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Contribution à la connaissance des Staphylinidae. XLVI. Sur quelques espèces de Berbérie

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... GÉNERO TIPO: Bledius Leach, 1819 Género Bledius Leach, 1819 (35 sp.,1ssp.) ESPECIE TIPO: B. tricornis (Herbst, 1784) Subgénero Astycops Thomson (Coiffait, 1982), Trogophloeus =evertsi (Bernhauer, 1915), Trogophloeus =lasti (Scheerpeltz, 1946), Trogophloeus =lindbergi (Scheerpeltz, 1937) boops ssp rondaensis (Fagel, 1957), andalusiacus (Fagel, 1957), Ancyrophorus =rivularis (Smetana, 1967) (Fagel, 1951), Ancyrophorus =ruteri (Jarrige, 1949), Ancyrophorus PENÍNSULA IBÉRICA: Primera cita en la península de Gerona por MAKRANCZY (2014) DISTRIBUCIÓN: Europea rosenhaueri (Kiesenwetter, 1850), Trogophloeus =corsicus (Fagel, 1956), Ancyrophorus =gracilis (Fagel, 1951), Ancyrophorus =jailensis (Scheerpeltz, 1950) (Rosenhauer, 1856), ...
... GÉNERO TIPO: Bledius Leach, 1819 Género Bledius Leach, 1819 (35 sp.,1ssp.) ESPECIE TIPO: B. tricornis (Herbst, 1784) Subgénero Astycops Thomson (Coiffait, 1982), Trogophloeus =evertsi (Bernhauer, 1915), Trogophloeus =lasti (Scheerpeltz, 1946), Trogophloeus =lindbergi (Scheerpeltz, 1937) boops ssp rondaensis (Fagel, 1957), andalusiacus (Fagel, 1957), Ancyrophorus =rivularis (Smetana, 1967) (Fagel, 1951), Ancyrophorus =ruteri (Jarrige, 1949), Ancyrophorus PENÍNSULA IBÉRICA: Primera cita en la península de Gerona por MAKRANCZY (2014) DISTRIBUCIÓN: Europea rosenhaueri (Kiesenwetter, 1850), Trogophloeus =corsicus (Fagel, 1956), Ancyrophorus =gracilis (Fagel, 1951), Ancyrophorus =jailensis (Scheerpeltz, 1950) (Rosenhauer, 1856), ...
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[Publicado originalmente en Boletín Sociedad Entomológica Aragonesa nº 43 (2008): 233-254]. ACTUALIZACIONES disponibles en web: http://bba.bioucm.es/cont/docs/RO_27.pdf , http://bba.bioucm.es/cont/docs/RO_94.pdf Actualizado hasta noviembre de 2022.
... DISTRIBUCIÓN: Hispánica asturiensis (Fagel, 1967), Sipalia =convadongaensis (Fagel, 1967) (Normand, 1945) Género Cantabrodytes Español, 1975 (1 sp) ESPECIE TIPO: C.vivesi Español, 1975 vivesi Español, 1975 Subgénero Adexiopisalia Pace, 1982ESPECIE TIPO: L. hummeleri Bernhauer, 1910 hummeleri Bernhauer, 1910 =hummeleri advena Pace, 1981 PENÍNSULA IBÉRICA: ...
... DISTRIBUCIÓN: Hispánica asturiensis (Fagel, 1967), Sipalia =convadongaensis (Fagel, 1967) (Normand, 1945) Género Cantabrodytes Español, 1975 (1 sp) ESPECIE TIPO: C.vivesi Español, 1975 vivesi Español, 1975 Subgénero Adexiopisalia Pace, 1982ESPECIE TIPO: L. hummeleri Bernhauer, 1910 hummeleri Bernhauer, 1910 =hummeleri advena Pace, 1981 PENÍNSULA IBÉRICA: ...
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El primer catálogo recogía datos hasta el 2005. En 2009 se realizó la primera actualización de la subfamilia y en 2014 la segunda. Desde entonces hasta el año 2019 se han producido una serie de modificaciones en los datos de esta subfamilia en la península Ibérica, siendo debidos a revisiones realizadas de diversos géneros (ASSING, GUSAROV, MAUS, TRONQUET) o a la publicación de resultados de la biodiversidad en diversas zonas (Andalucía: LÓPEZ-PÉREZ, GAMARRA et al., OUTERELO & GAMARRA; BALEARES: PINYA et al., PONS et al., RIDDIFORD et al.; Castilla León: PENIN; Cataluña: VIÑOLAS et al.; Galicia: NOVOA et al., PÉREZ-VALCÁRCEL et al.; La Rioja: PÉREZ-MORENO & MORENO, PÉREZ-MORENO et al.; Navarra: OUTERELO et al.) o resultados de muestreos en medios concretos como cuevas (ASSING, BELLÉS, GINÉS, GONZÁLEZ, PÉREZ, PÉREZ et al., VADELL et al.) o carroña (BEGOÑA, DÍAZ, GARCÍA et al., PÉREZ-BOTE et al.) o setas (OUTERELO et al.). Además, se han conseguido publicaciones anteriores al año 2014 como las de QUEDENFELDT, 1881 y 1884. Se incorporan aquellos cambios taxonómicos recogidos por SCHÜLKE & SMETANA en el capítulo de Staphylinidae en el Catalógo Paleártico editado por LÖBL & LÖBL (2015). Actualmente la subfamilia comprende 86 géneros y 548 especies. Las principales novedades en la presente actualización son: Las principales novedades en la presente actualización son: Erratas detectadas en la publicación original y su primera actualización: En anteriores versiones se recogía Aleochara (s.str.) haematodes Kraatz, 1862 citada de este modo de Portugal (DE LA FUENTE, 1924), en realidad la especie fue descrita por Schaum y es sinonimia de Aleochara (s.str.) haematoptera Kraatz, 1858, esto ya se recogía en las anteriores versiones. Geostiba (Sibiota) hispalensis Pace, 1990 se marcaba en el mapa Málaga SCHÜLKE & SMETANA, en su catálogo pone Sevilla: Sierra de Grazalema (sic) cuando esta sierra se localiza entre Málaga y Cádiz. PACE (1990) localiza la especie en Andalousie, Sierra Grazalema, sin más datos. Por tanto en los mapas marcamos Málaga y Cádiz. Geostiba (Tropogastrosipalia) consobrina Assing, 2003 se marcaba en el mapa Estremadura siendo lo correcto Beira Alta. La especie Leptotheta blascoi Tronquet, 2002 en las versiones anteriores se recogía por error como L. thamiaraea Tronquet, 2002. Tectusa estrelae Assing, 2003 se marcaba en el mapa Estremadura cuando tenía que ser Baixo Alentejo. ii Especies y subespecies descritas nuevas para la ciencia. Alevonota alboculata Assing, 2018 de Murcia Aloconota sulcifrons ssp. cantabrica Tronquet, 2014 de Santander Anaulacaspis iberica Assing, 2016 de Pontevedra y Jaén. Cousya struyvei Assing, 2018 de Jaén. Oxypoda (Bessopora) caespita Assing, 2018 de Granada (Sierra Nevada) Especies citadas nuevas para la Península Ibérica. Cypha rubripennis (Pandellé, 1869) por ASSING (2014b) de Murcia. Euryusa sinuata Erichson,1837 por PÉREZ-MORENO & MORENO-GRIJALBA (2009), de La Rioja. Gyrophaena (Phaenogyra) polita (Gravenhorst, 1802) por OUTERELO, GAMARRA & PAGOLA-CARTE. (2018), de Guipuzkoa. Gyrophaena (s.str.) nana (Paykull, 1800) por OUTERELO, GAMARRA & PAGOLA-CARTE. (2018), de Guipuzkoa Oxypoda pellax (Peyerimhoff,1919) por ASSING (2018d) de Jaén. Phloeopora teres (Gravenhorst, 1802) por PÉREZ-MORENO & MORENO-GRIJALBA (2009), de La Rioja. Placusa (Placusa) atrata (Mannerheim, 1830) por OUTERELO, GAMARRA & PAGOLACARTE. (2018), de Guipuzkoa. Especies citadas nuevas para España. Oxypoda (Mycetodrepa) alternans (Gravenhorst, 1802) por OUTERELO, GAMARRA, SAN MARTÍN & RECALDE (2016b) de Navarra. Especies citadas de “España” en el Catálogo de Schülke & Smetana. Estos autores en algunas especies hemos encontrado que en su distribución citan España y nosotros no tenemos ninguna referencia para poder concretar dicha cita genérica. Atheta (Microdota) boreella Benick, 1981 Atheta (Mocyta) negligens (Mulsant & Rey, 1873) Atheta (s.str.) hypnorum (Kiessenwetter, 1850) Atheta (s.str.) laevicauda Sahlberg, 1876 Blepharhymenus (Blepharhymenus) paulinoi (Skalitzky, 1884), Brundinia splendens Benick, 1940 Diglotta sinuaticollis (Mulsant & Rey, 1870) Gnypeta rubrior Tottenham, 1939 Gyrophaena (Phaenogyra) strictula Erichson, 1839 Leptusa (Gallopisalia) lapidicola Brisout de Barneville, 1863 Notothecta (s.srt.) flavipes (Gravenhorst, 1806) Oligota (s.str.) granaria Erichson, 1837 Oxypoda (Bessopora) ambigena Fauvel, 1869 Phytosus (Actosus) nigriventris (Chevrolat, 1843) Especies citadas de “Portugal” en el Catálogo de Schülke & Smetana. Estos autores en algunas especies hemos encontrado que en su distribución citan Portugal y nosotros no tenemos ninguna referencia para poder concretar dicha cita genérica. Atheta (Mycetota) mucronata (Kraartz, 1859) Atheta (Mycetota) consueta (Mulsant & Rey, 1873 Oligota (Holobus) apicatus (Erichson, 1837), Ocalea skalitzkyi Bernhauer, 1902 Oxypoda (Bessopora) caespita Assing, 2003 Phloeopora teres (Gravenhorst, 1802) Phloeopora testacea (Mannerheim, 1830) Dilacra luteipes (Erichson, 1837) Gnypeta rubrior Tottenham, 1939 iii Especies de las que se concreta su distribución en la Península Ibérica. Placusa (Placusa) pumilio (Gravenhorst, 1802). Material identificado, por los autores. de Cádiz. Placusa (Placusa) pumilio (Gravenhorst, 1802). Material identificado, por los autores. de Cádiz. Thamiaraea hospita (Märkel, 1845),.Material identificado de Cádiz. Thiasophila angulata (Erichson,1837) Material identificado, por los autores, de Lleida. Cambios taxonómicos que afectan a la fauna de la Península Ibérica. En el Catálogo de SCHÜLKE & SMETANA el subgénero de Atheta (Mocyta) es resucitado dejando de ser sinonimia de Acrotona y algunas especies que se incluían en el género Acrotona pasan al subgénero Mocyta: Ath. clientula (Erichson, 1840), Ath. fungi (Gravenhorst, 1806), Ath. haematica (Eppelsheim, 1884), Ath. orbata (Erichson, 1837) y Ath. orphana (Erichson, 1839). En el mismo Catálogo el Género Alaobia es considerado subgénero de Atheta. El Género Dimetrota es considerado subgénero de Atheta. El subgénero Derocala de Oxypoda se eleva a género. Muchas especies cambian de subgénero. Acrotona fuscipes (Heer, 1839) se sinonimiza con Liprocorrhe anceps (Erichson, 1837) Aleochara peezi Scheerpeltz, 1957 se sinonimiza con A. lanuginosa Gravenhorst, 1802 Atheta. complana (Mannerheim, 1831) se considera nomen dubium.y la especie válida pasa a ser Ath. picipes (Thomson, 1856). Atheta macroptera Bernhauer, 1913 se sinonimiza con Ath.(Alaobia) spelaea (Erichson, 1839) Atheta (s.str.) pertyi (Heer, 1839) se sinonimiza con Ath.castanoptera (Mannerheim, 1830) Atheta (Microdota) steineri Scheerpeltz, 1958 sinonimia de Ath. luctuosa (Mulsant & Rey, 1853). Cousya nitidiventris Fagel, 1958 sinonimia de Cousya nigrata (Fairmaire, 1854) Oxypoda ambigena Fauvel, 1869 deja de ser considera sinonimia de Oxypoda (Bessopora) haemorrhoa (Mannerheim, 1830) Oxypoda (Bessopora) filiformis Redtembacher, 1848 de O. (B.) ferruginea Erichson, 1840 Oxypoda (Bessopora) telifera Assing, 2008 sinonimia de O. (B.) determinata Scriba, 1870 Oxypoda (Bessopora) virgata Assing, 2008 pasa ser sinonimia de Oxypoda (Bessopora) hispanica Bernhauer, 1914. Especies que causan baja en el catálogo de la península Ibérica. Dimetrota picipennis (Mannerheim, 1843) es sustituida por hasta ahora su sinonimia Dimetrota aeneipennis (Thomson, 1856), ya que GUSAROV (2003) considera que D.picipennis tiene una distribución Holártica mientras que las citas paleárticas son en realidad de D.aeneipennis. Localidades del material identificado por los autores. [en gris en los mapas] que implica ampliación de la distribución en la Península Ibérica. A Coruña (Meirama, Cerceda) Juan Carlos Otero leg. A Coruña Juan Carlos Otero leg. Almería (Isla de San Andrés, Carboneras). M.A. Gómez de Dios leg. Almería (La Mojonera) MªD. Alcáraz leg. Almería (Peñón de Bernal, El Egido). M.A. Gómez de Dios leg. Almería (Pista de la Tetica, Bacares, Sierra de Filabres). M.A. Gómez de Dios leg. Almería (Velez Blanco) ). M.A. Gómez de Dios leg. Cádiz (Los Barrios) Antonio Verdugo leg. Cádiz (Los Barrios) José Luis Torres leg. Cádiz (Valdeinfierno) Antonio Verdugo leg. iv Girona (La Molina) G. Zsolt Muláes leg. Granada (Cenes de la Vega, Güejar Sierra, Huetor Sierra y Hoya de la Mora en Sierra Nevada) (H. Hernando & D. Hita leg.) Granada (Dehesa del Generalife y Deifontes) M. Morente, F. Ruano, M.Campos y A. Tinaut leg. Guadalajara (Sigüenza) G.González-Jurado leg. Guipuzkoa (Parque Natural Aiako Harria) S.Pagola-Carte leg. Huesca (Altorricón, Tamaite de Llitera, Fraga) Oscar Alomar leg. La Rioja (Muro en Cameros) Pérez-Moreno, San Martin y Recalde leg. Lleida (Aitona, Alcarrás, Almenar, Gimenells, Sunyer, Torres de Segre) Oscar Alomar leg. Navarra (El carrascal) S,Pérez Martínez Pontevedra (Moscoso) R.Outerelo leg. Pontevedra Mondariz, O Cabalo, O Cabo da Raña, Soutomaior (María Calviño leg.) Valladolid (Laguna de Duero) Adela López leg.
... This species is confined to Asturias and Galicia in Spain (Fagel 1967;Coiffait 1982 (s. str.) lohaji Anlaş, 2023 ...
... Neobisnius prolixus (Erichson, 1840) Horion, 1965; (Ganglbauer, 1905;Smetana, 1953;Öncüer, 1991;Kesdek et al., 2009; Kesdek et al., 2009; (Smetana, 1953;Kesdek et al., 2009 (Smetana, 1953;Fagel, 1963;Abacıgil et al., 2013;Kesdek et al., 2009 Smetana, 1953Öncüer, 1991;Kesdek et al., 2009;Abacıgil et al., 2013). ...
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The focus of this study is to provide the faunistic, ecological, and zoogeographical status of Staphylininae (Coleoptera: Staphylinidae) fauna of the Central Anatolian Region of Turkey. The study material comprises specimens collected from field studies between 2009 and 2012 and also specimens that were previously collected by other researchers. In total, 90 taxa of 26 genera belonging to 3 tribes of Staphylininae were recorded. Number of specimens, collection habitats and methods, months, and altitudes are given in a table and figures for collected species. Distributions of species in other regions of Turkey and zoogeographical regions are also given in the table and figures for all species. Zoogeographical statuses of all species are discussed.
... Hemos encontrado dos especies que aparecen citadas erróneamente como pertenecientes a la fauna Ibérica. Achenium brevipenne Quedenfeldt, 1881, del Sur de España por Coiffait (1982 al considerar como Península Ibérica Marruecos español" y Scopaeus biskrensis Fagel, 1957 descrita de Argelia apareciendo citada por error de España por Frisch (2002) y posteriormente por Smetana (2004). ...
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Resumen: Se presenta el catálogo actualizado hasta el año 2007 de los Staphylinidae Staphylininae iberobaleares, tomando como punto de partida la obra de J.M. de La Fuente (1922). En total se incluyen 298 especies (con 6 subespecies), 38 géne-ros, cuatro subtribus y cuatro tribus. Se recogen 937 sinónimos. De cada una de las especies y subespecies se indica la distri-bución iberobalear mediante mapas, y se indican las categorías mundiales de su distribución. Se citan nueve especies como nuevas para la Península Ibérica, y tres para España. Se concreta la presencia de cinco especies en España y la localización de otras tres. Catalogue of the Ibero-Balearic Staphylininae (Coleoptera: Staphylinidae) Abstract: A catalogue of the Ibero-Balearic Staphylinidae Staphylininae is presented, updated to 2007 and with J.M. de la Fu-ente (1922) as the starting point. 298 species (with 6 subspecies), 38 genera, 4 subtribes and 4 tribes are included. 937 synonyms are given. For each species and subspecies there is a map with its Ibero-Balearic distribution, and information is also given on its world distribution. 9 species are newly recorded from the Iberian Peninsula, and 3 from Spain. The presence of 5 species in Spain is confirmed, and the distribution of a further 3 is established.
... Gümüşhane (Zigana) [34]. [5]. ...
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This study was carried out with excursions which species belonging to the family Staphylinidae (Coleoptera) were collected in May-July, 2003, May-August, 2008 and April-June, 2009 in Ankara province. As a result of the study, 28 species belonging to 15 genera and five subfamilies (Aleocharinae, Omaliinae, Oxytelinae, Tachyporinae, Paederinae) were identified. Sampling localities, distributions of the species in Turkey and World are given. Specimen numbers, vertical distributions, collecting months and habitats, distributions of species in other geographical regions of Turkey and Zoogeographic regions are shown. According to present literature 18 of all detected species are new records for Ankara and Mycetoporus baudueri Mulsant & Rey, 1875 is a new record for Turkey.
... Ankara, Antalya, Aydın, Balıkesir, Denizli, Hatay (Antakya), İstanbul, Izmir, Kahramanmaraş, Konya, Manisa, Mersin, Muğla, Niğde (Fagel, 1970;Bordoni, 1980;Assing, 2004Assing, , 2004aAssing, , 2006Assing, , 2007bLöbl & Smetana, 2004;Anlaş & Çevik, 2008). ...
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In this study, Staphylinidae species collected by light trap method in Kemalpaşa district, Izmir province, Western Turkey during the years of 2005-2007 have been evaluated. A total of 16 species belonging to 13 genera have been recorded. Among those seven species [Tachyporus nitidulus (Fabricius, 1781), Aleochara laticornis Kraatz, 1856, Aleochara tristis Gravenhorst, 1806, Anotylus inustus (Gravenhorst, 1806), Carpelimus bilineatus Stephens, 1834, Carpelimus insularis (Kraatz, 1858), Homaeotarsus chaudirii Hochhuth, 1851] were reported for the first time from Aegean Region of Turkey.
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The revision of the subfamily Paederinae (Coleoptera: Staphylinidae) of the Portuguese fauna is concluded with this contribution, in which data of 69 species within 22 genera belonging to eight subtribes are provided. In this article, one genus and 10 species are newly recorded from Portugal, while three of them are in need of confirmation. Resumo: Sobre alguns géneros de Paederinae da fauna portuguesa (Coleoptera: Staphylinidae: Paederinae). A revisão da subfamília Paederinae (Coleoptera: Staphylinidae) da fauna portuguesa fica terminada com esta contribuição, na qual se procuram dados de 69 espécies distribuídas em 22 géneros pertencentes a oito subtribos. Neste artigo, um género e 10 espécies são assinalados como novidade para Portugal, enquanto três delas aguardam confirmação.
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The generic classifications of the paederine subtribes Scopaeina Mulsant and Rey, 1878, and Sphaeronina Casey, 1905, are revised. Sphaeronina, revised status, is resurrected from synonymy. Keys to the included genera of both subtribes are included. Newly discovered characters in both subtribes are discussed and illustrated. The Scopaeina now includes Scopaeus, Hyperscopaeus, Micranops, Orus, and Trisunius. The account for each genus includes its diagnostic characters, a description, summary of the general distribution, and list of the included species and specimens examined. Scopaeus Erichson, 1839, has a revised definition and is now restricted to species that have not only a constricted neck and a trichobothrium adjacent to and at about the middorsal margin of the eye, but also a metathoracic/mesofemoral stridulum comprised of a lateral, metaventral file and mesofemoral plectral ridges, slender, apically acute, metakatepisternal processes, and a middorsally fused median lobe of the aedeagus. The stridulum, redefined herein as a file and plectrum that when rubbed together produce stridulation in insects. The metaventral file and mesofemoral plectral ridges of Scopaeus, is, heretofore, unknown in the Staphylinidae or perhaps, even the Coleoptera. Variations of the stridulum and metakatepisternal processes are illustrated and described for each species group. Five genus-group names in the Western Hemisphere, Scopaeomerus Sharp, 1886, and Euscopaeus Sharp, 1886, are new synonyms of Scopaeus; Scopaeodera Casey, 1886, Scopaeoma Casey, 1905, and Scopaeopsis Casey, 1905, are revised status junior synonyms of Scopaeus. The species in those generic groups are now included in species groups of Scopaeus. Hyperscopaeus Coiffait, 1984, new status, is elevated to genus from subgeneric status in Scopaeus. Trisunius Assing, 2011, new subtribal assignment, is moved from the Medonina to the Scopaeina. Typhloscopaeus Jarrige, 1951, incertae sedis, formerly a subgenus of Scopaeus, is of unknown placement, but the species and generic names are retained in Scopaeus awaiting study of the type. Orus cervicula Casey, 1905, revised combination, is returned to Orus from Scopaeus. Orus femoralis (Sharp, 1887), new combination, is transferred from Scopaeus. There are now three named species of Orus with narrow necks. Scopaeus chiriquensis (Sharp, 1886), S. guatemalensis (Sharp, 1886), S. obscurus (Sharp, 1886), and S. palmatus (Sharp, 1886), new combinations, are transferred to Scopaeus from Scopaeomerus. Medon mexicanus (Bernhauer, 1910), new combination, is transferred to Medon from Scopaeomerus. Scopaeus crassitarsis (Sharp, 1886), S. gracilicornis (Sharp, 1886), S. impar (Bierig, 1935), new combinations, are transferred to Scopaeus from Euscopaeus. The following names are transferred from Scopaeus to Hyperscopaeus as new combinations: Hyperscopaeus admixtus (Fagel, 1973), H. albertvillensis (Fagel, 1973), H. allardianus (Fagel, 1973), H. andrewesi(Cameron, 1931), H. angolanus (Fagel, 1973), H. bamaniaensis (Fagel, 1973), H. borneensis (Cameron, 1941), H. bredoanus (Fagel, 1973), H. calidus (Bernhauer, 1932), H. confusoides (Fagel, 1973), H. confusus (Fagel, 1973), H. consimilis (Fagel, 1973), H. convexiceps (Bernhauer, 1932), H. corpulentus(Fagel, 1973), H. decelleanus (Fagel, 1973), H. dolosus (Fagel, 1973), H. endrodyanus (Fagel, 1973), H. errans (Fagel, 1973), H. erraticus (Fagel, 1973), H. fageli (Levasseur, 1981), H. fallaciosus (Fagel, 1973), H. filicornis (Fagel, 1973), H. flavidulus (Fagel, 1973), H. flavocastaneus (Lea, 1923), H. fluviatilis (Fagel, 1973), H. fossiceps (Eppelsheim, 1885), H. fuliginosus (Fagel, 1973), H. fulvescens (Motschulsky, 1858), H. fusculus (Motschulsky, 1858), H. gigantulus (Bernhauer, 1929), H. girardianus (Fagel, 1973), H. hova (Fauvel, 1905), H. hulstaertianus (Fagel, 1973), H. intermixtus (Fagel, 1973), H. kaszabianus (Fagel, 1973), H. katanganus (Fagel, 1973), H. kivuanus (Fagel, 1973), H. lamtoensis (Fagel, 1973), H. leleupianus (Fagel, 1973), H. leopoldvillensis (Fagel, 1973), H. lescuyeri (Delaunay, Coache, and Rainon, 2019), H. levasseuri (Lundgren, 1982), H. longiusculus (Fagel, 1973), H. machadoanus(Fagel, 1973), H. major (Eppelsheim, 1885), H. methneri (Bernhauer, 1932), H. minutulus (Fagel, 1973), H. mulongoensis (Fagel, 1973), H. nitidiceps (Fagel, 1973), H. nitidicollis (Fagel, 1973), H. opacicollis (Bernhauer, 1942), H. overlaetianus (Fagel, 1973), H. parvicornis (Fauvel, 1900), H. procerus (Kraatz, 1859), H. pruinosulus (Eppelsheim, 1885), H. pseudomethneri (Fagel, 1973), H. puberulus (Kraatz, 1859), H. reduncus (Fagel, 1973), H. ripicola (Fagel, 1973), H. rubricollis (Fagel, 1973), H. rubrotestaceus (Kraatz, 1859), H. ruguliceps (Fagel, 1973), H. ruziziensis (Fagel, 1973), H. semifuscus (Kraatz, 1859), H. senegalensis (Fagel, 1973), H. seydeli (Cameron, 1952), H. simillimus (Fagel, 1973), H. simulator (Fagel, 1973), H. spathiferus (Coiffait, 1970), H. spinosophallatus (Frisch, 2012), H. subconfusus (Fagel, 1973), H. subprocerus (Coiffait, 1978), H. surdus (Fagel, 1973), H. suspectus (Fauvel, 1907), H. tchapembanus (Fagel, 1973), H. thoracicus (Motschulsky, 1858), H. tristis (Bernhauer, 1929), H. vagans (Fagel, 1973, and H. voltae (Fagel, 1973). Sphaeronina, revised status, is resurrected from synonymy and now includes Sphaeronum Sharp, 1876, Tripectenopus Lea, 1918, Typhloleleupius Fagel, 1964, and Coecoscopaeus Coiffait, 1982; the last three genera are new assignments to the subtribe. Sphaeronina is redefined by the presence of a hypopharyngeal peg, an enlarged protibial concavity with combs, a ventral denticle on the left mandible, and a groove on the outer edge of the mandibles; additional possible diagnostic characters are discussed. Sphaeronum, Tripectenopus, Typhloleleupius, and Coecoscopaeus are redescribed; the genera are found, respectively, in the American tropical and subtropical regions, Australia, southern Africa and perhaps Madagascar, and Tunisia. Few African and Australian were available for study. Scopaeodracus Scheerpeltz, 1935, is a new synonym of Tripectenopus. Tripectenopus handschini (Scheerpeltz, 1935), new combination, is transferred from Scopaeodracus; Tripectenopus australiae (Fauvel, 1878), T. microps (Lea, 1923), T. pectinatrix (Lea, 1923), and T. torrensensis (Blackburn, 1891), new combinations, are transferred from Domene.
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Publicado originalmente en Boletín Sociedad Entomológica Aragonesa nº 37(2005): 1-81 http://bba.bioucm.es/cont/docs/143.pdf. Primera actualización en el 2018: http://bba.bioucm.es/cont/docs/RO_61.pdf Y esta actualización es hasta AGOSTO del 2023. (PDF) SEGUNDA actualización del CATÁLOGO IBEROBALEAR DE LOS TACHYPORINAE (Coleoptera: Staphylinidae). Available from: https://www.researchgate.net/publication/374548559_SEGUNDA_actualizacion_del_CATALOGO_IBEROBALEAR_DE_LOS_TACHYPORINAE_Coleoptera_Staphylinidae [accessed Oct 09 2023].
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The focus of this study is to make contributions to Staphylininae (Coleoptera: Staphylinidae) fauna of Turkey with additional material which was composed of specimens collected from Eastern Black Sea Region between 2013 to 2015. Totally, 71 taxa of 19 genera belonging to 3 tribes of Staphylininae were recorded. Among them, Ocypus almensis Coiffait, 1967 and Quedius edmundi Coiffait, 1969 are collected from Turkey for the first time. Moreover, first detailed localities for Philonthus picimanus (Mntris, 1832), Philonthus splendens sideropterus Kolenati, 1846 and Tasgius globulifer (Geoffroy, 1785) are given with this study. General distribution pattern of the species depending on collecting months, altitudes and habitats, and species diversity of the region are presented and compared. Distribution of species in other regions of Turkey and zoogeographical regions are included, and the zoogeographical statuses of species are discussed. European and Middle Eastern species compose the most of the collected species. Since the region is transition between Caucasia and Europe through the Anatolian peninsula, species composition reflects the regional characters, and consequently, represents this pathway.
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Here we report on the faunistic, ecological and zoogeographical composition of the subfamily Steninae (Coleoptera: Staphylinidae) of the Eastern Blacksea Region of Turkey. Forty two species, belonging to two genera, were detected by field studies between years 2013-2015 in the Eastern Blacksea Region. It is concluded that among these, five species from Stenus are new records for Turkey. Four species are endemic for Turkey and seven species are new records for Eastern Blacksea Region.
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Die Typen und weiteres Material der Arten der Gattungen Callicerus Gravenhorst, Pseudosemiris Machulka und Saphocallus Sharp werden revidiert. Auf der Grundlage eines morphologischen Vergleichs unter Berücksichtigung der wahrscheinlich nahverwandten Taxa Aloconota Thomson, 1858 und Disopora Thomson, 1859 werden Callicerus, Pseudosemiris und Saphocallus als distinkte Gattungen betrachtet und redeskribiert. 7 valide Callicerus-, 7 Pseudosemiris-Arten sowie eine Art der Gattung Saphocallus werden erkannt und beschrieben: C. obscurus Gravenhorst, C. muensteri Bernhauer, C. atricollis (Aubé), C. appenninus sp. n., C. sparsicollis Bernhauer, C. fulvicornis Eppelsheim, C. rigidicornis (Erichson), P. kaufmanni (Eppelsheim), P. breiti Scheerpeltz, P. circassica Fagel, P. fulgida sp. n., P. granulosa Fagel, P. stricticornis Fagel, P. zanettii sp. n. und S. parviceps Sharp. P. velox Jablokow-Khnzorian wird als species dubia betrachtet. Die folgenden Taxa werden synonymisiert: Callicerus Gravenhorst, 1802 = Semiris Heer, 1839, resyn., = Sphaerotaxus Bernhauer, 1915, syn. n.; Callicerus obscurus Gravenhorst, 1802 = Callicerus stolfai Scheerpeltz, 1956, syn. n., = Callicerus ibericus Fagel, 1958, syn. n.; Calodera atricollis Aubé, 1850 = Callicerus clavatus Rottenberg, 1870, syn. n.; Callicerus fulvicornis Eppelsheim, 1883 = Callicerus gagliardii Scheerpeltz, 1956, syn. n.; Homalota rigidicornis Erichson, 1839 = Callicerus mandli Scheerpeltz, 1956, syn. n., = Callicerus beieri Scheerpeltz, 1959, syn. n.; Homalota gregaria Erichson, 1839 (nomen protectum) = Aleochara caliginosa Stephens, 1832, syn. n. (nomen oblitum); Atheta montenegrina Bernhauer, 1899 = Callicerus smetanai Scheerpeltz, 1967, syn. n. Atheta toroenensis Bernhauer, 1943, die zuvor Callicerus zugeordnet worden war, wird in die Gattung Homoiocalea Bernhauer, 1943, stat. n. gestellt. Sowohl die Gattung als auch H. toroenensis (Bernhauer), comb. n. werden redeskribiert. Für Callicerus obscurus Gravenhorst wird ein Neotypus, für Calodera atricollis Aubé, Callicerus muensteri Bernhauer, C. obscurus var. pedemontanus Baudi, C. clavatus Rottenberg, C. sparsicollis Bernhauer, C. fulvicornis Eppelsheim und C. kaufmanni Eppelsheim werden Lectotypen designiert. Die Differentialmerkmale aller untersuchter Gattungs- und Artengruppentaxa sowie die Ergebnisse morphometrischer Analysen werden illustriert. Die Diagnosen werden durch Angaben zur intraspezifischen Variabilität und zur Unterscheidung von ähnlichen Arten sowie durch Bestimmungstabellen für die Arten der Gattungen Callicerus und Pseudosemiris ergänzt. Die verfügbaren Daten zur Verbreitung und Bionomie werden ausgewertet. Die Arten der Gattungen Callicerus und Pseudosemiris sind danach univoltin; die Fortpflanzungsperiode beginnt im Frühjahr, und die Präimaginalentwicklung ist im Frühsommer abgeschlossen. Reproduktion und Überwinterung finden offenbar in einem unterirdischen, bisher aber unbekannten Habitat statt. Dies würde erklären, warum praktisch alle Arten selten bis extrem selten nachgewiesen wurden. Die Phänologien von C. obscurus und C. rigidicornis werden durch Diagramme veranschaulicht. Verbreitungskarten illustrieren die Verbreitungsgebiete der weniger seltenen Arten.StichwörterColeoptera, Staphylinidae, Aleocharinae, Athetini, Callicerus, Pseudosemiris, Saphocallus, Homoiocalea, Aloconota, Palaearctic region, Europe, taxonomy, biogeography, ecology, intraspecific variation, life history, new species, new synonymy, new combination, neotype designation, lectotype designation.Nomenklatorische Handlungengregaria (Erichson, 1839) (Aloconota), nom. protectum described as Homalota gregariamontenegrina (Bernhauer, 1899) (Aloconota (Disopora)), Lectotype described as Atheta (Disopora) montenegrinacaliginosa Stephens, 1832 (Bolitochara), nom. oblitum; syn. n. of Aloconota gregaria (Erichson, 1839): nom. protectumappenninus Assing, 2001 (Callicerus), spec. n.atricollis (Aubé, 1850) (Callicerus), Lectotype described as Calodera atricollisclavatus Rottenberg, 1870 (Callicerus), Lectotype; syn. n. of Callicerus atricollis (Aubé, 1850)fulvicornis Eppelsheim, 1883 (Callicerus), Lectotype described as Callicerus atricollis var. fulvicornismuensteri Bernhauer, 1900 (Callicerus), Lectotypeobscurus Gravenhorst, 1802 (Callicerus), Neotypesparsicollis Bernhauer, 1915 (Callicerus), Lectotypestolfai Scheerpeltz, 1956 (Callicerus), syn. n. of Callicerus obscurus Gravenhorst, 1802smetanai Scheerpeltz, 1967 (Callicerus (Callicerus)), syn. n. of Atheta (Disopora) montenegrina (Bernhauer, 1899)beieri Scheerpeltz, 1959 (Callicerus (Semiris)), syn. n. of Callicerus rigidicornis (Erichson, 1839)gagliardii Scheerpeltz, 1956 (Callicerus (Semiris)), syn. n. of Callicerus fulvicornis Eppelsheim, 1883ibericus Fagel, 1958 (Callicerus (Semiris)), syn. n. of Callicerus obscurus Gravenhorst, 1802mandli Scheerpeltz, 1956 (Callicerus (Semiris)), syn. n. of Callicerus rigidicornis (Erichson, 1839)pedemontanus Baudi, 1869 (Callicerus obscurus var.), Lectotype now a synonym of Callicerus atricollis (Aubé, 1850)toroenensis (Bernauer, 1943) (Homoiocalea), comb. n. hitherto Atheta (Homoiocalea) toroenensisfulgida Assing, 2001 (Pseudosemiris), spec. n.kaufmanni (Eppelsheim, 1887) (Pseudosemiris), Lectotype described as Callicerus kaufmannizanettii Assing, 2001 (Pseudosemiris), spec. n.Homoiocalea Bernhauer, 1943 (Staphylinidae), stat. n. hitherto a subgenus of AthetaSemiris Heer, 1839 (Staphylinidae), syn. n. of Callicerus Gravenhorst, 1802Sphaerotaxus Bernhauer, 1915 (Staphylinidae), syn. n. of Callicerus Gravenhorst, 1802
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The scope of this study is to evaluate the faunistic, ecological and zoogeographical composition of the subfamily Steninae (Coleoptera: Staphylinidae) of the Central Anatolian Region of Turkey. Thirty-eight species, belonging to two genera, were detected by field studies between 2009-2011 in the Central Anatolian Region. It is concluded that among these, 12 species represent a first record for Central Anatolian Region and one species (Stenus distortus Assing, 2006) is endemic for Turkey. In previous studies, five species belonging to Stenus were recorded from Turkey, however, it was not clear that which provinces they were collected from. With this study, detailed localities are provided (Stenus ater Mannerheim, 1830, S. atratulus Erichson, 1839, S. carbonarius Gyllenhal, 1827, S. crassus Stephens, 1833, S. formicetorum Mannerheim, 1843).
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p>(*) 191° contribution to the knowledge of the Staphylinidae. The catalogue includes all the species described and/or referred to Cyprus and Asia Minor until 2007 inclusive. The bibliography on the species description to that of 2001-2002 and the geonemical citations is generally available in Herman (2001) and Smetana (2002); that of the species described or mentioned in contributions appeared after 2002 can be found in the bibliography at the hend of this catalogue. Described species of Cyprus and/or Asia minor are accompanied by the following informations: type locality or list of the localitys mentioned in the description, institutions or collections where is preserved the type, whether is male or female or if you do not know the sex; if it has been designed the lectotype and the author of the designation, the distribution. Some synonymies are proposed which will be discussed elsewhere, and some general remarks on the genus Geostiba Thomson.</p
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All names of species and higher taxa proposed in the tribe Lomechusini (Staphylinidae: Aleocharinae) up to the end of the year 2010 are cataloged, with a full bibliography of original descriptions. The original combination of the described species, original citation of their type localities and their world distributions are given. The following replacement names are proposed for newly discovered primary and secondary junior homonyms: Brachysipalia elgon nom. nov. (= B. elgonensis Pace), Drusilla sculpticollides nom. nov. (=Drusilla sculpticollis Pace), Macrogerodonia bolivianides nom. nov. (=M. boliviana Pace), Orphnebius curticornides nom. nov. (= O. curticornis Pace 2008), Pachorhopala ituri nom. nov. (= P. gerardi Bernhauer 1932), Zyras (Camonia) aethiopicides nom. nov. (=Zyras (Camonia) aethiopicus Pace) and Zyras (Zyras) neoparageminus nom. nov. (= Zyras (Zyras) parageminus Pace). The following secondary homonyms are reinstated as valid: Drusilla opacicollis Cameron (= D. vedda Pace), and Zyras (Grammodonia) erraticus Last (= Z. (G.) adjectus Last). The following new synonymies are proposed (junior synonym first): Aenictobiina Kistner (= Aenictoteratini Kistner), Athexeniina Pace (= Myrmedoniina Thomson), Thoracophagus Kistner (= Drusilla Leach), Urodonia Silvestri (= Pedinopleurus Cameron), Lomechusoides hosodai K. Sawada (= L. amurensis (Bernhauer)), Diplopleurus notabilis Pace (= Trachydonia incredibilis Bernhauer), and Acanthoglossa badia Motschulsky (= Zyras (Glossacantha) affinis Kraatz) (the last-mentioned being a reversal of precedence). The following new combinations are established: Aenictocleptis antennarius (Pace) ex Zyras (Glosacantha); Drusilla congoensis (Kistner) ex Thoracophagus; Pedinopleurus notabilis (Silvestri) ex Urodonia; Pedinopleurus reptans (Pace) and Pedinopleurus tumorfrontalis (Pace ) ex Zyras (Diaulaconia); Pseudastilbus fortispina (Pace) ex Zyras (Glosacantha) and Deroleptus draco (Assing) ex Orphnebius. A lectotype is designated for Lomechusoides ganglbaueri (Bernhauer) to fix this name as a synonym of L. amurensis (Wasmann). The following genera are newly included into Lomechusini, subtribe Myrmedoniina: Aenictophila Seevers, Athexenia Pace, Cephaplakoxena Pace, Malaybergius Kistner, Mimaenictus Kistner & Jacobson, Procantonnetia Kistner & Jacobson, Steysborgia Kistner & Jacobson and Weissflogia Kistner. The genus Myrmigaster Sharp is removed from Lomechusini to Oxypodini. Nomenclatural problems with the names Lomechusa Gravenhorst, Lomechusoides Tottenhamand Atemeles Dillwyn, and recent taxonomic changes to the genus Orphnebius Motschulsky, are also discussed. Lomechusini as currently constituted includes 207 genera and 2205 species or subspecies placed in the subtribes Lomechusina (3 genera, 40 species or subspecies), Termitozyrina (11 genera, 16 species) and Myrmedoniina (193 genera and 2149 species or subspecies). Their regional distribution patterns and extensive associations with ants and termites are briefly summarized.
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The Catalogue of Palaearctic Coleoptera provides information about all beetles occurring in Europe, North Africa and Asia north of the tropics. © 2015 by Koninklijke Brill nv, Leiden, The Netherlands. All rights reserved.
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A synopsis of adventive species of Staphylinidae (Coleoptera) in Canada is provided. Of the over 1654 rove beetle species recorded from Canada, 156 species in 76 genera and 13 subfamilies are reported as adventive, constituting about 9.4% of the fauna. The majority of adventive species belong to two subfamilies, Staphylininae (51 species) and Aleocharinae (49), and the genera with the most species are Philonthus (16), Atheta (8), Aleochara (7) and Quedius (7). All 156 adventive species are reviewed, including diagnostic features (supported by 156 colour habitus photos and 230 black and white images of diagnostic characters), native range, detailed distribution in Canada (with 151 distribution maps), general distribution in the USA, early Canadian and North American records and habitat information. New jurisdictional records are Atheta longicornis (Gravenhorst) from Alberta, Philonthus sanguinolentus (Gravenhorst) from eastern Canada (Ontario) and Xantholinus longiventris Heer from Canada (British Columbia).
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Emendations are cited or one of multiple original spellings adopted in Bledius, Bryoporus, Carpelimus, Edaphus, Eleusis, Gauropterus, Indosorius, Lispinus, Lithocharodes, Medon, Osorius, Philonthus, Piestus, Pinophilinus, Platydracus, Priochirus, Prognathoides, Quedius, Stenus, and Trigonurus. New combinations are proposed in Arrhenopeplus (3), Dialycera (2), Eusphalerum (14), Nacaeus (19), Pseudoxyporus (4), Homalotrichus (1), Thinodromus (2), Gabrius (1), Platydracus (57), Torobus (9), Bryophacis (7), Ischnosoma (16), Lordithon (66), Ryvkinius (1), and Sepedophilus (244). Fifty-nine new synonyms are listed for Micropeplus (1), Anthophagus (2), Eusphalerum (1), Phloeonomus (1), Eleusis (1), Leptochirus (1), Neolosus (1), Osorius (5), Priochirus (1), Anotylus (1), Bledius (1), Carpelimus (1), Ochthephilus (1), Apoquedius (1), Diatrechus (1), Erichsonius (2), Gabrius (2), Gastrisus (1), Hesperus (2), Heterothops (2), Leptacinus (1), Notolinus (1), Phallolinus (1), Philonthus (6), Quedius (5), Staphylinus (2), Tasgius (1) Xantholinus (4), Stenus (7), Sepedophilus (1), and Tachinus (1). Polyphematiana E. Strand is a junior synonym of Trigonopselaphus Gemminger and Harold. Eleven names are resurrected from synonymy, one each in Phyllodrepoidea, Anotylus, Oxytelus, Loncovilius, Philonthus, Platydracus, Lordithon, Mycetoporus, and Tachinus and two in Stenus. Under provisions of article 23.9.1 of the Code, 28 junior synonyms are protected in Amphichroum (1), Carpelimus (1), Deleaster (1), Proteinus (1), Bisnius (1), Leptacinus (1), Megalinus (1), Neobisnius (1), Ocypus (1), Philonthus (3), Quedius (4), Tasgius (2), Stenus (5), Bolitobius (1), Lordithon (1), Mycetoporus (1), and Tachyporus (2). Provisions of the same article protect 5 junior homonyms in Eusphalerum (1), Xylodromus (1), Quedius (1), Tachinus (1), and Tachyporus (1). Under provisions of article 23.9.3 of the Code, 15 junior synonyms will be referred to the Commission for rulings under the plenary power. Pending the outcome of these cases, use of the junior name is maintained. The affected species are in Eusphalerum (1), Lesteva (1), Omalium (1), Phloeostiba (1), Xylodromus (1), Anotylus (1), Bledius (1), Carpelimus (1), Ocypus (1), Philonthus (1), Quedius (4), and Lordithon (1). Under provisions of article 23.9.5 of the Code, 64 junior primary homonyms will be referred to the Commission for rulings under the plenary power. Pending the outcome of these deliberations, use of the junior name is maintained. The species are in Eusphalerum (1), Mannerheimia (1), Omaliomimus (1), Omaliopsis (1), Omalium (4), Phyllodrepa (1), Pycnoglypta (1), Xylodromus (1), Anotylus (1), Bledius (1), Carpelimus (1), Oxyporus (1), Belonuchus (2), Bisnius (1), Cafius (2), Cheilocolpus (1), Diatrechus (1), Endeius (1), Gabrius (5), Hesperus (2), Leptacinus (1), Nordus (1), Paederomimus (1), Philonthus (10), Platydracus (3), Quedius (2), Staphylinus (6), Xantholinus (2), Xanthopygus (1), Xenopygus (2), Carphacis (1), Coproporus (1), Tachinomorphus (1), and Tachinus (3). Replacement names are proposed for 90 preoccupied names in 37 genera: Euaesthetus (1), Leptotyphlus (1), Micropeplus (1), Eusphalerum (2), Lesteva (1), Olophrum (1), Omalium (2), Clavilispinus (1), Eleusis (3), Holotrochus (3), Leptochirus (1), Lispinus (4), Osorius (3), Priochirus (2), Thoracochirus (1), Anotylus (2), Carpelimus (2), Platystethus (1), Atanygnathus (1), Bisnius (1), Diochus (1), Gabrius (3), Hesperus (2), Heterothops (1), Leptacinus (3), Philonthus (19), Quedius (4), Staphylinus (3), Tasgius (1), Dianous (1), Bolitobius (1), Carphacis (1), Coproporus (3), Lordithon (3), Sepedophilus (5), Tachinus (4), Tachyporus (1). Eight names are resurrected to replace preoccupied names in Anotylus (1), Philonthus (1), Quedius (1), Stenus (2), Lordithon (1), Tachinomorphus (1), and Tachinus (1). Twenty-six senior homonyms, in Hapalaraea (1), Omalium (2), Phyllodrepa (1), Bledius (1), Oxytelus (1), Bisnius (1), Quedius (1), Staphylinus (15), Carphacis (1), Sepedophilus (1), and Tachinus (1), are regarded to be “nomina dubia” and the junior names conserved by fiat, without sanction by the Code. Psephidonus is the older name, but application to the Commission will be made to reject that name in favor of the younger Geodromicus. Tachyporiniformes is an unavailable name.
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A cladistic analysis of the West Palaearctic Scopaeina Mulsant & Rey, 1878 (Coleoptera, Staphylinidae: Paederinae) is presented along with bionomic and biogeographic information. A total of 76 morphological characters were coded for the 88 currently known West Palaearctic species, except for S. bifossicapitata (Outerelo & Oromi, 1987). Results show that Scopaeina comprises two well-supported monophyletic groups in the West Palaearctic, Micranops Cameron, 1913 and Scopaeus Erichson, 1840, which are considered to represent distinct genera. Phylogenetic relationships to Orus Casey, 1884, distributed in North and South America, are briefly discussed. Whereas Micranops is only represented by M. pilicornis (Baudi, 1869) in the region under study, 87 species of Scopaeus are currently known from the West Palaearctic. Within Scopaeus, the cladistic analysis yielded many well-supported monophyletic species groups, most of which are restricted to the West Palaearctic. However, except for Hyperscopaeus Coiffait, 1984, they are not in agreement with the widely used subgeneric concept sensu Coiffait (1952–1984). The following polyphyletic subgenera are consequently synonymized: Alloscopaeus Coiffait, 1968, Anomoscopaeus Coiffait, 1968, Geoscopaeus Coiffait, 1960, and Hyposcopaeus Coiffait, 1960 synn. n. = Scopaeus Erichson, 1840. Nivorus Herman, 1965, and Microscopaeus Coiffait, 1981 synn. n. = Micranops Cameron, 1913. The monotypical genus Coecoscopaeus Coiffait, 1984, established for C. coecus (Peyerimhoff, 1906), is excluded from Scopaeina. Scopaeus mitratus perroti Ochs, 1953 is raised to species rank, and S. nigellus Wollaston, 1864, formerly a synonym of S. minimus Erichson, 1939, is revalidated. Finally, we present a catalogue of species and synonyms of West Palaearctic Scopaeina along with distributional data and five new synonymies of species group names: S. bordei Peyerimhoff, 1914 syn. n. = S. portai Luze, 1910; S. tassiliensis Jarrige, 1958, S. mauretanicus Coiffait, 1960 synn. n. = S. crassipes Wollaston, 1867; S. saoudiensis Coiffait, 1981 = S. sinaicus Coiffait, 1970; S. mateui Coiffait, 1953 syn. n. = S. didymus Erichson, 1840. A lectotype is designated for S. didymus Erichson, 1840.See also Electronic Supplement (Parts 13) at http://www.senckenberg.de/odes/02-02.htm
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