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Accepted by M. Pellinen: 13 Oct. 2015; published: 3 Dec. 2015
ZOOTAXA
ISSN 1175-5326 (print edition)
ISSN
1175-5334
(online edition)
Copyright © 2015 Magnolia Press
Zootaxa 4052 (2): 251
–
296
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http://dx.doi.org/10.11646/zootaxa.4052.3.1
http://zoobank.org/urn:lsid:zoobank.org:pub:775796D2-BEDF-459B-B937-FAC0EC457B92
Contribution to the knowledge on the Palaearctic and Oriental taxa of the
Meganola s.l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with
descriptions of 4 new genera and 11 new species
GYULA M. LÁSZLÓ
1,4
, GÁBOR RONKAY
2
& LÁSZLÓ RONKAY
3
1
Fadrusz u. 25, H-1114 Budapest, Hungary.
2
Heterocera Press Ltd., Szt. István krt. 4, H-1137 Budapest, Hungary.
3
Hungarian Natural History Museum, Department of Zoology, Baross u. 13, H-1088 Budapest, Hungary.
4
Corresponding author. E-mail: gyula.m.laszlo@gmail.com
Abstract
Present paper contains the descriptions of four new genera distinguished from the collecting genus Meganola s.l. (Hamp-
sonola, Wittonola, Fragilonola and Maculonola gen. n.) and 11 new species (Meganola pseudobasalactifera, Hampsono-
la diehli, H. subbasirufa, H. angustifasciata, H. stueningi, H. micra, Wittonola latifasciata, Nanola rothschildi,
Fragilonola fragilis, F. parentela and Maculonola dolokmerangirensis spp. n.) from South East Asia. Based on genital
morphology the concept of the genus Nanola is extended, involving several species treated earlier as Meganola. Meganola
yakovlevi László, Ronkay & Ronkay, 2010 is synonymised with M. tetrodon (de Joannis, 1928), and M. pekarskyi László,
Ronkay & Ronkay, 2014 is synonymised with Nanola liaoningensis (Han & Li, 2008), syn. n. 45 new combinations and
2 new stati are established. With 48 colour photos and 42 genitalia figures.
Key words: Meganola, Nanola, Nolini, Nolinae, Nolidae, Noctuoidea, taxonomy, new genera, new species, new synon-
ymy, new combination, new status, South East Asia
Introduction
The genus Meganola was established by Dyar (1898) for a Nearctic species Meganola conspicua Dyar, 1898. The
generic name has been introduced for the Old World fauna by Poole (1989) who applied it for practically all taxa
previously treated by authors as Roeselia Hübner, 1825 except e.g. Sarbena Walker, 1862 and Proneca Swinhoe,
1890, which were considered by Poole as distinct genera. It is worth to note that Roeselia is synonymous with Nola
Leach, 1815 due to its unfortunate type-species designation. Grote choose Phalaena Tinea cucullatella Linnaeus,
1758 as type-species of Roeselia, since the type-species of Nola is, by monotypy, Phalaena Noctua palliola [Denis
& Schiffermüller], 1775. As these two names refer to the same species, both genera have the common type-species,
therefore Roeselia is a mere synonym of Nola.
The delineation of this major genus has long been problematic, lacking a proper and consistent generic
diagnosis. The quadrifine hindwing venation has long been deemed as a distinctive morphological character of
Meganola (s.l.), but this concept produced an undoubtedly paraphyletic assemblage. Holloway (2003) clarified
first the taxonomic position of the (otherwise also extraordinarily species rich) genus Manoba, correcting the
erroneous traditional treatment of the group which considered Manoba as synonymous with the lithosiine genus
Stictane Hampson, 1900, and restored its status as a valid noline genus based on the trifine hindwing venation as
main diagnostic character besides the characteristic configuration of the genitalia. The species belonging to the
other large branches of this generic complex remained, however, in the paraphyletic and diverse generic unit of
Meganola (s.l.). It is worth to note that concerning the hindwing venation of the Meganola complex some
discrepancies between publications of several authors have been found. Holloway (2003) divided Meganola and
Manoba based on the quadrifine hindwing venation (with M
3
and CuA
1
stalked) of the former and trifine one (with
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·
Zootaxa 4052 (2) © 2015 Magnolia Press
M
3
and CuA
1
fused) of the latter genus, while Zahiri et al (2013) refers to Meganola having trifine hindwing
venation with fused M
3
and CuA
1
. To get a clearer picture we investigated the hindwing venation of species of
several genera of the Meganola s.l. complex; the results are summarised in the following:
Ctenane: M
2
present, M
3
+CuA
1
stalked (variably long within the genus, the stalking section is much longer in the
trianguloquelinea-group than in the labuana-group).
Fragilonola: M
2
present, M
3
+CuA
1
fused.
Hampsonola: M
1
+R
1-5
stalked, M
2
present, M
3
+CuA
1
fused.
Maculonola: M
2
present, M
3
+CuA
1
stalked (with medium-long stalking section).
Manoba: M
2
present, M
3
+CuA
1
fused.
Meganola: M
2
present, M
3
+CuA
1
fused in M. conspicua (the type-species of the genus) but M
3
+CuA
1
is long stalked in many
Oriental species which can explain the discrepancy between the statements of Holloway (2003) and Zahiri et al. (2013).
Nanola: M
2
present, M
3
+CuA
1
stalked (relatively shortly).
Proneca: M
2
present, M
3
+CuA
1
stalked.
Sarbena: M
2
present, M
3
+CuA
1
stalked.
Selca: M
2
present, M
3
+CuA
1
fused.
Wittonola: M
2
present, M
3
+CuA
1
fused.
According to our investigation the quadrifine and trifine type of hindwing venation both widespread in the generic
complex, the clarification of the phylogenetic importance of this character requires examination of significantly
more taxa. According to the present knowledge and literature references the authors of the present paper do not
deem hindwing venation as from a taxonomical point of view so important character in the reconstruction of the
generic level system of the Nolini as it is supposed in the former works. We refer, therefore, to the hindwing
venation in the diagnostic parts of the genera discussed in this paper without further comment.
Twelve years ago a comprehensive taxonomic research project on the rather neglected tribe Nolini was
initiated by Mr Thomas Witt and the authors of the present paper. The access of the vast collection material of the
Museum Witt, Munich promoted very much the investigations at the beginning of the project. Later, during the
progress of the study, more and more museum collections have been involved to the investigation, resulting in a
continuous discovery of new taxa. One of the main target groups of this study was the Meganola genus-group
(László, Ronkay & Witt 2004, 2005, 2006, 2007a, b, 2010, László, Ronkay & Ronkay, 2013a, b, c, d, 2014a, b, c,
d, e, f, 2015). The examination of a large Nolini material from eastern and south-eastern Asia and the thorough
study of the type specimens provided the discovery of a large number of new Meganola species and led to the
establishment of a number of new genera. Due to the increase of the newly discovered taxa, more and more
morphological characters have been learnt which allowed to delineate smaller taxonomic units within the
Meganola (s.l.) lineage. These studies also provided additional information for the assessment of such historic
genera as Selca Walker, 1866, Proneca Swinhoe, 1890, Ctenane Swinhoe, 1905, etc. and the clarification of their
phylogenetic relationship with the lineages of Meganola (s.l.) (László, Ronkay & Ronkay, in press).
The species content of the Meganola lineage has been rapidly increased after the basic works of Inoue (1958,
1970, 1991, 1996, 1998, 2001), due to the studies of the authors and the other experts of this field (Hu et al. 2013a,
b, 2014, Han & Li 2008, Shao, Li & Han 2009, and Hacker et al. 2012). This last work has an eminent importance
due to the designation of the lectotype of M. conspicua, the type species of the genus, and the illustration of its
genitalia.
As a result of the works of the above mentioned authors, Meganola (s.l.) has become one of the most species
rich noline genera in the Palaearctic, Oriental and Afrotropical zoogeographical regions, especially in the forested
habitats including both the continental areas and the Archipelago. Thus, a complete taxonomic revision of this
difficult generic complex would be highly desirable. In spite of the thorough and exhausting examinations standing
behind the present work as well as the studies of the above mentioned authors, it is not yet possible, by the
following reasons. This extraordinarily species-rich and diverse generic complex is distributed worldwide,
therefore morphotaxonomic investigation of all known taxa from each continent is inevitable – this task requires
investigations of further several thousand specimens and numerous primary types scattered in museum and private
collections worldwide, which is beyond the actual possibilities of the authors. Moreover, the clarification of the
phylogenetic relationships within the generic complex (as well as between all supraspecific Nolini taxa) requires
the application of the modern molecular methods. This extensive molecular taxonomic study can only be a topic of
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PALAEARCTIC AND ORIENTAL MEGANOLA
a new project with firm institutional background where the funds and the technical conditions are given to acquire
molecular genetic data in large numbers. A molecular-based phylogenetic concept of Nolinae is published recently
by Zahiri et al. (2013), outlining two well established lineages within the subfamily, the Nola-Meganola-Manoba
and Melanographia-Membranola-Alcanola clades. The number of samples involved were only 82 specimens
representing all subfamilies of Nolidae, which is definitely not eligible for a sound phylogenetic analysis of lower
level units as e.g. tribes or genus groups. This should be the reason of the ignorance of numerous genera of the
Nolini (in the old sense; e.g. Aeneanola, Porcellanola, Barnanola, Casminola, Inouenola, Mokfanola etc.). This
publication pinpoints the urgent need of the synthetic assessment of the classic and the molecular taxonomic
studies. The involvement of substantially more morphotaxa to the molecular genetic studies would produce
apparently more and more reliable results on the knowledge of the phylogeny of the entire subfamily. The mutual
benefit of the cooperation would be a better founded delineation of the main Nolini clades, leading to the
establishment of a complete cladistic system being supported by both the morphological and DNA information.
The main aim of this present work is to give a clearer picture on the taxonomy of the Oriental taxa of the
Meganola s.l. complex, attempting to identify supposedly monophyletic units within the diverse collecting genus
based on morphotaxonomic information gained by our recent investigations. As a result, four main lineages have
been delineated based on genital morphology and described here as new genera, and further 11 new species are
described alongside with numerous new taxonomic statements. The newly described species and all new
combinations are provided following the diagnosis of the six distinguished genera of the generic complex.
Material and methods
The conclusions of the present work are based on, besides the thorough study of the literature, the examination of
more than 8.000 Nolinae specimens preserved in the following museum collections: Hungarian Natural History
Museum, Budapest; Natural History Museum, London; Museum Witt, Munich; Zoological State Collection,
Munich; Zoological Research Museum Alexander Koenig, Bonn; Naturalis Biodiversity Center, Leiden; State
Museum of Natural History, Karlsruhe; Zoological Museum of the University, Copenhagen; Natural History
Museum, Vienna. Some 3000 specimens have been photographed during the studies using Nikon D90 camera. The
images were adjusted by Adobe Photoshop CS3 software. During the studies more than 2500 specimens have been
dissected using the standard method. The detached abdomens had been macerated in 10% KOH and boiled in
water-bath for several minutes. The softened abdomens were cleared of hairs, scales and intestine residuals, then
the genital apparatus had been dissected off, soaked in lactic acid, then cleared and flattened in the standard
position using 96% ethanol for desiccation. The properly prepared genital apparatures have been stained with
eosine red and mounted on microscopic slide in euparal. During the dissection and mounting Nikon SMZ745
stereomicroscope has been used. The genitalia photographs have been taken using a Nikon Eclipse 80i compound
microscope and a Nikon DS-Fi1 digital camera.
Abbreviations
BM — number of genital slide of BMNH
BMNH — British Museum of Natural History, London
HNHM — Hungarian Natural History Museum, Budapest
LGN — Number of genital slide made by Gy.M. László
MWM — Museum Witt, Munich
NEFU — Northeast Forestry University, Harbin
RMNH — Royal Museum of Natural History (Naturalis Biodiversity Center), Leiden
SCAU — South China Agriculture University, Guangzhou
SMNK — State Museum of Natural History, Karlsruhe
USNM — Smithsonian Institution, National Museum of Natural History, Washington D.C.
W — Number of genital slide of Museum Witt
ZFMK — Zoological Research Museum Alexander Koenig, Bonn
ZMUC — Zoological Museum of the University, Copenhagen
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Taxonomic part
Characterization of the genus Meganola Dyar, 1898 and description of 4 new genera and 11 new species
Genus Meganola Dyar, 1898
(Plate 1, Figs 1–4; gen. figs 1–3)
Meganola Dyar, 1898, Journal of the New York Entomological Society 6: 43. Type-species: Meganola conspicua Dyar, 1898,
ibidem, 6: 43, by original designation by monotypy.
Synonymy
Antennola de Freina & Witt, 1984, Entomofauna 5(23): 267. Type-species: Nola impura Mann, 1862, Wiener Entomologische
Monatschrift 6: 368, pl. 3, fig. 3, by original designation by monotypy;
Poliothripa Hampson, 1900, Annals of the South African Museum 2: 311. Type-species: Poliothripa niphostena Hampson,
1902;
Metanola van Son, 1933, Annals of the Transvaal Museum 15: 203. Type-species: Metanola myriostigma van Son, 1933;
Paranola van Son, 1933, Annals of the Transvaal Museum 15: 222. Type-species: Paranola bipartita van Son, 1933.
Remarks. Hacker et al. (2012) erroneously referred Poecilonola Hampson, 1900 as synonym of Meganola, as the
type-species of the former genus, P. plagiola Hampson, 1989, belongs to the genus Evonima Walker, 1865. Thus
Poecilonola is a synonym of this latter genus as it is referred correctly by Holloway (2003).
Characterization of the genus. The type-species of the genus was described from Texas, USA, nevertheless
the genus is widespread in all continents. The hindwing venation, used generally for the characterisation of the
Nolini genera, is variable throughout the genus, therefore the general statements about this feature (Holloway 2003,
Hacker et al. 2012, Zahiri et al. 2013) are to be unified as the veins M
3
and CuA
1
are either fused or variably long
stalked. The external features show so great variability for practically all morphological characters even in the
restricted sense of Meganola (due to the high number of species, see, for instance, the Plates 6–11 in László,
Ronkay & Ronkay 2014b) that none of them can be used for the general characterisation of the genus but their
species-groups only.
The generic characters are expressed in the genitalia as follows (for the genitalia figures of the type species see
Hacker et al. 2012).
Male genitalia. Uncus always present, variably long, slender to medium thick, apically pointed; socii absent;
tegumen simple, without processes (except in M. scriptoides Holloway, 2003); valva unilobate, in some species
groups with short ventro-basal dilatation, elongate, apically variably rounded or sometimes truncate, generally
slightly curved; harpe present, variable in shape and size, arising from basal plate of valva, being located rather far
from ventral margin; sacculus generally weakly developed, in most taxa without processes; vinculum well
developed, variably long, U- or V-shaped. Aedeagus tubular, with variable length and thickness; apical process
(carinal extension) may be present. Vesica with or without cornuti; armature of cornuti variable in number and size.
Female genitalia. Ovipositor short, conical, apophyses short or medium-long, eighth tergite short,
quadrangular or medially tapering; ostium bursae and antrum generally strongly sclerotized; ductus bursae usually
long, variably thick, with often strongly sclerotized distal section; corpus bursae large, ovoid, generally with a pair
of longitudinally aligned, acute, crest- or process-like signa, which in some species connected with ribbon-like
scobinated plate.
List of the Palaearctic and Oriental species of the genus Meganola s. str., with taxonomic and nomenclatural
novelties
Remarks. The taxon described as Roeselia oblita by Wileman & West in 1929 is combined by Poole (1989) with
the genus Meganola. This taxon is, however, synonymous with Manoba phaeochroa (Hampson, 1900), syn. n.,
therefore it is deleted from the following checklist of Meganola.
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PALAEARCTIC AND ORIENTAL MEGANOLA
PLATE I. Adults of Meganola and Nanola species.
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Meganola conspicua Dyar, 1898
Meganola conspicua Dyar, 1898, Journal of the New York Entomological Society 6: 43. Type-locality: USA,
Arizona, Fort Grant. Lectotype: female, in coll. USNM.
Meganola gigantula (Staudinger, 1878)
Roeselia gigantula Staudinger, 1878, Horae Societatis Entomologicae Rossicae 14: 328. Type-locality: [Turkey]
Amasia. Holotype: female, in coll. ZMHU.
Meganola tenebrosa (Hampson, 1896)
Nola tenebrosa Hampson, 1896, Fauna of British India, Moths 4: 504. Type-locality: Bhutan. Holotype: female, in
coll. BMNH.
Synonymy
Manoba greenwoodi Holloway, 2003, The Moths of Borneo 18: 44, pl. 2, fig. 58. Type-locality: Borneo, Sarawak,
Gunong Mulu National Park, Mulu, 500 m. Holotype: male, in coll. BMNH.
Meganola guentereberti László, Ronkay & Ronkay, 2014
Meganola guentereberti László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 38–39, pl. 9, figs 5–6; gen.
figs 24–25. Type-locality: Nepal, Annapurna Himal, Tadapani. Holotype: male, in coll. MWM.
Meganola margalla László, Ronkay & Ronkay, 2014
Meganola margalla László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 38, pl. 9, figs 3–4; gen. figs 22–
23. Type-locality: Pakistan, Islamabad, Margalla Hills. Holotype: male, in coll. MWM.
Meganola basisignata Inoue, 1991
Meganola basisignata Inoue, 1991, Tyo to Ga 42(2): 69, figs 9, 10a, 10b. Type-locality: Japan, Miyagi pref.,
Sakunami Spa. Holotype: male, in coll. BMNH.
Meganola suffusata (Wileman & West, 1929)
Nola suffusata Wileman & West, 1929, Annals and Magazine of Natural History 10(3): 189. Type-locality:
[Taiwan] Formosa, Kanshirei. Holotype: female, in coll. BMNH.
Meganola eleagni Stshetkin, 1980
Nola elaeagni Stshetkin, 1980, Izvestija Akademii Nauk Tadshikskoj SSR, Otdelenie biologitsheskih Nauk 4(81):
98–100. Type-locality: Tajikistan, Dushanbe. Holotype: male, in coll. ZIN.
Meganola arenbergeri Hacker, 2012
Meganola arenbergeri Hacker, 2012, Esperiana 17: 229. An objective replacement name for Roeselia nanula
Wiltshire, 1961, Beiträge zur Naturkundischen Forschung in Südwestdeutschland 19: 338, pl. 2, figs 4, 6, 7, text
figs 2–7. Type-locality: Afghanistan, vicinity of Kabul. Holotype: male, in coll. ZSM.
Meganola impura
impura (Mann, 1862)
Nola impura Mann, 1862, Wiener Entomologische Monatschrift 6: 368, pl. 3, fig. 3. Type-locality: [Turkey] Asia
Minor, Brussa.
Synonymy
Nola impura wiltshirei Warnecke, 1939, in Ellison and Wiltshire, Transactions of the Entomological Society of
London 88: 26. Type-locality: Lebanon, Beyrouth. Syntypes: two males, in coll. BMNH.
Meganola impura centralis (Warnecke, 1938)
Nola impura ssp. centralis Warnecke, 1938, Entomologische Rundschau 55: 295. Type-locality: [China] [Xinjiang]
Kuldja. Syntypes: in coll ZMHU.
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PALAEARCTIC AND ORIENTAL MEGANOLA
FIGURES 1–3. Male genitalia of Meganola species.
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Synonymy
Nola silvicola Stshetkin, 1957, Izvestija Otdelenija Estestvennih Nauk AN Tadjikiskoi SSR 21: 124–128. Without
type designation.
Meganola togatulalis (Hübner, 1796)
Pyralis togatulalis Hübner, 1796, Sammlung Europäischer Schmetterlinge 6: pl. 20, fig. 131. Type-locality:
Europe.
Synonymy
Nola togatulana Duponchel, [1845] 1844, an unjustified emendation of Pyralis togatulalis Hübner, 1796.
Meganola pulverata (Wileman & West, 1929)
Rhynchopalpus pulverata Wileman & West, 1929, Annals and Magazine of Natural History 10(3): 193. Type-
locality: [Taiwan] Formosa, Kanshirei. Holotype: male, in coll. BMNH.
Meganola ascripta (Hampson, 1894)
Selca ascripta Hampson, 1894, Fauna of British India, Moths 2: 144. Type-locality: [India] [Nagaland] Naga Hills,
5000–7000 feet. Holotype: male, in coll. BMNH.
Meganola subascripta Hu, László, Ronkay & Wang, 2013
Meganola subascripta Hu, László, Ronkay & Wang, 2013, Zootaxa 3608 (7): 596, figs 1–4. Type-locality: China,
Guangdong, Nanling. Holotype: male, in coll. SCAU.
Meganola argentescens (Hampson, 1895)
Pisara argentescens Hampson, 1895, Catalogue of the Lepidoptera Phalaenae in the British Museum 5: 296. Type-
locality: [India] Sikkim. Holotype: female, in coll. BMNH.
Meganola jinghongensis (Shao, Li & Han, 2009) comb. nov.
Manoba jinghongensis Shao, Li & Han, 2009, Tinea 20 (5): 307–308. Type locality: China, Prov. Yunnan,
Jinghong city. Holotype: male, in coll. NEFU, Harbin.
Meganola tarka László, Ronkay & Witt, 2010
Meganola tarka László, Ronkay & Witt, 2010, Esperiana 15: 45, pl. 11, figs 16–17; gen. fig. 46. Type-locality:
Thailand, Chantaburi, Khao Soi Dao, 300 m. Holotype: male, in coll. BMNH.
Meganola tomkovichi László, Ronkay & Ronkay, 2014
Meganola tomkovichi László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 37, pl. 8, figs 7–8; gen. figs 20–
21. Type-locality: India, Daitari, Orissa. Holotype: male, in coll. HNHM.
Meganola gigas (Butler, 1884)
Nola gigas Butler, 1884, Annals and Magazine of Natural History 5(13): 274. Type-locality: [Japan] Yesso.
Holotype: male, in coll. BMNH.
Synonymy
Nola maculata Staudinger, 1877, Mémoires sur les Lépidoptéres 3: 180. Type-locality: [Russia] [Far East] Suifun.
Syntypes: two males, in coll. ZMHU;
Roeselia nigromaculata Nagano, 1918, Insect World 22: 414. Type-locality: Japan.
Meganola protogigas (Inoue, 1970)
Roeselia protogigas Inoue, 1970, Bulletin of the Japan Entomological Academy 6: 3, pl. 1, figs 1, 35. Type-
locality: Japan, Shimane Prefecture, Ichibata. Holotype male, in coll. BMNH.
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PALAEARCTIC AND ORIENTAL MEGANOLA
FIGURES 4–6. Genitalia of Meganola species.
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Meganola bicoloria László, Ronkay & Ronkay, 2014
Meganola bicoloria László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 41, pl. 10, figs 5–6; gen. figs 33–
34. Type-locality: China, Chekiang, West-Tien-Mu-Shan. Holotype: male, in coll. ZFMK.
Meganola subgigas (Inoue, 1982)
Meganola subgigas Inoue, 1982, in Inoue et al., Moths of Japan 1: 665, pl. 229, figs 45–46, pl. 354, fig. 8, pl. 351,
fig. 10. Type-locality: Japan, Hokkaido, Oshima, Nakayama Pass. Holotype: male, in coll. BMNH.
Meganola major major (Hampson, 1891)
Nola major Hampson, 1891, Illustrated Typical Specimens of the Lepidoptera Heterocera in the Collection of the
British Museum 8: 4, 48, pl. 139, fig. 13. Type-locality: India [Tamil Nadu], Nilgiri district. Syntypes: in coll.
BMNH.
Meganola major caesiopennis (Inoue, 1982)
Meganola major caesiopennis Inoue, 1982, in Inoue et al., Moths of Japan 1: 666. Type-locality: Japan, Amami-
Oshima Island, Nishninakama. Holotype: male, in coll. BMNH.
Meganola major fulvusana Sung-Hwan Oh, 1991 stat. n.
Meganola fulvusana Sung-Hwan Oh, 1991, Systematics of the family Nolidae (Lepidoptera) in Korea: 66, figs 18,
50, 81, 112. Type-locality: Korea (South), Prov. Gangweon, Yangyang Holotype: male, in coll. Kangwon National
University, Chuncheon.
Remarks. Oh (1991) has published several new taxa in his PhD theses. Although the validity of these taxa can be
disputed due to the publication process may be insufficient for taxonomic statements but as the theses are available
freely, we treat here these taxa as valid items.
Meganola major phaea (Hampson, 1900) stat. n.
Nola phaea Hampson, 1900, Catalogue of the Lepidoptera Phalaenae in the British Museum 2: 35, pl. 19, fig. 2.
Type-locality: China, Ichang. Holotype: female, in coll. BMNH.
Meganola major takasago Inoue, 1982
Meganola major ssp. takasago Inoue, 1982, in Inoue et al., Moths of Japan 1: 666. Type-locality: [Taiwan]
Formosa. Holotype: male, in coll. BMNH.
Synonymy
Nola formosana Wileman & West, 1929, Annals and Magazine of Natural History 10(3): 190. Type-locality:
Philippines, Negros Island. Holotype: female in coll. BMNH. A junior secondary homonym of Roeselia formosana
Wileman & West, 1928; the replacement name is Meganola major ssp. takasago.
Meganola major mahera László, Ronkay & Ronkay, 2014
Meganola major mahera
László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 39, pl. 9, figs 7–8; gen. figs
26–27. Type-locality: Moluccas, Halmahera, Talagaranu Mt. Holotype: female, in coll. MWM.
Meganola brunellus (Hampson, 1893)
Roeselia brunellus Hampson, 1893, Illustrations of Typical Specimens of Lepidoptera Heterocera in the Collection
of the British Museum 9: 15, 89, pl. 158, fig. 31. Type-locality: [Sri Lanka] Ceylon, Pundaloya. Type(s): in coll.
BMNH.
Synonymy
Nola brunella Hampson, 1900, Catalogue of the Lepidoptera Phalaenae in the British Museum 2: 34. An
unjustified emendation of Nola brunellus Hampson, 1893;
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FIGURES 7–9. Genitalia of Meganola and Hampsonola species.
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FIGURES 10–12. Genitalia of Hampsonola and Xenonola species.
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Meganola pseudohypena Inoue, 1982, in Inoue et al., Moths of Japan 1: 667, pl. 154, figs 26–27, pl. 351, fig.
1, pl. 353, fig. 9. Type-locality: Japan, Ishigaki Island, Mt. Banna. Holotype: female, in coll BMNH;
Celamoides corticella van Eecke, 1926, Zoologische Mededeelingen Leiden 9: 39, pl. 2, fig. 10. Type-locality:
[Indonesia] Sumatra, Fort de Kock. Syntypes: nine specimens, in coll. RMNH Leiden.
Meganola flexilineata (Wileman, 1916)
Roeselia flexilineata Wileman, 1916, Entomogist 56: 132. Type-locality: Philippines, Luzon, Subprov. Benguet,
Palali, 2500 feet. Holotype: male, in coll. BMNH.
Synonymy
Celama scriptrix van Eecke, 1926, Zoologische Mededeelingen Leiden 9: 46, pl. 2, fig. 15. Type-locality:
[Indonesia] Sumatra, Fort de Kock, 920 m. Holotype: female, in coll. RMNH Leiden.
Meganola triangulalis (Leech, 1890)
Nola triangulalis Leech, 1890, Proceedings of the Zoological Society of London 1888: 608, pl. 31, fig. 12. Type-
locality: Japan, Satsuma. Holotype: male, in coll. BMNH.
Meganola albula albula ([Denis & Schiffermüller], 1775)
Noctua albula [Denis & Schiffermüller], 1775, Ankündung eines Systematischen Werkes von den Schmetterlinge
der Wiener Gegend 1775: 69. Type locality: [Austria] Vienna region. Type(s) destroyed.
Synonymy
Pyralis albulalis Hübner, 1796, Sammlung Europäischer Schmetterlinge 6: pl. 3, fig. 14. An unjustified
emendation of Noctua albula [Denis & Schiffermüller], 1775;
Roeselia albulana Hübner, [1825] 1816, Verzeichniss bekannter Schmettlinge: 397. An unjustified emendation of
Noctua albula [Denis & Schiffermüller], 1775;
Nola albula var. nivalis Caradja, 1934, Deutsche Entomologische Zeitschrift Iris 40: 189. Type-locality: Romania.
Type(s): in coll. Antipa Museum, Bucharest.
Meganola albula formosana (Wileman & West, 1928)
Roeselia formosana Wileman & West, 1928, Entomologist 61: 276. Type-locality: [Taiwan] Formosa, Kanshirei.
Holotype: male, in coll. BMNH;
Meganola albula mesotherma (Hampson, 1914)
Roeselia mesotherma Hampson, 1914, Catalogue of the Amatidae and Arctiidae (Nolinae and Lithosiinae) in the
Collection of the British Museum 1914: 427, pl. 24, fig. 23. Type-locality: China [Sichuan], Chunking. Holotype:
male, in coll. BMNH.
Meganola albula pacifica (Inoue, 1958)
Roeselia albula ssp. pacifica Inoue, 1958, Kontyu 26: 234. Type-locality: Japan, Shizuoka Pref., South Izu,
Nashimoto. Holotype: male, in coll. BMNH.
Meganola daminga Hu, Han & Wang, 2013
Meganola daminga Hu, László, Ronkay & Wang, 2013, Zootaxa 3608 (7): 600, figs 15–16. Type-locality: China,
Guangxi, Damingshan. Holotype: male, in coll. SCAU.
Meganola hoenei László, Ronkay & Ronkay, 2014
Meganola hoenei László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 42, pl. 11, fig. 3; gen. fig. 38. Type-
locality: China, Prov. Hunan, Hoeng-shan. Holotype: male, in coll. MWM.
Meganola ohsunghwani László, Ronkay & Ronkay, 2014
Meganola ohsunghwani László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 40, pl. 10, figs 1–2; gen. figs
29–30. Type-locality: China, Prov. Shaanxi, Mien-shan. Holotype: male, in coll. MWM.
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Meganola costalis (Staudinger, 1887)
Nola costalis Staudinger, 1877, Mémoires sur les Lépidoptéres 3: 179, pl. 10, fig. 3. Type-locality: [Russia] [Far
East] Askold [Island]. Syntypes: one female and three males, in coll. ZMHU.
Synonymy
Roeselia melanocosta Inoue, 1961, Check List of the Lepidoptera of Japan, (6), 1961: 683. Type-locality: Japan,
Gumma Pref., Kumanotaira. Holotype: male, in coll. BMNH.
Meganola postmediana László, Ronkay & Witt, 2010
Meganola postmediana László, Ronkay & Witt, 2010, Esperiana 15: 48, pl. 7, fig. 6; gen. fig. 44. Type-locality:
North Thailand, Prov. Chiang Mai, between Chiang Dao and Kariang, 900 m, 98°48’E, 19°25’N. Holotype: male,
in coll. MWM.
Meganola nitida (Hampson, 1894)
Selca nitida Hampson, 1894, Fauna of British India, Moths 2: 147. Type-locality: [India] Manipur. Holotype:
female, in coll. BMNH.
Meganola subnitida László, Ronkay & Witt, 2007
Meganola subnitida László, Ronkay & Witt, 2007, Entomofauna 28(14): 160, figs 5–7, 15. Type-locality: North
Thailand, Prov. Nan, Doi Phuka NP, between Pua and Bo Luang, 1350 m, 101°05’E, 19°12’N. Holotype: male, in
coll. MWM.
Meganola mediofusca László, Ronkay & Witt, 2007
Meganola mediofusca László, Ronkay & Witt, 2007, Entomofauna 28(14): 161, figs 8–10, 16, 20. Type-locality:
Taiwan, Ilan County, Ming Chyr Forest Recreation Area, 1200 m. Holotype: male, in coll. HNHM.
Meganola fuscimarginalis (Wileman, 1914)
Nola fuscimarginalis Wileman, 1914, Entomologist 47: 161. Type-locality: [Taiwan] Formosa, Garambi. Holotype:
male, in coll. BMNH.
Meganola punctilinea (Wileman & South, 1919)
Pisara punctilinea Wileman & South, 1919, Entomologist 52: 270. Type-locality: Philippines, Mindanao, Lanao,
Kolambugan. Holotype: female, in coll. BMNH.
Meganola calligrapha László, Ronkay & Witt, 2005
Meganola calligrapha László, Ronkay & Witt, 2005, Entomofauna 26(11): 208. Type locality: N. Thailand, Prov.
Chiang Mai, between Chiang Dao and Kariang. Holotype: male, in coll. MWM.
Meganola zolotuhini László, Ronkay & Witt, 2010
Meganola zolotuhini László, Ronkay & Witt, 2010, Esperiana 15: 47, pl. 7, figs 4–5; gen. fig. 43. Type-locality:
North Thailand, Prov. Chiang Mai, 1600 m, between Fang and Nor Lae, 99º06’E, 20º02’N. Holotype: male, in coll.
MWM.
Meganola ruficostata (Hampson, 1896)
Selca ruficostata Hampson, 1896, Fauna of British India, Moths 4: 507. Type-locality: Bhutan. Holotype: female,
in coll. BMNH.
Meganola cuneifera (Walker, 1862)
Melia cuneifera Walker, 1862, Journal of the Proceedings of the Linnean Society of London (Zoology) 6: 127.
Type-locality: Borneo, Sarawak. Holotype: male, in coll. UM Oxford.
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FIGURES 13–15. Genitalia of Hampsonola species
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Meganola brechlini László, Ronkay & Ronkay, 2014
Meganola brechlini László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 41–42, pl. 10, fig. 8; gen. fig. 35.
Type-locality: China, Shaanxi, Tapaishan Mts, Tsinling Mt, Houzhenzi. Holotype: male, in coll. MWM.
Meganola shimekii (Inoue, 1970)
Roeselia shimekii Inoue, 1970, Bulletin of the Japan Entomological Academy 6: 5, figs 15–17, 37, 46. Type-
locality: Japan, Tokushima Pref., Tokushima City, Kuramoto. Holotype: male, in coll. BMNH.
Meganola albiscripta László, Ronkay & Witt, 2005
Meganola albiscripta László, Ronkay & Witt, 2005, Entomofauna 26(11): 210, figs 1, 19, 24. Type-locality:
Northern Vietnam, Fan-si-pan Mts, Sa Pa, 2400 m. Holotype: male, in coll. MWM.
Meganola mikabo (Inoue, 1970)
Roeselia mikabo Inoue, 1970, Bulletin of the Japan Entomological Academy 6: 4, pl. 1, fig. 2. Type-locality: Japan,
Gumma Pref., Mikaboyama. Holotype: female, in coll. BMNH.
Meganola cenwanga Hu, László, Ronkay & Wang, 2013
Meganola cenwanga Hu, László, Ronkay & Wang, 2013, Zootaxa 3608 (7): 597, figs 9–10. Type-locality: China,
Guangxi, Cenwanglaoshan. Holotype: male, in coll. SCAU.
Meganola mediofascia (Inoue, 1958)
Celama mediofascia Inoue, 1958, Kontyu, 26: 236, pl. 2, figs 3, 8. Type-locality: Japan, Yamanashi Pref., Yashajin
Pass. Holotype: male, in coll. BMNH.
Meganola manoboides Holloway, 2003
Meganola manoboides Holloway, 2003, The Moths of Borneo 18: 29, pl. 1, figs 26, 40. Type-locality: Borneo,
Sarawak, Gunong Mulu NP, Mulu, 1000 m. Holotype: male, in BMNH.
Meganola strigivena (Hampson, 1894)
Selca strigivena Hampson, 1894, Fauna of British India, Moths 2: 147. Type-locality: [India] Sikkim. Holotype:
male, in coll. BMNH.
Meganola tetrodon (de Joannis, 1928)
Nola tetrodon de Joannis, 1928, Bulletin de la Société Entomologique de France 97: 250. Type-locality: [N
Vietnam] Tonkin, Hoang su phi. Holotype: male, in MNHN Paris.
Synonymy
Meganola yakovlevi László, Ronkay & Witt, 2010, Esperiana 15: 44, pl. 6, fig. 16; gen. fig. 41. Type-locality:
North Thailand, Prov. Chiang Mai, Mt. Doi Phahompok, 16 km NW of Fang, 2000 m. Holotype: male, in coll.
MWM, syn. n.
Meganola zegzugminta László, Ronkay & Ronkay, 2014
Meganola zegzugminta László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 42–43, pl. 11, figs 5–6; gen.
figs 39–40. Type-locality: Nepal, Annapurna Himal, Poon Hill. Holotype: male, in coll. MWM.
Meganola zirkalmashka László, Ronkay & Ronkay, 2014
Meganola zirkalmashka
László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 43, pl. 11, figs 7–8; gen. figs
41–42. Type-locality: Nepal, Koshi, Taplejung Area. Holotype: male, in coll. MWM.
Meganola strigula ([Denis & Schiffermüller], 1775)
Noctua strigula [Denis & Schiffermüller], 1775, Ankündung eines Systematischen Werkes von den Schmetterlinge
der Wiener Gegend 1775: 69. Type locality: [Austria] Vienna region. Type(s) destroyed.
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FIGURES 16–18. Genitalia of Hampsonola species.
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Synonymy
Phalaena aspersalis de Villers, 1789, Caroli Linnaei Entomologica, Faunae Suecicae Descriptionibus Aucta, 2:
451. Type-locality: Europe;
Pyralis strigulalis Hübner, 1796, Sammlung Europäischer Schmetterlinge 6: pl. 3, fig. 16. Type-locality:
Europe, an unjustified emendation of Noctua strigula [Denis & Schiffermüller], 1775;
Phalaena corticosa Vallot, 1802, Mémoires pour sevir a l’Historie de Insectes, 1802: 39. Type-locality: No
locality given;
Pyralis monachalis Haworth, 1911, Lepidoptera Britannica, sistens Digestionem novam Insectorum
Lepidopterorum que in Magna Britannia Reperiuntur, Larvarum pabulo, Temporeque Pascendi; Expansione
Alarum; Mensibusque Volandi; Synonymis Atque Locis Observationibusque Varris 1: 386. Type-locality: Great
Britain. Syntypes;
Roeselia strigulana Hübner, [1825] 1816, Verzeichniss bekannter Schmetterlinge: 397. An unjustified
emendation of Noctua strigula [Denis & Schiffermüller], 1775;
Nola lineolalis Eversmann, 1844, Fauna Lepidopterologica Volgo-Uralensis Exhibens Lepidopterorum
Species quas per Viginti quinque Annos in Provinciis Volgam Fluvium inter et Montes Uralenses sitis Observavit et
Descripsit, 1844: 474. Type-locality: [Russia] Ural Mts;
Nola zelleriana Fischer von Waldheim, 1824, Entomographia Imperii Rossici 1824.
Meganola kolbi (Daniel, 1935)
Roeselia strigula ssp. kolbi Daniel, 1935, Deutsche Entomologische Zeitschrift Iris 49: 33. Type-locality: Hungary,
Somogy megye, Balatonszentgyörgy. Lectotype: male, in coll. MWM, designated by de Freina and Witt, 1984.
Synonymy
Roeselia pannonica Kovács, 1947, Folia Entomologica Hungarica (New Series) 2: 67. Type-locality: Hungary.
Syntypes: in coll. HNHM;
Roeselia taurica Daniel, 1935, Deutsche Entomologische Zeitschrift Iris 49: 42. Type-locality: [Turkey]
Amanus, Dül-Dül dagh. Holotype: female, in coll. MWM.
Meganola strigulosa strigulosa (Staudinger, 1877)
Nola strigulosa Staudinger, 1877, Mémoires sur les Lépidoptéres 3: 180, pl. 10, fig. 4. Type-locality: [Russia] [Far
East] Vladivostok; Askold I. Syntypes: in coll. ZMHU.
Meganola strigulosa satoi (Inoue, 1970)
Roeselia satoi Inoue, 1970, Bulletin of the Japan Entomological Academy 6: 4, pl. 1, figs 19–21, 34, 49. Type-
locality: Japan, Niigata Pref., Nitsu City, Mt. Akiba. Holotype: male, in coll. Inoue.
Meganola tsinlinga László, Ronkay & Ronkay, 2014
Meganola tsinlinga
László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 39, pl. 10, fig. 7; gen. fig. 28.
Type-locality: China, Shaanxi, Qinling Mts. Holotype: male, in coll. MWM.
Meganola seima László, Ronkay & Ronkay, 2014
Meganola seima László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 44, pl. 12, fig. 1; gen. fig. 44. Type-
locality: Cambodia, Prov. Mondolkiri, Seima. Holotype: male, in coll. MWM.
Meganola pseudobasalactifera sp. n.
(Plate 1, Figs 3, 4; gen. figs 1, 2)
Holotype. Male, [Indonesia] Sumatra, Prapat, HW 3 [= “Holzweg 3”], 12.VIII.1983, leg. Dr. Diehl, slide No.: LGN 2158 (coll.
HNHM).
Paratypes. Indonesia, Sumatra: 1 male, from the same locality as the holotype, 31.I.1984; 1 male, from the same site, 4.X.1983;
2 males, from the same site, 28.X.1983; 1 male, from the same site, 28.IV.1985; 1 male, Sitahoan, 24.XI.1981, leg. Dr.
Diehl (coll. HNHM); 1 male, Dolok Merangir, 180 m, 2–13.VII.1970, leg. Dr. E. Diehl, slide No.: LGN 2062; 1 male,
Doulou, 1250 m, 18.IX.1969, leg. Dr. E. Diehl, slide No.: LGN 2061 (coll. SMNK).
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Diagnosis. The new species resembles externally mostly Nanola basalactifera and N. subbasalactifera (Plate 1,
Figs 5–8) having very similar forewing pattern. Their typical marking is the light basal part bordered by broad,
sharply defined and evenly arcuate reddish-brown stripe in the medial area of forewing included the blackish, also
evenly arcuate medial line. Despite this superficial similarity, M. pseudobasalactifera shows no closer relationship
with the two Nanola species, the large genital differences (gen. figs 1–7) support placing these three species into
two distinct genera.
Meganola pseudobasalactifera displays closer relationship, according to the configuration of the male
genitalia, with M. seima. The new species can be distinguished from M. seima (Plate 1, Figs 1, 2) by its
conspicuously larger whitish basal area extending from the costa to the ventral margin and more distally positioned
dark medial area (in M. seima the pale basal part is only an ovoid patch in the middle of the wing and the dark area
is present only between the antemedial and medial lines). The median fascia of M. pseudobasalactifera is evenly
arcuate, sharply defined, and blackish, while that of M. seima is diffuse, poorly visible.
In the male genitalia, the new species (gen. figs 1, 2) has, in comparison with M. seima (gen. fig. 3), slightly
longer uncus, somewhat narrower valva with shorter and narrower basal dilation, and considerably longer
vinculum. The aedeagi of the two species have the same basic configuration, but that is somewhat longer in the
new species than in M. seima.
Female unknown.
Meganola suisharyonensis (Strand, 1917)
Roeselia fumosa ab. suisharyonensis Strand, 1917, Archiv für Naturgeschichte 82A(1): 130. Type-locality: Taiwan,
Suisharyo. Two syntypes, in coll. DEI Müncheberg.
Meganola parki Sung-Hwan Oh, 1991
Meganola parki Sung-Hwan Oh, 1991, Systematics of the family Nolidae (Lepidoptera) in Korea: 69, figs 25, 27,
88, 118. Type-locality: Korea (South), Prov. Gyeonggi, Gwangneung. Holotype: male, in coll. Kangwon National
University, Chuncheon.
Meganola flexuosa (Poujade, 1886)
Nola flexuosa Poujade, 1886, Bulletin de la Société Entomologique de France 1886: 27, pl. 18, fig. 26. Type-
locality: [China] [Sechuan] Tibet, Moupin. Holotype: female, in MNHN Paris.
Meganola nanlinga Hu, László, Ronkay & Wang, 2013
Meganola nanlinga Hu, László, Ronkay & Wang, 2013, Zootaxa 3608 (7): 596, figs 5–8. Type-locality: China,
Guangdong, Nanling. Holotype: male, in coll. SCAU.
Meganola bryophilalis bryophilalis (Staudinger, 1887)
Nola bryophilalis Staudinger, 1877, Mémoires sur les Lépidoptéres 3: 181, pl. 10, fig. 5. Type-locality: [Russia]
[Far East] Askold [Island], Raddefka. Syntypes: four males, in coll. ZMHU.
Synonymy
Roeselia basifascia Inoue, 1958, Kontyu 26: 235, figs 2, 7. Type-locality: Japan, Hokkaido, Kushiro, Shibecha.
Holotype: male, in coll. BMNH.
Meganola bryophilalis hondoensis (Inoue, 1970)
Roeselia basifascia ssp. hondoensis Inoue, 1970, Bulletin of the Japan Entomological Academy 6: 6, pl. 1, figs 9–
12. Type-locality: Japan, Tokushima Pref., Tokushima City, Kuramoto. Holotype: male, in coll. BMNH.
Meganola scripta scripta (Moore, 1888)
Roeselia scripta Moore, 1888, Proceedings of the Zoological Society London 1888: 393. Type-locality: [Pakistan]
Kangra. Holotype: male, in coll. BMNH.
Meganola scripta csoevarii László, Ronkay & Witt, 2007
Meganola scripta csoevarii László, Ronkay & Witt, 2007, Entomofauna 28(2): 19, figs 3–4. Type-locality: China,
Prov. Shaanxi, Daba Shan, 15 km S of Shou Man village, 1800 m. Holotype: male, in coll. MWM.
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Meganola implicata László, Ronkay & Witt, 2007
Meganola implicata László, Ronkay & Witt, 2007, Entomofauna 28(2): 19, figs 5–6. Type-locality: North
Thailand, Prov. Mae Hong Son, 1250 m, between Pa Pae and Khun Sa, 98°39¢E, 19°08¢N. Holotype: male, in coll.
MWM.
Meganola fumosa (Butler, 1879)
Nola fumosa Butler, 1879, Entomologist’s Monthly Magazine 14: 9, pl. 43, fig. 2. Type-locality: Japan, Yokohama.
Holotype: male, in coll. BMNH.
Meganola nitidoides Holloway, 2003
Meganola nitidoides Holloway, 2003, The Moths of Borneo 18: 27, pl. 1, figs 31, 32. Type-locality: Borneo,
Sarawak, Gunong Mulu NP, Mulu, 1000 m. Holotype: male, in BMNH.
Meganola simulata László, Ronkay & Witt, 2007
Meganola simulata László, Ronkay & Witt, 2007, Entomofauna 28(14): 163, figs 12–13, 18, and 22. Type-locality:
Indonesia, W Sumatra, 7 km E Panti, 1000 m, 0°12’N, 100°01’E. Holotype: male, in coll. MWM.
Meganola scriptoides Holloway, 2003
Meganola scriptoides Holloway, 2003, The Moths of Borneo 18: 27, pl.1, figs 21, 41. Type-locality: Borneo,
Sarawak, Gunong Mulu NP, Mulu, 1000 m. Holotype: male, in BMNH.
Descriptions of newly established genera separated from Meganola s.l.
Hampsonola gen. n.
(Plate 2, Figs 1–4, 7, 8, Plate 3, Figs 1–8, Plate 4, Figs 1, 2; gen. figs 8–10, 13–24)
Type species: Selca indistincta Hampson, 1894, Fauna of British India, Moths 2: 147. Type-locality: [India] [Nagaland] Naga
Hills. Holotype: male, in coll. BMNH.
Diagnosis. The externally very diverse species of this new genus has been treated so far as Meganola, in spite of
their conspicuously different, apomorphic genitalia structures of both sexes. In the hindwing venation, the veins
M
1
+R
1–5
stalked, and the M
3
+CuA
1
are entirely fused.
The diagnostic features of the male genitalia are the characteristically bilobate valva, the much slenderer (or in
some cases reduced) uncus, the more uniform, arched and apically acute harpe and the simple, short or medium-
long aedeagus with the vesica lacking cornuti; those of the female genitalia are the poorly sclerotized, usually
simple, V-shaped ostium bursae and the much narrower, membraneous ductus bursae (the female genitalia of
Meganola have usually much broader, more strongly sclerotized ostium bursae and conspicuously thicker and
more sclerotized ductus bursae).
The genus Hampsonola displays closer relationship with the South African genus, Vandamia van Son, 1933,
due to the similar configuration of their (otherwise dissimilarly bilobate) valvae, but Hampsonola has much
slenderer and generally longer uncus (with the exception of those taxa where the uncus is reduced) while the uncus
of Vandamia is more robust, thicker and shorter. An additional difference between the two genera that the dorsal
valval lobe of Hampsonola lacks hair tuft, while Vandamia has a rather dense hair tuft in the narrowest part of the
dorsal lobe of valva.
Description. Due to the high diversity of the external morphological features observable within the genus
(including the body size, wing shape, colouration and wing pattern etc.), a general characterization of the external
features of the genus is hardly presented, thus the description is focused on the genitalia characters.
Male genitalia. Uncus variable in length, it may be fully reduced (basinigra species-group) or long to very long
and slim (indistincta species-group); tegumen with variably developed medial process; valva bilobate, medial
incision between dorsal and ventral lobes generally deep, dorsal lobe generally longer than ventral one and apically
may be dilated; ventral lobe rounded or triangular, may bear androconial hair scales at its distal margin (sijthoffi
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and rufa species-groups); dorsal valval lobe with sclerotized costal margin projecting partly to apex; ventral valval
lobe membraneous. Harpe well developed, strongly sclerotized, elongate, most often slightly curved, claw- or
beak-like; sacculus very short, without processi; vinculum relatively short, apically rounded. Aedeagus generally
straight, sometimes S-shaped curved (e.g. in H. rufa, H. longshengensis, H. angustifasciata), relatively short and
thin, without apical carinal process, coecum penis rather variable, apically bilobate; vesica without cornuti.
Female genitalia. Ovipositor short, relatively broad, conical, apophyses relatively long, thin; ostium bursae
relatively broad, simple, V- or U-shaped; ductus bursae narrow in its full length, membraneous, only its short distal
section may be somewhat stronger sclerotized; corpus bursae generally large, ovoid, with a pair of remote,
generally triangular signa.
Species content
Hampsonola indistincta (Hampson, 1894) comb. n.
(gen. figs 8, 9)
Selca indistincta Hampson, 1894, Fauna of British India, Moths 2: 147. Type-locality: [India] [Nagaland] Naga
Hills. Holotype: male, in coll. BMNH.
Hampsonola harutai (László, Ronkay & Ronkay, 2014) comb. n.
Meganola harutai László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 44, pl. 11, fig. 4; gen. fig. 43. Type-
locality: Nepal, Annapurna Himal, Chitre. Holotype: female, in coll. MWM.
Hampsonola diehli sp. n.
(Plate 2, Figs 3, 4; gen. fig. 10)
Holotype. Male, [Indonesia] Sumatra, Kebon Balok, NNW of Medan, 24.XI.1968, leg. Dr. E. Diehl, slide No.: LGN 2057 (coll.
SMNK).
Paratypes. Indonesia, Sumatra: 1 male, Dolok Merangir, Najaradja, 280 m, 8.XII.1969, leg. Dr. E. Diehl (coll. SMNK).
Malaysia: 1 male, Sabah, Gunung Kinabalu, Sayap, 950 m, 116°34’E, 06°10’N, primary forest edge, in front of ranger
house near Sungai Kemantis, 8.III.2001, at light, J.P. Duffels & M.A. Schouten, slide No. MEV 001; 1 male, from the
same area, 1000 m, Gua Malayu trail, clearing in primary forest, 13.III.2001, J.P. & M.J. Duffels (coll. RMNH).
Diagnosis. The new species resembles superficially solely the rather remote species Xenonola limbata (Plate 2,
Figs 5, 6; gen. figs 11, 12) due to the unusual and in Hampsonola unique forewing pattern, the extensive blackish
inner part of the forewing, but the two species differ in several details of the forewing shape and markings. The
most conspicuous differences, besides a number of fine differences in the wing pattern, are the much broader
forewing and more expanded blackish part of the wing reaching the tornus in H. diehli, while the blackish forewing
area is shorter in X. limbata, being terminated at the proximal two-thirds of ventral margin
There are no other similar species known in Meganola or in any Eurasian noline genus. The male genitalia of
the new species display, surprisingly, a closer relationship with H. indistincta (Hampson, 1894) (gen. figs 8, 9),
according to the following distinctive features: H. diehli has conspicuously shorter, apically rounded uncus (that of
H. indistincta much longer, apically pointed), somewhat thicker tegumen without process (H. indistincta has
rounded, finger-like process on tegumen), narrower, apically less dilated dorsal and considerably shorter ventral
lobe of valva; similarly shaped, but medially less bent harpe; much shorter, broadly rounded vinculum (it has
triangular apical process in H. indistincta); the configuration of the aedeagus is practically identical in both species.
Female unknown.
Hampsonola basirufa (de Joannis, 1928) comb. n.
(gen. figs 15, 16)
Celama basirufa de Joannis, 1928, Bulletin de la Société Entomologique de France 97: 249. Type-locality: [N
Vietnam] Tonkin. Syntypes: two females, in coll. MNHN Paris.
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FIGURES 19–21. Genitalia of Hampsonola species.
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PLATE II. Adults of Hampsonola and Xenonola species.
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Hampsonola subbasirufa sp. n.
(Plate 2, Figs 7, 8; gen. figs 13, 14)
Holotype. Male, China, Prov. N. Yunnan, Li-kiang, 7.VIII.1934, leg. H. Höne, slide No.: LGN 1946 (coll. ZFMK).
Paratypes. China, Yunnan. A series of both sexes from the same locality as the holotype, with the following collecting data:
19.VII.1934, 20.VII.1934, 3.VIII.1934, 8.VIII.1934, 9.VIII.1934, 10.VIII.1934, 15.VIII.1934, 27.VIII.1934, 13.VIII.1934,
14.VIII.1935, 17.VIII.1935, 18.VIII.1935, 25.VIII.1935, 28.VIII.1935, 1.IX.1935, 5.IX.1935, 6.IX.1935, 9.IX.1935, slide
Nos: LGN 1947, LGN 2021 (females) (coll. ZFMK); 2 males, from the same site, 29.VII.1935 and 2.VIII.1935 (coll.
SMNK). China, Shaanxi: 1 male, Tapaishan in Tsinling, ca. 3000 m, 10.VIII.1936, leg H. Höne (coll. ZFMK).
Diagnosis. Hampsonola subbasirufa is similar externally to H. basirufa (de Joannis, 1928) (Plate 3, Figs 1, 2), due
to their similar elongate reddish patch covering the basal area of forewing, extending broadly on the costal margin
towards the middle of forewing. The main distinctive external characters are as follows: the new species has
graphite-grey head, collar and tegulae, darker, unicolorously grey forewing ground colour with only traces of
medial and postmedial lines and the darker, pale greyish hindwing while H. basirufa has bright white head and
collar, reddish tegulae, whitish forewing ground colour, sharply defined transverse lines and fine, dense darker
suffusion in the terminal area; in addition, the hindwing is paler, whitish-grey.
In the male genitalia the new species has, in comparison with H. basirufa (gen. figs 13–16), considerably
shorter uncus, shorter apical extension of tegumen, narrower and shorter medial dilatation of tegumen, somewhat
narrower dorsal lobe of valva, somewhat longer, more elongate harpe, slightly longer apical process of vinculum
and somewhat thicker, but similarly simple, short and tubular aedeagus. There are considerable differences in the
female genitalia as the new species has broad V-shaped ostium bursae, considerably smaller, thorn-like signa and
somewhat thicker apophyses compared to those of H. basirufa, which has narrower U-shaped ostium bursae.
Hampsonola longshengensis (Hu, Han, László, Ronkay & Wang, 2014) comb. n.
Meganola longshengensis Hu, Han, László, Ronkay & Wang, 2014, Florida Entomologist 97(3): 1067, figs 9–10.
Type-locality: China, Longsheng, Guangxi. Holotype: male, in coll. SCAU.
Hampsonola angustifasciata sp. n.
(Plate 3, Figs 3, 4; gen. figs 17, 18)
Holotype. Male, China, Prov. N. Yunnan, Li-kiang, 15.VIII.1934, leg. H. Höne, slide No.: LGN 1948 (coll. ZFMK).
Paratypes. China. 1 female, with the same data as the holotype, slide No.: LGN 1949; 1 male, from the same site, 14.VIII.1934
(coll. ZFMK); 1 female, from the same site, 29.III.1935, leg. H. Höne, slide No.: LGN 2160 (coll. SMNK).
Diagnosis. The new species is closely related to H. longshengensis (Hu, Han, László, Ronkay & Wang, 2014), but
is easily distinguishable by the following characters: the new species has very conspicuous, arcuate and tapering
blackish medial stripe on the whitish-grey forewing, while H. longshengensis has only a pale quadrangular costal
patch and the forewing ground colour is somewhat darker, more greyish.
In spite of the largely different external appearance, the two species have very similar configuration of male
genitalia with the following differences: H. angustifasciata has somewhat shorter apical process of tegumen,
considerably narrower, medially less dilated dorsal lobe of valva and slightly longer and thinner harpe compared to
those of H. longshengensis. The almost identical configuration of the curiously S-shaped aedeagus also supports
the close relationship of the two species.
As the female of H. longshengensis is still unknown, the female genitalia of the new species is compared with
H. subbasirufa. H. angustifasciata has considerably thicker and shorter apophyses, much narrower V-shaped
ostium bursae, somewhat longer ductus bursae and conspicuously larger, claw-like signa bursae compared to those
of H. subbasirufa.
Hampsonola stueningi sp. n.
(Plate 3, Figs 5, 6; gen. figs 19, 20)
Holotype. Male, China, Prov. N. Yunnan, Li-kiang, 3.VII.1934, leg. H. Höne, slide No.: LGN 1971 (coll. ZFMK).
Paratypes. China, Yunnan. 2 females from the same site as the holotype, but collected at 16.VIII.1934 and 12.VII. 1934, slide
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Nos: LGN 1972, LGN 1988; 1 male, from the same site, 28.VII.1935, slide No.: Dufay gen prep. 764; 1 female, from the
same site, 19.VIII.1934 (coll. ZFMK).
Diagnosis. The closest relative of H. stueningi is H. angustifasciata, the differences between them are the
following: the new species has pale grey forewing ground colour with rather broad darker grey medial part
bordered by the somewhat sharper defined antemedial and postmedial crosslines, while H. angustifasciata has
much paler whitish-grey forewings with darker, almost blackish, and much narrower, tapering medial stripe.
Comparing the male genitalia of the two sister-species, H. stueningi has somewhat longer uncus,
conspicuously different, shorter and more dilated, apically truncate, more or less triangular dorsal valval lobe (that
is evenly rounded in H. angustifasciata) and longer, narrower, apically pointed ventral valval lobe (being rounded
in the related species), and considerably shorter harpe. The aedeagus of the new species is curved once only in its
basal third, while that of H. angustifasciata is characteristically S-shaped.
The rather similar configuration of the female genitalia of H. stueningi and H. angustifasciata supports the
close relationship of the two species, with the following distinctive features: H stueningi has considerably thinner
and somewhat longer apophyses, funnel-like ostium bursae (that is V-shaped in H. angustifasciata) and somewhat
thinner but similarly claw-like signa bursae.
Hampsonola wilbarka (Hu, Han & Wang, 2013) comb. n.
Meganola wilbarka Hu, Han & Wang, 2013, Zootaxa 3608 (7): 599–600, figs 11–14. Type locality: China,
Longsheng, Guangxi. Holotype: male, in coll. SCAU.
Hampsonola hiranoi (Inoue, 1991) comb. n.
Nola hiranoi Inoue, 1991, Tyo to Ga 42(2): 66, figs 1g, 1h, 7, 8. Type-locality: Japan, Nagano prefecture,
Ookuchizawa, Toyoshina. Holotype: male, in coll. BMNH.
Hampsonola honeyi (László, Ronkay & Witt, 2010) comb. n.
Meganola honeyi László, Ronkay & Witt, 2010, Esperiana 15: 41, pl. 6, fig. 6; gen. fig. 38. Type-locality: North
Thailand, Prov. Chiang Mai, 1800 m, 4 km S of Kop Dong, 99º03’E, 19º52’N. Holotype: male, in coll. G. Ronkay.
Hampsonola tarkabarka (László, Ronkay & Witt, 2010) comb. n.
Meganola tarkabarka László, Ronkay & Witt, 2010, Esperiana 15: 40, pl. 6, fig. 1; gen. fig. 34. Type-locality:
Thailand, Prov. Nan, 25 km N of Bo Luang, 1150 m. Holotype: male, in coll. MWM.
Hampsonola micra sp. n.
(Plate 3, Figs 7, 8; gen. figs 21, 22)
Holotype. Male, Indonesia, Sumatra, Prapat, HW 2, 15–19.V.1985, leg. Dr Diehl, slide No.: LGN 2567 (coll. HNHM).
Paratypes. Indonesia, Sumatra. 1 female, from the same locality as the holotype, but collected at 26.V.1985, slide No.: LGN
2214 (coll. HNHM); 1 female, HW II, 28 km S Pematang Siantar, near Tigadoluk, 1050 m, 02°45’52”N, 099°58’20”E,
10.II.2002, leg. K. Larsen & M. Fibiger, slide No.: LGN 1654 (coll. ZMUC); 1 male, Dolok Merangir, 180 m, 6.X.-
5.XI.1970, leg. Dr. E. Diehl, slide No.: LGN 2157 (coll. SMNK).
Diagnosis. The closest relative of the new species is H. tarkabarka (László, Ronkay & Witt, 2010) (Plate 4, Figs 1,
2), despite the large differences in size (wingspan of H. micra and H. tarkabarka are 13–14 mm and 18–19 mm,
respectively) and wing pattern. Their sister species relationship is indicated by the genitalia characters in both sexes
(gen. figs 21–24), with a number of distinctive features which are as follows: The new species has, in comparison
with H. tarkabarka, somewhat thicker uncus, less dilated tegumen, considerably longer, apically more dilated
dorsal, and basally much broader ventral lobe of valva, much thinner, more curved harpe and considerably longer
vinculum, while the aedeagi and vesicae of the two related species are practically identical.
In the female genitalia, the new species has considerably shorter and thinner apophyses, narrower, ovoid
ostium bursae (that of M. tarkabarka is broad U-shaped) and shorter ductus bursae, and a conspicuous
quadrangular dilatation at cervix bursae (which is absent in M. tarkabarka). In addition, the configuration of the
conspicuous, thorn-like pair of signa is different in the two species: the signa of H. micra are considerably smaller,
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armed with thinner, more stick-like processes, and the distal one is larger than proximal one, while the signa are
larger, broader, more triangular in M. tarkabarka, and the proximal one is bigger than the distal one.
Hampsonola paramediana (Hu, Han, László, Ronkay & Wang, 2014) comb. n.
Meganola paramediana Hu, Han, László, Ronkay & Wang, 2014, Florida Entomologist 97(3): 1064, figs 1–2.
Type-locality: China, Mt. Daming, Guangxi. Holotype: male, in coll. SCAU.
Hampsonola nankunensis (Hu, Han, László, Ronkay & Wang, 2014) comb. n.
Meganola nankunensis Hu, Han, László, Ronkay & Wang, 2014, Florida Entomologist 97(3): 1064–1066, figs 3–
4. Type-locality: China, Mt. Nankun. Holotype: male, in coll. SCAU.
Hampsonola zillii (László, Ronkay & Ronkay, 2014) comb. n.
Meganola zillii László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 45, pl. 12, figs 3–4; gen. figs 46–47.
Type-locality: Vietnam, Bao Loc, Rung Cat Tien. Holotype: female, in coll. MWM.
Hampsonola semmiminta (László, Ronkay & Ronkay, 2014) comb. n.
Meganola semmiminta László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 44, pl. 12, fig. 2; gen. fig. 45.
Type-locality: Vietnam, Mt Fan-si-pan, Sa Pa. Holotype: male, in coll. MWM.
Hampsonola semirufa (Hampson, 1894) comb. n.
Pisara semirufa Hampson, 1894, Fauna of British India, Moths 2: 146. Type-locality: [India] Sikkim. Holotype:
female, in coll. BMNH.
Hampsonola latiscripta (László, Ronkay & Witt, 2005) comb. n.
Meganola latiscripta László, Ronkay & Witt, 2005, Entomofauna 26(11): 219. Type locality: Vietnam, Bach-Ma
NP, 1200 m, 16°10’N, 107°54’E. Holotype: male, in coll. MWM.
Hampsonola galsworthyi (László, Ronkay & Witt, 2010) comb. n.
Meganola galsworthyi László, Ronkay & Witt, 2010, Esperiana 15: 41, pl. 6, fig. 5; gen. fig. 37. Type-locality:
Thailand, Prov. Chiang Mai, 4 km SE of Pang Faen, 1100 m. Holotype: male, in coll. MWM.
Hampsonola hollowayi (László, Ronkay & Witt, 2010) comb. n.
Meganola hollowayi László, Ronkay & Witt, 2010, Esperiana 15: 40, pl. 6, fig. 2; gen. fig. 35. Type-locality: N.
Thailand, Chiang Mai Prov., between Chiang Dao and Kariang, 900 m, 98°48’E, 19°25’N. Holotype: male, in coll.
G. Ronkay.
Hampsonola mediana (László, Ronkay & Witt, 2010) comb. n.
Meganola mediana László, Ronkay & Witt, 2010, Esperiana 15: 40, pl. 6, figs 3–4; gen. fig. 36. Type-locality:
Thailand, Prov. Chiang Mai, Mt. Doi Phahompok 16 km NW of Fang, 2000 m. Holotype: male, in coll. MWM.
Hampsonola gigantoides (Inoue, 1961) comb. n.
Roeselia gigantoides Inoue, 1961, Check List of the Lepidoptera of Japan, (6), 1961
: 682. Type-locality: Japan,
Gumma Pref., Kumanotaira. Holotype: male, in coll. BMNH.
Hampsonola donglashanensis (Hu, Han, László, Ronkay & Wang, 2014) comb. n.
Meganola donglashanensis Hu, Han, László, Ronkay & Wang, 2014, Florida Entomologist 97(3): 1066–1067, figs
7–8. Type-locality: China, Donglashan, Sichuan. Holotype: male, in coll. SCAU.
Hampsonola basinigra (Pellinen, 2012) comb. n.
Nola basinigra Pellinen, 2012, Tinea 22(1): 64–66, figs 2, 7, 8, 10. Type-locality: Thailand, Lampang, Muban
Phichai. Holotype: male, in coll. BMNH.
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PLATE III. Adults of Hampsonola species.
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Hampsonola mimetica (László, Ronkay & Ronkay, 2014) comb. n.
Meganola mimetica László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 45, pl. 12, fig. 5; gen. fig. 48.
Type-locality: Nepal, Annapurna region, Naya Pul at Birethanti. Holotype: male, in coll. MWM.
Hampsonola sijthoffi (van Eecke, 1920) comb. n.
Nola sijthoffi van Eecke, 1920, Zoologische Mededeelingen Leiden 5: 119, fig. 7. Type-locality: Java, Preanger.
Syntypes: three males and two females, in coll. RMNH Leiden.
Synonymy
Nola sijthoffi var. sumatrana van Eecke, 1926, Zoologische Mededeelingen Leiden 9: 48, pl. 2, fig. 18. Type-
locality: [Indonesia] Sumatra, Fort de Kock. Syntypes: three females, in coll. RMNH Leiden.
Roeselia monticola Roepke, 1948, Tijdschrift voor Entomologie 89: 211, pl. 14, fig. 8. Type-locality:
[Indonesia] Sumatra, Mt. Tanggamus. Holotype: female, in coll. RMNH Leiden.
Hampsonola benescripta (László, Ronkay & Witt, 2010) comb. n.
Meganola benescripta László, Ronkay & Witt, 2010, Esperiana 15: 42, pl. 6, fig. 9; gen. fig. 340. Type-locality:
Thailand, Chiang Mai Prov., between Chiang Dao and Kariang, 900 m, 98°48’E, 19°25’N. Holotype: male, in coll.
G. Ronkay.
Hampsonola csorbagabori (László, Ronkay & Ronkay, 2014) comb. n.
Meganola csorbagabori László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 46, pl. 12, figs 7–8; gen. figs
49–50. Type-locality: Cambodia, Prov. Mondolkiri, Seima. Holotype: male, in coll. MWM.
Hampsonola sapatagka (László, Ronkay & Ronkay, 2014) comb. n.
Meganola sapatagka László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 46, pl. 13, figs 1–2; gen. figs
51–52. Type-locality: Cambodia, Prov. Mondolkiri, Seima. Holotype: female, in coll. G. Ronkay.
Hampsonola geoffmartini (László, Ronkay & Witt, 2010) comb. n.
Meganola geoffmartini László, Ronkay & Witt, 2010, Esperiana 15: 42, pl. 6, fig. 9; gen. fig. 40. Type-locality:
North Thailand, Prov. Chiang Mai Prov., between Chiang Dao and Kariang, 900 m, 98°48’E, 19°25’N. Holotype:
male, in coll. G. Ronkay.
Hampsonola andamana (László, Ronkay & Ronkay, 2014) comb. n.
Meganola andamana László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 47, pl. 13, figs 3–4; gen. fig. 53.
Type-locality: India, Andaman Islands, Baratang Island. Holotype: male, in coll. MWM.
Hampsonola rufa (Hampson, 1900) comb. n.
Celama rufa Hampson, 1900, Catalogue of the Lepidoptera Phalaenae in the British Museum 2: 8, pl. 18, fig. 2.
Type-locality: [Sri Lanka] Ceylon, Pundaloya. Holotype: female, in coll. BMNH.
Hampsonola kerala (László, Ronkay & Ronkay, 2014) comb. n.
Meganola kerala László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 47, pl. 12, fig. 6; gen. fig. 54. Type-
locality: India, Kerala State, Munnar. Holotype: male, in coll. MWM.
Wittonola gen. n.
(Plate 4, Figs 3–6; gen. figs 25, 26)
Type-species: Wittonola latifasciata sp. n.
Diagnosis. The external appearance of the type-species of the new genus is rather unique, resembling somewhat
Ctenane labuana (Swinhoe, 1904) (Plate 4, Figs 7, 8; gen. figs 27, 28) due to the presence of the similar, oblique
dark forewing medial stripe and the postmedial line which is replaced by a wavy row of blackish dots. This
similarity is only superficial, the genitalia of both sexes show the closer relationship of Wittonola not with Ctenane
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but with the Meganola lineage, according to the robust, long, evenly tapering, apically pointed uncus, the simple,
relatively narrow, slightly arcuate valva with parallel margins and the harpe arising rather medially on the basal
part of valva, however the very short, button-like harpe is unknown in Meganola. The generic level separation of
Wittonola from Ctenane is supported also by the hindwing venation as in Wittonola the vein M
3
+CuA
1
is entirely
fused while in Ctenane they are variably long stalked.
The configuration of the female genitalia displays further distinctive characters between Wittonola and
Meganola. Wittonola has narrow, elongate papillae anales (this phenomenon is also known in certain species-
groups of Nola) with extraordinarily long apophyses; these characters are unknown in Meganola. The single,
thorn-like signum of Wittonola resembles rather to a signum of the Manoba species than to any other Meganola
taxa.
The genus Wittonola shares certain genitalia characters with Meganola and Manoba (Plate 5, figs 1–2 and gen.
figs 29–30) but, due to the above combination of the external and genital characters, it cannot be unified with any
of them. Thus, this lineage is considered here as a distinct genus.
Description. The only known species of the genus is rather small in size with the wingspan 11–15 mm, length
of forewing 5–7 mm. Head and eyes relatively small; antennae bipectinate in male, filiform in female; frons,
vertex, tegulae and thorax covered with pale brown scales, collar dark brown. Forewing elongate, apically rounded;
ground colour pale brown with reddish sheen; costal margin dark brown from the base to the middle; median area
with conspicuous, thick, oblique dark brown stripe; postmedial and terminal areas dark brown; all but one
crosslines deleted, postmedial line present but replaced by a row of small dark brown dots. Hindwing pale brown,
without discal spot and transverse lines.
Male genitalia. Uncus rather long, robust, evenly tapering, apically pointed, tegumen short, relatively broad,
valva simple, relatively narrow, slightly arcuate, margins parallel, costal sclerotization rather narrow and weak.
Harpe very short, button-like, arising in the middle of basal part of valva, vinculum well developed, pointed, V-
shaped. Aedeagus simple, tubular, relatively short and thin, without apical carinal process; vesica with small,
finger-shaped, finely scobinated section.
Female genitalia. Ovipositor relatively long, very narrow, apically expanded, papillae anales very long, thin,
eighth tergite rather short, ostium bursae relatively broad, cup-shaped, ductus bursae medium-long, distal half
strongly sclerotized, proximal half membraneous, corpus bursae elongate, distal half gradually broadening,
proximal one ovoidal; signum bursae single, relatively large, acute, thorn-like.
Species content
Wittonola latifasciata sp. n.
Wittonola latifasciata sp. n.
(Plate 4, Figs 3–6; gen. figs 25, 26)
Holotype. Female, Cambodia, Mondolkiri Prov., Seima Biodiversity Conservation Area, between Seima and O’Rang,
12º12’12”N, 107º01’09”E, 300 m, 30.I.2006, leg. G. Csorba & G. Ronkay, slide No.: LGN 1734 (W 22330) (coll. MWM).
Paratypes. Cambodia: 46 specimens of both sexes, with same data as the holotype, slide Nos: LGN 1731 (W 22327), 1733 (W
22329) (males), LGN 1732 (W 22328) (female); 23 specimens of both sexes, Seima Biodiversity Conservation area,
between Seima and O'Rang, 360 m, 12º15’44”'N, 107º03’49”'E, 27–29.I.2006, leg. G. Ronkay (coll. MWM); 1 male, with
the same data (coll. HNHM). Indonesia, Sumatra: 2 females, Rimbo Panti, 250 m, 00º20’50”N, 100º04’07”E, 24–
25.II.2002, leg. K. Larsen & M. Fibiger, slide Nos: LGN 1638, LGN 1639 (coll. ZMUC).
Diagnosis. The diagnosis of W. latifasciata is given in detail in the diagnosis of the genus Wittonola.
Nanola László, Ronkay & Witt, 2010
(Plate 1, Figs 5–8, Plate 5, Figs 3–7; gen. figs 4–7, 31–34)
Nanola László, Ronkay & Witt, 2010, Esperiana 15: 38. Type-species: Nanola hluchyi László, Ronkay & Witt, 2010, by
original designation.
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PLATE IV. Adults of Hampsonola, Wittonola and Ctenane species.
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Taxonomic notes. When the genus was described in the monographic work on the Nolinae fauna of Thailand
(László et al. 2010), only the type species was considered as member of the genus, due to the restricted area of that
study. The detailed analysis of the genital morphology of Meganola s.l. revealed the existence of a distinct phyletic
lineage being associated with Meganola by the former authors. According to the recent investigations, the
taxonomic content of Nanola has been extended remarkably, comprising more than a dozen of species. The
hindwing venation of the Nanola species is “quadrifine”, with the veins M
3
+CuA
1
are stalked.
Diagnosis. The main generic characters of this lineage are the relatively short, apically pointed uncus, the
generally broadly opened, simple, rather narrow and apically rounded valva, the origin of harpe from the ventral
margin of the valval plate as a direct continuation of sacculus, unlike in Meganola, where the harpe is arising more
medially from the valval plate (this feature displays closer connections between Nanola and Manoba Walker,
1864), and the usually short, relatively thick, simple aedeagus with or without cornuti in the vesica.
The configuration of the female genitalia, at least in those taxa where the female is known, is also rather
curious. The main diagnostic features are the extraordinarily thick, strongly sclerotized ductus bursae and the
unusal (and very variable) shape of the signum (it is a large quadrangular plate in N. liaoningensis, a long, narrow,
evenly arcuate transverse sclerotization in N. franzdanieli, horseshoe-shaped in N. patkosigna and a pair of
scobinated rounded plates in N. lesarbena); or even fully reduced like in N. hluchyi). In spite of the rather large
external diversity, all species are characterised by the above mentioned combination of genital characters,
indicating their close relationship.
Species content
Nanola hluchyi László, Ronkay & Witt, 2010
(Plate 5, Figs 3, 4; gen. figs 31, 32)
Nanola hluchyi László, Ronkay & Witt, 2010, Esperiana 15: 38, pl. 5, figs 16–17; gen. fig. 33. Type-locality:
Thailand, Prov. Nan, 5 km N of Bo Luang, 1000 m. Holotype: male, in coll. MWM.
Nanola rothschildi sp. n.
(Plate 5, Figs 5–7; gen. figs 33, 34)
Holotype. Female, [Philippines] Luzon, Palali, 2000 m, Benguet, 27.XII.1912, leg. A.E. Wileman, slide No.: LGN 2013 (coll.
BMNH).
Paratypes. Philippines: 1 male, Tawi Tawi, Lapid Lapid at Manalik Channel, 19.XI.1961, Noona Dan Exp. 61–62, Caught by
Mercury-light 19.00–02.00, slide No.: LGN 2133 (coll. ZMUC). Indonesia, Sumatra: 1 male, Dolok Merangir,
12.III.1981, leg. Dr. Diehl, slide No.: LGN 2175 (coll. HNHM).
Diagnosis. The closest relative of N. rothschildi is N. hluchyi. The two species display easily recognisable external
and genital differences which are as follows: The new species has pale grey head, collar and tegulae, pale brownish
forewing ground colour with elongate quadrangular whitish basal patch and narrow, elongate whitish apical patch,
continuing in a relatively broad, oblique, whitish line bordered proximally with the dark brownish, diffuse, rather
wavy postmedial line towards the middle of the ventral margin of forewing. In comparison, N. hluchyi has whitish
head and thorax, much darker brown forewing ground colour with larger, more rounded whitish basal and apical
patches, lacking the oblique white line in the postmedial area.
Comparing the male genitalia of the two species (gen. figs 31–34), N. rothschildi has considerably longer
uncus with slightly shorter acute apical process, somewhat shorter medial dilatation of tegumen, somewhat
narrower apical part of valva, thicker, more robust, more dentate harpe, and more broad-based, short, triangular
cornutus of vesica.
In the female genitalia, the new species has narrower, less sclerotized basin-shaped ostium bursae (that of N.
hluchyi is much broader, strongly sclerotized and wrinkled, funnel-like), broader, only partially sclerotized,
proximally more swollen ductus bursae, in addition, N. rothschildi has, well-developed, finger-like signum bursae,
while the related species lacks the signum.
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FIGURES 22–24. Genitalia of Hampsonola species.
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PLATE V. Adults of Manoba, Nanola and Fragilonola species.
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Nanola laevis (László, Ronkay & Ronkay, 2014) comb. n.
Meganola laevis László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 34, pl. 7, fig. 5; gen. fig. 10. Type-
locality: Vietnam, Fan-si-pan Mts, Sa Pa. Holotype: male, in coll. MWM.
Nanola franzdanieli (László, Ronkay & Ronkay, 2014) comb. n.
Meganola franzdanieli László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 35, pl. 8, figs 1–2; gen. figs
13–14. Type-locality: [Vietnam], Tonking. Holotype: female, in coll. MWM.
Nanola liaoningensis (Han & Li, 2008) comb. n.
Meganola liaoningensis Han & Li, 2008, Tinea 20 (3): 153–154, figs 1–3. Type-locality: China, Prov. Liaoning,
Jianchang, Mt. Bailang. Holotype: male, in coll. NEFU.
Synonymy
Meganola pekarskyi László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 36–37, pl. 8, figs 5–6; gen. figs
17–18. Type-locality: China, Prov. Kiangsu, Nanking, Lungtan. Holotype: male, in coll. ZFMK; syn. n.
Nanola lesarbena (László, Ronkay & Ronkay, 2014) comb. n.
Meganola lesarbena László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 37, pl. 9, fig. 1; gen. fig. 19.
Type-locality: Lower Burma. Holotype: female, in coll. MWM.
Nanola patkosigna (László, Ronkay & Ronkay, 2014) comb. n.
Meganola patkosigna László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 34–35, pl. 7, figs 7–8; gen. figs
11–12. Type-locality: [Indonesia, Sulawesi], Lindoe Paloe. Holotype: male, in coll. BMNH.
Nanola amboni (László, Ronkay & Ronkay, 2014) comb. n.
Meganola amboni László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 35–36, pl. 8, fig. 3; gen. fig. 15.
Type-locality: Indonesia, Ambon, Air Koluar. Holotype: male, in coll. MWM.
Nanola moluccana (László, Ronkay & Ronkay, 2014) comb. n.
Meganola moluccana László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 36, pl. 8, fig. 4; gen. fig. 16.
Type-locality: Moluccas, Halmahera, Talagaranu Mt. Holotype: male, in coll. MWM.
Nanola wangi (Hu, Han, László, Ronkay & Wang, 2014) comb. n.
Meganola wangi Hu, Han, László, Ronkay & Wang, 2014, Florida Entomologist 97(3): 1066, figs 5–6. Type-
locality: China, Jianfengling, Hainan. Holotype: male, in coll. SCAU.
Nanola basalactifera (Holloway, 2003) comb. n.
(Plate 1, Figs 5, 6; gen. figs 4, 5)
Meganola basalactifera Holloway, 2003, The Moths of Borneo 18: 29, pl. 1, figs 34, 35. Type-locality: Borneo,
Sarawak, Gunong Mulu NP, Mulu, 160 m. Holotype: male, in coll. BMNH.
Nanola subbasalactifera
(László, Ronkay & Ronkay, 2014) comb. n.
(Plate 1, Figs 7, 8; gen. figs 6, 7)
Meganola subbasalactifera László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 33, pl. 6, figs 7–8; gen.
figs 6–7. Type-locality: Philippines, Palawan, Mt. Salokot. Holotype: male, in coll. MWM.
Nanola promelaena (Hampson, 1914) comb. n.
Nola promelaena Hampson, 1914, Catalogue of the Amatidae and Arctiidae (Nolinae and Lithosiinae) in the
Collection of the British Museum 1914: 417, pl. 24, fig. 7. Type-locality: [Taiwan] Formosa, Kanshirei. Holotype:
male, in coll. BMNH.
Nanola mirabilis (László, Ronkay & Ronkay, 2014) comb. n.
Meganola mirabilis László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 34, pl. 7, figs 3–4; gen. fig. 9.
Type-locality: [Indonesia], Java, Nongkodjadjar. Holotype: male, in coll. BMNH.
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FIGURES 25–27. Genitalia of Wittonola and Ctenane species.
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FIGURES 28–30. Genitalia of Ctenane and Manoba species.
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FIGURES 31–33. Genitalia of Nanola species.
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Nanola submelaena (László, Ronkay & Ronkay, 2014) comb. n.
Meganola submelaena László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 33, pl. 7, figs 1–2; gen. fig. 8.
Type-locality: [India] Khasis. Holotype: male, in coll. BMNH.
Nanola klondykei (László, Ronkay & Ronkay, 2014) comb. n.
Meganola klondykei László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 32–33, pl. 6, fig. 5; gen. fig. 5.
Type-locality: Philippines, Luzon, Prov. Benguet. Holotype: male, in coll. BMNH.
Fragilonola gen. n.
(Plate 5, Fig. 8, Plate 6, Figs 1–6; gen. figs 35–40)
Type species:‘Meganola’ igorkostjuki László, Ronkay & Witt, 2010, Esperiana 15: 51, pl. 7, fig. 7; gen. fig. 45. Type-locality:
North Thailand, Prov. Chiang Mai, 1100 m, 4 km SE of Pang Faen. Holotype: male in coll. MWM.
Diagnosis. The external appearance of the members of the new genus is rather conspicuous; all known taxa are
colorful, tiny species with the wingspan 12–16 mm. The forewings of Fragilonola have broad orange-brown or
pale brownish basal field, red-brown or greyish-brown terminal area and dark costal margin, the postmedial line is
conspicuous, almost straight, oblique, broad, and whitish. Fragilonola species are easily recognizeable due to this
combination of characters, while the configuration of their genitalia shows close relationship with Nanola, sharing
the characteristic origination of the harpe at the ventral margin of valva in both genera.
The main distinctive character between Fragilonola and Nanola is the considerably longer and narrower uncus
and the fine but conspicuous, needle-like apical carinal process of the aedeagus of the former genus, while Nanola
has generally shorter, more robust uncus and the carina of aedeagus lacks the apical process. In the female
genitalia, the differences are even larger as Fragilonola has the longer, thinner, less sclerotized ductus bursae and
double signum, while the species of Nanola have rather thick, swollen, strongly sclerotized ductus bursae and a
single signum of very variable shape.
Comparing the hindwing venation of the two genera, the hindwing of Fragilonola is “trifine” with the vein
M
3
+CuA
1
being entirely fused, while in Nanola it is “quadrifine” as the vein M
3
+CuA
1
is variably long stalked.
Description. Wingspan 12–16 mm, length of forewing: 6–8 mm. Head relatively small; eyes relatively large;
male antenna bipectinate with ciliae gradually shortening towards tip, female antenna filiform. Forewing relatively
narrow, apically rounded, ground colour brownish-grey, basal and terminal area lighter grey or orange-brown,
costal margin dark brown. Transverse lines deleted, except postmedial line which is rather broad, oblique, more or
less straight, and whitish. Hindwing pale grey, without discal spot and transverse lines.
Male genitalia. Uncus relatively long, thin, slightly arcuate, apically pointed. Tegumen rather broad, medially
slightly dilated. Harpe bifurcate, erected at the ventral margin of valva. Vinculum long and pointed in the type
species, relatively short, rounded in F. fragilis. Aedeagus tubular, medium-long, relatively thin, apical carinal
process, if present, straight, thorn-like apical; vesica without cornuti but with fine scobination.
Female genitalia. Ovipositor short, conical, apophyses medium-long, relatively thin, eighth tergite very short,
ostium bursae narrow, cup-shaped, ductus bursae long or relatively short, corpus bursae large, ovoid, signum
bursae double, represented by a pair of rather remote, rounded, scobinated plates or short triangular processes.
Species content
Fragilonola igorkostjuki (László, Ronkay & Witt, 2010) comb. n.
(Plate 6, Figs 1–3; gen. figs 35, 36)
‘Meganola’ igorkostjuki László, Ronkay & Witt, 2010, Esperiana 15: 51, pl. 7, fig. 7; gen. fig. 45. Type-locality: North
Thailand, Prov. Chiang Mai, 1100 m, 4 km SE of Pang Faen. Holotype: male, in coll. MWM.
Additional material examined: 1 male, Indonesia, NE Sumatra, Dolok Merangir, 280 m, 10.I.1970, leg. Dr. E. Diehl, slide No.:
LGN 2047; 1 female, from the same locality, collected in 1977, slide No.: LGN 2035 (coll. SMNK).
Description of the female genitalia (gen. fig. 36). Ovipositor short, conical, apophyses medium-long, relatively
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thin, eighth tergite very short. Ostium bursae narrow, cup-shaped; ductus bursae rather long, distal section strongly
sclerotized, posteriorly straight and smooth, anteriorly curved and wrinkled, membraneous proximal part dilated,
with rounded lateral protuberation; corpus bursae large, ovoid; signum bursae double, represented by a pair of
rather remote rounded scobinated plates, distal one much larger than proximal one.
Remarks. The species was described from Northern Thailand and it was known to occur only in its type
locality. This is the first record of the species from Sumatra.
Fragilonola fragilis sp. n.
(Plate 6, Figs 4–6; gen. figs 37–39)
Holotype. Male, [Indonesia] Sumatra, Sitahoan, 21–22.XI.1981, leg. Dr. Diehl, slide No.: LGN 2154 (coll. HNHM).
Paratypes. Indonesia, Sumatra: 1 female, Prapat, HW 3, 31.I.1984, leg. Dr. Diehl, slide No.: LGN 2156 (coll. HNHM); 2 males,
Dolok Merangir, 180 m, collected in 1971, and 5–25.XI.1973, leg. Dr. E. Diehl, slide Nos LGN 2076, LGN 2153 (coll.
SMNK).
Diagnosis. The new species is similar to F. igorkostjuki but is easily distinguished by the following characters: F.
fragilis has considerably longer ciliation of male antenna, somewhat shorter forewing, much darker colouration of
forewing with less light areas, and conspicuously broader whitish postmedial line.
In the male genitalia the new species has considerably thicker uncus, longer tegumen, broader valva, larger,
more robust bifurcate harpe with more unequal processes and conspicuously shorter vinculum than in F.
igorkostjuki. The aedeagus of F. fragilis lacks the apical carinal process, while the carina of F. igorkostjuki
possesses relatively short but well developed thorn-like apical process. The vesica is with fine scobination in both
species.
The female genitalia of the new species has somewhat shorter apophyses, much broader, more strongly
sclerotized ostium bursae, considerably shorter and thinner ductus bursae being sclerotized in its full length (that of
F. igorkostjuki is sclerotized only in its distal section), somewhat larger corpus bursae, and the signa are a pair of
remote, more or less equally small, short triangular processes, while the signa of F. igorkostjuki are inequal,
relatively small, rounded scobinated plates.
Fragilonola parentela sp. n.
(Plate 5, Fig. 8; gen. fig. 40)
Holotype. Male, Philippines, N. Palawan, S. Vicente, 20 km NEE Roxas, 10º21’N, 119º10’E, 600 m, 12.–17.I.1988, leg. Cerny
& Schintlmeister, slide No.: LGN 2155 (coll. HNHM).
Diagnosis. Fragilonola parentela appears as an allopatric sister species of F. igorkostjuki occurring in the
Philippines. The two sister species are externally almost identical, the distinctive characters are expressed in the
configuration of the male genitalia. F. parentela has, in comparison with F. igorkostjuki, somewhat shorter and
thicker uncus, considerably shorter and broader valva, and slightly larger bifurcate harpe with apically rounded
arms (these arms are apically pointed in F. igorkostjuki). Last but not least, the aedeagus of the new species is
somewhat shorter than that of F. igorkostjuki.
Maculonola gen. n.
(Plate 6, Figs 7, 8; gen. figs 41, 42)
Type species: Meganola apiensis Holloway, 2003, The Moths of Borneo 18: 30, pl. 1, fig. 20. Type-locality: Borneo, Sarawak,
Gunong Mulu NP, Api, 900 m. Holotype: male, in coll. BMNH.
Diagnosis. The habitus of the type-species of the new genus is unique within the whole tribe. The pale greyish-
brown forewing lacks all but one crosslines except the oblique, straight, pale, but well visible brownish postmedial
line. The other markings of the wing are conspicuous blackish patches on the basal and medial part of the costa, an
oblique row of three dots in the terminal area and a larger patch at the middle of the ventral margin. The hindwing
venation is typically „quadrifine”, with the M
3
+CuA
1
long stalked.
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FIGURES 34–36. Genitalia of Nanola and Fragilonola species.
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FIGURES 37–39. Genitalia of Fragilonola species.
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FIGURES 40–42. Male genitalia of Fragilonola and Maculonola species.
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PLATE VI. Adults of Fragilonola and Maculonola species.
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The basic configuration of the male genitalia indicates the closer relationship of Maculonola rather with
Nanola than Meganola according to the rather narrow-elongate valval shape and the configuration of harpe. The
main autapomorphic character of the new genus is the presence of a rather curious, long, slim, strongly arcuate
process of valva arising basally from the sclerotized costal margin, covered densely with short, fine, hair-like
cornuti. This character is unknown in other species of the tribe, being eligible to delineate a distinct genus.
Description. Wingspan 19–20 mm, length of forewing 9–10 mm. Head and eyes relatively large, frons, vertex,
collar and tegulae covered with pale greyish-brown hair-scales. Forewing elongate, apically rounded, ground
colour pale greyish-brown, all but one transverse lines deleted, postmedial line present, as an oblique, straight, pale
but well visible, narrow brownish stripe. Forewing costal margin with conspicuous blackish patches at the basal
and medial sections and oblique row of three relatively large blackish dots in the terminal area; the mid-section of
the ventral margin is marked by a larger patch. Hindwing shining greyish brown, somewhat paler than forewing,
discal spot and transverse lines absent.
Male genitalia. Uncus short and broad, with relatively long, acute, fine medial process apically. Tegumen
narrow, without medial dilatation; valvae long, rather narrow, apically barely dilated, rounded. Harpe arising at the
ventral margin of valva, short ribbon-like with parallel margins, slightly arcuate, apically either with short, pointed
process arising at the distal margin, or fully truncate. Valva is armed with long, slim, caudally curved process
arising basally at the costal margin, covered densely with short, fine hair-like cornuti; vinculum relatively short,
rounded. Aedeagus tubular, relatively short and narrow, without apical carinal process, coecum penis
conspicuously short; vesica without cornuti.
Female unknown.
Species content
Maculonola apiensis (Holloway, 2003) comb. n.
(Plate 6, Fig. 7; gen. fig. 41)
Meganola apiensis Holloway, 2003, The Moths of Borneo 18: 30, pl. 1, fig. 20. Type-locality: Borneo, Sarawak, Gunong Mulu
NP, Api, 900 m. Holotype: male, in coll. BMNH.
Maculonola dolokmerangirensis sp. n.
(Plate 6, Fig. 8; gen. fig. 42)
Holotype. Male, [Indonesia] Sumatra, Dolok Merangir, 3.IX.1980, leg. Dr. Diehl, slide No.: LGN 2023 (coll. HNHM).
Diagnosis. The new species is a twin species of M. apiensis with easily recognisable external and genital features
which are as follows: The forewing ground colour of M. dolokmerangiri is somewhat paler with more brownish
shine (M. apiensis is more greyish); the forewing patches of the new species are more blurred, except the large one
at the ventral margin, which is larger and more sharply defined than in M. apiensis.
In the male genitalia, the new species (gen. fig. 42) has, compared with M. apiensis (gen. fig. 41), slighly
narrower valva, fully truncate harpe (that of M. apiensis has short, pointed apical process arising at distal margin)
and much longer basal costal process. The aedeagus of the two species are rather similar, but the short coecum
penis is quadrangular in the new species, and rounded in M. apiensis.
Female unknown.
Acknowledgements
We would like to express our sincere thanks to the following colleagues provided extensive help during our studies:
Mr Thomas J. Witt (MWM, Munich), Mr Martin Honey and Mr Geoff Martin (BMNH, London), Dr Erik van
Nieukerken and Mr Rob de Vos (RMNH, Leiden), Dr Ole Karsholt (ZMUC, Copenhagen), Dr Dieter Stüning
(ZFMK, Bonn), Dr Robert Trusch (SMNK, Karlsruhe), Dr Martin Lödl and Dr Sabine Gaal-Haszler (NHM,
Vienna).
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This research received support from the SyntheSys Project http://www.synthesys.info/ which is financed by
European Community Research Infrastructure Action under FP6 „Structuring the European Research Area”
Programme, Grant Nos GB-TAF-2644 (G. Ronkay), GB-TAF-5432 (Gy.M. László) and NL-TAF-4023 (Gy.M.
László).
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