NEW DATA ON RARE AND INSUFFICIENTLY KNOWN SPECIES OF GROUND BEETLES (COLEOPTERA: CARABIDAE: NEBRIINAE, CARABINAE, OMOPHRONINAE, ELAPHRININAE, SCARITINAE, BROSCINAE) IN LITHUANIA

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NAUJOS IR RETOS LIETUVOS VABZDŽIŲ RŪŠYS. 27 tomas 47
NEW DATA ON RARE AND INSUFFICIENTLY KNOWN SPECIES OF
GROUND BEETLES (COLEOPTERA: CARABIDAE: NEBRIINAE,
CARABINAE, OMOPHRONINAE, ELAPHRININAE, SCARITINAE,
BROSCINAE) IN LITHUANIA
VYTAUTAS TAMUTIS1,2, BRIGITA TAMUTĖ1, ROMAS FERENCA1
1Kaunas T. Ivanauskas Zoological Museum, Laisvės al. 106 LT-44253 Kaunas, Lithuania.
2Aleksandras Stulginskis University, Studentų 11, Akademija, Kaunas distr., LT-53361,
Lithuania.
E-mail: dromius@yahoo.com
Introduction
The ground beetles (Carabidae) is comparatively well studied beetle’s family in
Lithuania. In most cases they were studied as bioindicators to determine the influence of
some ecological factors to both agricultural (Tamutis, 2000; Tamutis et al., 2004, 2007)
and forest (Žiogas & Vaičikauskas, 2007a, 2007b; Dapkus & Tamutis, 2008; Lynikienė,
2006) ecosystems. Some studies were devoted to the investigation of species
composition and distribution in some important areas (Sharova & Grüntal, 1973,
Dvilevičius et al., 1988) or deal with exclusively faunistics and ecology (Ogijewicz,
1933; Pileckis & Monsevičius, 1995; Tamutis & Barševskis, 2014). Lots of single
faunistic records are scattered in numerous publications and communications (Tamutis et
al., 2011). Despite the attention of entomologists to this beetles’ family in Lithuania the
knowledge on distribution of some species is still insufficient.
In current paper we present the new information on some rare or insufficiently known
species of Nebriinae, Carabinae, Omophroninae, Elaphrininae, Scaritinae, Broscinae
subfamilies.
Material and methods
The data is based on the material collected by the authors Vytautas Tamutis (V.T.),
Brigita Tamutė (B.T.), Romas Ferenca (R.F.) and other collectors: Kęstutis Arbačiauskas
(K.A.), Teisutis Ašmenavičius (T.A.); Dalius Dapkus (D.D.), Elena Gaidienė (E.G.),
Artūras Gedminas (A.G.), Povilas Ivinskis (P.I.), Saulius Karalius (S.K.), A. Kaulinis
(A.K.), L. Kontenytė (L.K.), Aleksandras Meržijevskis (A.M.), Vidmantas Monsevičius
(Vidm.M.), Virginijus Monsevičius (Virg.M.), Povilas Mulerčikas (P.M.), Norbertas
Noreika (N.N.); Henrikas Ostrauskas (H.O.), Rimantas Pankevičius (R.P.), Simonas
Pileckis (S.P.), Algimantas Ragelis (A.R.), Gintautas Steiblys (G.S.), Gintaras
Slavinskas (G.Sl.), K. Tešerskis (K.T.), Vytautas Uselis (V.U.). All available published
and unpublished data of findings was used to present the pattern of seasonal distribution
of collected specimens in Lithuania. The main mass of material was collected using
pitfall traps, other specimens mostly of such species as: Notiophilus germinyi, Omophron
limbatum, Blethisa multipunctatum, Elaphrus aureus, Clivina collaris and Miscodera
arctica, were collected surveyng specific habitats. For identification of species we used
the keys suggested by K. Hůrka (1996) and G. Müller-Motzfeld (2004). The

48 NEW AND RARE FOR LITHUANIA INSECT SPECIES. Volume 27
nomenclature follows Carabidae of the World database (Anichtchenko, 2007–2015) and
Catalogue of Palaearctic Coleoptera (Löbl & Smetana, 2003). The material is deposited
in the collection of Kaunas Tadas Ivanauskas Zoological museum.
List of localities
Localities aDistricts Coordinates (WGS)
Akademija env. Kaunas 54°54'16"N; 23°49'28"E;
Antalgė Utena 55°30'08"N; 25°28'11"E;
Ariogala env. Raseiniai 55°15'17"N; 23°28'44"E;
Aukštosios Miškas f. Šakiai 54°59'05"N; 22°49'56"E;
Aukštumala bog Šilutė 55°23'50"N; 21°20'43"E;
Ąžuolų Būdos Miškas f. Kazlų Rūda 54°43'13"N; 23°31'31"E;
Baluoša peat bog Švenčionys 54°54'19"N; 25°51'09"E;
Birštonas env. Birštonas 54°36'39"N; 24°01'58"E;
Braziūkai env.1 Kaunas 54°54'09"N; 23°29'00"E;
Braziūkai env.2 Kaunas 54°54'06"N; 23°29'01"E;
Braziūkai env.3 Kaunas 54°54'16"N; 23°28'47"E;
Bubių Miškas f. Šiauliai 55°50'54"N; 23°08'36"E;
Čepkeliai Nat. R.1 Varėna 54°01'38"N; 24°28'04"E;
Čepkeliai Nat. R.2 Varėna 54°00'50"N; 24°35'04"E;
Čižiškės env. Vilnius 54°56'28"N; 25°37'26"E;
Čižiūnai env. Trakai 54°36'11"N; 24°34'00"E;
Daukšiai env. Joniškis 56°21'08"N; 23°20'26"E;
Dituva env. Klaipėda 54°33'33"N; 21°15'18"E;
Druskininkai env. Druskininkai 54°02'09"N; 23°58'55"E;
Dubravos Miškas f.1 Kaunas 54°50'10"N; 24°09'08"E;
Dubravos Miškas f.2 Kaunas 54°49'32"N; 24°04'03"E;
Dubravos Miškas f.3 Kaunas 54°50'45"N; 24°04'02"E;
Dubravos Miškas f.4 Kaunas 54°50'02"N; 24°01'18"E;
Dūkštų ąžuolynas f. Vilnius 54°49'59"N; 24°57'12"E;
Dumaičiai env. Mažeikiai 56°12'25"N; 22°04'10"E;
Dzidai env. Šiauliai 55°49'52"N; 23°09'45"E;
Eržvilkas env. Jurbarkas 55°15'33"N; 22°41'36"E;
Ežerėlis env.1 Kaunas 54°53'11"N; 23°35'20"E;
Ežerėlis peat bog Kaunas 54°51'43"N; 23°40'43"E;
Gimbetiškė env. Akmenė 56°08'43"N; 22°33'18"E;
Girionys env. Kaunas 54°51'51"N; 24°03'16"E;
Girininkai env. Kaunas 54°50'31"N; 23°33'18"E;
Glitis bog Tauragė 55°08'49"N; 22°28'44"E;
Glitiškės env. Vilnius 54°59'01"N; 25°13'26"E;
Grybaulia env. Varėna 53°59'51"N; 24°21'14"E;
Gyvoliai env. Mažeikiai 56°12'59"N; 22°33'14"E;
Janaslavas env. Lazdijai 54°09'50"N; 23°28'36"E;
Jiesia landscape rezerve Kaunas 54°51'11"N; 23°56'16"E;
Joniškis env. Joniškis 56°14'25"N; 23°36'39"E;
Josvainiai env. Kėdainiai 55°14'33"N; 23°49'54"E;

NAUJOS IR RETOS LIETUVOS VABZDŽIŲ RŪŠYS. 27 tomas 49
Juknaičiai env. Šilutė 55°18'08"N; 21°40'54"E;
Juodeikiai env. Joniškis 56°14'21"N; 23°13'07"E;
Juodkrantė env. Neringa 55°32'46"N; 21°06'06"E;
Juškinės Miškas f. Šakiai 55°01'14"N; 23°26'53"E;
Kajackaraistis env.1 Kazkų Rūda 54°47'56"N; 23°37'22"E;
Kajackaraistis env.2 Kazkų Rūda 54°48'10"N; 23°37'24"E;
Kalnelis env. Joniškis 56°14'31"N; 23°33'23"E;
Kalviai env. Jurbarkas 55°04'21"N; 23°21'21"E;
Kamanos Nat. R. Akmenė 56°16'15"N; 22°37'09"E;
Kamšos Miškas f. Kaunas 54°53'37"N; 23°50'26"E;
Karmazinai env. Vilnius 54°49'02"N; 24°56'09"E;
Katra env. Varėna 53°59'27"N; 24°35'30"E;
Kaunas1 Kaunas 54°52'31"N; 23°57'15"E;
Kaunas2 Kaunas 54°54'03"N; 23°55'10"E;
Kazlų Rūda env. Kazlų Rūda 54°44'26"N; 23°28'58"E;
Kepalių Miškas f. Joniškis 56°13'55"N; 23°39'38"E;
Kiminiškė env. Mažeikiai 56°17'43"N; 22°11'47"E;
Krivasalis env. Ignalina 55°18'53"N; 25°53'46"E;
Kruonis env. Kaišiadorys 54°44'53"N; 24°14'28"E;
Kuodiškiai env. Rokiškis 56°05'14"N; 25°17'05"E;
Lapnugaris landscape R. Neringa 55°34'38"N; 21°07'47"E;
Linkaičiai env. Joniškis 56°11'38"N; 23°28'29"E;
Lyduvėnai env. Raseiniai 55°30'01"N; 23°05'26"E;
Margininkai Kaunas 54°47'35"N; 24°02'54"E;
Maželių Miškas f. Joniškis 56°17'26"N; 23°35'33"E;
Meištinės Miškas f. Kaunas 54°53'18"N; 23°35'09"E;
Merkinė Varėna 54°10'03"N; 24°12'35"E;
Mickiškė env. Tauragė 55°17'40"N; 22°31'58"E;
Miknaičiai env. Šakiai 54°44'23"N; 23°09'02"E;
Nagliai Nat. R. Neringa 55°29'08"N; 21°06'01"E;
Naujasis Daugėliškis env. Ignalina 54°36'11"N; 24°34'00"E;
Nemajūnų Miškas f. Birštonas 54°45'26"N; 25°26'13"E;
Nemunėlio Radviliškis env. Biržai 56°23'20"N; 24°45'56"E;
Nida env. Neringa 55°18'21"N; 21°00'28"E;
Noreikiškės env. Kaunas 54°53'12"N; 23°50'13"E;
Novaraistis peat bog Kaunas 54°56'34"N; 23°24'58"E;
Pabariukai env. Kelmė 55°37'18"N; 23°04'25"E;
Pabradė env. Švenčionys 54°59'54"N; 25°45'25"E;
Padumblys env. Ignalina 55°17'49"N; 26°02'28"E;
Paežerių Miškas f. Šiauliai 55°57'35"N; 22°49'05"E;
Pakalniškiai env. Akmenė 56°16'56"N; 22°58'05"E;
Paketuriai env. Kupiškis 55°50'57"N; 24°58'06"E;
Palanga airport env. Palanga 55°59'22"N; 21°05'49"E;
Paliepės Miškas f. Varėna 54°06'43"N; 24°18'57"E;
Paminėlė env. Švenčionys 55°14'45"N; 25°44'57"E;
Panemunės Šilas f. Kaunas 54°52'51"N; 23°58'50"E;
Paryžinės Miškas f. Kaunas 54°57'38"N; 23°26'51"E;

50 NEW AND RARE FOR LITHUANIA INSECT SPECIES. Volume 27
Pašilių Miškas f. Kėdainiai 55°14'12"N; 23°57'26"E;
Pažaislis env. Kaunas 54°52'27"N; 24°01'15"E;
Perloja env. Varėna 54°14'22"N; 24°25'01"E;
Pervalka env. Neringa 55°25'02"N; 21°06'49"E;
Plikiai env. Klaipėda 55°46'25"N; 21°14'19"E;
Plokštinė Plungė 56°01'01"N; 21°51'23"E;
Pogarenda env. Varėna 53°56'09"N; 24°28'33"E;
Punios Šilas f. Alytus 54°30'55"N; 24°02'59"E;
Puvočiai env. Varėna 54°06'53"N; 24°18'03"E;
Raguviškiai env. Kretinga 55°51'37"N; 21°23'21"E;
Ramoniškė env. Mažeikiai 56°11'05"N; 22°36'09"E;
Ramuldava env. Anykščiai 55°35'21"N; 25°12'07"E;
Randamonys env. Varėna 54°03'19"N; 24°13'01"E;
Ratnyčia env. Druskininkai 53°59'43"N; 24°02'07"E;
Ratnyčios Miškas f. Druskininkai 53°59'21"N; 24°01'06"E;
Reibiniškių Miškas f. Joniškis 56°15'46"N; 23°20'06"E;
Reiskiai env. Plungė 55°51'35"N; 21°35'22"E;
Renavas env. Mažeikiai 56°13'43"N; 22°03'46"E;
Ringovė Entomological R. Kaunas 55°03'03"N; 23°31'08"E;
Romainiai park Kaunas 54°55'30"N; 23°49'30"E;
Rubuliai env. Kretinga 55°53'52"N; 21°25'10"E;
Saldutiškis env. Utena 55°21'07"N; 25°49'07"E;
Satkūnų Miškas f. Joniškis 56°16'59"N; 23°39'16"E;
Šeimyniškiai Anykščiai 55°30'37"N; 25°04'17"E;
Senasis Daugėliškis env. Ignalina 55°20'40"N; 26°16'34"E;
Sėlų Miškas f. Šalčininkai 54°22'15"N; 25°16'34"E;
Šilėnai env. Vilnius 54°45'26"N; 25°26'13"E;
Simokai env. Šakiai 54°44'21"N; 23°09'07"E;
Siponys Birštonas 54°31'24"N; 23°59'51"E;
Šiauduva env. Šilalė 55°34'34"N; 22°13'51"E;
Šimonių Giria f. Anykščiai 55°41'26"N; 25°10'17"E;
Škėvonys env. Birštonas 54°36'16"N; 24°00'11"E;
Skriaudžių Miškas f. Prienai 54°45'11"N; 23°37'39"E;
Slydžiai env. Šiauliai 55°49'55"N; 23°09'27"E;
Smalininkai env. Jurbarkas 55°04'17"N; 22°36'07"E;
Smiltynė env.1 Klaipėda 55°42'54"N; 21°06'03"E;
Smiltynė env.2 Klaipėda 55°40'46"N; 21°06'34"E;
Tabariškiai env. Kaunas 54°52'18"N; 23°46'58"E;
Tartokių Miškas f. Vilnius 54°49'41"N; 25°21'15"E;
Tervydoniai env. Šakiai 55°01'38"N; 23°26'36"E;
Trakai env. Trakai 54°38'42"N; 24°54'59"E;
Traklaukas env. Tauragė 55°16'22"N; 22°32'27"E;
Užgiriai env. Marijampolė 54°27'22"N; 23°21'27"E;
Užuožerės Miškas f. Varėna 54°01'31"N; 24°25'38"E;
Vaišniūnai env. Ignalina 55°23'23"N; 26°06'05"E;
Varėna env. Varėna 54°13'20"N; 24°32'18"E;
Varlaukis env. Jurbarkas 55°19'10"N; 22°40'23"E;

NAUJOS IR RETOS LIETUVOS VABZDŽIŲ RŪŠYS. 27 tomas 51
Varliškės env. Elektrėnai 54°46'37"N; 24°52'28"E;
Varniai env Telšiai 55°44'28"N; 22°21'53"E;
Varnikai env. Trakai 54°39'10"N; 24°58'46"E;
Ventės Ragas env. Šilutė 55°20'36"N; 21°11'47"E;
Verkiai env. Vilnius 54°46'00"N; 25°18'07"E;
Vidzgiris Botanical R. Alytus 54°22'46"N; 24°01'57"E;
Viešvilė Nat. R.1 Tauragė 55°10'35"N; 22°28'18"E;
Viešvilė Nat. R.2 Jurbarkas 55°07'11"N; 22°24'53"E;
Viešvilė Nat. R.3 Jurbarkas 55°08'22"N; 22°27'11"E;
Viešvilė Nat. R.4 Jurbarkas 55°07'20"N; 22°25'01"E;
Viešvilė Nat. R.5 Tauragė 55°08'24"N; 22°29'39"E;
Viršužiglis env. Kaunas 54°50'09"N; 24°11'59"E;
Vytautas park Kaunas 54°53'52"N; 23°56'08"E;
Vytinėlės Miškas f. Ukmergė 55°13'35"N; 24°44'55"E;
Zapyškis env. Kaunas 54°55'33"N; 23°39'26"E;
Žeimiai env. Joniškis 56°15'05"N; 23°18'00"E;
Želsva env. Marijampolė 54°26'24"N; 23°26'39"E;
Zervynos Varėna 54°06'56"N; 24°30'14"E;
Žiegždrių Miškas f. Kaunas 54°53'13"N; 24°05'13"E;
Žydeikiškės env. Kaišiadorys 54°48'21"N; 24°20'10"E.
aThe term ‘districts’ herein stands for all types of administrative units used in
Lithuania since 2000 (i.e. municipalities, district municipalities and town municipalities)
Annotated list of species
Family CARABIDAE
Subfamily Nebriinae Laporte, 1834
Tribe Nebriini Laporte, 1834
Nebria brevicollis (Fabricius, 1792)
Akademija env., 16 08 2007, 1 spec. (V.T.); Birštonas env., 06 06 2001, 1 spec.
(R.F.); Čižiūnai env., 17 06 1999, 2 spec. (P.I.); Daukšiai env., 15-30 08 2007, 7
spec. (T.A.); Dubravos Miškas f.1, 09 08 1981, 1 spec. (R.F.); Dūkštų ąžuolynas f., 1
07 1995, 1 spec. (H.O.), 18 06 2009, 1 spec. (V.T.); Dumaičiai env., 9 05 1991, 1
spec. (Vidm.M.); Dzidai env., 12 07 1992, 05 04 2003, (1, 1) spec. (V.T.); Eržvilkas
env., 21 08 1995, 1 spec. (G. Sl.); Girininkai env., 06 06 1994, 1 spec. (V.T.);
Girionys env., 17 05 1989, 1 spec. (A.M.); Janaslavas env., 28 05 1990, 1 spec.
(G.Sl.); Jiesia landscape rezerve, 13 06 1981, 07 08 1981, 07 07 1982, 13 08 1982, 28
07 1984, 30 07 1986, 19 08 1993, 24 06 2000 (5, 2, 1, 1, 2, 1, 2, 1) spec. (R.F.);
Joniškis env., 15-30 08 2007, 1 spec. (T.A.); Josvainiai env., 13 06 1995, 1 spec.
(G.Sl.); Juodeikių env., 15-30 08 2007, 12 spec. (T.A.); Juodkrantė env., 8 09 2005,
18 10 2005, (3, 7) spec. (P.I.); Kalnelis env., 15-30 08 2007, 1 spec. (T.A.); Kamanos
Nat. R., 16 05 1991, 01-16 06 1991, 16 08 1991, 01-16 09 1991, (2, 3, 1, 4) spec.
(Vidm.M.); Kamšos Miškas f., 21 05 1992, 29 03 2003, 21 04 2003, 15 06 2008, (3,
1, 1, 3) spec. (V.T.); Kaunas1, 09 06 1977, 27 07 1986, (1, 4) spec. (R.F.); Kaunas2,
02 04 2002, 2 spec. (V.T.); Kepalių Miškas f., 15-30 08 2007, 4 spec. (T.A.);
Linkaičiai env., 15-30 08 2007, 1spec. (T.A.); Margininkai, 16 05 1999, 20 06 1999,

52 NEW AND RARE FOR LITHUANIA INSECT SPECIES. Volume 27
(1, 3) spec. (V.T.); Maželių Miškas f., 15-30 08 2007, 1spec. (T.A.); Merkinė env.,
10 08 1981, 1 spec. (G.Sl.); Noreikiškės env., 03 08 1991, 15 06 2006, 25 05 2009,
(1, 2, 2) spec. (V.T.); Paežerių Miškas f., 16 07 1998, 1 spec. (R.F.); Palanga aeroport
env., 09 05 2001, 2 spec. (R. F.); Panemunės Šilas f., 19 08 1982, 1 spec. (R.F.);
Pažaislis env., 27 07 2002, 1 spec. (V.T.); Perloja env., 28 05 1982, 1 spec.
(Anonimous); Pogarenda env., 03 08 1977, 1 spec. (Vidm.M.); Punios Šilas f., 27 06
2008, 2 spec. (V.T.); Puvočiai env., 20 07 1979, 1 spec. (K.A.); Ramoniškė env., 23
06 1993, 1 spec. (Vidm.M.); Reibiniškių Miškas f., 15-30 08 2007, 8 spec. (T.A.);
Reiskiai env., 10 09 1994, 15 07 2001, (1, 5) spec. (S.K.); Renavas env., 01 07 1991,
21 07 1992, (2, 3) spec. (Vidm.M.); Saldutiškis env., 06 07 1998, 1 spec. (V.T.);
Satkūnų Miškas f., 15-30 08 2007, 5 spec. (T.A.); Senasis Daugėliškis env., 10 08
1991, 1 spec. (Vidm.M.); Šiauduva env., 31 07 1991, 1 spec. (G.Sl.); Škėvonys env.,
16 09 1981, 1 spec. (G.Sl.); Slydžiai env., 22 07 1991, 06 05 1992, 03 08 2002, 09 03
2008, (4, 1, 1, 1) spec. (V.T.); Smiltynė env.2, 04-15 07 1983, 5 spec. (S.K.); Tartokių
Miškas f., 01-20 06 2005, 11 07 2005, (4, 2) spec. (P.I.); Užgiriai env., 05 08 1999,
05 09 1999, (1, 1) spec. (Vidm.M.); Ventės Ragas env., 23 08 1988, 1 spec. (R.F.);
Vidsgiris Botanical rezerve, 05 06 1998, 17 08 2001, (2, 2) spec. (R.F.); Viešvilė Nat.
R.1, 12 06 1992, 1 spec. (Vidm. M.); Viešvilė Nat. R.4, 29 07 2008, 1 spec. (V.T);
Vytautas park, 07 08 2007, 1 spec. (A.M.); Vytinėlės Miškas f., 21 06 2004, 28 08
2004, (6, 7) spec. (N.N.); Zapyškis env., 24 05 2009, 1 spec. (Anonymous); Žeimiai
env., 15-30 08 2007, 9 spec. (T.A.); Žydeikiškės env., 09 09 2001, 1 spec. (V.T.).
Fig. 1. Number of records and recorded specimens of Nebria brevicollis F. in
Lithuania
Notes. This species is widely distributed in Europe, also noted for Siberia and Far
East, Turkey, Algeria (Farkač & Janata, 2003), introduced to North America and is
rapidly expanding its range in the western part of the country (LaBonte, 2011). The
expansion of the distribution range of this species to the European north east and its
population growth in the last decades have been detected in Europe as well, but still not
widely disscussed (O. Aleksandrowich personal comment). The assigning of this species
as rare in Lithuania by Pileckis & Monsevičius (1995) is probably based on previous old
single records (Ogijewicz, 1933; Pileckis, 1960, 1963). Assuming all available data on
distribution of this species some tendencies of its population growth could be perceptible
in Lithuania too (Fig. 1). Presently this species is frequent and widely distributed in
0
20
40
60
80
100
120
140
160
180
200
till 1960 1961-1990 1991-2015
Periods
Number of records
Number of specimens

NAUJOS IR RETOS LIETUVOS VABZDŽIŲ RŪŠYS. 27 tomas 53
Lithuania; inhabits various habitats suchs as forests (preferably deciduous and mixed),
water banks, meadows, arable lands, shruby areas and likewise. Observations of adults in
Lithuania: III: 2; IV: 4; V: 17; VI: 44; VII: 41; VIII:80; IX: 15; X: 7.
Leistus rufomarginatus Duftschmid, 1812
Jiesia landscape rezerve, 28 08 2006, 01 07 2009, 29 04 2011 (1, 1, 1) spec. (R.F.);
Kamšos Miškas f., 27 02 2000, 1 spec. (V.T.), 01–15 07 2005, 11 spec. (P.M.),
Ringovė Entomological reserve, 18 07 1996, 1 spec. (A.M.); 05 07 1998, 1 spec.
(V.T.), 27 08 2003, 23 08 2005 (1, 2) spec. (R.F.); Romainiai park, 24 07 2007, 1
spec. (A.M.); Smalininkai env., 27 07 1995, 1 spec. (V.T.); Šeiminyškiai env., 15 08
2000, 1 spec. (Anonimous); Tervydoniai env., 29 08 2015, 1 spec. (R.F.); Vytautas
park, 07 08 2007, 1 spec. (A.M.).
Notes. This species is natively distributed in central Europe and Caucasus, northern
border of range reaches southern part of Sweden, Denmark, northern part of Poland and
eastern Prussia (nower days – Kaliningad region) (Lindroth, 1992, Bercio & Folwaczny,
1979). However new records of this species in Lithuania and Latvia (Barševskis, 2001,
Ferenca et al., 2002; Ivinskis et al., 2006) extends its distribution range to the northeast.
Since the first record of this species in the western part of Lithuania in 1992 it spread to
the east as far as 25° of eastern longitude. Almost all newly recorded specimens of L.
rufomarginatus in Lithuania were found in the stands of deciduous trees. Observations of
adults in Lithuania: II: 1; IV: 1; VI: 2; VII: 14; VIII: 8; X: 3.
Leistus piceus Frölich, 1799
Bubių Miškas f., 10 07 1999, 1 spec. (G.Sl.); Ramoniškė env., 23 06 1993, 1 spec.
(Vidm.M.).
Notes. It is poor known and seems to be extremely rare species in Lithuania;
previously recorded from central part of the country (Šablevičius & Ferenca, 1995). The
previous records of this species from Vilkija env. (Kaunas distr.), Juodkrantė and Preila
environs (Neringa mun.) (Šablevičius, 2003; Ferenca 2014), Stalgėnai env. (Plungė
distr.) (Šablevičius, 2003) were based on misidentifications and related to Leistus
ferrugineus Linnaeus, 1758. Leistus piceus is distributed in central Europe (Farkač &
Janata, 2003), and is considered as typical montane species (Burakowski et al., 1973;
Fritze & Blick, 2012); it is rare in lowlands, occasionally found in northwestern France,
Belgium, northeastern Poland, Belarus, Latvia, Estonia, western Russia (Amiet, 1962;
Desender, 1986; Burakowski et al., 1973; Aleksandriwicz, 2014; Barševskis, 2003;
Haberman, 1968; Alekseev, 2008; Kataev, 2004; Egorov, 2010).
Tribe Notiophilini Motschulsky, 1850
Notiophilus germinyi Fauvel, 1863
Antalgė, 03 04 1977, 1 spec. (A.K.); Baluoša peat bog, 02–15 06 1999, 1 spec.
(D.D.); Čepkeliai Nat. R.1, 15-29 07 1976, 19 08 1976, 02 09 1976, 6 06 1977, 01–20
06 1981, 11–17 08 1981, 04 08 1984, (2, 1, 1, 1, 2, 2, 1) spec. (Vidm.M.); 15 07
1985, 1 spec. (R.F.); Druskininkai env., 22 07 1995, 1 spec. (V.T.); Dubravos Miškas
f.1, 04 10 1984, 1 spec. (R.F); Ežerėlis env., 06 08 1982, 1 spec. (R.F.); Juodkrantė
env., 24 08 1967, 1 spec. (E.G.), 08 10 2005, 1 spec. (P.I.); Merkinė, 26 06 2002, 1
spec. (A.M.); Viešvilė Nat. R.1, 15 07 2001, 1 spec. (V.U.); Viešvilė Nat. R.2, 12 06
2008, 1 spec. (V.T. & B.T.); Viešvilė Nat. R.3, 09 05 2008, 12 06 2008, 29 07 2008,
(4, 1, 1) spec. (V.T. & B.T.); Zervynos, 15 07 1976, 16 06 1977, (1, 1) spec.
(Vidm.M.).
Notes. This typical xerophilous species occurs in sandy or gravely, weakly shaded or

54 NEW AND RARE FOR LITHUANIA INSECT SPECIES. Volume 27
open habitats: lichen rich pine heaths, dry meadows with low vegetation. Despite wide
distribution range of this species (Farkač & Janata, 2003) in Lithuania it was found fairly
locally and was known only from several districts (Ogijewicz, 1933; Barševskis, 2001,
Ferenca, 2014). Observations of adults in Lithuania: IV: 1; V: 6; VI: 8; VII: 8; VIII: 13;
IX: 7; X: 1.
Subfamily Carabinae Latreile, 1802
Tribe Carabini Latreile, 1802
Carabus clatratus Linnaeus, 1761
Aukštosios Miškas f. (Baltkojai forestry), 10 06 1960, 1 spec. (S.P.); Ąžuolų Būdos
Miškas f., August 1958, 1 spec. (S.P.); Baluoša peat bog, 13 05 1999, 1 spec. (D.D.);
Čepkeliai Nat. R.2, 25 05 1976, 19 06 1976, 19 06 1978, 25 06 1983, (1, 1, 1, 1) spec.
(Virg.M.); Dubravos Miškas f.3, 15 07 1955, 1 spec. (S.P.); Glitiškės env., August
1960, 1 spec. (S.P); Juknaičiai env., 10 06 1997, 12 05 1998, (1, 1) spec. (A.G.);
Kazlų Rūda env., 17 07 1956, 1 spec. (E.G.); Nemunėlio Radviliškis env., June–
August 1958, 2 spec. (S.P.); Padumblys env., 15 07 1955, 1 spec. (S.P.); Plokštinės
Miškas f., 06 08 1997, 1 spec. (H.O.); Varėna env., June 1959, 1 spec. (S.P.); 02 06
1976, 1 spec. (Anonymous).
Notes. This species was considered as not rare in Lithuania and noted for central,
eastern and southern parts of Lithuania without concrete data (Pileckis & Monsevičius,
1995). Full finding data previously was published only from few localities: Žuvintas Nat.
R. (Zajančkauskas & Pileckis, 1968; Sharova & Grüntal 1973), Tyruliai bog (Ferenca,
2006), Trakai and Vilnius environs (Ogijewicz, 1933), Juodkrantė env. (Ferenca, 2003,
2014; Ivinskis et al., 2004). Population of this species in Lithuania seems to decline; just
six specimens have been detected during the last 35 years. This species is also regarded
as rare and local in all Fenoscandian countries (Lindroth, 1992), in some countries of
Europe it is included into the lists of officially protected species (Cinītis, 1998;
Pawłowski et al., 2002; Belova, 2004; Curikov, 2006; Ødegaard et al., 2010; Rizun,
2011). Observations of adults in Lithuania: IV: 1; V: 5; VI: 12; VII: 4; VIII: 5; IX: 1.
Carabus convexus Fabricius, 1775
Čižiškė env., 06 06 2005, 02 07 2005, (1, 1) spec. (P.I.); Dubravos Miškas f.1, 03 08
1984, 10 08 1984, 23 08 1984, (1, 1, 1) spec. (R.F.); 27 04 1987, 1 spec. (A.R.);
Karmazinai env., 29 07 2003, 1 spec. (R.F.); Kruonis env., 19 05 1991, 1 spec.
(A.M.), Nagliai Nat. R., 30 05 2009, 1 spec. (R.F.); Palanga aeroport env., 09–30 05
2001, 3 spec. (R.F.); Randamonys env., 09 10 2001, 1 spec. (A.G.); Smiltynė env.1,
14 07 1982, 12 07 1983, 03 08 1983, 09 07 1984, 05 08 1984, (1, 1, 1, 1, 2) spec.
(S.K.); Smiltynė env.2, 04–23 06 1995, 06–07 07 1995, (8, 4) spec. (V.T.); Šilėnai
env., 29 07 2003, 1 spec. (R.F.); Šimonių Giria f., 08-11 06 1997, 1 spec. (R.P.);
Tartokių Miškas f., 01–20 06 2005, 22 08 2005, (2, 3) spec. (P.I.); Vaišniūnai env.,
14 06 1994, 1 spec. (R.P.); Verkiai env., 06 06 2005, 3 spec. (P.I.) Žiegždrių Miškas
f., 20 08 1982, 1 spec. (R.F.).
Notes. This species is widely distributed in Central Europe and Caucasus, but rare in
the northern areas (Lindroth, 1992; Cinītis, 1998; Pawłowski et al., 2002; Hyvärinen et
al., 2010; Ljunberg et al., 2010; Ødegaar et al., 2010). In Lithuania this species is local
and almost everywhere sporadic. It occurs either in dry or semidry deciduous and
coniferous forests. Most numerous specimens were observed in Pine forests of Curronian
Spit (Ferenca 2014). Observations of adults in Lithuania: IV: 16; V: 18; VI: 37; VII: 27;

NAUJOS IR RETOS LIETUVOS VABZDŽIŲ RŪŠYS. 27 tomas 55
VIII: 19; X: 2; XI: 6.
Carabus nitens Linnaeus, 1758
Aukštumala bog, June 2003, 1 spec. (V.U.); Dituva env., 3 07 1985, 1 spec. (S.K.);
Druskininkai env., 15 06 1996, 1 spec. (V.T.); Dubravos Miškas f.4, 6 06 1999, 1
spec. (R.F.); Ežerėlis peat bog, 19 05 1985, 1 spec. (R.F.); Glitis bog, 5 06 1996, 1
spec. (V.U.); Kajackaraistis env.1, 27 07 1986, 1 spec. (R.F.); Kamanos bog, 16 05
1984, 1 spec. (K.A.); Novaraistis peat bog, 20 08 2001, 1 spec. (A.G.); Paryžinės
Miškas f., 31 05 2009, 1 spec. (G.S.); Plikiai env., 1 05 1987, 1 spec. (S.K.).
Notes. This European species is widely distributed in the country, but everywhere is
sporadic; listed in the Red Data book of Lithuania as rare species (Ferenca, 2007).
Almost all specimens of this species have been detected in dry, open or slightly shaded
places with sand or peat soils: heaths, moors, clear cuts, openings; no catches in
agricultural lands. The populations of this remarkable species have declined severely in
recent decades in many countries of Central Europe (Assman & Janssen, 1999), and are
officially protected in many Euoropean countries. Observations of adults in Lithuania:
IV: 4; V: 50; VI: 53; VII: 20; VIII: 3; X: 2; XI: 6.
Carabus violaceus andrzejuscii Fischer von Waldheim, 1823
Braziūkai env.2, 27 07 2015, 1 spec. (V.T. & B.T.); Braziūkai env.3, 21 08 2011, 2
spec. (V.T. & B.T.); Čepkeliai Nat. R.1, 09 06 1977, 08 06 1978, 30 10 1978 (1, 1, 4)
spec. (Vidm.M.); Juškinės Miškas f., 20 05 2000, 25 08 2011 (2, 1) spec.. (R.F.);
Krivasalis env., 20 07 1974, 2 spec. (Vidm.M.); Meištinės Miškas f., 21 06 2006, 1
spec. (A.M.); Nagliai Nat. R., 16 06 2005, 08 09 2005 (2, 8) spec. (P. I.); Nida env.,
16 05 2002, 1 spec. (R.F.); Pabradė env., 13 07 1996, 10 spec. (H.O.); Paliepės
Miškas f., 17 05 2012, 2 spec. (K.T. & L.K.); Paminėlė env., 11 08 1991, 1 spec.
(Vidm.M.); Pogarenda env., 10 07 1977 1 spec. (Vidm.M.); Ramuldava env., 12 08
1988, 1 spec. (S.K.); Ratnyčios Miškas f., 10–21 05 1995, 1–27 06 1995 (4, 43) spec.
(V. T.); Smiltynė env.1, 29 07 1983, 12–14 08 1983 (1, 4) spec. (S.K.); Vaišniūnai
env., 14 07 1975, 2 spec. (Vidm. M.); Varliškės env., 03–24 07 1991, 03–27 07 1993,
21 06 1993, 14 07 1993 (4, 5, 4, 5) spec. (H.O.); Varnikai env., 29 06 1994, 1 spec.
(H.O.); Vidzgiris Botanical reserve, 30 07 1988, 17 08 2001 (1, 1) spec. (R.F.).
Notes. This subspecies is noted just for ten countries in Central Europe: Belarus,
Central European Russia, Estonia, Latvija, Lithuania, Moldova, Poland, Romania,
Slovakia and Ukraina (Bousquet et al., 2003). It is typical forest species considered as
rare in Lithuania (Pileckis & Monsevičius 1995), included into the Red lists of some
European countries (Kataev & Krivohatskij, 2004; Trautner et al., 2005,
Aleksandrowicz, 2014). In Lithuania this species is distributed widely but not evenly,
most frequently observed in Curronian Spit (Žiogas & Zolubas, 2005; Ferenca 2014) and
Southern part, but remains sporadic in other areas; occurs mainly in pure pine stands or
with a small admixture of Picea or Betula species. Observations of adults in Lithuania:
II: 1; V: 14; VI: 79; VII: 71; VIII: 21; X: 5.
Subfamily Omophroninae Bonelli, 1810
Tribe Omophronini Bonelli, 1810
Omophron limbatum (Fabricius, 1777)
Ariogala env., bank of Dubysa r., 05 06 1992, 1 spec. (V.T.); Braziūkai env.1, bank of
sand pit, 01 06 2008, 1 spec. (A.M.); Dubravos Miškas f.2, bank of sand pit, 15 05
1994, 3 spec. (R.F.); Katra env., 01 08 1984, 1 spec. (Vidm.M.); Eržvilkas env., bank
of Šaltuona r., 11 05 2012, 4 spec. (V.T.); Girininkai env., bank of sand trench, 04 06

56 NEW AND RARE FOR LITHUANIA INSECT SPECIES. Volume 27
1992, 1 spec. (V.T.); Gyvoliai env., bank of Virvytė r., 25 07 1999, 5 spec.
(Vidm.M.); Grybaulia env., 1 08 1984, 3 spec. (Vidm.M.); Jiesia lanscape rezerve,
bank of Jiesia r., 28 04 1994, 2 spec. (R. F.); Kajackaraistis env.2, bank of sand pit, 11
06 1992, 1 spec. (R.F.); Kalviai env., bank of Nemunas r., 19 07 1999, 1 spec. (R.F.);
Kiminiškė env., 02 08 1991, 3 spec. (Vidm.M.); Lyduvėnų env., bank of Dubysa r.,
26 07 2006, 2 spec. (V. T.); Mickiškė env., bank of Šešuvis r., 31 07 2008, 4 spec.
(V.T. & B.T.); Simokai env., bank of Šešupė r., 09 05 2012, 4 spec. (V.T. & B.T.);
Nemajūnų Miškas f., bank of sand pit, 20 06 2009, 1 spec. (G.S.); Nida env., 24 06
1991, 30 06 2007, beach of Baltic Sea (4, 1) spec. (V.T); Pakalniškiai env., bank of
sand pit, 20 06 2015, 2 spec.. (V.T. & B.T.); Paketuriai env., 23 05 1994,1 spec.
(R.P.); Pervalka env., 13 07 2000, 1 spec. (R.F.); Puvočiai env., 30 06 1993, 1 spec.
(Anonimous); Reiskiai env., 16 07 1994, 1 spec. (S.K.); Siponys env., bank of
Nemunas r, June 1980, 1 spec. (G.Sl.); Škėvonys env., banko f Nemunas r., 28 05
2005, 1 spec. (V.U.); Tabariškiai env., banko f sand trench, 01–05 06 1992, 6 spec.
(V.T); Traklaukas env., bank of Šešuvis r., 11 07 2008, 1 spec. (V.T.); Varniai env.
July 1966, 1 spec.( S.P.); Viršužiglis env., bank of Kauno marios pond, 14 07 1994,
14 06 2006, (1, 1) spec. (V.T.); Želsva env., 04 08 1999, 2 spec. (Vidm.M.).
Notes. West Palaearctic species widely distributed Central and Southern Europe
(Hůrka, 2003), but rare in northern border of its distribution range (Lindroth, 1992;
Pawłowski et al., 2002). It was considered as rare species in Lithuania as well (Pileckis
& Monsevičius, 1995) and was mentioned just for several localities before (Sharova &
Grüntal, 1973; Ferenca, 2004, 2006; Ivinskis et al., 2009; Ferenca 2014). Assuming new
findings we conclude that this species is quite frequent and widely distributed in
Lithuania; inhabits sandy banks of various water bodies, sometimes even in relatively
small or newly made ones, like sand quarries, pits, trenches and etc. Observations of
adults in Lithuania: IV: 2; V: 20; VI: 22; VII: 25; VIII: 10.
Subfamily Elaphrinae Erichson, 1837
Tribe Elaphrini Erichson, 1837
Blethisa multipunctata (Linnaeus, 1758)
Aukštumala bog, 03–31 05 2003, 1 spec. (V.U.); Juknaičiai env., 10 06 1997, 1 spec.
(A.G.); Lapių Miškas f., 28 05 2009, 1 spec. (R.F.); Margininkai, 05 06 1994, 1 spec.
(V.T.); Smiltynė env.1, 12 07 1985, 1 spec. (S.K.); Trakai env., 23 04 1966, 1 spec.
(E. G.).
Notes. This species is widely distributed in temperate and boreal zones of Eurasia and
North America (Goulet et al., 2009). The data on distribution of this species in Lithuania
is poor. Previous records were known only from Curronian Spit (Šablevičius, 2003;
Ivinskis et al., 2004; Ferenca, 2014), Žuvintas Nature reserve and Tyruliai bog (Ferenca,
2006). As rare species it is listed in the Red lists of some European countries (Pawłowski
et al., 2002). This stenotopic hygrophilous species occurs on the banks of stagnant or
slow-flowing fresh or brackish waters (Lindtroth, 1992). Lithuanian catches: IV: 1; V: 7;
VI: 6; VII: 2; VIII: 3; X: 5.
Elaphrus aureus P.W.J. Müller, 1821
Ariogala env., 09 05 1979, 1 spec. (Anonymous); Eržvilkas env., bank of Šaltuona r.,
11 05 2012, 1 spec. (V.T.); Gimbetiškė env., 17 05 1991, 2 spec. (Vidm. M.); Jiesia
Landscape rezerve, bank of Jiesia r., 05 08 1981, 28 041994, (1, 1) spec. (R.F.);

NAUJOS IR RETOS LIETUVOS VABZDŽIŲ RŪŠYS. 27 tomas 57
Pabariukai env., bank of Dubysa r., 12 07 2008, 2 spec. (V.T.); Pašilių Miškas f.,
bank of Ašarėna r., 18 05 1994, 1 spec. (R.F.); Raguviškiai env., bank of Minija r., 12
06 2000, 2 spec. (S.K.); Ramoniškė env., bank of Uogys r., 26 06 1991, 09 08 1993,
(1, 1) spec. (Vidm.M.); Rubuliai env., bank of Minija r., 14 04 1989, 4 spec. (S.K.);
Viešvilė Nat. R.4, bank of Viešvilė r., 31 07 2008, 5 spec. (V.T. & B.T.); Varlaukis
env., bank of Šešuvis r., 07 05 2012, 6 spec. (V.T. & B.T.).
Notes. This European species is widely distributed in Central Europe (Goulet, 2003),
rare in northern border of its distribution range (Barševskis, 2003; Günther & Hölscher,
2004; Aleksandrowicz, 2014). In Lithuania this species was considered as very rare
(Pileckis & Monsevičius 1995). Just few records from Zarasai environs (Pileckis, 1968),
Lomena Lanscape reserve (Gedminas et al., 1996) and Kartenalė Entomological reserve
(Ferenca et al., 2002) have been published previously. Newly found specimens were
discovered in open (generally sun exposed) or slightly shaded with sparse vegetation
river banks on sandy or sandy-loamy soil. Observations of adults in Lithuania: IV: 6; V:
11; VI: 4; VII: 10; VIII: 4.
Subfamily Scaritinae Bonelli, 1810
Tribe Clivinini Rafinesque, 1815
Clivina collaris (Herbst, 1784)
Jiesia landscape reserve, bank of Jiesia r., 08 08 1981; 27 05 1999, (1, 1) spec. (R.F.);
Miknaičiai env., bank of Šešupė r., 09 05 2012, 7 spec. (V.T. & B.T.); Sėlų Miškas f.,
bank of Visinčia r., 19 06 1985, 1 spec. (R.F.); Varlaukis env., bank of Šešuvis r., 07
05 2012, 1 spec. (V.T. & B.T.).
Notes. It is poorly known species in Lithuania. To this day only few records have
been documented in the country (Ogijewicz, 1933, Sharova & Grüntal, 1973; Ferenca,
2006). Newly found specimens were discovered in loamy and slimy, without vegetation
river banks, close to water line. Observations of adults in Lithuania: IV: 1; V: 12; VI: 5;
VIII: 2.
Subfamily Broscinae Hope. 1838
Tribe Broscini Hope. 1838
Miscodera arctica Eschscholtz, 1830
Čižiškė env., 20 06 2005, 11 07 2005, 02 08 2005, (2, 4, 2) spec. (P.I.); Druskininkai
env., 24 05 1996, 08 08 1996, (1, 1) spec. (V.T.); Girionys env., 30 07 1983, 1 spec.
(R.F.); Kajackaraistis env.2, 24 08 1984, 1 spec. (R.F.); Kuodiškiai env., 29 09 1982,
1 spec. (G.Sl.); Viešvilė Nat. R.3, 09 05 2008, 2 spec. (V.T. & B.T.); Randamonys
env., 07 09 2001, 1 spec. (A.G.); Ratnyčia env., 24 05 1996, 1 spec. (V.T.);
Skriaudžių Miškas f., 02 07 1991, 1 spec. (A.M.); Užuožerės Miškas f., 28 06 1984, 1
spec. (R.F.).
Notes. This species is distributed in boreal zone in Eurasia and North America
(Bousquet, 2003), also is known from mountain region of Central Europe (Hůrka, 1996).
In the plains of Central Europe M. arctica is distributed very localy and sporadicaly, and
is considered as a rare species (Pawłowski et al., 2002; Pileckis & Monsevičius, 1995;
Gürlich et al., 2011; Aleksandrowicz, 2014). In Lithuania just few records from
Dubravos Miškas f. (Pileckis, 1968), Varėna district (Pileckis & Monsevičius, 1995) and
Baluoša peat bog (Ferenca et al., 2002) have been published previously. Newly found
specimens were discovered in dry forests with sandy soils, generally in clearings or

58 NEW AND RARE FOR LITHUANIA INSECT SPECIES. Volume 27
forest edges. Observations of adults in Lithuania: V: 4; VI: 6; VII: 10; VIII: 4; IX: 4; XI:
1.
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62 NEW AND RARE FOR LITHUANIA INSECT SPECIES. Volume 27
Žiogas A. & Vaičikauskas S. 2007a. Ground beetles (Coleoptera, Carabidae)
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Nauji duomenys apie retas ir mažai žinomas žygių rūšis (Coleoptera: Carabidae:
Nebriinae, Carabinae, Omophroninae, Elaphrininae, Scaritinae, Broscinae)
Lietuvoje
V. TAMUTIS, B. TAMUTĖ, R. FERENCA
Santrauka
Straipsnyje pateikiami nauji faunistiniai duomenys apie 13 žygių rūšių: Nebria
brevicollis (F.), Leistus rufomarginatus Duft., Leistus piceus Fröl., Notiophilus germanyi
Fauv., Carabus clatratus L., Carabus convexus F., Carabus ritens L., Carabus violaceus
andrzejuscii Fisch., Omophron limbatus (F.), Blethisa multipunctata (L.), Elaphrus
aureus Müll., Clivina collaris (Hbst.), Miscodera arctica Esch. Lietuvoje. Apžvelgiami
šių rūšių paplitimas ir ekologijos ypatumai.
Received: 20 October, 2015