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Paranotocoris Ahmad & Shadab, 1973, a new synonym of Phyllomorpha Laporte, 1833 (Hemiptera: Heteroptera: Coreidae)

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The tribe Phyllomorphini Mulsant & Rey, 1870 of the family Coreidae, subfamily Coreinae, is represented in the Palaearctic Region by three genera (Pephricus Amyot & Serville, 1843, Phyllomorpha Laporte, 1833, and Tongorma Kirkaldy, 1900) and four species. Two of them, Pephricus subtilis Montandon, 1913 and Tongorma latreillii (Guérin-Méneville, 1839) are known only from Yemen and represent Afrotropical faunal elements in the Arabian Peninsula. The only truly Palaearctic genus is therefore Phyllomorpha, comprising two species—P. lacerata Herrich-Schäffer, 1835 and P. laciniata (Villers, 1789)—distributed in southern Europe, North Africa, and western Asia from Turkey to Oman, Central Asia, Afghanistan, and Pakistan (Dolling 2006). Ahmad & Shadab (1974) described another monotypic genus from the Pakistani province of Baluchistan, Pronotocoris Ahmad & Shadab, 1974 (type species Paranotocoris echinus Ahmad & Shadab, 1974). Recent study of the original descriptions of both of the taxa drove us to the conclusion that Pronotocoris is a junior subjective synonym of Phyllomorpha, and Pronotocoris echinus a junior subjective synonym of Phyllomorpha lacerata.
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Accepted by C. Schaefer: 15 Apr. 2013; published: 17 May 2013 397
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Paranotocoris Ahmad & Shadab, 1973, a new synonym of Phyllomorpha Laporte,
1833 (Hemiptera: Heteroptera: Coreidae)
PETR KMENT1) & DÁVID RÉDEI2)
1) Department of Entomology, National Museum, Kunratice 1, CZ-14800 Praha 4, Czech Republic. E-mail: sigara@post.cz
2) Institute of Entomology, Nankai University, 300071 Tianjin, Weijin Road 94, China. E-mail: david.redei@gmail.com
The tribe Phyllomorphini Mulsant & Rey, 1870 of the family Coreidae, subfamily Coreinae, is represented in the
Palaearctic Region by three genera (Pephricus Amyot & Serville, 1843, Phyllomorpha Laporte, 1833, and
Tongorma Kirkaldy, 1900) and four species. Two of them, Pephricus subtilis Montandon, 1913 and Tongorma
latreillii (Guérin-Méneville, 1839) are known only from Yemen and represent Afrotropical faunal elements in the
Arabian Peninsula. The only truly Palaearctic genus is therefore Phyllomorpha, comprising two species—P.
lacerata Herrich-Schäffer, 1835 and P. laciniata (Villers, 1789)—distributed in southern Europe, North Africa, and
western Asia from Turkey to Oman, Central Asia, Afghanistan, and Pakistan (Dolling 2006). Ahmad & Shadab
(1974) described another monotypic genus from the Pakistani province of Baluchistan, Pronotocoris Ahmad &
Shadab, 1974 (type species Paranotocoris echinus Ahmad & Shadab, 1974). Recent study of the original
descriptions of both of the taxa drove us to the conclusion that Pronotocoris is a junior subjective synonym of
Phyllomorpha, and Pronotocoris echinus a junior subjective synonym of Phyllomorpha lacerata.
Phyllomorpha Laporte, 1833
Phyllomorpha Laporte, 1833: 47. Selected as correct original spelling and placed on the Official List of Generic Names in
Zoology (ICZN 2008: Opinion 2198). Type species: Coreus hystrix Latreille, 1817 (= Phyllomorpha laciniata (Villers,
1789)) by subsequent designation (Reuter 1888: 414).
Phyllomorphus: Laporte (1833: 41). Incorrect original spelling, placed on the Official Index of Rejected and Invalid Generic
Names (ICZN 2008: Opinion 2198).
Craspedum Rambur, 1839: 138. Unnecessary new name for Phyllomorpha Laporte, 1833.
Phyllophya Stål, 1873: 80. Synonymized by Horváth (1889: 326). Type species: Phyllomorpha persica Westwood, 1841 (=
Phyllomorpha lacerata Herrich-Schäffer, 1835) by monotypy.
Paranotocoris Ahmad & Shadab, 1974: 181, syn. nov. Hereby fixed as correct original spelling under Article 24.2.3 of the
International Code of Zoological Nomenclature (ICZN 1999). Type species: Paranotocoris echinus Ahmad & Shadab,
1974, by original designation.
Paranotocornis: Ahmad & Shadab (1974: 181). Incorrect original spelling.
Paranotocoris: Ahmad et al. (1977: 6, 24) (key to genera, catalogue); Ahmad (1979: 7) (key to tribes and genera); Ahmad
(1980: 122, 131) (checklist).
Taxonomy. Ahmad & Shadab (1974) described the new genus, provided illustrations of habitus, metathoracic
scent gland peritreme, hemelytron, hindwing, abdominal trichobothrial pattern, and male genitalia (pygophore,
paramere, and phallus), and compared it to other related genera. According to them Paranotocoris ‘... can be easily
distinguished from all other genera of the tribe Phyllomorphini by the following characters: body with large and
small spines all over, except on third and fourth antennal segments and distal portion of tibiae and tarsi; basal
antennal segment very long, longer than third and fourth segments combined; and labium reaching beyond the
anterior margin of fourth abdominal venter.’ However, characters of the antennae (antennal segment I long,
antennal segments II and III unarmed) and legs (tibiae unarmed except of two spines frequently present at their
extreme base), considered as of generic significance by Ahmad & Shadab (1974), perfectly fit Phyllomorpha
lacerata, and they are universally considered in the literature as diagnostic for Ph. lacerata at the species level,
KMENT & RÉDEI
398 · Zootaxa 3652 (3) © 2013 Magnolia Press
differentiating it from Ph. laciniata (e.g., Stichel 1959, Gidayatov 1982, Moulet 1995). The apex of labium extends
from base of abdominal sternite III to middle of abdominal sternite IV in various specimens of Ph. lacerata
examined by us, therefore P. echinus falls within the variability range of Ph. lacerata, and otherwise this character
obviously does not have generic importance. Comparison of the description and illustrations of P. echinus with
specimens of Ph. lacerata deposited in the collections of the National Museum, Praha, and Hungarian Natural
History Museum, Budapest left no doubt that the two are conspecific. For this reason, we consider Paranotocoris
Ahmad & Shadab, 1974, syn. nov., a junior subjective synonym of Phyllomorpha Laporte, 1833.
FIGURE 1. Habitus of Phyllomorpha lacerata Herrich-Schäffer, 1835. Specimen from Syria, Bludan, 6.v.2008, V. Skoupý lgt.,
P. Kment det., coll. P. Horsák (Photo: Z. Chalupa).
Nomenclature. The genus name is spelled Paranotocoris throughout the paper (Ahmad & Shadab 1974), but
an alternative spelling Paranotocornis occurs once on page 181, apparently as a result of inadvertent error.
Therefore Paranotocoris is hereby fixed as the correct original spelling under Article 24.2.3 of the International
Code of Zoological Nomenclature (ICZN 1999).
Note. Burmeister (1835: 310), acting as First Reviser, selected Phyllomorphus as the correct original spelling.
Westwood (1840: 484) validly designated Cimex paradoxus Sparrmann, 1777 (= Pephricus paradoxus
Zootaxa 3652 (3) © 2013 Magnolia Press · 399
PARANOTOCORIS, SYNONYM OF PHYLLOMORPHA
(Sparrmann, 1777)) as the type species of Phyllomorphus (Dolling 2006, Kerzhner 2007). These acts were set aside
by Opinion 2198 (ICZN 2008).
Phyllomorpha lacerata Herrich-Schäffer, 1835
Phyllomorpha laceratus Herrich-Schäffer, 1835: 41, 73.
Phyllomorpha persica Westwood, 1841: 8. Synonymized by Horváth (1889: 326).
Paranotocoris echinus Ahmad & Shadab, 1974: 181–186, figs 1–8, syn. nov. Holotype: male, Pakistan, Baluchistan, Mekran, 8
mile to Turbat on Kalat and Turbat Highway, 24.iv.1969, I. Ahmad lgt. (coll. Natural History Museum, Department of
Zoology, University of Karachi).
Paranotocoris echinus: Ahmad et al. (1977: 14: figs 4, 24, 37) (catalogue, habitus figure); Ahmad (1979: 42) (checklist);
Ahmad (1980: 131) (checklist).
Taxonomy. As it was discussed above, the original description and illustrations of Paranotocoris echinus fit
perfectly the characters of Ph. lacerata. In spite of occurrence of Ph. lacerata in the neighbouring regions of
Afghanistan and Iran (including the Iranian province of Sistan and Baluchistan) (see Wagner 1961, 1968;
Kiritshenko 1963; Brown 1966; Hoberlandt 1989; Hoberlandt & Švihla 1990; Linnavuori 2007, 2012; Ghahari et
al. 2012), the latter species was omitted by the authors when diagnosing P. ech inu s. We propose the following new
subjective synonymy: Paranotocoris echinus Ahmad & Shadab, 1974, syn. nov. = Phyllomorpha lacerata Herrich-
Schäffer, 1835.
Distribution. Southern Europe (Albania, Greece, Italy, ?Yugoslavia), North Africa (Algeria), Near East (Asian
Turkey, Iraq, Israel, Jordan, Lebanon, Syria), Transcaucasia (Azerbaijan, Armenia), Central Asia (Asian
Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan, Uzbekistan), Iran, Afghanistan, and Pakistan (Dolling 2006).
There seems to be no reliable record published from the former Yugoslavia (see Proti 2001).
Acknowledgements
We are greatful to Mick D. Webb and William R. Dolling (Natural History Museum, London, United Kingdom) for
sharing part of their manuscript of the Coreoidea World Catalogue. We are also indebted to Zden k Chalupa and
Petr Horsák (both Kladno, Czech Republic) for the permission to include their photo of Ph. lacerata. This study
was partly supported by the grant of the Ministry of Culture of the Czech Republic (DKRVO MK-S 760/2013/12
OVV) to P. Kment and the China Postdoctoral Science Foundation (20110490769) to D. Rédei.
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... Vertreter der Tribus Phyllomorphini, die zu den Coreinae innerhalb der Coreidae zählt, kommen in Afrika, Asien und im südlichen Europa vor (Kment & Rédei 2013). Tongorma ist in Afrika und Asien vertreten. ...
... Tongorma latreillii (Guérin- Méneville, 1839) wurde in Namibia (Stål, 1858), im Senegal (Blöte 1935, Dolling 2006) sowie in Südafrika nachgewiesen und gilt als südlich der Sahara vorkommendes afrotropisches Faunenelement . Allerdings kommt T. latreillii rezent auch im Jemen vor (Dolling 2006, Kment & Rédei 2013), wo die Art ein afrotropisches Faunenelement der Arabischen Halbinsel darstellt (Kment & Rédei 2013). Die bei Blöte (1935) zudem zu findende Angabe " S. America " ist bereits in seiner Arbeit mit einem Fragezeichen versehen und muss als falsch angesehen werden. ...
... Tongorma latreillii (Guérin- Méneville, 1839) wurde in Namibia (Stål, 1858), im Senegal (Blöte 1935, Dolling 2006) sowie in Südafrika nachgewiesen und gilt als südlich der Sahara vorkommendes afrotropisches Faunenelement . Allerdings kommt T. latreillii rezent auch im Jemen vor (Dolling 2006, Kment & Rédei 2013), wo die Art ein afrotropisches Faunenelement der Arabischen Halbinsel darstellt (Kment & Rédei 2013). Die bei Blöte (1935) zudem zu findende Angabe " S. America " ist bereits in seiner Arbeit mit einem Fragezeichen versehen und muss als falsch angesehen werden. ...
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A list of 204 species of the Miridae from Gilan and the adjacent prov- inces of Iran is provided. The following 58 species are new for Iran: Adelphocoris vandalicus (Rossi, 1790), Anapus freyi (Fieber, 1864), Atractotomus mali (Meyer- Dür, 1843), Blepharidopterus angulatus (Fallén, 1807), Brachycoleus caucasicus (Poppius, 1912), Camptotylus linae (Puton, 1881), Campylomma annulicorne (Signoret, 1865), Compsidolon (Apsinthophylus) absinthii (Scott, 1870), C. (A.) nanno Linnavuori, 1971, Deraeocoris (Deraeocoris) rutilus (Herrich-Schaeffer, 1839), D. (D.) trifasciatus (Linnaeus, 1767), Ectagela aspera Linnavuori, 1984, Ephippiocoris lunatus Poppius, 1912, Eumecotarsus milidius Seidenstücker, 1980, Europiella albipennis (Fallén, 1829), E. alpina (Reuter, 1875), E. artemisiae (Becker, 1864), E. nigrocunealis (V. G. Putshkov, 1975), Euryopicoris nitidus (Meyer-Dür, 1843), Excentricus planicornis (Herrich-Schaeffer, 1836), Globiceps (Globiceps) coryli V. G. Putshkov, 1970, G. (G.) sphaegiformis (Rossi, 1790), G. (Kelidocoris) fulvicollis Jakovlev, 1877, Halticus apterus apterus (Linnaeus, 1758), H. darbandikhanus Linnavuori, 1984, H. macrocephalus Fieber, 1858, Lepidargyrus putshkovi Drapolyuk, 1993, Leptopterna putshkovi Vinokurov, 1982, Macrotylus (Alloeonycha) horvathi (Reuter, 1876), M. (Macrotylus) talhou- ki Wagner, 1976, Megacoelum infusum (Herrich-Schaeffer, 1839), M. pistaciae V. G. Putshkov, 1976, Nanopsallus carduellus (Horváth, 1888), Oncotylus (On- cotylus) nigdensis Linnavuori, 1961, O. (O.) viridifl avus longipes Wagner, 1954, Orthocephalus fulvipes Reuter, 1904, O. parvulus Reuter, 1891, O. saltator (Hahn, 1835), O. vittipennis (Herrich-Schaeffer, 1835), Orthonotus kiritshenkoi Josifov, 1964, Orthops (Orthops) basalis (A. Costa, 1853), Orthotylus (Orthotylus) margi- nalis Reuter, 1883, O. (O.) nassatus (Fabricius, 1787), O. (O.) populi Drapolyuk, 1991, Phytocoris (Ktenocoris) ulmi (Linnaeus, 1758), Pilophorus cinnamopterus (Kirschbaum, 1856), P. perplexus (Douglas & Scott, 1875), Pinalitus cervinus (Herrich-Schaeffer, 1841), Plagiognathus (Plagiognathus) zuvandiensis V. G. Putshkov, 1978, P. (Zophocnemis) bicolor (Jakovlev, 1880), Psallus (Mesopsallus) ambiguus (Fallén, 1807), P. (Psallus) cruentatus (Mulsant & Rey, 1852), P. (P. ) gidajatovi Drapolyuk, 1987, P. (P. ) salicis (Kirschbaum, 1856), P. (P. ) varians va-
A revision of the superfamilies Coreoidea and Pentatomoidea (Heteroptera: Pentatomomorpha) from Pakistan, Azad Kashmir and Bangladesh. Part 1: Additions and corrections of Coreid and Pentatomid fauna with phylogenetic considerations
  • I Ahmad
Ahmad, I. (1979) A revision of the superfamilies Coreoidea and Pentatomoidea (Heteroptera: Pentatomomorpha) from Pakistan, Azad Kashmir and Bangladesh. Part 1: Additions and corrections of Coreid and Pentatomid fauna with phylogenetic considerations. Supplement of the Entomological Society of Karachi, 4 (1), 1-113.
Insect fauna of Pakistan and Azad Kashmir-some groups within the order Hemiptera
  • I Ahmad
Ahmad, I. (1980) Insect fauna of Pakistan and Azad Kashmir-some groups within the order Hemiptera. Proceedings of the Pakistan Congress of Zoology, 1, 115-155.
A new genus and a new species of Phyllomorphini (Hemiptera: Coreidae) from Pakistan, with phylogenetic considerations
  • I Ahmad
  • M U Shadab
Ahmad, I. & Shadab, M.U. (1974) A new genus and a new species of Phyllomorphini (Hemiptera: Coreidae) from Pakistan, with phylogenetic considerations. Pakistan Journal of Zoology, 5 (1973), 181-187. [For year of publication see Ahmad (1980).]