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Lectotype designations and taxonomic corrections on species of Lygaeoidea (Hemiptera: Heteroptera) described by Gustav Breddin

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Types of selected species of Lygaeoidea (Hemiptera: Heteroptera) described by Gustav Breddin, so far of unknown depository, were re-examined and the taxonomic and nomenclatural problems that existed among those species are discussed and resolved as required. The holotype of Dieuches schultheissi Breddin, 1906 is documented. Lectotypes are designated and documented for the following taxa: Lygaeidae: Aethalotus horni Breddin, 1907; Aspilogeton nubicola Breddin, 1901; Lygaeus hospes Fabricius, 1794 var. celebensis Breddin, 1901; Oxycarenidae: Oxycarenus musculus Breddin, 1907; Rhyparochromidae: Lispochroa blandula Breddin, 1907; Notochilaster teres Breddin, 1907; Dieuches horni Breddin, 1906; D. jacobsoni Breddin, 1906; D. nudipes Breddin, 1906; D. villosulus Breddin, 1906; Poeantius brevicollis Breddin, 1907; Pamera recincta Breddin, 1901; Paromius robustior Breddin, 1907. The following taxonomic changes are proposed: Naphius schultzei (Breddin, 1913), new combination (transferred from Rhyparochromus); Graptostethus verticalis (Dallas, 1852) = Pyrrhobaphus scutellatus Breddin, 1907, new subjective synonym; Naphius schultzei (Breddin, 1913) =Naphius capensis Scudder, 1971, new subjective synonym.
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Accepted by M. Malipatil: 28 Apr. 2015; published: 29 May 2015
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Lectotype designations and taxonomic corrections on species of Lygaeoidea
(Hemiptera: Heteroptera) described by Gustav Breddin
ELŐD KONDOROSY
1
& DÁVID RÉDEI
2,3
1
Dept. of Animal Science, Georgikon Faculty, University of Pannonia, Deák F. u. 16, H-8360 Keszthely, Hungary.
E-mail: kondorosy.ee@gmail.com
2
Institute of Entomology, College of Life Sciences, Nankai University, Weijin Road 94, Tianjin, 300071, China.
E-mail: david.redei@gmail.com
3
Department of Zoology, Hungarian Natural History Museum, H-1088 Budapest, Baross u. 13, Hungary
Abstract
Types of selected species of Lygaeoidea (Hemiptera: Heteroptera) described by Gustav Breddin, so far of unknown de-
pository, were re-examined and the taxonomic and nomenclatural problems that existed among those species are discussed
and resolved as required. The holotype of Dieuches schultheissi Breddin, 1906 is documented. Lectotypes are designated
and documented for the following taxa: Lygaeidae: Aethalotus horni Breddin, 1907; Aspilogeton nubicola Breddin, 1901;
Lygaeus hospes Fabricius, 1794 var. celebensis Breddin, 1901; Oxycarenidae: Oxycarenus musculus Breddin, 1907;
Rhyparochromidae: Lispochroa blandula Breddin, 1907; Notochilaster teres Breddin, 1907; Dieuches horni Breddin,
1906; D. jacobsoni Breddin, 1906; D. nudipes Breddin, 1906; D. villosulus Breddin, 1906; Poeantius brevicollis Breddin,
1907; Pamera recincta Breddin, 1901; Paromius robustior Breddin, 1907. The following taxonomic changes are pro-
posed: Naphius schultzei (Breddin, 1913), new combination (transferred from Rhyparochromus); Graptostethus verticalis
(Dallas, 1852) = Pyrrhobaphus scutellatus Breddin, 1907, new subjective synonym; Naphius schultzei (Breddin, 1913) =
Naphius capensis Scudder, 1971, new subjective synonym.
Key words: Lygaeoidea, Breddin, lectotype, new synonymy, Oriental Region, Neotropical Region
Introduction
Gustav Breddin (1864–1909) was a prolific author of Heteroptera having a significant impact primarily on
Pentatomoidea, Coreoidea, Reduviidae, and Lygaeoidea. Except of a remembrance containing a list of his most
important publications (Schröter 1910) and some brief obituaries (Dixey 1910, Kuhnt 1910), no detailed biography
or complete bibliography of Breddin has ever been published.
Breddin described 75 species and 7 infrasubspecific taxa (“varieties”) of Lygaeoidea. The majority of the types
of these taxa are now deposited in the Senckenberg Deutsches Entomologisches Institut in Müncheberg (DEIC)
(catalogued by Gaedike 1971), but several other types are scattered at the Senckenberg Museum in Frankfurt am
Main (Schröder 1964), the Zoological Museum in Hamburg (Weidner 1972), the Natural History Museum in Berlin
(Museum für Naturkunde, former Zoological Museum of the Humboldt University Berlin, ZMHB) (J. Deckert,
pers. comm.), the Zoological Collection of the Martin Luther University in Halle-Wittenberg (MLUH) (K.
Schneider, pers. comm.), the Hungarian Natural History Museum in Budapest (HNHM), and the Museum and
Institute of Zoology of Polish Academy of Sciences in Warsaw (MZPW) (T. Huflejt, pers. comm.). Probably type
material has been deposited elsewhere, too. The above three papers and a personal communication of J. Deckert
(ZMHB) list type materials of 55 lygaeoid taxa; type depositories of an additional 27 taxa have remained unknown
so far.
Descriptions by Breddin are usually detailed and therefore most of the species described by him are easy to
recognize. However, Breddin rarely indicated the number of the specimens on which his descriptions were based,
therefore most of his types have to be considered as syntypes following Recommendation 73F (“Avoidance of
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assumption of holotype”) of the International Code of Zoological Nomenclature (ICZN 1999). Depositories of the
type(s) and other specimens examined by him were usually not mentioned in his works, or depositories are
incorrect now (S.M. Blank, pers. comm.).
A further problem is that Breddin, particularly in case of species described in his later papers (e.g. Breddin
1906, 1907a, b), frequently did not provide type specimens with type labels, sometimes not even with identification
labels. After Breddin’s unexpected death, Ernst Ewald Bergroth completed, edited and published some of his
unfinished manuscripts, but Bergroth also left these type specimens unlabelled. The above circumstances
frequently render recognition of Breddin’s types difficult.
In some cases specimens are present among identified or unidentified materials in the usual depositories of
Breddin (most importantly DEIC) which morphological characters, label data, and other features fit well with the
original description of some of Breddin’s taxa, and it is reasonable to suppose that these represent specimens on
which the original descriptions were based, thus have type status. A careful consideration of the concerning
specimens is, however, necessary in each case.
During visits to the DEIC by the senior and to the RMNH by the junior author apparently overlooked syntype
specimens were located and studied. Re-examination of some of these types revealed unresolved taxonomic
problems. Discussion of these problems and proposals of solutions are the subjects of the present paper.
Material and methods
Label data are cited verbatim, lines on a single label are divided by a slash (/), comments on label data are provided
in square brackets. Printed text (pr) is not indicated except if the preceding text is handwritten (hw). In addition to
the original labels each specimen has also been provided with our lectotype or paralectotype labels. The symbol ‘>’
between the original and final depository indicates specimen transfer between two collections.
Abbreviations for depositories:
BMNH Natural History Museum, London, UK;
DEIC Senckenberg Deutsches Entomologisches Institut, Müncheberg, Germany;
HNHM Hungarian Natural History Museum, Budapest, Hungary;
RMNH Naturalis Biodiversity Centre (formerly Rijksmuseum van Natuurlijke Historie), Leiden, The
Netherlands;
ZMHB Natural History Museum (formerly Zoological Museum) of the Humboldt University, Berlin,
Germany.
Family Lygaeidae
Lygaeus hospes var. celebensis Breddin, 1901
(Figs. 1–2)
Lygaeus hospes var. celebensis Breddin, 1901a: 17, 78. Syntypes: [Indonesia:] Celebes [= Sulawesi], Lompobatang; depository
not stated.
Type material examined. Lectotype (present designation): ♀, “Celebes / Lompobatang X XI 95 / (Gipfelzone
über 2500 m) / Dres. Sarasin” [hw], “Coll. Breddin”; pinned, intact (HNHM) (Figs. 1–3).
Remarks. This species was described from unknown number of specimens (Breddin 1901a). The original
description and label data provided by Breddin (1901a) perfectly fit with the specimen examined during the present
study therefore it is recognized as a syntype and is hereby designated as lectotype. Four specimens of this variety
are present in DEIC (S.M. Blank, pers. comm.), those are probably syntypes too. They were not examined during
the present study.
This variety has apparently not been cited after its original description (Slater 1964). The single examined
specimen indeed represents Spilostethus hospes (Fabricius, 1794), a species widely distributed from Iran to
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Vanuatu. The variety celebensis has no taxonomic value and it should be considered as a junior synonym of S.
hospes.
FIGURES 1–3. Lectotype of Lygaeus hospes Fabricius, 1794 var. celebensis Breddin, 1901. Fig. 1, dorsal view; Fig. 2, lateral
view; Fig. 3, labels. © HNHM. Scales in mm.
Aspilogeton nubicola Breddin, 1901
(Figs. 4–5)
Aspilogeton nubicola Breddin, 1901a: 18, 79. Syntypes: [Indonesia:] Celebes [= Sulawesi], Lompobatang; depository not
stated.
Type material examined. Lectotype (present designation): ♀, “Celebes / Lompobatang X XI 95 / (Gipfelzone
über 2500 m) / Dres. Sarasin” [hw], “Coll. Breddin”; pinned, intact (DEIC) (Figs. 4–6). Paralectotype: with labels
identical with those of lectotype (1 ♀ DEIC).
Remarks. This species was described from several specimens (Breddin 1901a). The original description and
label data provided by Breddin (1901a) perfectly fit with the two specimens examined during the present study
therefore they are recognized as syntypes and one of them is hereby designated as lectotype. The species is
currently valid; it has apparently not been cited after its original description (Slater 1964).
Pyrrhobaphus scutellatus Breddin, 1907
(Figs. 7–8)
Pyrrhobaphus scutellatus Breddin, 1907a: 43. Lectotype (Gaedike 1971: 120): Ceylon [= Sri Lanka]: Nalanda; DEIC.
Current status. Junior subjective synonym of Graptostethus verticalis (Dallas, 1852), new synonymy.
Type material examined. Lectotype: ♂, “Nalanda / CeylonHorn”, “Pyrrhobaphus / scutellatus / Bredd.”
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[Breddin’s hw], “Lectotypus” [red], “H. Gaedike des. 196 [pr] 9 [hw]”; micro-pinned, tarsus of right fore leg
lacking (DEIC) (Figs. 7–8).
FIGURES 4–12. Type specimens of Lygaeoidea species described by G. Breddin. Fig. 4, Aspilogeton nubicola Breddin, 1901,
lectotype, dorsal view; Fig. 5, same, lateral view; Fig. 6, same, label; Fig. 7, Pyrrhobaphus scutellatus Breddin, 1907,
lectotype, dorsal view; Fig. 8, same, lateral view; Fig. 9, Aethalotus horni Breddin, 1907, lectotype, dorsal view; Figs. 10a–b,
same, labels; Fig. 11, Oxycarenus musculus Breddin, 1907, female paralectotype (Hemi-00039 of DEIC), dorsal view; Figs.
12a–b, same, labels. © DEIC; Figs. 7–8 by courtesy of S. Blank. Scales in mm.
Remarks. The species was described from an unspecified number of specimens of both sexes from
Anuradhapura, Horrowapatam and Nalanda, Ceylon [= Sri Lanka]. A lectotype (Figs. 7–8) was selected and
paralectotypes were listed by Gaedike (1971). Re-examination of the type material of both P. scutellatus and G.
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verticalis (deposited in BMNH) by the senior author concluded that they are conspecific, therefore the above new
subjective synonymy is proposed. Graptostethus verticalis was described from unknown locality. Later Distant
(1910) recorded it from northeastern India and Slater (1979) from Sri Lanka.
Aethalotus horni Breddin, 1907
(Fig. 9)
Aethalotus horni Breddin, 1907a: 42. Syntypes: ♂, ♀, Ceylon [= Sri Lanka]: Puttalam and Anuradhapura; depository not
stated.
Type material examined. Lectotype (present designation): ♀, “Ceylon / W. Horn”, “Anu- / radhapura”; mounted
on card, segment IV of both antennae lacking (DEIC) (Figs. 9–10). Paralectotype: “Puttalam / Ceylon Horn”;
mounted on card, segments III–IV of left and segment IV of right antenna lacking (1 ♀, DEIC).
Remarks. According to the original description the syntype series included male(s) too, therefore it can be
stated with certainty that a part of the specimens (paralectotypes) remains undiscovered. A. horni is a valid species
distributed in southern India and in Sri Lanka. Its diagnostic characters were discussed and illustrated and a
detailed description was presented by Scudder (1963).
Family Oxycarenidae
Oxycarenus musculus Breddin, 1907
(Fig. 11)
Oxycarenus musculus Breddin, 1907b: 219. Syntypes: ♂, ♀, Ceylon [= Sri Lanka]: Trincomalee and Anuradhapura; depository
not stated.
Current status. Junior subjective synonym of Oxycarenus bicolor Fieber, 1852 (synonymized by Horváth 1926:
136).
Type material examined. Lectotype (present designation): ♂, “Ceylon / W. Horn”, “Anu- / radhapura”,
“Coll. Breddin” (Hemi-00041, DEIC). Paralectotypes: with labels identical with those of lectotype (1 ♀, Hemi-
00040, DEIC); “Trincomalee / Ceylon Horn”, “Oxycarenus / lugubris Motsch.” [Breddin’s hw], “Coll. Breddin” (1
♀, Hemi-00039 DEIC) (Figs. 11–12).
Remarks. The species was described from an unspecified number of specimens from two localities in Ceylon
[= Sri Lanka]. Specimens from both localities were found in DEIC. Although one of them was identified by
Breddin as “Oxycarenus lugubris Motsch.”, the specimens fit well with the diagnosis of O. musculus provided by
Breddin (1907) therefore they are recognized as syntypes of the latter species, and one of them is designated as
lectotype. We confirm the subjective synonymy between O. musculus and O. bicolor proposed by Horváth (1926).
Although they are morphologically similar, O. bicolor and O. lugubris are not conspecific.
Family Rhyparochromidae
Subfamily Rhyparochrominae
Tri be Lethaeini
Lispochroa blandula Breddin, 1907
(Fig. 13)
Lispochroa blandula Breddin, 1907b: 216. Syntypes: ♂, ♀, Ceylon [= Sri Lanka]: Anuradhapura and Kanthaley; depository not
stated.
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FIGURES 13–19. Type specimens of Rhyparochromidae species described by G. Breddin. Fig. 13, Lispochroa blandula
Breddin, 1907, male paralectotype (Hemi-00044), dorsal view; Fig. 14, same, labels; Fig. 15, Notochilaster teres Breddin,
1907, lectotype, dorsal view; Fig. 16, same, labels; Fig. 17, Aphanus schultzei Breddin, 1913, holotype, dorsal view; Fig. 18,
same, lateral view; Fig. 19, same, labels. © DEIC (Figs. 13–16), ZMHB (Figs. 17–19); Figs. 17–19 by courtesy of J. Deckert.
Scales in mm; Figs. 14–16 not to scale.
Current status. Diniella blandula (combination proposed by Scudder 1962: 767).
Type material examined. Lectotype (present designation): ♂, “Kanthaley / Ceylon Horn” (DEIC).
Paralectotypes: with label identical with that of lectotype (8 ♂♂ 4 ♀♀ DEIC, 1 ♂ DEIC>HNHM); “Ceylon / W.
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Horn”, “Anu- / radhapura” (5 ♂♂ 5 ♀♀); “Ceylon / W. Horn”, “Anu- / radhapura”, “Diniella sp. [hw] / J. A. Slater
det., 19 [pr]” (1 ♂, Hemi-00044, DEIC) (Figs. 13–14).
Remarks. This species was described from an unspecified number of specimens of both sexes from
Anuradhapura and Kanthaley in Ceylon [= Sri Lanka]. Several specimens from both localities fitting with the
original description were examined and recognized as syntypes of L. blandula. One of them is here selected as the
lectotype. This species has only been recorded from Sri Lanka (Distant 1910; Scudder 1962; Slater 1964, 1979).
Tri be Drymini
Notochilaster teres Breddin, 1907
(Fig. 15)
Notochilaster teres Breddin, 1907b: 212. Syntype(s): ♀, Ceylon [= Sri Lanka]: Nalanda; depository not stated.
Type material examined. Lectotype (present designation): ♀, “Nalanda / CeylonHorn”, “Mizaldus sp. [hw] / J. A.
Slater det., 19 [pr] 62 [hw]”; mounted on triangle, head and segments I–III of right antenna broken off and mounted
on the triangle separately, left antenna and segment IV of right antenna lacking (DEIC) (Figs. 15–16).
Remarks. The species was described from an unspecified number of female(s) from Nalanda, Ceylon [= Sri
Lanka]. The labels of the specimen examined during the present study indicate that it is part of the original syntype
series. Notochilaster Breddin, 1907 is certainly not identical with Mizaldus Distant, 1901 (Kondorosy 2006).
Therefore we regard J.A. Slater’s identification label as erroneous. So far this species has only been known from
Sri Lanka (Slater 1964, 1979; Kondorosy 2006).
Tri be Rhyparochromini
Aphanus schultzei Breddin, 1913
(Figs. 17–18)
Aphanus schultzei Breddin in Schumacher, 1913: 69. Holotype: ♀, [South Africa:] Cape flats; ZMHB.
Naphius capensis Scudder, 1971: 1545. Holotype: ♀, South Africa: Cape Town; BMNH. New subjective synonym.
Current status. Naphius schultzei (Breddin, 1913), new combination.
Type material examined. Holotype: ♀, “Cape flats / L. Schultze / 764. IX. 04.” [dark blue, hw], “S.764”
[hw], “Aphanus / Schultzei / Bredd.” [Breddin’s hw], “Type” [red], “Zoolog. Museum / Berlin”; pinned, intact,
both antennae oligomerous, with three segments (ZMHB) (Figs. 17–19).
Remarks. Schumacher (1913: 49) explicitly stated that the authorship of this species (and several other species
described in the same paper) should be attributed to Breddin. The species was described from a single specimen
(holotype) that is deposited in ZMHB. The species has remained greatly omitted in the literature. Slater (1964:
1289) transferred it to Rhyparochromus Hahn, 1826.
Based on the re-examination of the holotype we disagree with the placement proposed by Slater (1964) and we
are convinced that this species belongs to Naphius Stål, 1874 therefore the above new combination is proposed.
This species is identical with Naphius capensis Scudder, 1971. Holotype and some paratypes of the latter species
were also examined by the senior author in connection with the present study. As a consequence the above new
synonymy is proposed.
Dieuches horni Breddin, 1906
(Figs. 20–21)
Dieuches horni Breddin, 1906: 326. Syntypes: ♂, ♀, Ceylon [= Sri Lanka]: Puttalam, Nalanda; depository not stated.
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Current status. Metochus horni (combination proposed by Bergroth 1915: 174).
Type material examined. Lectotype (present designation): ♂, “Puttalam / CeylonHorn”, “Museum Leiden.
[pr] / Dieuches [hw] / horni Bredd. [hw] / det: [pr] Blöte [hw]” [with black frame]; pinned, segments II–IV of left
antenna, left hind tarsus, left mid and hind legs lacking (RMNH) (Figs. 20–22). Paralectotype: “Nalanda /
CeylonHorn”, “Museum Leiden. [pr] / Dieuches [hw] / horni Bredd. [hw] / det: [pr] Blöte [hw]” [with black frame]
(1 ♀ RMNH).
Remarks. See D.
jacobsoni
below.
FIGURES 20–25. Lectotypes of Rhyparochromidae species described by G. Breddin. Fig. 20, Dieuches horni Breddin, 1906,
dorsal view; Fig. 21, same, lateral view; Fig. 22, same, labels; Fig. 23, Dieuches jacobsoni Breddin, 1906, dorsal view; Fig. 24,
same, lateral view; 25, labels. © RMNH. Scales in mm.
Dieuches jacobsoni Breddin, 1906
(Figs. 23–24)
Dieuches jacobsoni Breddin, 1906: 329. Syntype(s): ♀, Java: Samarang; depository not stated.
Current status. Metochus jacobsoni (combination proposed tentatively by Eyles 1973: 37).
Type material examined. Lectotype (present designation): ♀, “Java, Sama- / rang, 33M / lg. Jacobson” [hw],
“Museum Leiden. [pr] / Dieuches [hw] / jacobsoni Bredd [hw] / det: [pr] Blöte [hw]” [with black frame]; pinned,
segments II–IV of both antennae, right fore leg, tarsi of left fore, mid and hind legs, tibiae and tarsi of left mid and
hind legs lacking (RMNH) (Figs. 23–25).
Remarks. Being unable to locate any type specimens, Eyles (1973) inferred from the original description that
Dieuches jacobsoni, D. nudipes, D. schultheissi and D. villosulus probably belong to the genus Metochus.
Examination of the hereby designated lectotypes and paralectotypes led us to the same conclusion for all four
species. Similarly, D. kreyenbergi Breddin, 1906 and D. procericornis Breddin, 1908 (lectotypes of these were
already designated by Gaedike (1971)) represent valid species, both belonging to Metochus (Scudder 1962). All of
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the above species are valid. A revision of the genus is in preparation by the senior author; the identity of the widely
distributed and often mentioned M. uniguttatus (Thunberg, 1822) was recently clarified by Kondorosy et al.
(2014).
Dieuches nudipes Breddin, 1906
(Figs. 26–27)
Dieuches nudipes Breddin, 1906: 325. Syntypes: ♂, ♀, Ceylon [= Sri Lanka]: Puttalam, Nalanda, Anuradhapura and
Kanthaley; depository not stated.
Current status. Metochus nudipes (combination proposed tentatively by Eyles 1973: 37).
Type material examined. Lectotype (present designation): ♂, “Kanthaley / CeylonHorn”; pinned, right
antennal segments II–IV and left hind leg lacking (DEIC) (Figs. 26–28). Paralectotypes: with label identical with
that of lectotype (1 1 ♀ DEIC); “Puttalam / CeylonHorn” (1 DEIC); “Nalanda / CeylonHorn” (1 ♀ DEIC);
“Ceylon / W.Horn”, “Anu- / radhapura”, “Museum Leiden. [pr] / Dieuches [hw] / nudipes Bredd [hw] / det: [pr]
Blöte [hw]” [with black frame] (1 ♂ RMNH).
Remarks. Specimens from Java mentioned in the original description are not part of the type series because it
was explicitly expressed that they are doubtfully attributed to the species (ICZN 1999, Art. 72.4.1). For further
remarks see above under D. jacobsoni.
Dieuches schultheissi Breddin, 1906
(Figs. 29–30)
Dieuches schultheissi Breddin, 1906: 330. Holotype: ♀, [Indonesia:] NO Sumatra, Tebing Tinggi; depository not stated.
Current status. Metochus schultheissi (Breddin, 1906), combination proposed tentatively by Eyles (1973: 37).
Type material examined. Holotype: ♀, “N.O. Sumatra / Tebing-tinggi / Dr.Schultheiss.”, “Coll. Breddin”;
pinned, left antennal segment IV lacking (DEIC) (Figs. 29–31).
Remarks. Although the number of the examined specimens was not given, the singular case of the German
noun Stück in the original description (Breddin 1906: 71) (“…beim vorliegenden Stück” = “…in the available
specimen”) implicitly indicates that it was based on a single individual. The examined specimen is therefore
considered as the holotype of D. schultheissi fixed by monotypy (ICZN 1999, Art. 73.1.2). For further remarks see
above under D. jacobsoni.
Dieuches villosulus Breddin, 1906
(Figs. 32–33)
Dieuches villosulus Breddin, 1906: 327. Syntypes: ♂, ♀, Ceylon [= Sri Lanka]: Anuradhapura and Horrowapatam; depository
not stated.
Current status. Metochus villosulus (Breddin, 1906), combination proposed tentatively by Eyles (1973: 37).
Type material examined. Lectotype (present designation): ♂, “Ceylon / W.Horn”, “Anu- / radhapura”;
pinned, intact, left antenna oligomerous, with three segments (DEIC) (Figs. 32–34). Paralectotype: “Ceylon /
W.Horn”, “Horromapotam [sic]” [hw], Cat.N
o
[pr] / 1 [hw]”, “Museum Leiden. [pr] / Dieuches [hw] / villosulus
Bredd. [hw] / det: [pr] Blöte [hw]” [with black frame] (♀, RMNH)
Remarks. See above under D. jacobsoni.
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FIGURES 26–36. Type specimens of Rhyparochromidae species described by G. Breddin. Fig. 26, Dieuches nudipes Breddin,
1906, lectotype, dorsal view; Fig. 27, same, lateral view; Fig. 28, same, label; Fig. 29, Dieuches schultheissi Breddin, 1906,
holotype, dorsal view; Fig. 30, same, lateral view; Fig. 31, same, labels; Fig. 32, Dieuches villosulus Breddin, 1906, lectotype,
dorsal view; Fig. 33, same, lateral view; Fig. 34, same, label; Fig. 35, Poeantius brevicollis Breddin, 1907, lectotype, dorsal
view; Fig. 36, same, label. © DEIC. Scales in mm.
Poeantius brevicollis Breddin, 1907
(Fig. 35)
Poeantius brevicollis Breddin, 1907b: 207. Syntypes: ♂, ♀, Ceylon [= Sri Lanka]: Trincomalee and Negombo; depository not
stated.
Current status. Junior subjective synonym of Poeantius lineatus Stål, 1874; synonymized by Distant (1910: 75).
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Type material examined. Lectotype (present designation): ♀, “Negombo / CeylonHorn”; mounted on card,
left antennal segment IV, both fore legs, left mid and hind legs lacking (Hemi-00046, DEIC) (Figs. 35–36).
Paralectotype: “Trincomalee / CeylonHorn”; segments III–IV of left and II–IV of right antenna, left mid tarsi,
right mid and both hind legs lacking (1 ♀, Hemi-00045, DEIC).
Remarks. According to the original description some of the syntypes are males. Since no male could be
located it can be stated with certainty that some of the specimens (paralectotypes) remain undiscovered. The
synonymy of P. brevicollis and P. lineatus proposed by Distant (1910) is hereby confirmed.
Tri be Myodochini
Paromius robustior Breddin, 1907
(Figs. 37–38)
Paromius robustior Breddin, 1907b: 204. Syntype(s): ♀, Ceylon [= Sri Lanka]: Anuradhapura; depository not stated.
Current status. Junior subjective synonym of Paromius exiguus Distant, 1883; synonymized by Distant (1910:
50).
Type material examined. Lectotype (present designation): ♀, “Ceylon / W.Horn”, “Anu- / radhapura”,
“Paromius / exiguus / Dist.” [Breddin’s hw]; pinned, left mid leg apicad of middle of tibia lacking (Hemi-00042,
DEIC) (Figs. 37–39).
Remarks. The species was described from an unspecified number of specimens from Anuradhapura. A single
female in DEIC is apparently part of the original syntype series and it is hereby designated as the lectotype.
FIGURES 37–42. Type specimens of Rhyparochromidae species described by G. Breddin. Fig. 37, Paromius robustior
Breddin, 1907, lectotype, dorsal view; Fig. 38, same, lateral view; Fig. 39, same, labels; Fig. 40, Pamera recincta Breddin,
1901, lectotype, dorsal view; Fig. 41, same, lateral view; Fig. 42, same, labels. © DEIC. Scales in mm.
KONDOROSY & RÉDEI
112
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Zootaxa 3964 (1) © 2015 Magnolia Press
Pamera recincta Breddin, 1901
(Figs. 40–41)
Pamera recincta Breddin, 1901b: 59. Syntypes: ♂, ♀, Ecuador; type depository not stated.
Current status. Neopamera recincta (Breddin, 1901), combination proposed by Harrington (1980: 107).
Type material examined. Lectotype (present designation): ♀, “-Palmar—/ (Ecuad.) / R. Haensch S.”, “Coll.
Breddin”, “D E I / EBERSWALDE”; pinned, left antennal segment IV and apical half of right antennal segment IV
lacking; deposited in DEIC (Figs. 40–42).
Remarks. According to the original description the syntype series included male(s). Since no male could be
located it can be stated with certainty that some of the specimens (paralectotypes) remain undiscovered.
Acknowledgements
We are grateful to Stephan M. Blank (DEIC), Yvonne van Nierop and Willem Hogenes (RMNH), Jürgen Deckert
(ZMHB) and Michael D. Webb (BMNH) for providing access to types and other materials; to Tomasz Huflejt
(MZPW) and Karla Schneider (MLUH) for information about specimens deposited in the collections under their
care; to S. M. Blank and to J. Deckert also for the photos reproduced in this paper, and to S. M. Blank and an
anonymous reviewer for several valuable comments on the manuscript. This study received financial support from
the National Natural Science Foundation of China (grant no. 31472024) and the One Hundred Young Academic
Leaders Program of Nankai University to DR.
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... The German entomologist Gustav Breddin (1864Breddin ( -1909 described eleven species and two infrasubspecific taxa ("varieties") of Neotropical Scutelleridae (Hemiptera: Heteroptera) (Breddin 1903a(Breddin , 1903b(Breddin , 1903c(Breddin , 1904a(Breddin , 1904b(Breddin , 1906(Breddin , 1914. Type specimens of taxa described by Breddin are frequently difficult to trace primarily because he usually did not indicate the number and depository of the specimens studied by him, and in many cases he did not provide specimens with identification or type labels (Kondorosy & Rédei 2015). Gaedike (1971) and Eger (1990) reported the presence of type material of eight of the above mentioned eleven Neotropical scutellerid species in the Deutsches Entomologisches Institut, Eberswalde (now Senckenberg Deutsches Entomologisches Institut, Müncheberg) (SDEI) and the Institut Royal des Sciences Naturelles de Belgique, Brussels, respectively. ...
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Lectotypes are designated and documented for the following species and infrasubspecific taxa: Dystus villosus Breddin, 1904; Lobothyreus breviceps Breddin, 1914; Pachycoris torridus (Scopoli, 1772) var. laetissimus Breddin, 1906 (originally proposed as var. laetissima); P. torridus var. moestissimus Breddin, 1906 (originally proposed as var. moestissima); Polytes speculiger Breddin, 1914. The identities of the taxa in concern are clarified. The following new junior subjective synonyms are proposed: Pachycoris torridus (Scopoli, 1772) = P. torridus var. laetissimus Breddin, 1906, syn. nov., = P. torridus var. moestissimus Breddin, 1906, syn. nov.
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Types of Rhyparochromidae (Hemiptera: Heteroptera: Lygaeoidea) species described by Carl Peter Thunberg, deposited in the Museum of Evolution (formerly Zoologiska Institut), Uppsala University, Uppsala, Sweden, were reexamined and the taxonomic and nomenclatural problems that existed among those species discussed and resolved as required. Lectotypes are designated for Cimex caffer Thunberg, 1784, Lygaeus ater Thunberg, 1822, Lygaeus biguttatus Thunberg, 1822, and Pendulinus guttatus Thunberg, 1825. The lectotype of Pendulinus (now Metochus) guttatus is designated as neotype of Pendulinus (now Metochus) uniguttatus Thunberg, 1822; as a result the former name becomes junior objective synonym of the latter. The following taxonomic changes are proposed: Lethaeus ater (Thunberg, 1822), new combination (from Lygaeus); Migdilybs biguttatus (Thunberg, 1822), new combination (from Lygaeus) = Migdilybs furcifer Hesse, 1925, new subjective synonym; Metochus uniguttatus (Thunberg, 1822) = Metochus bengalensis (Dallas, 1852), confirmed subjective synonym = Metochus yeh (Dohrn, 1860), confirmed subjective synonym; Raglius alboacuminatus (Goeze, 1778) = Cimex caffer Thunberg, 1874, confirmed subjective synonym. Lethaeus barberi Slater, 1964 does not belong to Lethaeus Dallas, 1852 but currently it cannot be placed with confidence in any existing genus.
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Hemiptera from the Bombay presidency
  • E E Bergroth
Bergroth, E.E. (1915) Hemiptera from the Bombay presidency. Journal of the Bombay Natural History Society, 24, 170-179.
Die Hemipteren von Celebes. Ein Beitrag zur Faunistik der Insel
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Breddin, G. (1901a) Die Hemipteren von Celebes. Ein Beitrag zur Faunistik der Insel. Abhandlungen der Naturforschenden Gesellschaft zu Halle, 24, 1-215, 1 plate.
Neue neotropische Wanzen und Zirpen (Fortsetzung)
  • G Breddin
Breddin, G. (1901b) Neue neotropische Wanzen und Zirpen (Fortsetzung). Societas Entomologica, 16 (8), 59-60.