/43). Species from the 2000 IUCN Red List of Threatened Species.
Malacosoma neustria (Linnaeus, 1758)
Studen Kladenets Dam, Sredna Arda Railway Station; Studen Kladenets Village; the
Iron Bridge on Arda River near Dolno Cherkovishte; Momina Skala Hut near Madjaro-
vo; Harmanli; Ivaylovgrad Dam, Arda Hut near Dabovets Village; Siv Kladenets Village.
Malacosoma castrensis (Linnaeus, 1758)
Studen Kladenets Dam, Sredna Arda Railway Station; the Iron Bridge on Arda River
near Dolno Cherkovishte; Momina Skala Hut; Ivaylovgrad Dam, Arda Hut near Dabovets
Village; Dolno Lukovo Village; Siv Kladenets Village; Odrintsi Village.
Lasiocampa trifolii ([Denis & Schiffermüller], 1775)
Lebed Village, Djebel District; Studen Kladenets Dam, Sredna Arda Railway Station;
Studen Kladenets Dam, Kroyatsi Hunting Farm near Nanovitsa Village; Studen Klad-
enets Village; Momina Skala Hut; the bridge on Arda River near Madjarovo; Borislavtsi
Village; Harmanli; Plevun Village; Ivaylovgrad Dam, Arda Hut near Dabovets Village;
Meden Buk Village; Belopolyane Village; Siv Kladenets Village.
Lasiocampa grandis (Rogenhofer, 1891)
The Iron Bridge on Arda River near Dolno Cherkovishte. Found only once in the
first half of August. As far as the present authors know, the praeimaginal stages have never
been illustratad before. Here they are illustrated for the first time (Figs 1-2.). In captivity the
larvae can be fed with Crataegus.
Macrothylacia rubi (Linnaeus, 1758)
Belite Brezi Hut near Ardino; Dishlik-Dere Valley near Oreshari Village; Ivaylovgrad
Dam, Arda Hut near Dabovets Village.
Dendrolimus pini (Linnaeus, 1758)
Belite Brezi Hut near Ardino; the Iron Bridge on Arda River near Dolno Cherko-
vishte.
Phyllodesma ilicifolia (Linnaeus, 1758)
Kroyatsi Hunting Farm near Nanovitsa Village; above the bridge on Arda River near
Madjarovo. Species from the 2000 IUCN Red List of Threatened Species.
Insecta: Lepidoptera 563
Beron page 563
Figs 1-2. Lasiocampa grandis (Rogenhofer, 1891). The Iron Bridge on Arda River near
Dolno Cherkovishte Village, larvae, ex ovo.
564 S. BESHKOV, M. LANGOUROV
Beron page 564
Phyllodesma tremulifolia (Hübner, [1810])
Belite Brezi Hut near Ardino; Studen Kladenets Dam, Sredna Arda Railway Station;
Studen Kladenets Dam, Kroyatsi Hunting Farm near Nanovitsa Village; Studen Klad-
enets Village; Krumovgrad; the Iron Bridge on Arda River near Dolno Cherkovishte;
Momina Skala Hut near Madjarovo; Madjarovo, 08.8.2003, at light trap, Peter Davey leg.
(as Phyllodesma kermesifolia). The flight period suggests that the species must be P. tremulifil-
ia, not P. ilicifolia; The bridge on Arda River near Madjarovo; Ivaylovgrad Dam, Arda
Hut near Dabovets Village; “Petko Bair” near Armira, Ivaylovgrad District; Mandritsa
Village; Siv Kladenets Village, Ivaylovgrad District. In two generations.
Gastropacha quercifolia (Linnaeus, 1758)
Studen Kladenets Dam, Kroyatsi Hunting Farm near Nanovitsa Village; Studen
Kladenets Village; Momina Skala Hut near Madjarovo; Ivaylovgrad Dam, Arda Hut near
Dabovets Village; Siv Kladenets Village, Ivaylovgrad District.
Odonestis pruni (Linnaeus, 1758)
Studen Kladenets Dam, Sredna Arda Railway Station; near Nanovitsa Village; Studen
Kladenets Dam, Kroyatsi Hunting Farm near Nanovitsa Village; Studen Kladenets Vil-
lage; Dishlik-Dere Valley near Oreshari Village; The Iron Bridge on Arda River near
Dolno Cherkovishte; between Momina Skala Hut and Kotlari Village; Momina Skala
Hut near Madjarovo; Madjarovo; The bridge on Arda river near Madjarovo; above the
bridge on Arda River near Madjarovo; 2.5 km from the bridge on Arda River near Mad-
jarovo to Borislavtsi Village; 4 km from the bridge on Arda River near Madjarovo to
Borislavtsi; Ivaylovgrad Dam, Arda Hut near Dabovets Village; Dolno Lukovo Village;
Siv Kladenets Village.
SATURNIIDAE
Saturnia pyri ([Denis & Schiffermüller], 1775)
Studen Kladenets Dam, Sredna Arda Railway Station; near Bivolene Village; Studen
Kladenets Village; the bridge on Arda River near Madjarovo; above the bridge on Arda
River near Madjarovo; Harmanli; Ivaylovgrad Dam, Arda Hut near Dabovets; the bridge
on Byala Reka River near Meden Buk Village; “Ladjata” near Ivaylovgrad; Mandritsa
Village; Belopolyane Village.
Saturnia pavonia (Linnaeus, 1758)
Studen Kladenets Dam, Sredna Arda Railway Station; Studen Kladenets Dam, OSO
Sport Center between Kaloyantsi and Gnyazdovo villages; Studen Kladenets Village; Ivay-
lovgrad Dam, Arda Hut near Dabovets Village; Svirachi Village, Ivaylovgrad district;
“Petko Bair” near Armira, Ivaylovgrad District.
Saturnia spini ([Denis & Schiffermüller], 1775)
Studen Kladenets Dam, Sredna Arda Railway Station; Studen Kladenets Dam, Kroyatsi
Hunting Farm near Nanovitsa Village; Studen Kladenets Village; Momina Skala
Hut near Madjarovo. Definitely local and rare species, in the near past considered extinct
in Bulgaria. Flies in early spring and comes to artificial light.
Insecta: Lepidoptera 565
Beron page 565
Perisomena caecigena (Kupido, 1825)
Near the Zlatnata Yama Cave near Kremen Village, Djebel District, 280m; Djebelska
Reka River below Tarnovtsi Village, Djebel District, 380m alt.; Kardjali; Momchilgrad;
Perperek Village; Between Shiroko Pole and Sedlovina villages, Kardjali District; Studen
Kladenets Dam, Kaloyantsi Village; Studen Kladenets Dam, Sredna Arda Railway Station;
Studen Kladenets Dam, OSO Sport Center between Kaloyantsi and Gnyazdovo villages;
Studen Kladenets Village; Svezhest Hut near Krumovgrad; The Iron Bridge on Arda River
near Dolno Cherkovishte; Momina Skala Hut; Madjarovo; The bridge on Arda River
near Madjarovo; between Lyubimets and Harmanli; the bridge on Byala Reka River near
Meden Buk Village; Arda Hut near Dabovets Village; Odrintsi Village; Siv Kladenets
Village, Ivaylovgrad District; Belopolyane Village. In most of the localities in oak forests
or near them it is very abundand in October. Thi s i s t he only n o c turnal L e p i dopt e ra
species, protected by law in Bulgaria. To have been considered a rare species in the past was
probably due to the late flight period (end of September-November).
LEMONIIDAE
Lemonia balcanica (Herrich-Schäffer, 1847)
Near the Zlatnata Yama Cave near Kremen Village, Djebel District; Studen Kladenets
Dam, Sredna Arda Railway Station; between Momchilgrad and Raven Village; Studen Klad-
enets Dam, OSO Sport Center between Kaloyantsi and Gnyazdovo villages; The crossroad
to Haskovo near Knizhovnik Village; Studen Kladenets Village; The Iron Bridge on Arda
River near Dolno Cherkovishte; Momina Skala Hut near Madjarovo; The bridge on Arda
River near Madjarovo; between Lyubimets and Harmanli; the bridge on Byala Reka River
near Meden Buk Village; Siv Kladenets Village, Ivaylovgrad District; Belopolyane Village.
SPHINGIDAE
Marumba quercus ([Denis & Schiffermüller], 1775)
Studen Kladenets Dam, Sredna Arda Railway Station; near Nanovitsa Village; Studen
Kladenets Village; Momina Skala Hut near Madjarovo; The bridge on Arda River near
Madjarovo; above the bridge on Arda River near Madjarovo; 2.5 km from the bridge on
Arda River near Madjarovo to Borislavtsi Village; Arda Hut near Dabovets Village; Siv
Kladenets Village.
Mimas tiliae (Linnaeus, 1758)
Belite Brezi Hut near Ardino; near Bivolene Village; Ayda Hut near Haskovo; Studen
Kladenets Village; The Iron Bridge on Arda River near Dolno Cherkovishte; the bridge on
Arda River near Madjarovo; above the bridge on Arda River near Madjarovo; “Petko
Bair” near Armira, Ivaylovgrad District; Dolno Lukovo Village; the bridge on Byala Reka
River near Meden Buk Village; Siv Kladenets Village.
Smerinthus ocellata (Linnaeus, 1758)
Studen Kladenets Village; Momina Skala Hut; The bridge on Arda River near Mad-
jarovo; above the bridge on Arda River near Madjarovo; Arda Hut near Dabovets Village;
the bridge on Byala Reka River near Meden Buk Village; Dolno Lukovo Village; “Petko
Bair” near Armira, Ivaylovgrad District; Siv Kladenets Village.
566 S. BESHKOV, M. LANGOUROV
Beron page 566
Laothoe populi (Linnaeus, 1758)
Belite Brezi Hut near Ardino; Studen Kladenets Village; above the bridge on Arda
River near Madjarovo; Arda Hut near Dabovets Village; Mandritsa Village; Siv Kladenets
Village; Odrintsi Village.
Agrius convolvuli (Linnaeus, 1758)
Belite Brezi Hut near Ardino; Near the Zlatnata Yama Cave near Kremen Village,
Djebel District, 280m, 22.10.2003; Djebelska Reka river below Tarnovtsi Village, Djebel
District, 380m alt., 21.10.2003; Studen Kladenets Dam, Kroyatsi Hunting Farm near Nano-
vitsa Village; Studen Kladenets Village; Momina Skala Hut near Madjarovo; Madjarovo;
The bridge on Arda River near Madjarovo; Arda Hut near Dabovets; Siv Kladenets Village.
Acherontia atropos (Linnaeus, 1758)
Momina Skala Hut near Madjarovo; The bridge on Arda River near Madjarovo.
Sphinx ligustri (Linnaeus, 1758)
Studen Kladenets Dam, Sredna Arda Railway Station; Studen Kladenets Dam, Kroyatsi
Hunting Farm near Nanovitsa Village; near Nanovitsa Village; Studen Kladenets Village; The
Iron Bridge on Arda River near Dolno Cherkovishte; Dishlik-Dere Valley near Oreshari Village;
Momina Skala Hut near Madjarovo; Arda Hut near Dabovets Village; Siv Kladenets Village.
Hyloicus pinastri (Linnaeus, 1758)
Belite Brezi Hut near Ardino; near Nanovitsa Village; above Dishlik-Dere Valley
near Oreshari Village; Strandjevo Village; Arda Hut near Dabovets Village; Dolno Lukovo
Village; Mandritsa Village.
Kentrochrysalis elegans (A. Bang-Haas, 1912)
Studen Kladenets Village; Arda Hut near Dabovets Village; Mandritsa Village. This
species has at least two generations and it is flies from April to August. Studen Kladenets
Village seems to be the westernmost point of the range of the species, previously known in
Bulgaria from the Southern Black Sea Coast and Sakar Mountain.
Macroglossum stellatarum (Linnaeus, 1758)
Djebelska Reka river below Tarnovtsi Village, Djebel District, 380m alt.; 22.10.2003;
Ostrovitsa Village; Lisitsite Village; Studen Kladenets Dam, Shiroko Pole Village; Sredna
Arda Railway Station; Studen Kladenets Dam, Kaloyantsi Village; Studen Kladenets Vil-
lage, Krumovitsa River; The Iron Bridge on Arda River near Dolno Cherkovishte; Dish-
lik-Dere Valley between Kulich Village, Potochnitsa District and Kotlari Village; Kotlari
Village; Madjarovo; Byal Gradets Village; between Zhelezino and Gugutka; Zhaltichalsko-
to Dere Valley near Meden Buk Village; Arda Hut near Dabovets Village; Siv Kladenets
Village, Ivaylovgrad District; Belopolyane Village.
Proserpinis proserpina (Pallas, 1772)
Belite Brezi Hut near Ardino; Arda Hut near Dabovets Village; the Iron Bridge on
Arda River near Dolno Cherkovishte; Dishlik-Dere Valley near Oreshari Village. A strict-
Insecta: Lepidoptera 567
Beron page 567
ly protected species from Appendix II of the Bern Convention. A species from Appendix
IV of the EEC Habitat Directive (92/43). Species from the 2000 IUCN Red List of
Threatened Species.
Rethera komarovi drilon Rebel & Zerny, 1931
Studen Kladenets Village; Dishlik-Dere Valley near Oreshari Village; the bridge on
Arda River near Madjarovo; above the bridge on Arda River near Madjarovo; Mandritsa
Village. Flies from the beginning of April (10.4.) until late May. Before these recent find-
ings in the Eastern Rhodopes, Rethera komarovi was known in Bulgaria from Kresna
Gorge and Slavyanska Mts only.
Hyles euphorbiae (Linnaeus, 1758)
Studen Kladenets Dam, Sredna Arda Railway Station; Studen Kladenets Village; 5
km north of Djanka Village; Dishlik-Dere Valley near Oreshari Village; the Iron Bridge
on Arda River near Dolno Cherkovishte; Momina Skala Hut near Madjarovo; Madjaro-
vo; the bridge on Arda River near Madjarovo; 4 km from the bridge on Arda River near
Madjarovo to Borislavtsi; the bridge on Byala Reka River near Meden Buk Village; Be-
lopolyane Village.
Hyles nicaea (Prunner, 1798) ?
Madjarovo, 11.8.2003, at a light trap, Peter Davey leg. The present authors are not
confident about the correct identification. It is very possible that the specimen/s of Peter
Davey are in fact large individual/s of Hyles euphorbiae. H. nicaea seems to be an extemely
rare species in Bulgaria, and the previous reports of this species for Bulgaria might also be
due to misidentification.
Hyles livornica (Esper, 1780)
Near Nanovitsa Village; Studen Kladenets Village; above the bridge on Arda River
near Madjarovo; Arda Hut near Dabovets Village; Dolno Lukovo Village.
Deilephila elpenor (Linnaeus, 1758)
Studen Kladenets Dam, Sredna Arda Railway Station; “Petko Bair” near Armira,
Ivaylovgrad District.
Deilephila porcellus (Linnaeus, 1758)
Belite Brezi Hut near Ardino; Studen Kladenets Dam, Sredna Arda Railway Station;
Dishlik-Dere Valley near Oreshari Village; the Iron Bridge on Arda River near Dolno Cherk-
ovishte; Arda Hut near Dabovets Village; Dolno Lukovo Village; Siv Kladenets Village.
Theretra alecto cretica Boisduval, 1827
The districts of Kardjali, without exact locality.
HESPERIIDAE
Thymelicus lineola (Ochsenheimer, 1808)
The bridge on Arda River near Madjarovo; Harmanli.
568 S. BESHKOV, M. LANGOUROV
Beron page 568
Thymelicus sylvestris (Poda, 1761)
Studen Kladenets Dam, Sredna Arda Railway Station; Studen Kladenets Dam, Kroyatsi
Hunting Farm near Nanovitsa Village; Studen Kladenets Village; the bridge on Arda River near
Madjarovo; Harmanli; Kamilski Dol Village; near Zhelezino Village; Byal Gradets Village.
Thymelicus acteon (Rottemburg, 1775)
Studen Kladenets Dam, Sredna Arda Railway Station; Studen Kladenets Village; the
Iron Bridge on Arda River near Dolno Cherkovishte; above Dishlik-Dere valley; between
Zhelezino and Gugutka.
Hesperia comma (Linnaeus, 1758)
Madjarovo, 09.8.2003, Peter Davey leg. Very possibly this report concerns Ochlodes
venatus, which is abundand in certain places within that region.
Ochlodes venatus (Bremer & Grey, 1853)
Studen Kladenets Village; the Iron Bridge on Arda River near Dolno Cherkovishte;
Momina Skala Hut near Madjarovo; between Zhelezino and Gugutka villages.
Erynnis tages (Linnaeus, 1758)
Studen Kladenets Dam, Sredna Arda Railway Station; Studen Kladenets Dam, Kroyatsi
Hunting Farm near Nanovitsa Village; Studen Kladenets Village; The Iron Bridge on
Arda River near Dolno Cherkovishte; Dishlik-Dere Valley between Dolno Cherkovishte
and Oreshari villages; Kulich Village, Potochnitsa District; Kotlari Village; Momina Ska-
la Hut near Madjarovo; Ivaylovgrad Dam, Arda Hut near Dabovets Village; Byal Gradets
Village; Dupkata Hill near Ivaylovgrad.
Carcharodus alceae (Esper, [1870])
Studen Kladenets Dam, Zvezdelina Village; Studen Kladenets Dam, Sredna Arda Rail-
way Station; Studen Kladenets Dam, Kroyatsi Hunting Farm near Nanovitsa Village; Studen
Kladenets Village; Krumovitsa River; Kotlari Village; Momina Skala Hut near Madjarovo;
Madjarovo; Harmanli; between Zhelezino and Gugutka villages; Siv Kladenets Village.
Carcharodus lavatherae (Esper, [1783])
Studen Kladenets Dam, Sredna Arda Railway Station; near Rabovo Village.
Carcharodus flocciferus (Zeller, 1847)
Dolno Lukovo Village; Siv Kladenets Village, Ivaylovgrad District.
Carcharodus orientalis Reverdin, 1913
Studen Kladenets Dam, Sredna Arda Railway Station; Studen Kladenets Village; Ivay-
lovgrad Dam, Arda Hut near Dabovets; Zhelezari Village.
Spialia orbifer (Hübner, [1823])
Komuniga Village; Studen Kladenets Dam, Sredna Arda Railway Station; Studen
Kladenets Village; Krumovitsa River; The Iron Bridge on Arda River near Dolno Cherk-
Insecta: Lepidoptera 569
Beron page 569
ovishte; above above Dishlik-Dere Valley; Momina Skala Hut near Madjarovo; the bridge
on Arda River near Madjarovo; Dolno Lukovo Village.
Pyrgus sidae (Esper, [1784])
Studen Kladenets Dam, Sredna Arda Railway Station; Studen Kladenets Dam, “Kroyat-
si” Hunting farm near Nanovitsa Village; Studen Kladenets Dam, Darets; the bridge on
the Arda River near Madjarovo; The bridge on Byala Reka River near Meden Buk Village;
Kamilski Dol Village.
Pyrgus malvae (Linnaeus, 1758)
Studen Kladenets Dam, Sredna Arda Railway Station; Garvanovo Village; Studen
Kladenets Village; above Dishlik-Dere Valley near Oreshari Village; the bridge on Arda
River near Madjarovo; Arda Hut near Dabovets Village; Kamilski Dol Village.
Pyrgus serratulae (Rambur, [1839])
Kotlari Village, 11.8.2003, Peter Davey leg.
Pyrgus cinarae (Rambur, [1839])
Near Rabovo Village; Between Momina Skala Hut and Madjarovo; on the road from
Madjarovo to Borislavtsi Village, 2.5 km from the bridge on Arda River near Madjarovo.
Pyrgus armoricanus (Ch. Oberthür, 1910)
Djebelska Reka river below Tarnovtsi Village, Djebel district, 380m; Studen Klad-
enets Dam, Sredna Arda Railway Station; Studen Kladenets Village; Rabovo Village; Gar-
vanovo Village; Arda Hut near Dabovets Village. The report on Pyrgus carlinae from Kot-
lari Village, 11.8.2003, Peter Davey leg., probably concerns P. armoricanus.
Pyrgus alveus (Hübner, [1803]) ?
Madjarovo, 09.8.2003, Peter Davey leg.; Krumovitsa River 08.8.2003, Peter Davey
leg. This report was very likely the result of a misidentification with another Pyrgus species,
for example P. armoricanus, which is a common species in the Eastern Rhodopes.
PAPILIONIDAE
Zerynthia polyxena ([Denis & Schiffermüller], 1775)
Studen Kladenets Village; Dishlik-Dere Valley between Kulich Village near Potoch-
nitsa; Kotlari Village; Madjarovo; the bridge on Arda River near Madjarovo; Arda Hut
near Dabovets Village; Kamilski Dol Village; Svirachi Village, Ivaylovgrad District; Siv
Kladenets Village. A strictly protected species from Appendix II of the Bern Convention. A
species from Appendix IV of the EEC Habitat Directive (92/43).
Zerynthia cerisy ferdinandi Stichel, 1907
Near Plazishte Village; Kardjali; Haskovo; Studen Kladenets Dam, Sredna Arda Rail-
way Station; Studen Kladenets Dam, Kroyatsi Hunting Farm near Nanovitsa Village; Studen
Kladenets Village; near Rabovo Village; the Iron Bridge on Arda River near Dolno Cherko-
vishte; Dolno Cherkovishte Village; 2 km from the bridge on Arda River near Madjarovo to
570 S. BESHKOV, M. LANGOUROV
Beron page 570
Borislavtsi Village; 4.4 km on Arda river from the bridge near Madjarovo to Borislavtsi
Village; Arda Hut near Dabovets Village; Meden Buk Village; Siv Kladenets Village.
Parnassius mnemosyne (Linnaeus, 1758)
Haskovo; Studen Kladenets Village; Madjarovo; the bridge on Arda River near Mad-
jarovo. A strictly protected species from Appendix II of the Bern Convention. A species
from Appendix IV of the EEC Habitat Directive (92/43).
Iphiclides podalirius (Linnaeus, 1758)
Kardjali; Haskovo; Sredna Arda Railway Station; Studen Kladenets Dam, Kroyatsi
Hunting Farm near Nanovitsa Village; Studen Kladenets Village; the Iron Bridge on Arda
River near Dolno Cherkovishte; Kotlari Village; Momina Skala Hut near Madjarovo;
Madjarovo; the bridge on Arda River near Madjarovo; Malo Gradishte Village; Arda Hut
near Dabovets Village; Kaleto above Mezek Village; Dupkata Hill near Ivaylovgrad; “Lad-
jata” near Ivaylovgrad; Kamilski Dol Village; Mandritsa Village; Siv Kladenets Village.
Papilio machaon Linnaeus, 1758
Studen Kladenets Village; Krumovitza River; the Iron Bridge on Arda River near
Dolno Cherkovishte; between Kulich Village, Potochnitsa District and the bridge on
Arda River near Dolno Cherkovishte Village; Kotlari Village; Madjarovo; Ivaylovgrad
Dam, Borislavtsi Village; Harmanli, Lyubimets; Arda Hut near Dabovets Village; Dupka-
ta Hill near Ivaylovgrad; Siv Kladenets Village.
PIERIDAE
Leptidea sinapis (Linnaeus, 1758)
Near Plazishte Village; Sredna Arda Railway Station; Garvanovo Village, Haskovo
District; Kroyatsi Hunting Farm near Nanovitsa Village; Studen Kladenets Village; Kru-
movgrad; Dishlik-Dere Valley between Kulich Village, Potochnitsa District and the Iron
Bridge on Arda River near Dolno Cherkovishte Village; the Iron Bridge on Arda River
near Dolno Cherkovishte; Oreshenskoto-Dere Valley near Odrintsi Village; Harmanli;
Kamilski Dol Village; between Zhelezino and Gugutka villages; Mandritsa Village. A com-
plex of two closely related species (Leptidea sinapis and L. reali Reissinger, 1990). For cor-
rect identification additional material is needed.
Leptidea duponcheli (Staudinger, 1871)
Studen Kladenets Village; Krumovitsa River; Kotlari Village; Madjarovo; Harmanli,
Svirachi Village, Ivaylovgrad District.
Colias erate (Esper, [1805])
Sredna Arda Railway Station; Garvanovo Village; Studen Kladenets Village; Kulich
Village near Potochnitsa; the Iron Bridge on Arda River near Dolno Cherkovishte; Mad-
jarovo; Arda Hut near Dabovets Village; Mandritsa Village.
Colias alfacariensis Ribbe, 1905
Harmanli.
Insecta: Lepidoptera 571
Beron page 571
Colias crocea (Frourcroy, 1785)
Near Sinchets Village; Ridino vilage; Djebelska Reka river below Tarnovtsi Village,
Djebel District, 380m alt.; 22.10.2003; Near Plazishte Village; Momchilgrad; Studen Klad-
enets Dam, Zvezdelina Village; Studen Kladenets Dam, Sredna Arda Railway Station;
Gorna Krepost Village; Perperek Village; Studen Kladenets Dam, Kaloyantsi Village; Studen
Kladenets Dam, Kroyatsi Hunting Farm near Nanovitsa Village; Studen Kladenets Vil-
lage; Krumovitsa River; Krumovgrad; 3 km SE of Krumovgrad on the road to Avren
Village; Southern of Madjari Village; the Iron Bridge on Arda River near Dolno Cherko-
vishte; Dishlik-Dere Valley between Kulich Village, Potochnitsa District and the Iron
Bridge on Arda River near Dolno Cherkovishte Village; Kotlari Village; Momina Skala
Hut near Madjarovo; Madjarovo; the bridge on Arda River near Madjarovo; near Topolo-
vo Village; Ivaylovgrad Dam, Borislavtsi Village; Arda Hut near Dabovets Village; Zhelezari
Village; Byal Gradets Village; Meden Buk Village; Zhaltichalskoto Dere Valley near Meden
Buk Village; Dolno Lukovo Village; near gallery between Lozen to Cherna Mogila villag-
es; Lyubimets; Kamilski Dol Village; Ivaylovgrad; Dupkata Hill near Ivaylovgrad; Man-
dritsa Village; Siv Kladenets Village.
Gonepteryx rhamni (Linnaeus, 1758)
Djebelska Reka river below Tarnovtsi Village, Djebel district, 380m alt.; 22.10.2003;
Studen Kladenets Village; Kulich Village near Potochnitsa; Kotlari Village; Dishlik-Dere
Valley between Dolno Cherkovishte and Oreshari villages; Madjarovo; Arda Hut near
Dabovets Village; Harmanli, Biser Village; Siv Kladenets Village, Ivaylovgrad District.
Anthocharis cardamines (Linnaeus, 1758)
Studen Kladenets Village; Dishlik-Dere Valley between Kulich Village, Potochnitsa
District and the Iron Bridge on Arda River near Dolno Cherkovishte Village; the Iron
Bridge on Arda River near Dolno Cherkovishte; Madjarovo; Ivaylovgrad Dam, Borisla-
vtsi Village; the bridge on Arda River near Madjarovo; Arda Hut near Dabovets Village;
Harmanli; Meden Buk Village; Oreshenskoto-Dere Valley near Odrintsi Village; Siv Klad-
enets Village, Ivaylovgrad District; Dupkata Hill near Ivaylovgrad; Svirachi Village, Ivay-
lovgrad District; Mandritsa Village.
Euchloe ausonia (Hübner, [1804])
Studen Kladenets Village; Dishlik-Dere Valley between Kulich Village, Potochnit-
sa District and the Iron Bridge on Arda River near Dolno Cherkovishte Village; the
Iron Bridge on Arda River near Dolno Cherkovishte; Madjarovo; Meden Buk Village;
Dupkata Hill near Ivaylovgrad; Kamilski Dol Village; Siv Kladenets Village, Ivaylovgrad
District.
Aporia crataegi (Linnaeus, 1758)
Between Ardino and Nedelino, 2 km before the crossroad to Nedelino; Near Plazishte
Village; Studen Kladenets Dam, Sredna Arda Railway Station; the Iron Bridge on Arda
River near Dolno Cherkovishte; Madjarovo; the bridge on Arda River near Madjarovo;
Arda Hut near Dabovets Village; Meden Buk Village; Siv Kladenets Village, Ivaylovgrad
District.
572 S. BESHKOV, M. LANGOUROV
Beron page 572
Pieris brassicae (Linnaeus, 1758)
Komuniga Village; Djebelska Reka river below Tarnovtsi Village, Djebel district,
380m alt.; Kardjali; Haskovo; Studen Kladenets Dam, Shiroko Pole Village; Studen Klad-
enets Dam, Zvezdelina Village; Studen Kladenets Dam, Sredna Arda Railway Station;
Studen Kladenets Dam, Kroyatsi Hunting Farm near Nanovitsa Village; Studen Klad-
enets Village; Krumovitsa River; Krumovgrad; between Kulich Village, Potochnitsa Dis-
trict and the Iron Bridge on Arda River near Dolno Cherkovishte Village; the Iron Bridge
on Arda River near Dolno Cherkovishte Village; Kotlari Village; Bryagovets Village; Momina
Skala Hut near Madjarovo; Madjarovo; the bridge on Arda River near Madjarovo; Meden
Buk Village; Arda Hut near Dabovets Village; Kamilski Dol Village; Siv Kladenets Village,
Ivaylovgrad District.
Pieris krueperi Staudinger, 1860
Studen Kladenets Dam, Sredna Arda Railway Station; Studen Kladenets Village; near
Rabovo Village; Momina Skala Hut near Madjarovo; Madjarovo; 2.5 km from the bridge on
Arda River near Madjarovo to Borislavtsi Village. Flies from April to July in two generations.
Pieris mannii (Mayer, 1851)
Djebelska Reka river below Tarnovtsi Village, Djebel District, 380m alt.; 22.10.2003;
Studen Kladenets Dam, Sredna Arda Railway Station; Studen Kladenets Dam, Kroyatsi
Hunting Farm near Nanovitsa Village; Studen Kladenets Village; near Rabovo Village;
Kotlari Village; Momina Skala Hut near Madjarovo; Madjarovo; 2.5 km from the bridge
on Arda River near Madjarovo to Borislavtsi Village; Harmanli, Zhelezari Village; Meden
Buk Village; Kamilski Dol Village.
Pieris rapae (Linnaeus, 1758)
Kardjali; near Sinchets Village; Studen Kladenets Dam, near Sredna Arda Railway
Station; Momchilgrad; Perperek Village; Studen Kladenets Dam, Kaloyantsi Village; Hasko-
vo; Studen Kladenets Dam, Kroyatsi Hunting Farm near Nanovitsa Village; Rupata Cave
near Egrek Village; Studen Kladenets Village; near Rabovo Village; Krumovgrad; the Iron
Bridge on Arda River near Dolno Cherkovishte; Dishlik-Dere Valley between Dolno Cherk-
ovisthe and Oreshari villages; Kulich Village, Potochnitsa District; Kotlari Village; Momina
Skala Hut near Madjarovo; the bridge on Arda River near Madjarovo; Madjarovo; Arda
Hut near Dabovets Village; Zhelezari Village; Harmanli, Lyubimets; Meden Buk Village;
Zhaltichalskoto Dere Valley near Meden Buk Village; Kamilski Dol Village; Ivaylovgrad
Dam, Kochash Village; Ivaylovgrad; Dupkata Hill near Ivaylovgrad; Mandritsa Village.
Pieris ergane (Geyer, [1828])
Garvanovo Village; Studen Kladenets Village; the Iron Bridge on Arda River near
Dolno Cherkovishte; above Dishlik-Dere Valley; Madjarovo; between Madjarovo and
Borislavtzi villages.
Pieris napi (Linnaeus, 1758)
Studen Kladenets Dam, Zvezdelina Village; Studen Kladenets Dam, Sredna Arda
Railway Station; Studen Kladenets Village; Krumovitsa River; Krumovgrad; Dishlik-Dere
Insecta: Lepidoptera 573
Beron page 573
Valley between Kulich Village, Potochnitsa District and the Iron Bridge on Arda River
near Dolno Cherkovishte Village; the Iron Bridge on Arda River near Dolno Cherkovish-
te; Kotlari Village; Momina Skala Hut near Madjarovo; the bridge on Arda River near
Madjarovo; Madjarovo; Harmanli; Arda Hut near Dabovets Village; Meden Buk Village;
Kamilski Dol Village; Mandritsa Village; Siv Kladenets Village.
Pieris balkana Lorkovic, [1970]
Garvanovo Village; Momina Skala Hut ner Madjarovo; Siv Kladenets Village, Ivay-
lovgrad District. Incorrect identification seems very possible, especially for the summer
generation.
Pontia edusa (Fabricius, 1777)
Komuniga Village; 2 km before Sussam Village from Gorski Izvor; Studen Kladenets
Dam, Shiroko Pole Village; Studen Kladenets Dam, Zvezdelina Village; Studen Kladenets
Dam, Sredna Arda Railway Station; Perperek Village; near Nanovitsa Village; Studen Klad-
enets Dam, Kroyatsi Hunting Farm near Nanovitsa Village; Studen Kladenets Village;
Krumovitsa River; Krumovgrad; near Topolovo Village; the Iron Bridge on Arda River
near Dolno Cherkovishte; between Kulich Village, Potochnitsa District and the Iron
Bridge on Arda River near Dolno Cherkovishte Village; Kotlari Village; Momina Skala
Hut near Madjarovo; Madjarovo; 2.5 km from the bridge on Arda River near Madjarovo
to Borislavtsi Village; Harmanli; Arda Hut near Dabovets Village; Zhelezari Village; Ly-
ubimets; Meden Buk Village; Zhaltichalskoto Dere Valley near Meden Buk Village; Ivay-
lovgrad Dam, Kochash Village; Siv Kladenets Village, Ivaylovgrad District.
Pontia chloridice (Hübner, [1813])
Studen Kladenets Dam, Shiroko Pole Village; Studen Kladenets Dam, Sredna Arda
Railway Station; Studen Kladenets Village; Siv Kladenets Village, Ivaylovgrad District.
Flies from March to September, probably in three generations.
LYCAENIDAE
Hamearis lucina (Linnaeus, 1758)
Between Ardino and Nedelino, 2 km before the crossroad to Nedelino; The Iron
Bridge on Arda River near Dolno Cherkovishte; Kotlari Village; Momina Skala Hut near
Madjarovo.
Lycaena phlaeas (Linnaeus, 1761)
Djebelska Reka river below Tarnovtsi Village, Djebel district, 380m alt.; 22.10.2003;
Near Plazishte Village; Studen Kladenets Dam, Sredna Arda Railway Station; Gorna Krepost
Village; Studen Kladenets Dam, Kroyatsi Hunting Farm near Nanovitsa Village; near Rupa-
ta Cave near Egrek Village; Studen Kladenets Village; Krumovitsa River; Krumovgrad; South
of Madjari Village; Dishlik-Dere Valley between Kulich Village, Potochnitsa District and the
Iron Bridge on Arda River near Dolno Cherkovishte Village; The Iron Bridge on Arda River
near Dolno Cherkovishte; Kotlari Village; Momina Skala Hut near Madjarovo; Madjarovo;
The bridge on Arda River near Madjarovo; Harmanli; near Zhelezino Village; Zhelezari
Village; Arda Hut near Dabovets Village; Meden Buk Village; Lyubimets; Ivaylovgrad Dam,
574 S. BESHKOV, M. LANGOUROV
Beron page 574
Kochash Village; Kamilski Dol Village; Ivaylovgrad; Dupkata Hill near Ivaylovgrad; Kodja-
Kaya Hill near Odrintsi Village; Mandritsa Village; Siv Kladenets Village.
Lycaena dispar ([Haworth], 1803)
Studen Kladenets Village; Lyubimets. Strictly protected by Appendix II of the Bern
Convention. A species from Appendix IV of the EEC Habitat Directive (92/43).
Lycaena virgaureae (Linnaeus, 1758)
Between Ardino and Nedelino, 2 km before the crossroad to Nedelino.
Lycaena ottomana (Lefebvre, [1830])
On the road from Madjarovo to Borislavtsi Village, 2.5 km from the bridge on Arda
River near Madjarovo; Ivaylovgrad Dam, Arda Hut near Dabovets illage; between Zhelez-
ino and Gugutka villages; Siv Kladenets Village, Ivaylovgrad District. Rare and local
species in Bulgaria and Europe with stable populations in the Eastern Rhodopes, abun-
dant in wet meadows near Siv Kladenets Village at the very beginning of May. Flies from
the end of April - beginning of May to the middle of July. Species from the 2000 IUCN
Red List of Threatened Species.
Lycaena tityrus (Poda, 1761)
Near Plazishte Village; Sredna Arda Railway Station; Garvanovo Village; 2 km be-
fore Sussam Village from Gorski Izvor; near Nanovitsa Village; Studen Kladenets Village;
the Iron Bridge on Arda River near Dolno Cherkovishte; Kotlari Village; Momina Skala
Hut; Madjarovo; the bridge on Arda River near Madjarovo; near Topolovo Village; Har-
manli; Byal Gradets Village; between Zhelezino and Gugutka villages; near Zhelezino
Village; Zhelezari Village; Meden Buk Village; Zhaltichalskoto Dere Valley near Meden
Buk Village; Siv Kladenets Village.
Lycaena alciphron melibaeus (Staudinger, 1879)
Between Ardino and Nedelino, 2 km before the crossroad to Nedelino; Sredna Arda
Railway Station; the Iron Bridge on Arda River near Dolno Cherkovishte; Momina Skala
Hut; the bridge on Arda River near Madjarovo; Meden Buk Village.
Lycaena candens (Herrich-Schäffer, [1844]
Between Ardino and Nedelino, 2 km before the crossroad to Nedelino.
Lycaena thersamon (Esper, [1784])
Garvanovo Village; 2 km before Sussam Village from Gorski Izvor; Kroyatsi Hunt-
ing Farm near Nanovitsa Village; near Nanovitsa Village; Studen Kladenets Village; Kru-
movitsa River; Kotlari Vilage; the bridge on Arda River near Madjarovo; near Topolovo
Village; between Zhelezino and Gugutka villages; Siv Kladenets Village.
Thecla betulae (Linnaeus, 1758)
3 km SE of Krumovgrad on the road to Avren Village; near Madjarovo. Found only
in October and November.
Insecta: Lepidoptera 575
Beron page 575
Favonius quercus (Linnaeus, 1758)
Komuniga Village; Studen Kladenets Dam, Kroyatsi Hunting Farm near Nanovitsa
Village; Studen Kladenets Village; Southern of Madjari Village; Momina Skala Hut; the
bridge on Arda River near Madjarovo; Kotlari Village; Zhelezari Village; Zhaltichalskoto
Dere Valley near Meden Buk Village.
Callophrys rubi (Linnaeus, 1758)
Between Ardino and Nedelino, 2 km before the crossroad to Nedelino; Studen Klad-
enets Village; above Dishlik-Dere Valley near Oreshari Village; Momina Skala Hut; Mad-
jarovo; the bridge on Arda River near Madjarovo; Arda Hut; Kamilski Dol Village.
Satyrium w-album (Knoch, 1782)
Studen Kladenets Dam, Kroyatsi Hunting Farm near Nanovitsa Village; Studen
Kladenets Village.
Satyrium spini (Fabricius, 1787)
Studen Kladenets Dam, Sredna Arda Railway Station; between Zhelezino and Gu-
gutka villages; Byal Gradets Village.
Satyrium ilicis (Esper, [1789])
Near Plazishte Village; Sredna Arda Railway Station; the Iron Bridge on Arda River
near Dolno Cherkovishte; Madjarovo; the bridge on Arda River near Madjarovo; 2 km
from the bridge on Arda River near Madjarovo to Borislavtsi; Harmanli.
Satyrium acaciae (Fabricius, 1787)
Studen Kladenets Village; Harmanli.
Leptotes pirithous (Linnaeus, 1767)
Garvanovo Village; Krumovgrad; the Iron Bridge on Arda River near Dolno Cherk-
ovishte; Kotlari Village; Ivaylovgrad Dam, Kochash Village.
Taru cus balkanicus (Freyer, [1844])
Garvanovo Village; Studen Kladenets Village; Rabovo Village; Krumovitsa River;
Kotlari Village; the bridge on Arda River near Madjarovo;. Sometime and somewhere
abundant in areas with Paliurus.
Cupido decoloratus (Staudinger, 1886)
Near Plazishte Village; Komuniga Village; the Iron Bridge on Arda River near Dolno
Cherkovishte.
Celastrina argiolus (Linnaeus, 1758)
Studen Kladenets Dam, Sredna Arda Railway Station; Studen Kladenets Dam, Kroyatsi
Hunting Farm near Nanovitsa Village; Studen Kladenets Village; near Rabovo Village; Kulich
Village near Potochnitsa; the Iron Bridge on Arda River near Dolno Cherkovishte; Kotlari
Vilage; Madjarovo; Byal Gradets Village; Gorata Ridge above Malo Gradishte Village.
576 S. BESHKOV, M. LANGOUROV
Beron page 576
Pseudophilotes vicrama schiffermuelleri (Hemming, 1929)
Studen Kladenets Dam, Sredna Arda Railway Station; Studen Kladenets Village; above
Dishlik-Dere valley; Dupkata Hill near Ivaylovgrad.
Scolitantides orion (Pallas, 1771)
The bridge on Arda River near Madjarovo, a single specimen in a Malaise trap.
Glaucopsyche alexis (Poda, 1761)
Studen Kladenets Village; Dishlik-Dere Valley between Kulich Village, Potochnitsa
District and the Iron Bridge on Arda River near Dolno Cherkovishte Village; Madjarovo;
the bridge on the Arda River near Madjarovo; 4.4 km from the bridge on Arda River near
Madjarovo to Borislavtsi Village; Arda Hut near Dabovets Village; Siv Kladenets Village.
Glaucopsyche iolas (Ochsenheimer, 1816)
The Iron Bridge on Arda River near Dolno Cherkovishte.
Glaucopsyche arion (Linnaeus, 1758)
The Eastern Rhodopes (without locality); Momina Skala Hut. Strictly protected by
Appendix II of the Bern Convention. A species from Appendix IV of the EEC Habitat
Directive (92/43). Species from the 2000 IUCN Red List of Threatened Species.
Plebeiuus pylaon sephirus Friveldsky, 1835
above Dishlik-Dere valley.
Plebeius argus (Linnaeus, 1758)
Near Plazishte Village; The crossroad to Manastir Village from the road Haskovo-
Kardjali; Garvanovo Village; Studen Kladenets Dam, Kroyatsi Hunting Farm near Nano-
vitsa Village; Studen Kladenets Village; Krumovitsa River; the Iron Bridge on Arda River
near Dolno Cherkovishte; Momina Skala Hut near Madjadovo; Siv Kladenets Village.
Plebeius argyrognomon (Bergsträsser, 1779)
Krumovitsa River, 07.8.2003, Peter Davey leg.; Ivaylovgrad Dam, Arda Hut near
Dabovets Village; Byal Gradets Village.
Plebeius agestis ([Denis & Schiffermüller], 1775)
Near Karangil Cave above Shiroko Pole Village, Kardjali district; Studen Kladenets
Dam, Sredna Arda Railway Station; Studen Kladenets Dam, Kroyatsi Hunting Farm near
Nanovitsa Village; Studen Kladenets Village; near Rabovo Village; Krumovitsa river; Kru-
movgrad; the Iron Bridge on Arda River near Dolno Cherkovishte; Kotlari Village; Mo-
mina Skala Hut near Madjarovo; Madjarovo; the bridge on Arda River near Madjarovo;
2.5 km from the bridge on Arda River near Madjarovo to Borislavtsi Village; Zhelezari
Village; Byal Gradets Village; between Zhelezino and Gugutka villages; Arda Hut; Dolno
Lukovo Village; Mandritsa Village; Siv Kladenets Village.
Insecta: Lepidoptera 577
Beron page 577
Plebeius anteros (Freyer, [1838])
Studen Kladenets Dam, Sredna Arda Railway Station; Studen Kladenets Village; Kru-
movitsa River; Krumovgrad; the Iron Bridge on Arda River near Dolno Cherkovishte;
Kotlari Village; Arda Valley, Bryagovets Village; Madjarovo.
Polyommatus semiargus (Rottemburg, 1775)
Between Ardino and Nedelino, 2 km before the crossroad to Nedelino; Studen Kladenets
Dam, Kroyatsi Hunting Farm near Nanovitsa Village; Arda Hut near Dabovets Village.
Polyommatus escheri (Hübner, [1823]
Above Dishlik-Dere valley.
Polyommatus amandus (Scheider, 1792)
Meden Buk Village.
Polyommatus thersites (Cantener, [1835])
Above Dishlik-Dere valley; Madjarovo, 09.8.2003, Peter Davey leg.
Polyammatus icarus (Rottemburg, 1775)
Near Plazishte Village; Near Karangil Cave above Shiroko Pole Village, Kardjali dis-
trict; Studen Kladenets Dam, Zvezdelina Village; Studen Kladenets Dam, Sredna Arda
Railway Station; Studen Kladenets Dam, Kroyatsi Hunting Farm near Nanovitsa Village;
Studen Kladenets Village; near Rabovo Village; Southern of Madjari Village; Krumovitsa
River; the Iron Bridge on Arda River near Dolno Cherkovishte; above Dishlik-Dere Val-
ley near Oreshari Village; Kotlari Village; Momina Skala Hut near Madjarovo; Madjaro-
vo; the bridge on Arda River near Madjarovo; near Topolovo Village; Arda Hut near
Dabovets Village; Byal Gradets Village; near Zhelezino Village; Zhaltichalskoto Dere Val-
ley near Meden Buk Village; the bridge on Byala Reka River near Meden Buk Village;
Kamilski Dol Village; Lyubimets; Ivaylovgrad Dam, Kochash Village; Ivaylovgrad; Dup-
kata Hill near Ivaylovgrad; Mandritsa Village; Siv Kladenets Village.
Polyommatus daphnis ([Denis & Schiffermüller], 1775)
Studen Kladenets Dam, Kroyatsi Hunting Farm near Nanovitsa Village; Krumovit-
sa River; Kotlari Village; above Dishlik-Dere Valley; the Iron Bridge on Arda River near
Dolno Cherkovishte; Momina Skala Hut near Madjarovo.
Polyommatus bellargus (Rottemburg, 1775)
Djebelska Reka river below Tarnovtsi Village, Djebel district, 380m alt.; Studen Klad-
enets Dam, Kroyatsi Hunting Farm near Nanovitsa Village; Studen Kladenets Village;
Krumovitsa River; Krumovgrad; Kotlari Village; above Dishlik-Dere Valley; the bridge on
Arda River near Madjarovo.
Polyommatus coridon (Poda, 1761)
Komuniga Village; Studen Kladenets Village; the Iron Bridge on Arda River near
Dolno Cherkovishte; Kotlari Village; Madjarovo.
578 S. BESHKOV, M. LANGOUROV
Beron page 578
Polyommatus admetus (Esper, [1783])
Studen Kladenets Village; Krumovitsa River, 08.8.2003, Peter Davey leg.; between
Zhelezino and Gugutka villages.
NYMPHALIDAE
Libythea celtis (Laicharting, 1782)
Momina Skala Hut; Madjarovo; On the road from Madjarovo to Borislavtsi Vil-
lage, 2.5 km from the bridge on Arda River near Madjarovo; Dupkata Hill near Ivay-
lovgrad.
Kirinia roxelana (Cramer, [1777])
Komuniga Village; Sredna Arda Railway Station; Studen Kladenets Dam, Kroyatsi
Hunting Farm near Nanovitsa Village; Studen Kladenets Village; Studen Kladenets Dam,
Darets; near Rabovo Village; above Dishlik-Dere Valley; the Iron Bridge on Arda River
near Dolno Cherkovishte; Momina Skala Hut near Madjarovo; the bridge on Arda River
near Madjarovo; Zhaltichalskoto Dere Valley near Meden Buk Village; Zhelezari Village.
Pararge aegeria (Linnaeus, 1758)
Studen Kladenets Dam, Sredna Arda Railway Station; Studen Kladenets Village; Kru-
movitsa River; the Iron Bridge on Arda River near Dolno Cherkovishte; Kotlari Village;
Momina Skala Hut near Madjarovo; Madjarovo; the bridge on Arda River near Madjaro-
vo, collected at a light trap twice; Byal Gradets Village; Zhelezari Village; Arda Hut near
Dabovets Village; Meden Buk Village.
Lasiommata megera (Linnaeus, 1767)
Djebelska Reka river below Tarnovtsi Village, Djebel District, 380m alt.; 22.10.2003;
Momchilgrad; Near Karangil Cave above Shiroko Pole Village, Kardjali district; Studen
Kladenets Dam, Sredna Arda Railway Station; Studen Kladenets Dam, Kroyatsi Hunting
Farm near Nanovitsa Village; Studen Kladenets Village; Krumovitsa River; the Iron Bridge
on Arda River near Dolno Cherkovishte; Momina Skala Hut near Madjarovo; the bridge
on Arda River near Madjarovo; Meden Buk Village; Lyubimets; Ivaylovgrad; Dupkata
Hill near Ivaylovgrad; Kodja-Kaya Hill near Odrintsi Village.
Lasiommata maera (Linnaeus, 1758)
Studen Kladenets Village; the Iron Bridge on Arda River near Dolno Cherkovishte;
Kotlari Village; Madjarovo; the bridge on Arda River near Madjarovo; near Gorno Pole
Village, Madjarovo District; Kamilski Dol Village.
Coenonympha arcania (Linnaeus, 1761)
Studen Kladenets Dam, Kroyatsi Hunting Farm near Nanovitsa Village; above Dishlik-
Dere Valley; Momina Skala Hut near Madjarovo; between Zhelezino and Gugutka villages.
Coenonympha leander (Esper, [1784])
Kamilski Dol Village; Mandritsa Village; Meden Buk Village.
Insecta: Lepidoptera 579
Beron page 579
Coenonympha pamphilus (Linnaeus, 1758)
The crossroad to Manastir Village from the road Haskovo-Kardjali; 2 km before
Sussam Village from Gorski Izvor; Sredna Arda Railway Station; Studen Kladenets Dam,
Kroyatsi Hunting Farm near Nanovitsa Village; Studen Kladenets Village; Krumovitsa
River; near Topolovo Village; the Iron Bridge on Arda River near Dolno Cherkovishte;
above Dishlik-Dere Valley near Oreshari Village; Kotlari Village; Momina Skala Hut near
Madjarovo; Madjarovo; the bridge on Arda River near Madjarovo; 4.4 km from the bridge
on Arda River near Madjarovo to Borislavtsi Village; Zhelezari Village; Arda Hut near
Dabovets Village; Meden Buk Village; Kamilski Dol Village; Lyubimets; Ivaylovgrad Dam,
Kochash Village; Mandritsa Village; Siv Kladenets Village.
Pyronia tithonus (Linnaeus, 1767)
Komuniga Village; Studen Kladenets Dam, Kroyatsi Hunting Farm near Nanovitsa
Village; Studen Kladenets Village; Krumovitsa River; The Iron Bridge on Arda River near
Dolno Cherkovishte; Kotlari Village; Momina Skala Hut near Madjarovo; Krumovgrad
District (without exactly pointed locality); Arda Hut near Dabovets Village; Ivaylovgrad
Dam, Kochash Village; Siv Kladenets Village, Ivaylovgrad District.
Maniola jurtina (Linnaeus, 1758)
Between Ardino and Nedelino, 2 km before the crossroad to Nedelino; near Sinchets
Village; Near Plazishte Village; Studen Kladenets Dam, Zvezdelina Village; Studen Klad-
enets Dam, Sredna Arda Railway Station; Studen Kladenets Dam, Kroyatsi Hunting Farm
near Nanovitsa Village; Studen Kladenets Village; Studen Kladenets Dam, Darets; near
Rabovo Village; Krumovgrad; near Topolovo Village; the Iron Bridge on Arda River near
Dolno Cherkovishte; Kotlari Village; Momina Skala Hut near Madjarovo; Madjarovo;
the bridge on Arda River near Madjarovo; Zhelezari Village; Arda Hut near Dabovets
Village; Byal Gradets Village; between Zhelezino and Gugutka villages; Meden Buk Vil-
lage; Kamilski Dol Village; Ivaylovgrad Dam, Kochash Village; Siv Kladenets Village.
Hyponephele lupina (Costa, [1836])
Studen Kladenets Village, 05.07.2004, S. Beshkov leg.; near Madjarovo, August, 2003,
A. Ignatov, perss. comm.; Krumovitsa River, 08.8.2003, Peter Davey leg.
Melanargia galathea (Linnaeus, 1758)
Near Sinchets Village; 2 km before Sussam Village from Gorski Izvor; Studen Klad-
enets Dam, Sredna Arda Railway Station; Studen Kladenets Dam, Kroyatsi Hunting Farm
near Nanovitsa Village; near Nanovitsa Village; Studen Kladenets Village; Southern of Mad-
jari Village; near Topolovo Village; the Iron Bridge on Arda River near Dolno Cherkovishte;
Momina Skala Hut near Madjarovo; the bridge on Arda River near Madjarovo; Byal Gra-
dets Village; between Zhelezino and Gugutka villages; near Zhelezino Village; Harmanli.
Melanargia larissa (Geyer, [1828])
Studen Kladenets Village; Momina Skala Hut near Madjarovo; Madjarovo; on the
road from Madjarovo to Borislavtsi Village, 2.5 km from the bridge on Arda River near
Madjarovo; Harmanli.
580 S. BESHKOV, M. LANGOUROV
Beron page 580
Minois dryas (Scopoli, 1763)
Komuniga Village.
Hipparchia syriaca (Staudinger, 1871)
Kardjali; Studen Kladenets Dam, Sredna Arda Railway Station; Studen Kladenets
Dam, Kroyatsi Hunting Farm near Nanovitsa Village; Studen Kladenets Village; above
Dishlik-Dere Valley; the Iron Bridge on Arda River near Dolno Cherkovishte; Momina
Skala Hut near Madjarovo; Zhelezari Village; Arda Hut near Dabovets Village; Meden
Buk Village; Mandritsa Village.
Hipparchia volgensis (Mazochin-Porshnjakov, 1952)
Studen Kladenets Dam, Sredna Arda Railway Station, 21.6.1991, 1 , 13.10.1991, 2
‡‡, 18.10.1990, 1 .
Hipparchia semele (Linnaeus, 1758)
Studen Kladenets Dam, Sredna Arda Railway Station.
Hipparchia statilinus (Hufnagel, 1766)
Studen Kladenets Dam, Sredna Arda Railway Station; Madjarovo; near Madjarovo
on the road to Borislavtsi Village (2-3 kilometers after the bridge on Arda River); Harman-
li; Zhelezari Village
Hipparchia fatua Freyer, [1844]
Studen Kladenets Dam, Sredna Arda Railway Station; Studen Kladenets Village;
Zhelezari Village; Arda Hut near Dabovets Village; Meden Buk Village.
Arethusana arethusa ([Denis & Schiffermüller], 1775)
Madjarovo, 09.8.2003, Peter Davey leg.
Brintesia circe (Fabricius, 1775)
Near Plazishte Village; Studen Kladenets Dam, Sredna Arda Railway Station;
near Nanovitsa Village; Studen Kladenets Dam, Kroyatsi Hunting Farm near Nano-
vitsa Village; Studen Kladenets Dam, Darets; Studen Kladenets Village; near Rabovo
Village; Southern of Madjari Village; the Iron Bridge on Arda River near Dolno
Cherkovishte; the bridge on Arda River near Madjarovo; Byal Gradets Village; Zhelezari
Village; Zhaltichalskoto Dere Valley near Meden Buk Village; Lyubimets; Siv Klad-
enets Village.
Chazara briseis meridionalis (Staudinger, 1886)
Komuniga Village; Studen Kladenets Dam, Kroyatsi Hunting Farm near Nanovitsa
Village; Studen Kladenets Village.
Apatura ilia ([Denis & Schiffermüller], 1775)
Harmanli.
Insecta: Lepidoptera 581
Beron page 581
Apatura metis Freyer, 1829
The Iron Bridge on the Arda River near Dolno Cherkovishte; Kotlari Village; Mad-
jarovo; Lyubimets. Flies in two generations. Strictly protected by Appendix II of the Bern
Convention. A species from Appendix II and IV of the EEC Habitat Directive (92/43).
Limenitis camilla (Linnaeus, 1764)
Haskovo; Krumovgrad; Kotlari Village, 11.8.2003, Peter Davey leg.; Ivaylovgrad Dam,
Arda Hut near Dabovets Village; Harmanli; Meden Buk Village; Kamilski Dol Village.
Limenitis populi (Linnaeus, 1758)
Madjarovo, 09.8.2003, Peter Davey leg.
Limenitis reducta schiffermuelleri Higgins, 1932
Sredna Arda Railway Station; Haskovo; Studen Kladenets Village; Krumovitsa Riv-
er; above Dishlik-Dere valley; the Iron Bridge on Arda River near Dolno Cherkovishte;
Kotlari Village; Madjarovo; 2 km from the bridge on Arda River near Madjarovo to
Borislavtsi; Byal Gradets Village; Harmanli; Zhaltichalskoto Dere Valley near Meden
Buk Village; Dolno Lukovo Village; Lyubimets; Siv Kladenets Village, Ivaylovgrad Dis-
trict; Oreshenskoto-Dere Valley near Odrintsi Village.
Vanessa atalanta (Linnaeus, 1758)
Near Karangil Cave above Shiroko Pole Village, Kardjali district; Sredna Arda Rail-
way Station; Kroyatsi Hunting Farm near Nanovitsa Village; Delishte Village; Studen
Kladenets Village; 3 km SE of Krumovgrad on the road to Avren Village; Dishlik-Dere
Valley between Kulich Village, Potochnitsa District and the Iron Bridge on Arda River
near Dolno Cherkovishte Village; the Iron Bridge on Arda River near Dolno Cherkovish-
te Village; Madjarovo; Harmanli; Ivanovo Village; Dolno Lukovo Village; Ladjata Suburb
of Ivaylovgrad; Dupkata Hill near Ivaylovgrad; Siv Kladenets Village, Ivaylovgrad Dis-
trict; Kodja-Kaya Hill near Odrintsi Village.
Va ne s s a c a r d u i (Linnaeus, 1758)
Sredna Arda Railway Station; Kroyatsi Hunting Farm near Nanovitsa Village; Studen
Kladenets Village; the Iron Bridge on Arda River near Dolno Cherkovishte; Momina Skala
Hut; the bridge on Arda River near Madjarovo; near Gorno Pole Village, Madjarovo District;
Madrets Village; Harmanli; Arda Hut near Dabovets Village; Meden Buk Village; Zhaltichal-
skoto Dere Valley near Meden Buk Village; Kamilski Dol Village; Dupkata Hill near Ivay-
lovgrad; between Zhelezino and Gugutka Villages; Siv Kladenets Village, Ivaylovgrad District.
Inachis io (Linnaeus, 1758)
Gyaur Hambar Cave near Ridino Village; Zlatnata Yama Cave near Kremen Village,
Podkova District, hibernating in the cave; Konevo Village; Studen Kladenets Village; Dish-
lik-Dere Valley between Kulich Village, Potochnitsa District and the Iron Bridge on Arda
River near Dolno Cherkovishte Village; Momina Skala Hut; Madjarovo; on the road
from Madjarovo to Borislavtsi Village, 2.5 km from the bridge on Arda River near Mad-
jarovo; Harmanli; Disused Mine Gallery in Gurgen Dere Valley near Gaberevo Village,
582 S. BESHKOV, M. LANGOUROV
Beron page 582
Madjarovo District; Svirachi Village, Ivaylovgrad District; Siv Kladenets Village, Ivay-
lovgrad District; Kodja-Kaya Hill near Odrintsi Village, hibernating in a pill-box.
Aglais urticae (Linnaeus, 1758)
Studen Kladenets Village; Momina Skala Hut near Dabovets Village; Meden Buk
Village; Kamilski Dol Village; Siv Kladenets Village, Ivaylovgrad District.
Polygonia c-album (Linnaeus, 1758)
Momchilgrad; Near Karangil Cave above Shiroko Pole Village, Kardjali district; Sredna
Arda Railway Station; Kroyatsi Hunting Farm near Nanovitsa Village; Studen Kladenets
Village; Krumovitsa River; Krumovgrad; Dishlik-Dere Valley between Kulich Village, Po-
tochnitsa District and the Iron Bridge on Arda River near Dolno Cherkovishte Village;
the Iron Bridge on Arda River near Dolno Cherkovishte Village; Kotlari Village; Madjaro-
vo; Harmanli; Arda Hut near Dabovets Village; Ivaylovgrad Dam, Kochash Village; Siv
Kladenets Village, Ivaylovgrad District.
Polygonia egea (Cramer, [1775])