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1. INTRODUCTION
In the Aydıncık area an autochtonous sequence of plat-
form carbonate rocks was deposited, ranging in age
from the Liassic to the Aptian. The sequence conform-
ably overlies Upper Triassic basal conglomerates, inter-
bedded with mudstones and sandstones (DEMI
.
R T AS¸-
LI, 1984), and is unconformably overlain by Upper
Campanian fore-reef carbonate breccia and limestone
(TASLI & EREN, 1999).
The biostratigraphy of Jurassic-Cretaceous carbon-
ate sequences of the Central Taurides has been studied
by the author since 1997. The present work on the Ay-
dıncık profile forms part of the ongoing studies, such as
TASLI (2000). The Late Jurassic section is particularly
rich in benthic foraminifera belonging to the families
Pfenderinidae (Kurnubiinae) and Valvulinidae. This
Benthic Foraminifera of the Upper Jurassic Platform Carbonate
Sequence in the Aydıncık (I
.
çel) Area, Central Taurides, S Turkey
Kemal TASLI
association corresponds to that of coeval facies of the
Mediterranean realm (HENSON, 1948b; SARTONI &
CRESCENTI, 1962; HOTTINGER, 1967; GU©I∆,
1969; NIKLER & SOKA», 1968; VELI∆, 1977;
SEPTFONTAINE, 1980) and is linked with shallow-
water, protected lagoon carbonates. Jurassic benthic
foraminifera from the Taurus Mountains were reported
from north of Isparta (GUTNIC & MOULLADE,
1967), north and northwest of Antalya (BASSOULLET
& POISSON, 1975), east of Kayseri (ALTINER &
SEPTFONTAINE, 1979) and Munzur Dag˘ (BASSOU-
LLET & BERGOUGNAN, 1981).
The aim of this paper is to describe some important
species of benthic foraminifera. For the present study,
approximately eighty random thin-sections from twen-
ty-five fossiliferous samples within the measured strati-
graphic section were studied. Micropalaeontological
determinations are based on the study of these random
thin-sections and a large number of successive acetate
peels and polished samples.
2. LOCATION AND FIELD DESCRIPTION OF
THE AYDINCIK PROFILE
The Aydıncık Upper Jurassic profile is located about 2
km along the road to Karaseki village west of Aydıncık
(I
.
çel) town (Fig. 1). It continuously overlies an alterna-
tion of dolomite and limestone of late Dogger age
(TASLI, 2000) and is about 50 m thick. Its upper limit
is marked by the reappearance of entirely dolomitic
beds representing the transition between the Jurassic
and the Cretaceous, which is easily recognizable in the
field because of its morphological difference.
The analyzed section consists of light brown, pre-
dominantly thick-bedded limestones. C l a d o c o r o p s i s
m i r a b i l i s FELIX, although occuring sporadically in the
underlying beds (i.e. in the Upper Dogger), is so fre-
quent and abundant that it seems to be lithologically
dependent.
3. BIOSTRATIGRAPHY
Stratigraphic distribution of the benthic foraminifera
and calcareous algae is shown in Fig. 2. The analyzed
Geologia Croatica 54/1 1 - 13 2 Figs. 1 Tab. 3 Pls. ZAGREB 2001
Key words: Benthic foraminifera, Carbonate plat-
form, Upper Jurassic, Central Taurides, Turkey.
Mersin Üniversitesi Jeoloji Mühendislig
ˇ
i Bölümü, TR-33342 I
.
ç e l ,
Türkiye.
e-mail: ktasli@mersin.edu.tr
Abstract
The Upper Jurassic sequence of the Aydıncık (I
.
çel) area consists of
platform limestones which were deposited in a subtidal, restricted
lagoon environment. Stratigraphic distribution of benthic foraminifera
and calcareous algae, examined in thin-sections, is shown in a range-
chart. The microfossil assemblage indicates the Salpingoporella sellii
subzone of the Kurnubia palastiniensis cenozone, corresponding
approximately to the lower part of the Malm. Some benthic
foraminifera with considerable stratigraphic value within the Meso-
zoic Tethys are described. Among the benthic foraminifera, taxa of
the family Pfenderinidae, especially the subfamily Kurnubiinae, are
dominant and frequent throughout the sequence. The planispirally
coiled taxa are represented by the families Nautiloculinidae, Charenti-
idae and Cyclamminidae (subfamily Bucciccrenatinae).
2 Geologia Croatica 54/1
limestones continuously overlie limestones and dolomi-
tes corresponding the underlying biostratigraphic unit,
Paleopfenderina salernitana cenozone of SARTONI &
CRESCENTI (1962) (TASLI, 2000). Kurnubia palas -
t i n i e n s i s HENSON, an index species of the cenozone
established by SARTONI & CRESCENTI (1962) in the
Apennines, is frequent and locally abundant throughout
the sequence. This species is known to occur through
the whole Malm of the Mediterranean realm (e.g.
VELI∆, 1977; SEPTFONTAINE, 1980). The presence
of Salpingoporella sellii (CRESCENTI) (Pl. III, Fig.
10), a dasycladacean alga used as an index species in
the Apennines (SARTONI & CRESCENTI (1962) and
Dinarides (NIKLER & SOKA», 1968; GU©I∆ et al.,
1971; VELI∆, 1977), indicates the S. sellii s u b z o n e
which corresponds approximately to the lower part of
the Malm.
4. MICROFACIES DEVELOPMENT AND
PALAEOENVIRONMENTAL SETTING
The limestones consist mainly of fossiliferous wacke-
stones and biopeloidal grainstones, with oncoids. Intra-
clastic grainstones with F a v r e i n a sp. only occur in the
uppermost part of the sequence. The presence of abun-
dant, well preserved Cladocoropsis mirabilis F E L I X ,
centimetre-sized oncoids and benthic foraminifera are
characteristic for these limestones. Benthic foraminifera
are most abundant regarding both the number of species
and individuals, whereas calcareous algae are subordi-
nate. Dasycladacean algae are represented by Salpingo -
porella annulata CAROZZI (Pl. III, Fig. 11), which is
locally common, and rarely by Salpingoporella sellii
(CRESCENTI). Thaumatoporella parvovesiculifera
(RAINERI) (Pl. III, Fig. 12) is frequent, but not abun-
Fig. 1 The geographic subdivisions of the Taurides (after
ÖZGÜL, 1984) and location of the Upper Jurassic section.
dant. Bioclastic fraction is mainly composed of frag-
ments of C l a d o c o r o p s i s, pelecypods and subordinately
brachiopods (punctate), ostracods and echinoids. Cal-
careous sponge spicules are locally abundant. Siliciclas-
tic material is totally absent.
Predominantly dense micrites and abundant oncoids
indicate low energy in a sheltered environment and
shallow-water conditions. Intraclastic grainstones with
Favreina sp. in the uppermost part of the sequence indi-
cate high energy conditions, resulting in the develop-
ment of a regressive facies.
5. SYSTEMATIC DESCRIPTIONS
Family Nautiloculinidae LOEBLICH & TAPPAN,
1985
Genus Nautiloculina MOHLER, 1938;
emend. BRÖNNIMANN, 1967
Type-species: Nautiloculina oolithica MOHLER,
1938
Nautiloculina circularis (SAID & BARAKAT), 1959
(Pl. I, Fig. 1)
1966 Nautiloculina circularis (SAID & BARAKAT).-
DERIN & REISS, Photo Nos. 70, 71, 83, 254,
263, 264, 271, 280, 283, 286-289, 309.
1968 Nautiloculina circularis (SAID & BARAKAT).-
BRÖNNIMANN, p. 64, fig. 3, pl. 1, figs. 1-8, pl.
2, figs. 1-6.
1977 Nautiloculina oolithica MOHLER.- VELI∆, pl.
VIII, figs. 7, 8.
1985 Nautiloculina circularis (SAID & BARAKAT).-
FOURCADE, ARAFA & SIGAL, pl. 3, fig. 4.
M a t e r i a l : Twelve random thin-sections with appro-
ximately eighty specimens.
Description and remarks : This form with lenticu-
lar-shaped, involute-planispirally coiled test shows the
umbilical fillings and axial depressions. BRÖNNI-
MANN (1968) distinguishes N. circularis from N .
o o l i t h i c a MOHLER by the marked subacute periphery,
axial depressions, and by the larger size and larger
numbers of the whorls in the former. ARNAUD-VAN-
NEAU & PEYBERNÈS (1978) give a comparison table
of the principal characters of the four species of N a u -
t i l o c u l i n a (including the two Cretaceous species). In
our material there are specimens which are assignable
to the species N. circularis and N. oolithica (Pl. 1, Fig.
2), with the dominance of the former.
3Tasli: Benthic Foraminifera of the Upper Jurassic Platform Carbonate Sequence...
Fig. 2 Stratigraphic column of the Aydıncık Upper Jurassic section and distribution of benthic foraminifera and other microorganisms.
4 Geologia Croatica 54/1
D i m e n s i o n s : The measurements are from axial sec-
tions only (12 specimens). Equatorial and axial diame-
ters vary between 0.55-0.80 mm, and 0.30-0.48 mm,
respectively. The ratio of equatorial/axial diameter osci-
llates around 1.8:1. The inner diameter of proloculus
ranges from 0.03 mm to 0.05 mm.
Family Charentiidae LOEBLICH & TAPPAN, 1985
Genus Karaisella KURBATOV, 1971
Type-species: Karaisella uzbekistanica KURBATOV,
1971
Karaisella aff. uzbekistanica KURBATOV, 1971
(Pl. I, Figs. 3-13)
?1958 H a p l o p h r a g m i u m aff. s u p r a j u r a s s i c u m S C H -
WAGER.- DUFAURE, pl. 1, figs. 21-22.
?1968 Haplophragmium cf. suprajurassicum SCHWA-
GER.- NIKLER & SOKA», pl. IX, fig. 7.
Material : Fifteen random thin-sections and ten suc-
cesive acetate peels with approximately ninety speci-
mens.
D e s c r i p t i o n : Test is free, lenticular shaped, thick-
ened towards the poles, with small axial depressions.
Peripheral margin is rounded to subrounded. Septal
sutures, observed in equatorial sections (Pl. I, Figs. 6,
7), are slightly depressed. Coiling is planispiral, occa-
sionally streptospiral, and involute, with a tendency to
become uncoiled in the later stage (Pl. I, Fig. 12). There
are two or at maximum three and a half whorls. The
number of chambers in the last whorl is nine to twelve.
Chamber interiors are simple. Septa are curved and
inclined in the direction of coiling, in continuity with
the outer wall. The base of the septum against the previ-
ous whorl is thickened and chomata-like (Pl. I, Figs. 8-
10). Septal spacing (= height of chamber) and height of
the whorls slightly increase during the ontogeny. Mega-
losphere is simple, spherical or slightly ovoid. Aperture
is simple and central (Pl. I, Figs. 6, 7). Wall is calcare-
ous microgranular (= finely agglutinated) with a kerio-
thecal structure, occasionally visible only in the last
whorl of large specimens (Pl. 1, Fig. 11).
D i m e n s i o n s : Dimensions (in mm) are shown in
Table 1.
R e m a r k s : Because of its streptospiral coiling, sin-
gle areal aperture, and the presence of chomata-like
thickening at the base of the septa (Pl. I, Fig. 5), this
form is herein assigned to the genus K a r a i s e l l a K U R-
BATOV, 1971 (in LOEBLICH & TAPPAN, 1988).
K a r a i s e l l a aff. u z b e k i s t a n i c a is a close homeomorph of
the Cretaceous genus C h a r e n t i a NEUMANN, 1965,
except its streptospiral coiling. The genus K a r a i s e l l a
was placed in the family Charentidae by LOEBLICH &
TAPPAN (1988) although they note that it did not show
a canaliculate wall structure (keriothecal structure of
HOTTINGER, 1967). Specimens in the Aydıncık mate-
rial do not show a sharp change in the plane of coiling
from early to later whorls and subacute periphery as in
the type species.
This form is similar to the genera B u l b o b a c u l i t e s
MAYNC, 1952 and H a p l o p h r a g m i u m REUSS, 1860 in
areal aperture and the streptospiral nature of coiling.
However, the wall is distinctly agglutinated and simple
in the former genus, and alveolar in the latter. More-
Sample Greater Inner diameter
No. (AY) Specimens equatorial Axial diameter of proloculus
diameter
38-1 1 (Pl.1, Fig. 4) 0.85 0.45 0.05-0.06
38-1 2 0.63 0.30 not observed
38-1 3 0.55 - 0.04
38-3 4 0.70 - 0.06
38-3 5 0.65 - not observed
38-3 6 (Pl.1, Fig. 10) 0.80 - not observed
38-3 7 0.85 - 0.12
38-4 8 0.72 - not observed
38-5 9 0.70 - 0.05
38-6 10 (Pl.1, Fig. 6) 0.48 0.30 0.06
38-6 11 (Pl.1, Fig. 11) 0.90 - not observed
38-6 12 (Pl.1, Fig. 9) 0.65 - 0.06
38-6 13 0.66 - 0.11
38-8 14 (Pl.1, Fig.6) 0.70 - 0.10
38-8 15 0.75 0.40 0.05
38-10 16 0.47 - 0.06
38-10 17 0.70 - 0.13-0.15
38-13 18 (Pl. I, Fig.7) 0.46 - 0.06
38-14 19 0.50 - 0.07
Table 1 Dimensions of Karaisella a f f .
uzbekistanica (in mm).
5Tasli: Benthic Foraminifera of the Upper Jurassic Platform Carbonate Sequence...
over, both genera have a marked rectilinear uniserial
stage which is only occasionally developed in our spec-
imens. Karaisella aff. uzbekistanica differs from anoth-
er Upper Jurassic species, Mesoendothyra izjumiana
DAIN, 1958 (with its wall consisting of large pores,
DAIN in BYKOVA et al., 1958), by the central posi-
tion of the aperture, the presence of chomata-like thick-
enings in the base of the septum and by a lacking of
distinct streptospiral coiling.
K a r a i s e l l a aff. uzbekistanica closely resembles the
genus B o s n i e l l a GU©I∆, 1977 by its keriothecal wall
structure and growth pattern, but differs from it in the
absence of a peneropline stage with cribrate aperture
and in having only a central aperture. In the latter, the
aperture is at first basal, then central and finally cribrate
as in “Mesoendothyra” croatica GU©I∆ (GU©I∆, 1969;
FURRER & SEPTFONTAINE, 1977). The genus B o s -
n i e l l a is represented by two species: B. oenensis f r o m
the Lower Jurassic of NW Bosnia (GU©I∆, 1977) and
B. fontainei from the Middle Jurassic of Thaïland
(BASSOULLET, 1994). It is considered as a junior
synonym of M e s o e n d o t h y r a DAIN (SEPTFONTAINE,
1988). Alternatively, BASSOULLET (1994) removed
“M e s o e n d o t h y r a ” c r o a t i c a GU©I∆ from the genus
M e s o e n d o t h y r a DAIN, because of its keriothecal wall
structure, and assigned it to the genus B o s n i e l l a G U -
©I∆, 1977.
O c c u r e n c e : It is abundant in biopeloidal grainsto-
nes with oncoids, in association with Nautiloculina cir -
c u l a r i s and Kurnubia ex. gr. p a l a s t i n i e n s i s , whereas it
is missing or rare in fossiliferous wackestones.
Family Pfenderinidae SMOUT & SUGDEN, 1962
Subfamily Kurnubiinae REDMOND, 1964
Genus Kurnubia HENSON, 1948
Type species: Kurnubia palastiniensis HENSON,
1948
After SMOUT & SUGDEN (1962), who assigned
HENSON’s (1948b) species Valvulinella jurassica a n d
V. wellingsi to the genus K u r n u b i a HENSON and after
SARTONI & CRESCENTI (1962), who considered the
species K. jurassica synonymous with Kurnubia palas -
t i n i e n s i s, REDMOND (1964) described, from isolated
specimens only, three new species of K u r n u b i a: K .
v a r i a b i l i s , K. bramkampi and K. m o r r i s i. MAYNC
(1965) included all these species in Kurnubia gr. palas -
t i n i e n s i s HENSON, except K. morrisi, considering the
existence of intermediate forms. GU©I∆ (1969) adopted
MAYNC’s (1965) opinion and considered the three
infrasubspecific taxa as “forms” j u r a s s i c a , p a l a s t i n i e n -
s i s and w e l l i n g s i. HOTTINGER (1967) redescribed K .
p a l a s t i n i e n s i s in detail, including K. jurassica , and
retained the three REDMOND’s (1964) species of Kur -
nubia. Later, this common Late Jurassic genus has been
recorded mostly under the name Kurnubia palastinien -
s i s HENSON in many studies (e.g. BASSOULLET &
POISSON, 1975; AZÉMA et al., 1977; VELI∆, 1977;
FOURCADE et al., 1985; LUPERTO-SINNI & MAS-
SE, 1994). In general, REDMOND’s (1964) species
seem not to be accepted, probably owing to difficulties
in comparing with isolated specimens.
The aim of the following descriptions is to con-
tribute more data to the existing knowledge on the sub-
family Kurnubiinae, avoiding the creation of new taxa.
Kurnubia ex. gr. palastiniensis HENSON, 1948b
(Pl. II, Figs. 1-7)
D e s c r i p t i o n : See description in HOTTINGER
(1967).
R e m a r k s : Our specimens display wide morpholog-
ic variations and considerable differences in size. They
vary from those smaller in size, only trochospirally,
having a weakly developed central column and possess-
ing a hypodermic network (SEPTFONTAINE, 1988)
with first order partitions (Pl. II, Figs. 1, 2, 5), to those
having a larger test with a marked central column, a
more or less developed uniserial stage, and possessing a
complete hypodermic network (Pl. II, Figs. 4, 6). The
former have a simple, ovoid proloculus, measuring
about 0.04 mm (inner diameter) and representing the
megalospheric generation. They are included in this
group due to the presence of the transitional forms. In
the latter forms, the proloculus is not visible. The cen-
tral column seems to be weakly developed in highly
conical specimens (Pl. II, Fig. 6), whereas it is well
developed in specimens having a relatively larger basal
diameter (Pl. II, Figs. 3, 4).
Kurnubia cf. morrisi REDMOND, 1964
(Pl. II, Figs. 8-12, 14)
?1964 Kurnubia morrisi new species.- REDMOND, p.
253, pl. 1, fig. 4.
1967 Kurnubia cf. m o r r i s i REDMOND.- HOTTIN-
GER, p. 93, pl. 19, figs. 35-37.
D e s c r i p t i o n : Test is fusiform, trochospirally coiled
throughout the ontogeny. Early chambers are not visi-
ble. Spiral sutures are depressed, at about 30°to the axis
of coiling. Septal sutures are obscure. The wall is cal-
careous, microgranular without agglutinated grains,
possessing a complete hypodermic network. The prima-
ry aperture is set in the inner margin of the peripheral
zone where the septa do not meet the central column
(Pl. 2, Fig. 11). It probably represents “intercameral
foramina” (SMOUT & SUGDEN, 1962). Prolongations
of the adjacent first order vertical partitions projecting
inward from the epidermis adjoin to each other and coa-
lesce with the interseptal pillars (Pl. II, Fig. 10). The
second order vertical partitions are restricted only to the
marginal zone of the chambers. The central zone has a
trochoidally laminated appearance (Pl. 2, Fig. 14)
6 Geologia Croatica 54/1
which recalls the apertural plates intergrown with pil-
lars in the Pfenderinidae. The base of the test is strongly
convex in the centre and very obliquely set to the axis
of coiling.
D i m e n s i o n s : Axial length varies from 2.1-2.5 mm,
measured in nearly axial sections. Basal diameter is ≥
0.8 mm and exceeds up to 1.0 mm. The ratio of
length/diameter oscillates around 2.5:1. The width of
the peripheral zone surrounding the central zone is
nearly constant throughout the adult stage, measuring
0.20-0.22 mm. The central column increases progres-
sively in diameter, up to 0.5 mm.
Remarks : K. cf. morrisi has a larger test and central
column, and a wider peripheral zone than all other
described species of Kurnubia and a complete hypoder-
mic network consisting of two generations of partitions
in the adult stage. Furthermore, specimens of Kurnubia
ex. gr. p a l a s t i n i e n s i s do not exceed 0.7 mm in basal
diameter. Purely because of the trochospiral coiling,
this form is not considered as K. w e l l i n g s i ( H E N S O N ) .
Six to eight tiers of chamberlets per chamber, men-
tioned by REDMOND (1964, p. 253), are not account-
able in random thin sections.
Genus Conicokurnubia SEPTFONTAINE, 1988
Type species Conicokurnubia orbitoliniformis
SEPTFONTAINE, 1988
Conicokurnubia orbitoliniformis SEPTFONTAINE,
1988
(Pl. II, Figs. 13, 15, 16)
Description : Test is sharply conical (Pl. II, fig. 13),
where chambers do not increase in diameter as added,
and broadly conical where chambers increase slowly in
diameter. The base is slightly to strongly convex in the
centre, with a narrow imperforate rim. The cone side is
straight. Proloculus is not visible. The trochospiral
arrangement of the early chambers is suggested by
traces of the spiral suture. Later and remaining larger
portion of the test consists of a co-axial series of ten to
seventeen very low chambers which increase slightly in
height as added. Septal sutures are distinct and depres-
sed. Each chamber has a peripheral zone with a com-
plete hypodermic network. Each septum is inwardly
thickened and then adjoins with the adjacent septum,
forming buttress-like interseptal pillars (Pl. II, Fig. 15).
The first order vertical partitions form a “reticulate
zone” (HENSON, 1948a) in the centre of the test as
seen in transverse sections (Pl. II, Fig. 16). Apertural
pores are not observable. The primary aperture consists
of an opening near the margin of the central zone (Pl.
II, Fig. 15).
D i m e n s i o n s : The broadly conical specimens have a
basal diameter of 1.25-1.50 mm and a height of 1.75-
1.85 mm, measured in nearly axial sections. The
sharply conical specimens have a basal diameter of
0.50-0.70 mm and a height of 1.25-2.0 mm, measured
in nearly axial sections. The height of the last chamber,
for both forms, is ≤ 0.1 mm. The width of the peripher-
al zone surrounding the central column is 0.10-0.12
mm.
Remarks : Specimens of this species from Aydıncık
are closely comparable with SEPTFONTAINE’s (1988)
figures (pl. II, figs. 12, 13) from the Oxfordian (?) to
Kimmeridgian of Western Taurus, Turkey. However,
the available axial and transverse sections are insuffi-
cient for a complete description of the species.
Conicokurnubia orbitoliniformis occurs throughout
the Aydıncık Upper Jurassic section, in association with
K u r n u b i a ex gr. p a l a s t i n i e n s i s. In the random thin-sec-
tions, specimens with a marked uniserial stage of the
latter might be confused with sharply conical specimens
of C. orbitoliniformis. The width of the peripheral zone
seems to be narrower than in Kurnubia palastiniensis
and Kurnubia aff. morrisi.
6. BIOSTRATIGRAPHIC REVIEW
AND CONCLUSIONS
The characteristics of the Upper Jurassic limestone sec-
tion from the Aydıncık (I
.
çel) area are the presence of
C l a d o c o r o p s i s , benthic foraminifera, calcareous algae
and the dominance of mudstones which indicate a sub-
tidal, protected lagoon environment.
Stratigraphic distribution of the benthic foraminifera
and calcareous algae is shown in a range-chart. Micro-
fossil assemblage corresponds to the S a l p i n g o p o r e l l a
s e l l i i subzone of Kurnubia palastiniensis c e n o z o n e ,
established by SARTONI & CRESCENTI (1962).
Ten species of benthic foraminifera are identified
and figured. Those of considerable stratigraphic value
within the Mesozoic Tethys are described.
7. REFERENCES
ALTINER, D. & SEPTFONTAINE, M. (1979): Micropaléon-
tologie, stratigraphie et environnement de déposition
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de Pınarbas¸ı (Taurus oriental, Turquie).- Rev. Micropalé-
ont., 22/1, 3-18.
ARNAUD-VANNEAU, A. & PEYBERNÈS, B. (1978): Les
représentants Éocrétacés du genre N a u t i l o c u l i n a M O H-
LER, 1938 (Foraminifera, Fam. Lituolidae?) dans les cha-
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Franco-Espagnoles. Revision de Nautiloculina cretacea
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AZÉMA J., CHABRIER, G., FOURCADE, E. & JAFFRE-
ZO, M. (1977): Nouvelles données micropaléontologiqu-
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7Tasli: Benthic Foraminifera of the Upper Jurassic Platform Carbonate Sequence...
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BASSOULLET, J.-P. & BERGOUGNAN, H. (1981): Faune
et facies typiques du domaine sud-tethysien: le Lias du
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BRÖNNIMANN, P. (1968): Re-examination of the morphol-
ogy of Nautiloculina circularis (SAID & BARAKAT),
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BYKOVA, N.K., BALAKHMATOVA, Y.T., VASSILEN-
KO, V.P., VOLOSHINOVA, N.A., GRIGELIS, A., DA-
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Manuscript received August 24, 2000.
Revised manuscript accepted April 23, 2001.
8 Geologia Croatica 54/1
PLATE I
scale bar: 0.2 mm
Fig. 1 Nautiloculina circularis (SAID & BARAKAT)
Axial section showing the acute periphery through the ontogeny, sample AY 38-9.
Fig. 2 Nautiloculina oolithica MOHLER
Axial section showing the rounded periphery, sample AY 38-9.
Figs. 3-13 Karaisella aff. uzbekistanica KURBATOV
3: axial section showing streptospiral coiling in the two early whorls, sample AY 38-8; 4: oblique axial
section showing spherical proloculus, sample AY 38-1; 5: subaxial section, sample AY 38-4; 6: axial
(megalospheric specimen) and oblique subequatorial sections, sample AY38-6; 7: equatorial section of
a megalospheric specimen, sample AY 38-13; 8: equatorial section showing the chomata-like thicken-
ings in the base of the septa, as the genus C h a r e n t i a NEUMANN, sample AY 38-8; 9: equatorial sec-
tion offset by a fracture, showing thinning of the septa towards the apertural area, sample AY 38-6; 10:
oblique subequatorial section, sample AY 38-3; 11: oblique equatorial section of a probable micros-
pheric specimen revealing the keriothecal wall structure in the last whorl (arrow), sample AY 38-6; 12:
oblique section of a probable microspheric specimen revealing terminally uncoiled chambers, sample
AY 38-14; 13: subaxial section, sample AY 38-5.
9Tasli
PLATE I
10 Geologia Croatica 54/1
PLATE II
scale bar: 0.2 mm
Figs. 1-7 Kurnubia ex. gr. palastiniensis HENSON
1: axial section of a megalospheric specimen revealing an ovoid proloculus, sample AY 50; 2: axial sec-
tion showing the first order horizontal partitions only and a weakly developed central column, sample L
4-3; 3: subaxial section, sample AY 38-12; 4: subaxial section, sample AY 50; 5: oblique axial section,
sample L 4-1; 6: nearly axial section of a highly conical specimen, sample AY 40; 7: oblique transverse
section, sample L 3-2.
Figs. 8-12, 14 Kurnubia cf. morrisi REDMOND
8: oblique axial section, sample L 4-2; 9: oblique transverse-tangential section, sample L 1; 10: trans-
verse section showing the first and second order vertical partitions in the peripheral zone, sample L 7;
11: oblique axial section showing the second order partitions which are missing in the deeper part of the
peripheral zone. Note that the septa do not meet the central column, leaving an opening near the margin
of the central column, sample AY 50; 12: oblique transverse section resembling Fig. 7, included in this
species because of its larger diameter, sample AY 49; 14: subaxial section showing curved thin plates in
the central column, sample AY 42.
Figs. 13, 15, 16 Conicokurnubia orbitoliniformis SEPTFONTAINE
13: subaxial section of a highly conical specimen revealing strongly convex base in the centre, sample
AY 45; 15: subaxial section showing labyrinthian appearance of the central zone occupied by intersep-
tal pillars. Note the early portion of the cone recalling trochospiral coiling during the early ontogeny,
sample L 3; 16: transverse section passed through the adult stage of a large conical specimen, sample
AY 41.
11Tasli
PLATE II
12 Geologia Croatica 54/1
PLATE III
scale bar: 0.2 mm
Fig. 1 Everticyclammina sp.
Subaxial section showing a broadly rounded periphery which is not known in the other Jurassic species,
E. virguliana (KOECHLIN), sample AY 47.
Fig. 2 Indet. Lituolidae
Equatorial-longitudinal section, sample L 4-3.
Fig. 3 Valvulina lugeoni SEPTFONTAINE
Subaxial section, sample L 7.
Figs. 4, 5 Pfenderina sp.
4: axial section of a broken specimen revealing the keriothecal wall structure, sample L 4-3; 5: trans-
verse section, sample AY 45.
Figs. 6, 7 Verneuilina sp.
6: subaxial section, sample AY 38-9; 7: transverse section, sample AY 50.
Fig. 8 Siphovalvulina sp.
Axial section showing the spheric proloculus and siphonal canal, sample AY 43.
Fig. 9 Aeolisaccus sp.
Sample AY 47.
Fig. 10 Salpingoporella sellii (CRESCENTI)
Sample L 4-2.
Fig. 11 Salpingoporella annulata CAROZZI
Sample AY 38-14.
Fig. 12 Thaumatoporella parvovesiculifera (RAINERI)
Sample AY 38-9.
13Tasli
PLATE III
14 Geologia Croatica 54/1
