Article

Tetrigoidea and Tridactyloidea (Orthoptera: Caelifera) and Addenda to OSF Vols. 1-5. Orthoptera Species File

Authors:
To read the full-text of this research, you can request a copy directly from the author.

No full-text available

Request Full-text Paper PDF

To read the full-text of this research,
you can request a copy directly from the author.

... Tetrix contractus: Hancock 1907bHancock 1909: 412;Hancock 1913: 52;Günther 1939: 124. Lamellitettigodes contractus: Günther 1939Steinmann 1970a: 228;Blackith 1992: 100;Paris 1994: 235;Yin et al. 1996: 878;Otte 1997 Notes.-Bolívar (1898) described the species under the section Metrodorae as belonging to Xistra, while earlier he described L. contractus within the section Tettigiae, assigned to the genus Paratettix. ...
... Moreover, I did not find reported differences when comparing the types. Otte (1997) gave the specific status to the subspecies only with the reason of occurrence on different islands. ...
... In the specimens of L. contractus from Borneo, these undulations are weaker. Otte (1997) gave the subspecies a specific status based on occurrence on different islands (as in L. tricristatus mentioned above). All these features are within the range of variation of the species and do not represent diagnostic traits. ...
Article
Full-text available
The genus Lamellitettigodes Günther, 1939 from Southeast Asia is reviewed. The genus currently includes seven species and is transferred to Tetriginae Rambur, 1838. Two new species are described: Lamellitettigodes novaeguineaesp. nov. from New Guinea and Lamellitettigodes karwinkelisp. nov. from Yunnan, People’s Republic of China. Lamellitettigodes palawanicus Günther, 1939 stat. nov. is no longer regarded as a subspecies of L. contractus , but a separate species. Two species are transferred from Euparatettix Hancock, 1904 to Lamellitettigodes : Lamellitettigodes sagittatus (Bolívar, 1887) comb. nov. and Lamellitettigodes cultratus (Bolívar, 1898) comb. nov. One species is transferred from Tetrix Latreille, 1802 to Lamellitettigodes - Lamellitettigodes signatus (Bolívar, 1887) comb. nov . Xistra tricristata sumatrana Bolívar, 1898 syn. nov. , Xistra tricristata Bolívar, 1898 syn. nov. , and Probolotettix corticolus Blackith & Blackith, 1987 syn. nov. are synonymized with Lamellitettigodes contractus (Bolívar, 1887). Probolotettix kevani Blackith & Blackith, 1987 syn. nov. is synonymized with Euparatettix personatus (Bolívar, 1887). Additionally, in the genus Lamellitettix Hancock, 1904, Lamellitettix acutus Hancock, 1904 stat. rev. is retransferred from a subspecies of Lamellitettix gallinaceus (Stål, 1877) to a separate species status.
... In the next revision of Australian Tetrigidae, one new species of Vingselina was described and the genus was placed in the subfamily Cladonotinae on the basis of the considerably widened frontal ridge (Rehn, 1952). But this point of view was not accepted, and Vingselina was again placed in the subfamily Batrachideinae (Otte, 1997;Cigliano et al., 2018). Vingselina willemsei Günther, 1937 was described from the Solomon Islands (Günther, 1937), but later this species was placed in the monotypic genus Vilma Steinmann, 1973; the differences of both genera are given in a key (Steinmann, 1973). ...
... Based on the morphology of the male genitalia of four species of the subfamily Tetriginae and two species of Batrachideinae, Podgornaya (2001aPodgornaya ( , 2001bPodgornaya ( , 2003 divided the latter subfamily into two tribes: Batrachideini and "Cassitettini". But a little earlier Otte (1997) seems to have correctly modified the original name of the latter group (subfamily "Cassitettinae" Yin, 1984) in Cassitettiginae. Nowadays Cassitettiginae and "Cassitettini" are considered as synonyms of Batrachidein-ae (Cigliano et al., 2018). ...
... Bufonides was placed in the "section Bufonidae" by Hancock (1907), but Steinmann (1970) transferred it in Cladonotinae. Later Otte (1997) allocated this genus to the tribe Tetrigini (subfamily Tetriginae), while Tumbrinck (2014) considered Bufonidinae as a synonym of Cladonotinae. Nowadays Bufonidinae is treated as a synonym of Batrachideinae (Tumbrinck & Skejo, 2017;Cigliano et al., 2018). ...
Article
Full-text available
The pygmy grasshopper subfamily Batrachideinae consists of three tribes: Batrachideini Bolívar, 1887; Cassitettigini Yin, 1984, nom. resurr.; Bufonidini Hancock, 1907, nom. resurr., stat. nov. The Australian pygmy grasshoppers of the tribe Bufonidini are reviewed. Three new genera are described, namely Paraselina gen. nov. (type species Vingselina multifora Rehn, 1952), Anaselina gen. nov. (type species Vingselina minor Sjöstedt, 1921), and Selivinga gen. nov. (type species Selivinga tribulata sp. nov.). Four new combinations are proposed: Paraselina multifora (Rehn, 1952), comb. nov.; P. trituberculata (Sjöstedt, 1932), comb. nov.; P. brunneri (Bolívar, 1887), comb. nov.; Anaselina minor (Sjöstedt, 1921), comb. nov. A key to Australian genera of Batrachideinae is provided.
... We present basis for further studies and understanding of the former, but for latter genera and their species we point the necessity of deep and detailed taxonomic and biogeographic revision, thus they are briefly discussed and their species are shortly listed in checklist with type locality data and notes. Genus Amphibotettix Hancock, 1906 Amphibotettix: Hancock 1906, Hancock 1907a, Kirby 1910, Hancock 1915, Günther 1942, Blackith 1992, Yin et al. 1996, Otte 1997, Storozhenko & Dawwrueng 2015. Amphybotettix (misspelling) : Steinmann 1970. ...
... Genus Eufalconius Günther, 1938 Eufalconius: Günther 1938a, Blackith 1992, Yin et al. 1996, Otte 1997 Type species: Eufalconius pendleburyi Günther, 1938 Composition and distribution. Monotypic genus, its only species (E. ...
... Euscelimena: Günther 1938a, Blackith 1992, Yin et al. 1996, Otte 1997, Gupta & Chandra 2018, Skejo et al. 2018. Abbasia: Kirby 1914, Hebard 1930. ...
Book
Full-text available
Pygmy grasshoppers (Tetrigidae) are a speciose group of complicated taxonomy, with many species requiring clarification on their species boundaries, and more still awaiting discovery. Two new species of Scelimena Serville, 1838 are described: (1) from S. discalis species group S. gombakensis sp. nov. and (2) from S. hexodon species group Scelimena marta sp. nov. Catalogue of Scelimenini genera (15) and species (100) is presented and taxonomy and biogeography of the tribe are discussed. New and resurrected combinations are: Falconius becvari (Buzzetti & Devriese, 2008) comb. nov. (of Gavialidium becvari), Gavialidium carli Hebard, 1930 comb. resurr. (of Bidentatettix carli), Indoscelimena india (Hancock, 1907) comb. nov. (of Scelimena india), Paragavialidium nodiferum (Walker, 1871) comb. nov. (of Platygavialidium nodiferum), Platygavialidium productum (Walker, 1871) comb. nov. (of Gavialidium productum), Scelimena hexodon (Haan, 1843) comb. resurr. (of Hexocera hexodon), Scelimena rosacea (Hancock, 1915) comb. resurr. (of Amphibotettix rosaceus), Tegotettix bufocrocodil (Storozhenko & Dawwrueng, 2015) comb. nov. (of Gavialidium bufocrocodil). New synonyms are: Gavialidium phangensum Mahmood, Idris & Salmah, 2007 syn. nov. (of Eufalconius pendleburyi), Gavialidium philippinum Bolívar, 1887 syn. nov. (of Platygavialidium productum comb. nov.), Hexocera Hancock, 1915 syn. nov. (of Scelimena), Paracriotettix Liang, 2002 syn. nov. (of Scelimena), Paracriotettix zhengi Liang, 2002 syn. nov. (of Scelimena melli), Scelimena mellioides Deng, 2016 syn. nov. (of Scelimena melli), Scelimena wuyishanensis Deng, 2016 syn. nov. (of Platygavialidium sinicum). The genus Scelimena is divided into six species groups. Eucriotettix neesoon Tan & Storozhenko nom. nov. is new name for homonym Eucriotettix guentheri Tan & Storozhenko, 2017. Finally, a tabular key to 15 Scelimenini genera, based on 16 morphological characters, is presented.
... The Tetrigidae are basal caeliferan Orthoptera distinguished by their small size (<15 mm), pronotum extended over the head and lack of arolia in the tarsal claws (Perez-Gelabert et al., 1998). The most recent world catalogue of Tetrigidae was published by Otte (1997) and included some 1195 species in 11 subfamilies. The Cladonotinae are distinguished by a widely forked frontal costa and for being wingless; this latter feature possibly being an important factor determining their limited dispersal capabilities. ...
... The Cladonotinae are mostly an Old World group with relatively few species in the New World. Over 150 species of Cladonotinae have been reported from the Ethiopian and Oriental regions (Otte, 1997), while only three genera and six species are known from South America where other tetrigid subfamilies are predominant (Steinmann, 1969). In contrast to their rarity in the New World, the Cladonotinae appear to be the dominant group of tetrigids in the Greater Antilles. ...
Article
Full-text available
The new genus and its type species Armasius iberianus, a remarkable cladonotine tetrigid, are described based on a single individual collected from Monte Iberia, Guantánamo Province, in eastern Cuba. Armasius is characterized for having a large pronotum flattened on top to a thin crest that is uniformly elevated only about half the body height, the pronotum covers the whole abdomen, narrowing posteriorly, and turning upwards at its posterior end. A character not yet seen in any other Antillean Cladonotinae is that the posterior angles of the lateral lobes of the pronotum project laterally, flattening to form a horizontal triangular spine.
... Firstly Tripetalocerinae was considered as separate subfamily of family Tetrigidae by H. Steinmann (1970). This point of view was accepted in majority of recent publications (Otte, 1997;Mahmood et al., 2007), but attempts to raise the taxonomic status of this subfamily to the family Tripetaloceridae was made (Liang & Zheng, 1998;Zheng, 2005). In the Orthoptera Species File (Eades et al., 2013) it is treated as subfamily Tripetalocerinae with a single tribe Tripetalocerini. ...
... 4, 5) studied and illustrated the holotype and additional specimens of K. apicalis Bolívar, 1909 from Sulawesi and emphasized that the shape and the number of segments of antennae in this species is the same as in the genus Discotettix (Fig. 46) and genus Kraengia must be placed in the section Discotettigiae. Though, still now Kraengia is considered as a member of Tripetalocerinae (Steinmann, 1970;Otte, D. 1997;Eades et al., 2013). Herein I support the Günther's point of view and again transferred Kraengia to the subfamily Discotettiginae based on the specific shape of antennae. ...
Article
Full-text available
Subfamily Tripetalocerinae consist of four genera distributed in Southeast Asia. Clinophaestus gen. nov. (type species: Clinophaestus robustus sp. nov.) and Tripetaloceroides gen. nov. (type species: Tripetalocera tonkinensis Gunther, 1938) are described. New tribe Clinophaestini trib. nov. (type genus: Clinophaestus gen. nov.) is established. Diagnosis and key to tribes and genera are given. An annotated list of species is provided. Genus Kraengia Bolivar, 1909 is transferred from Tripetalocerinae to Discotettiginae.
... America in particular is extremely diverse and yet simultaneously poorly documented. Indeed, only 19 tridactyloid species are recorded from Mesoamerica compared to over 100 known from South America (Otte 1997; Günther 1980; Eades et al. 2011). Work on Mesoamerican tridactyloids began with Saussure's (1859) description of Ripipteryx mexicana from Oaxaca, Mexico and continued with his contribution to Biologia Centrali-Americana (Saussure 1896) in which he described 17 additional species. ...
Article
Full-text available
A new species of the genus Ripipteryx (Orthoptera: Tridactyloidea: Ripipterygidae) from the Toledo District of southern Belize is described and illustrated. Ripipteryx mopana sp. n. is placed in the Scrofulosa Group based on its elaborately ornamented frons and is readily distinguished from its congeners by the fusion of the superior and inferior frontal folds to form a nasiform median process, the epiproct with both anterior and posterior margins emarginate, the subgenital plate with distinct lateroapical depressions either side of the median line, the basal plate of the phallus strongly bilobed apically, and the development of well-demarcated denticular lobes in the dorsal endophallic valves. A preliminary key to the species of the Scrofulosa Group is provided.
... The genus currently consists of 38 described species (Hebard, 1930;Günther, 1938;Blackish, 1992;Zheng, 2005;Li et al., 2014;Storozhenko & Dawwrueng, 2015;Cigliano et al., 2016). Majority of them are the locally endemic (Zheng & Ou, 2001;Otte, 1997;Deng et al., 2012). Only seven species of Eucriotettix are more or less widely distributed in Asia: E. spinilobus (Hancock, 1904) is known from Sri Lanka and India, E. maculatus (Kirby 1914) from Myanmar and India, E. dohertyi (Hancock, 1915) from India and China, E. tricarinatus (Bolívar, 1887) from Sri Lanka, India, Malaysia and Philippines, E. grandis (Hancock, 1912) from India, Nepal, Thailand and China, E. oculatus (Bolívar, 1898) from Indonesia, Malaysia, Vietnam, China, Taiwan and Japan (Ryukyu Islands), and E. ridleyi Günther, 1938 is recorded from Singapore, China and Indonesia (Shishodia, 1991;Ichikawa, 1994;Blackish & Blackish, 1987;Liang & Zheng, 1998;Ingrisch, 2001;Mahmood at al., 2007;Kim & Pham, 2014;Storozhenko & Dawwrueng, 2015). ...
Article
Limno-terrestrial pygmy grasshoppers are mostly recognised using morphology but characters used are not always clear-cut for species identification. From a faunistic survey in Singapore, Eucriotettix were collected and two new species are identified and described here: E. guentheri sp. nov. and E. simulans sp. nov. This was done by iteratively incorporating morphological, molecular, morphometric, and ecological data. Mitochondrial COI trees, COI cluster analysis, microhabitat preference, and morphometric analyses provide strong enough indirect evidence that the specimens comprise a pair of species. A key to species of Eucriotettix from Singapore is also provided.
... These diminutive insects are primarily ground-dwelling and most commonly encountered among leaf litter on the forest floor, or in wet, marshy habitats bordering rivers, streams or standing water. With more than 1,700 species in over 250 genera, the Tetrigidae are, among Caelifera, second only to Acrididae in terms of species diversity (Steinmann 1962(Steinmann , 1969(Steinmann , 1970(Steinmann , 1971Otte 1997). The family has a cosmopolitan distribution and is most diverse in the tropics (Hancock 1907;Rentz 1991;Heads 2009a). ...
Article
Full-text available
A new genus and species of pygmy grasshopper (orthoptera: Tetrigidae) is described from Early Miocene (Burdigalian) Dominican amber. Electrotettix attenboroughi Heads & Thomas, gen. et sp. n. is assigned to the subfamily Cladonotinae based on the deeply forked frontal costa, but is remarkable for the presence of tegmina and hind wings, hitherto unknown in this subfamily.
... Notes. Recently two replacement names, Ergatettix guentheri Steinmann, 1970 and Ergatettix paranodulosus Otte, 1997, proposed for Euparatettix nudulosus are set aside (Tumbrinck, 2015). ...
Article
Full-text available
An annotated list of 39 species in 25 genera and seven subfamilies of the pygmy grasshoppers (Orthoptera: Tetrididae) from Thailand is given; from these 18 species are recorded from this country for the first time. Five new species are described: Cotysoides gaponi sp. nov. (subfamily Metrodorinae), Eucriotettix anisyutkini sp. nov., Gavialidium bufocrocodil sp. nov., Scelimena bellula sp. nov. (subfamily Scelimeninae) and Phaesticus uvarovi sp. nov. (subfamily Discotettiginae). One species is transferred from Scelimena to Amphibotettix and a new combination is proposed: Scelimena hafizhaii Mahmmod, Idris et Salman, 2007 = Amphibotettix hafizhaii (Mahmmod, Idris et Salman, 2007), comb. nov. The previously unknown male of Falconius tschernovi Storozhenko, 2014 is described.
... All these genera developed similar morphological traits, probably as an adaptation to deter predators. Two genera with similar general appearance to Arulenus are shown on Figure 2. Hancock 1907, Kirby 1910, Blackith 1992, Yin et al. 1996, Otte 1997 Material examined. (1 known specimen): Holotype: (1/1) 1♀ Ins. ...
Article
Full-text available
Arulenus miae Skejo & Caballero sp. nov. is described from Buknidon and Davao, Mindanao, the Philippines. The species was serendipitously found in an amateur photo posted in Orthoptera Facebook group by Leif Gabrielsen. Holotype and paratype are deposited in Nederlands Centrum voor Biodiversiteit in Leiden, the Netherlands. Detailed comparison with Arulenus validispinus Stål, 1877 is given. A new diagnosis of the genus and A. validispinus is given. The paper is part of the revision of the subfamily Discotettiginae. This study provides a good example of how social networks can be used as a modern tool of discovering biodiversity if the regulations of the International Code of the Zoological Nomenclature are followed. A brief insight into habitat and ecology of this rainforest and mountainous species is presented.
... The type specimen and generic placement of Tridactylus galla Saussure, 1895 (1896), Lucas (1898) and Kirby (1906), but aside from a brief mention by Günther (1995), is entirely overlooked by subsequent authors and is absent from Otte's (1997) catalogue. During the course of compiling an annotated catalogue of the Orthoptera described by Saussure (Hollier and Heads, 2012) we were able to relocate the type of Tridactylus galla in the collection of the Museo Civico di Storia Naturale "Giacomo Doria" in Genova, Italy. ...
Article
Tridactylus galla was described by Henri de Saussure (1895) on the basis of a single adult female collected during Vittorio Bottego’s first expedition to the Horn of Africa in 1892 and 1893. The species appears in lists compiled by Fenizia (1896), Lucas (1898) and Kirby (1906), but aside from a brief mention by Gunther (1995), is entirely overlooked by subsequent authors and is absent from Otte’s (1997) catalogue. During the course of compiling an annotated catalogue of the Orthoptera described by Saussure (Hollier and Heads, 2012) we were able to relocate the type of Tridactylus galla in the collection of the Museo Civico di Storia Naturale “Giacomo Doria” in Genova, Italy. Our examination of the specimen confirmed Gunther’s (1995) assertion that its placement in Tridactylus Olivier, 1789 is erroneous, and the species is herein formally transferred to the genus Xya Latreille, 1809.
... Even GÜNTHER (1938), who reviewed the Discotettiginae seven years later, overlooked the publication. OTTE (1997), in the fi rst printed version of the Orthoptera Species File, listed this species as of uncertain placement within Tetrigidae. The type specimen was listed in the catalogue of Bolívar's collection (PARÍS 1994). ...
Article
Full-text available
The genus Rosacris Bolívar, 1931 is a pygmy grasshopper (Orthoptera: Tetrigidae) genus hitherto known only from a holotype male of its type species Rosacris antennata Bolívar, 1931 from Makiling Mt. (Luzon island, the Philippines) and originally assigned to the Discotettiginae. Redescription of the genus and its type species are based on the holotype and a new photographic record of the species (Isarog Mt.) found in Flickr (www.flickr.com). The holotype is found to be male nymph rather than an adult animal. The species is morphologically similar to members of the genus Metamazarredia Günther, 1939 from the Philippines. Therefore the genus is transferred from the subfamily Discotettiginae to Metrodorinae. Assumption exists that the specimens hitherto identified as R. antennata represent nymphs of Metamazarredia fuscipes (Stål, 1877).
... The subfamily Scelimeninae presents a predominantly tropical distribution, appearing in Southeast Asia, Australia, Africa, and Central and South America (Otte 1997;Deng et al. 2007;Cigliano et al. 2016). It is mainly characterized by having the filiform antenna, a relatively small divergence of the rami of the frontal costa not forming a frontal scutum. ...
Article
Full-text available
A new species, Hebarditettix longipennis n. sp. is described with detailed illustrations of external morphology. Comparison between the species H. longipennis n. sp. and allied species is compiled into a table for identification. The distribution and an updated identification key to all known species of the genus are given. http://zoobank.org/urn:lsid:zoobank.org:pub:01FDB52B-4EF0-4E2A-9912-42AD58AA0DBB
... The subfamily Cladonotinae is one of the most diverse groups of pygmy grasshoppers (Orthoptera: Tetrigidae) in the Oriental region (Blackith 1992;Otte 1997;Tumbrinck 2014;Bhaskar et al. 2020;Cigliano et al. 2021). Three genera of this subfamily are probably closely related. ...
Article
The species composition of the genus Epitettix (Tetrigidae: Cladonotinae) is discussed. The differential diagnosis of the genus is given. New synonymies are proposed: Epitettix Hancock, 1907 = Vaotettix Podgornaja, 1986, syn. nov., = Pseudepitettix Zheng, 1995, syn. nov. Two new taxa are described, namely Epitettix mikhailovi sp. nov. from Vietnam (Gia Lai Province) and Epitettix punctatus montanus subsp. nov. from Malaysia (Sabah, North Borneo). Seven new combinations are established: Epitettix guibeiensis (Zheng et Jiang, 1995), comb. nov., E. hainanensis (Deng, 2020), comb. nov., E. linaoshanensis (Liang et Jiang, 2004), comb. nov., E. nigritibis (Zheng et Jiang, 2000), comb. nov., E. pimkarnae (Storozhenko et Dawwrueng, 2014), comb. nov., E. strictivertex (Deng, 2020), comb. nov., and E. parallelus (Podgornaja, 1986), comb. nov. The unknown male of the latter species is also described and illustrated.
... Saussurella borneensis, Saussurella cornifrons, Saussurella curticornu, Saussurella indica; Saussurella brunneri was in need of a replacement name. Grant (1966) reviewed the genus and S. brunneri Hancock, 1912 and S. cucullifera (Walker, 1871) were regarded as synonyms of S. cornuta (Haan, 1843); S. cornifrons S. sumatrensis Bolivar, 1898 were regarded as synonym of S. decurva Brunner von Wattenwyl, 1893;Podgornaya (1992) described Saussurella inelevata; Cassitettix longiptera was transferred into Saussurella by Otte (1997); Liang and Zheng (1998) described Saussurella acuticornis Zheng; Zheng et al. (2013) described Saussurella xizangensis; and S. inelevata Podgornaya, 1992 was regarded as a synonym of S. decurva Brunner von Wattenwyl, 1893 by Zheng et al. (2013). To date, the genus includes nine species endemic to southeast Asia. ...
Article
Full-text available
Taxonomy of a tetrigid genus Saussurella Bolivar is reviewed. One new species, Saussurella brachycornis sp. nov. is described from China and an updated identification key to all known species of the genus is given, as well as brief comments on phylogenetic relationships and distributions of the genus.
... The tribe Xerophyllini includes 15 genera and 30 species, which are distributed mainly in tropical Africa and Madagascar. The nominotypical tribe contains about 60 genera and near 220 species, distributed as follows: South America and The Caribbean islands (18 genera, 29 species), Africa and Madagascar (6 genera, 31 species), Australia (2 monotypic genera), the small Melanesian islands (2 genera, 4 species), temperate Asia (7 genera, 27 species), but majority of taxa are known from South-East Asia -32 genera and 126 species (Bolívar 1887, Günther 1979, Blackith 1992, Otte 1997, Tumbrinck 2014, Cigliano et al. 2017. Previously only one species of Cladonotini, Potua coronata Bolívar, 1887, was known from the Peninsular Malaysia (Bolívar 1887). ...
Article
Full-text available
The monotypic genus Hancockitettix Storozhenko et Pushkar, gen. Nov. (type species Hancockitettix humeratus Storozhenko et Pushkar, sp. Nov.) from the Malay Peninsula is described. A new genus belongs to the subfamily Cladonotinae. It resembles the genera Hancockella Uvarov, 1940 from India and Austrohancockia Günther, 1938 from China, Taiwan, Japan and Vietnam in the general body appearance, in the large lobules on the upper and lower margins of the fore and mid femora, and in the relatively narrow scutellum, but differs from them by the presence of fully developed tegmen, hind wing and tegminal sinus, by the triangular plate-like projected shoulder of pronotum, and by the considerable distance between the most outstanding parts of shoulders which exceeds the pronotum lobe width (in other mentioned genera the hind wing and tegminal sinus are absent, the tegmen is absent or rudimentary, lateral side of shoulder is broadly rounded or obtusely angular, and the distance between shoulders is less than the pronotum lobe width).
... nov.). Subsequent authors followed Rehn's application of these names (Günther 1939(Günther , 1959(Günther , 1970Yin et al.1996;Otte 1997). This was a serendipitous error, as Rehn (1929) did not have a possibility to check what Serville (1838) described under the name H. lucifer. ...
Article
Full-text available
Madagascar is home to some of the largest and most colorful pygmy grasshoppers (Tetrigidae) in the world, known as ‘Malagasy Metrodorinae’. Among them, Devil’s pygmy grasshoppers (genus Holocerus Bolívar, 1887) are unique in having two long spines on the back, which are modified internal lateral pronotal carinae. The genus Holocerus was composed of two species – H. lucifer (Serville, 1838) and H. taurus Rehn, 1929 syn. nov. , but here it is evidenced that the latter represents a junior synonym of the former. Simultaneously, H. devriesei sp. nov. is described as a species new to science. Holocerus lucifer is a northern species of paler coloration and longer spines (distributed from Marojejy and Maroantsetra in the north to Zahamena in the south), whereas H. devriesei sp. nov . represents the southern and darker species (distributed from Vohimana and Andasibe-Mantadia in the south to the Antongil Bay in the north). There are potential overlaps in the distribution of the two species, but without more georeferenced localities, it is impossible to discriminate whether they occur only sympatrically or also syntopically.
... Following Storozhenko & Paik's (2011) division of the subfamily into two tribes, Cladonotini are characterized with a convex vertex, whose middle part is the highest in frontal view, while fastigial horns are lacking or at least, are not the highest part of the vertex. Bolívar 1887, Hancock 1904, 1907a, 1915, Kirby 1910, 1914, Günther 1938, Sandrasagara 1950, Blackith 1992, Otte 1979, 1997, Yin et al. 1996, Hollier 2013 Distribution and habitat. Sri Lanka: Sinharaja rainforest. ...
Article
Full-text available
Wide nosed pygmy grasshoppers (Tetrigidae: Cladonotinae) are grasshoppers in which the frontal costa bifurcates into strongly divergent facial carinae, between which there is wide scutellum. Altogether seven genera and thirteen species inhabit India and Sri Lanka. Tribe Cladonotini is represented in this region by five genera and nine species (Cladonotus— 4 spp., Deltonotus—2 spp., Gignotettix—1 sp., Hancockella—1 sp., and Yunnantettix—1 sp.), while the tribe Xerophyllini by two genera, Tettilobus (3 sp.) and Potua (1 sp.). For this Potua species it is not clear if it belongs to this genus. A simple identification key to Cladonotinae of India and Sri Lanka is provided. New synonyms are: Deltonotus humilis syn. nov., Epitettix tamilus syn. nov. and Potua aptera syn. nov., synonymous with Deltonotus gibbiceps. Deltonotus cristatus should be regarded nomen nudum. A new species, assigned to the genus Tettilobus is described from the Western Ghats of Kerala, Tettilobus trishula Skejo, Bhaskar et Stermšek sp. n. Furthermore, we present the first records of D. subcucullatus from India (Kerala). Key words: Cladonotini, Xerophyllini, Western Ghats, rainforest, endemism, synonyms
... The monotypic genus Bolivaritettix was established by , it was considered a member of the subfamily Metrodorinae (Tetrigidae) based on head not exserted, antennae filiform and pronotum truncate anteriorly. The genus Bolivaritettix Günther is a large-sized genus in the Metrodorinae with 98 previously known world species Steinmann, 1962;Kevan, 1966;Shishodia, 1991;Blackith, 1992;Yin, 1996;Otte, 1997;Jiang & Zheng, 1998;Liang & Zheng, 1998;Zheng, 2005;Ingrisch, 2006;Deng, 2016;Cigliano et al., 2017) which are distributed over India, Vietnam, Palawan, Cambodia, Myanmar, Philippines, China, Nepal, Indonesia, Malaysia and Thailand. A total of 71 species of Bolivaritettix Günther occurring in China are systematically researched in this paper. ...
Article
Full-text available
The tetrigid genus Bolivaritettix Günther, 1939 from China is taxonomically reviewed. The genus now includes 71 species from China, in which two new species are described and illustrated, namely: Bolivaritettix baoxingensis Deng sp. nov. and Bolivaritettix strictifemura Deng sp. nov. Synonymy. Their distribution and an annotated identification key to Chinese species are provided.
... Remarks. Otte (1997) erroneously recorded the type species of this genus as based on monotypy, while in the original description type species was really designated by Hancock (1909). Composition and distribution. ...
Article
Full-text available
There are numerous pygmy grasshoppers (Tetrigidae) that exhibit leaf-like appearance. Leaf-mimic species can be found mainly in the subfamily Cladonotinae (tribes Cladonotini, Xerophyllini). Two leaf-mimic pygmy grasshopper species found in India, Pakistan (Oxyphyllum pennatum Hancock, 1909) and Borneo (Paraphyllum antennatum Hancock, 1913) are the only macropterous leaf-like species of Asia and were traditionally assigned to subfamily Cladonotinae. Our study present new records of these species, as well as updated descriptions. Oxyphyllum pennatum is recorded for the first time from Chhattisgarh (Central India), and furthermore, records from Pakistan were reviewed and confirmed. Paraphyllum antennatum is reported from a few localities in the mountains of Borneo (East Malaysia). New records found in online social media (Flickr, iNaturalist) were implemented. Brachypronotal specimens of P. antennatum are reported. After comparison with Cladonotini and Xerophyllini members, we conclude that Oxyphyllum and Paraphyllum should not be regarded Cladonotinae members. Traditional assignment was based on superficial resemblance to leaf-like Cladonotinae genera (e.g. Phyllotettix, Hymenotes, Holoarcus, Xerophyllum). However, comparison of morphological characters reveals that Oxyphyllum shares morphology with Tetriginae genera, while Paraphyllum shares morphology with Asian Metrodorinae. Here, we assign the genus Oxyphyllum to Tetriginae and the genus Paraphyllum to Metrodorinae.
... The genus Formosatettix was considered a member of the subfamily Tetriginae (Tetrigidae) based on head not exserted, antennae filiform and lateral lobes with the posterior angle broadly rounded. Formosatettix Tinkham is a large-sized genus of the Tetriginae with 62 previously known species in the world (Tinkham 1937;Bey-Bienko 1951;Storozhenko 1981;Blackith 1992;Storozhenko and Ichikawa 1993;Yin et al. 1996;Otte 1997;Jiang and Zheng 1998;Liang and Zheng 1998;Ingrisch 2001;Kim and Kim 2004;Zheng 2005Zheng , 2012Deng et al. 2007;Deng 2016;Cigliano et al. 2018) which are distributed over China, Russia, Korea, Japan, Pakistan and Nepal. In this report, the utility of the characters used to diagnose the genus is redetermined. ...
Article
Full-text available
Formosatettix serrifemora Deng sp. nov., Formosatettix zhejiangensis Deng sp. nov. and Formosatettix guposhanensis Deng sp. nov. are described with detailed illustrations of external morphology. New synonymy is proposed: Formosatettix tianmushanensis Zheng and Li = Formosatettix xiai Zheng, Shi and Yin, syn. nov. An updated key to species of the genus is given. http://www.zoobank.org/urn:lsid:zoobank.org:pub:D64FA763-EBA7-453C-9D76-7209F5659478
... Remarks. Otte (1997) erroneously recorded the type species of this genus as based on monotypy, while in the original description type species was really designated by Hancock (1909). Composition and distribution. ...
... This material is not divisible into two separate genera, so the variability of pronotal morphology points that there is a single leaf-like Cladonotinae genus in the Philippines, and the name Hymenotes has nomenclatural priority. Thus, all the leaf-like pygmy grasshoppers from the Philippines belong to Hymenotes, hence the new combination Hymenotes westwoodi (Bolívar, 1887) Bolívar 1887, Hancock 1904, 1907, 1915, Kirby 1910, 1914, Günther 1938, Sandrasagara 1950, Blackith 1992, Otte 1979, 1997, Yin et al. 1996, Hollier 2013, Tumbrinck 2014, Bhaskar et al. 2020 Type species. tettix (Cladonotus) humbertianus Saussure, 1862. ...
Article
Full-text available
The paper describes a new species of a Sri Lankan twighopper, genus Cladonotus Saussure, 1862 (Orthoptera: Tetrigidae: Cladonotini), C. bhaskari sp. n., based on a single female specimen collected and photographed by T. Kirschey in 2016 in Sinharaja rainforest (SW Sri Lanka). The new species is clearly distinguished from other species of the genus by its long and spine-like frontomedial projection and cockscomb-shaped promedial projection. Species of this genus resemble tiny twigs, hence the name ‘twighopper’. Our new specimen is the first known female of the genus, and we also present the first photograph of the member of this genus in the natural habitat. An annotated identification key for the Cladonotus species is provided. Furthermore, we synonymize genus Hypsaeus Bolívar, 1887 syn. nov. with Hymenotes Westwood, 1837, based on the pronotal variability of the leaf-like pygmy grasshoppers in the Philippines and introduce one new combination—Hymenotes westwoodi (Bolívar, 1887) comb. nov..
... The species has not been reported since. The original description was later only cited in taxonomic catalogues (Blackith 1992, Yin et al. 1996, Otte 1997, Cigliano et al. 2016. The genus and its only species were assigned to Scelimeninae on account of its similarity to Platygavialidium Günther, 1938 (Günther 1938, Cigliano et al. 2016. ...
Article
Full-text available
Evidence is presented on the synonymy of Hancockitettix humeratus Storozhenko et Pushkar, 2017 syn. nov., (a recently described species from the Cameron Highlands of the Malay peninsula (the only recorded specimen, a female holotype, caught in 2014)) and Eufalconius pendleburyi Günther, 1938, a long-time-no-see species from Bukit Kutu of the Malay peninsula (the only recorded specimen, a female holotype, caught in 1926)-100 km S of the Cameron Highlands and part of the same massif (Tenasserim range: Titiwangsa Mts.). The holotypes of the two species are conspecific, almost identical in morphology. As a result Hancockitettix Storozhenko et Pushkar, 2017 syn. nov. becomes a junior synonym of Eufalconius Günther, 1938. The original description of Hancockitettix syn. nov. and H. humeratus syn. nov. contains much more data than that of Eufalconius and E. pendleburyi and serves as a very good 're-description'. New records of E. pendleburyi found on the social media photo database Flickr from the same massif (Endau Rompin NP, Bukit Fraser) are presented herein. Detailed comparison is given between the genera Austrohancockia Günther, 1938, Discotettix Costa, 1864, Eufalconius, Gibbotettix Zheng, 1992, Hancockella Uvarov, 1940, Paragavialidium Zheng, 1994, and Platygavialidium Günther, 1938.
Article
An Si3N4 sample was irradiated with an Nd-YAG laser. The species present in the plume were detected by a quadrupole mass spectrometer (QMS). The species observed were Si, SiN, Si2, Si2N, Si3, Si3N, Si4, Si3N4, Si6, Si7, and N. Weak signals were observed at m/e = 126 and 280. The analysis of the velocity of the different species indicates the existence of thermal equilibrium in the vaporization process.
Article
The preoccupied genus name of Eurasian grasshopper Primnoa Fischer-Waldheim 1846 (Arthropoda:Orthoptera) (nec Lamouroux 1812 [Cnidaria:Alcyonacea]) was replaced by its oldest synonym, Prumna Motschulsky 1859. The current work reviews the authorship and availability of Primnoa and its type species. Additionally, it generally resolves the nomenclatural issues associated with these names, one of which includes providing Prumnini nom. nov. as a replacement name for the tribe Primnoini syn. nov. Subtribe Primnoae becomes a new synonym of Prumnini. Sharovia Otte 1997 is synonymized with Sharovella Gorochov 1985 and Archibrunnea Otte 1997 syn. nov. with Brunneus Hong 1983. Linhagla nom. nov. is provided as a replacement name for Parahagla Lin 1982 and Phyllophora pizaerosifolia nom. nov. replaces Ph. erosifolia Piza 1981, primary homonym.
Article
Pseudoxistrella eurymera, sp. et gen. nov is described from Hainan Island, South China.
Article
Characters used to diagnose the subfamily Metrodorinae (Bolívar, 1887) (Orthoptera: Tetrigidae) are largely variable or homoplastic. Relevant traits used to distinguish amongst subfamilies of Tetrigidae, including sensorial structures, mouth parts, and genitalia are not present in any description of representatives of the subfamily Metrodorinae. The identity of Metrodorinae remains thus at stake. Aiming to fill the current lack of knowledge, a complete anatomical description of adult and last instar nymphs of Allotettix simoni (Bolívar, 1890) is provided. Allotettix simoni is a controverted taxon, originally described in the genus Paratettix and subsequently synonymized with Allotettix peruvianus. The revision of type specimens revealed that A. simoni and A. peruvianus are different species. As neither keys nor descriptions of nymphs of Neotropical Tetrigidae exist, A. simoni nymphs were identified comparing mitochondrial sequences of the cytochrome c oxidase subunit I (COI) gene with adult specimens. The anatomical description of A. simoni expanded by comparisons with material of Metrodora sp. nov., revealed that the subfamilies Metrodorinae and Tetriginae are morphologically very similar, whereas molecular analyses of the COI data set do not support the monophyly of the subfamily Metrodorinae with respect to Tetriginae. Consequently Metrodorinae might not represent an independent entity with respect to Tetriginae. © 2012 The Linnean Society of London, Zoological Journal of the Linnean Society, 2012, 164, 52–70.
Article
Full-text available
The order Cnemidolestida (Insecta: Perlidea) is revised. A total of 11 families from the Carboniferous and Permian of Eurasia, North America, and South America and from the Triassic of Europe are included in the order: Cnemidolestidae Handlirsch, 1906, Spanioderidae Handlirsch, 1906, Gerapompidae Handlirsch, 1906, Cymenophlebiidae Pruvost, 1919, Emphylopteridae Handlirsch, 1922, Tillyardembiidae G. Zalessky, 1938, Psoropteridae Carpenter, 1976, Sylvabestiidae Aristov, 2000, Prygidae Aristov et Rasnitsyn, 2014, Neraphidiidae Aristov, fam. n., and Ctenoptilidae Aristov, fam. n. New synonymies are established: Cnemidolestidae = Aetophlebiidae syn. n., = Narkemidae syn. n., = Narkeminidae syn. n.; Emphylopteridae = Guloidae syn. n.; and Spanioderidae = Anthraconeuridae syn. n., = Carpenteropteridae syn. n. The family Cymenophlebiidae and genus Dieconeurites Handlirsch, 1906, previously considered synonyms, are re-established as valid. Two new genera (both from the Permian of Russia) are described in the family Sylvabestiidae: Ivkinus Aristov, gen. n. (type species Mezenalicula conjucta Aristov et Storozhenko, 2011 = Ivkinus conjuctus comb. n.) and Kityakia Aristov, gen. n. (type species Kityakia attrita Aristov, sp. n.); a new combination is proposed: Permula edemskii Aristov et Storozhenko, 2011 = Sojanopermula edemskii (Aristov et Storozhenko, 2011), comb. n. The new genus Anarkemina Aristov, gen. n. (type species Narkemina winsdoriensis Lewis, 1979 = Anarkemina winsdoriensis comb. n.) is described from the Carboniferous of the United States in the family Cnemidolestidae. Two new genera are included in the family Neraphidiidae Aristov, fam. n.: Fodinopenna Aristov, gen. n. (type species Fodinopenna abakanica Aristov, sp. n. from the Carboniferous of Russia) and Yaria Aristov, gen. n. (type species Yaria arenaria Aristov, sp. n. from the Permian of Russia). The family Ctenoptilidae Aristov, fam. n. includes a single genus, Ctenoptilus Lameere 1917 from the Upper Carboniferous of France.
Article
Full-text available
Vietnamese species of the genus Bolivaritettix Günther, 1939 are reviewed and illustrated. Altogether ten species are reported from Vietnam, of which five are new to science, namely—B. abruptus sp. nov., B. anaglypticus sp. nov., B. constrictus sp. nov., B. similis sp. nov., and B. vietnamensis sp. nov., while three species, B. sculptus (Bolívar, 1887), B. humeralis Günther, 1939 and B. celaenotus Zheng, Zhang et Dang, 2009, are recorded from Vietnam for the first time. A key to Vietnamese species is provided.
Article
Full-text available
A new genus of the orthopteran subfamily Batrachideinae from New Guinea is described: Wiemersiella gen. nov. The subfamily of the Batrachideinae is clearly characterized by the dorsal sulcated frontal and middle femur (fi g. 1). Outside the Neotropis (twelve genera) and Africa (two genera) we only know four genera of Batrachideinae. The difference between Palaioscaria Grant, 1966 (GRANT 1966), the only known genera from New Guinea, as well as Saussurella Bolívar, 1887 (BOLÍVAR 1887) and the new genus are the absent wings. In difference to Vingselina Sjöstedt, 1921 (SJÖSTEDT 1921) as well as Vilma Steinmann, 1973 (STEINMANN 1973) Wiemersiella, the new genus, has a rostrad upturned tip of the pronotum. The two new described species Wiemersiella highlandensis sp. nov. and Wiemersiella morobensis sp. nov. are differentiated by the the longer frontal tip of the pronotum and the broadened scutellum with curved lateral keels (fi gs 12 and 13). The disjunct distribution area for both species is shown in a map below.
Article
Full-text available
Taxonomic review of the genus Yunnantettix Zheng, 1995 is given. It is similar to Aspiditettix Liang, 2009, Pseudepitettix Zheng, 1995, and Epitettix Hancock, 1907. Yunnantettix is the most similar to Aspiditettix in the general appearance, the rugose disc of the pronotum, and the bisinuate lateral lobe of the pronotum, and diff erent from it by the completely reduced hind wing and the position of the antennal socket. Yunnantettix is similar to Pseudepitettix and Epitettix in the moderately widened frontal ridge and low median carina of pronotum, but diff erent from the latter by the presence of the narrow tegmen and a shallow yet distinct tegminal (upper) sinus on the pronotal lateral lobe. Originally, Yunnantettix is a monotypic genus (type species: Yunnantettix bannaensis Zheng, 1995 from South China). Two species are additionally included to this genus: Yunnantettix elytratus (Günther, 1939), comb. n. (= Epitettix elytratus Günther, 1939) from India and Yunnantettix thaicus, sp. n. from Thailand. The new species diff ers from all other species of this genus by the shallow excision on apex of the posterior pronotal process and by the external lateral pronotal carina, arch-like and strongly curved upward above the tegmen. A key to the species and redescription of the genus Yunnantettix are provided.
Article
The South American genus Diponthus Stål comprises species with cryptic and probably aposematic colorations. It belongs to a group of grasshoppers, tribe Romaleini, where color polymorphism is common. Morphometric techniques were used to clarify the taxonomic limits and status of six nominal species of Diponthus: D. permistus (Serville), D. argentinus Pictet & Saussure, D. pycnostictus Pictet & Saussure, D. communis Bruner, D. pictus (Bolivar) and D. schulzi Bruner. Results of cluster (CA) and principal component (PCA) analyses showed that D. pycnostictus can be defined by a unique combination of characters, while the remaining nominal species represent different color morphs of a single taxon, here recognized as D. argentinus. We also used statistical analyses to examine whether the discontinuities in body coloration of the polymorphic D. argentinus could be explained by geographic variation. The distribution patterns of body coloration of D. argentinus do not appear to be restricted to any particular biogeographic region and no character correlation resulted with latitude. Furthermore, the results show that color morphs display high levels of overlap across the geographic range, as well as intra-population variation. We conclude that in D. argentinus body coloration varies considerably across the geographic distribution range but without any noticeable color gradient correlated with the latitude. According to the phylogenetic species concept (PSC) we consider D. pycnostictus Pictet & Saussure and D. argentinus Pictet & Saussure as valid species. The following synonymies are proposed under D. argentinus: D. communis Bruner n. syn., D. schulzi Bruner n. syn., D. pictus (Bolivar) n. syn., and D. permistus (Serville) n. syn. We declare D. permistus (Serville) a nomen oblitum.
Article
Full-text available
Seven new species of the genus Zhengitettix Liang, 1994 are described: Z. hosticus sp. nov., Z. mucronatus sp. nov. and Z. spinulentus sp. nov. from Vietnam; Z. albitarsus sp. nov. and Z. extraneus sp. nov. from Thailand; Z. palawanensis sp. nov. and Z. taytayensis sp. nov. from the Philippines. Two species, Z. curvispinus Liang, Jiang et Liu, 2007 and Z. obliquespicula Zheng et Jiang, 2005 are firstly recorded from Vietnam. An annotated check-list and key to species of the genus Zhengitettix are given. Position of Zhengitettix within the family Tetrigidae is briefly discussed.
Article
Full-text available
Two species of the genus Austrohancockia are found in Vietnam. A. orlovi sp. nov. is described from northern and central parts of this country. A key to Vietnamese species of Austrohancockia is provided.
Article
Full-text available
The genus Xistrella Bolívar, 1909 (Orthoptera: Tetrigidae) is taxonomically reviewed. It can be easily recognised from other genera in SE Asia, and especially in the People’s Republic of China. It is distinctive by antennae inserted below the inferior margin of eye and considerably long; pronotum between the shoulders strongly elevated to obtuse gibbosity or slightly elevated in the posterior part of shoulders. This genus is distributed mainly in China, India, Indonesia, Sri Lanka, Myanmar, Nepal, Malaysia, and the Philippines. One new species, X. hainanensis Deng sp. nov. (China: Hainan), is described, and a new record of X. dubia (Brunner von Wattenwyl, 1893) from China is given. An annotated identification key to all species of the genus Xistrella is provided.
Article
Full-text available
Nine species of the genus Hebarditettix are found in Vietnam; H. chamensis sp. nov., H. fuscus sp. nov., H. gibbus sp. nov., H. intermedius sp. nov., H. magnus sp. nov., and H. oculatus sp. nov. are described from this country. A key to Vietnamese species of Hebarditettix is provided.
Article
Taxonomy and bio-ecology of Tetrigidae in Cameroon remain poorly studied. This study reports for the first time the presence of Thibron cf. tectatus Günther, 1979 in Cameroon and provides morphological and bio-ecological characteristics of Thibron illepidus (Karsh, 1893) and T. cf. tectatus. The specimens studied were collected between February 2016 and Febru-ary 2018 in three regions of Cameroon. In each region, species were collected monthly using sweep-nets and pitfalls in four vegetation types. Some morphological characters formerly omitted in previous descriptions were considered in this study. In addition to the characters already known, T. illepidus differs from other species of the same genus in terms of length of body and interocular space, posterior thigh width, median femur length and width and finally the length of the three tarsomeres of posterior tarsus only in T. illepidus. In addition, for T. cf. tectatus, the length of the head, fastigium of vertex, eyes are longer than in T. illepidus. The later was more abundant in the Centre region (79%), less abundant in the Southwest region (18%) and rare in the West (3%). Meanwhile, Thibron cf. tectatus is rare in the Centre region (4%). Pitfalls (92%) were more effective than sweep-nets (8%) in capturing T. illepidus. Results obtained could contribute to improve the knowledge of the taxonomy of both species in Cameroon.
ResearchGate has not been able to resolve any references for this publication.