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Clepsis trifasciata sp. n. with notes on some Lepidoptera Tortricidae from Kirgizstan

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Pasquale Trematerra at University of Molise
Pasquale Trematerra
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A new species of tortricid moth, Clepsis trifasciata sp. n., from Kadzhi- Sai, Issik-Kul province (Kirgizstan) is described. Externally the new species is characterised by its wing pattern, whilst the male genitalia is close to Clepsis violacea Razowski, 1966, which is widely distributed in the Mongolian steppe region, and to Clepsis laetitiae Soria, 1997, from Aranjuez (Central Spain). Moreover, some notes on some interesting tortricid moths which were collected in the mountainous localities of Kirgizstan are reported.
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Clepsis trifasciata sp. n. with notes on some Lepidoptera Tortricidae from Kirgizst
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J. Entomol. Acarol. Res.
Ser. II, 42 (1): 1-10 30 April 2010
P. TREMATERRA
Clepsis trifasciata sp. n. with notes on some Lepidoptera Tortricidae
from Kirgizstan
Abstract - A new species of tortricid moth, Clepsis trifasciata sp. n., from Kadzhi-
Sai, Issik-Kul province (Kirgizstan) is described. Externally the new species is char-
acterised by its wing pattern, whilst the male genitalia is close to Clepsis violacea 
Razowski, 1966, which is widely distributed in the Mongolian steppe region, and 
to Clepsis laetitiae Soria, 1997, from Aranjuez (Central Spain). Moreover, some 
notes on some interesting tortricid moths which were collected in the mountainous 
localities of Kirgizstan are reported.
Riassunto - Clepsis  trifasciata sp. n. con note su alcuni lepidotteri tortricidi del
Kirgizstan
Viene descritta Clepsis trifasciata sp. n., una nuova specie di Lepidoptera Tortrici-
dae, trovata in Kirgizstan a Kadzhi-Sai, nella provincia di Issik-Kul. La presenza 
di tre fasce alari è una caratteristica tipica della nuova specie, mentre negli apparati 
genitali maschili è simile a Clepsis violacea Razowski, 1966, presente nelle steppe 
della Mongolia, e  a  Clepsis laetitiae Soria,  1997,  raccolta  ad Aranjuez  (Spagna 
centrale). Inoltre si riportano brevi note faunistiche su alcuni interessanti tortricidi 
trovati in località montane del Kirgizstan.
Key words: Lepidoptera Tortricidae, Clepsis trifasciata, new species, Kirgizstan, 
fauna.
INTRODUCTION
In the present paper rare and interesting Tortricidae (Lepidoptera) are reported as 
found in amongst material collected in the mountainous localities of Kirgizstan, taken 
in 2000, by Sergei V. Churkin (Moscow, Russia), moreover, Clepsis trifasciata sp. n. 
is described. Externally, the new species is characterized by the typical wing coloration 
in the fasciae; the genitalia of C. trifasciata is similar to that of C. violacea Razowski, 
1966, which is widely distributed in the Mongolian steppe region, and to C. laetitiae
from Aranjuez, Central Spain (Razowski, 1979 and 2002; Soria, 1997).
Amongst other tortricids of particular interest were: Cochylimorpha pirizanica Ra-
zowski, 1963, Cochylimorpha woliniana (Schleich, 1868), Eana samarcandae Razow-
Journal of Entomological and Acarological Research, Ser. II, 42 (1), 2010
2
ski, 1958, Cnephasia cupressivorana (Staudinger, 1871), Cnephasia longana (Haworth, 
[1811]), Pandemis cinnamomeana (Treitschke, 1830), Celypha ermolenkoi Kostyuk, 
1980, Pelochrista medullana (Staudinger, 1879) and Eucosma lugubrana (Treitschke, 
1830) (Kuznetsov, 1989;  Razowski,  2002, 2003  and  2009;  Brown,  2005; Alipanah, 
2009; Trematerra, 2010).
LIST OF TAXA
LEPIDOPTERA TORTRICIDAE
SUBFAMILY TORTRICINAE
Tribe Cochylini
Phtheochroa inopiana (Haworth, [1811]) 
Material exaMined: 3 males, Kirgizstan, Suusamyr mts., Kokemeren r, 5km N Kyzyl-
Oi, 1800m, 24-25.06.2000,  S. Churkin leg.;  1 male and  1  female, Kirgizstan, E 
Suusamyr mts., Kovjuksu r., 1650m, Kyzyl-Oi, 26.07.00, S. Churkin leg.
distribution: Palaearctic: ranging from the Iberian Peninsula and the British Isles to 
Japan.
Cochylimorpha asiana (Kennel, 1899)
Material exaMined: 1 male, Kirgizstan, S. Issyk-kul L., Kadzhi-Sai v., 1.07.00, 1800m, 
S. Churkin leg.
distribution: SE Europe: Ukraine and Russia; NW Africa: Libya; Caucasus; Turkme-
nistan; from Kazakhstan to Central Asia and Mongolia; Iran; Afghanistan; China.
Cochylimorpha pirizanica Razowski, 1963
Material exaMined: 1 female, Kirgizstan, S Issyk-Kul L., Ottuk v, 1650, 29.06.2000, 
S. Churkin leg.; 2 females, (Kirgizstan) Fergana valley, 27km S. Osh, 19.07.2000, 
1200m S. Churkin leg.
distribution: Iran: Fars Province, Pirezan and Muk road.
Cochylimorpha woliniana (Schleich, 1868)
Material exaMined:  1  female,  Kirgizstan,  S.  Issyk-kul  L.,  Kadzhi-Sai  v.,  1.07.00, 
1800m, S. Churkin leg.
distribution: Known from Europe, mainly from its southern and middle parts, S Russia.
Phalonidia contractana (Zeller, 1847) 
Material exaMined: 1 male, Kirgizstan, S. Toktogul L., 15km NE Karakul v., 1300-
1400m, 17.06.2000, S. Churkin leg.; 2 males, Kirgizstan, South Chatkal, 5km. E 
3
P. Trematerra: Clepsis trifasciata and notes on some Tortricidae from Kirgizstan
Aatun v., Karasu r., 1350m, 19.06.2000, S. Churkin leg.
distribution: Widely distributed in Southern and Central Europe, found also in Asia 
Minor, Israel; Iran; Afghanistan; Kashmir; China (Xinjiang).
Agapeta hamana (Linnaeus, 1758) 
Material exaMined: 1 female, Kirgizstan, S Issyk-Kul L., Ottuk v, 1650m, 29.06.2000, 
S. Churkin leg.; 1 male and 1 female, Kirgizstan, E Suusamyr mts., Kovjuksu r., 
1650m, Kyzyl-Oi, 26.07.00, S. Churkin leg.
distribution: Throughout  the  west  Palaearctic  region, excluding  the  southern  areas, 
north to 66°, Europe (from Iberian Peninsula to Ural Mts): W and S Siberia; Cauca-
sus; Kazakhstan; Transcaucasia; Asia Minor; C Asia; Iran; Afghanistan; Mongolia; 
W China; N India.
Eupoecilia sanguisorbana (Herrich-Schäffer, 1860) 
Material exaMined: 1 male, Kirgizstan, Suusamyr mts., Kokemeren r, 5km N Kyzyl-
Oi, 1800m, 24-25.06.2000, S. Churkin leg. 
distribution: Europe (excluding southern areas) from France to Scandinavia and Ro-
mania; E Europe to Ural Mts; W Kazakhstan; China (Heilongjiang).
Aethes xanthina Falkovitsh, 1963
Material exaMined: 1 female, Kirgizstan, Suusamyr mts., Kokemeren r, 5km N Kyzyl-
Oi, 1800m, 24-25.06.2000, S. Churkin leg.; 1 female, Kirgizstan, E Suusamyr mts., 
Kovjuksu r., 1650m., Kyzyl-Oi, 26.07.00, S. Churkin leg.
distribution: SE Europe (Ural Mts); Kazakhstan; Turkmenia; Tadzhikistan; Uzbeki-
stan; Iran.
Aethes dilucidana (Stephens, 1852) 
Material exaMined: 1 male and 1 female, Kirgizstan, South Chatkal, 5km. E Aatun 
v., Karasu r., 1350m, 19.06.2000, S. Churkin leg.; 1 female, Kirgizstan, S Issyk-Kul 
L., Ottuk v, 1650m, 29.06.2000, S. Churkin leg.
distribution: NW Africa (Algeria); Europe; S Siberia; Kazakhstan; Turkmenia.
Aethes bilbaensis (Rössler, 1877) 
Material exaMined: 1 male, Kirgizstan, South Chatkal, 5km. E Aatun v., Karasu r., 
1350m, 19.06.2000, S. Churkin leg.; 1 female, Kirgizstan, SE Toktogul L, Sargata 
v., 10.07.1999, 1100m, leg. S. Churkin.
distribution: NW Africa; Europe (from Iberian Peninsula to Greece and S Ural Mts); 
Asia Minor, N Lebanon; Iran; Crimea; Caucasus; Transcaucasia; W Kazakhstan; 
Afghanistan; Turkmenia. 
Journal of Entomological and Acarological Research, Ser. II, 42 (1), 2010
4
Cochylidia moguntiana (Rössler, 1864) 
Material exaMined: 3 males and 3 females, Kirgizstan, S. Issyk-kul L., Kadzhi-Sai v., 
1.07.00, 1800m, S. Churkin leg. 
distribution: Europe from Spain to Balkan Peninsula, C Europe to Ural Mts; Siberia, 
Buryatya, Chita; Afghanistan; China; N Pakistan.
Cochylis posterana Zeller, 1847 
Material exaMined: 1 female, Kirgizstan, Toktogul area, Karasù L., 2100m, 16.06.2000, 
S. Churkin leg.; 1 male, Kirgizstan, S Issyk-Kul L., Ottuk v, 1650, 29.06.2000, S. 
Churkin leg. 
distribution: Europe, Ukraine, European Russia east as far as Transkaspia.
Cochylis piana (Kennel, 1919) 
Material exaMined: 1 male, Kirgizstan, Suusamyr mts., Kokemeren r, 5km N Kyzyl-
Oi, 1800m, 24-25.06.2000, S. Churkin leg.
distribution: Kazakhstan;  Tadjikistan; Uzbekistan;  Mongolia; Iran; Afghanistan;  C 
Asia; Tadzhikistan; Djarkent; China (Shaanxi, Sinjiang).
Cochylis amoenana Kennel, 1899 
Material exaMined: 1 male and 2 females, Kirgizstan, Toktogul area, Karasù L., 2100m, 
16.06.2000, S. Churkin leg.
distribution: Asia Minor; N Caucasus; Transcaucasia; Tadzhikistan; Afghanistan; Pa-
kistan; Iran.
Tribe Cnephasiini
Eana samarcandae Razowski, 1958
Material exaMined: 1 male and 1female, Kirgizstan, S. Toktogul L., 15km NE Karakul 
v., 1300-1400m, 17.06.2000, S. Churkin leg.; 4  males  and  8  females, Kirgizstan 
South Chatkal, 5km. E Aatun v., Karasu r., 1350m, 19.06.2000, S. Churkin leg.
distribution: Recorded from Samarkand (Uzbekistan).
Cnephasia cupressivorana (Staudinger, 1871) 
Material exaMined: 1 female, Kirgizstan, S Issyk-Kul L., Ottuk v, 1650, 29.06.2000, 
S. Churkin leg.; 1 female, Kirgizstan, Inn. Tian-Shan, Naryn r. vall, Kazarman vic., 
1400m., 21.07.2000, S. Churkin leg.
distribution: S Europe and Asia Minor.
Cnephasia longana (Haworth, [1811]) 
Material exaMined: 1 female, Kirgizstan, SE Toktogul L, Sargata v., 10.07.1999, 1100m, 
leg. S. Churkin.
distribution: Europe and east to western European Russia.
5
P. Trematerra: Clepsis trifasciata and notes on some Tortricidae from Kirgizstan
Tribe Archipini
Archips betulanus (Hübner, [1787]) 
Material exaMined: 1 female, Kirgizsky, Mts S.slopes, Karakol r., Balyktyr., 2300m, 
13.07.1999, leg. S. Churkin. 
distribution: Palaearctic species, probably excl. the south.
Archips rosanus (Linnaeus, 1758) 
Material exaMined: 2 males, Kirgizstan, Tash-kumyr v., 1000m., 18.06.2000, S. Chur-
kin leg.; 1 female, Kirgizstan, South Chatkal, 5km. E Aatun v., Karasu r., 1350m, 
19.06.2000, S. Churkin leg.
distribution: Palaearctic species, introduced to North America.
Pandemis cinnamomeana (Treitschke, 1830)
Material exaMined: 2 females, Kirgizstan, Toktogul area, Karasù L., 2100m, 16.06.2000, 
S. Churkin leg.
distribution: From W Europe to Ural Mts; Siberia; Mongolia; Russian Far East; NW 
China; Korea; Japan.
Pandemis chondrillana (Herrich-Schäffer, 1860) 
Material exaMined: 1 female, Kirgizstan, S. Toktogul L., 15km NE Karakul v., 1300-
1400m, 17.06.2000, S. Churkin leg.
distribution: Europe, known from France, Romania, Bulgaria; Ukraine and southern 
part of European Russia; Asia Minor; Iran; Afghanistan; Kazakhstan through Siberia, 
Mongolia, C Asia and NW China.
Clepsis moeschleriana (Wocke, 1862)
Material exaMined: 1  male,  Kirgizstan, S Issyk-Kul L., Ottuk v, 1650, 29.06.2000, 
S. Churkin leg. 
distribution: Europe (Polar  Ural), Altai Mts,  in  Nearctics spread from  Labrador  to 
Alaska, other parts of Canada, USA.
Clepsis praeclarana (Kennel, 1899) 
Material exaMined:  1  female,  Kirgizstan,  S.  Issyk-kul  L.,  Kadzhi-Sai  v.,  1.07.00, 
1800m, S. Churkin leg.
distribution: SE Europe; Georgia; SW Siberia, Saisan and Mongolia.
Clepsis neglectana (Herrich-Schäffer, 1851) 
Material exaMined: 5 males and 1 female, Kirgizstan, Suusamyr mts., Kokemeren r, 
5km N Kyzyl-Oi, 1800m, 24-25.06.2000, S.  Churkin  leg.;  1 male, Kirgizstan, S 
Issyk-Kul L., Ottuk v, 1650, 29.06.2000, S. Churkin leg.
distribution: Europe; N Africa; Cyprus; Lebanon; C Asia  reaching  Kyrgyzstan and 
Tuva.
Journal of Entomological and Acarological Research, Ser. II, 42 (1), 2010
6
Clepsis trifasciata sp. n.
Material exaMined: 1 male, holotype, labelled as follows: Kirgizstan, S Issik-Kul L., 
Kadzhi-Sai v., 1.07.2000, 1800 m, S. Churkin leg.; paratype, 1 male with same label.
adult: Wing expanse: male 18-19 mm (Fig. 1). Antennae greenish-brown. Labial palpi 
cream spotted light brown. Frons and vertex brownish-cream. Thorax and tegula 
brownish sprinkled with cream. Ground colour of forewing creamy-white. Mark-
ings are distinct; brownish with a greenish hue; basal blotch sub-triangular, median 
fascia parallel to the last mentioned, terminal fascia conuent with median fascia 
on to tornus. Cilia concolorous with the colour of the markings, being browner at 
the apex and at the tornus. Hindwing whitish, browner on the periphery, cilia con-
colorous with the wing base.
Male genitalia: shown in  gures  2-4.  Uncus  broad ovate;  tegumen broad;  arm  of 
gnathos fairly slender except for terminal portion, termination strong. Distal part of 
valva short, sinuated. Sacculus strongly sclerotized, provided with two ventral promi-
nences: one long situated postbasally, the second smaller and ill-dened terminally. 
Labis with strong dentate part. Aedeagus broad and long armed with dorsal denticles 
accompanied by small proximal spines, terminal part slender curved downwards. 
Cluster of small spines and one very thin cornutus present. 
FeMale genitalia: Unknown.
distribution: Known only from the type locality: Kadzhi-Sai, Issik-Kul province (Kir-
gizstan).
Host: Unknown.
biology: Moths collected on early July at the altitude 1800 m.
reMarks: Fore-wings  present fasciae in C. trifasciata, which in C. violacea are uni-
colorous; ochre with a weak pinkish-violet hue, whereas in C. laetitiae the forewing 
ground colour is ferruginous-yellow with darker shades, these markings are much 
darker being a brownish-rust edged with refractive spots (Razowski, 1979 and 2002; 
Soria, 1997). 
The male genitalia of C. trifasciata is similar to that of C. violacea which is widely 
distributed in the steppes of Mongolia and to C. laetitiae from Central Spain. How-
ever, they differ in the sacculus, in the distal margin of the valvae, and in the aedea-
gus. The sacculus in C. trifasciata is similar in shape to C. violacea and C. laetitiae, 
but the ventral post-basal prominence is stronger. The distal part of the valves are 
sinuate in C. trifasciata and in C. violacea whilst they are convex in C. laetitiae. In 
contrast to C. violacea, the aedeagus is longer in C. trifasciata.
The type specimens of C. trifasciata are deposited in the Trematerra Collection, Uni-
versity of Molise, Campobasso, Italy.
etyMology: The new species is named according to its characteristic wing pattern.
7
P. Trematerra: Clepsis trifasciata and notes on some Tortricidae from Kirgizstan
Fig. 2-4 - Clepsis trifasciata sp. n.: holotype, genitalia (1-4) (2), aedeagus (3-4).
Fig. 1 - Clepsis trifasciata sp. n.: holotype, adult.
1
3
2
4
Journal of Entomological and Acarological Research, Ser. II, 42 (1), 2010
8
SUBFAMILY OLETHREUTINAE
Tribe Bactrini
Bactra bactrana (Kennel, 1901) 
Material exaMined: 1 male, Kirgizstan, Suusamyr mts., Kokemeren r, 5km N Kyzyl-
Oi, 1800m, 24-25.06.2000, S. Churkin leg.
distribution: Canary Is; S Spain; S Italy; Morocco; Algeria; Malta; Egypt; Asia Minor; 
Arabia; Mesopotamia; Iraq; Iran; Caucasus; Afghanistan; Caspian area; Tadzhikistan; 
Uzbekistan; Pakistan; India; Ethiopian regions.
Tribe Olethreutini
Celypha ermolenkoi Kostyuk, 1980 
Material exaMined: 1 female, Kirgizstan, E Suusamyr mts., Kovjuksu r., 1650m, Kyzyl-
Oi, 26.07.00, S. Churkin leg.
distribution: Ukraine, Crimea.
Celipha cespitana (Hübner, [1814-1817])
Material exaMined: 2 males, Kirgizstan, Toktogul area, Karasù L., 2100m, 16.06.2000, 
S. Churkin leg.; 1 male and 3 females, Kirgizstan, South Chatkal, 5km. E Aatun v., 
Karasu r., 1350m, 19.06.2000, S. Churkin leg.; 10 males, Kirgizstan, Suusamyr mts., 
Kokemeren r, 5km N Kyzyl-Oi,  1800m, 24-25.06.2000, S. Churkin leg.; 1 male, 
Inn. Tian-Shan Naryn r. vall, Kazarman vic., 1400m., 21.07.2000, S. Churkin leg.
distribution: W Europe to Ural Mts; Transcaucasia; Asia Minor; Near East; Iran; Rus-
sia (Siberia, C Asia, Cisbaikal, Tuva, Far east: Primorsk); NE China (Manchuria); 
Korea; Japan (Hokkaido, Honshu); Noth America. 
Tribe Eucosmini
Thiodia torridana (Lederer, 1859)
Material exaMined: 2 females, Kirgizstan, Toktogul area, Karasù L., 2100m, 16.06.2000, 
S. Churkin leg.
distribution: Europe; Transcaucasia; Asia Minor; Kazakhstan; Turkmenia; Russia (S 
Siberia, Amur-Zeya area, Priamure, S Primorsk, S Sakhalin, Kuril Is).
Thiodia lerneana (Treitschke, 1835)
Material exaMined: 3 females, Kirgizstan, Suusamyr mts., Kokemeren r, 5km N Kyzyl-
Oi, 1800m, 24-25.06.2000, S. Churkin leg.
distribution: Spain, Italy, Austria to Hungary and Balkan Peninsula, southern parts of 
E Europe; Transural; Kazakhstan; Turkmenia.
9
P. Trematerra: Clepsis trifasciata and notes on some Tortricidae from Kirgizstan
Epinotia thapsiana (Zeller, 1847)
Material exaMined: 1 male, Kirgizstan, Suusamyr mts., Kokemeren r, 5km N Kyzyl-
Oi, 1800m, 24-25.06.2000, S. Churkin leg.
distribution: Canary Is; southern and median belts of Europe to E European Russia; 
Transcaucasia; Asia Minor; Iran; S Siberia; Kazakhstan; Tadzhikistan; Turkmenia; 
Korea.
Pelochrista medullana (Staudinger, 1879)
Material exaMined: 2 males, Kirgizstan, Toktogul area, Karasù L., 2100m, 16.06.2000, 
S. Churkin leg.; 1 male, Kirgizstan, Suusamyr mts., Kokemeren r, 5km N Kyzyl-Oi, 
1800m, 24-25.06.2000, S. Churkin leg.
distribution: Balkan Peninsula, Asia Minor and Iran. 
Eucosma lugubrana (Treitschke, 1830) 
Material exaMined: 1 female, Kirgizstan, E Suusamyr mts., Kovjuksu r., 1650m, Kyzyl-
Oi, 26.07.00, S. Churkin leg.
distribution: Europe from France to Hungary, SE Europe and Greece; Kazakhstan.
Eucosma conterminana (Guenée, 1845) 
Material exaMined:  2  males  and  12  females,  Kirgizstan,  S  Issyk-Kul  L.,  Ottuk  v, 
1650m, 29.06.2000 S. Churkin leg.; 2 females, (Kirgizstan) Fergana valley, 27km 
S. Osh 19.07.2000, 1200m, S. Churkin leg.
distribution: Europe from Portugal to Caucasus, Ural Mts; Transcaucasia; Asia Minor; 
Iran; Kazakhstan; C Asia; S Siberia; Mongolia; Far East; China.
Eucosma metzeneriana (Treitschke, 1830) 
Material exaMined: 1  male,  Kirgizstan, S Issyk-Kul L., Ottuk v, 1650, 29.06.2000, 
S. Churkin leg.; 1 male, Kirgizstan, Inn. Tian-Shan Naryn r. vall, Kazarman vic., 
1400m., 21.07.2000, S. Churkin leg.
distribution: N Africa; W E Europe; Trascaucasus; Asia Minor; Iran; Kazakhstan; C 
Asia; Mongolia; Russia (Siberia, Far East); China; Korea; Japan.
Eucosma ephedrana (Christoph, 1877) 
Material exaMined: 1 male, Kirgizstan, S. Issyk-kul L., Kadzhi-Sai v., 1.07.00, 1800m 
S. Churkin leg.
distribution: Trans-Caucasus; C Asia; Iran; Afghanistan.
Eucosma pupillana (Clerck, 1759) 
Material exaMined: 1 male, Kirgizstan, S. Toktogul L., 15km NE Karakul v., 1300-
1400m, 17.06.2000, S. Churkin leg.; 2 males, Kirgizstan, South Chatkal, 5km. E 
Aatun v., Karasu r., 1350m, 19.06.2000, S. Churkin leg.; 1 female, Kirgizstan, S 
Issyk-Kul L., Ottuk v, 1650, 29.06.2000, S. Churkin leg.
distribution: Europe, Ural Mts; Asia Minor; Iran; Kazakhstan; Tadzhikistan.
Journal of Entomological and Acarological Research, Ser. II, 42 (1), 2010
10
Gypsonoma oppressana (Treitschke, 1835) 
Material exaMined: 1 male, Kirgizstan, E Suusamyr mts., Kovjuksu r., 1650m, Kyzyl-
Oi, 26.07.00, S. Churkin leg.
distribution: Europe; Transcaucasia; Kazakhstan; Tadzhikistan.
Epiblema foenellum (Linnaeus, 1758) 
Material exaMined: 1 female, Kirgizstan, South Chatkal, 5km. E Aatun v., Karasu r., 
1350m, 19.06.2000, S. Churkin leg.
distribution: Europe, S  Russia; Caucasus; Siberia;  Kazakhstan; C Asia;  Mongolia; 
Russian far east; China (Manchuria, Kuansien, Zhejiang); Korea; Japan; India; China; 
Taiwan; Vietman.
ACKNOWLEDGEMENTS
I wish  to express  my sincere  thanks  to  Giuseppe  Spina  (University  of Molise, 
Campobasso, Italy) for technical assistance offered. Also to Gareth King (Universidad 
Autónoma de Madrid, Spain) for checking the English language as used in the manuscript.
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Slamka, Bratislava: 1-195.
soria S., 1997 - Clepsis laetitiae sp.n., una nueva especie del género Clepsis, Guenee, 1845 (Lep.
Tortricidae) en Aranjuez (España). - Bol. San. Veg. Plagas, 23: 63-71.
treMaterra p., 2010  -  Lepidoptera Tortricidae from SE European Russia with description of 
Ceratoxanthis saratovica sp. n. - J. Entomol. Acarol. Res. Ser. II, 42 (1): 19-26.
proF. pasquale treMaterra - Università degli Studi del Molise, Dipartimento di Scienze Animali, 
Vegetali e dell’Ambiente, Via de Sanctis, I-86100 Campobasso (Italy). trema@unimol.it
Accepted 30 March 2010
... Part of the material was trapped during the daytime, and a few species were raised from larvae. Among other species, earlier, from the same material were described Ancylis barabashi and Ancylis ussuricana (Trematerra & Li, 2009), Parapammene komaiana , Statherotmantis olga (Trematerra & Spina, 2009), Clepsis trifasciata (Trematerra, 2010a), and Ceratoxanthis saratovica (Trematerra, 2010b). ...
... The list of species considered is reported below, for the most interesting taxa photographs of wing adult patterns and genital characteristics are provided (Razowski, 2003(Razowski, , 2009Kuznetzov, 1989;Brown, 2005;Alipanah, 2009;Trematerra, 2010a and2010b). Materials are deposited in the Trematerra Collection (University of Molise, Campobasso, Italy). ...
FAUNISTIC NOTES ON FEW TORTRICID MOTHS FROM CENTRAL ASIA AND RUSSIAN FAR EAST (LEPIDOPTERA TORTRICIDAE)
Article
  • Jun 2022
Trematerra P., Li H.H. – Faunistic notes on few tortricid moths from Central Asia and Russian Far East (Lepidoptera Tortricidae). In the paper are reported some species of Lepidoptera Tortricidae collected in Central Asia and Russian Far East territories. Distribution of these species are described; for the most interesting taxa photographs of wing adult patterns and genitalia are provided. Key Words: Lepidoptera Tortricidae, fauna, Central Asia, Russian Far East.
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... Archips 108 X X X X X Brown 2005; Byun et al. 1998;Chapman 1973;Diakonoff 1941c;1951;1952;1976;1982;Duncan 2006;Franclemont 1986;Freeman 1958;Hoebeke et al. 2008;Jinbo 2006;Kruse 2000;Kruse and Sperling 2001;Liu and Li 2002;MacKay 1962;Maier 2003;Razowski 1977;Robinson et al. 1994;Trematerra 2010a;2010b Argyrotaenia 99 X X X Brown and Cramer 1999;Brown 2005;Chapman 1973;Chapman and Lienk 1971;Duncan 2006;Freeman 1944;Heppner 1989;Liu and Li 2002;MacKay 1962;Obraztsov 1961;Powell 1960;1964;Razowski 2001;Razowski and Becker 2000;Trematerra 2010b;Trematerra and Brown 2004 Aristocosma Dang et al. 1996;Diakonoff 1957;1976;Dombroskie and Brown 2009;Duncan 2006;Freeman 1958;Jürivete and Õunap 2008;Kearfott 1907;Liu and Li 2002;MacKay 1962;Obraztsov 1962;1968;Razowski 1979a;1979b;2002a;Trematerra 2010a;Wang et al. 2003 Coeloptera 3 Diakonoff 1941a;1941b;1941c;1941d;1948;1952;1967;1982;Hulcr et al. 2007 Brown 2005;Diakonoff 1939;1941a;1941c;1941d;1948;1952;1953;Liu and Li 2002;Robinson et al. 1994 Labidosa 2 Franclemont 1986;Jinbo 2000;Liu and Li 2002;Obraztsov 1962;Powell 1962b;Razowski 2001;2002a;Razowski and Trematerra 2010; Lozotaeniodes 3 X Bradley et al. 1973;Brown 2005;Razowski 2002a Lumaria 10 X X Brown 2005;Diakonoff 1941;1948;Razowski 2002b;Robinson et al. 1994 Mantua ...
... Archips 108 X X X X X Brown 2005; Byun et al. 1998;Chapman 1973;Diakonoff 1941c;1951;1952;1976;1982;Duncan 2006;Franclemont 1986;Freeman 1958;Hoebeke et al. 2008;Jinbo 2006;Kruse 2000;Kruse and Sperling 2001;Liu and Li 2002;MacKay 1962;Maier 2003;Razowski 1977;Robinson et al. 1994;Trematerra 2010a;2010b Argyrotaenia 99 X X X Brown and Cramer 1999;Brown 2005;Chapman 1973;Chapman and Lienk 1971;Duncan 2006;Freeman 1944;Heppner 1989;Liu and Li 2002;MacKay 1962;Obraztsov 1961;Powell 1960;1964;Razowski 2001;Razowski and Becker 2000;Trematerra 2010b;Trematerra and Brown 2004 Aristocosma Dang et al. 1996;Diakonoff 1957;1976;Dombroskie and Brown 2009;Duncan 2006;Freeman 1958;Jürivete and Õunap 2008;Kearfott 1907;Liu and Li 2002;MacKay 1962;Obraztsov 1962;1968;Razowski 1979a;1979b;2002a;Trematerra 2010a;Wang et al. 2003 Coeloptera 3 Diakonoff 1941a;1941b;1941c;1941d;1948;1952;1967;1982;Hulcr et al. 2007 Brown 2005;Diakonoff 1939;1941a;1941c;1941d;1948;1952;1953;Liu and Li 2002;Robinson et al. 1994 Labidosa 2 Franclemont 1986;Jinbo 2000;Liu and Li 2002;Obraztsov 1962;Powell 1962b;Razowski 2001;2002a;Razowski and Trematerra 2010; Lozotaeniodes 3 X Bradley et al. 1973;Brown 2005;Razowski 2002a Lumaria 10 X X Brown 2005;Diakonoff 1941;1948;Razowski 2002b;Robinson et al. 1994 Mantua ...
Phylogeny of the tribe Archipini (Lepidoptera: Tortricidae: Tortricinae) and evolutionary correlates of novel secondary sexual structures
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... The present paper provides new data on the distribution of some Tortricidae (total number of species 69). Among other tortricids of particular interest were: Phtheochroa kenneli (Obraztsov), Cochylimorpha halophilana (Christoph), C. alternana (Stephens), C. emiliana (Kennel), C. pirizanica (Razowski), and C. woliniana (Schleich), Phalonidia contractana (Zeller), Agapeta zoegana (Linnaeus), Eugnosta lathoniana (Hübner), Cochylidia rupicola (Curtis) and C. moguntiana (Rössler), Eana samarcandae (Razowski), Cnephasia cupressivorana (Staudinger) and C. longana (Haworth), Clepsis moeschleriana (Wocke) and C. praeclarana (Kennel), Endothenia hebesana (Walker), Celypha ermolenkoi Kostyuk, Pelochrista griseolana (Zeller) and P. medullana (Staudinger), Pammene fasciana (Linnaeus) (Razowski, 1979(Razowski, , 2002(Razowski, , 2003(Razowski, and 2009Kuznetsov, 1989;Brown, 2005;Alipanah, 2009;Trematerra, 2010a and2010b (Kuznetsov, 1975) ...
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Faunistic data of some Lepidoptera Tortricidae collected in moun-tainous localities of Kazakhstan, Kyrgyzstan and Turkmenija, are reported. The total number of species recorded is 69; some of them are of special biogeographical interest.
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Lepidoptera of Pakistan based upon the Info-system of the Cesa II Preliminary annotated checklist of the species recorded in Pakistan.
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For the first time, 1476 species of the Lepidoptera of Pakistan are listed all together here. Among them, 334 butterfly species of 9 families are known. These are as follows; Papilionidae, Pieridae, Argynnidae, Libytheidae, Satyridae, Danaidae, Riodinidae, Lycaenidae, and Hesperiidae. Totally 1142 moth species are recorded in Pakistan. Their families are as follows; Adelidae, Aganaidae, Arctiidae, Autostichidae, Bombycidae, Brachodidae, Brahmaeidae, Carposinidae, Coleophoridae, Cosmopterigidae, Cossidae, Ctenuchidae, Drepanidae, Endromidae, Epermeniidae, Eriocottidae, Ethmiidae, Eupterotidae, Gelechiidae, Geometridae, Gracillariidae, Lasiocampidae, Lecithoceridae, Limacodidae, Lymantriidae, Momphidae, Noctuidae, Notodontidae, Psychidae, Pterophoridae, Pyralidae, Saturniidae, Scythridae, Sesiidae, Sphingidae, Stathmopodidae, Stenomidae, Thaumetopoeidae, Thyatiridae, Tineidae, Tortricidae, Yponomeutidae, Zygaenidae. In this study, 862 publications on the Lepidoptera of Pakistan are evaluated from the taxonomical, faunistical, ecological standpoints, and listed as author index. Besides, the material of the Lepidoptera preserved in the Cesa collection are partly examined. The images of the adults of the collection, as well as the images of the male genitalia in the archive are added to some species.
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Lepidoptera Tortricidae from SE European Russia with description of Ceratoxanthis saratovica sp. n.
Article
Full-text available
  • Apr 2010
Faunistic data of some Lepidoptera Tortricidae collected in Southern Russian territory are reported; moreover, the new species Ceratoxanthis saratovica sp. n., from Saratov region is described. Externally C. saratovica resembles to Ceratoxanthis argentomixtana (Staudinger, 1870); in the male genitalia is closed to Ceratoxanthis iberica Baixeras, 1992.
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1989 -Family Tortricidae (Olethreutidae, Cochylidae) - Tortricid Moths Keys to the insects of the European Part of the USSR, Volume IV, Lepidoptera , Part I
  • 279-991
  • I Kuznetsov V
kuznetsov v.i., 1989 -Family Tortricidae (Olethreutidae, Cochylidae) - Tortricid Moths. In: Medvedev G.S., Keys to the insects of the European Part of the USSR, Volume IV, Lepidoptera, Part I. - E.J. Brill, Leiden: 279-991.
Vegetali e dell'Ambiente, Via de Sanctis, I-86100 Campobasso (Italy). trema@unimol.it Accepted
  • Apr 2010
  • Dipartimento Pasquale Trematerra-Università Degli Studi Del Molise
  • Di Scienze
  • Animali
proF. pasquale treMaterra - Università degli Studi del Molise, Dipartimento di Scienze Animali, Vegetali e dell'Ambiente, Via de Sanctis, I-86100 Campobasso (Italy). trema@unimol.it Accepted 30 March 2010
In: World Catalogue of Insects 5
  • Jan 2005
  • 1-741
  • J W Brown
brown J.w., 2005 - Tortricidae (Lepidoptera). In: World Catalogue of Insects 5. - Apollo Book Aps, Stenstrup, Denmark: 1-741.
Revision of the Genus Clepsis Guenée (Lepidoptera, Tortricidae). Part I
  • Jan 1979
  • 101-198
razowski J., 1979 - Revision of the Genus Clepsis Guenée (Lepidoptera, Tortricidae). Part I. -Acta Zool. Cracov., XXIII (8): 101-198.
Clepsis laetitiae sp.n., una nueva especie del género Clepsis
  • Jan 1997
  • 63-71
  • S Soria
soria S., 1997 - Clepsis laetitiae sp.n., una nueva especie del género Clepsis, Guenee, 1845 (Lep. Tortricidae) en Aranjuez (España). - Bol. San. Veg. Plagas, 23: 63-71.