... However, changes in behavior may also be used as a bioindicator, as species may change their behavior and daily activities under altered conditions due to direct and indirect human disturbances (Sih, Stamps, Yang, McElreath, & Ramenofsky, 2011;Sih, 2013;Wong & Candolin, 2015;Fontúrbel, Candia, Malebrán, Salazar, González-Browne, & Medel, 2015;Costa, Madureira, & Zalmon, 2019). For example, population abundance of some crab species is commonly used as bioindicators of various human disturbances such as urbanization, mining, and contamination Jonah, Agbo, Agbeti, Adjei-Boateng, & Shimba, 2015;Schlacher et al., 2016;Wildsmith et al., 2009), and some of these species additionally alter burrowing behaviors in disturbed sites (Weis & Perlmutter, 1987;Bartolini, Penha-Lopes, Limbu, Paula, & Cannicci, 2009;Culbertson et al., 2007;Gül & Griffen, 2018a). Species may also alter their feeding habits (Griffiths et al., 2017;Jokimäki, Suhonen, Jokimäki-Kaisanlahti, & Carbó-Ramírez, 2016) and trophic interactions (Costa, Tavares, Suciu, Rangel, & Zalmon, 2017;Gray, Baldauf, Mayhew, & Hill, 2007) in areas with human disturbance, and thus these changes in behavior and daily activities can result in changes to the energy balance and physiological state (Chandurvelan, Marsden, Glover, & Gaw, 2015;Spellerberg, 2005). ...