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Weurauch’s type localities: a clarification; with illustrations of types of Orthalicoidea (Mollusca, Gastropoda, Stylommatophora) in the Tucumán museum

January 2012

Authors:

Naturalis Biodiversity Center

Additional, more precise locality data are provided for the species described by Wolfgang K. Weyrauch from South America. Most of the holotypes in the Tucumán museum pertaining to the Orthalicoidea are illustrated. Additionally, a list is provided of new species described by various authors and based on Weyrauch's material, illustrating the importance of the Weyrauch collection.

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Folia conchyliologica, n°17, Mars 2012

W ey ra uc h’ s ty pe l o ca li ti es : a cl ar i f ic a ti on ;

with illustrations of typ es of O rthalicoidea (Mollusca , Gastropoda,

Sty lo mmatophora) in the Tucumán museum

by Abraham S.H. B

REURE

Abstract.

—

Additional, more precise locality data are provided for the species described by Wolfgang K. Weyrauch from

South America. Most of the holotypes in the Tucumán museum pertaining to the Orthalicoidea are illustrated. Additionally, a

list is provided of new species described by various authors and based on Weyrauch’s material, illustrating the importance of

the Weyrauch collection.

Key words

. —

Weyrauch, Argentina, Bolivia, Brazil, Ecuador, Peru, georeferenced localities.

Introduction

Wolfgang K. Weyrauch worked for many years in Peru and Argentina. He collected many shells,

especially in central and northern Peru; based on his collection he described 197 new taxa, which have been

listed by B

ARBOSA

et al. (2008). However, the type localities mentioned in that paper only give “C. Peru”, etc.,

and are too vague to be useful for further references. During revisionary work I found that a more precise

(geo)reference to these localities is often needed to interprete the status of these taxa. The following list is

therefore an addition to the publication mentioned; only those (164) taxa are herein mentioned for which

Weyrauch published type localities (thus excluding new names and taxa above the species level). A best guess

has been added in notes (bold numbers in parenthesis; see last section) whenever I was unable to locate the type

locality with certainty, e.g. when Weyrauch used local names for places that are not included in the gazetteers

used. The localities as given by Weyrauch are all translated into English. Additonally, most of the holotypes of

the Orthalicoidea (sensu B

REURE

& R

OMERO

, in press) present in the collection of the Tucumán museum are

illustrated (referred to by a figure number in parenthesis at the end of the data for the taxon); these species were

classified with the families Amphibulimidae and Orthalicidae by B

ARBOSA

et al. (2008). For the types in the

Senckenberg collection, see N

EUBERT

& J

ANSSEN

(2004).

The Weyrauch collection remains an important source of material for our understanding of Neotropical

snails, especially species from Peru, as is shown by the section listing 55 (sub)species based on his material but

described by others, and other papers which made use of his collection (e.g., P

ILSBRY

, 1944a-c, 1945, 1949;

AAS

, 1947, 1948, 1951, 1952, 1955, 1959, 1966; Z

ILCH

, 1954; L

OOSJES

, 1957; L

OOSJES

& L

OOSJES

VAN

EMMEL

, 1966; B

REURE

& E

SKENS

, 1977, 1981; B

REURE

, 1978; S

ALAS

-O

ROÑO

, 2007; P

IZÁ

& C

AZZANIGA

2009). Moreover, Weyrauch has distributed many shells under manuscript names to various museums in the

world. Some of this material has been revised (B

REURE

, 2008; B

REURE

& N

EUBERT

, 2008). In the Tucumán

museum additional, unpublished material has been found which awaits a further, critical review.

Coordinates are according the GNS gazatteer (http://geonames.nga.mil/ggmagaz/); in some cases the

Fuzzy Gazetteer (http://isodp.hof-university.de/fuzzyg/query/) has been used. Elevations, if not given by the

author, have been extracted from Google Earth (v5.2.1.1588, http://earth.google.com). Abbreviations used for

type depositories are: ANSP, Academy of Natural Sciences, Philadelphia, U.S.A.; FML, Fundación Miguel Lillo,

Tucumán, Argentina; FMNH, Field Museum of Natural History, Chicago, U.S.A.; MACN, Museo Argentino de

Ciencias Naturales, Buenos Aires, Argentina; MRCN, Museu Rio-Grandense de Ciências Naturais, Porto Alegre,

Brazil; SMF, Senckenberg Natur-Museum, Frankfurt am Main, Germany. Taxa belonging to the Orthalicoidea

are re-classified according to the systematic arrangement presented in B

REURE

& R

OMERO

(in press). Shell

heights in the legends are abbreviated with H.

A.S.H. Breure, Netherlands Centre for Biodiversity Naturalis, P.O. Box 9517, NL-2300 RA Leiden, the Netherlands

(ashbreure@gmail.com)

Folia conchyliologica, n°17, Mars 2012

Published names by Wolfgang K. Weyrauch

acobambensis, Drymaeus W

EYRAUCH

, 1967b: 482, pl. 3 figs 24-25. Type locality: Peru, [Dept. Junín] Río Tarma

valley, between Tarma and Arcobamba, Huailahuichán [11°21'30" S 075°36'11" W], 2900 m. Holotype

SMF 155694. Bulimulidae.

aequistriata, Gracilinenia W

EYRAUCH

, 1956a: 110, pl. 6 fig. 2. Type locality: Peru, [Dept. Junín] Chanchamayo

valley, on the road from Hacienda Naranjal [11°08'37" S 075°24'00" W] near the village of San Ramón to

the mine “Pichita-Caluga” (1); elevation not measured, but probably about 1300 m. Holotype FML 1531.

agitata, Pseudoglandina W

EYRAUCH

, 1967b: 486, pl. 4 fig. 53. Type locality: Peru, [Dept. Junín] Chanchamayo

valley, between La Merced [11°03'21" S 075°19'43" W] and San Ramón [11°07'17" S 075°21'11" W],

1100 m. Holotype FML 1066 (2). Bulimulidae.

agueroi, Bostryx (Peronaeus) W

EYRAUCH

, 1960c: 126, pl. 12 figs 39-41. Type locality: Peru, [Dept. Lima] right

margin of Río Cañete, road from Cañete to Yauyos, Fundo Yacca [12°30'41" S 075°55'04" W], 2300 m.

Holotype SMF 162150. Bulimulidae.

aguilari, Bostryx (Bostryx) W

EYRAUCH

, 1967a: 349, pl. 1 figs 2-6. Type locality: Peru, [Dept. Lima] ‘lomas’ de

Amancaes [12°00'38" S 077°02'25" W], 200-250 m. Holotype SMF 162163. Bulimulidae.

albicolor, Peruinia W

EYRAUCH

, 1957a: 13, pl. 1 fig. 11. Type locality: Peru, [Dept. Huánuco] left margin of Río

Tulumayo, 23 km NE Tingo Maria, near bridge in the road to Pucallpa, 670 m [09°11'25" S 075°57'26" W

(3)]. Holotype SMF 155719.

albocostata, Temesa (Temesa) W

EYRAUCH

, 1964b: 270, pl. 1 figs 1-2. Type locality: Peru, [Dept. Lima] right

margin of Río Alis, near Alis [12°16'43" S 075°47'13" W], 3300 m. Holotype SMF 162108.

altispira, Euglandina W

EYRAUCH

, 1960a: 25, pl. 4 fig. 21. Type locality, Peru, [Dept. Cajamarca,] 25 km NE

Cajamarca [sic, (4)], on path from Sócota [06°18' S 078°44' W] to San Andrés [de Cutervo; 06°13' S

078°40' W], 2400 m. Holotype SMF 162003.

altorum, Mesembrinus (Mormus) expansus W

EYRAUCH

, 1958: 129, pl. 7 fig. 12-13. Type locality: Peru, [Dept.

Junín] Chanchamayo valley, right side of Río Chanchamayo, between La Merced and San Ramón, above

Puente Herrería, Quimiri Sur [11°05'01" S 075°17'24" W, 1430 m], 1100-1300 m. Holotype SMF 156295.

Bulimulidae.

altorum, Systrophia (Systrophia) W

EYRAUCH

, 1967a: 428, pl. 6 fig. 80. Type locality: Peru, [Dept. Junín]

Chanchamayo valley, Pichita Caluga (1), 2200 m. Holotype FML 10668.

andivagus, Naesiotus W

EYRAUCH

, 1956b: 56, pl. 1 fig. 2. Type locality: Peru, [Dept. Lima] Río Huaura valley,

between Huaura and Cerro de Pasco, Cabracancha [10°49'55" S 076°53'23" W], 2200 m. Holotype FML

1461 (2). Bulimulidae.

angelmaldonadoi, Bostryx (Bostryx) modestus W

EYRAUCH

, 1960a: 31, pl. 3 figs 11-12. Type locality: Peru,

[Dept. Lima] left side of Río Lurin, ‘lomas’ E of Pachacamac [12°15'14" S 076°47'38" W], 450 m.

Holotype SMF 155595. Bulimulidae.

angiportus, Newboldius W

EYRAUCH

, 1960b: 53, pl. 8 figs 5-6. Type locality: Peru, [Dept. Junín] tributary of Río

Chanchamayo, 2 km of Pichita Caluga Mine (1), 2200 m. Holotype SMF 162045. Bulimulidae.

angispira, Bostryx (Bostryx) obliquiportus W

EYRAUCH

, 1960c: 123, pl. 11 fig. 19. Type locality: Peru, [Dept.

Lima] left side of Río Cañete, 5 km of Laraos, [Cerro] Quichao [12°20'45" S 075°48'53" W], 3450 m.

Holotype SMF 162183. Bulimulidae.

angustus, Bulimulus (Bulimulus) vesicalis W

EYRAUCH

, 1966: 45, fig. 5. Type locality: Brazil, Edo. Rio Grande

do Sul, São Leopoldo, near “Clube Airmore” stadium [29°46'52" S 051°09'25" W]. Holotype MRCN

1576a. Bulimulidae.

araozi, Bulimulus (Bulimulus) W

EYRAUCH

, 1956c: 149, pl. 11 fig. 8. Type locality: Peru, [Dept. Huánuco] left

margin of Rio Huallaga, near Tingo Maria [09°19'17" S 076°00'55" W], 670 m. Holotype SMF 155304.

Bulimulidae.

argentinus, Pupoides (Ischnopupoides) chordatus W

EYRAUCH

, 1964a: 37, fig. 1. Type locality: Argentina, Prov.

Jujuy, near Tilcara [23°34' S 065°22' W], Garganta del Diablo, 2900 m. Holotype FML 522a.

bambamarcaensis, Naesiotus (Naesiotus) W

EYRAUCH

, 1960a: 37, pl. 6 fig. 38. Type locality: Peru, [Dept.

Cajamarca] Río Llaucan valley, 4 km NW Bambamarca [06°41' S 078°32' W], Cerro Machaipungo,

3000 m. Holotype SMF 156220. Bulimulidae.

Folia conchyliologica, n°17, Mars 2012

basiplanata, Epiphragmophora W

EYRAUCH

, 1960a: 43, pl. 6 fig. 39. Type locality: Peru, [Dept. Pasco] road

from Cerro de Pasco to Huánuco, valley of Río Payanchan, 3800 m (5). Holotype SMF 162033.

beltrani, Bostryx (Peronaeus) agueroi W

EYRAUCH

, 1964a: 52, figs. 9-10. Type locality: Peru, [Dept. Lima] left

side of Río Cañete, between Laraos [12°20'48" S 075°47'11" W] and Tintín [12°18'10" S 075°48'40" W],

3450 m. Holotype SMF 162152. Bulimulidae.

bequaerti, Drymaeus W

EYRAUCH

, 1956c: 154, pl. 11 figs 12-14. Type locality: Peru, [Dept. Huánuco] near

Tingo Maria, right side of Río Monzón near its confluence with Rio Huallaga [09°19'17" S 076°00'55"

W], 670 m. Holotype SMF 155309. Bulimulidae.

bequaerti, Temesa (Neniatracta) W

EYRAUCH

, 1957: 25, pl. 1 fig. 12. Type locality: Peru, [Dept. Cajamarca]

1 km NE Bambamarca [06°41' S 078°32' W], 2800 m. Holotype SMF 155720.

bermudezae, Bostryx (Pseudoperonaeus) W

EYRAUCH

, 1958: 111. pl. 9 figs 38-40. Type locality: Peru, [Dept.

Lima] Rio Cañete valley, Laraos [12°20'48" S 075°47'11" W], 3600 m. Holotype SMF 156350.

Bulimulidae.

bicolor, Naesiotus (Naesiotus) W

EYRAUCH

, 1967a: 408, pl. 6 fig. 83. Type locality: Peru, [Dept. Cuzco] Sicuani

[14°16'10" S 071°13'34" W, 3500 m], 3200 m. Holotype FML 3987 (Pl. 2 figs 17-18). Bulimulidae.

birabenorum, Bostryx (Lissoacme) W

EYRAUCH

, 1965a: 71, figs 1-3. Type locality: Argentina, Prov. Tucumán,

near Tafí del Valle [26°51'10" S 065°42'35" W], El Infernillo, 3042 m. Holotype FML 985a (Pl. 1 fig. 1).

Bulimulidae.

cajamarcana, Steeriana (Steeriana) W

EYRAUCH

& Z

ILCH

in Z

ILCH

, 1954: 73, fig. 6, pl. 5 fig. 8. Type locality:

Peru, [Dept. Cajamarca] 18 km from Cajamarca on the road to San Marcos, Hacienda Cochambul,

2800 m (6). Holotype SMF 135516.

camachoi, Neopetraeus W

EYRAUCH

, 1967a: 418, pl. 5 figs 68-70. Type locality: Peru, [Dept. Cajamarca] left

margin of Río Chotano, Cuesta de Chuguid, 1500 m (7). Holotype FML 1240a (Pl. 3 figs 29-31).

Bulimulidae.

celendinensis, Drymaeus W

EYRAUCH

, 1956c: 151, pl. 11 figs 10-11. Type locality: Peru, [Dept. Cajamarca] hill

2 km W Celendín [06°52'03" S 078°09'52" W, 2815 m], 2700 m. Holotype SMF 155307. Bulimulidae.

celendinensis, Steeriana (Steeriana) W

EYRAUCH

& Z

ILCH

in Z

ILCH

, 1954: 70, fig. 3, pl. 5 fig. 6. Type locality:

Peru, [Dept. Cajamarca] km 113 road [Cajamarca-]Balsas, 3 km SE Celendin [06°53'12" S 078°07'40" W,

2900 m], 2700 m. Holotype SMF 135517.

cerrateae, Epiphragmophora (Karlschmidtia) W

EYRAUCH

, 1960a: 47, pl. 6 fig. 40. Type locality: Peru, [Dept.

Ancash] 280 km N Lima, Río Pativilca [valley], 10 km N Chiquián, Aquia [10°04'27" S 077°08'39" W],

3350 m. Holotype SMF 162030.

cerrateae, Hemicena W

EYRAUCH

, 1958: 96, figs 1-2, pl. 6 fig. 9. Type locality: Peru, [Dept. Ancash] 280 km N

Lima, Río Pativilca [valley], 10 km N Chiquián, Aquia [10°04'27" S 077°08'39" W], 3350 m. Holotype

SMF 140714.

cerrateae, Naesiotus (Raphiellus) [sic, = Rhaphiellus] W

EYRAUCH

, 1967a: 410, pl. 5 fig. 77. Type locality: Peru,

[Dept. Ancash] 280 km N Lima, Río Pativilca [valley], 10 km N Chiquián, Aquia [10°04'27" S

077°08'39" W], 3350 m. Holotype SMF 162024. Bulimulidae.

chamayensis, Naesiotus (Naesiotus) subcostatus W

EYRAUCH

, 1967a: 409, pl. 7 figs 97-99. Type locality: Peru,

[Dept. Cajamarca,] left margin of Río Chamaya, near Chamaya [05°49' S 078°42' W], 550-660 m.

Holotype SMF 162190. Bulimulidae.

chiletensis, Scutalus (Scutalus) W

EYRAUCH

, 1967a: 373, pl. 2 figs 24-30. Type locality: Peru, [Dept. Cajamarca]

left margin of Río Jequetepeque, mountain opposite Chilete [07°13'18" S 078°50'40" W], 850-950 m.

Holotype FML 1236a (Pl. 5 figs 46-47). Orthalicidae.

chusgonensis, Bostryx (Bostryx) W

EYRAUCH

, 1960a: 30, pl. 3 figs 16-17. Peru, [Dept. La Libertad] right margin

of Río Chusgon, Hacienda Santa Elena, 1550 m [07°30'26" S 077°47'26" W]. Holotype SMF 162013.

Bulimulidae.

cleliae, Zilchogyra W

EYRAUCH

, 1965b: 124, pl. 7 fig. 2 [sic, fig. 3]. Type locality: Argentina, Buenos Aires, near

the Zoological Garden, park between Avenida Sarmiento and Avenida Figueroa. Holotype FML 977.

Folia conchyliologica, n°17, Mars 2012

combinai, Mesembrinus (Ornatimormus) W

EYRAUCH

, 1958: 135, pl. 8 fig. 16. Type locality: Peru, [Dept. Junín]

Chanchamayo valley, between La Merced and San Ramón, Quimiri Sur [11°05'01" S 075°17'24" W],

1300 m. Holotype SMF 156201. Bulimulidae.

compactus, Bostryx (Bostryx) zilchi W

EYRAUCH

, 1960c: 125, pl. 12 figs 27-28. Type locality: Peru, [Dept. Lima]

left margin of Río Cañete, footpath between Laraos [12°20'48" S 075°47'11" W] and Tintín [12°18'10" S

075°48'40" W], 3450 m. Holotype SMF 162156. Bulimulidae.

costatus, Bostryx (Bostryx) pygmaeus W

EYRAUCH

, 1960c: 122, pl. 11 figs 12-16. Type locality: Peru, [Dept.

Lima] right margin of Río Alis, near Alis [12°16'43" S 075°47'13" W], 3300 m. Holotype SMF 162098.

Bulimulidae.

costifer, Bostryx (Elatibostryx) imeldae W

EYRAUCH

, 1960c: 129, pl. 12 figs 35-38. Type locality: Peru, [Dept.

Lima] 5 km from Laraos on footpath to Yauyos, left margin of Río Mayo [Río Laraos], [Cerro] Quichao

[12°20'45" S 075°48'53" W], 3500 m. Holotype SMF 162101. Bulimulidae.

costulatus, Scutalus (Vermiculatus) W

EYRAUCH

1967a: 395, pl. 3 fig. 45. Type locality: Peru, [Dept. Junín] near

Junín, Huarmipuquio [11°10'09" S 076°02'25" W], 4200 m. Holotype SMF 162068. Bulimulidae.

crassicostata, Temesa (Temesa) decimvolvis W

EYRAUCH

1958: 102, pl. 6 fig. 2. Type locality: Peru [Dept. Junín]

road La Oroya-Huancayo, left side of Río Mantaro, Curicaca (8). Holotype SMF 156231.

crenulatus, Lopesianus W

EYRAUCH

, 1958: 121, pl. 6 figs 7-8. Type locality: Brazil, Edo. Rio de Janeiro, Cabo

Frio [22°53' S 042°00' W], Arraial-Praínha. Holotype SMF 156356. Bulimulidae.

crucilineatus, Bostryx (Peronaeus) W

EYRAUCH

, 1967a: 361, pl. 1 fig. 14. Type locality: Peru, [Dept. Lima,]

valley of Río Santa Eulalia, 2500 m (9). Holotype FML 1241 (not photographed). Bulimulidae.

cuencaensis, Temesa (Neniatracta) adusta W

EYRAUCH

, 1964c: 152, figs 7-8. Type locality: Peru, [Dept.

Huancavelica] Río Mantaro valley, near Mejorada, Cuenca [12°26' S 075°02' W], 2960 m. Holotype SMF

156217.

cuzcoensis, Scutalus (Vermiculatus) W

EYRAUCH

, 1967a: 396, pl. 4 figs 47-48. Type locality: Peru, [Dept. Cuzco]

7 km S Cuzco, [Cerro] Huancaro [14°21' S 072°01' W], Kachuna, 3500 m. Holotype FML 1225 (pl. 6

figs 55-56). Bulimulidae.

cylindricus, Bostryx (Pseudoperonaeus) W

EYRAUCH

, 1960c: 127, pl. 11 fig. 3. Type locality: Peru, [Dept. Lima]

right margin of Río Mayo [Río Laraos], 1 km from Laraos [12°20'48" S 075°47'11" W], 3500 m.

Holotype SMF 162107. Bulimulidae.

debilisculptus, Scutalus (Scutalus) coraeformis W

EYRAUCH

, 1967a: 376, pl. 2 figs 20-23, pl. 9 figs 137-138.

Type locality: Peru, [Dept. Cajamarca] Río Jequetepeque, 16 km N Tembladera on the road Pacasmayo-

Cajamarca, near Quinden, 600-700 m (10). Holotype FML 1984 (Pl. 5 figs 49-51). Orthalicidae.

debilisculptus, Thaumastus (Thaumastiellus) occidentalis W

EYRAUCH

, 1960a: 30, pl. 3 fig. 15. Type locality:

Peru, [Dept. Cajamarca] road Chiclayo-Cutervo, ca. 85 km NE Chiclayo, near Llama [06°30'52" S

079°07'13" W], 2000-2250 m. Holotype SMF 162029. Megaspiridae.

decimvolvis, Temesa (Temesa) W

EYRAUCH

, 1957: 21, pl. 1 fig. 13. Type locality: Peru, [Dept. Junín] left side of

Río Mantaro, road La Oroya-Huancayo, Acaya [11°48'08" S 075°35'53" W], 3450 m. Holotype SMF

156215.

dedicata, Andinia (Ehrmanniella) W

EYRAUCH

& Z

ILCH

in Z

ILCH

, 1954: 68, fig. 5, pl. 5 fig. 5. Type locality:

Peru, [Dept. Pasco] road from Cerro de Pasco to Huánuco, valley of Río Payanchan, 3800 m (5).

Holotype SMF 135515.

densestrigatus, Mesembrinus (Ornatimormus) henrypilsbryi W

EYRAUCH

, 1958: 134, pl. 8 fig. 20. Type locality:

Peru, [Dept. Junín] Chanchamayo valley, right margin of Río Chamchamayo, between La Merced and

San Ramón, Quimiri Sur [11°05'01" S 075°17'24" W], above Puente Herrería, 1300-1400 m. Holotype

SMF 156293. Bulimulidae.

elegantulus, Naesiotus W

EYRAUCH

, 1956b: 4, pl. 1 fig. 1. Type locality: Peru, [Dept. Cajamarca] W of Celendín,

Cerro San Isidro [06°56'15" S 078°08'10" W], 2750-2800 m. Holotype SMF 162023. Bulimulidae.

eliseoduartei, Systrophia (Scolodonta) W

EYRAUCH

, 1966: 46, fig. 6. Type locality: Uruguay, [Prov. Maldonado]

80 km E Montevideo, Cerro de las Ánimas [34°42' S 055°19' W], 300 m. Holotype FML 10686a.

Folia conchyliologica, n°17, Mars 2012

fernandezae, Naesiotus (Maranhoniellus) W

EYRAUCH

, 1958: 122, pl. 9 figs 45-46. Type locality: Peru, [Dept.

Amazonas] right margin of Río Marañon, near Balsas [06°50'41" S 078°01'41" W], 850 m. Holotype

SMF 157277. Bulimulidae.

flavilabrum, Drymaeus (Mormus) expansus W

EYRAUCH

, 1967b: 484, pl. 3 fig. 29. Type locality: Peru, [Dept.

Ucayali,] road Tingo Maria-Pucallpa, Río Aguaytia [09°02'35" S 075°30'09" W, 275 m]. Holotype FML

1197 (Pl. 4 figs 40-42). Bulimulidae.

florezi, Floreziellus W

EYRAUCH

, 1967b: 489, pl. 1 figs 7-17. Peru, [Dept. Apurimac,] 60 km W Cuzco, on the

road to Abancay, near Puente Cunyac [13°33'45" S 072°34'29" W], 2100 m. Holotype FML 10671a (Pl. 1

figs 4-5). Bulimulidae.

florezi, Happia (Happia) W

EYRAUCH

, 1965a: 75, fig. 6. Type locality: Peru, [Dept. Cuzco, Prov.] Espinar,

District Pichigua [14°40'53" S 071°25'01" W], Katahuihuayca, 4400 m. Holotype FML 10644a.

florezi, Incania W

EYRAUCH

, 1964b: 41, fig. 3. Type locality: Peru, [Dept. Cuzco] NE Cuzco, km 122 road

Paucartambo-Cosñipata, near a tunnel, 2800 m. Holotype FML 1218.

florezi, Phenacotaxus (Ataxellus) W

EYRAUCH

, 1967a: 369, pl. 4 figs 49-50. Type locality: Peru, [Dept. Cuzco]

25 km [NE] Cuzco [by road], Pisac [13°25'21" S 071°50'48" W], 3100 m. Holotype FML 10522a (Pl. 1

figs 8-9). Bulimulidae.

florezi, Radiodiscus W

EYRAUCH

, 1965c: 105, fig. 1. Type locality: Peru, [Dept. Cuzco] 190 km S Cuzco, [Prov.]

Espinar, Palpata [= Hector Tejada, 14°53'02" S 071°13'15" W], Accoyo-Chacsilla, 4700 m. Holotype

FML 10468a.

florezi, Systrophia (Scolodonta) W

EYRAUCH

, 1967a: 433, pl. 6 fig. 86. Type locality: Peru, [Dept. Cuzco] 210 km

NE Cuzco, near Cosñipata [13°04' S 071°11' W], Coloradito, 990 m. Holotype FML 10621a.

franzi, Zilchogyra W

EYRAUCH

, 1965c: 112, fig. 4. Type locality: Argentina, Prov. Buenos Aires, Sierra de la

Ventana, E of Bahía Blanca, 200 m (11). Holotype FML 1217.

geophilus, Naesiotus (Naesiotus) W

EYRAUCH

, 1967b: 477, pl. 3 figs 26-28. Type locality: Peru, [Dept.

Amazonas] Río Marañon valley, W of Cruce Bagua Grande [05°41'59" S 078°31'08" W], near the

bifurcation of the road from Chiclayo to Bagua Grande and Bagua Chica, 600 m (12). Holotype FML

1074a (2). Bulimulidae.

giganteus, Radiodiscus W

EYRAUCH

, 1958: 105, pl. 9a fig. 51. Type locality: Peru, [Dept. Pasco] road from Cerro

de Pasco to Huánuco, valley of Río Payanchan, 3800 m (5). Holotype SMF 156342.

globosus, Bostryx (Lissoacme) W

EYRAUCH

, 1967a: 355, figs 11-12. Type locality: Peru, [Dept. Ica] Paredones de

Nazca [14°51' S 074°56' W, 660 m]. Holotype FML 1226 (Pl. 1 fig. 2). Bulimulidae.

glomeratus, Bostryx (Bostryx) zilchi W

EYRAUCH

, 1960c: 124, pl. 12 figs 29-34. Type locality: Peru, [Dept.

Lima,] 1 km from Laraos on footpath to Yauyos, [Cerro] Quichao [12°20'45" S 075°48'53" W], 3500 m.

Holotype SMF 162104. Bulimulidae.

golbachi, Radiodiscus W

EYRAUCH

, 1965c: 106, fig. 2. Type locality: Argentina, Prov. Tucumán, Sierra del

Nogalito [26°22' S 065°00' W], 1400 m. Holotype FML 724a.

gracilis, Bostryx (Scansicohlea) W

EYRAUCH

, 1967a: 360, pl. 1 fig. 1. Type locality: Peru, [Dept. Lima] right

margin of Río Rimac, near [below] Tambo [de] Viso [11°48'02" S 076°20'51" W, 3340 m], 2750 m.

Holotype FML 1102 (Pl. 1 figs 10-11). Bulimulidae.

gracillimus, Naesiotus W

EYRAUCH

, 1956b: 6, pl. 1 fig. 3. Type locality: Peru, [Dept. Huánuco] near Tingo

Maria, Río Monzón near its confluence with Rio Huallaga [09°19'17" S 076°00'55" W], 670 m. Holotype

SMF 155696. Bulimulidae.

grandiportus, Bostryx (Bostryx) bromeliarum W

EYRAUCH

, 1958: 109, pl. 8 figs 18-19. Type locality: Peru,

[Dept. Lima] right margin of Río Rimac, near [below] Tambo [de] Viso [11°48'02" S 076°20'51" W,

3340 m], 2700 m. Holotype SMF 156367. Bulimulidae.

grandiportus, Zilchiella W

EYRAUCH

, 1957: 10, pl. 1 figs 5-10. Type locality: Peru, [Dept. Cajamarca] Río

Llaucan, Peña Rota [? 06°37'22" S 078°28'38" W], 2550-2750 m. Holotype SMF 155710.

grandiventris, Scutalus W

EYRAUCH

, 1960a: 42, pl. 5 figs 27-33. Type locality: Peru, [Dept. Cajamarca] between

Trujillo and Cajamarca, ca. 100 km NE Trujillo, above Cascas [07°29' S 078° 49' W], 1400 m. Holotype

SMF 155690. Orthalicidae.

Folia conchyliologica, n°17, Mars 2012

granulatus, Scutalus (Scutalus) chiletensis W

EYRAUCH

, 1967a: 375, pl. 2 figs 31-32. Type locality: Peru [Dept.

Cajamarca] right margin of Río Jequetepeque, hill N of Chilete [07°13'02" S 078°50'26" W], 850 m.

Holotype FML 1362a (Pl. 5 fig. 48). Orthalicidae.

granulosa, Epiphragmophora W

EYRAUCH

, 1960a: 46, pl. 6 fig. 37. Type locality: Peru, [Dept. Cuzco] 20 km N

Cuzco, Pisac [13°25'21" S 071°50'48" W], 3200-3300 m. Holotype SMF 162041.

haasi, Bostryx (Bostryx) W

EYRAUCH

, 1960a: 33, pl. 5 fig. 35. Type locality: Peru, [Dept. Lima] Río Rimac,

below Tambo [de] Viso [11°48'02" S 076°20'51" W, 3340 m], 2600 m. Holotype SMF 156370.

Bulimulidae.

haasi, Columbinia (Pfeifferiella) W

EYRAUCH

, 1957: 4, pl. 1 figs 1-2. Type locality: Peru, [Dept. Cajamarca]

85 km NE Chiclayo, near Llama [06°30'42" S 079°07'07" W], Cerro Chunrun, 2350 m. Holotype SMF

155715.

haasi, Llaucanianus W

EYRAUCH

, 1967a: 421, pl. 3 figs 34-36. Type locality: Peru, [Dept. Cajamarca] 8 km NE

Bambamarca, left margin of Río Llaucán, Peña Rota, 2700 m [? 06°37'22" S 078°28'38" W]. Holotype

SMF 162046. Bulimulidae.

haasi, Naesiotus W

EYRAUCH

, 1956b: 7, pl. 1 fig. 5. Type locality: Peru, [Dept. Cajamarca] right margin of Río

Jequetepeque, highway Pacasmayo-Cajamarca, mountain near Chilete [07°13'02" S 078°50'26" W],

850 m. Holotype FML 2862 (2). Bulimulidae.

haasi, Systrophia (Systrophia) W

EYRAUCH

, 1960a: 28, pl. 3 fig. 7. Type locality: Peru, [Dept. Cajamarca]

between Cutervo and Chota, Río Chotano, El Túnel, 2100 m. Holotype SMF 162000.

hernandezae, Littorina W

EYRAUCH

, 1963a: 251, [pl. 1] figs 1-2. Type locality: Peru, [Dept. Cuzco] Cuzco-Las

Salinas, 3600 m (subfossil). Holotype FML 1001a.

hyltonscottae, Zilchogyra W

EYRAUCH

, 1965c: 114, fig. 5. Type locality: Argentina, Prov. Tucumán, valley Río

Pueblo Viejo, 600 m. Holotype FML 976.

imeldae, Bostryx (Elatibostryx) W

EYRAUCH

, 1958: 113, pl. 9 fig. 37. Type locality: Peru, [Dept. Lima] 135 km

ESE Lima, Rio Cañete valley, Laraos [12°20'48" S 075°47'11" W], 3600 m. Holotype SMF 156347.

Bulimulidae.

inflatiportus, Bostryx (Bostryx) obliquiportus W

EYRAUCH

, 1960c: 123, pl. 12 figs 22-26. Type locality: Peru,

[Dept. Lima] right margin of Río Alis, near Alis [12°16'43" S 075°47'13" W], 3300 m. Holotype SMF

162144. Bulimulidae.

isidroensis, Steeriana (Steeriana) W

EYRAUCH

& Z

ILCH

in Z

ILCH

, 1954: 72, fig. 4, pl. 5 fig. 7. Type locality:

Peru, [Dept. Cajamarca] near Celendín, Cerro San Isidro [06°56'15" S 078°08'10" W], 2750-2800 m.

Holotype SMF 135511.

klappenbachi, Helicina (Trichohelicina) W

EYRAUCH

, 1966: 42, figs 1-3. Type locality: Argentina, Prov.

Misiones, [Arroyo] Soberbio [27°17'20" S 054°11'40" W, 137 m]. Holotype FML 11215a.

lachayensis, Scutalus (Scutalus) versicolor W

EYRAUCH

, 1967a: 383, figs 109-115. Type locality: Peru, [Dept.

Lima] 90 km N Lima, near Chancay, ‘lomas’ de Lachay [11°21'52" S 077°21'30" W], 300-350 m.

Holotype FML 147a (2). Orthalicidae.

laraosensis, Bostryx (Bostryx) obliquiportus W

EYRAUCH

, 1960c: 124, pl. 11 figs 20-21. Type locality: Peru,

[Dept. Lima] Río Cañete valley and on the left margin of Río Mayo [Río Laraos], 5 km from Laraos, on

foothpath to Yauyos, [Cerro] Quichao [12°20'45" S 075°48'53" W], 3500 m. Holotype SMF 162187.

Bulimulidae.

laraosensis, Temesa (Temesa) pilsbryi W

EYRAUCH

, 1960c: 119, pl. 11 fig. 7. Type locality: Peru, [Dept. Lima]

right margin of Río Mayo [Río Laraos], 1 km from Laraos [12°20'48" S 075°47'11" W], 3500 m.

Holotype SMF 162117.

latecolumellaris, Naesiotus (Naesiotellus) W

EYRAUCH

, 1967a: 415, pl. 5 figs 71-72. Type locality: Peru, [Dept.

Lima] right margin of Río Cañete, 1 km above Puente El Auco [12°34'56" S 075°57'27" W], 2100 m.

Holotype FML 1076 (Pl. 2 figs 19-20). Bulimulidae.

latestriata, Temesa (Temesa) W

EYRAUCH

, 1958: 100, pl. 6 fig. 1. Type locality: Peru, [Dept. Junín] road Tarma-

Jauja, above Acolla [11°43'55" S 075°32'47" W], 3700 m. Holotype SMF 156232.

lateumbilicatus, Radiodiscus W

EYRAUCH

, 1966: 44, fig. 4. Type locality: Argentina, Prov. Mendoza, nearby and

above Villavicencio [32°35'08" S 068°59'03" W], 1900 m. Holotype FML 11005a.

Folia conchyliologica, n°17, Mars 2012

lizarasoae, Bostryx (Pseudoperonaeus) W

EYRAUCH

, 1967a: 363, pl. 3 figs 38-39. Type locality: Peru, [Dept.

Huánuco] Río Huallaga valley, near Ambo [10°07'51" S 076°12'17" W], Cerro Arcupumco, 2200 m.

Holotype SMF 162009. Bulimulidae.

longispira, Bostryx (Pseudoperonaeus) W

EYRAUCH

, 1960c: 128, pl. 11 figs 4-5. Type locality: Peru, [Dept.

Lima,] right margin of Río Cañete, road from Magdalena to Yauyos, 2600 m [12°29'08" S 075°54'48" W].

Holotype SMF 162112. Bulimulidae.

macedoi, Scutalus (Vermiculatus) W

EYRAUCH

, 1967a: 398, pl. 3 fig. 42-44. Type locality: Peru, [Dept. Junín]

Lake Junín, Capillacocha, 4150 m. Holotype SMF 162070. Bulimulidae.

mantaroensis, Temesa (Temesa) decimvolvis W

EYRAUCH

, 1963b: 277, pl. 1 fig. 7. Type locality: Peru, [Dept.

Junín] left margin of Río Mantaro, near bridge in the road [La] Oroya-Huancayo, near Llocllapampa

[11°49'04" S 075°37'26" W], 3460 m. Holotype SMF 156237.

marasensis, Scutalus (Vermiculatus) cuzcoensis W

EYRAUCH

, 1967a: 398, pl. 2 fig. 33. Type locality: Peru, Dept.

Cuzco, Maras [13°20'06" S 072°09'24" W], 3000 m. Holotype FML 10641a (Pl. 6 fig. 57). Bulimulidae.

maximus, Thaumastus (Quechua) salteri W

EYRAUCH

, 1967a: 347, pl. 9 fig. 135. Type locality: Peru, [Dept.

Cajamarca] 25 km NE Cutervo [by road], trail between Sócota [06°18'59" S 078°44'37" W, 2464 m] and

San Andrés [de Cutervo; 06°13' S 078°40' W, 1940 m], Peña Blanca, 2600 m. Holotype SMF 156381.

Orthalicidae.

minor, Bostryx (Bostryx) haasi W

EYRAUCH

, 1960a: 35, pl. 5 fig. 34. Type locality: Peru, [Dept. Lima] road

Lima-La Oroya, 90 km NE Lima, right margin Río Rimac, above San Mateo [11°45'34" S 076°18'02" W],

3300 m. Holotype SMF 156371. Bulimulidae.

minor, Steeriana (Steeriana) celendinensis W

EYRAUCH

, 1958: 92, pl. 6 fig. 6. Type locality: Peru, [Dept.

Cajamarca] Río Yanguate valley, ca. 5 km E Celendín, ca. 2150 m. Holotype SMF 156212.

minor, Temesa (Temesa) decimvolvis W

EYRAUCH

, 1963b: 276, pl. 1 figs 9-10. Type locality: Peru, [Dept. Junín]

right margin of Río Mantaro, N of [La] Oroya, near Pacha, 3500 m. Holotype FML 1046a.

mirabilis, Epiphragmophora W

EYRAUCH

, 1960c: 129, pl. 12 fig. 42. Type locality: Peru, [Dept. Cajamarca,

30 km NE Cutervo, above San Andrés [de Cutervo; 06°13' S 078°40' W, 1940 m], entrance to Guacharo

cave, 2650 m. Holotype SMF 162064.

mirandoi, Potamopyrgus W

EYRAUCH

, 1963a: 244, [pl. 1] figs 3-4. Type locality: Peru, Dept. Cajamarca, Prov.

Cajabamba, District Citacocha, Hacienda Jocos, 2600 m. Holotype FML 1000a.

multiguttatus, Drymaeus (Ornatimormus) W

EYRAUCH

, 1964a: 55, fig. 8. Type locality: Peru, [Dept. Huánuco]

road Tingo Maria-Pucallpa, Boquerón de Abad [08°49'60" S 075°55'00" W], 550 m. Holotype FML 1200

(Pl. 3 figs 32-34). Bulimulidae.

obesus, Drymaeus (Diaphanomormus) coelestini W

EYRAUCH

, 1964a: 58, fig. 11. Type locality: Peru, [Dept.

Huánuco] road Tingo Maria-Pucallpa, Boquerón de Abad [08°49'60" S 075°55'00" W], 550 m. Holotype

FML 1196a (Pl. 3 figs 35-36). Bulimulidae.

obesus, Neopetraeus arboriferus W

EYRAUCH

, 1967a: 416, pl. 5 figs 62-63. Type locality: Peru [without further

specification]. Holotype FML 1244a (Pl. 2 figs 23-25). Bulimulidae.

obliquiportus, Bostryx (Bostryx) W

EYRAUCH

, 1958: 110, pl. 9 fig. 35-36. Type locality: Peru, [Dept. Lima] 155

ESE Lima, Rio Cañete valley, Laraos [12°20'48" S 075°47'11" W], 3600 m. Holotype SMF 156353.

Bulimulidae.

occidentalis, Thaumastus (Thaumastiellus) W

EYRAUCH

, 1960a: 28, pl. 3 figs 13-14. Type locality: Peru, [Dept.

Cajamarca] 110 km NNE Trujillo, near Contumazá [07°22'06" S 078°48'19" W], 2750 m. Holotype SMF

162026. Megaspiridae.

omissa, Temesa (Temesa) W

EYRAUCH

, 1957: 18, pl. 1 fig. 4. Type locality: Peru, [Dept. Junín] road Tarma-Jauja,

above Tarmatambo [11°28'15" S 075°41'12" W, 3450 m], 3400-3700 m. Holotype SMF 140715.

omissus, Scutalus (Vermiculatus) W

EYRAUCH

, 1967a: 400, pl. 8 figs 116-120. Type locality: Peru, [Dept.

Ancash] few km SE Huaráz, Shaurama [09°32' S 077°32' W], 3200-3300 m. Holotype SMF 155586.

Bulimulidae.

orcesi, Mesembrinus (Mormus) expansus W

EYRAUCH

, 1958: 130, pl. 7 fig. 15. Type locality: Ecuador, [Prov.

Pastaza] 270 km SW [sic, SE] Quito, Rio Bobonaza valley, tributary of Río Pastaza, Montalvo [02°04' S

076°58' W], 314 m. Holotype SMF 156292. Bulimulidae.

Folia conchyliologica, n°17, Mars 2012

orcesi, Thaumastus (Thaumastus) W

EYRAUCH

, 1967b: 473, pl. 1 fig. 2. Type locality: Ecuador, Prov. Pichincha,

35 km NE Quito, region of Nanegal [00°07' N 078°40' W], ca. 1500 m. Holotype FML 3165 (Pl. 6 figs

59-61). Megaspiridae.

ormeai, Epiphragmophora W

EYRAUCH

, 1956c: 158, pl. 11 fig. 19. Type locality: Peru, [Dept. La Libertad] ca.

140 km E Huamachuco, between [Quebrada La] Sata [08° 24' 34'' S 077° 06' 35'' W] and Parcoy

[07°58'16" S 077°25'44" W]. Holotype SMF 155315.

ortizi, Bostryx (Bostryx) W

EYRAUCH

, 1967a: 352, pl. 1 fig. 13. Type locality: Peru, [Dept. ?] between

Huamboyacu and La Colmena, Sotopampa, 1600 m (13). Holotype FML 1126 (Pl. 1 fig. 3). Bulimulidae.

ortizpuentei, Scutalus (Scutalus) W

EYRAUCH

, 1967a: 378, pl. 7 fig. 100. Type locality: Peru, [Dept. Cajamarca]

between Chiclayo and Chota, valley of Río Chanchay [ca. 06°38' S 079°05' W], 80 km N Quinden.

Holotype FML 1234 (Pl. 5 figs 52-54). Orthalicidae.

palizae, Solariopsis (Psadariella) W

EYRAUCH

, 1956c: 160, pl. 11 figs 15-17. Type locality: Peru, [Dept. La

Libertad] road Huamachuco-Parcoy, Chagual [07°50'08" S 077°38'03" W], ca. 1300 m. Holotype SMF

155311.

paucistrigatus, Neopetraeus arboriferus W

EYRAUCH

, 1967a: 417, pl. 5 figs 65-67. Type locality: Peru, [Dept. La

Libertad] road Huamachuco-Parcoy, Chagual [07°50'08" S 077°38'03" W], ca. 1300 m. Holotype FML

1239a (Pl. 2 figs 26-28). Bulimulidae.

peranoi, Littoridina W

EYRAUCH

, 1963a: 252, [pl. 1] figs 5-6. Type locality: Argentina, Prov. Tucumán, Serranía

del Timbó. Holotype FML 41a.

peruvianus, Pupoides (Pupoides) albilabris W

EYRAUCH

, 1960c: 117, pl. 11 figs 1-2. Type locality: Peru, [Dept.

Lima] 50 km NE Lima, road Lima-La Oroya, Torcona, 1250 m. Holotype SMF 162160.

peterseni, Zilchogyra W

EYRAUCH

, 1965b: 125, pl. 7 fig. 3 [sic, fig. 2]. Type locality: Peru, [Dept. Tumbes]

middle reaches of Río Tumbes. Holotype SMF 165245a.

pichitacalugaensis, Mesembrinus (Ornatimormus) henrypilsbryi W

EYRAUCH

, 1958: 135, pl. 8 fig. 17. Type

locality: Peru, [Dept. Junín] Chanchamayo valley, 2 km of Pichita Caluga Mine (1), 2200 m. Holotype

SMF 156388. Bulimulidae.

pilosus, Scutalus (Vermiculatus) W

EYRAUCH

, 1967a: 403, pl. 3 fig. 40-41. Type locality: Peru, [Dept. Junín]

43 km W Junín, [Cerro] Incapirca [11°00'39" S 076°12'15" W], 4200 m. Holotype SMF 162065.

Bulimulidae.

pilsbryi, Drymaeus W

EYRAUCH

, 1956c: 153, pl. 11 fig. 7 (14). Type locality: Peru, [Dept. Junín] Rio Tarma

valley, [Quebrada] Pan de Azucár [11°10'42" S 075°27'09" W, 1580 m], 1350 m. Holotype SMF 155303.

Bulimulidae.

pilsbryi, Naesiotus W

EYRAUCH

, 1956b: 6, pl. 1 fig. 4. Type locality: Peru, [Dept. La Libertad] 210 km E Trujillo,

road Huamachuco-Parcoy, near Chagual [07°50'08" S 077°38'03" W], Pampa Calquiche, 1200 m.

Holotype SMF 155698. Bulimulidae.

pilsbryi, Systrophia (Systrophia) obvoluta W

EYRAUCH

, 1958: 106, pl. 6 figs 4-5. Type locality: Peru, [Dept.

Huánuco] Tingo Maria, 670 m. Holotype SMF 156344.

pilsbryi, Temesa W

EYRAUCH

, 1956c: 146, pl. 11 figs 1-4. Type locality: Peru, Dept. Lima, right margin Río

Rimac, El Infernillo [11°42'22" S 076°16'51" W], 3360-3370 m. Holotype SMF 155296.

planispira, Systrophia (Systrophia) W

EYRAUCH

, 1967a: 429, pl. 6 fig. 82. Type locality: Peru, [Dept. Cajamarca]

road Chiclayo-Cutervo, near Llama [06°30'52" S 079°07'13" W], Cerro Churán, 2350 m. Holotype FML

1266a.

primigenia, Temesa (Temesa) pilsbryi W

EYRAUCH

, 1960c: 120, pl. 11 fig. 8. Type locality: Peru, [Dept. Lima]

right margin of Río Alis, near Alis [12°16'43" S 075°47'13" W], 3300 m. Holotype SMF 162147.

pygmaea, Temesa (Temesa) albocostata W

EYRAUCH

, 1963b: 272, pl. 1 figs 3-4. Type locality: Peru, [Dept. Lima]

left margin of Mayo Creek [Río Laraos], tributary to Río Cañete, 5 km from Laraos on the trail to Yauyos,

[Cerro] Quichao, 3500 m [12°20'39" S 075°48'47" W]. Holotype SMF 162111.

pygmaeus, Bostryx (Bostryx) W

EYRAUCH

, 1960c: 121, pl. 11 figs 10-11. Type locality: Peru, [Dept. Lima] right

margin of Río Alis, 1 km from Alis [12°16'43" S 075°47'13" W], on the road to Tintín, 3250 m. Holotype

SMF 162127. Bulimulidae.

Folia conchyliologica, n°17, Mars 2012

quadritaeniatus, Drymaeus (Orodrymaeus) farrisi W

EYRAUCH

, 1956c: 150, pl. 11 fig. 9. Type locality: Peru,

[Dept. La Libertad (?)] right margin of Río Chusgon, Hacienda Santa Elena, 1550 m [07°30'26" S

077°47'26" W]. Holotype SMF 155306. Bulimulidae.

rehderi, Bostryx (Elatibostryx) W

EYRAUCH

, 1960a: 35, pl. 3 figs 4-5. Type locality: Peru, [Dept. Lima] Río

Huaura, 2-3 km NNE Churin [10°48'39" S 076°52'36" W], on the road to Oyón, 2400-2450 m. Holotype

SMF 156386. Bulimulidae.

rodriguezae, Bostryx (Bostryx) W

EYRAUCH

, 1967b: 475, pl. 3 figs 33-48. Type locality: Peru, [Dept. Lima] left

margin of Mayo Creek [Río Laraos], tributary to Río Cañete, 4 km from Laraos on the trail to [Cerro]

Quichao, 3500 m [12°20'39" S 075°48'47" W]. Holotype FML 1125a (Pl. 1 fig. 13). Bulimulidae.

scotophilus, Bostryx (Bostryx) W

EYRAUCH

, 1967a: 353, pl. 1 fig. 10. Type locality: Peru, [Dept. Lima] Río

Chillón, Puente El Diablo, 2300 m [? 12°30'00" S 075°54'45" W]. Holotype FML 1130 (Pl. 1 fig. 15).

Bulimulidae.

semiclausa, Epiphragmophora diluta W

EYRAUCH

, 1960a: 45, pl. 6 fig. 36 (15). Type locality: Peru, [Dept.

Cuzco] 150 km SE Cuzco, Sicuani [14°16'10" S 071°13'34" W, 3560 m], 3200 m? Holotype SMF

162040.

shutcoensis, Temesa (Temesa) pilsbryi W

EYRAUCH

, 1960c: 119, pl. 11 fig. 6. Type locality: Peru, [Dept. Lima]

above Laraos, left margin of Río Shutco, 3700 m. Holotype SMF 162114.

silvaevagus, Naesiotus W

EYRAUCH

, 1960a: 36, pl. 4 fig. 20. Type locality: Peru, [Dept. Junín] Río Chanchamayo

valley, [Cerro Pichita, 11°06'22" S 075°26'30" W] 2 km of Pichita Caluga Mine (1), 2200 m. Holotype

SMF 162001. Bulimulidae.

silvaevagus, Obeliscus (Microbeliscus) W

EYRAUCH

, 1964a: 40, fig. 2. Type locality: Peru, [Dept. Huánuco] right

margin of Río Monzón, near Tingo Maria, Cueva de las Lechuzas, 670 m. Holotype FML 1219.

similis, Littoridina W

EYRAUCH

, 1963a: 256, [pl. 1] figs 13-14. Type locality: Peru, [Dept. Puno] E Huancane,

0.5 km from Lago Titicaca, Vilque Chico, 3850 m. Holotype FML 1002a.

souzalopesi, Drymaeus (Drymaeus) W

EYRAUCH

, 1965a: 74, figs 4-5. Type locality: Brazil, Edo. Goiás,

Planaltina Nova (16). Holotype FML 10622a (Pl. 4 figs 43-45). Bulimulidae.

subterranea, Columbinia (Pfeifferiella) W

EYRAUCH

, 1957: 6, pl. 1 fig. 3. Type locality: Peru, [Dept. Cajamarca]

85 km NE Chiclayo, near Llama [06°30'52" S 079°07'13" W], Cerro Chunrún, 2350 m. Holotype SMF

155718.

superba, Peruinia flachi W

EYRAUCH

, 1960a: 24, pl. 3 fig. 1-2. Type locality: Peru, [Dept. Junín] Río

Chanchamayo valley, [Cerro Pichita, 11°06'22" S 075°26'30" W] 2 km of Pichita Caluga Mine (1),

2200 m. Holotype SMF 156235.

superbus, Bostryx (Multifasciatus) W

EYRAUCH

, 1967a: 365, pl. 1 figs 15-19. Type locality: Peru, [Dept. Lima]

left margin of Mayo Creek [Río Laraos], tributary to Río Cañete, 4 km from Laraos on the trail to [Cerro]

Quichao, 3500 m [12°20'39" S 075°48'47" W]. Holotype FML 1137a (Pl. 1 fig. 12). Bulimulidae.

tarmensis, Naesiotus (Reclasta) W

EYRAUCH

, 1967b: 479, pl. 4 figs 51-52. Type locality: Peru, [Dept. Junín]

valley Río Tarma, Cerro Huayuncayo[c] near La Florida [11°22'13" S 075°40'30" W], 3000-3100 m.

Holotype SMF 162017. Bulimulidae.

tenuissimus, Plekocheilus (Orcesiellus) W

EYRAUCH

, 1967b: 469, pl. 2 fig. 23, pl. 4 fig. 50. Type locality:

Ecuador, [Prov. Pichincha] 20 km W Quito, eastern slope of Cerro Pichincha, Tandayapa [00°01' S

078°46' W], 2500 m. Holotype FML 3364 (17). Amphibulimidae.

terrestris, Steeriana (Cylindronenia) maranhonensis W

EYRAUCH

, 1964a: 44, figs 4-5. Type locality: Peru, [Dept.

Amazonas] near Santa Rosa, Quebrada Aguaturbia [05°47'42" S 078°19'43" W], 1500 m. Holotype FML

702a.

thomei, Bulimulus (Rhinus) W

EYRAUCH

, 1967b: 481, pl. 1 figs 3-5. Type locality: Brazil, Edo. Rio Grande do

Sul, [Santana de] Livramento [30°53' S 055°31' W, 190 m], Experimental Fruit Station. Holotype MRCN

1021a. Simpulopsidae.

thomei, Radiodiscus W

EYRAUCH

, 1965b: 121, pl. 7 fig. 1. Type locality: Brazil, [Edo.] Rio Grande do Sul,

Municipio Cambará, Itaimbèzinho, 900 m. Holotype MRCN 1073a.

Folia conchyliologica, n°17, Mars 2012

translucidus, Drymaeus (Drymaeus) W

EYRAUCH

, 1967a: 422, pl. 6 figs 78-79. Type locality: Peru, [Dept.

Huánuco] road Tingo Maria-Pucallpa, Boquerón de Abad [08°49'60" S 075°55'00" W], 500 m. Holotype

FML 1202a (Pl. 3 figs 37-39). Bulimulidae.

tridentata, Zilchistrophia W

EYRAUCH

, 1960a: 27, pl. 3 fig. 6. Type locality: Peru, [Dept. Junín] Chanchamayo

valley, Pichita Caluga (1) [~2200 m]. Holotype SMF 162006.

turritus, Naesiotus (Raphiellus) [sic, Rhaphiellus] W

EYRAUCH

, 1967a: 412, pl. 5 figs 64-64a. Type locality: Peru,

[Dept. Cajamarca] Marañon river basin, valley of Río Yanguate, 5 km NE Celendín [06°49'53" S

078°06'40" W, 3000 m], 2250 m. Holotype FML 1229a (Pl. 2 fig. 16). Bulimulidae.

unilamellatus, Ptychodon (Unilamellatus) W

EYRAUCH

, 1965b: 134, pl. 7 fig. 4. Type locality: Peru, [Dept.

Junin] road Tarma-Jauja, above Tarmatambo, 3500 m. Holotype FML 3863a.

vilchezi, Bostryx (Bostryx) W

EYRAUCH

, 1960a: 32, pl. 3 figs 8-10. Type locality: Peru, Dept. Cajamarca, Río

Guineamaya, 20 km NE Cutervo (1950 m), Sócota [06°18'59" S 078°44'37" W, 1820 m]. Holotype SMF

155704. Bulimulidae.

willinki, Bostryx (Bostryx) W

EYRAUCH

, 1964a: 54, fig. 12. Type locality: Argentina, Prov. Catamarca, Quebrada

de Tinogasta [28°02'39" S 067°33'45" W, 1210 m]. Holotype FML 121a (Pl. 1 fig. 7). Bulimulidae.

woytowskii, Leptariota W

EYRAUCH

, 1960c: 130, pl. 11 fig. 9. Type locality: Peru, [Dept. Cajamarca] near Jaën

[05°42'29" S 078°48'17" W], 700 m. Holotype SMF 164000.

wygodzinskyi, Radiodiscus W

EYRAUCH

, 1965c: 110, fig. 3. Type locality: Argentina, Prov. Tucumán, road

Tucumán-Tafí del Valle, valley Río Los Sosa, 900 m. Holotype: FML 4148a.

zilchi, Bostryx (Bostryx) W

EYRAUCH

, 1958: 108, pl. 9 figs 41-42. Type locality: Peru, [Dept. Lima,] 5 km from

Laraos on footpath to Yauyos, [Cerro] Quichao [12°20'45" S 075°48'53" W], 3600 m (18). Holotype SMF

156348. Bulimulidae.

zilchi, Epiphragmophora W

EYRAUCH

, 1960a: 44, pl. 6 figs 41-44. Type locality: Peru, [Dept. Arequipa] km 651

Panamericana Lima-Arequipa (220-500 m), E Chala, Lomas de Capac [15°52' S 074°09' W]. Holotype

SMF 156391.

zilchi, Naesiotus W

EYRAUCH

, 1956b: 9, pl. 1 fig. 6. Type locality: Peru, [Dept. Cajamarca] Río Jequetepeque

valley, Quinden, 600 m (10). Holotype FML 1908 (Pl. 2 figs 21-22). Bulimulidae.

zilchi, Scutalus (Vermiculatus) culmineus W

EYRAUCH

, 1967a: 393, pl. 9 figs 139-140. Type locality: Peru, [Dept.

Lima] Río Cañete basin, right margin of Río Alis, near Alis [12°16'43" S 075°47'13" W, 3275 m].

Holotype FML 3328a. Bulimulidae.

zilchi, Stenostylus W

EYRAUCH

, 1956c: 156, pl. 11 fig. 8. Type locality: Peru, [Dept. Ancash] 280 km N Lima,

road Barrancas-Huaráz, Tapacocha [10°01' S 077°35' W], 3250 m. Holotype SMF 155314. Bulimulidae.

zilchi, Systrophia (Systrophia) W

EYRAUCH

, 1967a: 431, pl. 6 fig. 81. Type locality: Peru, [Dept. Arequipa] near

Chala, Lomas de Atiquipa [15°47'46" S 074°21'49" W], 900 m. Holotype SMF 162007.

zilchi, Temesa (Temesa) W

EYRAUCH

, 1963b: 285, pl. 1 fig. 14. Type locality: Peru, [Dept. Lima] right margin of

Río Alis, near Alis [12°16'43" S 075°47'13" W], 3300 m. Holotype FML 1053.

zischkai, Nenia (Columbinia) W

EYRAUCH

, 1956a: 113, pl. 6 figs 3-3a. Type locality: Eastern Bolivia, [Dept.

Cochabamba,] Yungas de[l] Palmar [17°08' S 065°30' W], 700 m. Holotype FML 1368.

Taxa based on the Weyrauch collection

acobambensis, Nenia P

ILSBRY

, 1945: 80, pl. 3 figs 3-4. Type locality: Peru, [Dept. Junín] near Tarma, Acobamba,

3000-3400 m. Holotype ANSP 180032.

altorum, Bulimulus (Scutalus) revinctus H

AAS

, 1951: 516, fig. 104. Type locality: Peru, ‘puna‘ between

Andahuaylas and Abancay, 4000 m. Holotype FMNH 30912. Bulimulidae.

andium, Radiodiscus P

ILSBRY

, 1944c: 30, figs 11-11a. Type locality: Peru, [Dept. Junín], near Tarma, Chancha,

3400 m. Holotype ANSP 180007.

angusta, Euglandina (Euglandina) cylindrus H

AAS

, 1951: 539, fig. 122. Type locality: Peru, [ Dept. Cajamarca]

Jaën, 700 m. Holotype FMNH 30911.

atahualpa, Epiphragmophora P

ILSBRY

, 1944b: 119, pl. 11 figs 4-5. Type locality: Peru, Andahuaylas, 3100 m.

Holotype ANSP 180206.

Folia conchyliologica, n°17, Mars 2012

bicolor, Temesa P

ILSBRY

, 1949: 4, figs 1, 8-8a. Type locality: Peru, [Dept. Junín] (near Tarma), Tarmatambo,

3400 m. Holotype ANSP 183793.

boettgeri, Nenia quadrata P

ILSBRY

, 1945: 81, pl. 3 figs 1-2. Type locality: Peru, [Dept. Junín] Rio Tarma,

Carpapata, 2200-2300 m. Holotype ANSP 180035.

bradina, Nenia flachi P

ILSBRY

, 1945: 84, pl. 3 fig. 6. Type locality: Peru, Oxapampa, 1600 m. Holotype ANSP

17999.

cereonitens, Systrophia (Systrophia) H

AAS

, 1951: 535, fig. 117. Type locality: Peru, west of Río Chinchipe, S of

San Ignacio, 1200 m. Holotype FMNH 30924.

cunjacayensis, Bostryx B

REURE

& N

EUBERT

, 2008: 323, fig. 6. Type locality: Peru, Dept. Ancash, Río Fortaleza,

Caracayoc near Cajacay (10°09'23" S 077°26'38" W, 2650 m). Holotype SMF 156395. Bulimulidae.

edmundi, Bostryx B

REURE

& N

EUBERT

, 2008: 324, figs 11-16. Type locality: Peru, Dept. Lima, Río Cañete

valley, Hacienda Yacca (12°32'50" S 075°55'02" W, 2110 m). Holotype SMF 331475 (19). Bulimulidae.

eka, Nenia P

ILSBRY

, 1945: 82, pl. 3 figs 7-7a. Type locality: Peru, [Dept. Junín] near Tarma, Acobamba, 3000 m.

Holotype ANSP 180034.

eusteirus, Drymaeus P

ILSBRY

, 1944c: 29, pl. 1 fig. 7. Type locality: Peru, Oxapampa, 1800 m. Holotype ANSP

180674. Bulimulidae.

flammulata, Andiniella L

OOSJES

, 1957: 229, figs 1-2. Type locality: Peru, [Dept. Junín] Cerro Huilcashpata near

Palta (Rio Tarma), 3100 m. Holotype FML 1310.

granulatus, Bostryx B

REURE

& N

EUBERT

, 2008: 325, fis 17-19. Type locality: Peru, Dept. Lima, Río Cañete

valley, Yauyos, 3100 m. Holotype SMF 162178. Bulimulidae.

harangoensis, Steeriana (Steeriana) Zilch, 1949: 94, pl. 6 fig. 24. Type locality: Peru, east of Río Chichipe,

between Santa Rosa and San José de Lourdes, Huarango, 1400 m. Holotype SMF 62565a.

hirsutus, Bostryx B

REURE

, 1978: 82, figs 125-127, pl. 12 fig. 1. Type locality: Peru, Dept. Cajamarca, Rio

Jequetepeque, near Chilete, Tambo Tingo, 1000 m. Holotype FML 6072 (Pl. 1 fig. 6). Bulimulidae.

huarazensis, Bostryx P

ILSBRY

, 1944b: 121, pl. 11 fig. 17. Type locality: Peru, [Dept. Ancash], Santa valley,

Huaráz, 3100 m. Holotype ANSP 180000. Bulimulidae.

incarum, Megalobulimus (Microborus) P

ILSBRY

, 1944c: 29, pl. 1 figs 8-9. Type locality: Peru, [Dept. Ancash]

Huaráz, 3000-3200 m. Holotype ANSP 180677.

inornata, Solaropsis (Psadara) H

AAS

, 1951: 527, fig. 113. Type locality: Peru, Río Chusgon, Hacienda Santa

Elena, 1600-2150 m. Holotype FMNH 30929.

leucobasis, Epiphragmophora (Epiphragmophora) H

AAS

, 1951: 524, fig. 111. Type locality: Peru, near Río

Pampas, above Chincheros, 3000 m. Holotype FMNH 30923.

megomphalus, Bostryx P

ILSBRY

, 1944b: 122, pl. 11 figs 15-16. Type locality: Peru, [Dept. Junín] (near Tarma)

Acobamba, 3200-3400 m. Holotype ANSP 180036. Bulimulidae.

microhelix, Radiodiscus (Radiodiscus) H

AAS

, 1951: 538, fig. 121. Type locality: Peru, Chanchamayo valley,

800 m. Holotype FMNH 30921.

minuscula, Nenia P

ILSBRY

, 1945: 83, pl. 3 figs 9-10. Type locality: Peru, [Dept. Junín] near Tarma, Acobamba,

3000 m. Holotype ANSP 180033.

minuta, Chilina H

AAS

, 1951: 542, fig. 125. Type locality: Chile, near Santiago, Baños Morales. Holotype FMNH

30941.

multilineatus, Bostryx B

REURE

, 1978: 104, pl. 7 figs 11-13. Type locality: Peru, Dept. Ancash, below Colca,

2100 m. Holotype 3080a (Pl. 1 fig. 14). Bulimulidae.

munda, Helicina (Helicina) H

AAS

, 1951: 543, fig. 126. Type locality: Peru, near Huánuco, Ayancocha, 2200 m.

Holotype FMNH 30937.

nanus, Obeliscus (Protobeliscus) H

AAS

, 1951: 541, fig. 123. Type locality: Peru, Chanchamayo valley, 800 m.

Holotype FMNH 30931.

obeliscus, Bostryx (Peronaeus) Z

ILCH

, 1954: 78, pl. 6 figs 12-16. Type locality: Peru, [Dept. Lima], near

Autisha, Rio Santa Eulalia, 2100-2200 m. Holotype SMF 136762. Bulimulidae.

Folia conchyliologica, n°17, Mars 2012

palassus, Drymaeus (Drymaeus) B

REURE

& E

SKENS

, 1981: 32. Type locality: Peru, Dept. Cajamarca, 30 km NE

Cutervo, 2650 m. Holotype SMF 156377a. Bulimulidae.

perforatus, Bulimulus (Ataxus) H

AAS

, 1951: 518, fig. 106. Type locality: Peru, Ninabamba on the Pampas river,

2000 m. Holotype FMNH 30906. Bulimulidae.

peruviana, Solaropsis (Psadara) monile H

AAS

, 1951: 528, fig. 114. Type locality: Peru, Urubamba valley,

Sahuayaco, 800 m. Holotype FMNH 30928.

pizarro, Psadara P

ILSBRY

, 1944b: 119, pl. 11 figs 6-6a. Type locality: Peru, near Huancapistana, Yanago,

1800 m. Holotype ANSP 180355.

planior, Happia (Happia) cuzcana H

AAS

, 1951: 537, fig. 120. Type locality: Peru, Chanchamayo, Oreja de

Capelo, 1600 m. Holotype FMNH 30933.

platysma, Systrophia (Systrophia) H

AAS

, 1951: 530, fig. 115. Type locality: Peru, close to Río Marañon, east side

of Río Chinchipe, Huarango, 1400 m. Holotype FMNH 30926.

pseudelatus, Drymaeus (Drymaeus) H

AAS

, 1951: 520, fig. 108. Type locality: Peru, Mantaro river, Anco,

2500 m. Holotype FMNH 30908. Bulimulidae.

pumilus, Bostryx spiculatus B

REURE

, 1978: 126, figs 187-190. Type locality: Peru, Dept. Cuzco, near Calca on

the left side of Rio Vilcanota (ca. 2900 m). Holotype FML 1707a (2). Bulimulidae.

punctilineatus, Bulimulus (Scutalus) H

AAS

, 1951: 517, fig. 105. Type locality: Peru, Urubamba valley,

Sahuayaco, 800 m. Holotype FMNH 30914. Bulimulidae.

rhabdotus, Bulimus (Protoglyptus) H

AAS

, 1951: 512, fig. 100. Type locality: Peru, near Huánuco, Ambo,

2000 m. Holotype FMNH 30915. Bulimulidae.

sagasteguii, Kionoptyx H

AAS

, 1966: 239, figs 57a-b. Type locality: Peru, Dept. La Libertad, Prov. Huamachuco,

Marcabal (07°37 S, 77°49 W). Holotype FMNH 131682. Bulimulidae.

satipoensis, Thaumastus robertsi P

ILSBRY

1944b: 121, pl. 11 fig. 1. Type locality: Peru, near Huancayo, Satipo,

600 m. Holotype ANSP 179990. ?Orthalicidae.

subcostatus, Bulimulus (Protoglyptus) H

AAS

, 1948: 190, fig. 39. Type locality: Peru, Dept. Cajamarca, Jaën,

1500-2100 feet [~ 460-640 m]. Holotype FMNH 29148. Bulimulidae.

subelatus, Bulimulus (Peronaeus) H

AAS

, 1948: 189, fig. 38. Type locality: Peru, Dept. Huánuco, Ambo near

Huánuco, 6300 feet [~1920 m]. Holotype FMNH 29146. Bulimulidae.

urubambensis, Nenia angrandi P

ILSBRY

, 1945: 83, pl. 3 fig. 8. Type locality: Peru, [Dept. Cuzco] (head of the

Urubamba valley), Vilcanota Valley, Huadqueña, 1500 m. Holotype ANSP 180029.

virgula, Bulimulus (Peronaeus) H

AAS

, 1951: 514, fig. 102. Type locality: Peru, Ninabamba on the Pampas River,

2700 m. Holotype FMNH 30917. Bulimulidae.

viridulum, Aperostoma (Aperostoma) H

AAS

, 1952: 112, fig. 16. Type locality: Peru, [Dept. Cuzco] Macchu Pichu

near Cuzco, on Rio Urubamba, 2100 m. Holotype FMNH 38379.

weyrauchi, Austroselenites (?) H

AAS

, 1951: 535, fig. 119. Type locality: Peru, Cuzco, 3500 m. Holotype FMNH

30939.

weyrauchi, Bostryx P

ILSBRY

, 1944a: 87, pl. 9 fig. 5. Peru, [Dept. Ayacucho] Ninabamba near Ayacucho, 2000 m.

Holotype ANSP 179979. Bulimulidae.

weyrauchi, Calaperostoma H

AAS

, 1952: 115, fig. 18. Type locality: Peru, Oxapampa, 1700 m. Holotype FMNH

38377.

weyrauchi, Helicina (Helicina) H

AAS

, 1948: 192, fig. 40. Type locality: Peru, Dept. Cajamarca, Jaën. Holotype

FMNH 29142.

weyrauchi, Nenia P

ILSBRY

, 1945: 82, pl. 3 fig. 5. Type locality: Peru, [Dept. Ayacucho] Ninabamba near

Ayacucho, 1900 m. Holotype ANSP 179978.

weyrauchi, Neopetraeus P

ILSBRY

, 1944a: 88, pl. 9 fig. 4. Type locality: Peru, [Dept. Ancash], Santa valley,

Huaráz, 3200 m. Holotype ANSP 179980. Bulimulidae.

weyrauchi, Plagiodontes P

IZÁ

& C

AZZANIGA

, 2009: 1443, figs 4-12. Type locality: Argentina, Prov. Catamarca,

Cuesta de la Chilca, 22 km E Andalgalá (27°38 19 S, 66°10 27 W), 1455 m. Holotype MACN-In

37465. Odontostomidae.

Folia conchyliologica, n°17, Mars 2012

weyrauch [sic, weyrauchi], Thaumastus (Scholvienia) P

ILSBRY

1944b: 121, pl. 11 figs 2-2a. Type locality: Peru,

[Dept. Junín] Rio Tarma, near Palca, Carpapata, 2300 m. Holotype ANSP 179996. Orthalicidae.

willinki, Naesiotus B

REURE

, 1978: 162, figs 262-269, pl. 12 fig. 2. Type locality: Argentina, Prov. Salta, Estancia

Lumbrera, 750 m. Holotype FML 1263a (2). Bulimulidae.

Notes

(1) Exact locality not known, but Cerro Pichita is located at 11°06'22" S 075°26'30" W; the mine is possibly

about 7.5 km W San Ramón (as the crow flies).

(2) Photograph failed.

(3) Given the elevation, the location is supposed to be at the coordinates presented here; the locality is

12.4 km (as the crow flies) NNE Tingo Maria.

(4) This is probably meant as ca. 25 km NE Cutervo by road; cf. W

EYRAUCH

1976a: 348.

(5) This locality may be around 10°33' S 076°15' W.

(6) Given the elevation, Hacienda Cochambul may be around 07°12' S 078°23' W.

(7) There is no such named locality in the GNS gazetteer. Given the altitude estimated, it could be around

06°18' S 078°58' W.

(8) There is no such named locality in the GNS gazetteer; however in the region indicated, Curicancha as a

locality is known. Given the altitude estimated, it could be around 11°39' S 075°52' W.

(9) Given the elevation, this locality is probably around 11°42'34" S 076°35'30" W.

(10) The reference by Weyrauch as “16 km N Tembladera” seems erroneous, as there is no place called

Quinden at 600-700 m in this area. Following the road from Tembladera to Cajamarca, there is a

location Quinden in the Río Jetequepeque valley around 07°10' S 079°00' W at ca. 630 m; this is

likely the type locality.

(11) See D

ELHEY

et al., 2010 for details on the locality.

(12) At the indicated locality, elevations of 450-500 m prevail.

(13) Sotopampa and Huamboyacu do not appear in the GNS gazetteer. The places called (La) Colmena in

northern Peru do not agree with the altitude given by Weyrauch. There could have been a misspelling

for Sosopampa (Dept. Lambayeque) or Satapampa (twice in Dept. La Libertad), but none of the places

is near the given altitude. Hence the precise locality where this species was found remains unclear.

(14) Not Drymaeus pilsbry Zetek, 1934; renamed Mesembrinus (Ornatimormus) henrypilsbryi in

EYRAUCH

, 1958: 134.

(15) Not Epiphragmophora semiclausa Hylton Scott, 1951; renamed E. diluta semiaperta in W

EYRAUCH

1964a: 169.

(16) Places of this name have not been found in the gazetteer; the name could either refer to Edo. Goiás,

Planaltino de Goiás (15°29'04" S 047°38'20" W), or Distrito Federal, Planaltino (15°37'02" S

047°40'00" W). These places are only 15 km apart.

(17) Figured by B

ORRERO

& B

REURE

, 2011: fig. 5A-C.

(18) Corrected by W

EYRAUCH

, 1960c: 124, footnote 6.

(19) See B

REURE

& M

OGOLLÓN

VILA

2010: 19.

Acknowledgements

The author is highly indebted to Gabriela Cuezzo and Eugenia Salas-Oroño (Tucumán), for all support

during a visit to the FML in early 2010. Thanks are due to Jan van Tol (NCB, Leiden) and the Diederik van

Schagen Fund for financial support during this trip.

Due to difficult lighting circumstances during my visit, not all photographs were of the quality expected

and some had to be discarded.

References

ARBOSA

, A. F., D

ELHEY

, V.K. & C

OAN

, E.V., 2008. — Molluscan names and malacological contributions by

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ORRERO

, F.J. & B

REURE

, A.S.H., 2011. — The Amphibulimidae (Mollusca: Gastropoda: Orthalicoidea) from

Colombia and adjacent areas. Zootaxa, 3054: 1-59.

REURE

, A.S.H., 1978. — Notes on and descriptions of Bulimulidae (Mollusca, Gastropoda). Zoologische

Verhandelingen Leiden, 164: 1-255.

Folia conchyliologica, n°17, Mars 2012

REURE

, A.S.H., 2008. — Mysterious or confusing: enigmatic species in the Orthalicidae. Basteria, 72: 241-252.

REURE

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SKENS

, A.A.C., 1977. — Observations on the formation of spermatophores in a bulimulid

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REURE

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SKENS

, A.A.C., 1981. — Notes on and descriptions of Bulimulidae (Mollusca, Gastropoda),

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OGOLLÓN

VILA

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EUBERT

, E., 2008. — A preliminary revision of Weyrauch’s unpublished names: taxa of the

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OMERO

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ELHEY

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IZÁ

, J. & B

URELA

, S., 2010. — Zilchogyra franzi Weyrauch, 1965 (Gastropoda: Charopidae), a

minute landsnail rediscovered in Sierra de la Ventana (Southern Pampas, Argentina). Zootaxa, 2450: 61-64.

AAS

, F., 1947. — Malacological notes, V. Fieldiana Zoology, 31: 171-188.

AAS

, F. 1948. — Three new land shells from Peru. Fieldiana Zoology, 31: 189-193.

AAS

, F., 1951. — Remarks on and descriptions of South American non-marine shells. Fieldiana Zoology, 31:

503-545.

AAS

, F., 1952. — South American non-marine shells: further remarks and descriptions. Fieldiana Zoology, 34:

107-132.

AAS

, F., 1955. — On some small collections of inland shells from South America. Fieldiana Zoology, 34: 361-387.

AAS

, F., 1959. — Inland mollusks from Venezuela, southern Brazil, and Peru. Fieldiana Zoology, 39: 363-371.

AAS

, F., 1966. — On some new non-marine mollusks from Columbia and Peru. Fieldiana Zoology, 44: 231-241.

OOSJES

, F.E., 1957. — On a new Andinia (Gastropoda, Clausiliidae) from Peru. Zoologische Mededelingen

Leiden, 35: 229-234.

OOSJES

, F.E. & L

OOSJES

VAN

EMMEL

, A.C.W., 1966. — Some anatomical, systematical and geographical data

on Neniinae (Gastropoda, Clausiliidae). Zoologische Verhandelingen Leiden, 77: 1-59.

EUBERT

, E. & J

ANSSEN

, R., 2004. — Die Typen und Typoide des Natur-Museums Senckenberg, 84: Mollusca:

Gastropoda: Pulmonata: Orthalicoidea: Bulimulidae (2), Orthalicidae, Placostylidae. Archiv für Molluskenkunde,

133: 193-297.

ILSBRY

, H.A., 1944a. — New Peruvian land mollusks. The Nautilus, 57: 87-88.

ILSBRY

, H.A., 1944b. — New Peruvian land mollusks, II. The Nautilus, 57: 118-122.

ILSBRY

, H.A., 1944c. — New Peruvian land mollusks, III. The Nautilus, 58: 28-30.

ILSBRY

, H.A., 1945. — New Peruvian land mollusks, IV: Clausiliidae. The Nautilus, 58: 79-84.

ILSBRY

, H.A., 1949. — Review of Peruvian species of Temesa (Mollusca, Clausiliidae). Notulae Naturae, 214: 1-8.

Folia conchyliologica, n°17, Mars 2012

IZÁ

, J. & C

AZZANIGA

, N., 2009. — A new species of Plagiodontes from Argentina, and new data on the

anatomy of four other species in the genus (Gastropoda: Orthalicidae, Odontostominae). Journal of Natural

History, 43: 1437-1471.

ALAS

-O

ROÑO

, E., 2007. — Taxonomic review of the Spixia piriformis species complex (Gastropoda:

Pulmonata: Odontostominae). Zootaxa, 1498: 1-25.

EYRAUCH

, W.K., 1956a. — Two new species of Clausiliidae from Peru and Bolivia. The Nautilus, 69: 110-115.

EYRAUCH

, W.K., 1956b. — The genus Naesiotus, with descriptions of new species and notes on other Peruvian

Bulimulidae. Proceedings of the Academy of Natural Sciences of Philadelphia, 108: 1-17.

EYRAUCH

, W.K., 1956c. — Neue Landschnecken aus Peru. Archiv für Molluskenkunde, 85: 145-162.

EYRAUCH

, W.K., 1957. — Sieben neue Clausiliiden aus Peru. Archiv für Molluskenkunde, 86: 1-28.

EYRAUCH

, W.K., 1958. — Neue Landschnecken und Neue Synonyme aus Südamerika, 1. Archiv für

Molluskenkunde, 87: 91-140.

EYRAUCH

, W.K., 1960a. — Zwanzig neue Landschnecken aus Peru. Archiv für Molluskenkunde, 89: 23-48.

EYRAUCH

, W.K., 1960b. — Zur Kenntnis von Newboldius (Bulimulidae). Archiv für Molluskenkunde, 89: 49-56.

EYRAUCH

, W.K., 1960c. — Siebzehn neue Landschnecken aus Peru. Archiv für Molluskenkunde, 85: 117-132.

EYRAUCH

, W.K., 1963a. — Cuatro neuva especies de Hydrobiidae de Argentina y Perú (Gastropoda,

Prosobranchia). Acta Zoologica Lilloana, 19: 243-259.

EYRAUCH

, W.K., 1963b. — Aporte al conocimento de Temesa, I (Clausiliidae, Mollusca).— Acta Zoologica

Lilloana, 19: 261-288.

EYRAUCH

, W.K., 1964a. — Nuevos gastrópodos terrestres y nuevos sinónimos de Sudamérica, II. Acta

Zoologica Lilloana, 20: 33-60.

EYRAUCH

, W.K., 1964b. — Aporte al conocimento de Temesa, II (Clausiliidae, Mollusca).— Acta Zoologica

Lilloana, 20: 145-162.

EYRAUCH

, W.K., 1965a. — Tres nuevos gastrópodos terrestres de Sudamérica. Neotropica, 11: 71-76.

EYRAUCH

, W.K., 1965b. — Neue und verkannte Endodontiden aus Südamerika. Archiv für Molluskenkunde,

94: 121-134.

EYRAUCH

, W.K., 1965c. — Cinco nuevos Endodontidos de Argentina y Perú (Gastropoda, Euthyneura).

Neotropica, 11: 105-115.

EYRAUCH

, W.K., 1966. — Gastrópodos terrestres de Argentina, Uruguay y Brasil. Neotropica, 12: 41-47.

EYRAUCH

, W.K., 1967a. — Treinta y ocho nuevos gastrópodos terrestres de Perú. Acta Zoologica Lilloana, 21:

343-455.

EYRAUCH

, W.K., 1967b. — Descripciónes y notas sobre gastrópodos terrestres de Venezuela, Colombia,

Ecuador, Brasil y Perú. Acta Zoologica Lilloana, 21: 457-499.

ILCH

, A., 1949. — Studien Paul Ehrmann’s über asiatische und südamerikanische Clausiliiden. Archiv für

Molluskenkunde, 78: 69-98.

ILCH

, A., 1954. — Landschnecken aus Peru, 2. Archiv für Molluskenkunde, 83: 65-78.

Folia conchyliologica, n°17, Mars 2012

Figures 1-15. Bostryx (sensu lato) species. 1, Bostryx (Lissoacme) birabenorum

Weyrauch, 1965, holotype

FML 958a (H = 18.0). 2, Bostryx (Lissoacme) globosus Weyrauch, 1967, holotype FML 1226 (H = 19.2). 3,

Bostryx (B.) ortizi

Weyrauch, 1967, holotype FML 1126 (H = 17.7). 4-5, Floreziellus florezi

Weyrauch,

1967, holotype FML 10671a (H = 26.8). 6, Bostryx hirsutus Breure, 1978,

holotype FML 6072 (H = 11.8).

7, Bostryx (B.) willinki Weyrauch, 1964, holotype FML 121a (H = 18.5). 8-9,

Phenacotaxus (Ataxellus)

florezi

Weyrauch, 1967, holotype FML 10522a (H = 17.7). 10-11, Bostryx (Scansicohlea) gracilis

Weyrauch, 1967, holotype FML 1102 (H = 11.5). 12, Bostryx (Multifasciatus) superbus

Weyrauch, 1967,

holotype FML 1137a (H = 33.3). 13, Bostryx (B.) rodriguezae Weyrauch, 1967, holoty

pe FML 1125a (H =

27.5). 14, Bostryx multilineatus Breure, 1978, holotype FML (H = 29.5). 15, Bostryx (B.) scotophilus

Weyrauch, 1967, holotype FML 1130 (H = 18.1). All figures x2.

Plate 1

Folia conchyliologica, n°17, Mars 2012

Plate 2

Figures 16-20. Naesiotus (sensu lato) species. 16, Naesiotus (Rhaphiellus) turritus

Weyrauch, 1967,

holotype FML 1229a (H = 10.6). 17-18, Naesiotus (N.) bicolor

Weyrauch, 1967, holotype FML 3987 (H =

9.2). 19-20, Naesiotus (Naesiotellus) latecolumellaris Weyrauch, 1967, holotype FML 1076 (H = 14.6). 21

22, Naesiotus zilchi Weyrauch, 1956, holotype FML 1908 (H = 12.6). All figures x2.

Figures 23-28. Neopetraeus species. 23-25, Neopetraeus arboriferus obesus

Weyrauch, 1967, holotype

FML 1244a (H = 43.5). 26-28, Neopetraeus arboriferus paucistrigatus

Weyrauch, 1967, holotype FML

1239a (H = 41.4). All figures x1.5.

Folia conchyliologica, n°17, Mars 2012

Plate 3

Figures 29-31. Neopetraeus camachoi Weyrauch, 1967, holotype FML 1240a (H = 35.3).

Figures 32-39. Drymaeus species. 32-34, Drymaeus (Ornatimormus) multiguttatus

Weyrauch, 1964,

holotype FML 1200 (H = 19.8). 35-36, Drymaeus (Diaphanomormus) coelestini obesus

Weyrauch, 1964,

holotype FML 1196a (H = 11.3). 37-39, Drymaeus (D.) translucidus

Weyrauch, 1967, holotype FML 1202a

(H = 24.7) and paratype. All figures x1.5.

Plate 3

Folia conchyliologica, n°17, Mars 2012

Plate 4

Figures 40-45. Drymaeus species. 40-42, Drymaeus (Mormus) expansus flavilabrum

Weyrauch, 1967,

holotype FML 1197 (H = 42.7). 43-45, Drymaeus (D.) souzalopezi

Weyrauch, 1965, holotype FML 10622a

(H = 40.0). All figures x1.5.

Folia conchyliologica, n°17, Mars 2012

Plate 5

Figures 46-51. Scutalus species. 46-47, Scutalus (S.) [c.] chiletensis

Weyrauch, 1967, holotype FML 1236a

(H = 33.1). 48, Scutalus (S.) chiletensis granulatus Weyrauch, 1967, holotype FML 1362a (H = 38.0). 49-

51, Scutalus coraeformis debilisculptus Weyrauch, 1967, holotype FML 1984 (H = 35.6). All figures x1.5.

Figures 52-54. Scutalus (S.) ortizpuenti Weyrauch, 1967, holotype FML 1234 (H = 48.3). Actual size.

Folia conchyliologica, n°17, Mars 2012

Plate 6

Figures 55-58. Kuschelenia (Vermiculatus) species. 55-56, Scutalus (Vermiculatus) [c.] cuzcoensis

Weyrauch, 1967, holotype FML 1225 (H = 35.3). 57, Scutalus (Vermiculatus) cuzcoensis marasensis

Weyrauch, 1967, holotype FML 10641a (H = 34.4). 58, Scutalus (Vermiculatus) culmineus zilchi

Weyrauch,

1967, holotype FML 3328a (H = 29.9). All figures x1.5.

Figures 59-61. Thaumastus (T.) orcesi Weyrauch, 1967, holotype FML 3165 (H = 49.4). All figures x1.5.

Salvation and documentation: additional (probable) type material of South American land-snail species (Gastropoda, Stylommatophora) in the Museum für Naturkunde Berlin

Article

Full-text available

Mar 2024
J CONCHOL

Type material of some South American species in the Museum für Naturkunde Berlin, belonging to different families is documented. It includes 16 species: Bulimus cuneus L. Pfeiffer, 1854, Bulimus proteus Broderip, 1832, Bulimus scalarioides L. Pfeiffer, 1867, Bulimus similaris J. Moricand, 1856, Andinia (Ehrmanniella) dedicata Weyrauch & Zilch, 1954, Helix hettneriana E. von Martens, 1897, Eurycampta hidalgonis Döring, 1877, Helix aequatoris L. Pfeiffer, 1860, Helix bituberculata L. Pfeiffer, 1853, Helix bourcieri L. Pfeiffer, 1853, Helix neogranadensis L. Pfeiffer, 1845, Cyclostoma (Cyclophorus) bourcieri L. Pfeiffer, 1854, and Helix platygyra Albers, 1857. The concept of the “salvation” of type material is explained.

South American terrestrial Gastropoda in the collection of the Auckland War Memorial Museum

Article

Full-text available

Mar 2023

The catalogued collection of South American terrestrial gastropods, including the Caribbean ABC islands and Trinidad & Tobago, of the Auckland War Memorial Museum (AM), New Zealand, is discussed here. In total, 264 specimen lots from South America were found in the AM collection, including eight type lots. Twelve of the specimens represent new geographic distribution data, including the first known precise locality for the species Bostryx luridus (L. Pfeiffer, 1863) and potentially for Drymaeus cf. waldoschmidti Parodiz, 1962 as well. The specimens of Lopesianus crenulatus Weyrauch, 1967 allowed for the revision of the monospecific genus Lopesianus Weyrauch, 1958, which is herein considered valid in the family Simpulopsidae.

On the rare Neotropical genus Ophionthus Bernhauer (Coleoptera: Staphylinidae: Staphylininae): redescription of the type species and description of a new species

Article

Full-text available

Feb 2021

Two female specimens of the Neotropical genus Ophionthus Bernhauer were found during museum collection surveys, representing different species of the genus. These species are Ophionthus serpentinus Bernhauer, 1908 and O. asenjoi sp. nov. The genus Ophionthus was monotypic until the present study and only known from a single male specimen from Central Peru. The genus is redescribed with the inclusion of female characters from both the type species and the new species here proposed. Additionally, the systematic position of Ophionthus within the Neotropical lineage of Philonthina and its geographical distribution in Peru are reassessed. Descriptions, diagnoses, illustrations, pictures, a cladogram, and a distribution map are provided.

Revision of the genera of Scolodontidae, part 3: Entodina Ancey, 1887, Keranella gen. nov., Martinella Jousseaume, 1887, †Patagocharopa Miquel & P.E. Rodriguez, 2016, Polygyratia Gray, 1847, Ridleyconcha Christensen, 2020, Smenodonta gen. nov., Systrophia L. Pfeiffer, 1855, and Zilchistrophia Weyrauch, 1960

Article

Full-text available

Feb 2025

With this study, we continue the revision of the genera of Scolodontidae (Gastropoda: Stylommatophora), focusing on the taxa with tightly coiled, multiwhorled shells that often bear apertural and/or internal barriers. We analysed type material from all type species and studied some misclassified Charopidae and Streptaxidae. We provide redescriptions and better circumscription of the following Scolodontid taxa: Entodina Ancey, 1887, Systrophia L. Pfeiffer, 1855, and Zilchistrophia Weyrauch, 1960. Martinella Jousseaume, 1887 is transferred from Streptaxidae to Scolodontidae, and the extinct Patagocharopa Miquel & P.E. Rodriguez, 2016 is transferred from Charopidae; Patagocharopa, dating from the Miocene, represents the earliest fossil record of the family. Keranella gen. nov. and Smenodonta gen. nov. are erected. The genera Polygyratia J.E. Gray, 1847 and Ridleyconcha Christensen, 2020, despite recently revised, are also included here for comparison with the other taxa.

From depressed to detached: extreme shell shape variation in some Peruvian Bostryx species (Gastropoda: Bulimulidae)

Article

Full-text available

Apr 2024

This paper investigates Bostryx species from Peru, which show variation in shells shape and are partly carinated, using both ecological and molecular data. The study area is situated in a valley at the western side of the Andes, with a total length of 25 km and an altitudinal range of almost 1,140 m, but focuses on two localities: El Infernillo at ca. 3,350 m and Tambo de Viso at ca. 2,700 m. The morphological variation of the species presents a rather complex picture that cannot completely be resolved by ecological data alone. Molecular data points to three taxa, two of which are variable in shell shape and each restricted to one of the localities mentioned; the third taxon is constant in shell shape and occurs over a large range of the study area. The species originally described as Bostryx multiconspectus Breure, 2008 and B. primigenius Breure, 2008 are redefined as a result. Résumé: Cet article traite des especes de Bostryx du Peŕou qui preśentent des variations dans la forme de leur coquille et sont partiellement careńeés, en utilisant à la fois des donneés ećologiques et molećulaires. La zone d'etude est situeé dans une valleé du côté ouest des Andes, avec une longueur totale de 25 km et une etendue altitudinale de pres de 1 140 m, mais se concentre sur deux localiteś : El Infernillo à env. 3 350 m et Tambo de Viso à env. 2 700 m. Les variations morphologiques des especes preśentent une evolution assez complexe que les seules donneés ećologiques ne suf isent pas à expliquer. Les donneés molećulaires indiquent trois taxons, dont deux ont une forme de coquille variable et chacune est limiteé à l'une des localiteś mentionneés ; le troisieme taxon est constant dans la forme de la coquille et est preśent sur une large etendue de la zone d'etude. Les especes initialement dećrites comme Bostryx multiconspectus Breure, 2008 et B. primigenius Breure, 2008 sont en consequence redé inies.

Land and freshwater molluscs of mainland Ecuador: an illustrated checklist

Article

Full-text available

Feb 2022

This comprehensive checklist of non-marine molluscs from the mainland of Ecuador pre- sents data on 331 species (of which 5 are considered as nomen dubium), compiled from the literature, a subset of museum databases, and some verified observations from the internet. For all taxa the original reference is given and photographs of type specimens are provided if these could be located and obtained from museums. Where these were not available the original figure(s) have been reproduced. In the introduction, a brief history of research in Ecuador (including the Galápagos archipelago) is presented. This checklist may serve as a baseline document for further research. It is not intended to be a revision- ary work, although we have recognised the following new synonyms: Neritina intermedia (var. β) minima K. Miller, 1879 = Clypeolum latissimum (Broderip, 1833); Aperostoma (Aperostoma) olivaceum Bartsch & Morrison, 1942 = Incidostoma quitense (L. Pfeiffer, 1852); Hemisinus pazi Tryon, 1866 = Hemisinus guayaquilensis (Petit de la Saussaye, 1853); Bulimus cuneus L. Pfeiffer, 1854 = Protobeliscus fairmaireanus (Petit de la Saus- saye, 1853); Synapterpes (Zoniferella) riveti Germain, 1907 = Mesembrinus vesperus Jousseaume, 1887 = Zoniferella albobalteata (Dunker, 1882); Synapterpes bicingulatus Fulton, 1908 = Zoniferella bizonalis (Germain, 1907); Thaumastus thompsonoides Ober- wimmer, 1931 = Kara thompsonii (L. Pfeiffer, 1845). The need for further field work is illustrated by the fact that for 60 species, only imprecise localities are known, while for 168 species no modern (i.e. last 50 years) records are available. There are 179 species considered as endemics for the Ecuadorian malacofauna. For both the terrestrial and fresh- water species the ecoregions where they occur are indicated where possible.

The Neotropical land snails (Mollusca, Gastropoda) collected by the ‘Comisión Científica del Pacífico’

Preprint

Full-text available

Dec 2016

The land snails collected by the ‘Comisión Científica del Pacifíco’ (CCP), a Spanish expedition to South and Central America from 1862–1866, are restudied and revised. The historical context of the expedition and the study of its collected material are described. Biographical data is given for the main persons involved. The land snails were previously studied by Joaquin Hidalgo between 1867 and 1893. A total of 3470 specimens belonging to 211 species are treated in this paper. Of 34 species mentioned by Hidalgo is his catalogue, the corresponding material could not be located. A lectotype is now designated for Drymaeus ( Drymaeus ) chanchamayensis (Hidalgo, 1870); Bulimus visendus Hidalgo, 1869 is now placed in the genus Synapterpes Pilsbry, 1896, a new combination.

The Neotropical land snails (Mollusca, Gastropoda) collected by the ‘Comisión Científica del Pacífico’

Preprint

Full-text available

Mar 2017

A review of Stenostylus Pilsbry, 1898 and Drymaeus Albers, 1850 (Mollusca: Gastropoda: Orthalicoidea: Bulimulidae) from Colombia, with description of new species.

Article

Full-text available

Jan 2019

Folia conchyliologica

Article

Full-text available

Dec 2019

The land snails of the genera Drymaeus Albers, 1850 and Stenostylus Pilsbry, 1898, both belonging to the family Bulimulidae, and occurring within northwestern South America are revised and notes on their distribution are given. 78 species of Drymaeus and two of Stenostylus are herein confirmed from Colombia, and are illustrated for comparison. Six new (sub)species are described: Drymaeus (Drymaeus) denticulus, D. (D.) duplexannulus, D. (D.) felix restrepoensis, D. (D.) iniurius, D. (D.) intermissus, D. (D.) luciensis. Drymaeus flexuosus megas Pilsbry, 1944 is now upgraded to species level. A lectotype is designated for Drymaeus roseatus montanus Pilsbry, 1901. The following nominal taxa are herein synonymised: Bulimus antioquiensis L. Pfeiffer, 1855 = B. baranguillanus L. Pfeiffer, 1853; Bulimus hachensis Reeve, 1850 = B. virgo Lea, 1838 = B. columbianus Lea, 1838; Drymaeus eversus alata Piaget, 1914 = Drymaeus eversus subula Piaget, 1914 = Bulimus violaceus Mousson, 1873 = B. confluens L. Pfeiffer, 1855; Drymaeus cantatus medinanus Pilsbry, 1935 = D. tusagasuganus Pilsbry, 1935 = Bulimulus (Drymaeus) plicatoliratus da Costa, 1898 = Bulimus convexus L. Pfeiffer, 1855; Drymaeus fallax chicoensis Breure, 1977 = Bulimus fallax L. Pfeiffer, 1853; Bulimus trivittatus Mousson, 1869 = B. felix L. Pfeiffer, 1862; Bulimus andicola L. Pfeiffer, 1847 = B. multilineatus Say, 1825; Bulimulus (Drymaeus) comis Preston, 1907 = Bulimus pealianus Lea, 1838; Drymaeus incognita da Costa, 1907 = D. bellus da Costa, 1906 = D. blandi Pilsbry, 1898 = Bulimulus (Drymaeus) smithii da Costa, 1898. For the following species, precise localities are given for the first time: Drymaeus (Drymaeus) angusta da Costa, 1906, D. (D.) auris (L. Pfeiffer, 1866), D. (D.) baranguillanus (L. Pfeiffer 1853), D. (D.) cognatus Pilsbry, 1901, D. (D.) geometricus (L. Pfeiffer 1846), D. (D.) inclinatus (L. Pfeiffer 1862), D. (D.) spadiceus da Costa, 1906, D. (Mesembrinus) koppelli (G.B. Sowerby III, 1892), D. (M.) muliebris (Reeve 1849). Newly recorded for the Colombian malacofauna are the following five taxa: Drymaeus (Drymaeus) fordii Pilsbry, 1898, D. (D.) glaucostomus (Albers, 1852), D. (D.) volsus Fulton, 1907, D. (Mesembrinus) interruptus (Preston, 1909). The following 27 taxa are excluded from the Colombian fauna as we consider them based on erroneous or doubtful records: Stenostylus meleagris (L. Pfeiffer, 1853), Drymaeus (Drymaeus) attenuatus (L. Pfeiffer, 1853), D. (D.) chimborasensis (Reeve, 1848), D. (D.) edmuelleri (Albers, 1854), D. (D.) linostoma (d’Orbigny, 1835), D. (D.) membielinus (Crosse, 1867), D. (D.) phryne (L. Pfeiffer, 1863), D. (D.) poecilus (d’Orbigny, 1835), D. (D.) protractus (L. Pfeiffer, 1855), D. (D.) rugistriatus Haas, 1952, D. (D.) strigatus (Sowerby, 1833), D. (D.) subinterruptus (L. Pfeiffer, 1853), D. (D.) D. (Mesembrinus) cactivorus (Broderip, 1832), D. (M.) deshayesi (L. Pfeiffer, 1845), D. (M.) dubius (L. Pfeiffer, 1853), D. (M.) flavidus (Menke, 1829), D. (M.) granadensis (L. Pfeiffer, 1848), D. (M.) liliaceus (Férussac, 1821), D. (M.) loxanus (Higgins, 1872), D. (M.) manupictus (Reeve, 1848), D. (M.) multifasciatus (Lamarck, 1822), D. (M.) nitidus (Broderip, 1832), D. (M.) pertristis Pilsbry, 1898, D. (M.) pervariabilis (L. Pfeiffer, 1853), D. (M.) studeri (L. Pfeiffer, 1847), D. (M.) translucens (Broderip, 1832).

Support and surprises: Molecular phylogeny of the land snail superfamily Orthalicoidea using a three-locus gene analysis with a divergence time analysis and ancestral area reconstruction (Gastropoda: Stylommatophora)

Article

Full-text available

Jun 2012

Phylogenetic relationships of Orthalicoidea, a highly diverse and dominant element in the Neotropics, were studied using nuclear and mitochondrial DNA sequences (ITS2/28S, CO1, H3). Specimens of various locations from the Southern Hemisphere (South America, Africa, Australia, New Zealand, Solomon Islands) were analysed (74 taxa, representing 30 genera). Our results support previously presented hypotheses, but also give surprises in terms of unexpected topologies. Phylogenetic trees were estimated using maximum likelihood and Bayesian inference, and compared with traditional classifications. Phylogenetic estimations using three loci gave a strong support for monophyly of Orthalicoidea, as well as for some clades within this group (Bulimulidae, Bothriembryontidae, Orthalicidae, Amphibulimulidae), but not for others (Odontostomidae and Megaspiridae). In the resulting revision of the classification scheme of the Orthalicoidea, the tribe Simpulopsini is raised to family rank. One new subfamily is recognized, the Bostrycinae. The family Bulimulidae Tryon 1867 is retained under ICZN Art. 35.5, despite the senior synonymy of Peltellinae Gray 1855. Our analysis supports an origin of the Orthalicoidea in South America, with subsequent radiations into other parts of the Neotropics and the Southern Hemisphere. The hypothesis that the distribution on the southern continents may be explained by vicariance due to break-up of Gondwana is only partially supported by divergence time analysis using fossil calibrations. Ancestral area reconstruction suggests various independent dispersals out of South America into Central America and the West Indies, and possibly two independent dispersals to explain the remaining relations between groups of taxa on the southern continents. Divergence time analysis further shows that the major diversification of extant taxa within the superfamily may have started around the Cretaceous-Paleogene boundary.

Molluscan names and malacological contributions of Wolfgang Karl Weyrauch (1907–1970) with a brief biography

Article

Full-text available

Jun 2008
MALACOLOGIA

Wolfgang Karl Weyrauch (1907−1970) studied land and freshwater gastropods of South America, mainly taxa belonging to the families Camaenidae, Charopidae, Clausiliidae, Endodontidae, Helicinidae, "Hydrobiidae", Orthalicidae, Pupillidae, Scolodontidae, Subulinidae, and Urocoptidae. Here we list the 198 molluscan names introduced by Weyrauch and all of his publications of malacological interest. A brief biography of Weyrauch and a list of taxa named for him are also provided. 265 RESUMEN Wolfgang Karl Weyrauch (1907−1970) se dedicó al estudio taxonómico de gasterópodos sudamericanos terrestres y dulceacuícolos, en su mayoria integrantes de las familias Camaenidae, Charopidae, Clausiliidae, Endodontidae, Helicinidae, "Hydrobiidae", Orthalicidae, Pupillidae, Scolodontidae, Subulinidae y Urocoptidae. Se presenta el inventario de los 198 nombres de moluscos introducidos por Weyrauch y una lista completa de sus trabajos de interés malacológico. Se incluye además una breve biografia de Weyrauch y los nombres de especies descriptas en su honor. Palabras clave: Mollusca, Gastropoda, Wolfgang Karl Weyrauch, bibliografía.

Notes on and descriptions of Bulimulidae (Mollusca, Gastropoda)

Article

Full-text available

Jan 1978

Bram Breure

The following new taxa are described: Plekocheilus (Eurytus) bruggeni, Thaumastus (Scholvienia) bambamarcaensis, T. (S.) gittenbergerorum, Bostryx aileenae, B. arcuatus, B. carinatus, B. c. trochiformis, B. circuliportus, B. c. rimacensis, B. coelhoi, B. cracentis, B. cunyacensis, B. frederici, B. fuligineus, B. hirsutus, B. huanucensis, B. h. poveli, B. inaquosum, B. kathiae, B. lentiformis, B. louisae, B. mordani, B. multilineatus, B. olmosensis, B. pauli, B. placitus, B. ploegerorum, B. spiculatus paucicostatus, B. s. pumilus, Bulimulus wiebesi, Sphaeroconcha, Naesiotus cleeforum, N. florschuetzi, N. subcostatus maranonensis, N. willinki, Scutalus (Vermiculatus), S. (V.) aureus, S. (V.) costifer naggsi, S. (V.) minutus, S. (V.) peakei, S. (V.) pyramidalis, S. (Suniellus), S. (S.) chillu, Discoleus, Neopetraeus catamarcanus orientalis. The following taxa are placed in synonymy: Bulimus lamarckianus Pfeiffer, 1848 = Plekocheilus (Eurytus) coloratus (Nyst, 1845); Bulimus scytodes Pfeiffer, 1853 = Plekocheilus (Aeropictus) veranyi (Pfeiffer, 1848); Antitragus Oberwimmer, 1931 = Plekocheilus (Eudolichotis) Pilsbry, 1896; Tholus Strebel, 1909 = Atahualpa Strebel, 1910 = Thaumastus (Thaumastus) Albers, 1860; Megalobulimus (Microborus) incarum Pilsbry, 1944 = Thaumastus (Paeniscutalus) crenellus (Philippi, 1867); Kionoptyx Haas, 1966 = Floreziellus Weyrauch, 1967 = Naesiotellus Weyrauch, 1967 = Bostryx Troschel, 1847; Bulimulus (Lissoacme) rudistriatus Haas, 1955 = Bostryx abancayensis Pilsbry, 1944; Bulimus reconditus Reeve, 1849 = Bostryx nigropileatus (Reeve, 1849); Bulimulus (Scutalus) punctilineatus Haas, 1951 = Naesiotus (Reclasta) tarmensis Weyrauch, 1967 = Bostryx pictus (Pfeiffer, 1855) ; Buliminus styliger Beck, 1837 = Bostryx vittatus (Broderip, 1832); Loboa Ihering, 1917 = Bulimulus Leach, 1814; Xenothauma Fulton, 1896 = Scutalus (Scutalus) Albers, 1850; Bulimulus (Drymaeus) baroni Fulton, 1897 = Scutalus (S.) cretaceus (Pfeiffer, 1855); Bulimus revinctus Hupé, 1857 = Scutalus (Vermiculatus) gayi (Pfeiffer, 1857); Bulimulus atahualpa Dohrn, 1863 = Neopetraeus tessellatus perincrassatus Pilsbry, 1898 = Neopetraeus weyrauchi Pilsbry, 1944 = Neopetraeus cremnobates 'Pilsbry' H . B. Baker, 1963 = Neopetraeus tessellatus (Shuttleworth, 1852); Drymaeus (Leiostracus) ruthveni H. B. Baker, 1926 = Leiostracus demerarensis (Pfeiffer, 1861); Drymaeus obliquus poecilogramma Ancey, 1901 = Leiostracus obliquus (Reeve, 1849); Vitrina salomonia Pfeiffer, 1853 = Simpulopsis (S.) rufovirens (Moricand, 1846); Pseudoglandina Weyrauch, 1967 = Simpulopsis (Eudioptus) Albers, 1860; Pseudoglandina agitata Weyrauch, 1967 = Simpulopsis (Eudioptus) citrinovitrea (Moricand, 1836). Anatomical data are presented for 94 species, new locality records are given for 50 species and 22 species are redescribed.

The Amphibulimidae (Mollusca: Gastropoda: Orthalicoidea) from Colombia and adjacent areas

Article

Full-text available

Oct 2011

The land snails of the family Amphibulimidae occurring within northwestern South America are critically examined and notes on their distribution are given. The total number of taxa discussed is 61 (7 Dryptus, 54 Plekocheilus); of these, four Colombian Dryptus and 32 Plekocheilus species are included. The following new species are described: Plekocheilus (P.) bigener, P. (P.) incognitus, Plekocheilus (Eurytus) camaritagua, P. (E.) labiosus, and P. (E.) paraguas. The following nominal taxa are now synonymized: Plekocheilus (P.) speciosus (Pfeiffer, 1854) = P. (P.) plectostylus (Pfeiffer, 1848); Plekocheilus (Eurytus) couturesi Ancey, 1900 = P. (P.) glandiformis (Lea, 1838); Plekocheilus (Aeropictus) latilabris (Pfeiffer, 1855) = P. (A.) succineoides (Petit de la Saussaye, 1840); Plekocheilus (Eurytus) virgatus (Pilsbry, 1935) = P. (E.) mabillei (Crosse, 1867) = P. (E.) pulicarius (Reeve, 1848). Also, Simpulopsis fulguratus Miller, 1878 is now consid-ered a junior subjective synonym of Simpulopsis (Eudioptus) citrinovitrea (Moricand, 1836). For the following species, precise localities are given for the first time: Dryptus stuebeli (Martens, 1885), Plekocheilus (Aeropictus) cathcartiae (Reeve, 1848), P. (E.) cardinalis (Pfeiffer, 1853), P. (Eurytus) episcopalis auriformis (Da Costa, 1904), P. (E.) episcopalis corticosus (Sowerby, 1895), and P. (E.) lynciculus (Deville & Huppé, 1850). Newly recorded for the Colombian malaco-fauna are the following taxa: Plekocheilus (P.) couturesi (Ancey, 1900), P. (Eurytus) piperitus (Sowerby I, 1837). The fol-lowing taxa are excluded from the Colombian fauna: Dryptus funckii (Nyst, 1843), Plekocheilus (Aeropictus) veranyi (Pfeiffer, 1848), Plekocheilus (Eurytus) jimenezi (Hidalgo, 1872).

Mysterious or confusing: enigmatic species in the Orthalicidae (Gastropoda, Pulmonata)

Article

Full-text available

Jan 2008

Bram Breure

Five enigmatic species that have remained unnoticed or proved confusing are treated: Bostryx alausiensis (Cousin), Bulimulus gelidus (Reeve), Stenostylus colmeiroi (Hidalgo), Stenostylus nigrolimbatus (Pfeiffer) and Simpulopsis (S.) simula (Morelet). Drymaeus (D.) cleefi Breure & Eskens is now synonymized with Stenostylus nigrolimbatus (Pfeiffer).

Taxonomic review of the Spixia pyriformis species complex (Gastropoda: Pulmonata: Odontostominae)

Article

Jun 2007

EUGENIA SALAS OROÑO

The genus Spixia (Pilsbry & Vanatta, 1898) is endemic from southern South America. Information about this group is scarce, particularly referring to its anatomy and shell ultrastructure. According to current classification the Spixia pyriformis complex includes the following species, S. doellojuradoi (Parodiz 1941), S. martensii (Doering 1874), S. pyriformis (Pilsbry 1901) and S. tucumanensis (Parodiz 1941). In the present study a complete redescription of the species component of this complex, as well as new information on shell ultrastructure and anatomy of the different organ systems is provided. The Spixia pyriformis complex has a pyriform-subpyriform shell shape that distinguishes it from the rest of the species of the genus. Three of the four species studied have a similar periostracum structure consisting of spiral striae and triangular lamellae. Radular morphology is similar in all species. Jaw structure is diagnostic for each species. The inner morphology of the penis is divided into five areas marked by differential pattern of sculpture. Four new anatomical characters, the presence of a pallial gland in the distal portion of the pulmonary roof, the presence of a penial gland in the penial complex, the position of the insertion of the penial retractor muscle in either penial area I or area II and the sculpture of penis inner wall, are identified as the most informative for species diagnosis.

Some Anatomical, systematical and Geographical data on Neniinae (Gastropoda, Clausiliidae)

Article

I. INTRODUCTION Up to the moment more than one hundred species and subspecies of Neniinae are known from Central and South America. They are incorporated into 22 genera and subgenera, almost exclusively based on characters of the shells. Only little is known of the anatomy of the animals (Binney, 1871; Binney & Bland, 1871; Baker, 1961; Hesse, 1925; Scott, 1954; Loosjes, 1957; Pilsbry, 1926; Polinski, 1922; Zilch, 1953, 1959). These authors described the radula and/or the genital organs of one or several species. Thus from eleven species the radula has been described and from five the genital apparatus. By the courtesy of Prof. Dr. W. Weyrauch we were able to study the anatomical features of a relatively large number (38) of Peruvian and Argentine Neniinae. We wish to express our most cordial thanks to him for his generosity and also to Dr. B. Hubendick, who provided us with a sample of Nenia tridens (Chemnitz) from Puerto Rico. In Thiele's (1929/31) classification the American Neniinae are grouped into three genera, Nenia, Peruinia and Temesa, based on characters of the shells and of the radula. It turned out impossible to fit the information we found, into this system as it indicates the presence of many more units. In this survey we will follow Zilch's (1960) classification, which enumerates a number of genera and subgenera, based on conchological characters, although the order will be altered according to the results obtained (conf. chapter IV). Thus conchological data of the genera or subgenera involved have to be studied in Zilch (1960), or eventually in Thiele (1929/31).

On a new Andinia (Gastropoda, Clausiliidae) from Peru

Article

Jan 1957

F. E. Loosjes

Some time ago Prof. Dr. W. Weyrauch at Lima, Peru, sent to me specimens of a species of the subfamily Neniinae that proved to be new to science. Already about 70 species of the subfamily have become known from Peru, for an important part discovered by Prof. Weyrauch himself. Andinia (Ehrmanniella) flammulata spec. nov. (Figs. 1, 2) Diagnosis. A small, rather fusiform species of the subgenus Ehrmanniella Zilch. The decollated shell is provided with rather irregular fine white striae. Especially below the suture these striae are thickened, strikingly white, and arranged in groups of about 5 to 15 on low nodules. These nodules alternate with brown, quite smooth, malleated patches. At some distance it looks like small white and brown areas running obliquely over the surface of the shell below the suture. Lunella more or less interrupted below the middle. The principal plica is only 1/4 whorl long. The receptaculum seminis and its duct unto the junction with the diverticulum are as long as the diverticulum itself. Description. The shell is decollate, somewhat fusiform, moderately strong, with rather convex whorls of which the penultimate is broadest. Whorls of the decollate shell 4 to 7; some shed tops (juvenile shells) with straight outlines consist of about 10 whorls; sculptured with low, but rather large nodules (which can be taken as corresponding with the upper parts of the obliquely running ribs of Polinski's Nenia wagneri1)) below the suture, provided with groups of about 5-15 clear white striae, between these nodules there are brown smooth patches, often running obliquely below the nodules; 1) „breite, niedrige, stumpfe und wulstartige Rippen, welche von links oben nach rechts unten verlaufen". the middle and lower parts of each whorl are usually rather irregularly

Observations On the Formation of Spermatophores in a Bulimulid Land Snail, Drymaeus Canaliculatus (Pfeiffer, 1845) (Mollusca, Gastropoda, Pulmonata)

Article

Jan 1976

In one specimen of Drymaeus canaliculatus (PFEIFFER, 1845) a spermatophore was observed in the phallus complex. The structure of the spermatophore is described using light microscopy observations and histological methods. A hypothesis is presented describing the formation of spermatophores in the epiphallus and flagellum The process of spermatophore-formation is initiated in the most distal part of the epiphallus and the proximal part of the flagellum.

A new species of Plagiodontes from Argentina, and new data on the anatomy of four other species in the genus (Gastropoda: Orthalicidae, Odontostominae)

Article

Jun 2009

Plagiodontes weyrauchi sp. nov. is among the smallest species in the genus Plagiodontes (Doering, 1877). Statistical multivariate analyses confirmed its distinctiveness from other species, mostly because of its high number of whorls (8–9.75), relatively small width, and low proportions of last whorl and aperture length to shell length; multivariate discriminant analysis applied to this species and its most similar relative, P. multiplicatus parvus, correctly identified 100% of specimens. Pallial and genital anatomy supports P. weyrauchi as a different species, mainly because of the position of the ureteric pore, the small general size of the genital system, the presence of an accessory lobe alongside the verge, and the vaginal internal structure. Internal anatomy of four other small-sized taxa in the genus was studied. Two of them have a verge with no accessory lobe [P. dentatus (Wood, 1828) and P. patagonicus (d'Orbigny, 1835)], while the remaining two have an accessory lobe with different characteristics [P. rocae Doering, 1881 and P. multiplicatus parvus (Hylton-Scott, 19528. Hylton-Scott , M I . 1952. Nuevos moluscos terrestres del norte argentino. Acta Zool Lilloana., 10: 5–29. View all references)]. Internal structure of the vagina also shows specific differences among Plagiodontes species; their phylogenetic significance will be further investigated.

Weurauch’s type localities: a clarification; with illustrations of types of Orthalicoidea (Mollusca, Gastropoda, Stylommatophora) in the Tucumán museum

Abstract

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