During the Carboniferous Period around 350 million years ago, lyco-phytes dominated the landscape, comprising about 50% of the world's flora. Today, however, lycophytes are mostly inconspicuous plants, representing only about 0.5% of the flora. Studying the morphology of the lycophytes is integral to our understanding of land plant evolution and development because the lycophytes are sister to
... [Show full abstract] all other vascular plants. They represent an evolutionary lineage that has been distinct from other vascular plants, a lineage that has evolved growth forms and organs analogous to vascular plants and other organs that are homologous to those found in their sister group. As such, they provide a fascinating comparison for evolutionary developmental studies. The overall morphology of the lycophytes is relatively simple, with the sporophyte body composed of root, stem, microphyll, and sporangium. Three families comprise the present-day lycophytes: (1) Lycopodiaceae, (2) Isoetaceae, and (3) Selaginellaceae. Their unifying feature is the presence of a single sporangium per leaf, and this sporangium is located on the adaxial side of the leaf. Here, I briefly discuss the morphology and development of key lycophyte structures including meristems, sporangia, microphylls, rhizophores, and ligules.