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Hepatics of Europe including the Azores: An annotated list of species, with synonyms from the recent literature

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... polymorpha R.M. Schust. was described by Schuster (1969) from Greenland, and was later elevated by Schljakov (1979) and Grolle (1983) to the species rank as Saccobasis polymorpha (R.M.Schust.) Schljakov and Tritomaria polymorpha (R.M.Schust.) ...
... The first interpretation has recently been generally accepted (Söderström et al., 2016, Hodgetts et al., 2020. However, as Schuster (1988: 110) correctly pointed out, both Schljakov (1979) and Grolle (1983) who accepted this treatment "neither has seen any of the original material". Moreover, as Schuster (1988) emphasized both Schljakov (1979) and Grolle (1983), elevated ssp. ...
... However, as Schuster (1988: 110) correctly pointed out, both Schljakov (1979) and Grolle (1983) who accepted this treatment "neither has seen any of the original material". Moreover, as Schuster (1988) emphasized both Schljakov (1979) and Grolle (1983), elevated ssp. polymorpha on the basis of a supposed gemmae difference, but the idea of differences in the shape of gemmae in these taxa was erroneous (see discussion). ...
... Later, the heterogeneity within Pellia s.l. was revealed in the study of karyotype organization [8] and peroxidase isozyme polymorphism [9]. In combination with morphological distinctions, this allowed the subgenus Apopellia Grolle [10] to be segregated within the genus Pellia. ...
... The genus Apopellia (Grolle) Nebel & D.Quandt was separated from the genus Pellia Raddi by Schütz et al. [11] after integrative study with three chloroplast DNA markers of this ancient thalloid taxon. Plants with relatively long polymorphous slime hairs near the apex on the ventral side, thalli in cross section without thickening bands, perichaetia forming a complete ring and ciliate-laciniate mouth were previously considered as section Pellia [7] or as subgenus Apopellia [10]. In addition to the well-known A. endiviifolia (Dicks.) ...
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An integrative study of expanded sampling of Apopellia species, including the topotype of Apopellia megaspora, made it possible to clarify the taxonomic position and distribution of the species of the genus. The ITS1-2 and trnL-F sequence data were obtained for 36 molecularly tested specimens, including the topotype Apopellia megaspora, that together with data previously deposited in GenBank, support the treatment of Apopellia as a separate genus and A. alpicola as a distinct species, as well as radically change the idea on the distribution of the species of the genus. It is shown that A. megaspora is an American-Asian species with single records in Europe, whereas A. alpicola is a West-American-Eurasian species widespread in western North America and occurring scattered in Eurasia. Both species occur in the mountains of western North America and south Siberia. A. endiviifolia is widespread in Europe, scattered in Asia and so far not confirmed for America. The expanded sampling of Apopellia spp. allows us to clarify the morphological features of the species of the genus, and microphotographs illustrate the more-important morphological features.
... Anthoceros neesii (Fig. 2B) has a limited global distribution, being recorded only in the Czech Republic, Poland, Germany, and Austria (Proskauer 1958;Düll 1983;Grolle 1983;Düll and Meinunger 1989;Koła and Turzańska 1993;Dierßen 2001;Weddeling 2002;Manzke 2004Manzke , 2005Meinunger and Schröder 2007;Fischer et al. 2008;Teuber and Göding 2009;Koval and Zmrhalová 2010;Kučera 2011;Schlüsslmayr 2011;Schröck et al. 2014;Köckinger 2017;Pöltl et al. 2020). Given its distribution, A. neesii is classified as a Central European endemic. ...
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In 2023, field research in south-western Poland led to the noteworthy discovery of two hornworts: Notothylas orbicularis , a species previously unrecorded in this country, and the rediscovery of Anthoceros neesii for the Polish bryoflora. These findings are significant as they suggest a response to climate change, which is facilitating the range expansion of hornworts within Central Europe. Detailed descriptions of the new localities for both species are provided, highlighting the specific environmental conditions and habitats where they were found. Distribution maps for Notothylas orbicularis and Anthoceros neesii in Poland are provided, as well as SEM micrographs of spores. Additionally, a key to the identification of Polish hornwort species is also included. Furthermore, a model projecting the potential future spread of these hornworts within Poland and the broader Central European region is presented. This model considers climatic variables and habitat availability, offering insights into possible range shifts. This study contributes to the growing body of evidence that climate change is a driving factor in the redistribution of bryophytes.
... The inventory included the log, the uprooting part of the log, the snag (if present), and the major branches of the crown (if present). The nomenclature used follows Corley et al. (1981) and Corley and Crundwell (1991) for mosses, and Grolle (1983) for hepatics. Plagiochila porelloides and P. asplenioides were not separated and Plagiothecium species follow Frisvoll et al. (1995). ...
Article
Species composition and diversity of bryophyte communities occurring on dead beech trees were analyzed in five European countries (Slovenia, Hungary, The Netherlands, Belgium and Denmark). Altogether 1.009 trees were inventoried in 19 beech dominated forest reserves. The differences in the composition of bryophyte communities were high among countries and it overwhelms the effect of decay stages, which affect the species com-position at local scale. Well decayed trees were dominated by epixylic species in Slovenia, while in other countries the proportion of opportunistic bryophytes was much higher and in Atlantic countries uproot species are also common. In Slovenia hepatics, in Hungary and Denmark pleurocarps, and in The Netherlands and Belgium acrocarps were the most fre-quent. Diversity of communities differed considerably among regions. Slovenian sites were the hotspot of bryophyte diversity characterized by high species richness (both tree and country level) and a high fraction of rare and threatened species. This richness is most likely caused by the combination of high air humidity combined with a very high degree of natu-ralness of the Slovenian sites. The diversity of Hungarian stands was intermediate reflecting the relatively high naturalness of the study sites, as well as a rather continental climate sub-optimal for epixylic liverworts. Atlantic forests have deteriorated bryophyte communities on dead trees, dominated by a few opportunistic species. This is partly a result of habitat frag-mentation and past and recent management of forest. Most of the stands the amount and quality of dead wood is not suitable for species rich epixylic bryophyte communities.
... Para la nomenclatura se ha consultado Corley et al. (1981), Corley & Crundwell (1991), Grolle (1983) y Casas (1991. Se ha seguido el criterio de Hedenäs (1997) para Campylium s.l., el de Blom (1996) para el complejo de Schistidium apocarpum y el de Zander (1993) para la familia Pottiaceae. ...
Article
Se ha estudiado la flora briofítica de la región de Los Serranos region (NW de la Comunidad Valenciana, E de España). Se ha encontrado 187 táxones, 165 musgos y 22 hepáticas. Los táxones más destacados son: Didymodon umbrosus, Didymodon rigidulus var. icmadophyllus, Pterygoneurum crossidioides, Syntrichia calcicola, Syntrichia caninervis var. gypsophila, Schistidium agassizii, Schistidium atrofuscum, Schistidium robustum, Schistidium singarense, Orthotrichum vittii y Riccia crozalsii. The bryophytic flora of Los Serranos region (NW Valencian County, East of Spain) is studied. 187 taxa are listed, 165 mosses and 22 liverworts. The more remarkable taxa are: Didymodon umbrosus, Didymodon rigidulus var. icmadophyllus, Pterygoneurum crossidioides, Syntrichia calcicola, Syntrichia caninervis var. gypsophila, Schistidium agassizii, Schistidium atrofuscum, Schistidium robustum, Schistidium singarense, Orthotrichum vittii, and Riccia crozalsii.
... Frullania teneriffae (Fig. 2) is widely distributed in the Macaronesian archipelagos, as well as on the western coasts of the British Isles. Outside these regions, however, the species is concentrated in small, relatively isolated areas: the sierras of Cádiz (sierras of Ojén, Saladavieja and de la Luna) and Málaga (Sierra del Aljibe) in southern Spain (Guerra et al., 2003;Brugués & Sérgio, 2021), the Sierras of Sintra and Arrábida in Portugal (Sérgio et al., 2013), the Breton Finistère (Schumacker, 1988;Durfort, 2015), Faeroe Isles and Iceland (Grolle, 1983), the Tanger-Tetouan-Al Hoceima region (Djebel Musa mountain) in Morocco (Jelenc, 1955), and it has recently been reported in the humid north of Turkey (Ordu province, Özdemir & Batan, 2016). The new area detected in Galicia could be added to this list of small nuclei of occurrence, although it will be necessary to pay attention to all Frullania growing on the Galician coasts before reaching that conclusion. ...
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Se recopilan las novedades florísticas y nuevas especies de musgos descritas, que afectan al territorio del proyecto científico/editorial Flora Briofítica Ibérica y no se encuentranrecogidas en los volúmenes correspondientes, ya que fueron publicadas con posterioridad a la aparición de los mismos. Un total de 27 especies se han añadido a la flora de musgos de la península Ibérica y Baleares desde 2006. New records and new species of mosses, which affect the territory of the Flora Briofítica Ibérica project, are compiled. These taxa are not included in the corresponding volumes, since they were reported or described after the publication of their respective monographs. In all 27 species have been added to the moss flora of the Iberian Peninsula and the Balearic Islands since 2006.
... In general, the nomenclature follows Grolle (1983) or Stotler and Crandall-Stotler (1977). Collection numbers without a collector's name are the senior author's. ...
... The nomenclature of Musci follows Vitt (1982) and Ireland et al. (1987), and that of Hepaticae follows Stotler & Crandall-Stotler (1977) and Grolle (1983aGrolle ( , 1983b. Voucher Reserve, 1990). ...
... Specimens from the park borrowed from the herbaria of ALTA, CU and FH, have also been incorporated. In general the nomenclature follow Grolle (1983aGrolle ( , 1983b and Stotler & Crandall-Stotler (1977). The voucher collection numbers for each taxon are the author's. ...
... This gives a rough estimate of abundance in each of the 3 general collecting areas, ic. Cape Fife (areas 1-5 and 11-14, north end of park in Fig. 1), Mayer Lake (6-8, southern end of the study area) and Pure Lake (9, central part of the study area).In general, nomenclature follows Stotler & Crandall-Stotler (1977) and Grolle (1983aGrolle ( , 1983b ...
... and Schofield (1968 Stotler (1977) and Grolle (1983aGrolle ( , 1983b [hampeana (Nees) Schiffn.] --shaded soil. ...
... The nomenclature of vascular plants, mosses, hepatics and lichens follow Lid (1985), Corley et al. (1981), Grolle (1983) and Krog et al. (1980), respectively. ...
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A reference site for vegetational and ecological monitoring of low-alpine vegetation has been established in Gutulia National Park, Engerdal, Hedmark, Norway. Fifty vegetation plots, each 1 m2, were distributed randomly along four open transects. Species abundance was recorded as frequency in 16 subplots. Each plot was supplied with measurements of 26 environmental variables. The first axes of DCA and LNMDS ordinations were closely similar. The first DCA axis was interpreted as a complex snow cover gradient. Several environmental variables varied along this gradient, among them unevenness, soil moisture and soil nutrient status. The cover of the bottom layer varied along the complex-gradient. DCA axis 2 was interpreted as a microclimatic gradient. Partitioning of the environmental· and spatial variation in species composition showed that the spatial variation in the data set was rather low. Monitoring alpine vegetation is considered important since alpine ecosystems may be particularly vulnerable to airborne pollution.
... (1991) et Grolle (1983). Celle des Diatomées a été réalisée à l'aide de Krammer & Lange-Bertalot (1986, 1988, 1991a, Krammer (1992) et Lange-Bertalot (1993). ...
... The nomenclature of the vascular plants followed EHRENDORFER (1973), of the mosses CORLEY et al. (1981), of the liverworts GROLLE (1983) and of the lichens WIRTH (1980). Different taxonomic concepts are adressed in the text, but for this paper, the author abstained from applying the accepted names of the EU-Nomen or Plant+Med database. ...
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Spring habitats in woodland and grassland areas are part of the fresh water system, marking the spots where ground water discharges and becomes surface water. Ecologically they are characterized by water temperatures that are relatively low in the summer and relatively high in the winter, supporting plant species that are tolerant of low temperatures around their roots at all times. Ground water, by nature, is poor in nutrients, so that net primary production is low. For this reason, the typical plant species in spring habitats do not suffer from strong competition. The most widespread communities of spring habitats belong to the Montio-Cardaminetea and contain only plant species with low demand on temperature and nutrient supply. Some 1,800 relevés, of which more than 900 were processed in vegetation tables, were the basis of this investigation into the vegetation of spring habitats in Germany. The comparison of the relevés lead up to a distinction on the high syntaxonomical level of orders, dividing spring vegetation in woodland areas, on one side, from spring vegetation in grassland habitats, on the other side. Spring vegetation in grassland habitats is subdivided into thee alliances with 11 associations a total of, whereas, spring vegetation in woodland areas comprises only one alliance with two associations. Relevant parts of the syntaxon descriptions are syntaxonomy and nomenclature, floristic characterization and differentiation. Furthermore, investigations of vegetation structure and ecology are presented to underpin the syntaxonomical differentiation. Further information is added on vegetation dynamics, distribution, economic importance, zoocenology, and nature conservation. For each alliance and for the class a vegetation table with constancy values is provided. In addition, tables with average ecological parameters describe the ecological conditions for the given syntaxa. An overview of the syntaxa and their diagnostic species simplifies the determination and shows the floristic peculiarities of Montio-Cardaminetea. Keywords: Montio-Cardaminetea, Germany, syntaxonomy, classification, vegetation structure, ecology, dynamics, distribution, biocenology, nature conservation.
... (2) Taxon: The nomenclature generally follows Grolle (1983) for hornworts and liverworts and Corley & al. (1981) and Corley & Crundwell (1991) for mosses. Some modifications were based on recent works: Gabriel (1994), Hedenäs (1992), Jovet-Ast (1986) and ECCB (in press). ...
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Fifty-seven bryophyte species, mostly Macaronesian endemics and threatened species in Europe were surveyed from natural and semi-natural areas on Terceira Island (Azores). Based on recent field data and previous information, their conservation status is briefly discussed. The distribution of these selected species was mapped using a I km x I km UTM grid. The richest species squares, considered hot-spots, were used as indicators to define sites ofspecial interest for bryophytes that should be included in conservation programmes.
... Niewielki fragment południowo-wschodniej części terenu podlega Nadleśnictwu Ujsoły. , a nazewnictwo wątrobowców -za "Hepatics of Europe including the Azores: an annotated list of species, with synonyms from the recent literature" (GROLLE, 1983). Gatunki podlegające ochronie podano według aktualnie obowiązującego rozporządzenia Ministra Środowiska z dnia 9 października 2014 r. w sprawie ochrony gatunkowej roślin (Dz. ...
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The Ochodzita Massif located in the south part of the Silesian Beskid is a subject to heavy anthropopression. A large part of the landscape of this area are fields and meadows. Dense development of buildings lower parts of the Massif, the current agricultural management and the existence of object connected with tourism are conductive to the development of ruderal plants. There is no form of nature protection. In the years 2010–2012 the phytosociological research was conducted in the Ochodzita massif aiming at recognition of the diversity of plant communities and the identification of the areas of eligibility for legal protection in terms of vegetation. 43 non-forest plant associations, 21 non-forest communities, 8 forest plant associations and 3 forest communities were distinguished as a result of the analysis of the collected material. 13 habitats of European importance were identified within the Ochodzita Massif, including 4 priority habitats. They are represented by 16 plant communities. Numerous species being under legal protection were found in the study area. Bearing in mind the beauty of the landscape and the presence of protected species and natural habitats of European importance, it was proposed to create protected areas in the form of 3 nature and landscape complexes: "Ochodzita", "Koczy Zamek" and "Nad Czadeczką", 1 ecological site "Kosarzyska” and 1 nature reserve called "Czerna".
... Die Flächengrößen liegen im oberen Bereich der für Graslandaufnahmen empfohlenen Spanne (DIERSCHKE 1994: 151). Folgende Kürzel werden in den Tabellen verwendet: -mehr als ein / zwei Mahdtermine im Jahr QW, SW, ZW -Beweidung durch Rinder, Schafe, Ziegen M1W, M2W -ein-/ zweimalige Mahd mit Vor-oder Nach-Beweidung Die Nomenklatur der Phanerogamen folgt K ER G U ELEN (1993), jene der Kryptogamen CORLEY et al. (1980), CORLEY & CRUNDW ELL (1991) [Laubmoose] und GRO LLE (1983) [Leber­ moose]. ...
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The oat-grass-meadows of the Aigoual mountain region (France, Gard and Lozère) are described in detail, based on 101 phytosociological relevés. A comparison of all published relevés of oat-grass-meadows in the Languedoc region shows that the meadows of the Aigoual mountain valleys and the meadows of Central Europe belong to the same association, the Arrhenatheretum elatioris Braun 1915. The syntaxonomical position of the mediterranean meadows growing near the city of Montpellier is defi­ned: they represent an association apart, the correct name of which is Gaudinio fragilis-Arrhenathere­tum elatioris Br.-Bl. et al. 1952 em. The Trifolium molinerii-Arrhenatherum elatius-community is also described. The meadows belon­ging to this community occupy an intermediate position between the Arrhenatheretum, which occurs mainly at lower altitudes, and the Polygono-Trisetion-meadows, which are restricted to higher altitudes. The regional differences in the floristically-defined subgroups of the associations are graphically com­pared with regard to altitude and soil-acidity.
... La nomenclature employée repose essentiellement sur les publications de référence dans ce domaine, à savoir Corley et al. (1981) et Corley et Crundwell (1991 pour les mousses et Grolle (1983) et Grolle et Long (2000) pour les hépatiques. D'autres publications, notamment Koperski et al. (2000) et Ochyra et al. (2003) sont également utiles dans l'évaluation de certains problèmes nomenclaturaux spécifiques. ...
... I Nomenclature selon Corley et at. (1981), Corley et Crundwell (1991) pour les mousses, selon Grolle (1983) pour les hçatiques et selon Purvis et al. (1992) Tableau I : Concentrations de quelques éléments traces métalliques dans les sols (< normaux > et dans la lithosphère (d'après Fagot, 1993). Fox & Tao' 1981 Tableau 3: Sites pollués recensés dans différents pays. ...
Thesis
Cette recherche est axée sur l'écotoxicité des éléments traces métalliques à différents niveaux d'organisation des Bryophytes. Ainsi, dans une approche écotoxicologique pluridisciplinaire de la population métallique, nous abordons chez les Bryophytes, considérés comme des sentinelles de l'environnement, des aspects communautaires, physiologiques et cellulaires. Nous avons pu ainsi établir une échelle de diagnostic de la contamination des sols par l'observation des communautés bryophytiques et bryolichéniques terricoles. Nous avons mis au point un bryocapteur des retombés atmosphériques à l'aide de Ceratodon purpureus. Cette Bryophyte ubiquiste présente une certaine capacité à tolérer les fortes concentrations en éléments traces métalliques mais arrive également à se développer sur des sols non contaminés. L'exposition de ce bryocapteur à des retombées en Cd, Pb et Zn (à proximité de Metaleurop, une usine de métaux ferreux située à Noyelles-Godault, Pas-de-Callais, France), nous a permis d'observer les réponses biologiques de cette Bryophyte par l'intermédiaire de biomarqueurs de stress oxydant. Le stress oxydant causé par les éléments traces métalliques au niveau cellulaire provoque la mise en place de mécanismes de compensation tels que l'augmentation de l'activité des superoxyde dismutases et de la biosynthèse du glutathion. Ce phénomènes sont très peu étudiés en bryologie et permettent la mise en place d'outils de diagnostic précoce de la présence et des effets de polluants dans l'environnement. Cette recherche s'est effectuée dans le cadre du programme de recherches concertées, inscrit dans le contrat de plan Etat-Région 1994-1999
... Synonymous names were verified in accordance with L.O.Tasenkevych [17]. Latin names and authors of mosses and liverworts are given according to Corley M.F.V. et al. [24], Grolle R. [27]. ...
Article
Багаторічні дослідження показали, що флора української частини регіону Розточчя налічує 1342 види рослин, на території новоствореного біосферного заповідника «Розточчя» виявлено 1112 видів судинних рослин і 210 видів мохів та печіночників. На підставі аналізу ботанічних матеріалів, зібраних у заповідних об’єктах, що входять в українську частину біосферного заповідника «Розточчя», визначено 144 види судинних рослин, які вимагають охорони, моніторингу за станом популяцій, і навіть реінтродукції. Проте на території новоствореного біосферного заповідника для охорони рідкісних видів рослин використовують тільки пасивні методи, які в кінцевому підсумку можуть призвести до зникнення видів з неактивною життєвою стратегією. У заповіднику повинні охоронятися цілі екосистеми, особливо ті, до складу яких входять рідкісні види рослин. Для потреб створення білатерального заповідника ми провели інвентаризацію рослинності з використанням методу Браун-Бланке та виділенням об’єктів мережі Natura 2000. Науковою основою збереження біорізноманітності регіону Розточчя є созологічна оцінка та категоризація об’єктів охорони згідно міжнародних природоохоронних стандартів. Для оцінки статусу раритетних таксонів та синтаксонів використано систему фітосозологічних індексів, розроблену J. Čeřovský (1977). За допомогою аутфітосозологічних індексів проведена созологічна оцінка видів рослин, що потребують охорони на Розточчі. Аутфітосозологічний індекс вище середнього мають 170 видів, синфітосозологічний індекс вище середнього мають 67 асоціацій рослинності. Проведений аналіз не тільки засвідчив високу репрезентативність фітобіоти регіону у європейській шкалі, а й створив систему базових критеріїв для підбору територій-ядер екомережі України. Наші спостереження показали, що функції і завдання білатерального заповідника не завжди можуть бути повністю реалізовані через розбіжності в природоохоронному законодавстві Польщі та України і можливі місцеві заходи, що проводяться на території лише однієї частини заповідника. Організація біосферного заповідника «Розточчя» і його подальше функціонування вимагають значної корекції двосторонніх юридичних документів та природоохоронних заходів щодо цієї території. Для правильного функціонування заповідника необхідно розробити загальну стратегію для збереження природи цілого регіону.
... En los sistemas de clasificación de Schuster (1951) el género Anastrophyllum se ubicó en la familia Lophoziaceae; mientras que en Grolle (1983), la familia Lophoziaceae se trató como subfamilia (Lophozioideae) de Jungermanniaceae. En los estudios moleculares de Schill et al. (2004) y De Roo et al. (2007 no se encontró relación entre las familias Lophoziaceae -Jungermanniaceae. ...
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En el estudio taxonómico del género Anastrophyllum para Colombia se registran cinco especies. A. auritum, A. nigrescens, A. piligerum, A. stellatum y A. tubulosum. El género Anastrophyllum presenta pigmentación pardo-rojiza, rojo-negruzca, ocasionalmente verde; tallos con corteza y médula diferenciadas; ramificación terminal tipo Frullania, lateral tipo Plagiochila y ventral tipo Bazzania; hojas bífidas, cóncavas, asimétricas, margen entera; células con trígonos pequeños a conspicuos. Dioicas; perianto emergente, cilindrico, plegado desde la base o solamente en la parte distal, con la boca hialina. Anastrophyllum leptodictyon es pasado a la sinonimia de A. auritum. Para cada una de las especies se proporciona una descripción detallada, ilustraciones, mapas de distribución y una clave donde se destacan principalmente caracteres del gametofito.
... * Видові назви печінкових мохів наведені за системою, прийнятою у "Hepatics of Europe including the Azores; an annotated list of species, with synonyms from the recent literature"[9] та "Зведеному списку антоцеротів та печіночників України"[1], листяних -за "Mosses of Europe and the Azores; an annotated list of species, with synonyms from the recent literature"[7]. ...
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... * Видові назви печінкових мохів наведені за системою, прийнятою у "Hepatics of Europe including the Azores; an annotated list of species, with synonyms from the recent literature"[9] та "Зведеному списку антоцеротів та печіночників України"[1], листяних -за "Mosses of Europe and the Azores; an annotated list of species, with synonyms from the recent literature"[7]. ...
... --+ + + П р и м і т к а . Видові назви печіночних мохів наведені за системою, прийнятою у "Hepatics of Europe including the Azores and annotated list of species, with synonyms from the recent literature" [27], листяних -за "Mosses of Europe and the Azores; an annotated list of species, with synonyms from the recent literature" [24]. робних, так і мезо-й олігогемеробних ступенів окультуреності [23]. ...
Article
En el presente trabajo se corrigen 4 citas anteriores y se aportan nuevas localidades de 51 especies. Dos de los táxones se citan por primera vez en La Rioja.
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We investigated the species richness, endemism, and similarities of the bryofloras on ten islands and archipelagos in the Atlantic Ocean to determine the floristic relationships among them using presence/absence. Cluster analysis was used to determine similarities among the bryofloras of ten islands and archipelagos. We examined the bryophyte species to address the following questions: (1) How many bryophyte species are there in the total area and on each island and archipelago? (2) Do mosses and liverworts share patterns of diversity? (3) What are their species distribution patterns? (4) How many endemic species are there on the islands? (5) Is there a high similarity among those bryophyte floras? We encountered 1,647 species, 325 genera, and 204 families, with the richest families (Lejeuneaceae, Lophocoleaceae, Plagiochilaceae, Brachytheciaceae, Bryaceae, Fissidentaceae, and Pottiaceae), representing 35% of all species. Over 12% of the bryophytes are African elements, 8% have global distributions, approximately 3% are Macaronesia elements, 5% are endemic, and 34% are included in other distributions. We present a checklist of 1,647 bryophyte species, of which 91 are endemic. Bryophyte richness and diversity differed significantly among the ten islands and archipelagos studied, and their low similarities illustrate their high degrees of heterogeneity. This paper is the first evaluation of bryophyte species on ten islands and archipelagos of the Atlantic Ocean, giving insight into their species richness, endemism and similarities, and providing information for analyzing species turnover, migration, invasion, etc., representing an increase in our general knowledge of the bryofloras of oceanic islands and archipelagos, contributing to the targets of the Global Strategy for Plant Conservation.
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A synthesis of the current knowledge on the bryophyte flora of the Natural Park of Serra da Estrela (PNSE) is presented. This study is based on 7500 specimens, from old records since 1804 but mainly on material collected recently. Three hundred and eighty three taxa (284 mosses and 99 hepatics) are reported from the area. Thirty three taxa are first reports to Beira Alta province, and eighteen species are new to the Natural Park. Data on the distribution of bryophyte species are updated, and distributional patterns of bryophytes in the Park are analysed. PNSE is the exclusive area of occurrence for 37 taxa in Portugal, which evidences the importance of the bryophyte flora in this area. Seven phytogeographical elements are recognized, among which the most represented ones are the oceanic, temperate and boreal elements. Potential threats to the bryoflora in this area are discussed.
Article
Ramudaria is a monospecific genus well-characterized by the longitudinal, serially arranged lamelliform strips on the surface of the stem. This character seems to be unique amongst the leafy liverworts. Ramudaria lamellicaulis is so far known only from Arunachal Pradesh and Sikkim in the Eastern Himalaya, India. The species is reported here for the first time for China. The locality recorded here for China is the northernmost and highest in altitude. A description and illustrations of R. lamellicaulis based on the Chinese specimen are provided, along with a distribution map. A discussion about the status of such a morphologically remarkable genus is presented and an author citation of Ramudaria is corrected.
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Forty mosses and twenty-six hepatics are reported for the first time from Fuerteventura, and fifteen mosses and nine hepatics are reported for the first time from Lanzarote, among them Gigaspermum mouretii Corb. which is new to Macaronesia. All taxa recorded so far are listed and their distribution outlined. Distribution maps are given for some of the more common species. The zonation of bryophytes is considered, and a survey of their phytogeographical affinities is given. some ecological factors are discussed, and it is suggested that the bryophyte flora of these two islands is primarily restricted by climatic factors.
Article
Introduction Many taxa have been added to the bryophyte flora of Britain and Ireland since the publication of the previous checklist in 2008. This has coincided with numerous taxonomic and nomenclatural changes in the European flora, especially as a consequence of molecular sequencing. With the publication of a new European checklist in 2020 it is now appropriate to provide an updated and annotated list for Britain and Ireland. Methods A working group was convened to consider and reach a consensus on taxonomic and nomenclatural matters arising from the European checklist. Special consideration was given to the citation of the authors of names. Taxa reported as new to Britain and Ireland since the previous checklist in 2008 were incorporated. Key results A new checklist of bryophytes (hornworts, liverworts and mosses) is presented. Based on the taxonomy adopted in the checklist, the bryophyte flora of Britain and Ireland comprises four hornworts (Anthocerotophyta), 299 liverworts (Marchantiophyta), and 795 mosses (Bryophyta) at species rank, with 42 additional infraspecific taxa.
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ABSTRACT: Portugal continental wetland aIder forests. Portuguese alder forests that occur in permanent riversides are included in OsmundoAlnion alliance of the Populetalia albae order, and Querco-Fagetea class. The mesomediterranic and supra-mediterranic aIder forests have strong continental character, with their optimum at the upper and middle courses of Carpetano-Ibérico Leoneses rivers. These formations are c1assified as Ga/io broteriani-Alnetum glutinosae association, while termo-mediterrranic alders, with high atlantic influence are included in Scrophulario scorodoniae-Alnetum glutinosae. These ones are found in the whole portuguese terrritory of Luso-Extremadurense and Ribatagano-Sadense provinces. During the fieldwork, it was confirmed that in the Nabão hydrographic basin the alders and willow grow in some riversides with flat margins permanently flooded, thus forming swamps. Conceming this it was proposed to include these yards inAlnetea glutinosae class, Alnetalia glutinosae order, Alnion glutinosae alliance. These formations are 15-20 meters tall and grow on soils with an organie horizon reaehing 40 em deep and a pH between 4,9 and 5,8. The relevés were earried out from November of 1999 to June of 2000 in Caxarias and Seiça riversides following the phytosoeiologieal methodology and were eompared with different lberie alders relevés ineluded in Querco-F agetea and Alnetea Glutinosae. Data analysis was supported with TWINSPAN c1assifieative method. This elassifieation eonfirrned the similarity between the relevés and the type relevé Carici lusitanicae-Alnetum glutinosae T.E. Díaz & F. Prieto 1994 (n" 17). On the other hand, the relevés earried out in Agualva de Baixo (Águas de Moura) and Marateea (n" 18, 19, 20) by COSTA et ai. (1996) previously identified as Scrophulario scorodoniae-Alnetum glutinosae, are also ineluded in the same group. Reeently, NETO eonfirrned the oeurrenee of swamp alders in Sado hydrographie basin, whieh grow in less flooding eonditions eompared with Carici lusitanicae-Salicetum atrocinereae C. Neto, Capelo, J.C. Costa & M. Lousã 1996.
Technical Report
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Dans le présent travail sont étudiés les groupements végétaux (syntaxons hytosociologiques) des sols acides appartenant aux formations dominées par les ligneux bas et moyens, communément appelées matorrals, maquis ou landes. A partir de publications récentes de référence, sont en premier lieu exposés les principes méthodologiques sur lesquels se base l’ensemble de l’étude. Une première analyse critique des publications spécialisées, sous forme de tableau synthétique des fréquences, met en évidence un certain nombre de syntaxons, de rang inférieur (associations) ou supérieur (alliances, ordre, classe), paraissant superflus. Il apparaît clairement que les classes regroupant les maquis et les landes nécessitent une révision moderne avec vision d’ensemble. Ensuite sont étudiés 165 relevés effectués durant les mois de mai et juin 2006 dans le secteur des vallées cévenoles du Parc national des Cévennes, sur la base d’un échantillonnage stratifié, orienté selon la ou les espèce(s) ligneuse(s) dominante(s) (typologie « Code CORINE Biotopes »). Deux approches différentes sont testées : la première, dite « sigmatiste » prend en compte tous les végétaux rencontrés dans une surface donnée ; la seconde, dite « synusiale » distingue les différents types biologiques (cryptogames, herbacées annuelles, herbacées vivaces, ligneux) et les traite séparément. Il est démontré que l’approche synusiale est plus appropriée pour l’étude des matorrals et landes, groupements composés de synusies inter-indépendantes et réagissant à des paramètres stationnels différents. Les relevés sont d’abord analysés par des méthodes statistiques (Analyses factorielles des correspondances). Les relevés ainsi pré-classifiés sont ensuite triés par affinité floristique. Sont ainsi distinguées 20 unités de végétation définies de façon soit positive, par une ou plusieurs espèces différentielles, soit négative, par l’absence de certaines espèces. Un groupe de relevés dépourvus d’espèces différentielles n’a pu être attribué à un quelconque syntaxon. Les unités sont avant tout différenciées en fonction de l’altitude, de la fertilité des stations et du stade dynamique. La classification obtenue est justifiée par un tableau synthétique des fréquences. Les unités distinguées sont comparées avec les associations décrites en littérature, également sous forme d’un tableau synthétique des fréquences. Finalement, l’intérêt patrimonial des différents groupements est évalué et hiérarchisé. Ce sont trois habitats typiquement méditerranéens qui arrivent en tête du classement, à savoir les « matorrals à bruyère sur suintements », les « matorrals à Cistus salviifolius » et les « matorrals à Cistus populifolius ».
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The Authors present a study of the bryological flora of some localities of Aspromonte (Calabria). A list of 128 taxa is given: 32 liverworts and 96 mosses; 20 species (5 liverworts and 15 mosses) are new findings for the Calabria region.
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A widely accessible list of known plant species is a fundamental requirement for plant conservation and has vast uses. An index of published names of liverworts and hornworts between 2015 and 2016 is provided as part of a continued effort in working toward maintaining an updated world checklist of these groups. The list herein includes 64 higher taxon names, 225 specific names, 35 infraspecific names, two infrageneric autonyms and 21 infraspecific autonyms for 2015 and 2016, including also names of fossils and invalid and illegitimate names. Thirty-three older names omitted in the earlier indices are included.
Article
A phylogenetic arrangement of the 130 liverwort genera, comprising 582 species, that occur in North America north of Mexico is followed by an alphabetical synopsis of genera, species, subspecies, and varieties currently recognized. The treatment of each taxon includes pertinent synonyms, excluded names, and explanatory comments regarding currently accepted changes in taxon names and authors. Such updating has required making nine new combinations, as follows: Fuscocephaloziopsis connivens (Dicks.) Váňa & L. Söderstr. var. bifida (R. M. Schust.) Stotler & Crand.-Stotl., comb. nov.; F. connivens (Dicks.) Váňa & L. Söderstr. var. compacta (Warnst.) Stotler & Crand.-Stotl., comb. nov.; F. pleniceps (Austin) Lindb. var. sphagnorum (C. Massal.) Stotler & Crand.-Stotl., comb. nov.; Isopaches bicrenatus (Schmidel ex Hoffm.) H. Buch var. immersus (R.M. Schust. & Damsh.) Stotler & Crand.-Stotl., comb. nov.; Mesoptychia badensis (Gottsche ex Rabenh.) L. Söderstr. & Váňa var. apiculata (R. M. Schust.) Stotler & Crand.-Stotl., comb. nov.; Neoorthocaulis hyperboreus (R. M. Schust.) L. Söderstr., De Roo & Hedd. subsp. helophilus (R. M. Schust.) Stotler & Crand.-Stotl., comb. nov.; Schistochilopsis grandiretis (Lindb. ex Kaal.) Konstant. subsp. proteidea (Arnell) Stotler & Crand.-Stotl., comb. nov.; Tritomaria capitata (Hook.) Stotler & Crand.-Stotl., comb. nov.; T. laxa (Lindb.) Stotler & Crand.-Stotl., comb. nov. The basionym citations for these new combinations can be found in the text treatments of these taxa. Thirty-four new heterotypic synonyms, distributed among 27 accepted taxa, have been proposed, primarily as a consequence of nomenclatural updating (see Appendix 1 for a list). Type species are indicated for all genera and basionyms are included, where appropriate. Distributional data, based primarily on literature reports, are also provided for each infrageneric taxon.
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This article reports the occurrence of the leafy liverwort Jungermannia gollanii Steph. from the Sohelwa forest in Sravasthi District located in Terai region of the state of Uttar Pradesh, India. The plants are found growing at low altitude. The species is characterized with creeping, simple plants bearing rhizoids arising ventrally all along stem, succubous and dorsally secund leaves, a non-beaked, long exerted, plicate perianth with contracted mouth and a lacking perigynium. The plants of this new locality have been retained under subgenus Luridae (Spruce) Amak. of genus Jungermannia L.
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Jungermannia leiantha Grolle n. sp. is proposed for J. lanceolata auct. (non L. 1753). The author has selected as type (lectotype) of J. lanceolata L. emend. Müller: 'Dill. musc. 486, t. 70, f. 10, Fig. A' [Herb. fol. 156, n. 10; caespites duo ad sinistrum; J. riparia Tayl. c. coles.; Oxford (Dillenius-Herbar)]. J. lanceolata L. emend. Müller is put in synonymy of J. tristis Nees (1836) beceause of art. 69 of I.C.B.N. (1961). J. lanceolata L. emend. Müller is established as lectotype of the genus Jungermannia L. (1753) emend. Dum. (1835). This typification enables to retain the customary use of Jungermannia L. and makes conservation of Solenostoma Mitt. (1865) against Gymnoscyphus Corda (1835) unnecessary, since both names become synonyms of Jungermannia L. emend. Dum.
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Plagiochila britannica is described as a new species. It differs from P. porelloides and P. asplenioides in many characters, especially its often more narrowly oval leaves with a more variable, often truncate to bilobed apex and larger cells, its wider antical stem cells and its diploid chromosome number. Differences from P. arctica var intermedia include its more variable leaf apex, generally more dentate leaves often with longer teeth, the greater maximum size of its leaf cells and its wider antical stem cells. Male and female inflorescences and sporophytes are known in P. britannica, which favours base-rich substrata and has been found in Wales, N. England, Scotland and N. Ireland. A key to the nine species of Plagiochila in the British Isles is included.
Article
Key to the genera Marchesinia, Dicranolejeunea and Lopholejeunea. D. johnsoniana (Mitt.) comb.nov. from Macaronesia is figured and described in detail. The South African Marchesinia chrysophylla is transferred to Dicranolejeunea and is regarded as conspecific with D. atra (Mitt.) Vanden Berghen, syn.nov. from Central Africa.
Article
Fossombronia pusilla is shown to be represented by two varieties based on differences between the gametophyte and the sporophyte; var. decipiens is reduced to synonymy with the widespread var. pusilla; a new variety, maritima, is described from heathy coastal habitats in southern and south-western regions of the British Isles. F. loitlesbergeri is deleted from the British list following an examination of authentic gatherings; Cornish material is referable to F. pusilla var. maritima. F. crozalsii, known only from two British localities, has been refound recently in Wiltshire; Cornish records are referable to F. pusilla var. maritima. An examination of the type specimen of F. mittenii indicates that it is most probably an abnormal form of F. wondraczekii. A comparison of material of F. husnotii with British F. husnotii var. anglica shows that the latter cannot be maintained as a distinct taxon.
Article
Part I deals briefly with the application of the Code rules to family and subfamily names.In Part II full references are given for all family and subfamily names of Hepaticopsida, some with nomenclatural or taxonomic notes added. There are three lists, each in alphabetical order: A. legitimate names (78 f., 62 subf.), B. illegitimate names (21 f., 10 subf.), and C. invalid names (only those derived from generic names; 17 f., 11 subf.); in total 116 family and 83 subfamily names. Five family names (Choneoleaceae Schust., Conocephalaceae K. Müll., Exormothecaceae K. Müll., Oxymitraceae K. Müll., Perssoniellaceae Schust.) and three subfamily names (Allisonioideae Schust., Cololejeuneoideae Herz., Odontosehismatoideae Buch) are validated by Latin diagnosis. Five subfamily names (Hygrobielloideae (Joerg.) Schust., Isotaehidoideae (Hatch.) Grolle, Makinooideae (Nakai) Grolle, Mastigophoroideae (Nees) Grolle, Pallavicinioideae (Migula) (Grolle) are proposed by change of rank, whereas five other subfamily names (Acromastigoideae Grolle, Blepharostomatoideae Grolle, Cyathodioideae Grolle, Lethocoleoideae Grolle, Notothyladoideae Grolle) are newly proposed.Part III is a taxonomic arrangement of the hepatic families and subfamilies. For each of the five orders of Hepaticopsida the accepted families (sixty-two in total) are listed alphabetically with full synonymy : Anthocerotales (1), Marchantiales (16), Metzgeriales (8), Calobryales (2), Jungermanniales (35). The accepted subfamilies with their synonyms are added to each family.
Article
There are two groups of Pellia peroxidase isozyme phenotypes, one composed of P. epiphylla, P. neesiana and P. borealis, the other of P. endiviifolia from Europe, P. endiviifolia from Japan and P. megaspora. The phenotypes within the second group differ supporting the suggestion that there are three taxa present.
Article
The genus Pellia, with three currently recognised species, includes a relatively primitive taxon, P. endiviifolia, in which the perichaetium is high and incised, with an enclosed calyptra. It is dioecious, produces anthocyanin wall pigments in exposed sites, reproduces asexually by repeatedly furcate, narrow fragmenting ‘autumnal innovations’, is freely branched and has relatively small spores. A species previously confused with this taxon, and here described as P. megaspora, is an endemic boreal American species, found from New York, Connecticut and Massachusetts north to Newfoundland and west to Minnesota. The plant has a tall perichaetium that is strikingly incised-ciliate to lacerate. It is dioecious but does not produce wall pigments and autumnal innovations, is simple or oncefurcate and has extremely large spores. Cytological and biochemical data support the specific recognition of P. megaspora. Biochemical data also suggest that Japanese populations currently assigned to P. endiviifolia are distinct and field observation suggests that these plants may represent an autonomous subspecies.
Article
The marked differences between Plagiochila spinulosa (Dicks.) Dum. and P. killarniensis Pears. pointed out by Paton (1977) are confirmed and added to by further ones. As a result the two species are placed in different sections Spinulosae and Zonatae, respectively. For each of the two species the full synonymy including new synonyms and new lectotypifications and a comprehensive list of studied specimens and a distribution map are presented. In appendix, P. porelloides (Torrey ex Nees) Lindenb. is recorded from Madeira, a species new to Macaronesia.
Article
A new species, Fossombronia fimbriata, is described, which has affinities with F. incurva but is characterized by unique deeply dissected fimbriate-lobed leaves and pseudoperianths.
Article
Anthoceros punctatus var. cavernosus Prosk. homon. illeg. is renamed A. agrestis Paton, redescribed, illustrated and compared with A. punctatus L. (A. husnotii Steph.). The thalli and sporophytes are usually smaller in A. agrestis than in A. punctatus and the antheridial bodies are significantly smaller. World distributions and brief descriptions of British habitats are included for both species showing that A. agrestis is more restricted than A. punctatus in its distribution and ecology.
Article
Anthoceros fuciformis Mont. (syn. nov. A. stuhlmanni Steph.), Jungermannia renauldii Steph., Plagiochila boryana Gottsche ex Steph. and P. drepanophylla Sande Lacoste are reported from Africa, and other extensions of range within Africa reported for Antheros incurvus Steph., Isotachis conistipula Steph., Notoscyphus belangerianus (Lehm.) Mitt. and Tylimanthus rnwenzorensis S. Arnell (syn. nov., T. aetiopica S. Arnell); for some of these descriptions and/or critical notes are provided. Bazzania papillosa S. Arnell is described. The date of publication of Radula macroloba is corrected to 1892, and the synonymy of Aneura congoana Steph. and A. congoensis Steph. amended.