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Flora and vegetation of Gävdos (Greece), the southemmost Euro-
pean island. I. Vascular flora and chorological relations
ERWIN BERGMEIER
RALF JAHN
&
ARMIN JAGEL
ABSTRACT
BERGMEIER. E.. R. JAHN & A JAGEL ( 1997). Flora and veeetation ol Cavdos (Creece), üe
\ourhmmor I üopean ßlu'l I vs\u r 5o's "-o cho'olo8'cdl ?|Jro1, t a4dol'?a '1 10\
153.In English, English md Fre.ch absbacß
457 vascular plä.t species (+ 12 infnsp.c1fic raxa) dc ßorded trcm C6vdos, üe soudemnost
Europe.n island, including 297 (+ ll) .es r*ords. M,'i.arrd ",Ed is a n* rocord for lhe
Afte* Aota; A*asolß epislattß, Ercdiu .iututiun *bsq bipirnott^, Lolirh subtlatud,
trD Ha?llrpua to xnd h'r 'ortd,uh.p ,D/ rr,'p, rtr.e1' T" 6rdirgs.o' olhmdionc.
b K rin o' üe qoL'\ Aegu ,,p/.,r,r s,rJd,,r r $demi. b C6\ da WLhin rhe Floo Furc-
pßa bordEn of GE.e, rhe fol lo*ire r. confi ned ro CAvdos: ,.{ r., nio tqha-o lbo. Ca ihc^e
ptl.^ru, Chlontd.phora hd?ntud md ,Ad€da odolaia. The occmn€ ot eeeial da nor col
lected on Glvdos sincs 1904 ß confimed dd connd6 addcd lor the chomloCically signihcst
me. Gävdos is, phltogeogBph ically, pad of the Cardaegean region Ther ft älso stug linls
!o Nonh Alncü Cyremica
RtsuME
BERGMEIER. 8.. R. JAHN & A. JAGEL ( 1997) FIor. cr v0ga6lion de cdvdos (cdce), l'ile la
plus 66ridiomlc d Eurape I Flore vasculoie el relaions chorclogiqus Candollea 52: 305 354
En lnClais, n:snn,! anglais d nanqais.
Pour la fior ds Grvdos, I ile la plus nöridionale d'EuoD€, a57 espi.6 (+ 12 srspdcs) de
planres vascubnes onr 6rC relevees. donr 29? (+ I I ) pour lä pftdiap rnis. Matriaria turer est
une nolveaur6 pour h nore gEcqß. ,l*asdlus epElorn. E adiun ci.uLa.iun slbst. bi,M(-
tu . Loliun subutatun, Utleiaela punil. d n.ia latiw ilb.D. onphüdry so.t des nouve
tes ou ont q leü pdsence co.fim6e soit pou.la CIäe, bir pou les iles dE h Mei Egde. Ddr
les linites de la G.Ace €bblies ddr Flon Eüop@, les *Daces suiv. es ne s Lrcuvenl que sü
l llc dc G'rdos: A ehisia herba-alba, Callrn he Nleba, Chla,tdoptaro nidenkla, Resedd
.a.'oto Bupbutun saüdnnuü en la rLe end{n que de G,j\do\ Laprsenedcphnr.!dod
. e\fl?n* d ara Flü ee rCnhee depJß leM s pu eoe 'onhm__ L'. d.pecE phqogrld?
phiqB de Cövdos et de cetuines espöces sonl mnnentds
(EI ,,/ORDS. Grcecc Crete Cävdos kland v$ul$ frora Chorol$t
Intmduction
The South Aegean island of Gävdos, though small, is of considerable ph).oseosraphical
inter€st due to ils outlying position as the southemmost Europm landmark towards lhe Libyan
Sea. Africa is in fact closer than Athens, but lherc are linls to both nainland Greece and Twkey
rG)r05 (rer) ETANAUES DE CENEVE I99
306
\ ra rhe sourh Aesea ßland m. ln an allempr lo sne a comprehenrrve dccounr oirhc borany and
land,cdpe ecolos-J olUavdos lhis conlnburion provides a cäralogue ol lhe !a{(ular flora olrhe
i.land fte c"rai6eue s based chrefl) on out o;n.ollecrron' anä trcld nore.. as $ell a" on tull
evalution of the botatical literature. Unpublished data dc included from the collections ofB
Snogcrup and H. Ruo€mek, Lund. The main chorolosical traits are analysed and discussed Flo-
ristia si;ilarities to neishbouring regional iloms are cälcutated ih order to cxpress phytogeogra-
Geographicrl backg.ound
cävdos is a Greek island administcred by the west Cretan Salinos district (eparchia) within
rhe Drelecrure (nömo\, Chaniä. I hc 'sldnd .ompn'es c. \) km dnd b rruäred belqeen t4'48 -
52.i \ and 24'02 5 -08 1 rn rhe Libyan Sea. Ir r' sepmlcJ hom lhe large ßland oI \Jili (C'ele)
lo rhe nonh by a din (e ot c l7 h;. No pan ot F;rope lres tunher 'o-ulh üan ( ape Tnp,l, on
G;\do". The ilonh Afncan Cyrenarca codv is c. 250 ltlomerrc' disrdnr. .eparared bv open "ea.
whercas the southwest Anatolim mainländ somewhat more than 400 km bul connected via Kdti.
the Karparhos archipelaso and Rödos, fomins the castem pan oflhe phlos€osraplically highly
sisnifrc;nr South Aisean island arc lrs $estcm pan exlends \ rr' Anrikilhira ro Kithua. and ends
on= Fldfonisidnd the sou'heasrern coasl olthe Peloponnese. somc )00lm from Gävdo' Deep
sea ,nair. oi.1500 m below 'ea level bctseen Libyd dnd Cä\dos dnd 1500 m beMeen Kriri and
Cirdos LPFANNE\slll-l . Iqö0, indicare rhe e\rreme submanne ropograph\ Cätdos irsell
riscs to 368 m above sea level.
The ist d has rhe 'haFe ola rrape/oid, $ ilh a long noahtrcsr lo soulheasl onenrared sourh
coa{r olr. l0 km whrch lorms dn rmpresi\e escarpmenl 200 lo more rhan 100 m high.
Nonheash{ard cnvdos shows a moderate inclinc over a distanc€ ofc. 5 km towdds the northem
clitrcoasts and sandy b€aches. Below 200 m äbove sea lev€l chieffy t€rtiary (middle mioce.€)
fomations with sandy soils occur. thus comprising the nonhem (noltheaslem) halfofthe island
fte morc clc\dled sourhweslem pad i{ lormed b} an older cDsrdlline l'meslone serie" mired
qirh lew patches olfl'sch. lhc d'eä around 'he nonheasr Cape Tsounos consisr" ol a mioccne
!olcano-sedrmenrary ophiolilic <erre\ w h a complc\ olereenrock'. 'zndsrones md bteccids.lhe
latter containing various schists, gseiss and quanzites. Coastal smds and dunes are well repre-
sent€d along tha nonh coast- Small alluvial deposils can be found ät tonenl oullets in vdious
Dan, ol rhe r.ldnd, one ol Ihcm, near cape I nprrr. bern8 pafli(ularly relevznr rn flori'ric rcms
ilue ro lhe.aInrty ol lhc sJbsnare lhegeolo$oirhci.ländr"de.cnbedtnderailblVlCfNTt
(1970) and mappcd by the Institule ofceolosy and Minenl Exploration (IGME, 1993).
lhc rlimate ol G;rdos rs chdaclcri/ed by d ver) pronoun(c.l dD summer penod oi 6 8
month" dnd moßI, ralher mild s'nrers fros' lempemturcs härdly. ifever. occur Thcrc de no
mereoroloqrcal records bur, concludrnA lrom thc ncatev !arnlall srzlron of Palcochotä in $urh-
ue"r Krniiscc (limaroCram rn Bl RCMtIER. laa5'. mean r'nnual prccipitärion can be e\necred
tobc lcss than 400 nm.In summcr nothem winds prevail whilc south winds are most frequent
during spnrgdme.
Therc are three small villases (Kashi, Vatsianä and Ämbelos) on Gävdos, eachwirhpatches
ofcultivated fields. Additionally, a nunber ofhamlels and isolatcd snall farm siles is scattered
over thc island mosl ofwhich arc now desened. Two morc settlcments, however. are Srowing
rapidly witir expanding lourism KaravÖ, the harbour site, and Sarakiniko, a sandy ar€a at the
ronh coast, wliich is already by far the most populated place in summer wilh beach lourists.
E. BERCMFIER & AL FLOR.{ AND VECETATION OF CAVDOS (CREECE) 307
Floristic explorltioD of Gdvdos
The firet docum€nted visit of a plant collccting naturalist on the islands ofGävdosdd Ga!-
dopoüla was lhat of Prospero Alpini in 1584, on his way back fron E8)"t to Padua (ALPINI,
l5a l, c'ted afler I ACK. Iqqb: 187). 84 our of 135 planr spe.te' shich $c'c laret de'c'ibed and
illu'traled rn _De planrte\olicia tAl PINI. lö2Ti oflgrnrre hom rhe C'ctdn dreä(LACK. lqq6)
Whilemosror rhepl rs ldcl p'ecße local,ry inlomarion. ar lcasr oneofrhem. denorared "Tru-
srcanrha , i\ crpli'll' rrdred ro be from rhe 'rull ßland{ 'outh of Krili I Al PINI. lö27: 5) 5 t).
Howc\er, dn' arlempr lo rdenrit) rhc illusranon wnh ä \pecrer presenr sould bc hdzatdou.
The fißt modem florislic invertigation on Gävdos was made in March 1904. when the Aus_
tri botani$ Ignaz Dörfer spenr sixdayson the island. The visitwas partofhis sixmonths'col_
lcrriunjoume) ro Knrr ,ponsorcd by üe viennd Academy ofSciences. lmvcldära dnd hir col-
Iecrioni uere'publrhed 6l VIfRH,iPPI R & RFCHINGFR (la15r. including 137 planr records
from Cavdo. $hich trcrc dl'o inLolporared rn RFCHINCI-R ' Flom Aegaed ( lo,l3i Rechrnger
himselfspent a short visit on the isla.d in June 1942. His field notes and galherings from the
notheastcm parl of Gävdos (mainly wmdy spccies due to the lale season) were publishcd by
RECHINGER (1944) and later uscd in his Phytogeographia A€8aea (RECHINGER & RECHIN-
GER-MOSER, l95l). More extensive collecting was done by H. Runenark and B. Snogerup in
May 1980. On this occasion the o.ly known endcmic of Gävdos was found a small annual
alpl"!run which was described by SNOGERUP (1984i as B. Saadia,uh. 'th.ir dala, for the
nost part. js published here for thc first time although a few rccords havc been cited by the
authoß ofvarious tdonomic and floristic treatnents (SNOCERUB 1984t CARLSTROM. 1985,
I986i GREUTER, MATTHAS & RISSE, ]984; RAUS, I99O; BORAT\NSKI & A],, 1992: VAN
SLAGEREN, 1994). Multi-disciplinary studies on Gävdos in the fields of nanfal and social his-
tory wcre iniliated, or encoumged, by various Greek institutions especially in the 1980s. The
rcporls ofVOKOL !lqc l) dd ICO\OMOU I ls8qr conrarn bur scadered flori'li( dära mosr of
$hrch rannor be acccpreJ due ro obv,ous mFrdenülicarionr.
We visited Gävdos durins 199.1 1996 each spring for sev€ral days, thereby erploring
dlmo{ all nan' oirhc isldn.l. The small rsland ofCJtdopoula 8lm nußhwe.r or Cardos whr.h
ha, apparcirrlr nor been v!.iled b) a bolanr{ incc Alpini\ rimes. remarns dlmosr unlnown llo
risticälly sincc inclernenl weather conditions did not permit our visit. Our specimens as well as
exte.sivc field observalions more than doublc the number of 169 taxa so far published on Gäv-
dos. The tolal number of taxa known (including subspec;es) is 470.
Thc relevant flonslic exploration on Cävdos (in brackelsr nurnber ofspecimens, followcd
by the herbarium acronym of thc rcspccdve collections) ca. be summarized 6 follows:
19.'24.3.1904 - L Dörfler (137i w)
6.-7.6.1942 K. H. Rechinscr (36; \V)
9.-12.5.1980 H. Runenark, B. Snog€rup (c. 220; LD)
17.7. and 2.10.1991 U. Mahhäs(7t B. C)
23.-2'7.5.1994 E. and U. Be'smeicr A. Jagel (C)
26.-29.5.1994 - R. Jahn (private herb., Resensburs, with duplicates in UPA)
27.4.-3.5.1995 E. Bersmeicr (C)
l.-5.4.1996 R. Jahn (totalfor l994and 1996: 175; privatc hcrb., Regensburg)
5.-l1.4.1996 E. and U- Bergneier (otal for 1994-1996. 170; C)
308
For the locality lisl and the noristic catalogue lhe following acronyns apply: AJ : Armin
Jagel;EB = ENin Bergmeieri RJ= RalfJalni R & BS = Hans Runemark & Britt Snogerupi UM
I ^ulz Vanhä'. In rhe Rorisrrc crralogxe rhe Jcronlms rre lollosed b) rhe ,|ecimen number
and lhe localiD code. I'ield nore\ are indicared by obs.-. lollosed b) lhe 8nd relerencets). I alr-
tude is symbolized by a lettcr (A-E), reprcsendng the coordinatcs fiom noth to soulh 34052'to
34o48'N; longitude by a one-digit nurnber (3 7), representing coordinates fiom west to east
24"03'to 24'07'E (see Fig. l). For our herbarium specim€ns. the locatities are encoded by the
srid refercnce followed by a cuned number (see locality list).Ifa localiry is unpr€cise or lies on
thc bordcr of a grid reference, lalitude and/or longitude are given by two lefiers (or numbers) (e.g.
CD67, D45). In the calalogue, quolations conceming Dörfler's specimens are taken from
VIERHAPPER & RECHINGER (1915), those r€fering to Rechinger's collections from
RECHINGER (1944). For each taxon the rrcords are siven in chronolosical order. Herbarium
specimens precede field obscrations. Thc lattcr dc sivcn for all srid referenccs except for those
with documentation from specinens. Farnilies, generä and species are anmged alphabetically
within the major classificatlm lnits, viz., Preridophlra, Gy rospernde, Dicoq,ledoneae, Mono
(ufle./onpJe Accepred nu me, are ser rn bold-fJce tlalics: lhe) follow lhe recenl flora ofrhe Cre-
ran area (JAHN & SCHONII-LDtR. lqor) and 'clcranr ta\onomrc lilerature. S)nonlm. or
nisapplied names us€d by VIERHAPPER & RECHINGER (1935) and RECHINGER (1943,
1944) are also given. Non-established taxa are designat€d by square brackets [ ]. taxa enoneously
cited in liteature for cävdos re eiven in rcund brackets ( ). Th€ frequency of all taxa observed
is approximat€d, b6cd on ourficld observations.
Lisr of coll€cting localities
dound As. Io nis (34o52'00-20"N/2.1o04'00 50"8), ak. 090 m, EB 25/5/1994.
315i995, 6t1/t996
N coast I km E Ag. Ioannis (34"52' l0"N/24"05' l0"E) alt. 10-50 m, RJ 2715/1994
A5-l
24'O2E Ot ty' 06' 08'07
51'
34'4A N
f_ig r The isldd ofcävdos and the sid code adopred in rhe rexi for indicrring locatiiies
E BERGMEIER & AL. FLOM AND VECETATION OF CAVDOS (OREECE)
Bl'l Potm6s bay (34'51'35'N/24'03'40"E), ak. 0-10 m, EB 29l.1/1995
84-l fam Pcrihcilä. al lhe end ofthe lrack froft Kastrinorthward (3405 l'00- 10"N/24"04'50-
59"E) alt. 180 rn. RJ 2/.{/1996
85-l Pot. Ag. Pavlou (34'51 '10-40"N/24"05'20-30"8), alt.60-100 n, EB 2615/1994
B5-2 w uppcr pan of Pot. Ag. Pavlou. 0.5'lkm NNE of Ag. Pmdeleimon
(34'51'10'N/24"05'10"E) alt. 100-ll0 m, RJ 2715/1994, 2/4/1996
86-l S akiniko bay (34'5 I '30-50'\/24'06'10-50"E). ah.0-10 m, EB 2315/1994,2'114/1995,
30/,1/ I 995. 3/5/ l 995, RJ 26 291st1994, 4l4t t996
86-2 Pot. Sarakinikou. S of Sarakiniko bay (34'51 '00-20"N/24'06'40"8), alt. 10-30 m, EB
23/5tt994, 2',7 t4tt995, 3/5I t995, N 26-29I5It994, 4/41t996
86-3 SW Sarakiniko (14'5I l0 'N/24o06'20"E), Ält.30 m, EB 30/4/1995, l1/4/1996
86-4 NW Sarakiniko (34"51 '50-52'00"N/24'06'20 30"8). alt. 20 m. EB 30/4/1995,6/4/1996
86-5 400 m W Sarakiniko bay (34'51 '55'N/24'06' l5"E) ah. l0m,RJ27/5/1994
87-l above Karavö bay (34'51 '00'N/24'07'00- l0"E). 20-50 m. EB 23/5/1994, 5/4/1996, RJ
872
B7-3
B7 -4
cl-l
c3-2
c4 I
29t5/1994, 4/4^996
between Karavö and Akr. Tsouhos (3.{"5 I ' 10'30 'N/24o07' l0-20"E), ält. 30-80 m, EB
27 /5rr994, 30141995, 5/41 t996
Karavö bay (34'5 I '55'N/24"07'00"E), alt. 5 n, RJ 4/4/1996
sidc valley E Sarakiniko bay (34"5I '25"N/24'07 05"8). alt. 30 m. RJ 4/4/1996
Ämbelos, cultivated lÄnd and adjacent phrygana (34'50'25 40"N/24o03 20-30'E), alt.
310-330 m. EB 2914/1995. 215t1995. 1014t1996. Rt 2ul5/t994.2141t996
NE slope of Fanari hill (34'50'30'N/24'03'40"E), alt, 100 m, RJ 28/5/1994
abandoned hamlet (Frageliana) between Kastri and Ambelos (34o50'20"N/24'04'40"E),
Ält. 300 m, EB 2914/l995. I ol4t)996. RJ 28151t994,2141t996
C4-2 near Ag. Pandeleimon (1.{'50'50"N/24'04 50"8). ak. 200 m, EB 2615/1994, 10/4/1996,
R.J 2/4/t996
c5 l Kastri, the village and adjacenr cultivared fields N ofit (34050'10 30 'N/24"05' l0-30"E),
alr. I 20-200 m, EB 2415/ I 991, 7141t996, Rt 281511994, 21411996
c5-2 0.5 km sw of Kastri (]4'50'00/24'05 00 E), alr.260 n, EB 28/4/1995
c5-3 ravine E of Kasrri (14"50' l0"N/24"05',20 8), alt. 1.10-180 n, EB 7/4/1996
C5-4 NE ofKaslri, slope Eofthe mvine (34's0'20-30"N/24"05'40-50"8), alt. 100-120rn,EB
9t4^996
C5 5 Kapän6lou region, 500 m E ofAg. Pandeleimon (34'50'55 'N/24o05' l0"E) alt. 120 m,
RJ 2',7 I 5 t 1994. 2t 4/ 1 996
C6-l Spida Papadias, above Korfos bay (14'50 20 55"N/24"06'30-45"E), alt. .10-100 ln, EB
21/511994, 27 /1t995, 8t1t t996, N 2615,L994, |4/1996
C6-2 2 km NE of Kastri, neü As. Christos (34'50'30-35'\/2.1'06'00-20"E), all. 80-120 m.
EB 24/54991, 29/4^995, PJ 28t5/\994
C6-4 between Karav6 ud Smkiniko, olive grove and roadside (34'50'40-
5I '00"N/24o06'30"8), all.50-60 m, EB 2615/1994,28/4/1995,l0/4/1995. RJ l/4/1996
C6-5 Pot. Samkiniko. between Kastri and Karave (34'50'30-55'\/24'06'10-30"E). 20-80 m.
EB 7 t1t t996, N 2t4/t996
310
C6-6 between Kastri and Ag. Georgios. cultivaied fields (34"50'00-10'N/24"06'00-30"E), 80-
120 m, EB 814t1996, RJ 114t1996
CD67 Korfos bay, smdy beach snd rocky coasl (34"49'00-50'50'N/24'06'50-07'30"E), alt. 0-
40 m, AJ 2615/1994
D5-l between Vatsianä and Kasri (14'49'20-50"N/24o05'10-40"E), alt. 220-250 m, EB
2415/1994
D5-2 W ofVatsianä (14'49'20"N/24o05'40"8), alt. 220 m, EB 1/5/1995
D56 Vatsiuä, the village and adjacent cultivated fields (34'49'20'N/24"05'50-06'10"8), alt.
220 m. EB 2814/1995, 8141t996
D6-l NE-faci.g slope N of vatsidä (34.49'20-50'\/24'06',00-20"E) alt. 100-200 m, EB
24ls/t994, 28/4/1995, I l5/1995 , 8/411996t RJ 26151t994, 3t4/t996
D6-3 As. G€orgios, op€n woodland and cultivated fields (34'49'55"N/24"06'30"E), alt.80 m,
EB 2814/1995,814/1996, RJ t/4A996
D64 E of Vatsianä (14"49',10"N/24"06'00"E), alt. 220 rn, EB l/511995, RJ 261511994,
3/41t996
D6-5 small mvine dove the E coast 1.25 km E ofvatsimä (34o49'25"N/24o06'55"E), alt. 80
m. RI3/4/1996
D6-6 Korfos bay E ofAg. Georgios (34"49'55'N/24"06'50"E). alt. 0-5 m, RJ 1/4/1996
D6,7 r km NE Vatshnä (34"49'35'N/24"06'35"E). ak. r00 m, RJ l/4/1996
E6-l I km SE Vatsianä (34'48'55'\/24'06'20"E) 240 m,Rt 2615/t994, 314/\996
E?-l S ofAliki, Limni, NE Akr. Tripiti (14'48'30/24"07'30"8), alt.0-10 m, EB 28/4/1995,
t/5/t995,RJ 31411996
E7,2 Ak Tnpiti (34.48'05'N/24'07'15 20"E). alt. 20 m. RJ 3/4/1996
PTENDOPHYTA
Adiartaceae
Adianlun .apilus-rercis I LB obs. 85 rarc
Asple iacede
Asplefiun ceterach L. EB obs- C4, C5 - rare
Grnnognnnacede
A"og,anno leptophy a (L.\ Lir,k - Dödler 65: "Felsen bei Kastri" (C5)
Seldgi"e a.ede
Selogiie o denticulara (L.) Spring - EB, RJ obs. D6 rare
E. BERCMEIER & AL. FLORA AND VECETATION OF GAVDOS (CREF,CE) 3ll
Sinop@n.laceae
Cheitonthes acnsnca (B^lbis) Tod. - R & BS 47777: "Kastri" (C5) (see aho ZIMMER, l99l:
231); RJ obs- C4 nre
GYMNOSPEßMAE
Cupres@ede
Jutipetus a.rc.orya Sm. Dd4.r 02tt besonders de' Nordkusle : AP.rrrgel /Jooji
''Sand'rrand dn der Buchr Saral'niko (Bol (ree also RLCHI\GiR & RECHNGfR-
MOSER. l95l: tables 7 a.d 28, and BORATINSKI & nl., t992: 134); EB & UM 3450
(17171993): 86; EB, RJ obs. 87, C3, C4, C6, E7; EB obs. A4, 4.6, 83, 84, D5; RJ Photo:
B5-2r RI obs.86, C5, D6 conmon
Juiipetus pho.ni.ca L. - Do'fer //(] ber Xenaris (D45). Rc, rragPl /J^62i \ di8c Hügel
'am K;p I'uno (87/(see'alsoRl_aHINCER& RE( HINGFR-tOSLR. l95l:65.table Il.
üd BORATYNSKI & al-, 1992: 136); EB, RJ obs. 87, C3, C5, C6, D6, E7l EB obs A4,
A5. A6. C4. D5: RI obs. 85,86. E6 common
Pina,:eae
Pinus btutia Ten. Döaet 627("P hdlepasis"): "meist nur ca. 2 m hohe Str:iuchei'; Äe.rl,-
sel 1J6lrr "sa.dise Hüsel an Kap Tsuno" (87) G€e also RECHINGER & RECHINGER-
MOSER, l95l: tables 7 and I I, the indication ofP /'4beai' in table 7 is doubtlesdy eno-
neous); EB, RJ obs. 86, C3. C4, C5, C6, D6; EB obs. A4,84, D5, E7; RJ Photo: B5-2; RJ
obs. E6 - common
DICOTYLEDONES
Acantha.eae
Acoathus spi osusL- EB obs. C5 rarc
Aizoaceae
Aizoon hispanicun L. R&BS477I0:"Alongthe roadfiontheharbourloKastri"(BC6);,B
& Ai91-34: c6-lt RJ 96040./-16i c6-l; EB obs. c5 very scattered
Population size depe.ds very much on sufficient rainfall; after the dry winter of I 994/95 no
nlanß;cuned rn 1995- bur in 1994 and 199ö man\ indtt idudl' were found rn several lmalirics.
The indicdr,on lorCärdos ol Zysupbtlud alhun'b\ Rl in JAIIN & SCHO\Fl-l DtR I las5,
is erroneous due to confusion with this species.
Me'enbryonth?hud no.liJlorun L. f,B & 4J 94-'8: D5'lt LR 3 AJ 01' I I 3 CD6-: La a5
7J l7-li RJ 960.03- 14 t7-liFBob,.C5 scanered
it2
Pislacia le iscus L. Rechinser 13652: "sandige Hügel äm Kap Tsuno" (B7) Gee also
RECHINGER & RECHINGER-MOSER. ]95I: IabI€ I I. and BORATI'IISKI & aI.. 1992:
178)j EB. RJ obs. 87. C3. C6, E7i EB obs. ,A4, ,46, 84, C4. D5t RJ obs.85. C5. D6 com-
Apocyndceae
Neiun oleandet L. s\tbsp. oleandet Rechi,ger I 364./: "Sandstrand an der Bucht Sarakiniko"
(86) (se€ also RECHINGER & RECHINGER-MOSER. l95l: table 28, and BORA-
TYNSKI & al., 1992: 16l); EB, RI obs.86, EB obs. A4, 84, 85 scattered
Asclepia.laceae
Petiplocaa gustifoli'dLablll. Rechiryer I3647: "sndige Hügel än Kap Tsuno" (87) Gee also
GREUTER. ]973:54 and RECHINGER & RECHINGER-MOSER. ]95I:Iables 1I and28.
d BORÄrYNSKI & al., t992: t67)t R & 83 47768i "3 kn S of Kasri" (D5); x & RS
478r./: 'Kastri. Thc ravinc at thc church" (C5); ,a &lJ 94 87: C5 t. RJ 940529-10: B'7-
3; EB, RJ obs. E7; EB obs. 86; RI obs. D6 - scattcrcd
Befietidacede
Leo^tic. leontopetalun L. stbsp. t.ontopetatun dd"/ r'i 'Acker bei Ambelos" (Cl)
älmost cetainly exlincr
Anchüsa amptioca (L.) DC. Dijrlkr 84 ("A. r4rtesdr4"): 'Karstfelsen arn Kap Kanarela"
(.EU R & AS 4774I: "24 km S-SW of Kastri" (D5); tR 96-8r': C6-6i EB obs. C5 rare
(Anchusa wtiegara (L.) LEhm.) Drrle. 84i belongs to A. aesptiaca (GP,EU-IER, 1965: 210)
Echiun areMtiun Gtss. - EB & AJ 91 43: C6-\ EA 96 43: A4l; zu obs. E7 scattered
Ruslossoides a .nsis (L.) L M. Johnston Dödlet 1135 (Lithospem n aflense L.\ "vnt€t
sÄäten bei Kastri" (c5); t &AJ94-49:C6-1;Rl96040./ 62i D6-3; EB obs. C3, C5 scar
N.otost no apuluü (L.) t. M. toh.ston Döfet t 1 50 (Lithospemun apulun L.): "Karstbo-
de]n";R&RS17723: wof Kastri" (c.l); ,'t & AJ94 j7 c6-rt EB 96-34: B6-4t RJ
960401-59: C6-1. RJ 9604r1-66j C6-6;EB obs. C3;RJ obs. C4, C5 scätter€d
E. BERGMEIER & AL FLORA AND VEGETATION OF GAVDOS (CREECE)
Cactaceae
lopsnia rtcus-indica (L.) Millerl - EB obs. C5, C6 (each singl€ plants which represent rclics of
3ll
Callittichaceae
Calliiiche putchro S.hotsmar, - Dö,fq 45 ("C. truncald Guss. "): "in wasserlachen bei Kastri";
ER 96 103:C4-1, Rl960401-67:D6.7. N 960402-16: C4-l - ße
For the taxonomy ofthe pr€sent sr'€cies see SCHOTSMAN (1967). This Meditetranean
endcmic is correctly slat€d as being confined to Gävdos and Cyrenaica by COOK (1981: 550)
but the respective dot in the distribution map is nisplaced (1983: 568), and C ru,rara subsp.
occrirentaltr is enoneously given as present on Gävdos (map on p. 551).
Canpanulaceae
Canpa&lla e nusL. Döllet 97:"Kaßrboden an der Südküste"; EB, zu obs. C4, C5, C6;EB
obs. C3, E7; RJ obs. 87 ralher common
Leqousio htlbida (L.\ Delzftrc Dö.fler 6l (Speculaia hyhtida DC.): "üttet Saabn bei Kas-
tri" (c5); t 96-9' (whire petals): D56i tA 96 ./r/ (blue petäls): C3-l scattered
Cappüaceae
Cappotis spinosaL. snbsp. rrlesrfli (Sm.) Nyman ER&A-19488:C'-I EBobs.alsoinnatu-
ralhäbitats: C5-3 rare
Caryoph!llaceae
AArustedna githaEo L. EBobs.C5 ore
Arc aia leptoclad.os (Reichenb.) Guss. - rr 96-lJ: A4-1; Rl960404-03: B7':l scattered
Arcna a üüatis (Link) Sqeneel R & BS 47773b:"1kn S ofKastri" (D5)i,R & rS 4zz78l
"Kastri. Th€ ravine at the church" (C5) (CARLSTROM, 1986: 362)
Cerustiun stoü.mtun L. - ER 96-62: A4-l mrc
H.niaia cinerq DC. ER 95-41: C6-tt ER & AJ 94 123: C1-21RJ 940527-07: As-t. RJ
9601r1-2J: C6-lj EB obs. C3 scatrered
314 CANDOLLEA 52. 1997
Mituütia bbida (Vill.) khischkin subsp. t/ö'ida Ä & aS ?2869: "Sdakiniko" (86); E'A
9J--r'6: C6-l scatter€d
Mituartia neditenanea (Link) K. Maly Döfiet t t 38: "rrtstboden bei Kasl,li"t RJ 960401
3l: C6-1: Rl960402-l l: C3'l scattered
PctrcrhaEia .tubia \Rif,n) C. Löpez & Romo Dnrf?, llJ- \Koht,rLt.hta \"lh a \Güs'.)
Rchb ). _unrer Sairen ber Kasrn (C5r:,q d a-t4_84^i I 4lm \\\ ofK6rri" (BC45). EB
obs. D56. C5i RJ obs. C4 scattered
Polrcaryon te,ßphrllutn (L.) L. Döüet 37 (\& diphr ün (Cav.\ DC.): "Karstbod€n bci
Varhyana" (D56)i tR 95-1r': B6-2t RJ 960401-34: C6-l; EB obs. A4, C4 - scattered
Dörfler's specimen and t 9J-.1? belong to subsp. dirnlx' (Cav.) O. Bolös & Font QüeL
Sasina natiti aG.Dor,-R&,S47817i'SrnallstreämneartheNWpoint"(A4)tEB95-t70:
A.4-l; ER 96-39: A4-1;EB obs. C4, C5, D6 scattered
silene behen L. Dörfet 56: "ütcl saatenbeil<as'J1" (c5); ER 96-67:c6-6;EB obs. C5, D5
Silene coloruto Poier s\$sp. colorara Döüet l2: "Dünen an dcr Norrlküst€"; l? & 3S ? 78lrj
'The NW poinf'(A34); EB & AJ 94-1:86-l], ER & AJ 94-96: A.4-1, ER 95-30:86-t.EB.
zu obs. B7 scattered
Silerctu,icosaL. R & BS 47798: "Kastl The ravine at the church" (C5)
Sileae Ea icaL. ER96103:C3J ftie
Silcne io.rurna L. Dö4let aJ _Kaßrboden, R& R\ 1--92b Ka.m lhe ravrne dr rhe
chu'ch (C5r, tA oJ-9l t 3-l; PJ 010520-28 Dö-4. FB. RJ ob'. C5: l-B ob'. Cö. Dö: Rl
obs. C4 scattered
Silew s.doides Ponet subsp. sedoides EB & AJ94-15: B6-lt ER 95-81: E'7-1; RJ960103-t7:
E7-2t EB obs. A4. 83 scattered
The taxonomy follows OXELMAN (1995).
Sir?r?s,rcclt"rra loFsl. sub,p.\u.culcnn Ra hikgpr 13544: 'Slnd\trand dn der B ucht Sara-
lrnrko-(Bb) ('ee also RICHINGFR & RECHINUER-VO5LR, ll}5l. täble 28),lR d AJ
4?8.r'0j '.The \W poini tA34r; R & 8s ?_800j _Sarakrniko (Bo):/-Ad.tJo4-riBo-l:
EB obs. A4 rare. locally freque.t
E. BERGMEIER & AL. FLORA AND VEGETATION Of CÄVDOS (GRTECE) 3 I 5
sir.r. lrrßorn (Moench) ceclc subsp. ra.t c4tpd Turill R?, rdael lleg Heoogl.lJo'q
(n/e,e(,.znal6 wib. subsp. dnsr'r4or, (Guss.r Rech. nl.i: berm DorfAmbelos (C3).
R&85477b6: 24lrnS SWofKasrD-rD5,if8oJ-oo D5ör l-8, Rl obs. C5, EB obs.
CJ, Cb - narrered
Sperg.hna boccoiei (Scheele) Asch. & Cr EB 95-149: E7'l;EB obs. c4, C5 - very scalte-
Speryulaio diandru (Gtss.) Boiss. Ddder Joi "Kulturbod€n bei Vathy i (D56); R & RS
' -.1_68Ji _Alonp rhe road lrom thc haöou lo Kash (Bab): R d 8J 47825. _Smdll srream
near the NU ;orni (A4); fB 3 AJ 94-a. BO-|| LA & AJ o4'o2: Cb-l ER q6+ 4 : 87 2l
N 9ö0401-ii Cb l: FB obr. ( 5i RJ obs 84 \cancred
Se ana pa ido (D\jllnon.\ ]lfntb. ER 96'107:C4-l; EB obs. A4 rare
For corect author's citation see MILL (1996).
Chenopo.liaceae
Reta adanensis Lellen R &,S-,178./6r "1 4 km NW of Kastri" (8C45)
Chenopodiun abün L. ER & AJ 94 35 : C6-la ER 95- I 42 : D56 - tue
Che"opoditd nünleL. Dötfü 1139: "Kulturboder' b€i Vathyana" (D56); EB. RJ obs. C6;
EB obs C5. D6 scatt€red
Salsolo kotiL. stßsp. kali AJ. RJ obs. 86- l, CD67 rare
Cistdceae
Cistus crc,icus L. Döfet 106 ("C. villosß L."): Kärstbod€n" (see also RECHINGER &
RECHINGER-MOSER. 1951. table ll, md BORAT"I'I'ISKI & al.. 1992: 58 as "Cirrüs
incanLe L;'\t ER & UM 34.12 86; EB, R.l obs. 87, C3, C4, C6, D6; EB obs.44,46, C5,
D5. E7; RJ obs- 85, E6 common
Cistu. panillotus L^rn. D/le, /r,a _Karsrboden . Rethnger lJ65b:"südige Hügelam Kap
I'uno (B7r isee al'o RICHI\GIR & RICHI\GI-R MOSER. l95l rable ll d
BOR?|TYNSKI & al-, 1992: 6l); EB & UM.l449i 86; EB, RJ obs. 87, Cl, D6; EB obs
A4. A6. C4. C5, E7: RJ Photo: C5-1, RJ obs. 85, C6, E6 common
Cisrus salviifolissL. Dö4et t05: "K^tstboden" (see also BORATYNSKI & al.,1992162). EB
& LIM 3448: R6,EB.RJ obs. C3, C6. D6: EB obs. C4, C5, D5, E7l RJ Photo:C5-1, RJ obs
85, 87. E6 common
Futnaho arabi.a \l .) Spach Dötfct vt: 'Kt-tt'oden der Südküsrc :,ta,inat IJ66^b:
''sandrße Hüsel m Kap lsuno- (87) (see äho RTCHINGIR & RECHINCER-MOSIR.
I95l:rablcllri Rd 8s?_t)8/i _Along the road ftomlhehärbourro Ka\ra (BCol: ad
85 1--40 '2 4 km S St\ of Ka,ni (D5)i l-8, RJ ob". Cl, D6; I'B ob\. 87, C5, E-; RJ
Photo: D6-l; RJ obs. C6. E6 rather common
Funaho ththifolia ll .t\ ebb Dö41c, lt]I:'Ktslbodcn dcrSüdküstc ; Rp.htnq l3bo-a
f\ü. 1d"r6 tcar.) Oro\sc' r: -dndige Hügel dm Kap Tsuno (Br). LB. Rl obs. Bö. 87.
C3, C5; EB obs. A6, E7; RJ obs. 85. C6. E6 common
Helia theftrn stipulatun (Forssk.) c. chr Ea&AJ94 132:87-2;Ea 9J.I.l0i 86-3-rN
n beruia guttata (L.)Four. s.l. EB obs. A4, 86. B7i RJ obs. 85 scattered
Plants with clehtogamous flow€rs and small rosettes were s€en as well as plmts with open
Coüpositae
Aetheoüiza bülbosa (L.) Cass. subsp. ,rtcrodeprdro R€ch. fiI. n&BS478r9r"1 4kmNw
ofKastri" (BC45);,{J 960r'02-./9ai C5-l; EB, RJ obs. 87, C6, D6, E7; EB obs. A4, 84,86.
Cl. C4, D5 comnon
Anthedis chiaL.- EB 96 6-1i C5-l (specimens belong to I üo.,4r4 Greuter & al.) very scaG
Atthenß ||gidaHeldt. srbsp. tigitttt Dö4let 40("A. c,errrd'): "auf Karstbode n" I Ea 95-78:
E7- I ; EB. RJ obs. C5j EB obs. A4j RI obs- 84, C6 rath€r common
Anenisia heüa-alba Asso A & tS.l/688r "Along the roäd beiween the harbour and Kastri"
(c6) (GREUTER, MATTHAS & RTSSE, t984 271). ER & AJ 94-82: C6-2; Rr 910528-
08, C6-2r RJ Pboto: C5 rare
As,eisct s spinosus lL.) C. H. Scbulrz EB obs. D5i RJ obs. C5 mr€
Atdctlis cancelata L. stbsp. .Mceltatu Ddd"r 28i "Karstfelsen b€i Vathymi' (D56); EB,
RJ obs. C4, C6; RI obs. C5, D6 rather cormon
Beuis asnuaL. EB 96-Ill: C4-2 ffie
Reliun ninüt'n (L.) L. Dö41er 77:"Felsen amKap Kamarela. sehr selt€n" (E7); n & aS
4-17816: "The NW point" (A34)a ER 95-84: E7-ltEB obs. A4. 83, 87 scatrered
IBtRuVt p&Al t-toRAANDVI(rl tCnO\ Ot uAVrrO\LCRLFT tJ .] r7
Cale uk anenskL. Dö41et 62:"Kaßrbode beiKasjli tER95 92:C3-ltER9699 C3-l;
EB obs. D5, D6; Rl obs. C4 scattcred
Ca mceluscae.leus(L.)C.Prcsl R & BS 47696: "Along the road from the harbour to Kas
t1" (c5); EB & AJ 94-86:C5-l; EB obs. Cl rare.locally liequent
Caduus pycnocepholusL. Dö4Iet 89: "Kß'lelser am Kap Kamarela" (87); EB obs. C3, C5;
RJ obs. C6 scattered
Carlino g.aecaHeldr. &Sart. EB. RJ obs. Cl, C5; EB obs. D6, E7; RJ obs. C4, c6 common
Catthanus leucocaulos Sm. ER & AJ 94 130: C6 4; EB obs. C5i RJ obs. D6 - scattered
Thc spccimcn mentioned above was enoneously assigned to C äoi$ieli Hal. in BERG-
MEIER & MATTHAS (I995),
Ce,taurca n.tirensis L. R & 8S 47767:"24kmS-SW ofKästri" (D5)ttR &,!J94-.19j C6-
l; ßJ 940526 10: C6-6;EB obs. C5 rare,locally ftequenl
Chlanydophotu tidenta,a (Delile) Less. Dö'lpl 1r "Karstfelsen und Meeresküste am Kap
Kajlnarcla" (E7):, ER 95-72: E'7J. EB 96-1t3: C5-t. RJ 96010t-58: C6-tt RJ 9604U-7A:
D6-6i EB obs. ,A4, 83 scattered
Chry nthenu corc\aiu L. Dörfer 16: "Umrer Saaren bej Kastn" (C5)i EB. RJobs. C6;
EB obs. C3, C5. D5, D6i RJ Photo: D6-4; RJ obs. c4 scattered, leally abundant
ChOsan henum segetun L. Döüet 47: "nftet Saten bei Kastri" (C5); EB, Rl obs. C5 rarc,
Cichoriun endi'ia L. snhsp. di"/rr.atür, (Schousboe) P. D. Sell ER&AJ94-3t:C6-1.ER
9J-?9, 86-2: EB obs. C3 scattered
Cichoriun spinosun L. AJ obs. 44; EB, zu obs. E7; RJ obs. D6 rare, locally ftequenl
Crepis oenca Boiss. - Dö.ller Tj "sandiger Bod€n unweit Sarakiniko" (B6)j R & rS 47721, 'W
ofKäslri" (c4)t Ä../ 941519-02: 8',7-1; RJ 960401-02: 87-ri EB, RJ obs. C4i EB obs. C3,
E7; RJ obs. 85. C5, D6 common
Crepl' hahiiora Sm. R &,S.l.l73J2i "The NW polrt" ( 34\:EB&AJ94-ll3:CD67;ER
95 120: B3-\t RJ 910526-33: E6-l; RJ 960101-01: Bl-3;EB obs. A4 scaltered
C'e"ü"asirra (Somrn.) Me:,y.n. EB96-37:A4-1tRI9614r]-2lrC6-liEBobs.C4 verysca!
318 caNDoLLEA 52. 1997
Ct.pis tybakiensis V|eft. t 9J-86: C5-2t EB, RJ obs. 87; RJ obs. D6 scattcred
Crupitro ctupi at:bun (Motis) vis. R & BS 47790 "Kastri. The ravine at th€ church" (C5)l
,B 9J-1?/. D6-ll RJ obs. C5 scattercd
Cynara comieeft Lindley n & aS47z]5r "w ofKasrd" (C4), ER & AJ 94-67: D6-t;EB obs.
C3j RJ obs. C4, C5; also recordcd fiom Gavdopoüla (WELTER-SCHULTES. 1995) scar
Did.ichia viscosa (L.) Greuter EB obs. 85 rare
Filogo aegaeaw^genitzstbsp- dnis,ara Wagenitz Rechinget 13671("E spdthulata !ü. siia"\:
_,andise Hüsel amKao [.uno (87)(WACF\lTZ. 1970: ItOrGREUfFR, lq'3: 50),R
ABSZ'80Jb 'l km\\ or Ka"rri (C45r:ßd 8S.l'8JJi -The Nw poini (AJ4rRd
a-t47807: Sdakinilo - tBö), E B & AJ 04 - I A8 A4J ; EB 95- I 5. B6-L A a4n5 2^-05 : C n
t; RJ 940526-14:D6-l RJ 960401-09: C6-4tEB,W obs. 87, C3, C5; EB obs. 45,46, E1j
zu obs. 84.85, E6 common
Filaso cottrocta (Bolss.) Chrrek & Holub R&8547731:"2,1kmS Sw ofKastri" (D5);R
& BS ?780Jc: "1 km Nw of Kastüi" (C4s). EB 95 111: C3-lt ER 96-54: Cs-lt RJ 96A401-
z3r C6-li EB obs. D6 scattercd
Filaso crctensis G^td. - ER 96-10: A4-1; EB 96-106: C4-t r rc
The snall collcction ti 96-]06 belongs to subsp. cr"teasir. whereas ,a 96'4, matches
sthsp. crcladun Wzee tz which is known hitherlo only from the Cyclades md the Karpathos
echipclago where it is chiefly found on smaller islands (WAGENITZ, 1970: 125f.). Fcw speci-
mens in lhis collection ar€ lrahsitional between the two subspccics. Such fonns were alreädy
mentioned by WAGENITZ (1970).
Filago .tiocephala Gßs. - EB & AJ 94-122: C4-2 we
Filaaopa i.al AdaSr'_00ri Alonsrhe road fiom rheharbourro Kdnn (BCol:Rda.S
r'-332 fhc NW porna lAlar; LB obs Bo. Co: RJ obs Cs scadered
Iilalo pf.oni,lata L Do,Jtet 8- \l' sparr,/.Id PreJ): Kaßrbodcn bci Kap Kamarelä : R d
8J 4-_2r. -WolKasrri (C4):R&8S1--51:-2 4kmS-SWol Ka'tn \D5).tB&,41
94-52: C6-1t ER 95 3: B6-2t RJ 960401-24: C6-l:EB, RJ obs. C5t EB obs. A4, Cl, D6
Geroposon hrbidus (L.) C. H. Schtltz ER & AJ 91-125: C6-4 mre
E. BERCMEIER & AL, FLOLA AND VECETATION OI CAVDOS (GREECE) 3t9
Hc.lwlois apadioloi.lcs ( 1. ) F. W Sch mrdl subsp. rr ogadioloid?. \incl. H. tubaeformi.t Ten \
" Do.fr 15 r'H. , retico lL , $illd. ,: _Dünen an det No:dlnJrc- t Dö41et 98 \'H tubac
&'ni Ten.'): "Südküste"; EB obs. A4, 85, 86. C3, C5, D6; RJ obs. BT. C4 .athercon-
Hedrpnois .hasodioloides (L.) F. W. Schnidt subsp. rorsPzt'errir (Murb.) Hay€k EB obs
A4, B4, E7; RJ obs. Cl, C4, C5, C6 common
Helichtlstd co,slobatun (Viv) Steudel Ä & as .l/822i "Small stream nezr tbe NW point"
(A4\: EB & tJ 94 66 : D6-l ; EB & AJ 91 94 : A.4-l : ER 95-l/8i 83 l; EB obs. 86. C5, C6
EyoseÄs scab tL. Dötfd 86: "Kaßrboden , RJ 960401'27: C6't. EB. Rt obs C5; EB obs
C3. D5 scattercd
Hwochaerh oültopho s L. Dötfl l l 5- t H) pocroe'i\ detrr.a6 lL.r Bentham & Hook r:
''Sandboden : EB. RJ obs. B-. C5, Dbi EB obr. Cl, L7; RJ ob'. C4, C6 common
t.artuüsedioloL. EB. AJ obs. 87 rare
Lanwpsis qnanides (LaIn.) Dittrich EB, RJ obs. C4r EB obs. C5; RJ obs. C6, D6 scatte-
Leontotton h.bensus L. - Dö.fier I 156: "Ktstboder,"; EB. RJ obs C3, C4, C5, D6; EB obs.
87, D5. E7; RJ obs. 85, C6 common
MatÄcaia aurct (L.) S.htltzBip. EB 96-tt2:C4-2t RJ 960402'09: B4't - r?'re, but locally
New to Greece.
Onopo un toutictd Wllld. R & BS 47856: "24 km N of K6tri" (BC5)
Phosnator era"cun Boiss. & Hetü. Döde,88tPh ures,?(L.r DC. subsp.8'üe.,' rBorss
.-& Held.'. ) Hd yek ) _Kaßl bodcn bei Kap Kä mi rcl^- \11 ): Re.hinget I Jo4 o _sddiSe H ügel
am Kao Tsuno" tBT, ('ce also RLCHnCI R & RFCHINCL R-MOSI-R. lq5l: rdble ll): ,4
d As;-08^ 'Aiong rhe road from lhe harbow ro Kasm rBCö):I'8.RJobs B'.Cl.Dhl
EB obs. 84, 86, E7; RJ obs. 85. c4, c5, C6, E6 common
R.icho.dia pictoides (L.) Rot\ ER & AJ 94-t00: A4-l 6rc
320
Rhasadiolus srelatus (L.) G^etu€r - Dödet 59 (Rh. rrel/4r6 (L.) Willd. var. ed!/is (Gaertn.)
DC. f. hebelaenus DC.): "\rnter Saaten b€i Kastri" (C5); tB 96-66i C5-t; EB obs. C3, C6
Fo. the synonymy of thc nme applied by VIERHAPPER & RECHINGER (1935) to Dör-
flcr's specimen see MEIKLE (1979), and lhe key provided by RECHINGER (1943).
Se ecio leucan,hen,lolit sPoiret!^t.ptgnaeus (DC.)FloÄ EB 95-tt6:B3-t. Rl96040t-72:
D6-6; EB obs. ,44; RJ obs. 85 scattered
Records ofs. aal/ids fron Crete and lhe Karpathos island group eilher belong here (W
l<lir|\ or ro vaf, leuca theürrlt&t (E Kriti, Ka'pathos group) (see ALEXANDER, I 979).
Senecio rukans L. - EB, RJ obs. C6; EB obs. C3, D6; zu obs. 84, C4, E7 - rath€r colnlnon
Sonchüs olenceusL. ER & AJ 91-l l7: CD6'7. Ea 95-2Jr 86-2: EB. RI obs. C5j EB obs. C3.
D5i RJ obs. 87, C6, D6. E6 rather common
Steptofiinphus tubensus (Jacq.) Grossh. RJ obs. C5t EB, RJ obs. D6 scanered
Totsocan sect. Scatiotu - R & 3S47812j "l-4kmNW ofK^slli" (8C45); RJ 960401 39:C6-
l; EB, RJ obs. C5; EB obs. 44, 85, C3, C4j RI obs. D6 - scättered
Tmgopogon sin/,/,tus A\e-L^tl. R & BS 47762: "24 km S Sw of Kasbi" (D5)i EB, RJ obs.
D6; EB obs. D5; zu obs. Ci - scattered
Tyin ts leucogtophus (L.l Cass. Dö$et tt7: "K\tlturboden be; Vathya.a" (D56)i A & RS
,f7784: "Kastri. The ravine at the church" (C5)j t &,1J94'7ziC5-l;RJobs.D6 scar
Unspemütl pictoides (L.) F. W. Schmidt D net 115l: "Kaßtboden an der Südküste": EB,
RJ obs. 86. 87, C5; EB obs. ,44,84, C3, C4, E7; RJ obs. C6, D6 commo.
Conwlrulaceae
Coawlealus althaeoid8 L. EB &AJ91-134:C6-4t ,4A 9J'J8i D6-3: EB obs. 87.C3. D5.87
Conroteutus otelfolius Desi ER 95 147: D6 4 tute
Cuyuta pala$tino Boi<.. R & BS 1'-bl -2 4 km S sw ofKastri rD5r: R & AJ r---Ji -l
t'nSofKasni (D5);ÄBd AJ0l-6 86-2: RJ o4052o-r] B7-l,fB,RJobs C5,D6;EB
obs. 86. 87: RJ obs. 85, Cl, C4. C6 common (freqU€mly on Sdtürcid tbnbn and Glo-
buldtid alypun)
E BERGMEIER & AL, FLOLA AND VEGETATION OF GAVDOS (GREECE)
Crutstlldceae
ruaea alatu viv. EB95-l7l:44-1;E896-4t:44-liRJ96040l-20:C6-ll'RJ960401-61:D6-
3a RJ960402-15:C4-l;EB obs. C5 scattered
mheo eoil/anniwilld Dödet l-5 "inWassetlachen (Kaßllii{hem) nächsr Kästri. Tn eiskal-
rem wassei (see also RfCHINCER & RfCHINGfR-MOSfR, lq5l: lq0), f,B q5-163
A4-1t EB 96-22: B7-2t RJ 960401-68: D6-7t EB obs. C4, C5, D5 scattered
Accordins to TURLAND & al. (1993) ther€ is no post-1930 record for the Cretan eea.
Sedun.rcti.uh C.Presl kl960403-18: D6-5:EB obs. C5: RJ obs. C4 mre
Sedun lirorctd Gtss. s!bsp. litouln Dörjbr 472 ("5. zärud (L.) Thell."): "Karstf€lsen
unweu Kap Kamarels {L7l rfor rhe rdenrrry ofrhe sp€rrmen see RUNEMARK & GRfU-
TIR. I o8l : 20 md GRFUTFR. MATTHAS & RISSI. Iq84: 276): A d AS,/---6 "Kas-
tri. Thc ravine at the chüch" (C5); ER 95-41: C6-l: ER 96 26, 96 98: Cs-4, RJ 940527
01: B5-2: RJ 910528-05: C5-\; RJ 960401-55: C6-l:EB. RJ obs. C4;EB obs.4.1 rather
Sedu6 bens L..ubsp. bets R3 854--26:'w ol Kastri (c4rrcARl STRÖM. lo85:
l12. "S. d./rüa (Vierh ) Carlstrom ). AB oJ-,/J C6-l;RJob".C4,D6 scanered
Se& sedifohne (J'rcq.\ Patr Rechinset (les. He:üod 13661 ('s' 4/tirsmrn Poir"): 'bei
Ambelos" (C3); x & rS4z8r3r "Kastri. The ravine atrhe church" (C5); RJ obs. 85 rare
Un'bilic s honzontalis (Grss.) DC. R & BS 47787i "Kastri. The mvine atthe church" (C5);
EB 95-100: C4-ltalso in natural habitals at the eastem Korfos bav cliffcoast. AJ obs. nre
Cruciferae
Risdllelladi.lymaL- Dörful 5 5 ("\ar ciliata (DC) Hal."): "unter Saaten b€i Kastri" (C5); Döi
fer 1I3j ("\ar cotw tae (Ten.) Hal.")r'\nter Saaten bei Xenakis" (D45);EB, RJobs. C6;
EB obs. 86. C3. C4. C5. D5. D6 mth€r cohmon
Cakile na tinoscop. R& BS478lJi "The NW point' (A3); ER & AJ91-20i 86-I (ct subsp.
a€grptacd (Willd.) Nyftan) rare. locally freque.t
Capwlla bußa-postotk lL.) Medicus EB obs. ,A4, C5; RJ obs. C4i C6 scattered
ClJpcola joathlospi L. Dö4et al Gub,p. pctnco (Jo'dd & l-oumau) vie'h.): Kesrboden
än der südkrisre i R d 8J 4--J, _2 4 km S Sw ot Kastn rD5,: EB obs. A4. c4. c5. RJ
obs. C6 scattered
321
Di.lesnts a@Dtius tL.\ Des\. Döder oö tqapßtrun üegptiun \L l Batll ):'Kaßlbodm d
.rdsiidEile:Ä&sS4TTSJ kasri. The ravine ar r hi ihurch \Cs,tEB&AJqa'J|:C6'
tt EB 95- I I 0: C3-lt ER, RJ obs. C5 - scatt€red
DiDlouxis viniwa tL.tDC. Dorr./oji_KarslbodenanderSüdkü"le:R&AS4-^8'i Along
-'' ir'" -ra f..-,h. tr*Uour ro Kastri {B0,lad BSr'781Ji l 4l'm Nw olKastri (84,: ÄA
& AJ94-8: B'l-l: RJ 96040.1'0Ji C6-4 $atter€d
Etophilo pto.cor ßlelent DC. E 8 aö- I 0o : C 4' I I N 04 0 5 2 6' 2 4 Db 4 : N ob04 0 I ) 2 C 6' I :
R I06040)-l2 Cl-l:EB.tbs. A4 - scanered
Ilißchfeuia irctna (L.)Lagr.-Foss. - EB, RJ obs C6; EB obs. C3 rare
Hwznotobus t|o.unbaß (1.) Tor. & A Gta) R d AS r'78'li The I'Iw poini rRAUs
' lcc0: rö); E8 9J-8J_ F1-1, RJ ob0403- 12: E7J we
Malcolnia ahicana (L.) R. *. Rl960101-12: C64 ne
Malcolüio chia (L) DC. - Dö.fle/ 72 (!ü xercphila Yierh ): "Kaßlboden bei Kastri"; Dö'fer
72a (var. reiihitaYie .it "Dünen bei Saßkiniko" (86)
Makotnia ß.'uosa lsm ) Sm. subsp. tat 'r'1(Recb fil rA Sro -tA& 4Jo4'l0l:r'.4-tl
rB & -A.t 9a-112, CIj6lt EB 95-l l7: B3-lt RJ a0040J-l5i F?-2 scanercd
Malcolnia nana (W:.) Boiss. - EB & AJ 94 17: B6'1. N 960404-04: 874; EB obs A4-1
Raphanß nphonisnun L. EB obs. C3, C6 rare
Sin@is alba L. sutJ.o. nab.i {H. Lindb. fij.) Maire - R & ,S ?-Ö90 _Along lhe rcad ftom lhe
'harboü Lo Kasü" (BCö); EB & AJ e4- 24: 86'2. EB obs. C5: RJ obs C6 scatrered
.siu a,ir @eusiJ L. rubsD. awnsis R & aS a1'47: 2 4 km S-Sw of Kasni (D5). tA 0J-
' I I 2 C3-1. EB 9o-56: \ 6-5: LB obs. D5, Do scanered. locally lrcqueot
SisyDbr .n ofrcinale L. EB obs. C6 rare
Sisrdb.t].t., ofientaleL. t 9J-.11./i C3-1: EB, RJ obs C6l EB obs D5 scattered
Thlaspi p.tloliarun L. rB&AJobs.C5 rN
L BfRCMlltR& al I I OR^ A\D \ LuL IATIO\ Ol CÄl'ttOS r(JRlts(Fl 323
DiPM.de@
Prencepholus pls'nosus (L.) Cültet - R & BS 4773ri "w of Kastri" (C4)i RJ obs. C5 - rare
Etica.ede
Erka naniDutiflora Sal;sb Re.hmAet I 366J _sandrge H ü8el anr Kap Tsuno - ( B7l ( \ee Älso
RtCH-TNGER & RECHINGER-VOSER, l05l: rable ll, and BORÄTYNSKJ & al , lqq2:
100); EB, RJ obs. 86, 87, C3; EB obs. A6, C4, C5, E7; RJ obs 85, C6 - common
Euphofiiaceae
Et/Dhoüia exiAua L Dod./ /Ji Sdndboden l R d aS r'7_/8: !\ ol Kasni (C4,. R d as
_ 4??51: _t 4 lm s sw ol Kasm'rD<r; Ea d AJ 94-10: C6-li La 3 AJ o{oa A4'lt RJ
96040t -07: C64t ER, RJ obs- D6; EB obs. A6, C3, E7i RJ obs B7,C5 comon
Euphorbio helioscopia L l-B ob'. Cl. C5 rare
Lupho.bio parulio. L- EB obs. 86. CD67 rare
EuDhübia peDlis L. Rc.hüAcr lJ665 _an der Buchr Sa'aliniko l86)(seeal'oRE(HINGFR
' & R-EaIiINGFR-vosER l05 | : rables 28 md 2q). tB & U u J 5 ) 3: B6t EB & AJ e4' t 04 :
Et phoüia peplus L. EB 95-13:86-2t ER 96'48: A'4't. RJ 960401-08: C6'4; EB, RJ obs. 87;
EB obs. C3, D6; RJ obs. C5 - common
Mercüiolit annua L. - R& rS 47781i "Kastri. The ravine at the church" (C5)l EB, RJ obs C5;
RJ obs. D6 - scattered
Ftutkeniaceae
Frunkenia hirsutaL. R & BS 17797: "Kastn."the nvine at lhe church" (C5); R & BS 47813:
"l-4 km NW of Kasri" (BC45); EB & AJ 94-97: A4' l; ru 94r52z-r8i A5-l; EB, RI obs
C5, E7; EB obs. C6; RJ obs. D6 scattercd
Ftaücnia putvcrulenta L fd 854_8a5i -The Nw poini tA34t:I:B&AJ94'I4:86'21EB
1t Ä.t'94 l7: C6-l: LB 95-8Ji E?-l: zu obs DO ;canered
i24
Rlackstonia petfotiara (L.) Hudson ra 9J,2i 86-2; EB, RJ obs. C5j EB obs. D5. E7; RJ obs.
85, C4, D6 ralher comnon
Centuunun pulche ln lswanz) D ce ER & AJ 94-29: B6-2 ftte
Ceituunud kiuilotuü (Hoffmams. & Linl) Fritsch stbsp. ,enuiflorub R&8517727:"W
ofKasrri" (c4);R & rs 17860i "sarakiniko" (86);tt & AJ 94 95. A4-rt RJ 940526-03:
C6-li EB, RJ obs. 87.C3.C5; EB obs. A6, E7; RJ obs. 85, 86, C4, D6 - colmon
Gerunidceae
Etodhln bottt$ lcav.)BenoL Dijrfer 79:"Kztstboden" (sec also DAHLGREN, 1980. disti,
Endiun cicutaiun (L.) I Her subsp. ärpn'at!' (Ca!-) Totnet ER & AJ 91-93: A4-1. ER
95-61: C6-1t EB 96-32: B6-4i EB 96-51: C6-5 s.atteted
N€w to the Cretan area.
E. cicutariun stbsp. bipi nar!ft is distinguished from subsp. .lcrralt n by the absence of
ä furrow below the distal pit ofthc mericarp, by the ddker red violet (insread oflilac) petals, the
less deeply incised leaffets with subobtuse pimules and the less dense indumenhrm.
Ercdiun cicutunun L., L'Her. stbsp. cictta,iud ER 95-123: 86-1; ER 96-33: 86-4; RJ
94052604: C6-1. EB obs. C3 - ratber comon
The following records cmot be assisned to either subspeciesr n?i:rtng?," /J6lJr "Buch(
Sarakiniko" (86) Gee also RECHINGER & RECHINGER,MdSER, 1951: rable 28. and DAHL-
GRfN. lo80.disniburionmap\tR&BS1-693.1--05'AlonBrhcroadfiomrheharbourroKa$
rn (BC6l: A d 8S a-8-J _israk'niko {Bor; tu 00040/ roiC6-4; RJ ob,. 84, Dö
Etodit t' g.sinun (L.) L'Hä. - R & 8S 47695: "Alon8 tb€ road from rhe harbour to Kastri"
(BC6)i ,A 9J-J6: D6-3t RJ obs. C6 scattered
E.odiu laciaiatun(Cay.) willd. subsp. /a.iriatum R & ES 17873pp: "Sa:]ak;niko" (86); t
95'126: B6-\t RJ 960402 o3i C5 l; EB obs. A4, 87 scattered
Ercdiun fralacoides (L.) L'Har. R & BS 47759:"24 km S-Sw of rstri" (D5);44 9J,6r:
D6 3j EB obs. C5, D56; RJ obs. 85, C4. E7 scattered
Geruniun nolleL. EB obs. C5, D6;RJ obs. C4, C6 - scattcred
Genniun puryu,etn vill. EB. RJ obs. D6i EB obs. C4, C5 scattered
Geranit n ntundifolit n L. EB obs. C3, C5, D6; RJ obs. C4 scatte.ed
E. BERCMEIER & AL FLOR-A AND \,€GETATION OF CAIDOS (CRTECE) 325
Globaldtidceae
Gtobulatia alwu L. Dölflet 2: "Kaßrboden der No.d- und ostküste'; A"chiryet 13659
''sandige Hügel am Kap Tsuno" (B7) (see also RECHINGER & RECHINGER-MOSER,
l95l: table ll. and BORATYNSKI & al., 1992: 120);R & RS l77rli "Along the road from
the harbour to Kasrri" (BC6). R & aS 47769:"3k]x.s ofKastri" (D5);rB &,4J91-J.l: C6-
l; EB, RI obs. 86, 87; EB obs. 46, C5; RJ obs.: 85, C3 common
Guttiferae
Hrpeficufr tiquetifotian Tuna EB obs. C3, D56 very scatl€red
Labidtae
,ajdso i'a (L.) Schreber R&B,t?78r?:"Kasr.Theravineatthechurch"(C5);ER&AJ94-
l3: B'7-1i Rl940527-05: C5 5t RJ 960401 ./l C6-l - scättered
According to TURLAND & al. (1993) there ee no published records for K-riti since 1930.
Coti.lothtnus capitatüs (L.) Reich€nb. 61. Dö4ü 103: "Kaßthoden': Rechinset 1365t:
"sändige Hügel am Kap Tsuno" (B7) (see also RECHINGER & RECHINGER-MOSER,
1951. table I l, a.d BORÄIYNSKI & a1.,1992: 70); tB & Uü Jsr.ri 86; EB, RJ obs. 86,
87, C3, C4, C5, D6; EB obs. A4, A6, 84. D5, E7; RJ Pbotoi B7-4; RI obs. 85, C6, E6
Laniud anpbn@uk L. EB, RJ obs. C4. D6i EB obs. C5 scattered
Man biun 'ulgoreL. Ea&AJ941A C6-ltEB obs.86, C5, D5 scattered
Ncpeta ncli.sifolia I am. Dör,t"/JJi_Ksllcl'enberKa'tn.RdBJr__r0 Alongrheroad
liom rhc hüboJr tu Ka,rn-' \\ 6), t B & AJ o1 -tt l C 5- l t RJ aruo) - l 3 a l-l ; l- B, Rl obs.
C4; EB obs. C3, C6 scaxered,locaUy frequent
Phlonis Iruricßa L. ,{ & RS 47788i "Kastri. The ravine at the church" (C5)i EB obs. C5
Pr6iud najus L. Dörfet 100 (f. crcticun Hausskn.); EB, RJ obs. 86. C3,C6, D6; EB obs.
A4. A6, 84. 87, C.{, E7j RJ obs. E6 common
Salria veüenauL. - ER 9J-J9i C6-l; EB, RJ C5j EB obs. C3. D5 scahered
Sol'ia 'iri.ti. L. Döüet 29: "l<atstfelsen bei Vathyani' (D56); tB 96-74: D6-t arc
326
Sainejo "ana (P. tt. Davis & Doroszenko) R. Jahn EB obs. D6 - rare
Satueia n?nosa Desl. - R d BJ ?-r'So AlonC lhe road lrcm the harboü ro Kasm (BCöl:R
d aJ 4__JJi -2--4 km S SU ofKa.rn tD5r; fB d ,.{J 04-4J Co-l: tA oÄ-rr. Cö-5r EB.
zu obs. C3, C5; EB obs. E7: RJ obs. D6 - rather common
Satur.iathwbruL - Dö:,tel 6., _Kaßlboden berKa'rn (scealioBORATINSKI&al,l0q2:
2Alri ? d 8s ? _öo-'_Alone rhe road fiom the häöour lo Ka.m (BC6r:Ad a-Sr'_/oi
_w of Ka"m (C4,r l B, Rl obs. c4, C5i EEobs A4, 84, Bö, CJ, Cö, F?, RJ obs. 85
Sidetitis ca"nidcdsSt^pl Döüer 21\'5. rcaoao L 'ubsp eü-nnana Hay.- t. Kaßrboden be|
Kdyri : fA 9r-4l Cö-tr EB. RJ obs. 87. C5i I B obs. 44, Cl; Rl obs. C4 scanered
Teu.iu"t b.clilotiun sclßbet Dörflet 85 an fehen un$eir Kap Kamdrcld (E?,. fB oJ-
l2 t. 864: RJ 94052'-02 Bö-5i FB, RJ obs. l-7; tB obs. Aö, 87, RJ obs. CO scanered
Teuciaü .li'aicatuü Hel&. srbsp. di'a catnn E8&AJ91 128:C6-4(singleplatlalafen-
ced olive srovc, not observed after I 994i possibly adventive)
Teruiun niuophfllu^ Desf. - EB, RJ obs. C3. D6: EB obs- 46, E7 scattered
Lee ninosde
Anwtb foe,ido L. fa 9J-.l6.li C3-l; EB obs. C5 rar€
Arth lis hemotniaeL Re,htag4 l3oob: südige Hügel am KapTsuno (87' undauf
schrefer obs. (RFaHlNGl-R & RECHINoER-MOSFR. l95l: lable Il. d BORA-
TYNSKI & al., 1992: 24); EB, RJ obs. 87, Cl, C6i EB obs. 86, C4, C5, E7; RJ Photor C6-
l: RJ obs. E6 common
Asiagalß epislonis L. stbsp. epigloflk ER 96-tt1:86-3 mrc
New to Kriti.
Aslrasatus hanosus L. R & aS 47709: "Alonerhe road from the hdbour to Kashi" (BC6)l
ta 9J-J9i C6-l; EB obs. A4, C3, D5; RJ obs. 85; D6 - scattered
Asbagolus sinoats Boiss. - ER & AJ 94-I3l: B6-t. EB 95-t28: B7-2t RJ960101-61: C6-6;EB
obs. ,A.4 very scattered
Biü'ninatut bituüinosa (L.) C. H. Stirton EB, Rl obs. C5, E7 scattered
E. BERGMEIER & AL FLORA AND VEGETATION OF GAVDOS (CREFCE)
Cslicotone 'i M (Polret\ Link EBobs. AB5 rre
327
Centonh sitioua l . Rechinserob. ('cc dlso RLCHI\CI-R & RL( Hncl R MOSfR. ls5l
rl)ble ll.;dBoRATYNSKI&a1., lqq2 52): lB. PJ obs. Do: lBobs cS.(6.17 scar'
Chiefly but not exclusiv€ly found close to (fonnerly) cullivated places
Coronilla KoDioi.les tL.\ Koch Dörfler o0 _unrer Seren bei Ka\rn (C5). R & Bs 4__/r
'Alonq rhi rodd ftom rhe hübou' io K,srri (B( b). fA & 4J o1 5 5 : 1 6' l. L B l]5' t'' : C 6'
4; EB, RJ obs. C3, C5, D6; EB obs. C6, D5l RJ obs. 85, 87 - common
Donctiun hit\utun \L., Set Dö4et IA2\\t hüta (Jord & l-om r Riklir" Ka$rboden
'bei Sarakinil,o tB6)i I d AJ.l--lJ Along rhc rudd trom lhe hdbour ro Kasln lBC6):
CB & 4J 94-l2a Ch.4, RJ e40528-01 8ö-2; fB obs Cl. C4: RJ obi. 85. C5. D6 'cac
Hedßarun \Dinos|linud L. - R d 8-S 4--r,3i '.Along lhe road tiom lhe harbour to Katri
'(BCö);/iA /ooj Ji Bo-2. LB. RJ obs. C5. tB ob. A4. 87. C3. 17. RJ obs. C6. D6 rather
Hbpocte|is biforu Sprcneel R & BS 47758: "24lrtt S SW ofKasrd" (D5);tA 9J'JJj D6-
l; EB, RJ obs. Cl, C4, C5;EB obs. 87 rather common
HiDDoo?Dit .ition wmd Dö:,/c/ 8 Sandboden i R d Bs 4_8//i l_4 I'm Nw of Kasrri '
" tBC'45r, fA oJ-//J Cl]-'- Rl 0105)6-25. Db-4. LB. RJ obs. B?. C3. LB obs Aö. 86. E7:
Rl obs. 85, C5, E6 - common
Eunenocauos .ircinnatur (1., Savi - DorÄ, ori [ahlboden : I d aS r__// _Along rhe
" road lr;m rhe harbour ro Katn lBCOii l-B,RJobs C3, C4, Cs, C6; LB obs D5: Rlobs.
Lafttrus anphicaryosL. ER 95'88: C5-2; FJ 960102-19i C5-l; RJ obs D6 scattered
Lath!.us aphaü!L. EB 96-86:D56. RJ 960403-19: D64 fric
Lathttut.icmL. Dorf., J2i unterSaaren beiK6tri (5).Rda-S4 09l _AIonC rhe rcad
'riom rhe ha.bour r; Kästri (BCor: LA d AJ 04-13. 81 l, ER 95'03 o\'llJ: C3-1, RJ
940526-16:D6-l EB obs. Ce siatterea
EB 9r-t8 rcDresenls a \ anmr wirh darkaponed \alves whrch oc.urs ficquenrly in 6elds in
rhe areai rhe spors are hardl) 'ccognrzable rn dried mareflal.
Lalhyus hierc.otydilaius Boßs Dörfle,50 -\rnt{ Selen bei Kavri rC<r: EA 05_04i ( l-
I FR 96-N9r D56 scaured
328
Lathlrus s@otitis (vent.ryis. R & BS 47793:"Kasldi. The ravin€ atthe church" (C5)i R & aS
l786Ji "Särakiniko" (86); EB 96-44: A4-t very scaltered
Lathrrus setilolit s L. R & BS .t7792 : "Kaslll T\e .avine at the church" (C5)j RJ obs. C6
Lath!rus sphaericus Refz. ER & AJ 94-23: B6-2; ER 9J 6i 86 2, EB obs. C5. C6, D5 verf
z'eas r,r?ricaff (Bieb.) Godrcn- ER 95 87: Cs-2t ER 96-75: D6-tt EB 96-94: C5-1; RJ 960402
?0: C5-l scattered
Lotu. .lti\oi.lci I Dd,,Aet 9211.,rcli.us L. .ub\p. .rriordp\ (1., Bnq ) _Krßlboden;nd
AS.,t-^04i Along rhe road lrom rhe harbou lo Kdsri (BCö): A di AS,/-er/ _ rhe NW
poinr (A34J. la d AJ 0. lll CD6lt U a4n5''-Ao | 5-5; RJ aö040t //i ( o 4; t-8, Rt
obs. C3, E7;RJ Photo: D6-4; RJ obs. C5 scattered
Lo.us e.lulß L. EB. RI obs. C3, 85; EB obs. E7; RJ obs. 87, C5, C6, D6 nther common
Lotu\ halophilus Boßs & Spruner Do'lel..r. Düncn bcr Saml,rnilo rBor: Rechrngerobs:
''smdigc Nrederun8en (see also RFCHINGER & RE( HINoER-\4OStR, Iq5 l : r;bles I l,
28 and29)t R & RS 17833i "The Nw point" (A34): äB & AJ 91-16: B6-lt EB & AJ 94-
107: A4-la EB 95-34: B6-1; RJ914529 05: B7 l; EB, RJ obs. 87 scattered, locally he-
Lotus onithopodioides L. R&a517716:"wofKasrri"(C4);EB,RJobs.C3,D6iRJobs.
C4. C5 scatlered
Lotus percgi rsL. Döder1132:"Sndkn*e' E895-t:86-2,RJ960101-65:C6-6;EBobs.
C3, D5 scattered
Lotus tetogonolobts L. R & BS 47746: "24l<n S SW of Kästri" (D5); ,A 9J-64: D56. EB,
RJ obs. C3; EB obs. C6; RJ obs. D6 scattered
Medicago corotrata (L.) Ban'^|. Döllet tt45:"Felsen an Kap Kamärela" (87); EB, RJ obs.
Cl. C4, C5t EB obs. D5, E7; RJ obs- 85, 87, C6, D6 comfton
Me.licaso littototis Loisel. Dölnet t1(yat. brevßetaDc.): Dünen an derNordküste" (see also
RECHINGER & RECHINGER,MOSER. 1951: tabte 29); ER & AJ 94 65: C6 t. ER 95-
./ 72r 86-lt EB obs. A4 - scattcrcd
Medicago na na L. 1942 Rechinser obs. (see also RECHINGER & RECHINGER MOSER,
I95 l : Dble, 28 and 20r:,4 & 8.S 4 TJöi _Thc Nlt poinr'- I A l4i: n d 85 a-8-2. SaratF
nil'o (86): FB. RJ obs 86: IB obs A4 scarlered, Iocally fiequenr
E BERCMEIER & AL. FLORA AND VEGETATION Of OAVDOS (CREECE)
Medicaeo Minino \l )L R & 85 4"29. \\ ol Ka'ni (C4l: A d AS ai8ll Small smam
near the NW poina LA4); RJ obs. C5 rnre
Medicaso nonspelioca (L.rTtNtv. Döüet 1l (Ttisone a nonspeliaca L. f- trptca Eig.): \an-
diger Boden'; EB, RJ obs. C3, C4, C5;RJ obs. 85, 87, C6, D6, E6 common
Medicaeo o iü.Iafis (L.\ Ranal Dö41et 34 : "K^istboden bei Vathyana" (D56)i EB obs. Cl.
Medicaso pollnoryha L. Dö4{ 32 (M. hi,pidd GätlJ,. v^. lappacea (Desr) Hal.):
"Vathyana" (D56); ta 9J-?8: 86-2; EB obs. C3, D5, D6 scahercd
Medicaso nsidula (L., All. - ER 95 79: E']-l; ER 132:C6-4 rarc
The specimens my bc assigncd to var. 4arertr Bumat.
Medicago rugoso De* ta 9J ./r3. C3-l; EB obs. D5, D6 - scattered
Me.licaso iuncatula (Retz.) Willd. Dölflet 33 (M. rribuloides De$.jt "Kaßtboden bei
Varhlana tDsol.Rd8S4_-Jo _2 4kmS S$ ol Ka'rn (D<)rRda.S4_-0^i Ka'rn.
lhe ravrne ar rhe.hurch ((-r)i Ea oJ.ödi D5ö; LB, Rl obs. c5; bB obs. c3, F 7; RJ obs.
C4, D6 scattered
Our specinen represents the b"e vei€ty.
Medicago ilberculata Gaertner Dnä{e/ 6J: "unt€r Saaten bei Kastri" (C5)
Metitotus sulcatus D.sf. - R & aS 17804:"14kr,tNW ofKastri" (BC,15);rB 96-58: C6-5; RJ
960102-02: C6-5;EB ol's.87, D6 scatlered
Onobrychis aequideitata (Sm.) D'Urv. R d 3S ?7Zl "2 4 km S SW of Kastri" (D5)i tB
9J J8i C6-l; EB, RJ obs. C3; EB obs. C5; RJ obs. 87 scattered
Onobrychis capul-go itL.tIam. Rc.hth4ct /J66vi \ddrge Hügclm h.dp T'uno ,B?r: R
d aS 4--16 AlonB rhc roa.l fiom rhe härbou ro Ka'rr (BC6). Ä, sJ-l6 86-2r fB. RJ
obs. 87;EB obs. C3, E7;RJ obs. C5. C6, D6 rather common
Ononis hispa caL.l1l.stbsp.hispanica-Rechinserlj6l6i'BuchtSarakiniko"(86)Geealso
RFCHINGER & RFCHINGF.R-MOSF.R l95l: tables 28 and 29. and FORTHER & POD-
LECH, l99l: 251);,q d 3S 1770Jj "Along the road from rhe harbour to Kasti" (86)t R &
RS lZ8J4r "The NW point" \A34\: ER & UM 3520: R6t ER & AJ 94-110: 86-l; ER ohs.
A4, 87; RJ obs. C6 scatter€d, but locally frequent
330 caNDoLLEAs2, 1997
Oronis ornithopo.tioi.tes L R3 85478oi _SaEkinrt(o (86':fB0J-l-J Bt''2: LB A AJ
01-8l tö-2, RJ voulql-5li ( ö- l, l_B ob". C5 'canered
Onon| rcctinüa L Rc.hnget I Joo'c t var nollr Heldt. r. _sand rge H ügel am Kap Truno '
\87| R & BS 4--03 _Älong lhe road lrom üe harbour lo Kdsrri (BCbr: n d 8J 4-_8,
'Ka,rn. l hc ra\ rne ar lhe church' (a5), R d Bs 4_82r "Small srream nca the \w pojnr
(A4\Ea9589:C5-2;RJ94052606:C6-\;RJ940526-t8:D6-t.8J910527-01:86-5;RJ
940528-01: Cs-t. RJ 940529 06:R7-1 RJ 96010t -63r D6-3; EB, RJ obs. C3: EB obs- E7;
RJ obs.85. C4, E6 comnon
Omnis siebe DC. - R & BS l77rlr "Alons thc road ftom the harbour to Kastri" (BC6); t &
AJ o1. t33:81-L RJ a405)o-01: C6-l . Äe
Olotis eero. Sttt. Döi,flpl -Ji felsen am Kap Kamarcla /$!$hen Gebu\h l}l):FadlJ
o4--. Db-t:ERA5-90 C5-L RJ a4x526 0-:Cb-tt R-l0ö04rj-niiD6.l:EB.RJobs 87.
C4. D6; EB obs. 86, Cl, D5. E7; RJ Pholo: D6-3; RJ obs. 85 comon
Ononis visosa L. s\tbsp. bpi,t"'a (DC.) Nyman EB & AJ 94 79: C6-2; EB obs. C5 rare
Prrrru ratirrd L. subsp. sot un ER95-158:C3-l rare. esl.ablish€d weed
Sco.piutus drti@tus L. var mwicatus aa 9J /JJi C3- l: EB obs. C5i RJ obs. D6 - scatte-
S.o,piurus nuricatus L rar 1!ör'7,s!s (L.l Fion Dötfer R2 tS s,öv'rosh L ) "Kar{'bo-
den ber Kap Kamarela- : FB. RJ obs. ( l. C5. Do: Fa ob'. 8?. E?. RJ obs. 86. C6 com-
S?cunperu secud.hru \l .) Descn & Ddrfler Do.de, ?^ Kmlboden bei Kasm (C5,: R d
BS 4-'r4 -2 4 km S sW or Kdsrn (D5)l l-8. RJ obs. Cö. LB obs. D5. l-- 'caneted
Ttifoliun oNeis. RJ obs. C6 rare
Ttifotiun canpes,re Schreber Döder 7I:"Felsenaft Kap Kmdcli'(E7);EB. RIobs BT,
C3, C4, C5; EB obs. D5, E7; RJ obs. 85, C6, D6 - conmon
Trifoliut iqadibpone,tü Grctrter R&8547731:"WofKastri"(C4)iEB.RJobs.C3,C5l
RJ obs. 85, C4, D6 rather common
friloliatn scobru l . Ddatcr 0 -Dünen an der \ordküslc : R d 8J ?_'r- 'Along rhe road
lrom lhe habou to Kasm tBCör: l-8. RJ obs. B?. ( l. C5, Col l-B obs. 44, 84, E7l RJ
obs. 85, C4, D6 common
E AERGMEIER & AL. FLORA AND VECETATION OF GÄVDOS (OREECE)
Ttilotiun ncllotu,t L. Dötict tl31 Ka\lboden bcivarhyana (D5br:tB.RJobs Cl a5:
EB obs t7l RJ ob,. 85, Cl, C6, D6 common
hifoliud stfoca,uü L. - RJ 96040t -40: c6-l; EB.RJ obs-c4,c5;EBobs A4, c4 scatre-
Trifolisn tonentosuü L. - RJ 96040t-11: C6'|EB.RJ obs C4, C5, D6; EB obs 86, C3, D5
raüer cotrmon
Ttho"dtd spitFa L Dold., //JJ äufsandboden:RdBS4--2? w olKastri r(4i:ta
o6-tg C(,-a RIq6O401 4):C6-1 mrc
Ttipodion t taphlllu'n tl t r oun. - Dö4Ier 8 l t PhNsanthr is ktraph' a (t 't Bo irs ): "Ka Fr-
boden : EB obs. C6i Rl obs. D6 r2ß
r,r.iz.'zr'.a Boßs. & Heldt. Ddrnet o- tv s,tunetiBoiss.,: _Kasni ( 5), n d AS J--7ii "l
kmSolKasni (DsJtRd A"S,t--8t,i "Kastri The ravinc ar the chuKh (C5t:E896'Jl
87; EB, zu obs. C4, C5. C6, D6;EB obs. 86, C3, D5, E7 cotnmon
lncialabaL.l EB, RI obs.r cultivated, locally escaped
ncia hrb.ido L. - PJ obs. Cs-t tu.
nciaDadifloruCa\. lR& AS4-o8.l Alone rheroad ircm lhe harbour ro Ka\tri (B(ö'.4&
äS a=8roi -The Nw poinr'' l{l4), f,B 0t-4: 86-2: EB. RJ obs Cbr EB obs. 87. CJ. C4.
C5. D5 rather common
yicia percsÄno L. Rl960402 05: Cs-l nrc
W.ia satiea L. srbsp. anphicarpa \L ) Ban. RJ a4052ö')o D64t RJ ob010l'ta C6'l: RJ
9ö0402-06: B5-)t RJ 90040)-ttt C\-tt PJ oo)41J-ooi D6-4, RJ obs Cs, D6 scarlered
Confirm.fion for the Cretrtr ar€a.
t1cia etiva L. subsp. c,'/4.4 (Hoppe) Asch. & Graebn€r Düfer 51:"Kastn" (C5tt RJ
96040t-56: C6-l - rarc
The following records of Z r.rit4 cannot be assigned wilh cerrainty to any subspecies: tB
& AJ 94-22: 86-2, EB 95-6J: C3-li t 9J-99: D56; EB obs. C6
Linaceae
Li,'un stictun L. var. spic^tud Peß. - EB 95-20: 86-2; EB, RJ obs 86, 87, C3, C4, C5. D6:
EB obs. A6, D5, E7; RJ obs. 85, C6 common
331
332 caNDoLLEA 52. l99i
Lythrue@
Lythrun hyssopilolio L. ER 96-83: C6-6; EB obs. A4 87, C4. C5 scatt€red
Mahaceae
Molva aegptiaL- tA 96-.lJr 86-4; EB obs. E7; RJ obs. C3, C4, D6, E6 scatt€red
Motvo cretica CAv. s\tbsp. crctica Dörflet 1146: "Felsen am Kap Kamarela" (I7); EB obs. E7
MaIw paniflonL. R & BS 47748: "24kms SW ofKastri (D5);tB &,1J 91-36:C6-t RJ
960401-30: C6-1l' EB obs. C5, D6 - scattered
Moncede
I[icus conca L.) - EB, Rl obs. C5, D3 scaler€d occunenc€s probably all du€ to cultivarion
Oleaceae
Otea eurcpaea L. var. srh'estris (Mill€r.i Lehr EB obs. 87, C5 rare
Young pl tations ofthe cultivated taxon in 86 and C6.
Orcbahchaceae
Onbaache puhesceß DiJ'nt.-UN. ER 95-148:E7-l; ER 96-72i A4-l; EB obs. C6 scatr€red
Orobonthe runosa L. "ubsp. nut.tii tf w. Schukz) Cour.- Död.r J8 (O rlrel, F. Schulu I
rpitsa Becl,). "KarsLboden bei Kasti \Cs)t Döfiet IIa4{A urrp, L sp6rd, felsm am
Kap Kamareld (F7):RdaS,t-8lni"l 4 km Nw ofKastn (Bc45); Eaq6--^i Do-J (on
Chrysanthenun comnanuu), EB obs. C6 v€ry scattered
Orcbarche mnosa L. stbsp. rara (Reucr) Cout. RJ 910529-07: B'7 -\ we
Otdlitlaceae
Oxolit pes-cqne L. EB obs. 85 (1994, one pldt)! EB. RJ obs. C6 (1996, nuerous under
several oliv€ lle€s in a plantation) rare neoph)tic weed, but spreading
E BEROMEIER & AL. FLOKA AMD I'EGETA'IION OF GAVDOS (GREECE) 333
paptueruceae
Funatia noc.ocarpa P^n. - ER 95 102: C3-t we
Funaria pa illoru Lan. EB 95-134: C6-4t RJ 960402-0t: C6-5 ffie
Hypeco!6 pto.tnb s L,ubtp. pto.unben\ nd 8-t ?-000i _Along tle road fiom rhe har
bou! to Kasni- (B( 6), EB ob ö3: C'-lt LB 06 102: C i-lt RJ oon4q)-08 ( 4-2 tue
PapoethybtidunL.,R&rS.r7857j"Kastri"(C5\;Eß&AJ94-54:C6-t-8896-68:C5-1
RJ 960101-32: C6-laEB obs. C3. D6 scattered
Papotet pt tpurconargittotun Kadercit - EB 96 36: A4-l we
Papowr thoeas L. Rd BSIZ8J8: "Kastri" (C5);EB, RJobs. C6;EB obs. C3, C5,D6 scar
Roeneia hJbrida (L.\ DC. Dötfut I t47: '.,rntet SBten bei Kastri" (C5)i tB 96 80: C6 5i EB
96 104: C1-t we
ptdhtdgihaceüe
Plontaso afta L. - Dörfü 1140(P psy iunL.):"Küstbode,": ]? &rSlz682i "Along the road
from rhe haöour to Kastri" (BC6\t EB 95 19: 96 2,E8, RJ obs. C6, D6; EB C3, C5, D5;
RJ obs. C4 tather common
Plantago albica sL. ER & AJ94-7: B6-2 EB obs. E7; RJ obs. C5, C6 scater€d
Plairqgo be otdii All. R & BS 17752: "24 \Jn S-SW of Kstri" (D5); EB, RJ obs. C3; RJ
obs. C5 scattered
Planugo lagop!\ I Döl,re,2_ _Karrlboden be' Vath)and (D56).nd 854__J2 u ofKa$
fi tC4,: LR & AJ e4-53 Co-lI I B. RJ obr. C4. ( 5. Do. EB obs. C3. D5. E7r RJ obs. 84.
85, Dö common
PlantaEo lüceolata L- ER&AJ9435:c5-t we
Plahtago squat.osa Mwmy Dötfct I t a8: "Dnnen M der \orclhjsre. ma5senha fi ; lRecrzser
/J048 sdndstrand an der Buchr Säralinilo- (86) (see also Rl-CHINGI-R & RL( HIN-
GER-MOSER, l95l: rable 28); R & 3S l7837i "The Nw poinr" (A3). R & BS 47862:
"Sarakiniko ' (86); tB & ,4 J 94-98: A,4-l ER 95-125:86-1;EB obs. B7 scattered, locally
334
Ptontoeo act.l.nii Rctchenb. 'nbsp. Det.lenü Dörjct l15\"P.undopus r: -Ka^lboden:R
.0-A\ r78J9, _sadkrntko tBn\.ER95'152 t7_liLBob' 44,C4.C5.RJobs B:l' C6 -
scatte.e.l
Plunbaginaceae
[ino iun arei (DeNot.)Brullo & Erben) BRULLO ( 1988) regards I e.rtorires a Wesl
Mcd'ienanean raon wh'ch rs reDlaced in the Cennal and Fdsr Med'rerane b) I dv(i Tlus
onlv I ar.r ß ui\en lor Greece: bc'ide marenal ol Anili lhere ß one rccord fiom Catdos quo-
ted'b! BRULL-O ( la8br. The iource ol rhis i' nor "pecrfied but i. almosl cenarnl) bd"ed on
Rc.hinper thaJ,hich tepresenl' rn f&l t. e, &i,r../esil-RBtN. in lrfi ' Accordints lo BRULLO
(la88t and FRAF\ (1978) / drer i' characrerr/ed by mu.ronare leavcs änd prlose calvces
unhn, mmro while üe leates ol1. echio1lps ncit}]'et have acumen no' muclo, rhe calvccs
be'ne slabrous or subqlzbrou' $irh lonq-e\serled unc'nate mucro lhc spe.lmens collecrcd b)
u. oä öa'ao" mre ,t'"-'"t o' 'häracteF -ry11rcal oi l e.rto,dP\, and üe same h nue for all mare-
aal hom rhe Creran ruin i'land collecled b) RJ and A \4aler {see MAYI-R. Iqq5) a' well a'
lor rhe rollowrns {pecimen\ 'een by fB trom rhe SouLh Aegean: _Kreta. I asithi. fp Siria: Ira
mo. rl5"l6.N r6dlö Fi. 6 4 t984. Rax( o//_ lC,t -Käso,, lnscl Annarhia. Srjdosrtüsre
r!5'25 10N2ö.52.t0'Ei.t94t98t.Adh-5oorC)i-KarDalhus,südo(küstenördl.Ammopr
rl5'28 40 \ 2-'12 30 1i , 2l 5 1q84, R4!r 9804 tcr' Hcnce. the occunence oll a,ei for rhe
Cretan area as gi\en rn Med-Chccllsr rCRl L ll-R & ä1.. ls8or srtll remains lo be proted.
Litiot iun cre.icun AnetaÄ ER & AJ 91-l t5: cD67 (det A. May€r) mrc
Lihoniud echioites (L)Millet Rechi get 13613: 'sardstrand an der Bucht Sarakiniko':(Bo
lsce älso RECHINGER & RECHINGER-MOSER. l95 t: table 28): ER & AJ 94 23: B6-2i
ba a At c,t-t1s, Iq-l RJ 940526 09: c6-6, N 940526-Jli E6-l; EB, RJ obs. 84. c3'
C5, E7; EB obs. B7t RJ obs. D6 mther common
Linot iun etaphonisictlt t A. M^yet - EB & AJ 91'106r A4-l (det A. Maver) - rare
Linoniun Fruecun lPot el) Rech. fi|. subsp. ßtz"art RectuhAa 13c13 _S€nd'rrand a dct
Bucht,-sadkmrko {86,{seedl'oRFCHINUI-R&RICHI\GFR_MOSLR, la5l:lable28):
R & BS 44-850 _The\\{ potnl (A34r.nd aJ4_3'/ _Sarakrnrlo tBnt. EB & LV
.l5??r 86 (dct. A. May€r)i tt 9J ./19i B3-l (d€t. A. Mayer). RJ940528 02: B6-l scane-
Polfgalaceae
Poltgala monspelia.a L- RI obs. C6 rdc
Potvsala te\utoso Sn. Dörler _8 ^i'chcn Buschserl am Kap Kdndrela (F7': ad tS
- ".rZZoO _: +t.S-su oiKJstri tD5,ifBoJ_oJ Db I;FB,RJob' Cr:LBobs Aö.8'
C4. C5, 87, Rl obs C6 rather common
335
Polßonaceae
E ex spinoso (L.) CMpd. - EB obs. A4, C5, D6i RJ obs. 86 scattered
PotysoMn naitinnn L. Re.hi"gd 13642: "SffiJsllar'd an der Bucht Saraki'iko" (86) (s*
also RECHINGER & RECHINGER-MOSER. 1951. tables 28 nd 29): EB. RJ obs. 86. EB
obs.,A4, E7 ree
Runex bu.ephrtophotus L. subsp. a"84"l.r Rech 6t Dörflpt 5't R. bu,ephalophoru; r
'Kmt. and Kuhuboden-, EB obs. 44, 86, 87; RJ obs. C4; C5 - scanered
Rune, pulchet L. s\tbsp. o'odo'tus (Hausskn.) Rech. frl. ER 95-97: C3-t we
According ro TURLAND & al. ( 1993) there is no published record for th€ Cretan area since
1930.
Runü pakhetl.slbJp. woor'sri(De Nor.) tucangeli R&8541-65:"2 4kmS SWofKa'
ti \D5| EB & AJ 94.'b: D56, EB obs. Dö scanered
A field record from C6 cmot be assigned to a subspecies.
E. BERCMEIER & AL. TLORÄ AND VEGETATION OF CAVDOS (CREECE)
Pinulaceae
A^aa.allis aneßis L. \ar. caerulea lL.) Goru r ER95 11: B6-2t R-I940529-0.r: 87-l: EB, RJ
obs. 86, C3, C5, C6, D6, E7i EB obs. A4,46, C4, D5; RI obs. 84. 85, E6 common
Asterclinon liiun-stelhtttn (L.\ Duby Dirfet 1119i "Ka6tboden"; EB, RJ obs. 87, C3t EB
obs. A4; RJ obs.86, C5, C6, D6 common
Rallesiaceae
Ct'tiius htpocktk (L.\ L. stbsp. crr.itt Nyman RJobs. C6-l (on Cßr'6 creltc,r) - rare
Cttilus hrpocittß (L.\L. subsp. ori."dlrie Wettst. - Dd?tel J0.lj "auf Clrrüs am Kap Kamarela"
(81); ER 95-t30:87-2 (ot C6r6panforu);RJ obs. 86-5 scatt€red
Ranutelaceae
Adonis niüocaryaDC. s\tbsp. crenc, (Huth) \4erh. Dö.fet 20 ("A. dpaniana")r auf Kaßl-
bodetq R & BS 17757: ','24 km S SW of Kastri" (D5); EB & AJ 94-59: Cs-t: RJ 940526-
22: D64i Rl960402-07: C4-2 - scatt€red
3:t6
An.none corcnaio L. ER 96-53: C6-5: RJ 96040 t 15: C6-t; 8J 964403-2Ai D6-4t EB. RJ
obs. C5t EB obs. Cl. D6 scatter€d
tR 96-JJ represents the common vdimt with violel-blue petals which is by far mor€ fre'
quert on Gävdos tlan plmts with scarler petials (var. co.tined (Jordan) Bum.), th€ latter were
obsewed as a segcttl weed in C3, C5, D6.
Clmans cinhosaL. R & a.S.lzz83i "Kastri. The rÄvine ät lhe church" (C5) (s€e also BORA'
TYNSKI & al., 1992: 63); EB obs. C5 rare
Nige\a danoscena L. n & tS.l778Jr "Kastri. The ravine at the church" (C5); EB obs. C5
NiEeIa doernerivieth. R & as 47736:'24kmS-Sw ofKasrri" (D5);R d as 4777t: "3 km
S ofKastri" (D5); Aa & .U 91 50: C6-l; ER 95-7:86-2i EB 95-82: E7-tt RJ 940526 17:
D6-1t Rl 910526 26: D6-4 RJ 960101-13: C6-4i EB obs. C3; RJ obs. C4 - scättered
Ranmcutus asiaties L. \ar. at 4s Hayek dtkl 66: "auf Kaßtboden bei Kastri '; ,a & ,.1J
s. n.:C6-11ER 96-21:B'7-1, N 96010t-36: Cb l; EB, RJ obs. C5, D6iEB obs. 86;RI obs.
85 scattered, locally abundant
Ranua.ut!\ ßiati.us L. \at sa,s,iu"&r (Mrlle4 DC. Dirllpt I \1. puhh cu! Dötrict ): ^tletber Ambclos (( 3), LB, RJ ob,. Do: l-B ob). c I. C5. D5 lcD 'cdncteJ
Ranlnculus bunatusL. k|960101-53: C6-l:EB.Rl obs. C5 scattcred
Ranunculus creticus L. EB obs. C5 rare
Ranüficulüs lcüia L. s\tbsp. .httsocephalus P.D. Sell EBobs.C3,C5,D6 very scatt€red
Ranunctlus poludosus Poirer Döflet 112 (R. fabe a|'r Dest): bei Xenakis (D45). EB obs.
C3. C6, D5. E7i Rl obs. D6 scatlered
R.seda.ede
Reseala lurea L. subsp. lr,sd Dö,fiet It6: "K\tltrtbode bei vathyua" (D56); Recr,rael
1J667di 'sandige Hüsel am Kap Tsuno'(87); Ä & ,S 47698i 'Along lhe toad fron lhe
harbour to Kastii" (Bc6); n & BS 47712:"24kms sw ofKasbi" (D5); tB &,JJ 9.1'JJi
C6-li RJ Photo: C5-lj EB obs. C5, D56 scattered
Rese.la odo.ota L Dö41.- ^o t'R o4ptrtdrs lvülls' Boiss. r: -Kaßrboden bei Varhyana'
(DsorrRda.S4___,l Kann. Tte ra\ine dl the chutch (C5\iER&AJa4-a c6-l:Ea
& AJ 94 89. A4-lt RJ 910529-09: B7-la RJ 960105-01:B7-l; EB obs- C5 scattcred
E. BERGMEIER & AL FLORA AND VECETATION OF CAVDOS (CREECE) lt7
Rosaceae
(Rosa cdniaa L.) is eEoneously siven for Gävdos in BORAT"INSKI & al. (1992:211)
Sangl.isofia ninot Scop. subsp. 'e"ücaso (Decn€) Holmboe ta 9J 3/i C6-l; RJ obs. C5
Sarcopote,iud spinosun (L-) Spach EB, RJ obs. C3; EB obs. E7; Rl obs. C4, C5, C6, D6 -
common on limestone
Rubiaceae
Aspenla isido Sm. Rechikset 13664:"sandige Hüsel am Kap Tsuo" (87),R&BS47717:
"w of Kast.i" (C4)i t & ,4J 9.1 -?/i 86-2; EB, RI obs. D6; EB obs. A6, 87, C3, C4, C5,
C6, E7; RJ obs. 85, E6 conmon
C ruciane a hnfoüa L. E R & AJ 9 4 -7 2 : D6-l. EB, RJ obs. 87. EB obs. E7i RJ obs. 85, C6
Galiun opaÄneL--EB,Rl obs. C5, C6 scatter€d
Galiun sraecun L. srbsp. sroecrn R& BS 47782i "Kastri. The ravine at lhe church" (C5);
EB obs. C4. C5; RJ obs. D6 very scatter€d
Galiun nurale (L.) All. Dödcr 39: "Kaßtboden bei Vathyana" (D56)i 4.,/960401-10:C6-4.
EB obs. A4, C3, C4, D5j RJ obs. 87; C5: D6 mther common
Galium setaces L^n. R & BS 47770:"3kmSof Kastri" (D5)l ta 9J-]0i B6-2i EB. RI obs.
B7t RJ obs. D6 scatt€red
Galh.n t conutun Dandy EB 95-136: C64: EA 96-82: C6'6 ftrc
Goliud ven cost n fi)dson ER 9693: D56], RJ 9644ß-07: D6-4 nte
According to TURLAND & al. (1993) there is no posr-I930 record for Kriti.
Sherardia arunsis L. - EB, RJ obs. D6; EB obs. C3, D5; RJ obs. C4, C5 rather common
ualantio hispi.Ia L. Dö:,/e/.16j "Kaßtboden bei Vathyana" (D56): tB 9J-7r: C6-l; EB, RJ
obs. 84, 87, C4, C5, D6; EB obs. 44, C3, E7 common
valat tia nünlß L. ER 95-42:C6-1. RJ 96010l-/Jr C6-4; EB, RJ obs. D6; EB obs. ,A.4, 84,
87. C3l RJ obs. 85. C4. C5. E6 common
338 CANDOLLEA 52, r99?
Rutdceae
k!,a chaLpensis L. silibsp. cholepensis R & BS 47800: "Kas6 "the nvine at ihe chuich"
(C5\; ER 96-70: C5-3 - we
Santal4ceoe
Thesiun berssizü.c. EB obs. D6, E7; RJ obs. 86, 87, C6 - scane'€d
Süiftasaceae
Soxiftasa bidactt'lites L. - EB 96-97: C54 tue
Sctophulatiaceae
Belardia tixago (L.l t\ll. RJ obs. C5-5 rare
Kbkxia ela.ine (L.) D\mc t subsp. rie6e', (R€ich€nb.) Hayek - ER & AJ 91'25: B6t; ER 96
96: C54] RJ 940528-06: C5-lt EB, RJ obs. C4 scatter€d
Lharia chalepensit (L.)Millet ER 96-81: C6-6 tM
Linüia üicrontha (cav.) HofRnams. & Link EB 95-131: c64: RJ 960401-28: c6-1 - ßrc
According to TURLAND & al. ( l99l) there is no published record for th€ Cretan ar€a since
1930.
Linada pelisse a"a (L.) Mlllet Dö4a I I 36: '\mrer Saalen bei Kastri" (C5)
Linaria sinple.x (willd.)DC. ER 96-27: B7'2, RJ 940529'04: B7'tt RJ 960401-29: C6-l
Lina a tiphrua (L.\ Millü - Dödet 48: '\ritü S?d'telbeiKaslJi" (Cs)t EB 9 5-5I, 96-91 : D56:
RJ960403-08: D64 rarc
Accordirs to TURLAND & al. ( 1930) theie is no published record for th€ Crelan area since
1930.
Misopates omanu lL.) Rafin R& AS4TT9l "Kaslri. Tle ra\ ine al rhe church rC5,:EB
lJ-9i B6-)i FB, RJ obs. cÖi FB obs. E?, RJ obs. 87 scaftered
PoMt nccllia larifolio (L.) Caruel EB obs- C4-l -rare
E BERGMEIER&AL. FLORA AND VECETATION OF GÄVDOS (GRTECE) 339
yefiascum sinuotuh L. EB, RJ obs. C5;EB obs. 86 very scattered
Vercnico cynbolaia Bodard EB obs. C4, C5 - very scatüer€d
Vennica pol aFnes ER 96-60: CS-l -ftrc
Solanaceae
Hyoscranus albusL. EB & AJ 94- 109: B'7 -1; EB, RJ obs. C5 rare
lLycopeßicon * eitun Mill€rl EB obs. culrivated and as rare garden €scap€
Manbagoru autunnalis Bertol. - RJ obs. C6 rare
Theligonaceae
Thelisonun cynocrunbe L. A & BS 4782Ji "Small stteam n€ar the NW poinf' (A4); EB obs.
A4, C5: RJ obs. D6 scattered
Unbelliferae
Biforu terrielato (L.) Schultes Ddl/le' Jr'i "unter Smtcn bei Kastri" (C5);tR 95-9Ji C3-1
Bu tnletulaceun Sibth. & Sm. -t 9J-62: D56 rare
Aupkurun sau.Iiotun Snogetup R d aS a--JJi 2+ Lm s SU or Krsrn rD5r: A d aS
4-8J4i l km \ olKasrri (C5) (SNOG| RUP lo84): fa oJ-,ti ab-2r la d 4J 04 61 D6-
l: ER 96-95: Cs-4; RJ 940526-13: D6-l; N 940528-07: C3-2t RJ 960401'01: C6-lj R'l
960103-02: D6-\t EB,RJ obs. C3;EB obs. C4, E7; zu obs.85, C5, D6 rather common
Eupterun latciloliun Homen \*. tonsiloliun \Dest., Hayek Do.de, /// unrer Seren
bei Xenakis (D45)
&uDl.utum semi.odposi.t n I Rc hnp4 lJ6oo: '{ändige Hügelam Kip I'uno tBTr( tar
' Elaudn \Rob & Ca.'.) \.\olf), R d BS 4780Jd 't km Nw or Kasni (C45r: R d 8J
,-85J 'l km \ Kastri (C5); R d 8S ./-86? _Samt(inrko" (Bb)t LB & aJ a4 l2: Bb-ll
ER 95-22: B6-2; RJ 910526-l.r: C6-6: EB obs. A4, 87, C3, C5; RJ obs. C6, E7 scatlered
Criflrnuddo.innu L. RJ obs. E7 .ar€
Dauus guflab.s Sm. R & BS 17755: '2 -4 km S-SW of Kastri" (D5); EB obs. C5 rare
Darcus inroltcratus Sm. R&8547755pp:"24km S SW ofKasbi" (D5); EB obs. C3, D5;
Rl obs. C4, C5, C6, D6 - scattered
Eryryiun natitihun L. EB, RI obs. 86 rare
Fo.niculun wlsarc Mill$ subsp. pie"trd (Ucria) CNt. - Rechinset (leg. Herzog) 13673:
"Brachen b€i Ambelos" (Cl); EB, zu obs. 86. C5, C6 scattered
Losoecia edifioittes L. Dötfet tt51: "t<aßtboden" (sec also RECHINGER & RECHIN-
GER-MOSER. l95l: table I l); EB, RJ obs. C3; EB obs. D5, E7; RJ obs. C4, C5. C6, D6
PseadorlaJa punila (L.) Gmnde Dö41et 23 (Daucß pmiiß (Gou.) Ball.): "Dünen bei Sarä-
kiniko" (86) Gee also RECHINGER & RECHINGER-MOSER, 1951: table 29); Ä & BS
l78.t8r "Th€ Nw point" (A34); aa 9J J6i 86-l; EB, RJ obs. 87; EB obs. A4 rar€, but
locally frequ€nt
Scolige a napilornis (Sprensel) Grande EB obs. C4, D6 rare
Scandix aßtralis L. subsp. aüs,tatit Dö,fet $ ("srbsp. ga ico Vi€rh-'): "KaßtMen b€i
Kaslni:ER95-133:C6-4.ER96-79:D6-t.RJ96010J,.liD6-l;RJobs.C4,C5 scatt€-
Sca ix pecten-wneß L. Dö41et 1t 3: "K',ln\bodet"i t 95-l7i C6-4;EB obs. C3, C5, D6
Totdytiun aplud L. - Dödel 99: "Karstboden an d€r Südhist€"; Rr 96r401-l.li C6-4; EB, RJ
obs. C5, D6; EB obs. C3, D5; RJ obs. C4 rath€r common
r"'/'i ,,'iri' (Jäcq.) Lassen lT leprcphlta dßt.) R d aS 4'-?8i w ot Ka'tri rcar: R d
8J r_-J6 _2 4lT S SU ol K6rn-(D5); AA 0J /JJi C6-4; EB, RJ obs. C3, RJ obs C4.
Nomenclature follows P. Lassen (i. RAUS, 1996)-
Torilis nodosa (L.\ Gaettner R & BS 47756pp,4776J: "2 4 km S-Sw ofKastri" (D5); tB &
AJ94-61: C5-l.EB obs. C3, C4, D5 scatler€d
Ufticaceae
Paietaia ue,ica L. Döüet 75:'.Ielsena'J,Kap Kamarela" (E7);EB obs. C5; RJ obs. C6,
D6, E7 scattered
anica piluliferaL. EB, RJ obs. C4, C5, C6 scattered
Udi.a o.'r L. - RJ obs. 84. C4 scattered
E, BERGMEIER & AL. - FLORÄ AND I'ECETANON OF CAVDOS (GXTECE) 341
t/aleianaceae
Centra^rhus calcibapae (L.) Dufresn€ srbsp. colci.ropoe Dö"te. ./8i "Karstboden b€i
Vaihyana" (D56);EB obs. C5; R-l obs. C4 scatler€d
Valeiarclla corcnota (L.) DC. - Düfet 91: "Kaßtboden an der Südküste"
VatedMeua discoidea (L.\Loisel. Döüet 1142: "lrt€r Saaüen bei Kastri" (C5); Ä/ 96040]-
7J: C6-l! EB obs. D5 - scattered
tat.tiü.lla ni.mcoryoLoisel. R&ASr''824i Small ,tream near the Nw poini (A4): R d
8s 478?8 fie Nw pornr" \A34), Ea 06-40: A4-l -ftre
Valeiane a nüi.a.a Loisel. ER 96-tt0: C4-1 rzrc
Yaleiaaela punila (L.) DC. EB 96-64: C5-r -.arc
N€w to the Aegertr srea.
ValeiorcIa eesica a (L.) l\tn"nch - EB obs. C3, C6, D56 scattered
MONOCOTYLEDONES
A dly iddceae
\arcitsu\ tozeaa L. - ER oJ-og. D56;LBobs C5 rare
Pancrotiuü naÄttnun L. EB. RJ obs. 86 ffe
Anerun üllCarc Taß.-"tozz. EB, RJ obs. C4, C5, D6; EB obs. A4, 87, C3, D5, E7; RJ obs.
85. 86, C6 comon
CypdKue
Cü6 haleiaM Asso EB & '4J 91-75: D6-l; ER 95-115: D6-l lare
Car6 ilegitina Cesati, ER 95- 146: D6-t - ne
Schoenus nisticans L-- ER & AJ 91-90: A4-t we
Grunikede
lesilops biuncialis vts. R& BS4--54 2 4kmS Sw olKastri (van SLACIRIN. lsc4'
156 and disrriburion map.): Ea & 4J 04'0: D6-l: EB obs. b7. RJ obs. Dö 'cafiered
Aesilopt sdielata Roth R & BS 17692:"Alongthe road fiom the harbour to Kastri" (BC6)
(vu SLAGEREN, 1994: 238 and distribution rop)
Aailo$ naÄsmfri \Ge\r et Hammer Rd a-S4"2Ji-wol Kastr' (C4){r,nSI ACER}N.
tas4. löö, 7? i:d"/ara L . dol ercneousl) omfted rn distriburron mapt EB & U 04 6a:
D6-lt EB obs- C3, E7 - scatered
Aelutupls lagopoides (L.) Thw^ires EB95 76:E7-lt RJ 960403-10:E'7-1 rare,locally lie
Aba elegrntissiha Sc.nv - R & BS 47807 : "14 vrr,}Iw of Kasri" (8C45); EB, RJ obs. C5; RI
obs. 85 - scatoered
Atubopogon .listochyß L. EB obs. C5-3 rar€
lvelinia nichelä (S^vi\ Pül - EB & AJ 94-124: C4-2; ER 96-65: Cs-t:8J 960102 04: Cs-ll
EB obs. 87 scattered
AreM bübara Liak stbsp. barbata ER &AJ94-t26bi c6-4; EB, RJ obs.87, C3, D6;EB obs.
E7; RJ obs.85, 86, C4, C5, C6 common
lAeena bt'zontina C. Kochl - ER & AJ 94-56: C6-t cultivated 6 field crop dd rare weed in
fallow fields
E BERGMEIER & AL. FLORA AND VECETATION OF GÄVDOS (GREECE)
A'era stetilis L. s\tbsp. sterilis - ER & AJ 94-126a: C6-4 - scanered
The following fi€ld notes de assigned to tle species: EB obs. C5; RJ obs- 87, C6
Btachypodiun distacl'yon (L.) P. Beauv. R & BS 17744: "24 kmS-SW ofKastri (D5)! ß./
i60102-tid: c3-l EB,Rl obs. D6; EB obs. A4, A6, 87, c3, E7;RJ obs. 86 rather com-
Bntu dtidaL. EB. RJ obs. C3i RJ obs. C4. C5. D6 scattered
Rtuüus alopecuns Poi.et ER 95-106: C3-1 tN
Arudus tosckulatu. C. Pftsl Dö,flp, _o Kap Kamarelä tl-?)t Dötfl?t 35 'VÄlhyana--
(D5i,; ta oJ JJi D6-l; Äa on-jai B?-2 tBoth specrmens ol which rhe former $as kindl)
rdenrrfrcd bv Prof H. Scholz represenl rrn! planß w'lh unusuall) small 'Pilclet( änd
zlume!, N' o^o'u t 16 Ctr-l : RJ aon4oJ U I : b6) t lB. RJ obs. C4. C5. FB obs Ba. 87.
C3. E7i RJ obs. 86 - common
Bronus iitennediüs G\tss. EA&AJ91-19:87-t:EB&AJ91'80:C6-2.E89521:86-2:EB,
R-J obs. C3: EB. obs. E7; Rl obs. 85, C4. C5, D6
BnnLt nadfidns L. srbsp. na.t tensis EB & AJ 94-81 : C5'1; EB 95- I 7 : B6'2t EB obs-
C3; RI obs. C4. E6 scattered
ßtot'us rubens L. R & ,S 4768.1i "Alons the road from the hdbour to Kastri" (BC6)i Ä & aS
42799i "Kastri. The nvine at the church" (C5); tA & -4J 94 11:87-l; ER 96-72: C6-l; RJ
910526 02: C6-1,EB.Rl obs. C3,C5; RJ obs. E6 scattcred
Brcnus scopttits L. - ER & AJ 94-60: D6-t -rlre
Given as "before 1930" record for Gävdos by TURLAND & al ( I 993: 379) althougl rher€
is apparently no literature basis fot this.
Rtoüls st tilis L. EB obs. C5 rare
Ero us tectorumL. EB96-92:D56 rre
Carapodiun natibuh (L.) C. E. Hubbard ER 96 23:87-2:ER obs. A4, A6, E7 scatteted
(atapo.tiun iei.lun \l .)C. E. H'nbb^td Dörict 38 tS,/e&poa,rarld (L r(ms I _KdrsLbo
den i Ea oJ /r)ri cl-r; RJ 940520 23 D6-4, u 0105)6 J0: f6-lt RJ 060401 0a: ('64i
RJ 960101-47: C6-1.E8 obs. C6, D5, D6; RJ obs. 85, B?, C4, C5 conmon
343
344
Cuttndia daritino (L.) w Barb€y Rechingel t3654: "Bucht Sarakiniko" (86) Gee also
RECHINGER & RECHINGER MOSER, l95l: table 28); A & RS llTSJrr "The NW
point" (A34);tB 9J-3?. 86-1, EB obs. A.l mre, locally ftequent
Crnodon doctrlon (L.') Pets. ER 95-122: B6-4 ßrc
Dacolis gloderuta L. s\tbsp. /,acfrrrit (Ascb. & Graebner) Ciferri & Giacomini tA & ,4J 94
l16: CD61 -wc
D.tcülis gloneruto L. srbsp. /,isparrdd (Roth) Nyman EB, RJ obs. C3, C4, D6; EB obs. D5,
E7; RJ obs. C6, E6 common
Echinoio capirota (L-) Desf. ER 95-52: D6-l. ER &,.U9t-51r C6-lj EB, RJ obs. C3;RIobs.
C5 - scattered
EWiSia junceo (L.) Nevski RJ 940528-01 : 86-l :are
Gasbidiuü phleoides (Nees & Meyen) C. E. Hubbard ER &AJ94-83: C6-2; Rl940526-15:
D6-l; EB, RJ obs. C3; EB obs. A4,86,87. E7; RI obs. 85. C4. C5 common
Hoüeu leponnun Link.\bsp lepoiwn Dörfpr Jl (H nuinun t. stb,p. lcpotlhuh
tlrnkrAsch&Gr.r:_Kaß'bodmberVathlana-(D56),RdBS4---a "Ka\tri. Thcra\rne
ar rhe church tc5r:tadltJ04-.6:c6-ltuo^n4ol-)u t6-l; Rl ob040l-5-:ctr-l,LB,
RJ obs. C5j EB obs. C3j RI obs. C.l - scattered
lHodeun wlgarcL. srbsp. t'ulgarel EB& 9442a:C6-ltER95 151:C3-l cultivat€d
as 6eld ffop and subsponlaneous in fallow fields
Hyparüenio hi a (L.\Srzpl Rechihset 1 3 6 5 3: "sandie€ Hügel am Kap Tsuno" (B7) Gee also
RECHINGER & RECHINGER-MOSER, l95l: table I l); EB obs. E7j RJ obs. 85, 87, D6
Laalr(s ovatus L. stbsD. oeao^ Döfiet 1t4: "Kaßtboden"; EB & AJ 91-47: C6-|,EB,PJ
obs. C3, C5; EB obs. ,A4, 87 ratler common
Lolit ti tisidun Gatdin stbsp. lepl".oides (Boiss.) Sennen & Mauricio R & BS 117829: "The
\$ pornr" rA14[ tu oör.r0l-50 C6-l .ate
Loliun isidun Gatdi^ sibsp. tigidun E B 95 - I 38 : C3-l j EB obs. C5, C6 scattered
Loliun subulatuüYis. ER95 104: C3-1t ER 96-69i C5-l rare
Confi.mrtio. for th€ Cr€taD rre!.
I BtRCV |-F & 4r FTORAAlDVICI l4llO\Ol üAVDO\LÜRrrrr'
Nor gren by IURLAND & al !loo3i and JAIIN & SaHÖ\Fl-l DLR ( lqq5i. bur repoc
led for Kriri bJ RLCHIN(JER (1s44, änd found by RÄUS (10q" er in lin ) on Kasos and KaF
parhos. fte spcctes i\ orherwi.e knosn lrom Cyprus. lsmel. Lebanon. Slna rnd Crcalra I ll-R
RILL, lqö8r. as uell as Anarol'a tMlll . la85r. I ollow,ng ILRRFI I ll9b8) spccic' !änl i.
considered appropriate herc for this ta{on which ditrers cleärly from the r. .iad,n subspecies
b' rhe rollowins combinduon ot char€creF: rulms uprighr. spil'c'.traighl. ax's rndurared and
eicarated ro panl' embla.rng Ihe sprlelel'. adpresscd Blumes longer Ihdn 'p,keler' rBFRC
MEIER & RAUS, in prep.).
Lol rm tehulenta^ L. Ä&aSl780J:"1 -4 kn Nw ofKastri" (BC45); t & AJ94-11:C6-
li t 9J Jor D56; EB obs. C3 scattered
Parupholi\ incuna tL ) C l-. Hubbard R d 85 1-680 Along rhe road frcm rhc harbuur ro
Ka,tn rB( o): tA d 4J w ab. ol 2tt: 86-1, tB & AJ a4-18 Cn-l: FB 3 AJ 04 l2l: 85-
l: ER & AJ 94-t02: A4-lt ER 95-75:E'7-1; RJ 910526-32: E6 t. BJ 960401 71: D6-6, EB
obs. Cl, C5 scattcrcd
Paropholis na.ßinatt k)nemark adas.l 8/Jri l4kmNqof Ka.h- I BC45 r: ß d 8J
a-8l8, Small srrem ncar rhe Nw poina (A:l): Rd a.Sr'-dloi Ihc\Wpoinr (Al4)r
ER &.1J94 103: A4-l very scax€red
Pholanr niior k tz. - R & BS 42818, "Small stream near the NW point" (A4). tB & /U 94-
2l:86-2, EB 95-23: C]-t:, EB 95-49: D56t EB 95'163i C6-4 scattered
Phala.is pondofl | Rd8S4".3:"2 4kms sWofKa'rnlD5):ßdASr_8r8i"l 4km
\\\ of Kastri (B(45),tA&AJa4-'l Db-l:1805 /08i C.l-l rery scdflered
Phleun crypsoides (D'Urr.) Franchet subsp. c'lpsoiles Dörnet 22 lMaillea usillei PatL):
''Dünen an der Nordküste" (see also RECHINGER & RECHINGER-MOSER, l95l: table
29); R & RS 17870: "Sü^kitiko" (B6); ER & AJ 94-2: B6-tt RJ 910527 09: A5-1; RJ
960401-60:C6'ltEB obs. A4; RI obs. B7i D6 scattered,l@ally fiequent
Phleun subulatt tn (SaviJ Asch. & Gr&bner - i &,S?7ZZJ: "Kastri. Th€ ravine at the church"
(c5)
Pipta,herun nitiaeun (L.) Coss. subsp. nn d."rrr EB obs. 86, C3, E7; RJ obs. C6, E6
Poa itj.naK]dfih Aa 96-.18&33i 1.4- 1, D56i R l 960402-09a: B1-t: RJ 96010 t 33: C6-lt EB
obs. C4. C5 scattered
civen 6 "bcfor. 1930" record for Gävdos by TURLAND & ä1. ( I 993: 389) allhough ther€
is apparently no literature basis for this.
Poop€t6ali H. Scholz R.I960402-11:C3-l -ftie
Field records of Pod ,5"/rosa ass. (EB obs. C6, D6) rnay belone here.
34s
346
Pouposon üaritinus Willd. subsp. ,rattt tt&r EB 96-71: C6-tt EB obs. C4, C5 scattered
Polyposot a n,,üs Willd. subsp. rr!ärpattac"ls (R€q.) Asch. & Gmcbner - t 9J-74i E?-I
Potwogol nonsrylieatis (L.)Desf. R&aS47827: Snall strean near ihe NW point" (A4);
EB & AJ 94-9t: A4 t; ER & AJ 94 120: B5-l: RJ 960401-52: C6-l scalüered
Psilutus incuryß (Go\an) Schinz & TheU. ER95-27:86-2.ER&'4J949a:862;Ea9673:
C6-\t RJ960401-35: C6-t. EB, RJ obs. 87, C3, C5; EB obs. E7; RJ obs. 85, C4, D6 corn-
Rosbaia cß,ato (L.) T^elev v1r. cristata 8895 21:86-2tRJ960101-37:C6-l;RJ96u0l-
Jl: C6-l; EB, RJ obs. 87, C3, C5; EB obs. E7; RJ obs. 85. D6 comrnon
Beside tbe 0?e variery var gtabrifoto (Tt^!tv.) M. DoEan was found on Gävdos (t 9J-
102: Cl-li EB obs- C5) which is distinguish€d in habil by the very long. lob€d inflorescenc€, a.d
was found as a segetal weed only. The following record refers to the sp€cies: Dö?t?r 41i "Karst
boden bei Vathyana" (D56).
Stipa brcno les (L.\ DöÄet R & BS 47791:"Knsrti. The ruvine at the church'(C5); EB obs.
D6-l rare
SrllDa.apz'sis Thunb. EB, RJ obs. C6; EB obs. E7; RJ obs. 85, 87, C5, D6. E6 rathercom-
'rnphchne iiteß (Gtss.)Lit|.k Rechinger 13658:"Bucht Saraki.iko'(86) (see also RECHIN
GER & RECHINGER-MOSER, l95l: tabtes 28 md 29); R & a.S.lzz.l2i "Alons the road
from 'he hrrbour to Kastri (Bcb); f d aJ 4_8/9i _smdll {rream near the NW pornr" LA4r;
R d AS.,tr-84-i l hc NW poini (A34l,,q d Bs 4 Töl Sarak,niko (Borl gA or- iJi Bt!
t; ER & AJ 94-3: B6-2 ER & AJ 94-92: A4-ta RJ obs. 87 scatt€red, locally fiequent
[Triticsn tt.gi&.n L. s\tbsp. d!'!fr (Desi) Husn.] ER 95-105: C3 \ Ea & AJ 91 42b: C6-
I cultivaled as field crop and subspo.taneous in fallow fields
The noncnclature follows van SLAGEREN (1994).
yulpia ciliaaaDur.non. s\tbsp. ciliatc EB, RJ obs. C4 scattercd
hlpiafascia.lato(Fotssk.\Ftirs.h Rechinget I3657i "Sandstrmd dcrBucht Sarakiniko" (86)
(s€e also RECHINGER & RECHINGER-MOSER, l95t: t^blc 28); R & BS 447844: "'lhe
Nw point' (A34); R & aS 4/868. "Sarakiniko" (B6)t tR 9J-J1: B6-lt EB obs. A4, 87
rre, but locally fiequent
E. BERGMEIER & AL FLORA AND VEGETATION OF CÄVDOS (GREECE) 347
Iridacede
Ghdiolus italicut f,/illü Düfel 19 (G. tegetun Gawl.\: "! et Saaten"; Rechinget 13669:
"b€i Ambelos'(C3)j EB, RJ obs. D6: EB obs. C3, C5, C6, D5 scattered, locally abundant
Gwat.lritis nolooh ta Kl^n Döfict83 -aufKaßrboden. mirder rFischen lorm rt e O
.i\trinütud, Iti! tisytitu hiun L. lat nonoph) llat \see also UOLDBLA tl lq80): LB. RJ
obs 85: EB ob'. A4. 87. Cl. Drör RJ Photo: D6-4: RJ obs C5, C6, l-6 mther comrnon
Gr andfirit sisrnnchiutr (L.) Parl. EB, zu obs. C5, D6; EB obs. Cl, C5, C6, D5 - scatt€red
Juncaceoe
Jtncas heldrcichia tt Patl. stfrsp. heldrcichiat üs ER 95-166: A4-l; EB obs ET-t nte
Jtn.us hubridusBtot. Dötfl4 40 tJ. bufonius L. lar .orlpvrr wahlenb.,: _an was-erlachen
ber Ka<n rsee al'o RI?HINGLR & Rl-cH l\Gt R-MosER. lq5l: ls0r: R d as.l-828i
_Small sream ned Lhe NW poini' \A4)t EB 3 AJ a4 l19: B5'l: EB 95'lr0: l1'lt EB 95'
165: L4-t, ER 96'24:R7-2; P.I960401-48:C6-lt RJ 96rl0J-1.li E7-l; EB obs. C5 - scat-
Liliaceae s.l.
IA iun cepa L.l ER & AJ 94- I 27 : C6-4 cultivated, sometimes peßistins after cultivation
Altiun loryanun Pamp. EB & AJ 94-68:D6-1. EB 95-t44: D6-l; RJ 960102']8r C5-li EB,
RJ Photo: D6-4; EB obs. C4, C6 scatt€red
.4 iu'n nipu^L. Rechinset I3675:"\t erSa^ea";Eß95-48:D56;EE 9J-9Ji C3'l; EB obs
Variable, plants with green and black ovary were obsewed.
AIiuü bro'ittatu''Boiss. &Hel&. - R & BS 47745i "2 4 km S-SW ofKastri (D5); Ä & aS
778.1?r "r 4km NW ofKästri" (84)t EB, RJ obs.87, C3;EB obs. A6, E7; RJ obs.85,86,
C5. C6, D6 comnon
AUiün nifoliatun Cnillo - EB 95-96: C3-t. RJ 960401 l8:C6'1;RJ960403'05: D6-41EB obs
C5, D5 scattered
Asparaqus qhruts L. stbsp. aäierralis (Bake') P H. D^vis - ER & AJ 91-t t4: CD67; EB obs
C5, E7 scattercd
148
Asphodelus runosus L. EB, RJ obs. C5, C6, D6; EB obs. C3, C4, D5, E7: RJ obs. E6 com-
Relkrolia brcvbedicelata\üÄll - ER 95-62: D6-l; N r. ,. (cultrvated in the Botanical GaF
den ofthe Univcrsiry Regensburg): E6-1; EB, RJ obs. 86, C4, C6; EB obs. E7 scatter€d
Colchicun püsiuun Sieber EB 95-127: B7-2t ER 96 25: B'7-2; RJ 940529-08: B7-l; RJ
96a101-03: C6-l; EB, Rl obs. C3, D6; EB obs. 44, 46, C5, D5. E7; RJ obs. 85, 86. C4
Thc Gävdos population differs from specime.s o. the opposite Cretan coast of Sfakiä by
their broäder leaves exceeding 5 mm in width. Similar plants, howcvcr are not uncommon in
olher pans of Kriti (e.9. ,a 96-140lron thc plain of Onalos)
Gasu lbrusa (Desl.) Schrlt€s & Schulles 61. ER96-t05:C3-t nte
Gasea sraeca (L.\ Tena'.c. lR & BS l78.l.i: "The Nw point" (Al)i tt 96 ,0i 87 l; EB obs.
C5;RJ photo: C5-li RJ obs. C6, D6 scattered, locally abundmt
Gasea pedu culais(I-&C. Presl) Pascher RJ 960403-06: D6-4 nrc
Muscai conosan (L.) |lillet E896 29: B'7-l;EB,PJ obs.C6;EBobs.C3,C5.D5.RJPboto:
D6-4 rathcr common
Mus.ai sprcirzenholeri (Osterm.) Vierh. RJ 960103-16: E1-2;EB obs. C3 ftre
Ot,ti hogalun drubicunL. ER 95-113: D5t w.
The status ofthis showy species which is said to be occasionally cultivatcd for omament
has oflen b€en considered doub6.rl inthc Acscän (RECHINGER, 1943), orexplicitly "subspon-
taneous or casual" in vriti (YANNITSAROS. 1991). For the latter island it is known only from
än old record by Gandoeer (RECHINGER, 1943) and a recent on€ ned Rcthimo (scc TUR-
LAND & al.. 1993) where it is said ro be Mturalized in olivc srovcs. Howevet there is no rea-
son to doubt the native status ofthc single snall population which was found on Gävdos.
Onithoeal' naüonense L. R & BS 17724 : "w of Kasni (C1tt EB & AJ 91 3 2 : C6-tt ER
95-57: D6-3t BJ 940526 2|i D6-l; EB obs. C3, C5, D5 scättered
Prcsperc au,lnnale (L.) Salisb. s.l. (= s./1d d!tun,d/N L.) - EB obs. A4,87. C5;RJ obs. C4,
C6, D6 rather comon
The nome.clature follows SPETA (1993).
Tulipa s&atilis Sqeryel Dödtet 161: "a\tf ^ckembei Anbelos" (c3); ta 9J ./ 09: C3't; ER
96-8J, D56j EB obs. C5'l very scattered
E. BERCMEIER & AL. FLOM AND VECETATION OF CAVDOS (GREECE)
Uryin.a üaritiht (L-) Baker s.l. EB, RJ obs. 86, 87, C5, C6, D6; EB obs. ,44, ,46, 84, C3,
C4, D5, E7; RI obs. 85, E6 - conmon
Orchidaceae
Acerus anthrupophorun (L.) Aiton fiI. - EB, RJ obs. C4, D6 - very scatt€red
Anacanptis pynnidalis (L.) L. C. M. Richard RJ 910526-12: D6-l:EB,PJ obs. C6;EB obs.
87 scatlered
Linodots abo irut lL.) Swanz EB obs. 86, C6t RI obs. D6 scaüered
Neon"ea naculots lDesf.) Steam EBobs.D5 rare
Ophrys apilen H\tdson R & BS 17826i "Small stream near lhe NW point" (A4); EB obs. C6
Ophryt epkcopalß Ponü Dö4er 1152 (O. luti|lo/a Rchb. f. var. dariDa Fleischm., FLEI-
SCHMANN. 1925): "Kaßtboden"; EB, RJ obs. D6; EB obs. C6i RJ Photoi D6-4 scatte-
Thc obscNcd plants had rclarivcly small flowcß.
Ophrys fuva a8;g. - Dörfel 80 C'D,pit4" sensu FLEISCHMANN, 1925): 'Karstbodct'\ Dörfer
a2 Ql/311904) ("\zt. funerea Viv.", FLEISCHMANN, 1925\: RJ 910526 19: D6-tt Rl
91052627: D6-4]' RJ 910527-0J: B5-2; EB, RJ obs. C4, C5. C6; EB obs. 87, E7 corn-
Allplanls seen bad a sftall. twice geniculate labellum and can thus be retened to Opiryr
'?inereaphila fusca " (PAULUS, 1988), a clearly distinsuishible East MediteF ean taxon
which has not yet been validly described, and which is widely distributed also on thc Crctan
mainland. The nam€ Opirysrne,'ea Vv refeß to a taxon endemic to Corsica and Sardinia and
can,ot be applied to this taxon (PAULUS & GACK, 1995).
Ophrys nannosa Desf. Döitet 171 (type of O. doedsri Fleischmann): "Kaßlfelsen am Kap
[amarcla (l-1) ('ee lLl ISCHVAft\, la25;CAMPBI-l I , IaR2; COI Z& Rl INHARD,
lq85).RdaS.l 8JJ. 2 4km\oiKötri (A5l:RJo105)o 20 D6 I:l-8.Rlobs.C5.
C6; EB obs. 86; RJ Photo: D6-5; RJ obs. 85 scattered
As stared in rhe cited works the rostrum of the anrheFconnective in these plants is conspi-
cuously long. Thus the Gävdos planrs resemble A r/anshr'cana Czen. Ophrys c/prrca (Vierh.)
Nelson to which the name "O. doe üet i ;s ofien rcleft ed do€s not occur or th€ island.
Ophry"s siculaTineo Döilet 42 ("Ophtls tüte1lCav")r "Karstbod€n"; Rl obs. Cl, C5 very
350
Ophry\ ten h,ediniferu Willd. EB. RJ obs. C6; RJ obs. C4, C5 v€ry scaltered
or./'ir cotlina Banks & Sol. RJ obs. C4. C5. D6 scattered
Orchis conophoto L. sibsp. tagtats (Pollini) Sudr€ - EB, RJ obs BT; EB obs A6, C6 scat'
Otchß irolicoPoier EB obs. C4 - rare
Otchis papilionacea L. RJ obs. D6 lare
Orchis quadtbunctoraTei. EB obs. C4 nrc
Serupiß beryotüE. G. Camus EBobs.C3.C6 ra.e
S.topi6 oientatis l. \elson lRdASr--i4i-2-4LtnS Su ofKa'ri (D5':fB9J-Jt Dö-
l, fB so-J2 C6.5; FB, Rl obs. C5: Rl obs. C4 - ralher common
Potano&eto acede s.l-
Posidonia oceanica (L.)Delile EB obs. B6i mentioned by ICONOMOU (1989) n'orn sea b€ds
near Cap€ Tripiti (E7), Gaiduronisi (A3), and Karavö (C6) scattered otr th€ coasts
Rüppio na,itida L. - RJ 960403- t t: E7-l - nrc
According to TURLAND & al. (1993) lhere is no published post-1930 reco.d for Krlti.
zosteru rnatinaL. mentioned by ICONOMOU ( 1989) ftom sandy sea beds neär Karavö (C6)
Zanrichelliaceae
Zamich. ia D..luncuttuo Retchenb. - Dörfer 44 l"/a nk hcllia palßtt\ I ) ln $6scrla-
chen beiKasri (see also RfCHINCLR & RFCHINGER-MOSI-R. Iq5l l00). ER ot-
l0l: C4-t. ER 96 49: A4-l] RJ 960402 17:C4-l v€ry scattered
Current flons generally refer io the Gre€k and A€gean rePresentative(s) of tle genxs asz
,d/lsrrir L Ta\onomv and nomenclaturc here are accordine ro TALAVERA & dl (1q86) The
'soecres rs ei'en lor Greece by lAl AVERA & al. ( lc86: 266). bdsed on a Heldreich collecüon
fiom Attiti bur is ceflainly much more wrdesprcad. d pe'haps lhe commonest Zaurri rellrc
E BERGMEIER & AL FLOR"A AND VECTJTATION OF GÄVDOS (GREECE) l5l
Chorological commenls
Alftost 60% of the flora of Gävdos belongs to the Mcditcrmnean or East Mediteranean
phJloecographicäl elemenr (l rß. 2). Chorolotsi'dl 'elälronshrps our\tde rhe Medrlen can ac
irrongc't wrth the lrano-lu€nian and thc Sdharo-Arabian regron. The vedircmncar'/Orien-
hl/Saharian elemeht includes alnost 20% ofthe flora. widely distribut€d Euro-Siberian species
äs well as th€ pluriregional d the neophytic elements are poorly rcprcscnted. The only recent
adventive species is Oxalß pes-caprue. Compared to Kriti (see JAHN & SCHONFELDER,
1995: 26) thc Aegeän endemic elem€nt and particuldly that ofth€ Cietan area is less promin€nt
on Gävdos. But this is easily €xplained since most ofthe small-area €ndemics ofthc Acgean are
resrricted to high mountains or to cliffs, th€ latter being poorly, thc former not at all represenled
AccddrngroGRlt ll R (lo'lrwhcn elaboml ing on h rs ph)4ogeograph ical conccpr.lu! the
Sodrh Aegean ared. Gavdo' loms pan both ofwhdr he defined a' rhe (drdaescm!csion (com-
pnsrns Cyclades and Sourh Aegean lslands e\cludtng Kirhim dJ Ru.lus) dnd ,he "Sourh
Aesean (Sourh Aeeedn i'ldnd" includrng Krrhttu and Rodo'). wirhin rhese phyrogeographical
region.. uä\dosjoin. the $err ( rerän subrcgion shich i\ the ned'esl pan oflhe marn iJand.
Gävdos haöours 2l taxa endemic to the Aegeat, oi parl thereol By adopting Greuter\
cdresories ol endemics ro rhcsc ura, they may be "ubditrded Io J s'cno-(ndemi.( (ie, ra\a
contr-ned ro rhc sc{ Creran subtegron)r 4 pluiendemrc' (confrned ro various ( ßran :ubregron\l:
6 Cddacgcan endemicsi 2 South Aegean endcmics; 6 Aegean endeftics. The three steno_cndc-
Belte\rttu b,eqr.lnllald. lno$n hrrheno lrom .oulh$e'rem Kr ili r{cc disrribulion mdpr
rn TURL AND & al., loo3, dd in PHITOS & al.. leaö: E5). i' ioun.l in mdn) pdns ol G;rdos.
in a variety of habitals, but hardly ever in lar8e numbers.
Bupleutun saudionun is rhe only €ndenic of Gävdos. This small annul was discovcrcd
b! Runemdrk and B. Snoserup in la80 dnd found b) us rhroughour rhe rsl d and locdlly abun-
dan'. Srnce I cross in a \üiery olborh open and shrubland habilars sc äre glad lo di.agree s irh
S. Snoeerup Lin PHITO\ & al . l006, sho had consi.lcreJ Ihii rpecre. as vulnemble.
Linoniun elaphonbxutu qas only recently described by MAYER ( I 995) fion the south-
wcsl Crelan coast close to the islad of Elaf6nisos. Ou' fi.ding is th€ sccond known locality.
A:peruh tteda. I Moniun ,rcticua. Mu'tu.i 'p,p 'ckholcn and Ononi, v.rc" arc plu-
riendemrc.. wh'le Crepis cretia. Crtpr Shal,telM. IIIoEo ' rytea:ß- Ncp?u nrl'n'toha.
NiEe a daerfleri and Sedun .rertrrn (uless thc latter is conspecific with S. qr€rdrrn Brullo
& Fumari from Libya as assumed by JAFRI & RATEEB, I 98 I ) ar€ restricted to thc Cardaegean
region. Onlytwo specics, viz.,Tet riun nicrcpb n d lrba r6artrs (extending slighdyto
soulhwestern Anatoliä) may be consider€d Soüth Aegean cndenics, whereas the following have
a somewhat wider distnbution, ranging to thc nothem Aegean regiot and/or to thc adjacent
m i,nlamds AreMria nu 1li.\, Crcpx nuhifura, Galiun gaecun s$sp. sruecun, Malcolnia
./€r,osd sttbsp. ndrensi.\, Pdpdtet purpureonaryinatuü and Phleth ctypsoi.les. Ranunculus
'.rert.,!s, believed to belons hire, howevir, occurs ln Libya as well (GREUTER & al., 1989). thus
represenrns a eroup olruä be'ng nsrtree\clunvel) !n rhc AcBean dnd \onh Alrrca lhrsgroup
rq particul ly sell repre{enled on Cjldos.
The sourh Aesean reßion, vil.. the dry "oulheaslem far' oI Kriri d thc islands Gd!do'.
{ hns, rci,.loxr.ni\ir anJ Koufon,n sourh ol Lhe main islad. ri tnowr for lhe contderable num
ber of;oufterly distributed taxa which are missins, or alnost so, in theadjacenr contin€ntal pats
ofcrcccc and Turkey. They comprise both Saharo-Arabian and Ira.o-Turanian taxa (i-c., widely
distributed in semidesen arcas in North Afticaor Southwest Asia, orboth)ddtruc South Medi-
tcrancan taxa which are, in Norlh Africa, confined to areas with Meditcmncan climales. viz.,
the Gebel Akbdar dea (Crenaica, Libya). BALDACCI in l9l2 had already noted the floristic
similarity of KJiti and Cyrenaica and mention€d Gävdos as an imporlant phytogeographical link.
152
5o/o
2%
3Vo
gqo
't9%
F€.2. The Dhytogeosraphical elnenß in rhe dora ol Ctvdos. I = Creb ma endemicsi 2 = aegem area and rdjrcmt
Tp'n-.,J ldnVeutrm.ea.,4 Veo.emne,niS ved easnedn On"nuL/<äl-r d1 6 Vedirmrcanrlempeore
l;.pe t lürcoca^olurhe'e$r,i n.üirsorz (!r"puhü'. 'nrodrcco chodlo"roldr', treRr1'efi) L^er
lrcm DAVIS (l9it5 1935), FEINBRIJN-DOTHAN (1936), GRELIIER & a1, Med Checklisi (1934, 1936, 1939),
PIGNAnI ( 1932) and ZOHARY ( 1966. 1972. 1973)
SA\DWITH & SIMPSON (lq4l, commcnrcd rhar the fiora ol Cyrenarca r"telared mlher ro
Crere änd Creece rhan ro the ddi&enr Jcscds . Rl_( HINCLR (lq4ar enumerared \ourhem spc-
cies" wbich rhe South Acsca islands had in comrnon with North Africa. GREUTER ( l97l) lis-
ted 28 species with South Aegean semidesert-tlpe habitats. Eleven of thcm occur on Gävdos.
Si'ce thcn additionäl taxä have been found in KJiti which supporl strong ecological, and possi-
bly historical phytog€ographical links bctwecn the South Aegea, ard Gebel Akhdar, and our
research on cävdos revealcd a fcw rnore. Four taxa represenl an elenent known exclusivcly liom
th€ South Acgcan and the opposite Nonh African coasti
Alliun lonaanum was considered a Libyü endemic by EI-GADI ( I 977: I 2) but has si'ce
become known from two cycladean islands and the far northeast of Kritj (STEARN. I 977, with
disbibution map). Our new findins on Gävdos consists of several but rather small populations
totallina a few hundred flowcring plänls.
Ca irri.he pulchrc oc.\!ts only in northeastem Libya (omittcd by JAFRI, 1984) and on
Gävdos where it had been collecred by Dö.fler (SCHOTSMAN, 1967). Our records some 90
years later ar€ the lißt confimation- Few populations exist on Gävdos which are wlnerablc du€
to the; small numbcr and the instability oftheir habitat. Turland in PHITOS & al. (1996: 108)
rightly draws attenlion to C pdrrra as a "Red Data species" but ccrtainly errs conceming dala
on ils habitat, €cology and potential threar.
Reseda odorsta is considered native only b Libya, and in Europe to Gävdos (GREUTER
& al., 1989). elsewherc @cunins naturalized or as a mre escape from cultivation beins srown as
E. BERCMEIER & AL FLORA AND VECETATION OF GAVDOS (GREECE) 353
an omanental and for irs fragrant flowcß. On Givdos. R. odo.ara occuß in both anthropogenic
habitats and semi-nahml rocky pläces.
Teu.tiun blefifoliun ]s known ftom Crenaica (and adjacent northwestem Esypt) and thc
South Aegeän extendjng to the southem Greek d southwcstcm Anatolian mainländs (BORA-
T\'I.ISKI & al., 1992: 259). In Kriti, it occurs mäinly in the eäst. The species is not common on
cävdos and the plants are heavily browsed by goats.
The following taxa de also rdc, and rcstricled in Greece to lhe Soulh Aegean, but tbe non-
I uro|en Ense is mu,h larBcr rhdn for rhe prevrourl) menroned spec,e'. e\rendrng from \onh
Atrica ro Suuth$eil Lurope änd or Sourhqesr Asia. or be)ond:
Aelulopus l,tgopoidet ls known in Eu.ope only liom Sicily, Mälta and the Cretan ärea
(sev€ral localiries in the Karpathos dchipclago by GREUTER & al.. 1983, and RAUS. 1990:
recent additions by MAYER. 1995, to lhe known occurences in Kriti see TURLAND & al.,
1993)- Thcre is a single population on Gävdos near Cape Tripiti which had nor previously bccn
Aizoon hispanican does nol occur on the Gr€ek mainl d nor on the islands cxccpt Kriti
where it was once recorded in the southc€ntral parl (JALAS & SUOMINEN, 1980: 102). It was
first found on cävdos by Runcmark and B. Snogerup, and since confirmed by us in few places
in the central and notheastem part ofthe island.
Artenisia herba-alba was also discovered on Gävdos by Runemark and B. Snogcrup
(reporred by GREUTER, MATTHAS & RIssE, 1984). This is the only creek populatid and
other tban its occurence in the lberio pcninsula, the only European one.It is restricted to a small
area north of Kastri whcrc it is threätened by potential roadmaking rath€r tla. by browsing as
stated by Tu.l d in PHITOS & al. (1996).
Asttuaalß sinaicus is very scattered in Greece and tle Aegea. and was previously known
in Kdti only from lhe small islandofChrisi otrthe southeast coast (RECHINGER, 1944). Howc-
ver. it also occuß alons the southcentral md southeast coast of Knd (IU, unpubl.). Our records
are tbe fißl for Givdos whcrc it grows in very srnall numbeß in various local;ties.
Astruaalß epiglaüis stbsp. ep€lorrs is widely dislributed in lhe South Mediten €an but
is apparcntly rare both in Greece and the Aegean. Reports for lhe Cretan area are bascd on a
Pichler collection from 1883 made o. Karpathos (RECHINGER, 1943:319); this occunence hns
not been confrmed since (GREUTER & al.. 1983i TURLAND & al., 1993). The single, ralher
small population found on Cävdos is cndugered by accidental destruclion ofils habilal iftepr€-
sents the lißt record for kiti.
Chldn)\lophora tridentata is found in North Africa from Tunisia to Egypt, cxtcnding to
Pale{rine, änd ß reponed, mnreoler, lrom Crprus and Rodos (GRJFRSON, la?5, CARL-
S lRÖM, lo8-r. Ih; occunence on oävdo'. frßr 'epoded by Ddrfler. dnd no$ conhmed aner
90 yeaß, is the only onc in the Flora Europaea area.
Elo.liün ci.utotiun subsp. ö?trrd&d belongs to a group ofclosely relat€d taxa in need of
lirnher taxonomic studies. It is treated by GUITTONNEAU (1972) as a sp€cies, r. dethiopicun
(Lan.) BruInh. & Thell., also by GREUTER & al. (1986), üd is said to bc distributed in
Morocco. Tunisja and Algeria. Another subspecies occurs on the Southwest European Adantic
coasrs. GHAFOOR(1978) mcntions forms ofÄ ctcrrartü4 withouta turowbelow rhepitofthe
mericarp which occur in Libya; these mighl represent subsp. ,ilrrur"n. For üe East MeditcF
ränean area, two records from the South Anatolian coasr md thc Eäst Aegean island ofKos are
cited by DAVIS (1967: 486) but not commcntcd on by DAHLGREN (1980) who gives only t
.i.rldlirn s. str. for thc Acgean. Our subspecies lnay nevertheless be more widespread in th€
South Aesean as it is cäsily confused with üe rype subspecies. On Gävdos, both taxa often srow
rogerher r .n r/d/ira subsp. biptrratua pteler .andy coa{dl hrbrral' (see al'o Cl I I lO\
Nl- 4lJ, lo-2. 105. D.ammophrle de. dune, liräes rclarnares dc' prnödc.) an.l DAVIS. l067:
486, "sand dunes, beaches").
354
Gwan.lnris do,lophylld anges ftom Cfenaica to Palcstine- ln Europe. it is known only
fron soulhem Greece (Attica, Egina, Kriti; see distribution map in GOLDBLATT, 1980: 255)
The Uävdos noDularion ul thrs ra\on rs rarher lage, ai are 'hose lrom rhe oppoile Creran coasr
,een b\ üs: bcr*een tlaloni.o. md Paleo.hom, and zlong rhe Slali; coa(r rBl RGMFIFR &
MArTirAS, 1995).
Helianth?aua 'uNlatun i{ a wide.predd \onh Airicdn-lano-Turantan raron s hrch onl)
iusr reache" Iuroneon lhc ßldnds(hirsr (Cdidouroni.i' dnd Koulbnßr (Rl-aHlNCER la'u:
TTjRLAND & al.- l99l I os {ell ar lhe coa{ ot sourhcdsr Krt' (VA\ I R. laa5) (bcside one old
record from ilrs Pcluponnese quo{ed b) R-E( HINULR. lo:14: s0. sub H. elliPtiun tDe'l t
Pers.). With our records on Gävdos, the species is known to occur on three islatds south of kiti
d !n one,raron on thc main rsland lhe Ca\dos plüls se'e lound rn lwo locarion' in lhe
nonhersr. rhe habirars berng endangered md likel) lo be desb_oled by uncontrolled 'ourislic
Matricana aured occrts in thc southem parl of the lberian peninsula, Sicily. Malta and
Lampcdusa, as wellas in Nonh Africa and SouthwestAsia. The closesl occurence to our rccords
whici are the 6ßt for Gre€cc is inLibva{ALAVI. 1983).lt wäs älso found on the southwcst Ana_
rolian Mamaris p€ninsula by CARLSTIIÖM (198?).
P"nplotd onsu,tilolio, otheru i'e knoqn fiom Spsrn. Stcrly. Malla dnd Nonh Afnca. ß rn
creecc risnrcred ö rhi r.l.nds oä\tlo' and Chri.r (RicHl\uFR, 1q44. BoRAI YNSKI & al.,
loq2 lö7). We found ir Lhroughoul rhe rJand bur ncvcr in lärge numbers. A|hough 'o sume
desree brossed, rhc Uä\do" po-pularron rs nor rhrearened b) grtrng sherca' building of loL,irsr
lacilrrre, may p"'e a rhrear io.hrub' in dune habira's lhe species is th('elore appropnarel)
considercd vulnerable 'n Grcec€ by Turland in PHITOS & al. (1996: 406t).
PIditdEo squanosu, n eastem Mcditenanean taxon, is in Europc confined lo Greece and
th€ Aegeanlrea. Th€ Cret records refer to the islands of Elnf6nisos and Gävd6. Thougb
occuning io large nunb€rs this psammophyte must bc considered threatened due to potcntial tou-
rist developmcnt in the area.
Sile e succuknta subsp. s!c.!/erta occuß on the North Africän and Near East coasts,
reaching Europe only in the southemmost Aegean rc8ion where il is reshcted to four islands
soulh, o; southwesr, of Xriti, viz., Elafönisos, Cävdos, Chrisi and Koufonisi (se€ distribulion mäp
in PHITOS & al., 1996: 473). S. succüLhta ts clearly threaten€d by increasing beacb towism.
Orher ra\a s irh ä chiefl) Sou'h ved'rerancan dßniburron mnge occur on G;!do" bur mo'r
are 'omeshar s ider di(mbuied and les' 'arc in lhe Aegean rhan rho'e prc\ iou'ly lßled The lol_
lo$rng Jesene.pecrr'l men tion. 4 a' hu'o o?gtthr' d. Codua' cllu: aetuleü\. Ccntauryd nch
tensß Cistus patufotß, Crtvolruhß okifolius, Crassula vailldntii, Clelis pßilld, Di.lesnus
r"gr'!tü'. Etirliun loriniütutu. Juaperß natrccorpu. lotu' hdloPhilß- Mal,ol4ia rh^rto
uilid dcsptia. Oaun' ht'pan.a suh,p. hßpüniJ. O n4!t acbcn. O,nithosülun a,aht.üa.
PldnrasoTlbictlhs, Silehe behen, Silene ltuticosa, Speryuldtia dian.lra anö Ttipldch'e nitets
uRFU ft Rt ( la-1, phyrogeogaphr.dl concepr ts suppoded by our comlaflson' oi florß-
lic simrlan||e: berseen cärdos and sunoundin8 rcgion. (Fig. JJ. Gä\do, ha" q4oi ol its lä\a in
conmon with westcm KJiti, and 9l% wilh thc ccntral and eastem parls ofthc nain island. Il
,hües scll o\er 80oo w,lh Karpalho' dnJ onl) .l'gh'l) less sirh R6dü'. Kilhiß in lhe nonhwesl
and rhc,ourh$e,r Analolian Mmnari" penin'ula tn rhe nodhcasl $hich is much more distanl
hd\ e romewhJl less 'pccies in common wrlh Cä\dos thän Libya ,n 'he.ourh. I'unhc' behind dre
rhe.ourhemmo.r Peldponnese $hrch is rcprc.enred in our fr8lie by rhc smdll i.länd ot t laroni'r.
abour 200 Ln fiom Cardos. and rhc Mdhe'e irland' almo.r I000 km ru the fat $e'r Allro\i
matcly 60% of tbe Gävdos spccics occur in each ofthem.
The percentage valu€s (and absolute numbers) as displäyed in Fig. I indicate that thc phy-
togeographical connections via the South A€gean island arc äre much slrong€r nonhcastwÄrd
rhän nonhuesvad. The hnk, ro \onh Alrica m more pronounced lhan onc mighl e^pecr lrom
rhe lacr lhdl rhere i( no ldnd beMeen Gä\Jus and lhe I ib)rn coasr. lhc 'oulhem conneclron rs
E BERGMSIER & AL IJLORA AND VEGETATION OF GÄVDOS (GREECE) 155
I'e Ph\rJreocmohLc. roared'ol oesvnCatdo and unoundin! eg,Jn. IhengLr.rnd'.de.b.oLenrrbe.
In_d oc(e1;aa\siuiofrheud'do' noo Oe'er' r rhe e.pecn\e arca\ \oLne.lFideLrplbli.l-edd.uolPJ AL &
JAtÄt rrc;io'_r. r^FRl d. 1..ÜaDri,a7-ro3ö, ll-ucDl Llo^b la0, 'ARlslRoMrl03-1.lRlt'lR&
Rt,HtNutRrtob_r.lRtrrlR&". -Ved- hdlhr"(lc{,l.l03ö o30).HASL{V&r 'la--'.IACrI llm2).
s(HLMBRI & \t I ra\A I,o3ar. tljrFLA\D & st I taol, d o \A.{N jAFu\ L,oÄol
rndeednorles'p'onouncedrhänrhenoflhwcslemoneAlrhoughKirhi'ai'lessdis€nrrhanClre
narca 180 v' )50 km and rhee i' Knri nnd Anrilrrhrra alois rhe roure, rhe flonsric simildiq.
hoseve' insumcrenr presenr kno$ledge olrhe respccrite floras ma) be. rs J'8hd) sealer rhan
w,rh L,bva. Srnce lhcrc $ere no land conneclron{ between lhe Afncan cononenr and lhe Sourh
Aesean:Rerthc rerum ofthe \4ediren can Sea 'n P|ocene. rhc 'tnking degree ol flori'ric
RD-resenlarion ofCavdos sprcres in northem L'bla ma) bc intcrpreled rather b) ecolo8i'al üan
bi hi'rori,dl ph'logeosraphy. Finall). the relar\el) hish pocentage ofC;!dos \pecics ptesenr
on rhe Mallese rdand\ ma) also be.au.ed b) 'imilar habilat condirion. and planl communrlre\'
The latter remain to bc described in the second part of this work.
ACKNOWLEDGEMENTS
We ato nüch rndebred io Dr. B. S.ogdp and Prci Il Runemal4 Lund, fot poviding us wirh unpublished d.b
fod lhct cdvdos visn in 1930. nä*s foideleiflindion of specinEns rre due io Pst H scholz, Bdlin (Bu trslu.!
',J!\,3ndlr' A Mdrc,lDrberg(7r'ni,,') wedgnatolro{ aßercnopuLU\ Arhcns r BeßTei"' wa;
D V Ddtunrr' ,lall'o, D' v lit'm. Vi 'ch. Dr D PeddToelou arhdc Dr' I Fau' BerLin Pmf A su d 'opcn_
hdrcn.Dl l\r l.r CoDenhä,c,. D' L lrnßlo. Pdn?., dnJ b ai rnodynou' rcrere. wlo p'!\'ded vaLab.e nrorg
o;o-^..."d.'a'o,."oi'f'eld.udi".orlBt{RJhaveoeeanrdep.\iblethdnl\rortu fnm'\eD r!Fe
Forshungtsnein{hln
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