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A late-surviving basal theropod dinosaur from the latest Triassic of North America

The Royal Society
Proceedings of the Royal Society B
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The oldest theropod dinosaurs are known from the Carnian of Argentina and Brazil. However, the evolutionary diversification of this group after its initial radiation but prior to the Triassic-Jurassic boundary is still poorly understood because of a sparse fossil record near that boundary. Here, we report on a new basal theropod, Daemonosaurus chauliodus gen. et sp. nov., from the latest Triassic 'siltstone member' of the Chinle Formation of the Coelophysis Quarry at Ghost Ranch, New Mexico. Based on a comprehensive phylogenetic analysis, Daemonosaurus is more closely related to coeval neotheropods (e.g. Coelophysis bauri) than to Herrerasauridae and Eoraptor. The skeletal structure of Daemonosaurus and the recently discovered Tawa bridge a morphological gap between Eoraptor and Herrerasauridae on one hand and neotheropods on the other, providing additional support for the theropod affinities of both Eoraptor and Herrerasauridae and demonstrating that lineages from the initial radiation of Dinosauria persisted until the end of the Triassic. Various features of the skull of Daemonosaurus, including the procumbent dentary and premaxillary teeth and greatly enlarged premaxillary and anterior maxillary teeth, clearly set this taxon apart from coeval neotheropods and demonstrate unexpected disparity in cranial shape among theropod dinosaurs just prior to the end of the Triassic.
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... The most significant event that determined the appearance of the new higher group of reptiles and the rise in the level of organization of its representatives was the appearance of first archosaurs in the large size class as active predators at the end of the Permian. They occupied the levels of higher-order consumers, i.e., the top of the food pyramid of the dominant terrestrial community (Sennikov, 1995). This was accompanied by significant morphological transformations in the first archosaurs with respect to their ancestors, small early diapsids, such as Eosuchia. ...
... It is very likely that the early small theropod Daemonosaurus (Sues et al., 2011;Nesbitt and Sues, 2021) with a very short skull and jaws and with a small number of strongly anisodont, very large teeth (Fig. 14) was adapted to macrophagy, like ornithosuchids. One should note a sharp difference in the structure of its jaw apparatus from that of another Triassic theropod, Coelophysis, which indicates different strategies for capturing and killing prey. ...
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... The most significant event that determined the appearance of the new higher group of reptiles and the rise in the level of organization of its representatives was the appearance of first archosaurs in the large size class as active predators at the end of the Permian. They occupied the levels of higher-order consumers, i.e., the top of the food pyramid of the dominant terrestrial community (Sennikov, 1995). This was accompanied by significant morphological transformations in the first archosaurs with respect to their ancestors, small early diapsids, such as Eosuchia. ...
... It is very likely that the early small theropod Daemonosaurus (Sues et al., 2011;Nesbitt and Sues, 2021) with a very short skull and jaws and with a small number of strongly anisodont, very large teeth (Fig. 14) was adapted to macrophagy, like ornithosuchids. One should note a sharp difference in the structure of its jaw apparatus from that of another Triassic theropod, Coelophysis, which indicates different strategies for capturing and killing prey. ...
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... Langer and Benton 2006;Upchurch et al. 2007; Yates 2017a, b;Nesbitt and Chatterjee 2008;Martínez and Alcober 2009;Alcober and Martínez 2010). Yet, other analyses continued to recover the more traditional theropodan position of E. lunensis (e.g.Ezcurra 2006Ezcurra , 2010Ezcurra and Novas 2007;Nesbitt et al. 2009;Langer et al. 2011;Novas et al. 2011;Sues et al. 2011). More recently, a third alternative was proposed, this time placing E. lunensis as one of the earliest branching sauropodomorphs.This hypothesis was supported by multiple subsequent studies (e.g. ...
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lots o measurements!