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7
Contribution to the Nolinae (Lepidoptera, Noctuidae) fauna of
North Thailand (Plates 1-11)
by
Gyula M. LászLó, Gábor Ronkay and Thomas J. Witt
Abstract
Present paper is the rst comprehensive summary of the Nolinae (sensu stricto) fauna of Northern Thailand. It contains the records
of 127 species occurring in this area, with the descriptions of 16 new genera and 46 new species, numerous new combinations,
stati and synonyms. With 11 colour plates and 58 genitalia gures.
Introduction
The interpretation and the taxonomic rank of the compact and monophyletic group called as Nolinae has been
changed several times but was mainly treated as a subfamily of the family Arctiidae. It has been upgraded to a
family rank rstly by HoLLoWay (2003) in the Moths of Borneo series, lumping them into a common phyletic unit with
several clades placed formerly into the family Noctuidae (e.g. Chloephorinae, Sarrothripinae and Eariadinae). This
opinion was accepted by FibigeR & LaFontaine (2005) in their comprehensive work on the taxonomy and systemat-
ics of the superfamily Noctuoidea.
The target objects of this study are all those species and generic rank taxa which belong to the subfamily Nolinae
(s. str.). The studies were based principally on the huge Nolinae collections of the Witt Museum Munich (MWM),
the Hungarian Natural History Museum Budapest (HNHM) and The Natural History Museum London (formerly
British Museum, Natural History; BMNH). Almost all specimens studied from Thailand originate from the northern
mountainous areas of the country, the few exceptional examples are found in the material of the BMNH.
The systematic part contains all records of the species from Thailand recognised in the material of the mentioned
three museums. In the cases of the newly established genera, the descriptions of all newly discovered species
are included, also those which have no records from Thailand (yet).
Abbreviations
LGN – slide of Gyula M. LászLó
BMNH – British Museum Natural History, London
HNMH – Hungarian Natural History Museum, Budapest
LS London – Linnean Society London
MNHP – Museum National d’Histoire Naturelle, Paris
MWM – Museum Witt, Munich
NHMW - Naturhistorisches Museum, Wien
NRS – Naturhistoriska Riksmuseet, Stockholm
RNH – Rijskmuseum van Natuurlijke Historie, Leiden
UMOX – University Museum, Oxford
ZFMK – Zoologisches Forschunginstitut und Museum Alexander Koenig, Bonn
Addresses of the authors:
Gyula M. LászLó: Fadrusz utca 25, H-1114 Budapest, Hungary. E-mail: laszkogy@t-online.hu
Gábor Ronkay: Szent István krt. 4., H-1137 Budapest, Hungary.
Thomas J. Witt: Tengstrasse 33, D-80796 München, Germany.
Esperiana Band 15: 7-125 Schwanfeld, 12. Januar 2010 ISBN 978-3-938249-10-9
8
ZIN – Zoological Institute, Russian Academy of Sciences, St. Petersburg
ZMHU – Zoologisches Museum der Humboldt-Universität, Berlin
ZMUC – Zoological Museum, University of Copenhagen
ZSM – Zoologische Sammlung des Bayerischen Staates (Zoologische Staatssammlung), München
Systematic part
Alcanola gen. nov.
Type species: Melanographia tympanistis Hampson, 1900
Species content:
A. tympanistis Hampson, 1900 comb. nov.
A. speideli spec. nov.
A. obscurata spec. nov.
Diagnosis. The external appearance of the species of the new genus resembles somewhat to certain species of
the rather diverse (and supposedly polyphyletic) genus Aquita WaLkeR, 1863 (e.g. A. acontioides (WaLkeR, 1862))
and of the recently described Porcellanola LászLó, Ronkay & Witt, 2006 by their shiny, „porcelain white” ground
colour and brownish terminal area of the forewing. The differential features are the absence of the whitish suffusion
in the apical area and the sharply dened, shiny blackish-bluish markings of the forewing. Alcanola has greyish
brown ground colour on the outer two-thirds of the forewing without conspicuous crosslines or other drawings.
An apomorphic feature of the new genus (unique within the whole family) is a special structure on the x forewing.
It appears as a dorsally convex hump within the cell, with sclerotised marginal ring and erected scale-tufts. The
ventral side of this structure is connected to conspicuously inated parts of the cellveins.
The genus comprises, according to the conguration of the x genitalia, two distinct phyletic lineages, the tympanis-
tis and the obscurata species-groups. The basic structure of the x genitalia of both lineages of Alcanola displays
closer relationship with Meganola Dyar, 1898, but obviously differs by the following features: in the tympanistis
species-group the elongate, medially tapering valva is terminated in a large, thorn-like costal process and the
well-developed harpe arising from the outer half of valva, while the valva of Meganola has no terminal spine, and
the harpe is always located basally. The valval structure of the obscurata species-group is more similar to that of
Meganola, the valva is medially only slightly tapering, apically broadly rounded, lacking the terminal costal proc-
ess. The obscurata species-group differs from Meganola by the position of the robust harpe in the outer third of
valva and the presence of sclerotized triangular costal lobe; these features indicate its common origin with the
tympanistis species-group. The w genitalia of Alcanola is rather unique within the whole subfamily having a pair
of proximally elongated lobes on the eighth sternite which are the modied apophyses anteriores, and a pair of
elongate, weakly sclerotized, ribbon-like signa.
Alcanola tympanistis (Hampson, 1900) comb. nov.
(Pl. 1, gs 1-3; gen. g. 1)
Melanographia tympanistis Hampson, 1900, Catalogue of the Lepidoptera Phalaenae in the British Museum 2: 49, text g. 13.
Type-locality: [India] Sikkim. Holotype: x, in coll. BMNH.
Type material examined:
Holotype of Melanographia tympanistis Hampson, 1900: x, „TYPE” (red ring type label), „Sikkim, 1800 ft, June 1897, Dudgeon”,
slide No. BM Noct 17721 (coll. BMNH).
Additional material examined:
Nepal. 1 x, Milke Danda, Gupha pass, 3000 m, 23.viii.2000, leg. T. Csővári & m. HRebLay, slide No. LGN 374 (W 7317); 3 xx,
3 ww, Tanahoun distr., Bimalnager village, 530 m, 11-12.iv.1995, leg. gy.m. LászLó & g. Ronkay, slide Nos LGN 372, LGN 946
(W 7318) (xx), LGN 373 (w); 2 xx, same locality, but collected at 12.x.1994 by g. CsoRba & L. Ronkay (coll. HNHM and MWM).
Myanmar. Prov. Shan: 1 x, Kalaw, 1400 m, 30.x.1999, leg. M. HRebLay, slide No. LGN 1048 (W 7319). Prov. Mandalay: 1 w, 3 km
SE of Kyause, 220 m, 29.x.1999, leg. M. HRebLay, slide No. LGN 518 (W 7320) (coll. MWM). Cambodia. Prov. Mondolkiri: 1 w,
9
Seima Biodiversity Conservation Area, between Seima and O’Rang, 12º12’12N, 107º01’09E, 300 m, 30.i.2006, leg. g. CsoRba &
g. Ronkay, slide No. 1047 (coll. HNHM).
Remarks. The species has not yet been recorded from Thailand.
Alcanola speideli spec. nov.
(Pl. 1, g. 4; gen. g. 2)
Holotype. x, „India mer., Kerala, 6 km N Munnar, 1700m, Kodalar Tea Estate, 10º06’N/77º04’E, 14-15.4.1997, leg. sCHintLm. &
siniaev, Bergregenwald 14°C”, slide No. LGN 402 (W 7321) (coll. MWM).
Diagnosis. Alcanola speideli is a sister species of A. tympanistis, the differences between the two species are
as follows: The new species has more whitish colouration on the whole forewing, while A. tympanistis has white
colour only in the basal and ventral areas of the forewing. The wingspan of the new species is 17-20 mm, the
length of forewing is 8-10 mm.
The x genitalia of A. speideli differ from those of A. tympanistis by their somewhat shorter and less arched valva
with more rounded apex and the somewhat shorter and straight, spine-like costal process (that of A. tympanistis
is longer and slightly arched). The harpe of the new species is somewhat thicker; the base of the harpe is situated
somewhat closer to the valval end than in A. tympanistis; in addition, the aedeagus of A. speideli is somewhat
shorter with conspicuously shorter coecum penis. The w is unknown.
Remarks. The species is known from the type-locality only.
Alcanola obscurata spec. nov.
(Pl. 1, g. 5; pl. 11, g. 1; gen. g. 3)
Holotype. x, „Thailand, Chiang Mai, 6 km SE of Pang Faen (1100m), 2001.07.09, leg: HentsCHeL, petRányi”, slide No. LGN 945
(W 7296) (coll. MWM).
Paratypes: Thailand. Prov. Nan: 1 x, 5 km N of Ban Luang, 350m, between Pi Nai and Pi Tai, 100º27’E, 18º56’N, 4.xi.2002, leg.
b. HeRCzig & g. Ronkay, slide No. LGN 762 (W 7297). Prov. Chiang Mai: 1 x, Mae Suai, 500m, Charin Garden
Resort, 99º35’E, 19º47’N, 2.xi.2002, leg. b. HeRCzig & g. Ronkay, slide No. LGN 796 (W 7298) (coll. MWM); 1 x,
700 m, Samoeng/Hang Dong Rd. Km 15, 5.vi.-12.vi.1988, leg. a.m. Cotton & i.J. kitCHing BM 1988-312; 1 w,
Doi Suthep, 1100 m, 22-23.viii.1990, leg. H. inoue, slide No LGN 1322 (BM Arct. 6187) (coll. BMNH).
Diagnosis. Alcanola obscurata represents a distinct phyletic lineage within the genus, the new species is, how-
ever, hardly distinguishable from its congeners by the external features, the satisfactory identication requires the
examination of the x genitalia. There are only slight differences in the external appearance: the outer area of the
forewing is somewhat darker and larger, the edge of the brown area in A. obscurata reaches the middle of the
ventral margin and the crosslines are fully reduced. The ventral margin of A. tympanistis and A. speideli is white
towards the tornus and the crosslines are recognisable. These characters are, however, clearly observable only
in the freshly emerged specimens. Wingspan 17-20 mm, length of forewing 8-10 mm.
The differences between the x genitalia of A. obscurata and the members of the A. tympanistis – A. speideli spe-
cies-pair are surprisingly large, the differences are as follows: A. obscurata has much broader, tongue-shaped
valva with broadly rounded apex and without costal process, while the valvae of A. tympanistis and A. speideli are
strongly tapering, having well developed, acute, thorn-like costal process. The harpe of A. obscurata is nger-like,
much shorter and broader than the thorn-like harpe of the related two species; in addition, the penicular lobes of
A. obscurata are somewhat broader than in the taxa of the A. tympanistis group.
The w genitalia of the new species lacks the elongated lobes from the eighth sternite which are typical of A.
tympanistis, there are only short, apically rounded processes. The apophyses anteriores of the new species are
shorter and simple, the ductus bursae is much longer and narrower, and the signa broader and longer than in A.
tympanistis, with a short strongly sclerotized section in their distal third.
Distribution. Thailand.
10
Barnanola gen. nov.
Type species: Barnanola atra spec. nov.
Species content:
B. atra spec. nov.
B. sumatra spec. nov.
B. bigeri spec. nov.
B. orbiculata spec. nov.
Diagnosis. Relatively large species, wingspan 14-21 mm, length of forewing 7-9 mm. The taxa belonging to the
new genus resemble the members of the genus Alcanola described above, by their whitish ventral suffusion on the
dark brown forewing but the whitish ventral area is considerably shorter in Barnanola than in Alcanola, extending
only to the middle of the ventral margin. The brownish coloration is much more extensive and darker in Barnanola
than in Alcanola. The special sclerotised hump, being characteristic for Alcanola, is absent from Barnanola.
The conguration of the x genitalia also indicates the relationship between the two genera, displaying conspicu-
ous differences. Barnanola has, in comparison with Alcanola, much more elongated tegumen with long, narrow,
undivided apical part, slightly arched and moderately tapering valva without costal sclerotization or process, short,
slightly curved saccular process reaching only the middle of valva, and, last but not least, much weaker vinculum.
In case of Alcanola, the tegumen is simple, the valva is either strongly tapered medially or tongue-shaped without
tapering part but always with sclerotized costal margin and differently developed costal process, the elongated
harpe exceeds the costa, and the vinculum is considerably stronger than in Barnanola. The aedeagus of Barnanola
is very short and straight, with one well-developed cornutus in the vesica, while the aedeagus of Alcanola is longer,
slightly arched, and the vesica lacks the cornuti.
The differences in the w genitalia of the two genera are also clearly recognisable as Barnanola has simple eighth
sternite with very short and ne but distinct apophyses posteriores, much shorter sclerotized part of ductus bursae
and large, elongated, „baguette-like” double signa. In Alcanola the eighth sternite has a characteristic shape, the
apophyses posteriores are strongly modied; and the signum is absent.
Barnanola atra spec. nov.
(Pl. 1, gs 6-7; gen. g. 4)
Holotype. x, „Djoenggo-Ardjoeno 1500m, 9.37., O.-Java, Kalis”, slide No. LGN 913 (coll. RNH, Leiden).
Paratypes. 1 x, with the same data as the holotype, slide No. LGN 914 (coll. RNH, Leiden); 3 xx, Djoenggo Ardjoeno, E Java,
4500’, May 1934, leg. J. p. a. kaLis (slide Nos LGN 1267 (BM Arct. 6087), LGN 1268 (BM Arct. 6088), LGN 1269
(BM Arct. 6089)) (coll. BMNH).
Diagnosis. The genus Barnanola comprises externally usually confusingly similar and closely related species,
thus the proper identication requires the examination of the genitalia. The only exception is B. orbiculata, which
is easily distinguished from its congeners by its large, rounded, bright white patch in the forewing median area
and a greyish white reversed triangular spot in the tornal area; these prominent light spots are absent from the
other species of the genus.
The x genitalia of B. atra, B. sumatra and B. bigeri are rather similar, but easily separable by the shape of the
valva and the conguration of the saccular process. B. atra has, in comparison with the other two species, the
most elongated valva with narrowest apical third, and the longest sacculus with the most arcuate saccular proc-
ess. The valva of B. sumatra is somewhat shorter, less tapering, apically more arcuate, the sacculus is somewhat
narrower with almost straight process, while the valva of B. bigeri is conspicuously shorter, broader, apically not
tapering and the sacculus is the shortest but broadest within the genus, with slightly arched process. The w of B.
atra is unknown.
Distribution. The species is known from Java.
11
Barnanola sumatra spec. nov.
(Pl. 1, g. 8; gen. g. 5)
Holotype. x, „Sumatra, Prapat HW2 9.9.1985, Dr. DieHL leg.”, slide No. LGN 440 (coll. HNHM).
Paratypes. Indonesia. North-Sumatra: 1 x, 4 km S. Sidikalang, 2º41’51N, 98º18’18E, 1250 m, 16.ii.2002, leg. k. LaRsen & m. FibigeR,
slide No. LGN 941 (coll. FibigeR). Thailand. 1 x, 7 km N of Ranong, Ch 9 TV relay stn., 350-500 m, 26-29.xi.1991,
i.J. kitCHing & a.m. Cotton, BM 1992-9, slide No. LGN 1344 (BM Arct. 6217) (coll. BMNH).
Diagnosis. The diagnostic features of the x genitalia of B. sumatra are as follows: the medium-long, distally less
tapering and apically rather rounded valva and the slender, almost straight saccular process. The detailed com-
parison of the three species of the genus is given in the diagnosis of B. atra. The w is unknown.
Distribution. The species has been found in Sumatra and in Thailand.
Barnanola bigeri spec. nov.
(Pl. 1, g. 9; gen. g. 6)
Holotype. x, „Indonesia, North-Sumatra HW II. Near Prapat, 1050 m, 2º45’52N, 99º58’20E, 04.iii.2002. leg. m. FibigeR”, slide No.
LGN 942 (coll. FibigeR).
Diagnosis. The diagnostic features of the x genitalia of B. bigeri are the remarkably short and broad, apically not
tapering valva and the short but broad sacculus with slightly arched saccular process. The w is unknown.
Distribution. The species is found only once in Sumatra.
Barnanola orbiculata spec. nov.
(Pl. 1, gs 10-11; pl. 11, g. 2; gen. g. 7)
Holotype. x, „Thailand, Changwat Mae Hong Song, 1 km S of Bahundanda, 1000 m, 6.ii.1998 leg. Márton HRebLay & Csaba
szabóky”, slide No. LGN 146 (W 7322) (coll. MWM).
Paratypes. Thailand. Prov. Nan: 1 w, 5 km N of Bo Luang, 1000 m, 4.ii.2000, leg. m. HRebLay & a. szabó, slide No. LGN 363 (W
7323) (coll. MWM). Indonesia. Sumatra: 1 w, Prapat HW 3, 2.vii.1983, leg. e. DieHL, slide No. LGN 435; 1 w,
from the same site, 27.vii.1984; 1 w, from the same locality, 25.xi.1984, slide No. LGN 436; 1 w, Prapat HW 3,
19.v.-2.iv.1983; 1 w, 15 km S of Dolok Merangir, Dolok Ulu, 3.viii.1979, slide No. LGN 437, leg. e. DieHL (coll.
HNHM).
Diagnosis. The diagnostic external feature of B. orbiculata is the presence of a large, rounded, bright white patch
in the forewing median area and a greyish-white reversed triangular spot in the tornal area. The genitalia of the
species cannot be compared with those of its congeners, B. atra, B. sumatra and B. bigeri, as B. orbiculata is
known by ww only while the type-series of the other three species are all xx. Thus, there is a chance that B.
orbiculata represents the w sex of one of the three other Barnanola species, despite of the rather large external
differences. As the scale of the possible sexual dimorphism cannot be estimated from the material examined, B.
orbiculata is treated tentatively as a distinct species; this question is to be revised when new Barnanola material
will be available.
Distribution. The species is found in the northern part of Thailand and in Sumatra.
12
Leucobaeta gen. nov.
Type species: Zia hemiphea Hampson, 1905
Species content:
L. hemiphaea (Hampson, 1905) comb. nov.
Diagnosis. The species of the presumably monotypical new genus is rather similar externally to those of Leuconola
gen. nov. by the rather broad white(ish) basal area, and the dark brownish-grey outer two-thirds of the forewing,
but are easily distinguishable by the considerably darker hindwing and the more uniformly dark brownish grey
forewing terminal area of Leucobaeta, this part of the wing is greyish white in Leuconola with relatively large pale
grey patches. Leucobaeta also resembles externally Alcanola and Barnanola but the white basal area is much
broader in Leucobaeta, extending towards the inner quarter of the costal margin, while in Alcanola and Barnanola
the basal area is much narrower and the white colour is restricted to the ventral part of the basal eld.
The x genitalia of Leucobaeta are unique within the whole Nolinae by their rather short and broad, apically truncate
valva with short apical costal process; the related genera mentioned above have the usual basic structure of the
valva with elongated, distally gradually tapering valvae and rounded apex. The tegumen of Leucobaeta is similar
to that of Barnanola, but the elongated distal part is much broader in the former genus. The armature of the vesica
of the generic complex also shows easily recognisable diagnostic features: Leucobaeta has two robust, but rather
short cornuti; Leuconola and Alcanola have no cornutus in the vesica, while the vesica of Barnanola is armed with
one large, elongated cornutus.
The w genitalia of Leucobaeta also display rather characteristic feature: a large appendix bursae is present, such
a structure is absent in the related genera.
Leucobaeta hemiphea (Hampson, 1905) comb. nov.
(Pl. 1, gs 16-17; gen. g. 8)
Zia hemiphea Hampson, 1905, Annals and Magazine of Natural History 7(15): 177. Type-locality: [Indonesia] Java, Arjuno. Holotype:
w, in coll. BMNH.
Zia leucobaeta WiLeman & West, 1928, Entomogist 61: 206. Type-locality: Philippines, Luzon, Rizal, Montalban. Holotype: x, in
coll. BMNH; syn. nov.
Type material examined:
Holotype of Zia hemiphea Hampson, 1905: w, „TYPE” (red ring type label), „Arjuno 3000 ft. W. DoHeRty 96-252”, slide No. LGN 724
(BM Noct. 18204) (coll. BMNH).
Holotype of Zia leucobaeta WiLeman & West, 1928: x, „TYPE” (red ring type label), „Montalban, Prov. Rizal, Luzon, 21.iv.1914, a.
e. WiLeman”, slide No. LGN 727 (BM Noct. 18207) (coll. BMNH).
Additional material examined:
Thailand. Prov. Chiang Mai: 1 x, between Chiang Dao and Kariang, 900 m, 98º48’E, 19º25’N, 26.x.2002, leg. b. HeRCzig & g.
Ronkay, slide No. LGN 761; 2 xx, 1100 m, 4 km SE of Pang Faen, 13.i.2004, leg. p. HentsCHeL & a. szabó, slide Nos LGN 1007
(W 7324), LGN 1008 (W 7325) (coll. G. Ronkay and MWM).
Leuconola gen. nov.
Type species: Leuconola herczigi spec. nov.
Species content:
L. herczigi spec. nov.
L. wilsonae (HoLLoWay, 2003) comb. nov.
13
Diagnosis. The new genus is supposedly related to Leucobaeta according to the external appearance of the species
of the two genera: the forewing of the moths has broad whitish basal, and dark brownish-grey median and marginal
areas. The external differences between the two genera are as follows: Leuconola has considerably lighter hindwing
and much paler greyish white terminal area of forewing with relatively large pale grey patches, while Leucobaeta
has much darker hindwing and almost uniformly dark brownish grey terminal area of forewing.
The conguration of the x genitalia of the two genera is rather different: Leuconola has elongated, distally taper-
ing valva with more or less sinuous margins, sclerotised costa with well-developed triangular costal process at
basal section, elongated, sword-like saccular process, and the vesica lacks the cornuti; in Leucobaeta, the valva is
rather short, not tapering, apically broadly truncate with straight margins, and with a short costal process apically,
the saccular process is shorter and broader, more or less button-like, and the vesica is armed with two robust,
relatively short cornuti.
The w genitalia of the new genus are rather unique within the whole subfamily. Leuconola has conspicuously long
eighth tergite, characteristic „M” shaped sclerotization at the ostium bursae, very short ductus bursae with strongly
sclerotized lateral and weakly sclerotized dorsal and ventral walls, relatively small but characteristic, sclerotized
cervix bursae, and the elongate-elliptical corpus bursae with a special, narrow, medium-long and tube-like append-
age. The corpus bursae bears a pair of rather remote signa, the distal one is large and more or less crescent-like,
while the proximal one is rather small, short and narrow nger-like (see the Fig. 9).
Leuconola herczigi spec. nov.
(Pl. 1, g. 18; gen. g. 9)
Holotype. x, „N. Thailand, Chiang Mai Prov, between Chiang Dao and Kariang, 900 m, 98˚48’E, 19˚25’N, 26.x.2002, leg. b. Herczig
& G. Ronkay”, slide No. LGN 760 (W 7299) (coll. MWM).
Diagnosis. Leuconola herczigi is a close relative of L. wilsonae (HoLLoWay, 2003) comb. nov., described from
Borneo, however the external appearance of the two species is conspicuously different. Comparing the two close
relatives, the new species has much darker forewing pattern, consisting of a relatively thick and oblique dark grey
median streak and pale greyish suffusion in the terminal area, without dened crosslines, while L. wilsonae has
conspicuous bright white forewing with ne brownish crosslines and a prominent dark grey triangular spot in the
medio-costal area. The differences in the x genitalia of the two species are less conspicuous than their external
appearance, supporting strongly their close relationship. The x genitalia of L. herczigi differ from those of L. wilso-
nae by the considerably shorter and thicker uncus, broader tegumen, longer and medially more dilated valva with
larger costal lobe and narrower apical part, and by the somewhat thicker saccular process. The vinculum is also
different in the Leuconola herczigi-L. wilsonae species-pair as the new species has rather narrow, elongate apical
process while the vinculum of L. wilsonae is simply V-shaped. Finally, the aedeagus of L. herczigi is considerably
longer than in L. wilsonae, with more sclerotized and shorter basal part. The w is unknown.
Distribution. Northern Thailand.
Leuconola wilsonae comb. nov.
(Pl. 11, gs 3-4; gen. g. 10)
„Nola” wilsonae HoLLoWay, 2003, The Moths of Borneo 18: 60, Plate 3, g. 80. Type-locality: [Borneo] Sarawak, Gunong Mulu.
Holotype: x, in coll. BMNH.
Type material examined:
Holotype of „Nola” wilsonae HoLLoWay, 2003: x: „Sarawak, Gunong Mulu Nat. Park, R.G.S. Exped. 1977-8 (J.D. HoLLoWay et
al.), Site 22, April, W. Melinau Gorge, 150 m. 421578, wet kerangas”, slide No. BM Noct. 17544.
Paratype: 1 w, Brunei, 15 m, Telisai, Sandy Heath Forest and Gymnostoma 13.12.1979, leg. R. FaiRCLougH, slide No. LGN 1278
(BM Arct. 6089) (coll. BMNH).
Distribution. The species is known from Borneo.
14
Remarks. The w genitalia of L. wilsonae have remained undescribed in the original description. The w paratype
has been dissected during this study and its genitalia are characterised in detail in the diagnosis of the new genus
(see the Fig. 10).
Dialithoptera gemmata (Hampson, 1896)
(Pl. 1, g. 21, Pl. 2, g. 1)
Pisara gemmata Hampson, 1896, Fauna of British India, Moths 4: 506, by original designation. Type-locality: [India] Sikkim. Holo-
type: x, in coll. BMNH.
Material examined:
Thailand, Prov. Chiang Mai: 4 specimens, between Chiang Dao & Kariang, 900 m, 98º48’E, 19º25’N, 08.xi.2002; 5 specimens,
same site, 26.x.2002; 1 specimen, Mok Fa Garden Resort, 450 m, 98º48’E, 19º06’N, 1.xi.2002, leg. b. HeRCzig & g. Ronkay (coll.
G. Ronkay and MWM).
Dialithoptera margaritha LászLó, Ronkay & Witt, 2007
(Pl. 1, gs 19-20)
Dialithoptera margaritha LászLó, Ronkay & Witt, 2007, Entomofauna 28(2): 17-32. Type locality: N. Thailand, Prov. Mae Hong Son,
Pa Pae. Holotype: x, in coll. HNHM.
Type material examined:
Holotype. x, „North Thailand, Prov. Mae Hong Son, 1250 m, between Pa Pae and Khun Sa, 98°39’E, 19°08’N, 31.x.2002, leg. b.
HeRCzig et g. Ronkay”, slide No. LGN 776 (coll. HNHM).
Paratypes. Thailand. 1 w, with the same data as the holotype, slide No. LGN 906. Vietnam. 1 w, Mt. Fan-si-pan, Sa Pa, 2400 m,
22°15’N, 103°46’E, 8-29.v.1993, leg. sinjaev & simonov, slide No. LGN 362 (W 8323) (coll. HNHM and MWM).
Casminola gen. nov.
Type species: Poecilonola chionobasis Hampson, 1901
Species content:
C. chionobasis (Hampson, 1901) comb. nov.
C. arminbecheri spec. nov.
C. pulchella (LeeCH, 1889) comb. nov.
C. yoshimotoi (inoue, 2000) comb. nov.
C. splendida spec. nov.
C. rubropicta spec. nov.
C. seminigra (Hampson, 1896) comb. nov.
C. johannstump spec. nov.
C. breviharpe spec. nov.
C. spinosa spec. nov.
Diagnosis. The species of the new genus is somewhat similar externally to the members of Calonola gen. nov.,
Alcanola, and certain species of Aquita (especially A. acontioides) due to the combination of the bright white ground
colour and the blackish ornaments in the outer and costal areas of the forewing. The new genus represents a rather
compact phyletic unit of the subfamily Nolinae, according to the apomorphic structures of the main genitalia.
The main external differences of the species of Casminola, in comparison with Calonola, Alcanola and Aquita, are
the signicantly smaller size (length of forewing: 5–8 mm, wingspan: 11–16 mm, the measures of Aquita, Calonola
15
and Alcanola always exceed 8 mm and 16 mm, respectively). The forewing markings of the new genus are similar
to those of several Nolinae groups; the forewing has dark brownish-blackish outer area and narrow but always
rather conspicuous pair of costal dashes or patches in most species.
The structure of the x genitalia is unique and coherent throughout the genus: a simplied genital capsule in all
characters, the typical features are the rather elongated, and the apically rounded valva, the well-developed, long
and narrow, more or less straight, apically pointed and sometimes hooked harpe situated in the middle of the valva
running parallel with the valval margins. The harpe of Calonola is much smaller, shorter, always strongly curved,
originating much closer to the base of valva than in Casminola; the harpe of Alcanola is directed towards costal
margin in right angle while the harpe of Casminola is always parallel with valval margins. The x genitalia of Aquita
acontioides differ from those of Casminola in several important characters, having much shorter and almost straight
uncus (that of Casminola is always considerably longer and more curved), basally rather broad and apically taper-
ing valva (the valval base is relatively narrow in Casminola and the valval margins are usually parallel, sometimes
slightly broadened) and differently shaped harpe. The harpe of A. acontioides is an elongated, well-sclerotized
and dentate basal crest with short, slightly arcuate apical process, while in Casminola the erect part of the harpe
is much longer in most cases, and the sclerotized basal plate is missing.
The w genitalia of the new genus are also easily distinguished from those of the related genera by the rather short
eighth segment and the infundibular, relatively long, sclerotized ostium bursae. The eighth segment is longer
in Calonola, Alcanola and Aquita acontioides and the ostium bursae is sclerotized but tubular in Alcanola and
Aquita acontioides while it is membranous and very short in Calonola. The signum bursae of Casminola are either
absent (C. chionobasis, C. arminbecheri, C. pulchella, C. splendida, C. yoshimotoi, and C. rubropicta) or forming
a well-developed, characteristically curved, reversed U-shaped, scobinate and dentate bar (C. seminigra and C.
johannstump). Alcanola lacks the signa while the signa of Aquita acontioides and Calonola are represented by
a pair of elongated, scobinate plates.
Casminola chionobasis (Hampson, 1901) comb. nov.
(Pl. 2, gs 4-5)
Poecilonola chionobasis Hampson, 1901, Annals and Magazine of Natural History 7: 178. Type-locality: [India, Himachal Pradesh]
Kangra valley. Holotype: x, in coll. BMNH.
Type material examined:
Holotype of Poecilonola chionobasis Hampson, 1901: x, „TYPE” (red ring type label), „Kangra valley, 4500 ft, June 1899, Dudgeon”,
slide No. LGN 856 (BM Noct. 18489) (coll. BMNH).
Additional material examined:
Thailand. Prov. Chiang Mai: 1 x, Mt. Doi Phahompok, 16 km NW of Fang, 2000 m, 18.xii.1998, leg. m. HRebLay, slide No. LGN
360 (W 7310) (coll. MWM). Pakistan. Margalla Hills: 1 w, Islamabad, Pir Sohawa, 33º48’56”N, 73º08’01”E, 1300 m, 29.vii.1994,
slide No. LGN 392 (coll. HNHM); 2 ww, 20 km N of Islamabad, Pir Sohawa, 72º55’E, 33º50’N, 8.vii.1994, leg. b. HeRCzig, gy.m.
LászLó & g. Ronkay (coll. MWM).
Casminola arminbecheri spec. nov.
(Pl. 2, g. 6; gen g. 11)
Holotype. x, „Thailand, Changwat Nan, 25 km N of Bo Luang, 1150 m, 11.xi.1999, leg. Márton HRebLay”, slide No. LGN 358 (W
7311) (coll. MWM).
Paratypes. Thailand, Prov. Nan: 1 x, 4 km W of Pha Lak, 750 m, 100º34’E, 19º21’N, 5.xi.2002, leg. b. HeRCzig & g. Ronkay, slide
No. LGN 1232. Indonesia. Sumatra: 2 xx, Prapat HW 2, 17.ix.1985, slide Nos LGN 439, LGN 444; 2 xx, same
site, but collected at 8.vi.1985, and 29.iii.1986, slide Nos LGN 443, LGN 445; 1 x, Medan vic., Tiga Dolok,
21.x.1989, slide No. LGN 1247, leg. DR. DieHL (coll. HNHM, G. Ronkay and W. speiDeL).
Diagnosis. The new species shows close relationship with C. chionobasis and C. yoshimotoi, but is easily distinguish-
able from its relatives by its conspicuously narrower dark brown outer area of the forewing and the characteristically
thinner, oblique, dash-like pair of spots at the costal margin. The costal spots of C. chionobasis are large, rounded
16
and usually fused; those of C. yoshimotoi are broader and more triangular.
The x genitalia of the new species are easily separable from C. chionobasis by their broader and much longer,
distally dilated valva, somewhat longer, apically more hooked harpe, considerably longer, more strongly curved
uncus and somewhat longer vinculum. The aedeagus of C. arminbecheri is very similar in shape to that of C.
chionobasis but the vesica lacks the cornuti eld while C. chionobasis has a rather small, but well-dened cornuti
eld consisting of very short and ne cornuti. The differences between the x genitalia of C. arminbecheri and C.
yoshimotoi are comparatively smaller, but the valva of the new species is apically much more dilated and more
broadly rounded, the uncus longer and medially more curved and the vinculum is conspicuously longer and more
robust than in C. yoshimotoi. w unknown.
Distribution. The known range of the species is disjunct: it has been recorded from Thailand and from Sumatra;
the real distribution is supposedly much larger and continuous throughout the Malayan peninsula towards the
north-westernmost large island of Indonesia.
Casminola pulchella (LeeCH, 1889) comb. nov.
(Pl. 11, g. 5)
Acontia pulchella LeeCH, 1889, Proceedings of the Zoological Society London 1889: 524, pl. 53, g. 10. Type-locality: [Japan]
Ohoyama. Holotype: x, in coll. BMNH.
Type material examined:
Holotype of Acontia pulchella LeeCH, 1889: x, „TYPE” (red ring type label), „Japan, H. pRyeR coll.”; slide No. BM Arct. 1837 (coll.
BMNH).
Additional material examined:
Korea. 1 w, Korea I.A.S., 2.viii.1986, slide No. LGN 390 (coll. HNHM).
Remarks. The species has not yet been recorded from Thailand.
Casminola yoshimotoi (inoue, 2000) comb. nov.
(Pl. 11, g. 6)
Rhynchopalpus yoshimotoi inoue, 2000, Transactions of the Lepidopterological Society of Japan 51(4): 252, gs 2-3, 7-8. Type-
locality: Taiwan, Hotso (= Lushan Spa), Nantou Hsien. Holotype: x, in coll. BMNH.
Distribution. The species was described from Taiwan and is presumably endemic to this island.
Casminola splendida spec. nov.
(Pl. 11, gs 8-9; gen g. 14)
Holotype. x, „Upper Burma: Hpimaw Fort, nr. Myitkyina 8000 ft., Capt. A. E. Swann. Brit. Mus. 1923-488.”, slide No. LGN 1329
(BM Arct. 6204) (coll. BMNH).
Paratypes. Burma [Myanmar]. 5 xx, 1 w, with the same data as the holotype, slide Nos LGN 1348 (BM Arct. 6205) (x), LGN 1330
(BM Arct. 6206) (w) (coll. BMNH). Vietnam. Mt. Fan-si-pan, Sa-Pa: 1 w, 22º17’N, 103º44’E, 20-30.iv.1995, leg.
v. sinjaev & local collector, slide No. LGN 359 (W 7312); 1 w, 16-1800 m, 22º20’N, 103º40’E, 10.vi.-6.vii.1994,
leg. A. sCHintLmeisteR, slide No. LGN 1234 (W 7313) (coll. MWM).
Diagnosis. The new species resembles externally C. pulchella but is easily distinguishable from the latter taxon
by its more shiny white forewing with much broader and more unicolorously brown terminal area, and the more
17
sharply dened and darker brownish costal patches. The postmedial line of C. splendida is similarly sinuous as in
C. spinosa and C. johannstump, but is considerably broader and more diffuse.
The x genitalia of C. splendida differ conspicuously from those of C. pulchella by their much longer valvae, more
basally located and apically less hooked harpe, and the presence of a large eld of numerous ne cornuti in the
vesica; from those of C. spinosa mainly by the armature of the vesica. The large eld of cornuti consisting of a large
number of ne, short and weak spinules is a prominent autapomorphy of C. splendida; in comparison, C. pulchella
lacks the cornuti eld, C. spinosa has a rather large and elongated cornuti eld built up from some dozens of relatively
large, long and strongly sclerotized cornuti, and C. johannstump has a few, ne, but well-developed cornuti.
The w genitalia of C. splendida display closer relationship with C. pulchella, according to their similarly sclerotized
ductus bursae, large and swollen cervix bursae and the absence of the signum bursae. The new species has,
however, longer, funnel-like sclerotization in the ostial section of ductus bursae, shorter ductus bursae and a more
elongated corpus bursae.
Distribution. The species has been found in two small areas in Indochina, in northern Myanmar and the northern-
most mountainous area of Vietnam.
Casminola rubropicta spec. nov.
(Pl. 11, gs 11-12; gen g. 16)
Holotype. w, „Vietnam, Lao Cai, prov. Sa Pa district, Cat-Cat village, Frontier Vietnam Base Camp, 22º19’36,4”N, 103º49’46,1”E,
1250m, 4-8.viii.1998, leg. a. kun”, slide No. LGN 1233 (coll. HNHM).
Paratype. Thailand. 1 w, Mae Hong Son, 1240m, Pai District, Doi Mae Ya, 18.vii.1990, i.J. kitCHing & a.m. Cotton BM. 1992-19,
slide No. LGN 1343 (BM Arct. 6207) (coll. BMNH).
Diagnosis. Casminola rubropicta has a very particular external appearance within the genus with its pale brown-
ish mixed red forewing ground colour and a single, very narrow, elongated blackish costal patch, while all other
Casminola species have bright white forewings with a pair of broader and less elongated costal patches. The
marginal area of the new species is much darker than in the other Casminola species, it is almost black, and the
tornal area is marked by a triangular blackish patch, which is absent on all other Casminola.
The conguration of the w genitalia of C. rubropicta supports its placement into Casminola, the shape of oviposi-
tor, the funnel-like sclerotized ostium bursae, the rather large, swollen cervix bursae and the absence of signum
bursae are typical features of the genus. The ovipositor and the 8th segment are almost identical in C. rubropicta,
C. chionobasis and C. splendida, the main distinctive features of the three species are as follows: C. rubropicta has
the shortest apophyses anteriores, and sclerotized funnel-like ostium bursae and the largest and swollen cervix
bursae. The ostium bursae of C. chionobasis is only slightly longer than in C. rubropicta, while that of C. splendida
is about twice as long as in the other two taxa. The shape of corpus bursae of the new species is similarly elongated
as in C. splendida, while that of C. chionobasis is more globular. The x is unknown.
Distribution. The species is found only twice, in Vietnam and in Thailand.
Casminola seminigra (Hampson, 1896) comb. nov.
(Pl. 11, g. 7)
Pisara seminigra Hampson, 1896, Fauna of British India, Moths 4: 505. Type-locality: Bhutan. Holotype: w in BMNH.
Type material examined:
Holotype of Pisara seminigra Hampson, 1896: w, „TYPE” (red ring type label), „Bhutan, 95-37, 2.vii.94” (coll. BMNH).
Additional material examined:
18
Nepal. East-Nepal, Milke Danda: 1 x, Gursa, 2100 m, 22.viii.2000, slide No. LGN 186 (W 7314); 1 x, 2 ww, Nesum, 1500 m,
21.viii.2000, leg. T. Csővári & m. Hreblay, slide No. LGN 187 (W 7315) (w). Annapurna Himal: 1 x, 1 w, Geirigan village, 1340
m, 83º45’E, 28º20’N, 31.v.1996, slide No. LGN 353 (x); 1 x, from the same site, 26.vii.1995, leg. gy.m. LászLó & g. Ronkay, slide
No. LGN 1231. Syangja district: 1 x, 1 w, Kahule village, 1600 m, 26.vii.1995, leg. g. CsoRba, slide No. LGN 354 (w) (coll. HNHM
and MWM).
Distribution. The species is known from Nepal and Bhutan; it has no record from Thailand.
Casminola johannstump spec. nov.
(Pl. 2, g. 2; gen g. 12)
Holotype. x, „Nepal, Annapurna Himal, Geirigan village 1340 m, 83º45’E, 28º20’N, 26.vii.1995, leg. Gy.M. LászLó & g. Ronkay”,
slide No. LGN 355 (coll. G. Ronkay).
Paratypes. Nepal. Tanahoun district: 1 w, Baisakhe Ghat, 10 km W Dulegounda, 630 m, 10.x.1994, leg. CsoRba & Ronkay, slide No.
LGN 356 (W 7378). Syangja district: 1 w, 2 km E Syangja, 1200 m, 22.vii.1995; 1 x, Saldanda village, Bhaincy
Pani, 1100 m, 22.vii.1995, leg. g. CsoRba. Annapurna Himal: 1 w, 1 km S Bahundanda, 1000 m, 6.vi.1996,
84º25’E, 28º20’N, leg. m. HRebLay & Cs. szabóky, slide No. LGN 412 (W 7379) (coll. MWM). Indonesia. Sumatra:
1 x, Dolok Merangir, 26.xii.1984; leg. DR. DieHL, slide No. LGN 438 (coll. HNHM). Thailand. 1 x, Chaiyaphum,
650 m, Phu Khieo WS, km 7, Sala Prom checkpoint, 4/9.xi.1991, i.J. kitCHing & a.m. Cotton, BM 1992-9, slide
No. LGN 1327 (BM Arct. 6208) (coll. BMNH).
Diagnosis. Casminola johannstump is closely related to C. seminigra. The forewing pattern of the two species
are rather similar, the distinctive feature is as follows: the dark brown forewing marginal area of the new species is
much narrower, covering the outer fth of the wing, that of C. seminigra is much broader, usually the entire outer
half of the wing is darkened and the dark area is fused with the outer costal spot.
The x genitalia of C. johannstump differ from that of C. seminigra by the shorter, narrower, slightly curved uncus,
shorter, narrower, apically less dilated and rounded valva, much thinner, apically more tapering harpe and the
much shorter fultura inferior having thinner arms. The uncus of C. seminigra is much thicker, somewhat ag-like,
the valva is longer, broader, distally more dilated, the harpe is considerably thicker, less cuneate, and the fultura
inferior is longer with stronger lateral arms. The aedeagus of the two related species are similar, but its apical end
is obliquely truncate in C. johannstump, while elongate-triangular in C. seminigra, and the cornuti in the vesica
are somewhat ner in the new species.
In the w genitalia, the signum bursae of C. johannstump and C. seminigra differs conspicuously: that of the new
species is rather short, slightly curved, with very ne scobination, while rather long, reversed U-shaped, strongly
scobinated in C. seminigra; in addition the new species has, in comparison with that of C. seminigra, somewhat
shorter, funnel-like sclerotized ostium bursae.
Distribution. The new species is found in certain areas of the Nepal Himalaya at lower altitudes, in northern Thai-
land and in Sumatra.
Casminola breviharpe spec. nov.
(Pl. 2, g. 3; gen g. 13)
Holotype. x, „East-Nepal, Milke Danda, Nesum, 1500 m, 21.viii.2000, leg. Csővári & Hreblay”, slide No. LGN 357 (W 7380) (coll.
MWM).
Diagnosis. The new species is externally very similar to C. johannstump, but has somewhat shorter and narrower
costal patches on the forewing. The x genitalia of the two species display conspicuous differences: C. breviharpe
has much shorter, more robust, almost straight uncus, much shorter and thicker harpe, and somewhat shorter
valva. The uncus of C. johannstump is comparatively longer, thinner and slightly curved, the valva is longer, and
the harpe is much longer and thinner than in the related species. The aedeagus is similarly shaped in the two
species but the vesica of the new species lacks the cornuti eld while C. johannstump has ne, but clearly visible
19
cornuti eld. The w is unknown.
Distribution. The species is known by its unique type specimen from Nepal, it has not been recorded from Thai-
land.
Casminola spinosa spec. nov.
(Pl. 11, g. 10; gen g. 15)
Holotype. x, „NW Thailand: 1460m, Chiang Mai, Doi Suthep-Pui NP, 26.iv-10.v.1989 a.m. Cotton, BM 1989-145”, slide No. LGN
1328 (BM Arct. 6209) (coll. BMNH).
Diagnosis: The forewing pattern of C. spinosa resembles C. johannstump, but the ground colour is much more
shining white, and the darker markings are sharply dened by a thin, but blackish line, which is absent in C. johan-
nstump. The new species is also similar to C. splendida by its shining white forewing ground colour, but is much
smaller in size (the wingspan of C. spinosa is 14 mm, with the length of forewing 6 mm, those of C. splendida are
15-18 mm and 7-9 mm, respectively) and the forewing is conspicuously narrower.
The specic autapomorphy of the x genitalia of C. spinosa is the long and narrow cornuti eld of the vesica consist-
ing of a large number (ca 80) of relatively long, strong, acute spines. The other species of the genus have either
smaller cornuti eld of smaller, weaker spinules or the cornuti eld is fully absent. The conguration of the clasping
apparatus of the C. spinosa is most similar to C. johannstump, but the valvae are slightly longer and the harpe is
situated closer to the valval base than in its close relative. The w is unknown.
Distribution. The new species is known from the type-locality only.
Porcellanola LászLó, Ronkay & Witt, 2006
Type species: Porcellanola minna LászLó, Ronkay & Witt, 2006
Species content:
P. minna LászLó, Ronkay & Witt, 2006
P. chakri LászLó, Ronkay & Witt, 2006
P. lanna LászLó, Ronkay & Witt, 2006
P. sukhothai LászLó, Ronkay & Witt, 2006
P. thonburi spec. nov.
P. thai LászLó, Ronkay & Witt, 2006
P. ayutthaya spec. nov.
P. langtangi spec. nov.
P. uthong spec. nov.
Distribution. Porcellanola langtangi is the only known species of the genus Porcellanola found outside of the
mountains of Northern Thailand, it was discovered in the Langtang Himal in Nepal. The presence of the genus
in the Central Himalayan massif suggests that further, yet undiscovered members of the genus may occur in the
chains between the eastern Himalayas and the northern mountain regions of Indochina.
Porcellanola minna LászLó, Ronkay & Witt, 2006
(Pl. 2, gs 7-8)
Porcellanola minna LászLó, Ronkay & Witt, 2006, Entomofauna 27(21): 268. Type locality: N. Thailand, Prov. Nan, Pua. Holotype:
x, in coll. MWM.
Type material examined:
20
Holotype. x, „Thailand, Changwat Nan, 30 km E of Pua, 1700 m, 13. XI. 1999, leg. Márton HRebLay”, slide No. LGN 882 (W 8297)
(coll. MWM).
Paratypes. Thailand. Prov. Nan: 1 x, from the same site as the holotype, 10.xi.1999, slide No. LGN 886 (W 8301). Prov. Chiang
Mai: 1 x, 3 ww, Mt. Doi Phahompok, 18 km NW of Fang, 2100 m, 15.xi.1999, slide Nos LGN 883, (W 8298) (x), LGN 884 (W8299),
LGN 885 (W 8300) (ww), leg. m. HRebLay (coll. MWM).
Porcellanola chakri LászLó, Ronkay & Witt, 2006
(Pl. 2, gs 9-10)
Porcellanola chakri LászLó, Ronkay & Witt, 2006, Entomofauna 27(21): 272. Type locality: N. Thailand, Prov. Chiang Mai, Fang.
Holotype: x, in coll. HNHM.
Type material examined:
Holotype. x, „North Thailand, prov. Chiang Mai, 1600 m, between Fang and Nor Lae, 99°06′E, 20°02′N, 12. 11. 2002, leg. b.
Herczig & G. Ronkay”, slide No. LGN 887 (coll. HNHM).
Paratypes. Thailand. Prov. Chiang Mai: 1 x, with the same data as the holotype; 1 x, same site, between Fang and Nor Lae, 1600
m, 99º06’E, 20º02’N, 7.xi.2002, leg. b. HeRCzig & g. Ronkay. Prov. Mae Hong Song: 1 x, 1250 m, between Pa Pae and Khun Sa,
98°39′E, 19°08′N, leg. b. HeRCzig & g. Ronkay, slide No. LGN 880 (W 8303). Prov. Nan: 1 x, 30 km E of Pua, 1700 m, 10.xi.1999,
leg. m. HRebLay, slide No. LGN 196 (W 8309) (coll. HNHM and MWM).
Porcellanola lanna LászLó, Ronkay & Witt, 2006
(Pl. 2, gs 11-12)
Porcellanola lanna LászLó, Ronkay & Witt, 2006, Entomofauna 27(21): 272. Type locality: N. Thailand, Prov. Chiang Mai, Doi
Phahompok. Holotype: x, in coll. MWM.
Type material examined:
Holotype. x, „Thailand, Changwat Chiang Mai, Mt. Doi Phahompok, 18 km NW of Fang, 2100 m, 7.ii.2000, leg. Hreblay & Szabó”,
slide No. LGN 881 (W 8302) (coll. MWM).
Paratypes. Thailand. Prov. Chiang Mai: 1 x, with the same data as the holotype, slide No. LGN 889 (W 8304); 1 x, 20 km NW
of Mae Ai, 1650 m, 26.i.1999, leg. a. szabó & z. CzeRe, slide No. LGN 163 (W 8305); 1 w, Mt. Doi Phahompok, 16 km NW of
Fang, 2000 m, 24.ii.1998, leg. m. HRebLay & Cs. szabóky, slide No. LGN 915 (W 8365); 2 ww, 20 km NW of Fang, 2100 m, 24. I.
2004; 1 w, same site, 2150 m, 22. I. 2004, leg. a. szabó (coll. MWM).
Porcellanola sukhothai LászLó, Ronkay & Witt, 2006
(Pl. 2, gs 13-14)
Porcellanola sukhothai LászLó, Ronkay & Witt, 2006, Entomofauna 27(21): 273. Type locality: N. Thailand, Prov. Chiang Mai, Doi
Phahompok. Holotype: x, in coll. MWM.
Type material examined:
Holotype. x, „Thailand, Changwat Chiang Mai, Mt. Doi Phahompok, 18 km NW of Fang, 2100 m, 15. XI. 1999, leg. Márton HRebLay”,
slide No. LGN 891 (W 8306) (coll. MWM).
Paratypes. Thailand. Prov. Nan: 2 xx, 30 km E of Pua, 1700 m, 10.xi.1999, leg. m. HRebLay, slide Nos: LGN 195 (W 8308), LGN
892 (W 8307) (coll. MWM).
21
Porcellanola thonburi spec. nov.
(Pl. 2, g. 15; gen. g. 17)
Holotype. x, „Thailand, Changwat Mae Hong Song, 1 km S of Bahudanda, 1000 m, 6. II. 1998, leg. Márton HRebLay & Csaba
szabóky”, slide No LGN 1134 (W 7326) (coll. MWM).
Diagnosis. Porcellanola thonburi has a rather unique external appearance within the genus, according to its long
and narrow, apically pointed forewing with orange-yellow ground colour and dark costal and marginal areas. De-
spite its unusual external appearance, the new species inevitably show the typical x genital features of the genus
Porcellanola; its closest relative is P. sukhothai. The distinctive character of the x genitalia is the conguration of the
aedeagus: the new species has very long, narrow, stick-like carina, while P. sukhothai has shorter, strongly dentate
carinal plate. In addition, the valva of P. thonburi is somewhat shorter than in P. sukhothai. The w is unknown.
Distribution. Northern Thailand.
Porcellanola thai LászLó, Ronkay & Witt, 2006
(Pl. 2, gs 16-17)
Porcellanola thai LászLó, Ronkay & Witt, 2006, Entomofauna 27(21): 274. Type locality: N. Thailand, Prov. Mae Hong Son, Pa
Pae. Holotype: x, in coll. HNHM.
Type material examined:
Holotype. x, „North Thailand, Prov. Mae Hong Son, 1250 m, between Pa Pae and Khun Sa, 98°39′E, 19°08′N, 31.x.2002, leg. b.
HeRCzig et G. Ronkay”, slide No. LGN 888 (coll. HNHM).
Paratypes. Thailand. Prov Mae Hong Song: 3 xx, 1 w, with the same data as the holotype, slide Nos: LGN 890 (x), LGN 905
(w). Prov. Chiang Mai: 3 xx, between Fang and Nor Lae, 1600 m, 99°06′E, 20°02′N, 12.xi.2002, leg. b. HeRCzig & g. Ronkay
(coll. HNHM and MWM).
Porcellanola ayutthaya spec. nov.
(Pl. 2, gs 18-19; gen. g. 18)
Holotype. x, „Thailand, Changwat Chiang Mai, 4 km SE of Pang Faen, 1100m, 26. I. 2004, leg.: Attila Szabó”, slide No. LGN
1039 (W 7288) (coll. MWM).
Paratypes. Thailand. Prov. Mae Hong Song: 1 w, 21 km NW of Pai, 1360 m, 7.ii.1998, slide No. LGN 462 (W 7289). Prov. Chiang
Mai: 1 w, Mt. Doi Inthanon, 10 km E of Mae Chaem, 1160 m, 5.ii.1998, slide No. LGN 461 (W 7290); 2 xx, 1600
m, between Fang and Nor Lae, 99˚06’E, 20˚02’N, 12.xi.2002, leg. b. HeRCzig & g. Ronkay, slide Nos LGN 1228,
LGN 1229. Prov. Nan: 1 w, 30 km E of Pua, 1700 m, 27.ii.1998, slide No. LGN 832 (W 7291), leg. m. HRebLay &
Cs. szabóky (coll. G. Ronkay and MWM); 1 x, Doi Phu Kha, Pua, 1680 m, 18.ii.1993, leg. W. speiDeL (coll. speiDeL);
1 x, Doi Suthep-Pui NP, 1380 m, 24.ii.1989, leg. a.m. Cotton & i.J. kitCHing BM 1989-57 (coll. BMNH).
Diagnosis. The new species is much smaller in size than Porcellanola thai LászLó, Ronkay & Witt, 2006 (wingspan
14-16 mm; length of forewing 7-8 mm, while those of P. ayutthaya are 11-12 mm and 5-6 mm, respectively), hav-
ing narrower and more acute forewing with mentionably less contrasting and differently shaped black and white
markings. The genitalia of both sexes clearly show the relationship of the two species although the differences are
easily recognisable. The new species has, in comparison with P. thai, slightly broader uncus, conspicuously nar-
rower valva and somewhat longer fultura inferior, lacking the ventro-medial process. The harpe-ampulla complex
is pincer-like in both species, but the apical processes of this complex are signicantly shorter and almost straight
in the new species while they are longer and more curved in P. thai; the ampullar process of P. ayutthaya is longer
than the harpe, the situation is just the opposite in P. thai. The aedeagi of the two species are surprisingly similar
but the whole apparatus of P. ayutthaya is somewhat shorter with less serrate dorso-lateral bar of the carina.
22
The w genitalia of the two taxa are very similar, but the new species has somewhat shorter and less sclerotized,
quadrangular ostium bursae and longer ductus bursae. The ostium bursae of P. thai is comparatively longer, more
sclerotized and infundibular, and the ductus bursae is shorter than in P. ayutthaya.
Distribution. All known localities of the new species lie in northern Thailand.
Porcellanola langtangi spec. nov.
(Pl. 2, gs 20-21; gen. g. 19)
Holotype. x, „Nepal, Langtang, 1950m, 1,5 km NE Dhunche, 85º18’E, 28º06’N, 24.ix.1994, leg. G. CsoRba & L. Ronkay”, slide
No. LGN 460 (coll. HNHM).
Paratype. 1 w, with the same data as the holotype, slide No. LGN 1227 (coll. HNHM).
Diagnosis: Porcellanola langtangi is an allopatric sister species of P. ayutthaya. The external appearance of the
two taxa is very similar, the mentionable differences are the more oblique inner margin of the dark outer half of the
forewing and the longer and broader dark costal patch of P. langtangi. The satisfactory separation of the two species
requires the examination of their genitalia which display surprisingly large and prominent differences. Comparing
the x genitalia of the two species, P. langtangi has much broader valva (similar to that of P. thai), elongated-trian-
gular fultura inferior with pointed distal apex (it is rather quadrangular in P. ayutthaya, with slightly concave distal
margin); much longer, slightly curved and apically pointed harpe (it is a very short, apically rounded process in
P. ayutthaya), considerably shorter and extremely thin, apically pointed, thread-like ampulla (it is a much thicker,
slightly curved, distally tapering and apically rounded process in P. ayutthaya), shorter and broader vinculum and
conspicuously narrower aedeagus with much shorter and less serrate dorso-lateral bar.
The synapomorphies of the w genitalia of the two species are the similar conguration of the ovipositor and corpus
bursae with the characteristic pair of nely scobinated ovoid signum-plates. The distinctive features of P. langtangi
are the considerably longer and narrower ventral plate of the penultimate segment, the much narrower ostium
bursae, and the somewhat longer and much slenderer ductus bursae, which is more sclerotised distally than the
otherwise much broader but weaker ductus bursae of P. ayutthaya.
Distribution. The new species is found only once in the Langtang Himal in Central Nepal. The taxon is most prob-
ably an allopatric sibling species of P. ayutthaya, its occurence in Thailand is, therefore, less probable.
Porcellanola uthong spec. nov.
(Pl. 3, gs 1-2; gen. g. 20)
Holotype. x, „North Thailand, Prov. Chiang Mai, 1800m, 4 km S of Kop Dong, 99º03’E, 19º52’N, 06.11.2002, leg. B. Herczig et
G. Ronkay”, slide No. LGN 1073 (W 7292) (coll. MWM).
Paratype. Thailand. Prov. Mae Hong Son: x, between Pa Pae and Khun Sa, 1250 m, 98º39’E, 19º08’N, 31.x.2002, leg. b. HeRCzig
& g. Ronkay, slide No. LGN 1074 (W 7295) (coll. MWM).
Diagnosis. The new species belongs, despite the rather unique external appearance, to the Porcellanola thai spe-
cies-group, this relationship is indicated by the structure of the x genitalia. Porcellanola uthong differs externally
from all other Porcellanola species by the ground colour of the forewing: it is the only known member of the genus
having graphite-grey suffused forewings in contrast to the porcelain-white ground colour of the other species.
The x genitalia of the new species display closer relationship with P. sukhothai LászLó, Ronkay & Witt, 2006, the
differences between the two species are as follows: the fultura inferior of P. uthong is much more complex, it is a
larger, sclerotised, dorso-medially deeply incised plate with strong, robust, thorn-like medial process originating
from the ventral edge and with a pair of short, medio-lateral crests. The fultura inferior of P. sukhothai is a more
23
simple and shorter, quadratic plate with a short ventro-medial fold only. The ampulla of P. uthong is bifurcate, with
rather robust, claw-like distal and shorter proximal processes, the ampulla of P. sukhothai is a simple, claw-like
process; the erect part of the harpe of P. uthong is a very ne prominence only.
Distribution. The new species occurs in northern Thailand.
Calonola gen. nov.
Type species: Zia ectrocta Hampson, 1907
Species content:
C. ectrocta (Hampson, 1907) comb. nov.
C. argyria (Hampson, 1894) comb. nov.
Diagnosis. The external appearance of the members of the new genus resembles somewhat the overwhelming
majority of the species of Porcellanola due to their shining white forewing ground colour with brownish terminal
area and the characteristic triangular costal patch. The costal patch of Calonola is larger and darker than in Porcel-
lanola and the terminal area has more sharply dened bordering lines. The diagnostic generic characters are found
in the conguration of the x genitalia. The copulatory organs of Calonola and Porcellanola differ conspicuously:
the valva of Calonola is much more elongated, the harpe is usually smaller, originating much closer to the valval
base than in Porcellanola, the fultura inferior is rather small, poorly sclerotized, while this structure of Porcellanola
is always well-developed, sometimes with large arms. The vinculum of Calonola is rather quadrangular with a
narrow, elongate medial process, that of Porcellanola is simply U- or V-shaped. The w genitalia of the two genera
are also easily distinguishable by the presence of a pair of elongated, scobinate signum-stripes of Calonola, while
Porcellanola lacks the signa.
The structure of the x genitalia of Calonola indicates its closer relationship with the genus Barasa, despite the rather
dissimilar external appearance of their species; their shared features are the elongated, slightly curved valva and
the usually small, arcuate, claw-like harpe. The main difference between the x genitalia of Calonola and Barasa is
the characteristic shape of the valva which is simple and undivided in Calonola while it is divided into two (ventral
and costal) parts in Barasa. The w genitalia of the two genera are also easily recognisable: Barasa lacks the signa
while Calonola has two well-developed signum-stripes.
Calonola ectrocta (Hampson, 1907) comb. nov.
(Pl. 1, gs 12-13; gen. g. 21)
Zia ectrocta Hampson, 1907, Annals and Magazine of Natural History 7(19): 229. Type-locality: [Sri Lanka] Ceylon, Haputale.
Holotype: x, in coll. BMNH.
Type material examined:
Holotype of Zia ectrocta Hampson, 1907: x, „TYPE” (red ring type label), „Ceylon, W-Haputale II.03. Mackwood 1904.4.”, slide No.
LGN 722 (BM. Noct. 18202) (coll. BMNH); holotype of Zia ectrocta ab. ectroctoides Strand (manuscript name): x, „Maskeliya,
Ceylon, May, de Moulray 1906. 181”, slide No. LGN 723 (BM Noct. 18203) (coll. BMNH).
Additional material examined:
Thailand. Prov. Chiang Rai: 1 x, 1 km SE of Khun-Kon Waterfall, 600 m, 15.xi.1998, leg. T. Csővári & l. mikus, slide No. LGN 156
(W 7327). Prov. Nan: 1 w, 25 km N of Bo Luang, 1150 m, 3.ii.2000, leg. M. HRebLay & a. szabó, slide No. LGN 893 (W 7328) (coll.
MWM). Prov. Chiang Mai: 1 specimen, between Fang and Nor Lae, 99º06’E, 20º02’N, 28.x.2002, leg. b. HeRCzig & g. Ronkay
(coll. G. Ronkay).
24
Calonola argyria (Hampson, 1894) comb. nov.
(Pl. 1, gs 14-15)
Pisara argyria Hampson, 1894, Fauna of British India, Moths 2: 145. Type-locality: [India] Sikkim. Holotype: x, in coll. BMNH.
Type material examined:
Holotype of Pisara argyria Hampson, 1894: x, „TYPE” (red ring type label), „Sikkim”, slide No. BM Arct. 1651 (coll. BMNH).
Additional material examined:
Thailand. Prov. Chiang Mai: 8 specimens, Mt. Doi Phahompok, 16 km NW of Fang, 2000 m, 24.ii.1998 leg. m. HRebLay & Cs.
szabóky; 2 specimens, same site, 14.viii.1999; 1 specimen, same site, 6-7.viii.1999, leg. T. Csővári & l. mikus; 2 specimens, same
site, 10.i.1999; 1 specimen, same site, 12.ix.1999, leg. a. szabó & z. CzeRe; 1 specimen, same site, 27.iii.1998, leg. T. Csővári &
p. stégeR; 1 specimen, Mt. Doi Phahompok, 17 km NW of Fang, 2100 m, 15.viii.1999, leg. T. Csővári & l. mikus; 4 specimens, Mt.
Doi Phahompok, 18 km NW of Fang, 2100 m, 25.ii.1998, leg. m. HRebLay & Cs. szabóky, slide No. LGN 165 (W 7329) (x); 2 speci-
mens, same site, 19-20.ix.1999; 2 specimens, same site, 18.i.1999, leg. a. szabó & z. CzeRe; 2 specimens, Mt. Doi Phahompok,
19 km NW of Fang, 1900 m, 3.iv.1998 leg. T. Csővári & P. sTéger; 1 specimen, Mt. Doi Phahompok, 20 km NW of Fang, 2150
m, 28-29.i.2004; 5 specimens, same site, 22-25.i.2004, leg. a. szabó; 6 specimens, Mt. Doi Inthanon, NP, 2300 m, 7-8.ix.1999; 1
specimen, same site, 6.i.1999, leg. a. szabó & z. CzeRe; 1 specimen, same site, 16.i.2004, leg. p. HentsCHeL & a. szabó; 1 speci-
men, same site, 19-20.xi.1998, leg. T. Csővári & l. mikus; 1 specimen, 4 km S of Kop Dong, 99º03’E, 19º52’N, 1800 m, 11.xi.2002;
8 specimens, same site, 6.xi.2002, leg. b. HeRCzig & g. Ronkay; 3 specimens, 20 km NW of Sop Kha, 2 km S of Kop Dong, 1800
m, 21.ii.1998, leg. m. HRebLay & Cs. szabóky; 4 specimens, same site, 27.i.1999; 3 specimens, same site, 8.i.1999, leg. a. szabó &
z. CzeRe; 1 specimen, 20 km NW of Mae Ai, 1650 m, 7.i.1999, leg. a. szabó & z. CzeRe. Prov. Nan: 1 specimen, 30 km E of Pua,
1700 m, 24.xi.1998; 1 specimen, same site, 18.xi.1998, leg. T. Csővári & l. mikus; 3 specimens, same site, 18.ii.1998; 1 specimen,
same site, 20.ii.1998; 5 specimens, same site, 27.ii.1998; 1 specimen, same site, 1.iii.1998; 3 specimen, same site, 20.ii.1998, leg.
m. HRebLay & Cs. szabóky; 1 specimen, same site, 22-23.ix.1999, leg. a. szabó & z. CzeRe; 2 specimens, same site, 10.xi.1999; 4
specimens, same site, 13.xi.1999, leg. m. HRebLay; 2 specimens, same site, 20.i.2004, leg. p. HentsCHeL & a. szabó; 1 specimen,
25 km N of Bo Luang, 1150 m, 19.ii.1998, leg. m. HRebLay & Cs. szabóky (coll. G. Ronkay and MWM).
Barasa acronyctoides WaLkeR, 1862
(Pl. 3, g. 18)
Barasa acronyctoides WaLkeR, 1862, Proceedings of the Linnean Society (Zoology) 6: 192. Type-locality: Borneo, Sarawak. Type:
in coll. UM, Oxford.
Barasa cana Hampson, 1893, Illustrations of Typical Specimens of Lepidoptera Heterocera in the Collection of the British Museum
9: 107. Type-locality: [Sri Lanka] Ceylon, Halina valley. Holotype: x, in coll. BMNH.
Material examined:
Thailand. Prov. Chiang Mai: 1 w, between Chiang Dao and Kariang, 900 m, 98º48’E, 19º25’N, 26.x.2002, leg. b. HeRCzig & g.
Ronkay (coll. G. Ronkay).
Barasa lunisigna (Hampson, 1898)
(Pl. 3, g. 21, Pl. 4, g. 1)
Nola lunisigna Hampson, 1898, Journal of the Bombay Natural History Society, 11: 441. Type-locality: [India] [Meghalaya] Khasis.
Holotype: x, in coll. BMNH.
Type material examined:
Holotype of Nola lunisigna Hampson, 1898: x, „TYPE” (red ring type label), „Khasis”, slide No. BM. Noct. 17729 (coll. BMNH).
Additional material examined:
Thailand. Prov. Chiang Mai: 1 x, between Chiang Dao and Kariang, 900 m, 98º48’E, 19º25’N, 26.x.2002, leg. b. HeRCzig & g.
Ronkay (coll. G. Ronkay); 1 x, 4 km SE of Pang Faen, 1100 m, 31.i.2004; 1 x, same site, 26.i.2004, 3 xx, same site, 27.i.2004,
leg. a. szabó, slide No. LGN 932 (W 7331); 1 w, same site, 13.i.2004, leg. p. HentsCHeL & a. szabó, slide No. LGN 931 (W 7330); 1
25
x, same site, 6.ii.2000, leg. m. HRebLay, y. sHeRpani & a. szabó; 1 x, Mt. Doi Inthanon, NP, 2300 m, 17.x.2000, leg. local collector,
slide No. LGN 519 (W 7332) (coll. MWM). Prov. Nan: 1 x, Doi Phuka NP, 1350 m, between Pua and Bo Luang, 101º05’E, 19º12’N,
3.xi.2002, leg. b. HeRCzig & g. Ronkay, slide No. LGN 930 (coll. G. Ronkay); 1 x, 30 km E of Pua, 20.i.2004, leg. a. szabó. Prov.
Mae Hong Son: 1 w, 25 km NE of Pai, 1560 m, 2.iii.1998, leg. m. HRebLay & Cs. szabóky, slide No LGN 151 (W 7333); 1 x, 10 km
NE of Pai, 1560 m, 3.xii.1998, leg. m. HRebLay, i. soós & y. sHeRpani, slide No. LGN 956 (W 7334) (coll. MWM).
Barasa subtilis spec. nov.
(Pl. 3, gs 19-20; gen. g. 22)
Holotype. x, „N. Thailand, Prov. Nan, 5 km N of Ban Luang, 350m, between Pi Nai and Pi Tai 100º27’E, 18º56’N, 4.xi.2002, leg.
b. HeRCzig & g. Ronkay”, slide No LGN 1025 (coll. G. Ronkay).
Paratypes. Thailand. Prov. Chiang Mai: 1 x, 4 km SE of Pang Faen, 1100 m, 27.i.2004, leg. a. szabó, slide No. LGN 933 (W
7336); 1 x, 10 km NW of Fang, 550 m, 14.xi.1998, leg. T. Csővári & l. mikus, slide No. LGN 903 (W 7335).
Prov. Nan: 1 w, 5 km E of Bo Luang, 610 m, 23.xi.1998, leg. T. Csővári & l. mikus, slide No. LGN 1042 (W
7337) (coll MWM); 1 w, 20 km NE Nan, 21.ii.1993, leg. W. speiDeL (coll. speiDeL). Cambodia. 1 w, Mondolkiri
Prov., Seima Biodiversity Conservation Area, between Seima and O’Rang, 12º12’12”N, 107º01’09”E, 300 m,
30.i.2006 leg. g. CsoRba & g. Ronkay, slide No. LGN 1046 (W 7338) (coll. MWM).
Diagnosis. The new species is externally rather similar to B. lunisigna, but is somewhat smaller in size (length of
forewing of B. subtilis 9-10 mm, that of B. lunisigna 10-12 mm), and the forewing is darker with brownish-red suf-
fusion in the marginal area, while the ground colour of the forewing of B. lunisigna is uniformly pale grey.
The conguration of the x genitalia of B. subtilis differs conspicuously from that of B. lunisigna: the new species
has rather broad and rounded distal ventral lobe of the valva with well-developed apical hump, somewhat shorter,
thinner harpe, and much shorter, broader tegumen. The valva of B. lunisigna is rather narrow and very long, without
apical process; the harpe is longer, thicker and the tegumen is narrower than in B. subtilis.
The w genitalia of the two related species show also remarkable differences. The papillae anales of the new spe-
cies are more elongated, the apophyses anteriores are much longer, the cervix bursae is much narrower and the
corpus bursae is much larger in the new species than in B. lunisigna.
Distribution. The species was found in Thailand and Cambodia.
Barasa alopha Hampson, 1896
(Pl. 4, gs 2-3)
Barasa alopha Hampson, 1896, Fauna of British India, Moths, 4: 525. Type-locality: [India] [Maharasthra] Bombay, Tanna; [Sri
Lanka] Ceylon. Syntypes: in coll. BMNH.
Zia grisea Hampson, 1914, Catalogue of the Amatidae and Arctiidae (Nolinae and Lithosiinae) in the Collection of the British Museum
1914: 438, pl. 25, g. 6. Type-locality: China, Hupeh, Lui-Shin-Tze. Holotype: w, in coll. BMNH; syn. n.
Material examined:
Thailand. Prov. Chiang Mai: 1 specimen, between Chiang Dao and Kariang, 900 m, 98º48’E, 19º25’N, 26.x.2002; 1 x, between
Fang and Nor Lae, 99º06’E, 20º02’N, 12.xi.2002, leg. b. HeRCzig & g. Ronkay. Prov. Mae Hong Son: 1 w, 10 km NE of Pai, 1560 m,
3.xii.1998, leg. m. HRebLay, i. soós & y. sHeRpani, slide No. LGN 168 (W 7339) (coll. MWM). Prov. Nan: 2 specimens, Doi Phuka N.P.,
between Pua and Bo Luang, 1350 m, 101˚05’E, 19˚12’N, 3.xi.2002, leg. b. HeRCzig & g. Ronkay (coll. G. Ronkay and MWM).
26
Barasa costalis Hampson, 1895
(Pl. 4, gs 4-5)
Barasa costalis Hampson, 1895, Transactions of the Entomological Society of London 1895: 305. Type locality: [India] Sikkim,
Bhutan. Syntypes: in coll. BMNH, London.
Material examined:
Thailand. Prov. Chiang Mai: 1 specimen, 4 km SE of Pang Faen, 1100 m, 26.i.2004; 2 specimens, same site, 13.i.2004, slide No.
LGN 957 (W 7340) (x); 1 x, same site, 27.i.2004, leg. a. szabó, slide No. LGN 958 (W 7341) (coll. MWM).
Ezishnola gen. nov.
Type species: Ezishnola fuscographa spec. nov.
Species content:
E. fuscographa spec. nov.
Diagnosis. The species of this monotypical new genus resembles certain species of Barasa (e.g. B. lunisigna and
B. subtilis) and Suerkenola (S. longiventris) by their pale grey forewing ground colour, but the species of the new
genus lacks the strongly dened crosslines and the costal streak of forewing, which are characteristic for Barasa
and Suerkenola.
The x genitalia of the new genus is rather remote from any other genera of Nolinae having rather long, apically
rounded, nger-like uncus covered apically densely with hairs, the basally narrow, distally much broader, apically
rounded (somewhat ham-shaped) valva, and the characteristic harpe-ampulla complex. This complex consists of
strong and straight, thorn-like ampulla and rather short, spine-like erect part of harpe, resembling slightly those of
certain, externally dissimilar, species of Porcellanola (e.g. P. thai).
The w genitalia of the new genus are also characteristic: they have a pair of long and broad, sclerotized signa
bands. These signum stripes are somewhat similar to those of Barnanola, while the ovipositor, the ostium bursae
and the tubular part of corpus bursae are rather simple, displaying no autapomorphic features.
Ezishnola fuscographa spec. nov.
(Pl. 3, gs 3-4; gen. g. 23)
Holotype. x, „Thailand, Changwat Nan, 30 km E of Pua, 1700m, 20.viii.1999, leg. T. Csővári & l. mikus”, slide No. LGN 953 (W
7342) (coll. MWM).
Paratypes. Thailand. Prov. Nan: 1 w, 5 km E of Bo Luang, 610 m, 23.xi.1998, leg. T. Csővári & l. mikus, slide No. LGN 971 (W
7343); 1 w, Mt. Doi Phukha, 30 km E of Pua, 1700 m, 20.viii.1999; 1 w, same site, 10.xi.1999, leg. m. HRebLay
& i. soós; 1 w, Mt. Doi Phukha, 5 km E of Bo Luang, 610 m, 23.xi.1998, leg. T. Csővári & l. mikus (coll. mWm).
Prov. Chiang Mai: 1 x, between Chiang Dao and Kariang, 900 m, 98º48’E, 19º25’N, 26.x.2002, slide No. LGN
954 (W 7344); 1 x, same site, 8.xi.2002, leg. b. HeRCzig & g. Ronkay, slide No. LGN 1054; 1 x, Mt. Doi Inthanon,
NP, 2300 m, 17.x.2000, leg. local collector, slide No. LGN 1265 (W 7345); 3 xx, 3 ww, 12 km NW of Chiang
Dao, 750 m, 12.xi.1998; 1 w, 10 km NW of Fang, 550 m, 14.xi.1998, leg. T. Csővári & l. mikus (coll. MWM and
G. Ronkay). Uthai Thani District: 1 x, Khao Nang Rum, 400 m, 6-8.vi.1986, leg. m.g. aLLen (coll. BMNH).
Diagnosis. Ezishnola fuscographa resembles slightly certain taxa of Barasa (especially B. lunisigna and B. subtilis)
but the forewing is pale grey, lacking the blackish costal spot and the well-dened lower section of postmedial line
which are characteristic for the mentioned Barasa species. The new species is easily distinguishable from the
externally somewhat similar Barasa costalis and Suerkenola longiventris by its much smaller size and the absence
of the conspicuous apical streak of the forewing, this latter feature is typical of S. longiventris.
27
The diagnostic features of the genitalia of the new species are given in the diagnosis of the genus.
Distribution. The new species is distributed in the northern areas of Thailand.
Suerkenola gen. nov.
Type species: Nola longiventris pouJaDe, 1886 comb. nov.
Species content:
S. longiventris (pouJaDe, 1886) comb. nov.
Diagnosis. The genus is monotypical. The typical external features are the pale grey forewing ground colour (like
in certain Barasa species, e.g. B. costalis and B. lunisigna) and the sharply dened, dark grey, slightly arched
apical streak in the forewing termen.
The x genitalia of Suerkenola are rather unique within the entire subfamily, displaying no closer relationship with
the above-mentioned taxa. The diagnostic features are the presence of large, strongly sclerotized saccular lobe
with characteristically dentate inner margin, the hump-like process of the harpe, and the well-developed, narrow,
slightly curved ampullar process. The w genitalia are characterisable by the large ostium bursae with a pair of
strongly sclerotized medial plates.
Suerkenola longiventris (pouJaDe, 1886) comb. nov.
(Pl. 3, gs 5-6; gen. g. 24)
Nola longiventris pouJaDe, 1886, Bulletin de la Société Entomologique de France 6(6): 151. Type-locality: [China, Sichuan] Tibet,
Moupin. Holotype: x, in coll. MNHN Paris.
Barasa sufda sWinHoe, 1919, Annals and Magazine of Natural History (Series 9) 4: 120. Type locality: [India] [Meghalaya] Khasia
Hills. Holotype: x, in coll. BMNH; syn. n.
Material examined:
Thailand. Prov. Nan: 2 specimens, 30 km E of Pua, 1700 m, 22-23.ix.1999, slide No. LGN 137 x (W 7347); 3 specimens, same site,
14-15.ix.1999, leg. a. szabó & z. CzeRe; 1 specimen, same site, 16.xi.1998, leg. T. Csővári & l. mikus; 1 x, same site, 13.xi.1999,
leg. m. HRebLay & i. soós, slide No. LGN 951 (W 7348); 2 specimens, 7 km W of Ban Bo Yuak, 1000 m, 25.xi.1998, leg. T. Csővári
& L. mikus; 2 specimens, Doi Phuka N.P., between Pua and Bo Luang, 1350 m, 101˚05’E, 19˚12’N, 3.xi.2002, leg. b. HeRCzig & g.
Ronkay. Prov. Chiang Mai: 1 specimen, Mt. Doi Phahompok, 18 km NW of Fang, 2100 m, 7.xii.1998, leg. m. HRebLay, i. soós & y.
sHeRpani; 1 x, 16 km NW of Fang, 2000 m, 6-7.viii.1999; 1 specimen, 7 km W of Pa Pae, 1230 m, 27.xi.1998; 1 specimen, 12 km
NW of Chiang Dao, 750 m, 12.xi.1998, leg. T. Csővári & l. mikus; 1 specimen, 4 km S of Kop Dong, 1800 m, 99º03’E, 19º52’N,
6.xi.2002, leg. b. HeRCzig & g. Ronkay; 1 specimen, 10 km W of Mae Ai, 1500 m, 26.ii.1998, leg. m. HRebLay & Cs. szabóky; 1 w,
20 km NW of Sop Kha, 2 km S of Kop Dong, 1800 m, 13.ix.1999, leg. a. szabó & z. CzeRe, slide No. LGN 171 (W 7346). Prov.
Mae Hong Son: 4 specimens, between Pa Pae and Khun Sa, 1250 m, 98º39’E, 19º08’N, 31.x.2002, leg. b. HeRCzig & g. Ronkay
(coll. MWM and G. Ronkay).
Mokfanola gen. nov.
Type species: Mokfanola crustaceata spec. nov.
Species content:
M. crustaceata spec. nov.
Diagnosis. The new genus is known as monotypical. The type-species resembles strongly Ezishnola fuscographa
28
by its similarly pale grey forewing ground colour and the rather diffuse transverse lines, but the freshly hatched
specimens are distinguishable by their darker grey spots in the median area, which are absent in Ezishnola. The
identication of the worn specimens requires, however, the examination of the x genitalia which show conspicu-
ous differences. The conguration of the x genitalia is unique within the subfamily Nolinae, being dissimilar to the
other genera. The diagnostic features of the genus are the very strongly sclerotized, elongated-triangular valva
with pointed apex, the presence of rmly attached (not easily removable) and strong setae covered densely the
ventral margin and the sacculus, and the large and long, strongly sclerotized, claw-like process of costal margin,
forming together with the valval apex a structure resembling crab’s pincers. The fultura inferior of Mokfanola is
also characteristic, appearing as an elongated, reversed V-shaped plate. Despite the unique x genitalia, the w
genitalia is rather simple, displaying a few specic characters only like the strongly sclerotized, discoidal ostium
bursae, the strongly bent ductus bursae and the conspicuously elongated, narrow, more or less stick-like, strongly
sclerotized signum.
Mokfanola crustaceata spec. nov.
(Pl. 3, g. 7, Pl. 11, g. 13; gen. g. 25)
Holotype. x, „North Thailand, Prov. Chiang Mai, 450m, Mok Fa Garden Resort, 98º48’E, 19º06’N, 01.11.2002, leg. b. HeRCzig &
g. Ronkay”, slide No. LGN 1059 (coll. G. Ronkay).
Paratypes. Thailand. 1 w, Prov. Mae Hong Son, between Pa Pae and Khun Sa, 1250 m, 98º39’E, 19º08’N, 31.x.2002, leg. b.
HeRCzig & g. Ronkay, slide No. LGN 1264 (W 7381). India. Assam: 1 x, Kaziranga, Wild Life Reserve, 27º08’N,
93º56’E, 200 m, 2-3.vii.1997, leg. v. siniaev, slide No. LGN 1266 (W 7349) (coll. MWM).
Diagnosis. The new species is very similar externally to Ezishnola fuscographa, the distinctive external and genital
features are summarized in the diagnosis of the genus Mokfanola.
Distribution. The species has been found in northern Thailand and in north-east India (Assam).
Sarbena lignifera WaLkeR, 1862
(Pl. 3, g. 8)
Sarbena lignifera WaLkeR, 1862, J. Proceedings of the Linnean Society (Zoology) 6: 137. Type-locality: Borneo, Sarawak. Lectotype:
x, designated (as type) by sWinHoe, 1900, Catalogue of Eastern and Australian Lepidoptera Heterocera in the Collection of the
Oxford University Museum 2: 589.
Material examined:
Thailand. Prov. Nan: 1 x, 5 km N of Ban Luang, 350 m, between Pi Nai and Pi Tai, 100º27’E, 18º56’N, 4.xi.2002, leg. b. HeRCzig
& g. Ronkay, slide No. LGN 751. Prov. Chiang Mai: 1 x, between Chiang Dao and Kariang, 900 m, 98º48’E, 19º25’N, 26.x.2002
leg. b. HeRCzig & g. Ronkay, slide No. LGN 749; 1 x, 4 km SE of Pang Faen, 1100 m, 18.i.2004, leg. p. HentsCHeL & a. szabó, slide
No. LGN 948 (W 7350) (coll. G. Ronkay and MWM).
Sarbena ustipennis (Hampson, 1895)
(Pl. 3, g. 9)
Cyphotopsyche ustipennis Hampson, 1895, Trans. ent. Soc. Ld. 1895: 297. Type-locality: Bhutan and Ceylon [Sri Lanka]. Lectotype:
x, designated (as type) by Hampson, 1900, Catalogue of the Lepidoptera Phalaenae in the British Museum 2: 52.
Material examined:
Thailand. Prov. Nan: 1 x, 5 km N of Ban Luang, 350 m, between Pi Nai and Pi Tai, 100º27’E, 18º56’N, 4.xi.2002, leg. b. HeRCzig &
29
g. Ronkay, slide No. LGN 750 (coll. G. Ronkay). Prov. Chiang Mai: 1 x, 4 km SE of Pang Faen, 1100 m, 31.i.2004, leg. a. szabó,
slide No. LGN 947 (W 7351) (coll. MWM).
Sarbena hollowayi LászLó, Ronkay & Witt, 2004
(Pl. 3, g. 10)
Sarbena hollowayi LászLó, Ronkay & Witt, 2004, Entomofauna 25(18): 285. Type locality: N. Vietnam, Prov. Nghe An, Que Phong,
Ban Khom. Holotype: x, in coll. MWM.
Material examined:
Thailand. Prov. Chiang Mai: 1 x, 4 km SE of Pang Faen, 1100 m, 27.i.2004, leg. a. szabó, slide No. LGN 949 (W 7352) (coll.
MWM).
Evonima aperta WaLkeR, 1865
(Pl. 3, gs 11-12)
Evonima aperta WaLkeR, 1865, List of the Specimens of Lepidopterous Insects in the Collection of the British Museum 32: 505.
Type-locality: Java. Syntypes: two ww, in coll. BMNH.
Material examined:
Thailand. Prov. Chiang Mai: 1 specimen, 4 km SE of Pang Faen, 1100 m, 18.i.2004, leg. p. HentsCHeL & a. szabó; 1 specimen,
between Chiang Dao and Kariang, 900 m, 98º48’E, 19º25’N, 26.x.2002 leg. b. HeRCzig & g. Ronkay (coll. MWM). Prov. Nan: 1
specimen, 4 km W of Pha Lak, 100º34’E, 19º21’N, 5.xi.2002, leg. b. HeRCzig & g. Ronkay (coll. G. Ronkay); 1 x, 1 w, Loei Province
Phu Luang Wildlife Sanctuary, 8-14.x.1984, 700-900 m, leg. kaRsHoLt, LomHoLDt & nieLsen, slide Nos LGN 396 (x), LGN 395 (w)
(coll. ZMUC).
Evonima xanthoplaga (Hampson, 1911)
(Pl. 3, gs 13-14)
Roeselia xanthoplaga Hampson, 1911, Annals and Magazine of Natural History 8(8): 399. Type-locality: [India] Sikkim. Holotype:
w, in coll. BMNH.
Material examined:
Thailand. Prov. Chiang Mai: 5 specimens, 4 km S of Kop Dong, 1800 m, 99º03’E, 19º52’N, 6.xi.2002, leg. b. HeRCzig & g. Ronkay;
1 specimen, Mt. Doi Inthanon, NP, 2300 m,16.i.2004, leg. a. szabó (coll. G. Ronkay and MWM). Prov. Nan: 2 xx, 30 km E of Pua,
1700 m, 10.xi.1999, leg. m. HRebLay & i. soós, slide Nos LGN 516 (W 7353), LGN 517 (W 7354) (coll. MWM).
Evonima spec.
(Pl. 3, g. 15)
Material examined:
Thailand. Prov. Chiang Mai: 1 x, 4 km SE of Pang Faen, 1100 m, 18.i.2004, leg. p. HentsCHeL & a. szabó, slide No. LGN 1009
(W 7355) (coll. MWM).
Remarks. The species is recorded by a rather worn single specimen. The examined specimen differs externally
from all other congeners occurring in Thailand, but the proper identication would require the entire revision of
the genus Evonima.
30
Evonima ochritincta (Hampson, 1901)
(Pl. 3, g. 16)
Poecilonola ochritincta Hampson, 1901, Annals and Magazine of Natural History 7: 178. Type-locality: [Sri Lanka] Ceylon, Puttalam.
Holotype: in BMNH.
Material examined:
Thailand. Prov. Chiang Mai: 1 x, 4 km SE of Pang Faen, 1100 m, 26-27.i.2004, leg. a. szabó, slide No. LGN 1152 (coll. G.
Ronkay).
Evonima unicolor spec. nov.
(Pl. 3, g. 17; gen. g. 26)
Holotype. x, „N. Thailand, Prov. Nan, 5 km N of Ban Luang, 350 m, between Pi Nai and Pi Tai, 100º27’E, 18º56’N, 4.11.2002, leg.
b. HeRCzig & g. Ronkay”, slide No. LGN 797 (W 7356) (coll. MWM).
Diagnosis. The new species differs from all other known species of Evonima by its unicolorously dark grey ground
colour of forewing, lacking any conspicuous, colourful markings, only some darker grey spots in the terminal
area are present. The other species of Evonima have yellowish-brown or dark brown forewings with extensive,
often vivid pattern. The x genitalia of the new species are typical for the genus, displaying only slight differences
comparing with those of the other taxa. They can be distinguished from the related species (especially E. aperta
and E. xanthoplaga) by their shortest and broadest apical part of tegumen and uncus, the ampullar process of E.
unicolor is the shortest within the related taxa and its apical end is not surpassing the ventral margin of the valva
like in the related species. The w is unknown.
Distribution. The species is known from the type-locality only.
Varganola gen. nov.
Type species: Roeselia avibasis (Hampson, 1900) comb. nov.
Species content:
V. avibasis (Hampson, 1900) comb. nov.
Diagnosis. The new genus is monotypical, its closest relative is Meganola DyaR, 1898. The external appearance of
the species of the genus is rather unique, according to its brownish grey forewing with blackish longitudinal streaks
along the veins, this forewing pattern resembles slightly the species of Sarbena WaLkeR, 1862. The diagnostic
features of the x genitalia are the very long, medially strongly tapering, apically broadened valva, the rather broad,
ribbon-like harpe originating very close to the base of the valva, and the presence of a series of large, robust tri-
angular cornuti in vesica. The valva of the species of Meganola is shorter and rather variable in shape, sometimes
with strongly broadened basal part, the harpe is longer, usually thorn-shaped or claw-like and the vesica has much
more simplied armature without cornuti or with a single, elongated cornutus. The w genitalia of Varganola do not
show apomorphic feature, their conguration displays the close relationship with Meganola.
Varganola avibasis (Hampson, 1900) comb. nov.
(Pl. 4, gs 6-7; gen. g. 27)
Roeselia avibasis Hampson, 1900, Catalogue of the Lepidoptera Phalaenae in the British Museum 2: 63, pl. 20, g. 3. Type-locality:
[Sri Lanka] Ceylon, Colombo. Holotype: x, in coll. BMNH.
31
Type material examined: holotype of Roeselia avibasis Hampson, 1900: x, „TYPE” (red ring type label), „Ceylon”, slide No. LGN
718 (BM Noct. 18198) (coll. BMNH).
Additional material examined:
Thailand. Prov. Chiang Mai: 3 xx, between Chiang Dao and Kariang, 900 m, 98º48’E, 19º25’N, 26.x.2002, slide Nos LGN 752 (W
7357), LGN 753 (W 7358), LGN 980; 1 w, between Fang and Nor Lae, 99º06’E, 20º02’N, 28.x.2002, leg. b. HeRCzig & g. Ronkay,
slide No. LGN 981 (coll. G. Ronkay and MWM).
Manoba lativittata (mooRe, 1888)
(Pl. 4, gs 8-9)
Roeselia lativittata mooRe, 1888, Description of new Indian Lepidopterous Insects from the Collection of the late Mr. W. S. Atkinson
1888: 286. Type-locality: [India] [Sikkim] Darjeeling. Holotype: x, in coll. ZMHU Berlin.
Material examined:
Thailand. Prov. Chiang Mai: 2 specimens, 4 km SE of Pang Faen, 1100 m, 13.i.2004, leg. p. HentsCHeL & a. szabó; 1 specimen,
same site, 4 km SE of Pang Faen, 1100 m, 27.i.2004, leg. a. szabó; 7 specimens, between Fang and Nor Lae, 1600 m, 99º06’E,
20º02’N, 12.xi.2002; 3 specimens, same site, 28.x.2002; 2 specimens, same site, 7.xi.2002; 2 specimens, 900 m, between Chiang
Dao and Kariang, 98º48’E, 19º25’N, 8.xi.2002; 1 specimen, same site, 26.x.2002; 1 fem, Mok Fa Garden Resort, 450 m, 98º48’E,
19º06’N, 1.xi.2002, slide No. LGN 961, leg. b. HeRCzig & g. Ronkay; 1 specimen, 20 km NW of Sop Kha, 2 km S of Kop Dong, 1800
m, 27.i.1999; 2 specimens, 20 km NW of Mae Ai, 1650 m, 26.i.1999, slide No. LGN 1153 (W 7359) (x); 1 x, same site, 7.i.1999; 1
x, same site, 9.ix.1999, slide No. LGN 959 (W 7360), leg. a. szabó & z. CzeRe. Prov. Nan: 1 specimen, 1700 m, 30 km E of Pua,
20.i.2004, leg. p. HentsCHeL & a. szabó; 1 specimen, 5 km N of Ban Luang, 350 m, between Pi Nai and Pi Tai, 100º27’E, 18º56’N,
4.xi.2002, leg. b. HeRCzig & g. Ronkay; 1 x, 30 km E of Pua, 1700 m, 18.ii.1998, leg. m. HRebLay & Cs. szabóky; 1 w, 5 km N of Bo
Luang, 1000 m, 10.ii.2000, slide No. LGN 962 (W 7361); 1 x, 25 km N of Bo Luang, 1150 m, 3.ii.2000, leg. m. HRebLay, y. sHeRpani
& a. szabó. Prov. Mae Hong Song: 7 specimens, between Pa Pae and Khun Sa, 1250 m, 98º39’E, 19º08’N, 31.x.2002, leg. b.
HeRCzig & g. Ronkay (coll. G. Ronkay and MWM).
Manoba gyulaipeteri spec. nov.
(Pl. 4, gs 10-11; gen. g. 28)
Holotype. x, „North Thailand, Prov. Chiang Mai, 1600m, between Fang and Nor Lae, 99º06’E, 20º02’N, 7.xi.2002, leg. B. Herczig
& G. Ronkay”, slide No. LGN 1090 (coll. G. Ronkay).
Paratypes. Thailand. Prov. Chiang Mai: 1 x, Doi Suthep-Pui NP, 1460 m, 4.ii.1989, leg. a.m. Cotton & i.J. kitCHing BM 1989-57
(coll. BMNH); 2 xx, 4 km S of Kop Dong, 1800 m, 99º03’E, 19º52’N, 6.xi.2002, leg. b. HeRCzig & g. Ronkay, slide
No. LGN 1028 (W 7362); 1 x, Mt. Doi Phahompok, 16 km NW of Fang, 2000 m, 15.ii.1998, leg. m. HRebLay & Cs.
szabóky. Prov. Nan: 1 w, 30 km E of Pua, 1700 m, 20.i.2004, leg. p. HentsCHeL & a. szabó, slide No. LGN 1029
(W 7363); 1 x, from the same site, 25.i.1999, leg. a. szabó & z. CzeRe. Prov. Mae Hong Song: 2 xx, 21 km NW
of Pai, 1360 m, 7.ii.1998, leg. m. HRebLay & Cs. szabóky. Nepal. Ganesh Himal: 3 xx, 1 w, above Nesim, 2720
m, 85º16’E, 28º08,5’N, 21.ix.1995, slide Nos LGN 22 (W 7364), LGN 255 (W 7365), (xx), LGN 23 (W 7366) (w);
2 ww, near Godlang, 2520 m, 85º17’E, 28º10’N, 13.ix.1995; 1 x, 7 km W Godlang, 2950 m, 85˚17’E, 28˚10’N,
14.ix.1995; 1 x, 2 km W Gholjong, 2420 m, 85˚18’E, 28˚11’N, 12.ix.1995, leg. b. HeRCzig & gy.m. LászLó; 1 x,
2 km W of Thangjet, 2300 m, 85˚17’E, 28˚10’N, 18.ix.1994, leg. m. Hreblay & T. Csővári; 1 x, Kamalang, 1850
m, 27.x.1995, 85˚11’E, 28˚06’N, leg. m. HRebLay & L. bóDi; 1 x, 2 km N of Dhunche, 2050 m, 15.ix.1995, leg. L.
németH. Langtang Himal: 1 x, 1,5 km NE Dhunche, 85º18’E, 28º06’N, 24.ix.1994, leg. G. CsoRba & L. Ronkay.
Mechi, Taplejung Area: 1 w, Kare Banjang, 2250 m, 87º56’E, 27º25’N, 2.xi.1996, leg. gy.m. LászLó & g. Ronkay,
slide No. LGN 256 (W 7367) (coll. HNHM, G. Ronkay and MWM).
Diagnosis. The new species belongs to the Manoba lativittata-tessellata species group, its closest relative is M.
lativittata. It can be distinguished easily from M. lativittata by their completely different forewing pattern: M. gyulai-
peteri has the forewing basal (and partly the medial) area uniformly covered with dark greyish brown hair-scales,
without lighter spots, the medial line closing this dark area is more or less straight, and the more or less uniformly
32
whitish grey medial and partly terminal area is much darker than in M. lativittata. The dark area of the forewing of
M. lativittata is more blackish, reaching the terminal line of forewing, but contains a bright whitish elongate patch
at the basal part of costal margin, a large rounded whitish patch at the tornus, and the marginal area has large
whitish medio-costal and apical patches.
Despite the conspicuous external differences, the two species have very similar genitalia of both sexes. The slight
differences between M. gyulaipeteri and M. lativittata are as follows: the new species has somewhat thicker and
shorter harpe, and the carinal spine of aedeagus is slightly shorter and straight, this spine is somewhat longer and
slightly curved in M. lativittata. The w genitalia of the two species differ in the shape of signum bursae: the typical
cup-like signum is somewhat more elongated in the new species than in M. lativittata.
Distribution. The new species is found in Northern Thailand and the central and eastern massifs of the Nepal
Himalaya.
Manoba tesselata (Hampson, 1896)
(Pl. 4, gs 12-13)
Nola tesselata Hampson, 1896, Fauna of British India, Moths 4: 504. Type-locality: [India] [Meghalaya] Khasis. Holotype: w, in coll.
BMNH.
Material examined:
Thailand. Prov. Chiang Mai: 4 specimens, between Fang and Nor Lae, 1600 m, 99º06’E, 20º02’N, 12.xi.2002; 5 specimens, same
site, 28.x.2002, slide No. LGN 964 (x); 2 specimens, between Chiang Dao and Kariang, 900 m, 98º48’E, 19º25’N, 26.x.2002 leg.
b. HeRCzig & g. Ronkay; 1 w, Mt. Doi Phahompok, 18 km NW of Fang, 2100 m, 8.xi.1999, leg. m. HRebLay & i. soós, slide No. LGN
965 (W 7369); 2 specimens, Mt. Doi Phahompok, 20 km NW of Fang, 2150 m, 24.i.2004, leg. a. szabó; 1 specimen, 16 km NW of
Fang, 2000 m, 24.ii.1998; 1 w, 15 km SW of Wiang Haeng, 1400 m, 9.ii.1998, leg. m. HRebLay & Cs. szabóky; 1 w, 20 km NW of
Mae Ai, 1650 m, 26.i.1999; 1 x, Doi Inthanon, 2300 m, 7-8.ix.1999, slide No. LGN 960 (W 7368), leg. a. szabó & z. CzeRe. Prov.
Nan: 1 specimen, 5 km N of Bo Luang, 1000 m, 4.ii.2000, leg. m. HRebLay, y. sHeRpani & a. szabó; 1 x, 30 km E of Pua, 1700 m,
25.i.1999, leg. a. szabó & z. CzeRe; 1 w, same site, 18.ii.1998, leg. m. HRebLay & Cs. szabóky (coll. G. Ronkay & MWM).
Manoba adriennae spec. nov.
(Pl. 4, gs 14-15; gen. g. 29)
Holotype. x, „Thailand, Changwat Nan, 1700m, 30 km E of Pua, 20.i.2004, Leg. Péter HentsCHeL & Attila szabó”, slide No. LGN
963 (W 7370) (coll. MWM).
Paratype. 1 x, Thailand. Prov. Chiang Mai, Mt. Doi Phahompok, 20 km NW of Fang, 2100 m, 24.i.2004, leg. a. szabó, slide No.
LGN 1244 (W 7382) (coll. MWM).
Diagnosis. The new species is closely related to M. tessellata. The two species have similar forewing pattern, but
are easily distinguishable by the following features: the forewing ground colour of M. adriennae is more brownish,
the whitish spots are less prominent, the white part of the basal area is shorter and the hindwing is signicantly
darker than in M. tessellata.
The x genitalia of the new species are in general very similar to those of M. tessellata but the valva is somewhat
more elongate and apically narrower, the aedeagus is slightly longer, slenderer, and the lateral processes of the
proximal tip of the aedeagus are much shorter. The differences in the w genitalia are larger, as M. adriennae has
somewhat longer apophyses anteriores and posteriores, conspicuously less elongate, more ovoid corpus bursae
and considerably smaller and shorter claw-like signum bursae in comparison with those of M. tessellata.
Distribution. Northern Thailand.
33
Manoba fasciatus (Hampson, 1894)
(Pl. 4, gs 16-17)
Rhynchopalpus fasciatus Hampson, 1894, Fauna of British India, Moths 2: 144. Type-locality: [India] [Nagaland] Naga Hills. Holo-
type: x, in coll. BMNH.
Material examined:
Thailand. Prov. Chiang Mai: 12 specimens, between Fang and Nor Lae, 1600 m, 99º06’E, 20º02’N, 12.xi.2002, slide Nos LGN
996 (W 7372) (x), LGN 1068 (w); 10 specimens, from the same site, 28.x.2002 and 7.xi.2002, slide No. LGN 1066; 1 specimen,
4 km S of Kop Dong, 99º03’E, 19º52’N, 1800 m, 6.xi.2002 leg. b. HeRCzig & g. Ronkay; 1 specimen, 4 km SE of Pang Faen, 1100
m, 31.i.2004, leg.: a. szabó; 1 x, 20 km NW of Sop Kha, 2 km S of Kop Dong, 1800 m, 27.i.1999, slide No. LGN 478 (W 7374); 1
x, 20 km NW of Mae Ai, 1650 m, 26.i.1999, leg. a. szabó & z. CzeRe, slide No. LGN 477 (W 7373). Prov. Nan: 1 specimen, 1700
m, 30 km E of Pua, 20.i.2004, leg. p. HentsCHeL & a. szabó; 1 x, 25 km N of Bo Luang, 1150 m, 3.ii.2000, slide No. LGN 995 (W
7371), leg. m. HRebLay, y. sHeRpani & a. szabó; 1 x, 30 km E of Pua, 1700 m, 25.i.1999 leg. a. szabó & z. CzeRe. Prov. Mae Hong
Son: 1 w, between Pa Pae and Khun Sa, 1250 m, 98º39’E, 19º08’N, 31.x.2002, slide No. LGN 997, leg. b. HeRCzig & g. Ronkay
(coll. G. Ronkay and MWM).
Manoba ronkaylaszloi spec. nov.
(Pl. 4, gs 18-19; gen. g. 30)
Holotype. x, „N. Thailand, Chiang Mai Prov., between Chiang Dao and Kariang, 900m, 98º48’E, 19º25’N, 26.x.2002 leg. b. Herczig
& G. Ronkay”, slide No. LGN 998 (W 7375) (coll. MWM).
Paratypes. Thailand. Prov. Chiang Mai: 1 x, with the same data as the holotype; 1 x, 4 km SE of Pang Faen, 1100 m, 6.ii.2000,
leg. m. HRebLay, slide No. LGN 1258 (W 7376). Prov. Nan: 1 x, 30 km E of Pua, 1700 m, 20.i.2004, leg. p.
HentsCHeL & a. szabó. Vietnam. Prov. Tuyen Quang: 1 x, Na Hang Nature Reserve, 300 m, 105º5’E, 22º3’N,
22.ii.-5.iii.1997, leg. g. CsoRba, slide No. LGN 480 (W 7377) (coll. MWM).
Diagnosis. The new species is the sister species of M. fasciatus. The two species are easily distinguishable by
their external appearance: the forewing ground colour of M. ronkaylaszloi is much paler whitish grey, without red-
brownish shade in the basal area, and the median fascia is much broader and more diffuse, consisting of a dark
grey lled area between antemedial and medial lines. The forewing ground colour of M. fasciatus is darker with
characteristic red-brown hue in the basal part, and the median fascia is rather sharply dened, but considerably
narrower than in M. ronkaylaszloi.
The x genitalia display better the close relationship between the two species. M. ronkaylaszloi has, in comparison
with M. fasciatus, somewhat longer and narrower valva and slightly shorter, basally broader, more pointed harpe,
and the armature of the vesica is more complex, consisting of a larger number of spines which are considerably
larger than in its sister taxon. The w is unknown.
Distribution. The species is found in the northern part of Indochina, in North Thailand and North Vietnam.
Manoba coxi HoLLoWay, 2003
(Pl. 4, gs 20-21)
Manoba coxi HoLLoWay, 2003, The Moths of Borneo 18: 41, pl. 2, g. 74. Type-locality: Borneo, Sarawak. Holotype: x, in coll.
BMNH.
Type material examined:
Holotype. x, „Sarawak: Gunong Mulu Nat. Park, R.G.S. Exped. 1977-8 (J.D. Holloway et al.), Site 27, April, G. Api, 1500 m, Pan-
danus Camp, 429541, scrub, Pandanus”, slide No. BM Noct. 17760 (coll. BMNH).
Additional material examined:
Thailand. Prov. Chiang Mai: 5 specimens, between Fang and Nor Lae, 1600 m, 99º06’E, 20º02’N, 7.xi.2002, slide No. LGN 977
34
(w); 4 specimens, same site, 12.xi.2002; 4 specimens, same site, 28.x.2002, slide No. LGN 1023 (W 7384) (w); 2 specimens,
between Chiang Dao and Kariang, 900 m, 98°48’E, 19°25’N, 26.x.2002, slide No. LGN 976 (W 7383) (x), leg. b. HeRCzig & g.
Ronkay (coll. G. Ronkay and MWM).
Manoba melanota (Hampson, 1900)
(Pl. 5, gs 1-2)
Nola melanota Hampson, 1900, Catalogue of the Lepidoptera Phalaenae in the British Museum 2: 35, pl. 19, g. 3. Type-locality:
[India] Sikkim. Syntypes: one x and ten ww, in coll. BMNH.
Rhynchopalpus erythromedia inoue, 1998, syn. n., Tinea 15 (Supplement 1): 93, pl. 140, g. 23-24, text g. 890-891. Type-locality:
Nepal, Changma, Remechhap, Janakpur. Holotype: x, in NSMT.
Type material examined:
Syntype of Nola melanota: x, „TYPE” (red ring type label), „Sikkim, 7000 August 1895, J.g. piLCHeR.” „97-81” „Nola melanoton
(sic!) x Hmpsn”, slide No. BM Arct. 1799 (coll. BMNH).
Additional material examined:
Thailand. Prov. Chiang Mai: 1 x, between Fang and Nor Lae, 1600 m, 99º06’E, 20º02’N, 28.x.2002, slide No. LGN 988; 1 x, from
the same site, 12.xi.2002, leg. b. HeRCzig & g. Ronkay; 1 w, Doi Inthanon, NP, 2300 m, 7-8.ix.1999, slide No. LGN 1010 (W 7386);
1 w, 20 km NW of Sop Kha, 2 km S of Kop Dong, 1800 m, 13.ix.1999, slide No. LGN 989 (W 7387), leg. a. szabó & z. CzeRe; 1
x, Mt. Doi Phahompok, 19 km NW of Fang, 1900 m, 3.iv.1998, slide No. LGN 1107 (W 7388); 1 w, 16 km NW of Fang, 2000 m,
27.iii.1998, leg. T. Csővári & P. sTéger. Prov. Nan: 1 x, 30 km E of Pua, 1700 m, 20-21.vi.1998, leg. I. soós & a. szabó, slide No.
LGN 974 (W 7385); 1 x, from the same site, 10.xii.1998, leg. m. HRebLay, i. soós & y. sHeRpani (coll. G. Ronkay and MWM).
Manoba phaeochroa (Hampson, 1900)
(Pl. 5, g. 3)
Celama phaeochroa Hampson, 1900, Catalogue of the Lepidoptera Phalaenae in the British Museum 2: 29, pl. 18, g. 24. Type-
locality: [India] Sikkim. Syntypes: four ww, in coll. BMNH.
Type material examined:
Syntype w of Celama phaeochroa Hampson, 1900: „TYPE” (red ring type label), „Sikkim, 7000 6. 1895, J.g. piLCHeR.” „97-31”
„Aradrapha phaeochroa w Hmpsn”, slide No. BM Arct. 896 (coll. BMNH).
Additional material examined:
Thailand. Prov. Chiang Mai: 1 x, 1 w, Doi Inthanon, NP, 2300 m, 14.xii.1998, leg. m. HRebLay, i. soós & y. sHeRpani, slide No. LGN
1105 (W 7389) (x), LGN 1106 (W 7390) (w); 1 x, Doi Inthanon National Park, km 43,5 road (N of) Chom Thong – summit 5,5
km above checkpoint 2, 2050 m, lower montane forest, 15-21.xi.1998 leg/ex coll. DR. R. bReCHLin, slide No. LGN 1118 (W 7391)
(coll. MWM).
Manoba melanomedia (inoue, 1991)
(Pl. 5, g. 4)
Meganola melanomedia inoue, 1991, Tyô to Ga 42(2): 76, gs 1A, 1B, 2A, 3A. Type-locality: [Taiwan] Chiayi County (= Chiai Shien),
Alishan. Holotype: x, in coll. NSMT.
Material examined:
Thailand. Prov. Chiang Mai: 1 x, 4 km SE of Pang Faen, 1100 m, 31.i.2004, leg. a. szabó, slide No. LGN 1026 (W 7392) (coll.
MWM).
35
Manoba lilliptiana (inoue, 1998)
(Pl. 5, g. 5)
Rhynchopalpus lilliptiana inoue, 1998, Tinea 15 (Supplement 1): 93, pl. 140, g. 22, text gs 888-889. Type-locality: Nepal, Chapauli,
Sindhuli. Holotype: x, in coll. inoue.
Material examined:
Thailand. Prov. Chiang Mai: 2 xx, between Fang and Nor Lae, 1600 m, 99º06’E, 20º02’N, 7.xi.2002, slide Nos LGN 1083, LGN
1004 (W 7394); 2 xx, 1 w, from the same site, 12.xi.2002, slide Nos LGN 1082, LGN 1132 (W 7397) (xx), LGN 1133 (W 7398)
(w); 1 x, from the same locality, 28.x.2002, slide No. LGN 1088; 1 x, 4 km S of Kop Dong, 1800 m, 99º03’E, 19º52’N, 6.xi.2002,
slide No. LGN 1002 (W 7393), leg. b. HeRCzig & g. Ronkay; 1 x, 6 km SE of Pang Faen, 1100 m, 29.i.1999, leg. a. szabó & z.
CzeRe, slide No. LGN 1108 (W 7395); 1 w, 4 km SE of Pang Faen, 1100 m, 18.i.2004, leg. p. HentsCHeL & a. szabó, slide No. LGN
1036 (W 7399). Prov. Nan: 1 w, 25 km N of Bo Luang, 1150 m, 14.i.1999, leg. a. szabó & z. CzeRe, slide No. LGN 1124 (W 7396)
(coll. G. Ronkay and MWM).
Manoba subfuscataria (inoue, 1998)
(Pl. 5, g. 6)
Rhynchopalpus subfuscataria inoue, 1998, Tinea 15 (Supplement 1): 93, pl. 140, gs 20-21, text gs 886-887. Type-locality: Nepal,
Chapauli, Sindhuli. Holotype: x, in coll. inoue.
Material examined:
Thailand. Prov. Nan: 1 x, 30 km E of Pua, 1700 m, 10.xii.1998, leg. m. HRebLay, i. soós & y. sHeRpani, slide No. LGN 1127 (W
7400) (coll. MWM).
Manoba chamberlaini HoLLoWay, 2003
(Pl. 5, gs 7-8)
Manoba chamberlaini HoLLoWay, 2003, The Moths of Borneo 18: 39, pl. 2, g. 69. Type-locality: Borneo, Sarawak. Holotype: x,
in coll. BMNH.
Material examined:
Thailand. Prov. Nan: 1 x, 4 km W of Pha Lak, 100º34’E, 19º21’N, 5.xi.2002, slide No. LGN 1003 (W 7401), leg. b. HeRCzig & g.
Ronkay; 1 x, 30 km E of Pua, 1700 m, 12-13.i.1999, slide No. LGN 1032 (W 7403); 2 xx, 1 w, from the same site, 25.i.1999,
slide Nos LGN 1035 (W 7404) (w), LGN 1103 (W 7408), LGN 1109 (W 7410) (xx), leg. a. szabó & z. CzeRe; 1 w, from the same
locality, 6.iv.1998, leg. T. Csővári & P. sTéger, slide No. LGN 1104 (W 7409); 1 x, from the same site, 18.ii.1998, leg. m. HRebLay
& Cs. szabóky, slide No. LGN 1128 (W 8369); 1 x, 5 km N of Bo Luang, 1000 m, 4.ii.2000, leg. m. HRebLay, y. sHeRpani & a. szabó,
slide No. LGN 1120 (W 8366). Prov. Chiang Mai: 2 xx, between Fang and Nor Lae, 1600 m, 99º06’E, 20º02’N, 28.x.2002, slide
Nos LGN 1089, LGN 990 (W 7402); 1 x, from the same locality, 12.xi.2002; 2 xx, 1 w, from the same site, 7.xi.2002, slide Nos
LGN 1005 (W 7407) (w), LGN 1081, LGN 1087 (xx); 2 ww, 4 km S of Kop Dong, 99º03’E, 19º52’N, 1800 m, 6.xi.2002, slide No.
LGN 1034 (W 7405), LGN 1033 (W 7406), leg. b. HeRCzig & g. Ronkay; 1 w, 20 km NW of Mae Ai, 1650 m, 26.i.1999, slide No.
LGN 1119 (W 7411), leg a. szabó & z. CzeRe. Prov. Mae Hong Song: 1 x, 1 w, 1 km S of Bahundanda, 1000 m, 6.ii.1998, leg. m.
HRebLay & Cs. szabóky, slide Nos LGN 1121 (W 8367) (w), LGN 1122 (W 8368) (x) (coll. G. Ronkay and MWM).
Manoba tristicta (Hampson, 1900)
(Pl. 5, gs 9-10)
Nola tristicta Hampson, 1900, Catalogue of the Lepidoptera Phalaenae in the British Museum 2: 37, pl. 19, g. 4. Type-locality:
[India] Sikkim. Syntypes: two xx and three ww, in coll. BMNH.
Material examined:
36
Thailand. Prov. Chiang Mai: 3 specimens, 4 km S of Kop Dong, 99º03’E, 19º52’N, 1800 m, 6.xi.2002; 3 specimens, between Fang
and Nor Lae, 1600 m, 99º06’E, 20º02’N, 12.xi.2002; 11 specimens, from the same site, 28.x.2002 and 7.xi.2002, leg. b. HeRCzig &
g. Ronkay; 1 specimen, 23 km NW of Sop Kha, 1 km E of Kop Dong, 1650 m, 5.viii.1999, leg. T. Csővári & l. mikus; 3 specimens,
from the same site, 14.i.2004, leg. p. HentsCHeL & a. szabó; 2 specimens, 20 km NW of Sop Kha, 2 km S of Kop Dong, 1800 m,
8.i.1999, slide No. LGN 1125 (W 8459) (x); 1 specimen, from the same locality, 27.i.1999, leg. a. szabó & z. CzeRe; 1 specimen, 4
km SE of Pang Faen, 1100 m, 13.i.2004; 2 specimens, from the same site, 26.i.2004 and 31.i.2004, leg. p. HentsCHeL & a. szabó;
1 specimen, Mt. Doi Phahompok, 20 km NW of Fang, 2450 m, 24.i.2004, leg. a. szabó; 1 specimen, Mt. Doi Phahompok, 20 km
NW of Fang, 2000 m, 15.i.2004, leg. p. HentsCHeL & a. szabó; Prov. Nan: 1 specimen, 30 km E of Pua, 1700 m, 20.ii.1998, leg. t.
Csővári & l. mikus; 3 specimens, from the same locality, 20-21.vi.1998, leg. i. soós & a. szabó; 1 specimen, from the same site,
27.ii.1998, leg. m. HRebLay & Cs. szabóky; 1 specimen, from the same locality, 6.iv.1998, leg. T. Csővári & P. sTéger; 1 w, from the
same site, 12-13.i.1999, leg. a. szabó & z. CzeRe, slide No. LGN 1126 (W 8470); 2 specimens, from the same site, 20.i.2004, leg.
p. HentsCHeL & a. szabó; 1 specimen, 22 km N of Bo Luang, 1100 m, 18-19.vi.1998, leg. i. soós & a. szabó. Prov. Chiang Rai: 2
specimen, 1 km SE of Khun-Kon Waterfall, 600 m, 15.xi.1998, leg. T. Csővári & l. mikus. Prov. Mae Hong Song: 1 specimen, 21
km NW of Pai, 1360 m, 7.ii.1998; 1 specimen, 1 km S of Bahundanda, 1000 m, 6.ii.1998, leg. m. HRebLay & Cs. szabóky (coll. G.
Ronkay and MWM).
Inouenola gen. nov.
Type species: Meganola diversalis inoue, 1991
Species content:
I. diversalis (inoue, 1991) comb. nov.
I. pallescens (WiLeman & West, 1929) comb. nov.
I. grisalis (Hampson, 1893) comb. nov.
Diagnosis. The type species of the new genus was originally described in the genus Meganola. The basic structure
of the x genitalia of Inouenola is, however, more similar to that of Manoba, displaying certain unique features. The
x genitalia of all Manoba species are rather similar, forming a very compact phyletic unit. The new genus differs
conspicuously from Manoba in the following details: Inouenola has somewhat shorter and thicker, less curved
uncus, shorter, more or less quadrangular or slightly tapering, apically truncate valva, more robust harpe with
more elongated saccular base, very long and narrow vinculum, and much longer aedeagus (resembling more to
certain species of Meganola, especially the members of the M. ascripta-cuneifera group). The valva of Manoba is
always gradually tapering and apically rounded or sometimes pointed; the harpe is thinner, situated more basally,
the vinculum is always medium long and broad, usually V-shaped, and the aedeagus is considerably shorter.
In the w genitalia, Inouenola has much longer and narrower ductus bursae, and extremely long and robust, dag-
ger-like double signa bursae, this latter feature is unique within the whole subfamily.
Inouenola diversalis (inoue, 1991) comb. nov.
(Pl. 5, gs 11-12; gen. g. 31)
Meganola diversalis inoue, 1991, Tyô to Ga 42(2): 79, gs 1C, 1D, 2C, 3C. Type-locality: [Taiwan] Rushan-Unchen (= Lushan Spa),
Nantou Shien (= Nantou County). Holotype: x, in coll. NSMT.
Type material examined:
Holotype x of Meganola diversalis inoue, 1991: „TYPE” (red ring type label), „Formosa Nantow: Rushan-Unchen 3-6.iv.1972 m.
Owada” „Holotype Meganola diversalis Inoue (1991)” „slide 12226 (x))” „Type status veried k. Buckmaster 1993”, „Inoue Coll
1992-71”, „Type photographed in colour” Slide No. BM Arct. 4408 (coll. BMNH).
Additional material examined:
Thailand. Prov. Chiang Mai: 1 specimen, between Fang and Nor Lae, 1600 m, 99º06’E, 20º02’N, 12.xi.2002, (coll. G. Ronkay); 1
x, same site, 7.xi.2002, slide No. LGN 992 (W 8469) leg. b. HeRCzig & g. Ronkay (coll. MWM).
37
Inouenola pallescens (WiLeman & West, 1929) comb. nov.
(Pl. 5, g. 13)
Nola pallescens WiLeman & West, 1929, Annals and Magazine of Natural History 10(3): 191. Type-locality: [Taiwan] Formosa,
Kanshirei. Holotype: x, in coll. BMNH.
Type material examined:
Holotype x of Nola pallescens: Formosa, Kanshirei, 17.iv.1906, a.e. WiLeman, slide No. BM Arct. 721.
Additional material examined:
Thailand. Prov. Chiang Mai:1 x, between Chiang Dao and Kariang, 900 m, 98º48’E, 19º25’N, 26.x.2002, leg. b. HeRCzig & g.
Ronkay, slide No. LGN 1094 (coll. G. Ronkay).
Inouenola grisalis (Hampson, 1893) comb. nov.
Nola grisalis Hampson, 1893, Illustrated typical Specimens of the Lepidoptera Heterocera in the Collection of the British Museum 9:
15, 88, pl. 158, g. 9. Type-locality: [Sri Lanka] Ceylon, Pundaloya. Lectotype: x, in coll. BMNH, here designated.
Lectotype of Nola grisalis Hampson, 1893, here designated: x, „TYPE” (red ring type label), „Pundaloya, Ceylon, March.”, „Coll.
gReen. 91.-26.”. Slide No. LGN 707 (BM Noct. 18187) (coll. BMNH).
Distribution. The species is known only from Ceylon; it has no record from Thailand.
Toerpenola gen. nov.
Type species: Nola laticincta Hampson, 1896 comb. nov.
Species content:
T. laticincta (Hampson, 1896) comb. nov.
T. siamica spec. nov.
Diagnosis. The type species of the new genus has long been treated as Nola. The examination of the x genitalia
proved that N. laticincta has a closer relationship with the species of Manoba, but distinguished from them on a
higher taxonomic level. The discovery of the rather similar sister species of N. laticincta, T. siamica in Thailand
conrmed the early segregation of the lineage, supporting to establish a new genus for the two species.
The new genus differs from Manoba by the following features of the genitalia. In the x genitalia, Toerpenola has
more complex structure of valva having sinuous and more strongly sclerotized costal margin and slightly broadened
apical half, much larger and heavily sclerotized sacculus with a characteristic saccular process, but the harpe is
absent. In Manoba, the valva is tapering distally, its costal margin is more or less straight and less sclerotized, the
saccular process is absent but the harpe is always present. In the w genitalia, the new genus has considerably
broader ovipositor with longer apophyses and much shorter ductus bursae, and the single signum of the bursa
copulatrix is much ner than in Manoba.
Toerpenola laticincta (Hampson, 1896)
Nola laticincta Hampson, 1896, Fauna of British India, Moths 4: 502. Type-locality: Bhutan. Holotype: x, in coll. BMNH.
Type material examined:
Holotype x of Nola laticincta Hampson, 1896: „TYPE” (red ring type label), „Bhutan. Dudgeon. 95-203. 2500 [feet], VIII. [18]95”,
38
slide No. BM Arct. 1801 (coll. BMNH).
Distribution. The species is known from the Himalaya Mts; it has no record from Thailand.
Toerpenola siamica spec. nov.
(Pl. 5, gs 14-15; gen. g. 32)
Holotype. x, „Thailand, Changwat Chiang Mai, 4 km SE of Pang Faen, 1100m, 31.i.2004, Leg.: Attila Szabó”, slide No. LGN 1021
(W 8458) (coll. MWM).
Paratypes. Thailand. Prov. Chiang Mai: 1 w, with the same data as the holotype, slide No. LGN 1020 (W 8468) (coll. MWM); 1 w, 7
km N of Ranong, 350-500 m, Ch9 TV relay stn, 26-29.xi.1991, leg. i.J. kitCHing & a.m. Cotton, BM 1992-9 (coll.
BMNH). Cambodia. 1 w, Prov. Kampot, Bokor N.P. Hill Station, 1025 m, 10º37’37”N, 104º01’33”E, 19-21.i.2006,
leg. g. CsoRba & g. Ronkay slide No. LGN 1243 (W 8457) (coll. MWM).
Diagnosis. The new species is closely related to T. laticincta. The two species are easily distinguished by their
external features, the main differences are as follows: T. siamica is slightly smaller in size (wingspan and length
of forewing of T. siamica are 13-16 and 7-9 mm, those of T. laticincta are 14-17 mm and 8-10 mm, respectively).
The dark brown forewing median area is considerably broader in T. siamica than in T. laticincta; the white marginal
area of the new species is more greyish, while that of T. laticincta is more yellowish-brownish suffused. The differ-
ences between the x genitalia of the two species are surprisingly small, the only remarkable feature is the different
conguration of the well developed saccular process: it is nger-like in T. siamica with broadly rounded apex, that
of T. laticinta is apically pointed, rather claw-like. The aedeagus of the new species has a ne but strong carinal
tooth, while T. laticincta lacks this carinal process.
The w genitalia of the two species cannot be compared, as the w of T. laticincta is yet unknown. The typical features
of the w genitalia of T. siamica are the relatively broad ovipositor with rather long apophyses, the broad, strongly
sclerotized ostium bursae with two lateral rounded lobes fused partly with the ventral margin of the 8th segment,
the very short, distally strongly and asymmetrically sclerotized ductus bursae, the relatively short, ovoid corpus
bursae and the single, conspicuously small, ne, pin-like signum bursae.
Nanola gen. nov.
Type species: Nanola hluchyi spec. nov.
Species content:
N. hluchyi spec. nov.
Diagnosis. The only known species of the new genus represents, according to its genital features, a transitional
stage between the genera Manoba and Meganola. The narrow, elongate, apically broadly rounded, more or less
tongue shaped valva resembles certain Meganola species (e.g. M. nitida and M. mediofusca), while the harpe of
Nanola is similar in shape and position to that of Manoba, being rather simple, broad at base, slightly arcuate, api-
cally pointed, but not claw- or spine-like and without dentate margins, which is characteristic for some Meganola
species. The aedeagus of Nanola is rather simple, without carinal processes, and the vesica is without a cornuti
eld or long spines, but with a short but robust triangular tooth. The w genitalia can be characterised by the rather
long and wrinkled, strongly sclerotized, funnel-like ostium bursae, the very thick but relatively short ductus bursae
with conspicuous, oblique sclerotization in the middle, and the rather simple, membraneous corpus bursae; the
signum is absent. The w genitalia of the related genera Manoba and Meganola have simple ostium bursae and
the signum bursae is always present.
Nanola hluchyi spec. nov.
(Pl. 5, gs 16-17; gen. g. 33)
Holotype. x, „Thailand, Changwat Nan, 5 km N of Bo Luang, 1000 m, 4.ii.2000, leg. Hreblay & Szabó”, slide No. LGN 966 (W
39
8467) (coll. MWM).
Paratypes. Thailand. Prov. Nan: 1 w with the same data as the holotype, slide No. LGN 967 (W 8456) (coll. MWM); 1 x, Doi Phu
Kha, Pua, 1680 m, 26.ii.1993, leg. W. speiDeL (coll. speiDeL). Vietnam. Prov. Tuyen Quang: 1 x, Na Hang Nature
Reserve, 300 m, 105º5’E, 22º3’N, 22.ii.-05.iii.1997, leg. g. CsoRba; 1 x, Mt. Fan-si-pan, Cha Pa, 2400 m, 22º15’N,
103º46’E, 8-29.v.1993, leg. sinjaev & simonov, slide No. LGN 1185 (W 8466). Myanmar. 1 x, 21 km E Putao, Nan
Sa Bon village, 550 m, 1-5.v.1998, leg. murzin & sinjaev, slide No. LGN 1184 (W 8455). Taiwan. Prov. Taitung: 1
x, 7 km N of Tupan, 120º52’E, 22º29’N, 500 m, 16.xii.1997, leg. s. simonyi & a. szabó, slide No. LGN 1256 (W
8465). Prov. Pingtung: 1 x, 10 km E of Mutan, 400 m, 7-8.iv.1997, leg. CsoRba & Ronkay, slide No. LGN 1183 (coll.
HNHM and MWM). Khasis: 1 x, Nov. 1895, Nat. Coll., RotHsCHiLD Bequest B.M. 1939-1. (coll. BMNH).
Diagnosis. N. hluchyi is somewhat similar externally to Manoba lativittata by its whitish basal and apical areas of
the forewing, but is easily distinguished by its more elongated forewing, with a larger dark blackish brown area
and the smaller whitish spots. The forewing ground colour of the new species is somewhat paler, more brownish
than the more blackish coloured M. lativittata, and the forewing pattern of this latter species is more contrasting
than in N. hluchyi.
The diagnostic features of the genitalia of N. hluchyi are given in the diagnosis of the genus Nanola.
Meganola semirufa (Hampson, 1894) comb. nov.
(Pl. 5, gs 19-20, Pl. 11, g. 14)
Pisara semirufa Hampson, 1894, Fauna of British India, Moths 2: 146. Type-locality: [India] Sikkim. Holotype: w, in coll. BMNH.
Material examined:
Thailand. Prov. Chiang Mai: 2 xx, 4 km S of Kop Dong, 99˚03’E, 19˚52’N, 1800 m, 6.xi.2002, slide No. LGN 987 (W 8454) (coll.
MWM), leg. b. HeRCzig & g. Ronkay. Prov. Nan: 1 x, Doi Phuka N.P., between Pua and Bo Luang, 1350 m, 101˚05’E, 19˚12’N,
3.xi.2002 leg. b. HeRCzig & g. Ronkay (coll. HNHM). Vietnam. 1 x, Mt. Fan-si-pan, Sa-Pa, 1600-1800 m, 22º20’N, 103º40’E, 10.vi.-
6.vii.1994, leg. sCHintLmeisteR, slide No. LGN 1162 (W 8464) (coll. MWM).
Meganola latiscripta LászLó, Ronkay & Witt, 2005
(Pl. 11, g. 15)
Meganola latiscripta LászLó, Ronkay & Witt, 2005, Entomofauna 26(11): 219. Type locality: Vietnam, Bach-Ma. Holotype: x, in
coll. MWM.
Type material examined:
Holotype. x, „Vietnam mer., BACH-MA Nat. Park, 1200 m, 16°10’N, 107°54’E, 26. 7.-6. 8. 1996, leg. v. siniaev & E. Afonin”,
slide No. LGN 160 (W 8287) (coll. MWM).
Paratypes. Vietnam. 2 xx, 1 w, with the same data as the holotype, slide No. LGN 161 (W 8288) (coll. MWM). China. Hainan: 1
w, Wuzhi Shan Mts, 18°53’N, 109°43’E, 1500 m, February-April 2001, local collector leg., slide No. LGN 770 (coll. HNHM).
Additional material examined:
Thailand. Prov. Chiang Mai: 1 x, Mt. Doi Inthanon, NP, 2300 m, 17.x.2000, leg. local collector, slide No. LGN 1249 (W 8453) (coll.
MWM).
Meganola indistincta (Hampson, 1894)
(Pl. 5, g. 21)
Selca indistincta Hampson, 1894, Fauna of British India, Moths 2: 147. Type-locality: [India] [Nagaland] Naga Hills. Holotype: x,
in coll. BMNH.
40
Material examined:
Thailand. Prov. Chiang Mai: 6 xx, between Fang and Nor Lae, 99º06’E, 20º02’N, 28.x.2002, slide No. LGN 985 (W 8463) (x); 3
xx, from the same site, 12.xi.2002, leg. b. HeRCzig & g. Ronkay. Prov. Mae Hong Song: 1 x, between Pa Pae and Khun Sa, 1250
m, 98º39’E, 19º08’N, leg. b. HeRCzig & g. Ronkay, slide No. LGN 1079 (coll. G. Ronkay and MWM).
Meganola tarkabarka spec. nov.
(Pl. 6, g. 1; gen. g. 34)
Holotype. x, „Thailand, Changwat Nan, 25 km N of Bo Luang, 1150m, 17.ii.1998, leg. Márton Hreblay & Csaba Szabóky”, slide
No. LGN 614 (W 8452) (coll. MWM).
Diagnosis. The new species belongs to the Meganola semirufa – M. indistincta species group, being externally
somewhat similar to M. semirufa due to its reddish-brown forewing. The specic features of M. tarkabarka are the
presence of a dark grey costal patch and grey spots in the terminal area which are absent in M. semirufa, and the
reduced transverse lines which are distinctly marked in M. semirufa.
The differences of the x genitalia of the two species are easily recognisable. The new species has, in comparison
with M. semirufa, longer and much narrower, apically pointed and hairless uncus, conspicuously shorter and nar-
rower, tongue-shaped apical valval lobe, much shorter, more robust harpe and much shorter and thicker aedeagus.
The uncus of M. semirufa is shorter, apically truncate, laterally covered with dense, relatively long hairs, the valval
lobe is much larger, broadly rounded, the harpe is thinner but longer, and the aedeagus is longer, slenderer than
in M. tarkabarka. The w is unknown.
Distribution. The new species is known from the type-locality only.
Meganola hollowayi spec. nov.
(Pl. 6, g. 2; gen. g. 35)
Holotype. x, “N. Thailand, Chiang Mai Prov., between Chiang Dao and Kariang, 900 m, 98°48’E, 19°25’N, 26.x.2002, leg. b.
Herczig & G. Ronkay”, slide No. LGN 1052 (coll. G. Ronkay).
Diagnosis. Meganola hollowayi is externally slightly similar to M. semirufa but lacks the reddish colouration from
the forewing. The new species resembles also M. indistincta, but has much paler forewing ground colour and
weaker crosslines.
The conguration of the x genitalia of M. hollowayi displays the close relationship of the new species with M.
indistincta, the main differences between the two taxa are as follows: M. hollowayi has somewhat shorter, slightly
broader uncus, with much longer lateral hair-cover, longer, apically tapering valval lobe, shorter harpe which is
situated more closely to the base of valva, the aedeagus has much narrower, rather elongated proximal end, and
more sclerotized, apically slightly lance-like distal end. The w is unknown.
Distribution. The unique known specimen was collected in Northern Thailand.
Meganola mediana spec. nov.
(Pl. 6, gs 3-4; gen. g. 36)
Holotype. x, „Thailand, Changwat Chiang Mai, Mt. Doi Phahompok 16 km NW of Fang, 2000m, 27.iii.1998, leg. Tibor Csővári &
Pál Stéger”, slide No. LGN 1098 (W 8462) (coll. MWM).
Paratypes. Thailand. Prov. Chiang Mai: 1 w, with the same data as the holotype, slide No. LGN 1099 (W 8451) (coll. MWM); 1 w,
Chiang Dao, 1450 m, San Pakia RFD Watershed Station, 29.iv.-1.v.1994, leg. i.J. kitCHing et al., BMNH 1994-97.
slide No. LGN 1347 (BM Arct. 6219) (coll. BMNH).
41
Diagnosis. Meganola mediana is most similar externally to M. indistincta but their external differences are rather
large. The new species is larger in size (wingspan and length of forewing of M. mediana are 19-20 mm and 9-10
mm, those of M. indistincta are 14-15 mm and 7-8 mm, respectively), the forewing ground colour is paler and
the median area is considerably more darkened than in M. indistincta. The hindwing of the new species is also
mentionably darker, more unicolorous than in its close relative.
In the x genitalia, the new species has much thicker, slightly shorter and much more hairy uncus, much broader
apical valval lobe, conspicuously shorter, thinner, slightly but evenly curved harpe, being much closer to the base of
the valva than in M. indistincta. The aedeagus of the new species is considerably longer than that of M. indistincta,
having elongate-tapering proximal end.
The w genitalia of the new species are typical of most Meganola species having a pair of strongly sclerotized,
thorn-like signum bursae. The characteristic features of the new species are as follows: apophyses rather short,
8th segment very short, ostium bursae short and conspicuously funnel-like, ductus bursae relatively long and nar-
row, membranous, cervix bursae slightly swollen, corpus bursae large, ovoid, with well-developed membranous
appendix bursae and with a pair of strongly sclerotized thorn-like signum bursae surrounded by ne scobination.
Distribution. Northern Thailand.
Meganola galsworthyi spec. nov.
(Pl. 6, g. 5; gen. g. 37)
Holotype. x, „Thailand, Changwat Chiang Mai, 4 km SE of Pang Faen, 1100m, 13.i.2004, leg. Hentschel & Szabó”, slide No. LGN
1022 (W 8461) (coll. MWM).
Paratype. Thailand. Prov. Chiang Mai: x, between Chiang Dao and Kariang, 900 m, 98°48’E, 19°25’N, 26.x.2002, leg. b. HeRCzig
& g. Ronkay (coll. G. Ronkay).
Diagnosis. Meganola galsworthyi resembles externally certain Nola species (e.g. N. atrocincta inoue, 1998, N.
ochrolopha (Hampson, 1911) or N. defreina spec. nov.) having rather elongated, apically acute forewing with dark
grey median area closed sharply by the postmedial line. Despite this external similarity, the new species belongs
to the genus Meganola, according to the typical features of the x genitalia, its closest relatives are the members of
the M. indistincta – M. semirufa group. The x genitalia of the new species differ from those of the related species
(M. indistincta, M. semirufa, M. hollowayi, M. mediana, M. tarkabarka) by the shortest, thickest uncus covered with
dense long hairs, and the longest, slightly curved, distally slightly dilated apical lobe of valva. The basal lobe of the
valva is rather narrow-triangular, that of the related species are much broader or rounded; the harpe is right-angled
medially, like in M. indistincta and M. hollowayi, but is somewhat shorter and thicker. The relatively short, simple
aedeagus of the new species is very similar to that of M. indistincta by its broad-quadrangular proximal end, but
the apical part is rounded, while the aedeagus of the related species is apically pointed. The w is unknown.
Distribution. The new species has been recorded twice in Northern Thailand.
Meganola honeyi spec. nov.
(Pl. 6, g. 6; gen. g. 38)
Holotype. x, „North Thailand, Prov. Chiang Mai, 1800m, 4 km S of Kop Dong, 99º03’E, 19º52’N, 06. 11. 2002, leg. B. HeRCzig et
G. Ronkay”, slide No. LGN 1024 (coll. G. Ronkay).
Diagnosis. Meganola honeyi resembles externally M. indistincta by its pale grey forewing ground colour and darker
grey wing pattern, but is smaller in size (wingspan and length of forewing 13 mm and 6 mm, those of M. indistincta
14-15 mm and 7-8 mm, respectively), the basal area of its forewing is whitish grey (that of M. indistincta is dark grey)
with much narrower an paler median area in comparison with the related species. The x genitalia also support the
close relationship of M. honeyi and M. indistincta within the species group, according to the conspicuously narrow
tegumen and uncus and the similar shape of valva. The differences between the two taxa are clearly recognisable,
as the new species has much shorter uncus, apically and basally more dilated valva, and only slightly curved harpe,
42
and narrower and more elongated aedeagus. The harpe of M. indistincta is characteristically curved medially in a
right angle and straight apically. The w is unknown.
Distribution. Northern Thailand.
Meganola benescripta spec. nov.
(Pl. 6, gs 7-8; gen. g. 39)
Holotype. x, “N. Thailand, Chiang Mai Prov., between Chiang Dao and Kariang, 900 m, 98°48’E, 19°25’N, 26.x.2002, leg. B.
HeRCzig & g. Ronkay”, slide No. LGN 1078 (coll. G. Ronkay).
Paratype. Thailand. Prov. Chiang Mai: w, with the same data as the holotype, slide No. LGN 1076 (W 8450) (coll. MWM).
Diagnosis. The new species has a unique external appearance within the genus Meganola by its graphite-grey
forewing ground colour and the ne wing pattern consisting of sharply dened, but thin crosslines and a row of
blackish dots in the terminal area. This forewing colouration and wing pattern appear mainly in the genus Nola,
especially in the old sense Pisara species.
Meganola benescripta is a close relative of M. geoffmartini, these two species represent, according to the features
of their x genitalia, a distinct lineage, which is characterized by the conspicuous dentation on the outer margin
of the proximal lobe of the bilobate valva, this feature is absent from the other known Meganola s.l. species. The
new species differs from its sister species, M. geoffmartini, by its much longer, narrow, slightly arched, apically
pointed uncus, shorter and basally somewhat broader distal and somewhat shorter, apically more pointed proximal
lobe of the valva, more dense dentation of the outer edge of the valva consisting of smaller teeth. The harpe of M.
benescripta is much shorter than that of M. geoffmartini, broader at base, than strongly tapering, curved medially,
slightly dilated apically with rounded apex, while the harpe of the related species is more robust, rather claw-like,
tapering and arched continuously towards the pointed apex. The aedeagus of M. benescripta is considerably longer
and slightly narrower than that of M. geoffmartini, with more elongated and narrower, stick-like proximal end. The
vesica of both species has extensive, ne scobination.
The w genitalia of the new species is characterized by the relatively long apophyses posteriores and rather short
apophyses anteriores, the very narrow and weakly sclerotized 8th segment, the simple, weak, quadrangular
ostium bursae, the relatively short, but thick ductus bursae, the conspicuously large, globular cervix bursae with
very thin ductus seminalis, and the elongate-ovoid corpus bursae with a relatively small, rounded triangular but
strongly sclerotized signum.
Distribution. The species is known from the type locality only.
Meganola geoffmartini spec. nov.
(Pl. 6, g. 9; gen. g. 40)
Holotype. x, “N. Thailand, Chiang Mai Prov., between Chiang Dao and Kariang, 900 m, 98°48’E, 19°25’N, 26.x.2002, leg. B.
HeRCzig & g. Ronkay”, slide No. LGN 1077 (coll. G. Ronkay).
Paratype. Thailand. Prov. Chiang Mai: x, with the same data as the holotype, slide No. LGN 1053 (W 8460) (coll. MWM).
Diagnosis. M. geoffmartini is the sister species of the above described M. benescripta. The two species are simi-
lar in size (wingspan of both species 14-15 mm, length of forewing 7-8 mm) and the forewing ground colour is
similarly graphite-grey, but the crosslines and the marginal spots are paler and more indistinct in M. geoffmartini.
The x genitalia of the two species differ conspicuously, the distinctive features are given in the diagnosis of M.
benescripta. The w is unknown.
Distribution. The new species occurs sympatrically with its sister species, M. benescripta, in Northern Thailand.
43
Meganola scriptoides HoLLoWay, 2003
(Pl. 6, gs 10-11)
„Meganola” scriptoides HoLLoWay, 2003, The Moths of Borneo 18: 27, pl.1, gs 21, 41. Type-locality: Borneo, Sarawak. Holotype:
x, in coll. BMNH.
Material examined:
Thailand. Prov. Chiang Mai: 5 specimens, 20 km NW of Sop Kha, 2 km S of Kop Dong, 1800 m, 8.i.1999; 4 specimens, from the same
site, 7-8.i.1999; 1 specimen, 6 km SE of Pang Faen, 1100 m, leg. a. szabó & z. CzeRe; 1 specimen, from the same site, 18.i.2004,
leg. p. HentsCHeL & a. szabó; 9 specimens, between Fang and Nor Lae, 1600 m, 99º06’E, 20º02’N, 28.x.2002; 8 specimen, from
the same locality, 7.xi.2002; 1 specimen, between Chiang Dao and Kariang, 900 m, 98º48’E, 19º25’N, 26.x.2002, leg. b. HeRCzig &
g. Ronkay; 1 specimen, Mt. Doi Phahompok, 16 km NW of Fang, 2000 m, 24.ii.1998, leg. m. HRebLay & Cs. szabóky; 1 specimen,
19 km NW of Fang, 1900 m, 26.iii.1998, leg. T. Csővári & P. sTéger; 1 specimen, 7 km W of Pa Pae, 1230 m, 21.xi.1998, leg. t.
Csővári & l. mikus; 3 specimens, 15 km SW of Wiang Haeng, 1400 m, 4.xii.1998, leg. m. HRebLay, i. soós & y. sHeRpani. Prov. Nan:
1 specimen, 30 km E of Pua, 1700 m, 10.xi.1999, leg. m. HRebLay; 4 specimens from the same locality, 20. and 27.ii.1998; 1 speci-
men, from the same site, 21.i.2004, leg. p. HentsCHeL & a. szabó; 1 specimen, 12 km NW of Chiang Dao, 750 m, 12.xi.1998, leg. t.
Csővári & l. mikus; 1 specimen, from the same site, 1.iii.1998, leg. m. HRebLay & Cs. szabóky; 1 specimen, 25 km N of Bo Luang,
1150 m, 11.xi.1999, leg. a. szabó & z. CzeRe; 1 specimen, 4 km W of Pha Lak, 820 m, 28.iii.1998, leg. T. Csővári & P. sTéger; 3
specimens, Doi Phuka NP, between Pua and Bo Luang, 1350 m, 101º05’E, 19º12’N, 3.xi.2002, leg. b. HeRCzig & g. Ronkay, slide
No.LGN 779 (x), LGN 778 (w). Prov. Mae Hong Song: 1 specimen, 21 km NW of Pai, 1360 m, 7.ii.1998; 1 specimen, 1 km S of
Bahundanda, 1000 m, 6.ii.1998, leg. m. HRebLay & Cs. szabóky; 2 specimens, between Pa Pae and Khun Sa, 1250 m, 98º39’E,
19º08’N, leg. b. HeRCzig & g. Ronkay (coll. G. Ronkay & MWM).
Meganola scripta (mooRe, 1888)
(Pl. 6, gs 12-13)
Roeselia scripta mooRe, 1888, Proceedings of the Zoological Society London 1888: 393. Type-locality: India [Himachal Pradesh],
Dharmsala. Holotype: x, in coll. BMNH.
Material examined:
Thailand. Prov. Nan: 2 specimens, 30 km E of Pua, 1700 m, 20.ii.1998, leg. T. Csővári & l. mikus; 1 specimen, from the same
site, 10.xi.1999, leg. m. HRebLay; 2 specimens, from the same locality, 20.i.2004, leg. p. HentsCHeL & a. szabó. Prov. Chiang Mai:
1 specimen, Mt. Doi Phahompok, 16 km NW of Fang, 2000 m, 6-7.viii.1999, leg. T. Csővár & l. mikus; 1 specimen, Mt. Doi Pha-
hompok, 18 km NW of Fang, 2100 m, 15.xi.1999, leg. m. HRebLay; 2 specimens, Mt. Doi Phahompok, 20 km NW of Fang, 2450
m, 24.i.2004; 2 specimens, Mt. Doi Phahompok, 20 km NW of Fang, 2150 m, 22-25.i.2004, leg. a. szabó; 2 specimens, Mt. Doi
Phahompok, 20 km NW of Fang, 2000 m, 15.i.2004, leg. p. HentsCHeL & a. szabó; 2 specimens, 20 km NW of Sop Kha, 2 km S of
Kop Dong, 1800 m, 27.i.1999, leg. a. szabó & z. CzeRe; 1 x, 4 km S of Kop Dong, 99º03’E, 19º52’N, 1800 m, 11.xi.2002, slide No.
LGN 780; 1 specimen, from the same site, 6.xi.2002; 2 specimens, between Fang and Nor Lae, 99º06’E, 20º02’N, 28.x.2002; 2
specimen, from the same locality, 12.xi.2002, leg. b. HeRCzig & g. Ronkay; 1 specimen, Mt. Doi Inthanon, NP, 2300 m, 16.i.2004,
leg. p. HentsCHeL & a. szabó; 1 specimen, 23 km NW of Sop Kha, 1 km E of Kop Dong, 1650 m, 14.i.2004, leg. p. HentsCHeL & a.
szabó (coll. G. Ronkay & MWM).
Meganola implicata LászLó, Ronkay & Witt, 2006
(Pl. 6, gs 14-15)
Meganola implicata LászLó, Ronkay & Witt, 2006, Entomofauna 28(2): 17-32. Type-locality: N. Thailand, Prov. Mae Hong Son,
between Pa Pae and Khun Sa. Holotype: x, in coll. HNHM.
Type material examined:
Holotype. x, „North Thailand, Prov. Mae Hong Son, 1250 m, between Pa Pae and Khun Sa, 98°39’E, 19°08’N, 31.10.2002, leg.
b. HeRCzig et g. Ronkay”, slide No. LGN 777 (coll. HNHM).
Paratypes. Thailand. Prov. Mae Hong Son: 2 xx, with the same data as the holotype. Prov. Nan: 1 w, 30 km E of Pua, 1700 m,
12-13.i.1999, leg. a. szabó & z. CzeRe, slide No. LGN 814 (W 8316); 1 w, from the same site, 21.i.2004, leg. a. szabó & p. Hent-
sCHeL. Prov. Chiang Mai: 1 w, between Fang and Nor Lae, 1600 m, 99°06’E, 20°02’N, 12.xi.2002, leg. b. HeRCzig & g. Ronkay; 1
x, Doi Inthanon, 2300 m, 16.i.2004; 1 x, 1 w, 23 km NW Sop Kha, 1650 m, 14.i.2004, leg. a. szabó & p. HentsCHeL (coll. HNHM
and MWM).
44
Meganola yakovlevi spec. nov.
(Pl. 6, g. 16; gen. g. 41)
Holotype. x, „Thailand, Changwat Chiang Mai, Mt. Doi Phahompok, 16 km NW of Fang, 2000m, 24.ii.1998 leg. Márton HRe-
bLay & Csaba Szabóky”, slide No. LGN 1030 (W 8399) (coll. MWM).
Paratypes. Thailand, Prov. Chiang Mai: 1 x, Mt. Doi Phahompok, 18 km NW of Fang, 2100 m, 9.i.1999, slide No. LGN 1102 (W
8409); 1 x, 20 km NW of Mae Ai, 1650 m, 7.i.1999, leg. a. szabó & z. CzeRe; 8 xx, Mt. Doi Phahompok, 20
km NW of Fang, 2100 m, 24.i.2004; 6 xx, from the same site, 22-25.i.2004; 1 x, 4 km SE of Pang Faen, 1100
m, 26.i.2004, leg. a. szabó. Vietnam, Fan-si-pan Mts: 2 xx, Sa Pa, 1600 m, 22º17’N, 103º44’E, 20-30.x.1995,
leg. sinjaev & afonin, slide Nos LGN 823 (W 8398), LGN 877 (W 8408); 2 ww, from the same locality, 1525 m,
28.X-3.xi.1994, leg. sinjaev & local collector, slide Nos LGN 824 (W 8397), LGN 873 (W 8407); 2 xx, 1 w, Sa
Pa, 22º15’N, 103º45’E, 2400 m, 2-4.iii.1995, leg. DR. R. bReCHLin, slide Nos LGN 874 (W 8396), LGN 875 (W
8406) (xx), LGN 876 (W 8395) (w) (coll. MWM); 1 w, Sa Pa, Prov. Lao-Cai, 1650 m, lieux herb. Second., Á la
lumiére, 24.ix.1963, leg. t. póCs (coll. HNHM).
Diagnosis. Meganola yakovlevi belongs to the M. scripta-M. implicata group, being externally similar to M. scripta,
but is easily distinguishable by its considerably larger size (wingspan and length of forewing 19-25 and 9-12 mm,
those of M. scripta 17-20 mm and 8-10 mm, respectively), paler, more whitish grey forewing ground colour, the
absence of the conspicuous blackish apical and tornal spots (which are typical of M. scripta), and the more curved
upper and the much more sinuous lower section of the postmedial line.
The specic differences are even more prominent in the conguration of the genitalia in both sexes. In the x geni-
talia, the new species has slenderer uncus, being thicker only in the basal third, then tapering, forming a strong
thorn-like apical part, a pair of ne, but well developed, nger-like, apically hairy subscaphial processes, a relatively
large and more or less trapezoidal fultura inferior, a slightly dilated distal valval lobe with almost parallel margins,
a longer, more robust harpe, being directed caudally and a considerably longer vinculum. The uncus of M. scripta
is rather broad in the basal two-third, only its apical third is thorn-like, the subscaphial processes are absent, the
fultura inferior is much smaller, cordiform, the distal lobe of valva is much broader, the proximal lobe of valva is
remarkably longer, the harpe is much shorter, directed ventrally and the vinculum is signicantly shorter than in
M. yakovlevi. The elongate, relatively narrow aedeagus of the new species is slightly sinuous medially, that of M.
scripta is continuously slightly arched; the characteristically elongated, rather thin carinal process is somewhat
longer in the new species. The vesica of M. yakovlevi is armed with one rather short, but strong thorn-like cornutus,
while M. scripta has a somewhat thinner but ve times longer cornutus.
The ovipositor of the new species is more conical, the 8th segment is somewhat longer when comparing with
those of M. scripta. The ostium bursae of M. yakovlevi is somewhat shorter and thicker, the lateral process is much
shorter, rather hump-like, and the two signa are fused into an elongated and scobinated plate, forming a short,
thick, arrow-like structure. The lateral process of the ductus bursae is characteristically very long in M. scripta, and
the two triangular signa are not fused.
Distribution. The new species is distributed in the northern mountainous areas of Indochina (Northern Thailand
and the Fan-si-pan region in North Vietnam).
Meganola spec. prope exuosa (pouJaDe, 1886)
(Pl. 6, gs 17-18)
Material examined:
Thailand. Prov. Mae Hong Song: 1 x, 10 km NE of Pai, 1560 m, 28.i.1999, leg. a. szabó & z. CzeRe, slide No. LGN 1100 (W 8405);
1 x, 1 km S of Bahundanda, 1000 m, 6.ii.1998, leg. m. HRebLay & Cs. szabóky, slide No. LGN 1014 (W 8403). Prov. Chiang Mai: 1
x, 2 ww, 20 km NW of Sop Kha, 2 km S of Kop Dong, 1800 m, 7.i.1999, leg. a. szabó & z. CzeRe, slide Nos LGN 1130 (W 8404)
(x), LGN 1101 (W 8394), LGN 1015 (W 8392) (ww); 1 x, between Fang and Nor Lae, 99º06’E, 20º02’N, 28.x.2002, slide No.
LGN 1131 (W 8391); 1 x, from the same site, 7.xi.2002 (LGN 1067), leg. b. HeRCzig & g. Ronkay; 1 x, 10 km E of Mae Chaem,
1160 m, 5.ii.1998, leg. m. HRebLay & Cs. szabóky, slide No. LGN 1013 (W 8393). Prov. Nan: 1 x, 25 km N of Bo Luang, 1150 m,
17.ii.1998, leg. m. HRebLay & Cs. szabóky, slide No. LGN 1113 (W 8402) (coll. G. Ronkay and MWM).
Remarks. The type of „Nola” exuosa, described from Sichuan, has not been examined yet. The species collected
in Thailand can be conspecic with M. exuosa according to the original description. Inoue (1998) published a single
w specimen from Nepal as Rynchopalpus exuosa, which is rather similar in wing pattern to the Thai specimens.
Thus, the identication of the Thai specimens is still tentative, according to the limited and unconrmed information,
the satisfactory identication requires the examination of the type specimen preserved in MNHN, Paris.
45
Meganola subnitida LászLó, Ronkay & Witt, 2007
(Pl. 5, g. 18)
Meganola subnitida LászLó, Ronkay & Witt, 2007 Entomofauna 28(14): 160. Type-locality: Thailand, Prov. Nan, Doi Phuka. Holo-
type: x, in coll. MWM.
Type material examined:
Holotype. x, „N. Thailand, Prov. Nan, Doi Phuka NP, between Pua and Bo Luang, 1350 m, 101°05’E, 19°12’N, 3. XI. 2002, leg. b.
Herczig & G. Ronkay”, slide No. LGN 925 (W 8353) (coll. MWM).
Paratypes. Thailand. Prov. Nan: 1 x, with the same data as the holotype. Prov. Chiang Mai: 1 x, between Chiang Dao and Kar-
iang, 900 m, 98°48’E, 19°25’N, 26.x.2002, slide No. LGN 879, leg. b. HeRCzig & g. Ronkay; 3 xx, 20 km NW of Mae Ai, 1650 m,
9.ix.1999, leg. a. szabó & z. CzeRe, slide Nos LGN 878 (W 8355), LGN 924 (W 8354). Malaysia. 5 xx, Pahang State, Cameron
Highlands, Tanah Rata, 21.iii.–2.iv.1995, ex coll. g. Ronkay, slide Nos LGN 264, LGN 782, LGN 763 (W 8357), LGN 826 (W 8356)
(coll. MWM and HNHM).
Meganola ascripta (Hampson, 1894)
(Pl. 6, g. 19; gen. g. 42)
Selca ascripta Hampson, 1894, Fauna of British India, Moths 2: 144. Type-locality: [India] [Nagaland] Naga Hills. Holotype: x, in
coll. BMNH.
Material examined:
Thailand. Prov. Nan: 2 xx, Nan, 5 km N of Bo Luang, 1000 m, 12.xi.1999; 1 x, 25 km N of Bo Luang, 1150 m, 11.xi.1999, leg. m.
HRebLay & i. soós; 1 x, 30 km E of Pua, 1700 m, 27.ii.1998, leg. m. HRebLay & Cs. szabóky; 1 x, Doi Phuka NP, between Pua and
Bo Luang, 1350 m, 101º05’E, 19º12’N, 3.xi.2002, slide No. LGN 754, leg. b. HeRCzig & g. Ronkay. Prov. Chiang Mai: 1 x, 4 km
S of Kop Dong, 99º03’E, 19º52’N, 1800 m, 11.xi.2002, slide No. LGN 755; 1 x, from the same site, 29-30.x.2002, slide No. LGN
775, leg. b. HeRCzig & g. Ronkay (coll. MWM and G. Ronkay). India. Assam, Khasis: 1 w, 5000 ft, Shillong 6.ix.1927, FLetCHeR coll.,
slide No. LGN 1325 (BM Arct. 6190) (coll. BMNH).
Description of the w genitalia. Meganola ascripta has long been known only by the x sex. The species was de-
scribed based on the x holotype, and the recent expeditions to Indochina had no success in collecting a w speci-
men. Fortunately, during the detailed examination of the accessory material of the Nolidae collection of the BMNH,
the authors found a beautiful w specimen of M. ascripta collected in Khasis in 1927. Thus, the w of the species is
gured rstly in the present paper, and the w genitalia is described here.
Ovipositor simple, conical, apophyses posteriores medium-long, slightly arched. 8th tergite rather short, ribbon-
like, apophyses anteriores relatively short, straight. Ostium bursae rather thin, slightly cup-shaped; ductus bursae
slender, medium-long, almost straight, strongly sclerotized. Cervix bursae well-developed but short, sac-like; corpus
bursae relatively large, ovoidal, appendix bursae arisen at the proximal end of corpus bursae, rather membraneous,
relatively small, globular. Signum bursae large, strongly sclerotized, broadly triangular.
Meganola tarka spec. nov.
(Pl. 11, gs 16-17; gen. g. 46)
Holotype. x, „SE. Thailand: 300m, Chantaburi, Khao Soi Dao 9.viii.1987 Col. m.g. Allen, B.M. 1987-364”, slide No. LGN 1324
(BM Arct. 6189) (coll. BMNH).
Paratypes. Indonesia. SW Sulawesi (Celebes): 1 x, Pangean, near Maros, 2,000 ft. March 1938, J.p.a. kaLis, B.M. 1938-397, slide
No. LGN 1326 (BM Arct. 6191); 3 xx, Tjamba, near Maros, 1,500 ft. February 1938. J.p.a. kaLis, B.M. 1938-310.
W Sulawesi (Celebes): 1 w, Lindoe Paloe, 3700’, April 1937 J.p.a. kaLis, RotsCHiLD Bequest B.M. 1939-1, slide
No. LGN 1357 (BM Arct. 6201); 1 x, Paloe, G. Rangkoenau, 1,800’ Dec. 1936, J.p.a. kaLis, RotsCHiLD Bequest
B.M. 1939-1. Sumatra: 1 x, Prapat, HW 3, III-IV.1982, leg. DR. DieHL, slide No. LGN 1358 (coll. HNHM); 1 x,
Huta Padang, 310 m, 99º11’E, 2º48’N, 15.iii.1993 leg. R. bReCHLin, slide No. LGN 561 (W 7308). Malaysia. 1 w,
Kuala Lumpur, Malay Peninsula C. Boden Kloss, RotsCHiLD Bequest, B.M. 1939-1, slide No. LGN 1355 (BM Arct.
6202). Philippines. 1 w, Montalban, Luzon, 15. January 1914, a.e. WiLeman, RotsCHiLD Bequest B.M. 1939-1, slide
No. LGN 1356 (BM Arct. 6203) (coll. BMNH). India. Assam: 1 x, Garampani, Nambor Reserve Forest, 26º00’N,
46
94º20’E 100 m, 21-29.xi.1997, leg. sinjaev & murzin, slide No. LGN 153 (W 7309) (coll. MWM).
Diagnosis. The external appearance of the new species is rather unique within the subfamily, no similar species
is known. The main external characters of M. tarka are as follows: the antenna is bipectinated in both sexes, the
forewing ground colour is pale yellowish brown with whitish-yellow basal area and darker, more blackish median
area. The scattered darker elements of the forewing pattern gives a strongly variegated appearance of the moths.
The postmedial line is very ne, sharply dened, characteristically sinuous, the other crosslines are rather diffuse,
shadow-like or deleted.
The genitalia structures indicate the close relationship between M. tarka and M. ascripta. The distinctive features
in the x genitalia are as follows: the new species has somewhat shorter uncus, with much broader basal part,
straight and apically slightly dilated valva, and much thinner and shorter, apically pointed harpe with variably strong
dentation on distal margin at its basal third. The uncus of M. ascripta is evenly arcuate, apically slightly tapering,
and the harpe is much more robust, apically dilated, with ne dentation along the whole distal margin. The shape
and structure of the fultura inferior, the vinculum and the aedeagus of the two species show no mentionable dif-
ferences (see the Figs 42 and 46).
The w genitalia of M. tarka and M. ascripta are also similar, with clearly recognisable specic differences. The new
species has much broader, elongated, slightly arched intersegmental sclerotization between the papillae anales
and the 8th tergite, and the signum is represented by a large, long sclerotized bar with two rather long, nger-like
processes. The intersegmental sclerotization of the ovipositor of M. ascripta is much shorter, button-like, and the
signum is triangular with a single process.
Distribution. A widely distributed species, its range extending from the Sundaland and the Philippines to northern
Indochina.
Meganola cuneifera (WaLkeR, 1862)
(Pl. 6, g. 20)
Melia cuneifera WaLkeR, 1862, Journal of the Proceedings of the Linnean Society of London (Zoology) 6: 127. Type-locality: Borneo,
Sarawak. Holotype: x, in coll. UM Oxford.
Material examined:
Thailand. Prov. Nan: 1 x, 5 km N of Ban Luang, 350 m, between Pi Nai and Pi Tai, 100º27’E, 18º56’N, 4.xi.2002, leg. b. HeRCzig
& g. Ronkay, slide No. LGN 771. Prov. Chiang Mai: 1 x, between Chiang Dao and Kariang, 900 m, 98º48’E, 19º25’N, 26.x.2002,
leg. b. HeRCzig & g. Ronkay, slide No. LGN 758 (coll. G. Ronkay).
Meganola rucostata (Hampson, 1896)
(Pl. 6, g. 21, Pl. 7, g. 1)
Selca rucostata Hampson, 1896, Fauna of British India, Moths 4: 507. Type-locality: Bhutan. Holotype: w, in coll. BMNH.
Material examined:
Thailand. Prov. Chiang Mai: 12 specimens, between Chiang Dao and Kariang, 900 m, 98º48’E, 19º25’N, 26.x.2002, slide Nos LGN
774, LGN 773, LGN 759 (xx), LGN 772 (w), leg. b. HeRCzig & g. Ronkay. Prov. Mae Hong Son: 2 specimens, between Pa Pae and
Khun Sa, 1250 m, 98º39’E, 19º08’N, 31.x.2002, leg. b. HeRCzig & g. Ronkay (coll. G. Ronkay and MWM).
Meganola trianguloquelinea (eeCke, 1920) comb. nov.
(Pl. 11, gs 18-19; gen. g. 47)
Roeselia trianguloquelinea eeCke, 1920, Zoologische Mededeelingen Leiden 5: 45, g. 11. Type-locality: [Indonesia] Java, Bajor.
Holotype: w, in coll. RNH Leiden.
47
Type material examined:
Holotype of Roeselia trianguloquelinea eeCke, 1920: w, „Java, Bajor”, slide No. LGN 859 (coll. RNH, Leiden).
Additional material examined:
Thailand. 1 x, Chantaburi Dist., 300 m, Khao Soi Dao NP, 23.xi.1987, m.g. aLLen, BM 1988-162, slide No. LGN 1333 (BM Arct.
6210) (coll. BMNH); 1 w, Khao-Yai, 20-24.viii.1990, leg. k.t. paRk, slide No. LGN 1334 (coll. HNHM).
Description of the x genitalia. Uncus rather short and broad, distally tapering and apically rounded, covered with
dense, ne hairs. Subscaphium well-developed, ribbon-like, apically rounded, with a dense row of strong spines.
Tegumen long, medial part with distinct, triangular lobe; fultura inferior relatively small, V-shaped; vinculum rather
short, pointed. Valva long, slightly curved, medially tapering, apically broadly rounded; saccular lobe well-developed,
elongated, apically rounded, its dorsal margin covered by a row of gradually increasing, strongly sclerotized robust
spines. Aedeagus medium-long, straight, apically pointed; vesica with a narrow, slightly arcuate, somewhat funnel-
like and less sclerotized cornutus, and with a small cornuti eld consisting of rather short, ne cornuti.
Meganola calligrapha LászLó, Ronkay & Witt, 2005
(Pl. 7, gs 2-3)
Meganola calligrapha LászLó, Ronkay & Witt, 2005, Entomofauna 26(11): 208. Type locality: N. Thailand, Prov. Chiang Mai, between
Chiang Dao and Kariang. Holotype: x, in coll. MWM.
Type material examined:
Holotype. x, “N. Thailand, Chiang Mai Prov., between Chiang Dao and Kariang, 900 m, 98°48’E, 19°25’N, 26.x.2002, leg. B. Herczig
& G. Ronkay”, slide No. LGN 757 (W 8277) (coll. MWM).
Paratypes. Thailand. Prov. Chiang Mai: 1 x, with the same data as the holotype, slide No. LGN 756; 1 x, from the same site,
8.xi.2002, leg. b. HeRCzig & g. Ronkay; 1 x, Mt. Doi Phahompok, 16 km NW of Fang, 2000 m, 19.i.1999, leg. a. szabó & z. CzeRe;
1 x, 7 km W of Pa Pae, 1230 m, 27.xi.1998, leg. T. Csővári & l. mikus; 1 w, 15 km SW Wiang Haeng, 1400 m, 9.ii.1998, leg. m.
HRebLay & Cs. szabóky. Prov. Nan: 1 x, 22 km N of Bo Luang, 1120 m, 24.i.1999, leg. a. szabó & z. CzeRe, slide No. LGN 607
(W 8278). Prov. Mae Hong Song: 1 x, 1 w, 10 km NE of Pai, 1560 m, 28.i.1999 leg. a. szabó & z. CzeRe, slide No. LGN 608 (W
8279); 1 specimen, 10 km E of Mae Chaem, 1160 m, 5.ii.1998, leg. m. HRebLay & Cs. szabóky (coll. G. Ronkay and MWM). Nepal.
Annapurna Himal: 1 x, Sudame, 1250 m, 24-25.iii.1995, 83°45’E, 28°20’N, slide No. LGN 609 (W 8280); 1 w, Dhumre, Bhimal
Nager, 500 m, 26-28.iii.1995, 84°26’E, 27°55’N, leg. m. HRebLay & L. németH, slide No. LGN 610 (W 8281) (coll. MWM). Taiwan.
Nantou County: 2 xx, 3 km E of Tili, 555 m, 8-9.ii.1997, leg. s. simonyi & p. stégeR, slide Nos LGN 612 (W 8282), LGN 805 (W
8283). Pingtung County: 1 x, Huang Lion Forest Recreation Area, 210 m, 6.iii.1996, leg. gy. Fábián & L. németH, slide No. LGN
605 (W 8284). Taoyuan County: 1 x, 16 km E of Fuhsing, 900 m, 121°27’E, 24°50’N, 30.xi.-1.xii.1997, leg. s. simonyi & a. szabó,
slide No. LGN 806 (W 8285) (coll. MWM). Indonesia. Borneo: 8 specimens, Sarawak Gunong Mulu Nat. Park, G. Api, 900 m; 2
specimens, Gunong Mulu Nat. Park, Mulu, 1000 m (coll. BMNH).
Meganola zolotuhini spec. nov.
(Pl. 7, gs 4-5; gen. g. 43)
Holotype: x, „North Thailand, Prov. Chiang Mai, 1600m, between Fang and Nor Lae, 99º06’E, 20º02’N, 28.x.2002, leg. B. Herczig
et G. Ronkay”, slide No. LGN 798 (W 8401) (coll. MWM).
Paratypes. Thailand. Prov. Chiang Mai: 1 x, between Fang and Nor Lae, 1600 m, 99º06’E, 20º02’N, 7.xi.2002, slide No. LGN 799;
1 w, 4 km S of Kop Dong, 99º03’E, 19º52’N, 1800 m, 6.xi.2002, slide No. LGN 800, leg. b. HeRCzig & g. Ronkay; 1
x, Mt. Doi Inthanon, NP, 2300 m, 21.i.1999, slide No. LGN 1112 (W 8390); 1 x, 1 w, 20 km NW of Mae Ai, 1650
m, 26.i.1999, slide Nos LGN 1137 (W 8400) (x), LGN 1123 (W 8379) (w); 2 ww, 20 km NW of Sop Kha, 2 km S of
Kop Dong, 1800 m, 27.i.1999, leg. a. szabó & z. CzeRe. Prov. Mae Hong Song: 2 ww, 1 km S of Bahundanda, 1000
m, 6.ii.1998, leg. m. HRebLay & Cs. szabóky. Prov. Nan: 1 x, 30 km E of Pua, 1700 m, 25.i.1999, leg. a. szabó & z.
CzeRe, slide No. LGN 807 (W 8389); 1 w, 5 km N of Bo Luang, 1000 m, 10.ii.2000, leg. m. HRebLay, slide No. LGN
1031 (W 8378); 3 ww, from the same site, 4.ii.2000, leg. m. HRebLay & a. szabó, slide Nos LGN 1111 (W 8388),
LGN 1115 (W 8377). Vietnam. 2 xx, Bach-Ma National Park, 1200 m, 16º10’N, 107º54’E, 26.vii.-6.viii.1996, leg.
48
v. sinjaev & e. afonin, slide Nos LGN 811 (W 8387), LGN 812 (W 8376). Laos. 1 x, Phu Soai Dao, 101º09’E,
18º30’N, 6-8.viii.1996, leg. steinke & LeHmann, slide No. LGN 810 (W 8386). Taiwan. Taitung County: 1 w, 3 km
W of Hungyeh, 350 m, 120º52’E, 22º28’N, 5.vii.1997, slide No. LGN 816 (W 8375); 1 w, 7 km E of Taimali, 710
m, 4.vii.1997, 120º53’E, 22º35’N, leg. T. Csővári & l. mikus; 3 xx, 7 km N of Tupan, 120º52’E, 22º29’N, 500
m, 16.xii.1997, slide Nos LGN 803 (W 8385), LGN 818 (W 8374); 1 x, same site, 9-10.xii.1997, leg. s. simonyi
& a. szabó. Pingtung County: 7 xx, on the Road No. 199, 500 m, 22º14’38”N, 120º51’51”E, 19.iv.1997, leg. L.
PeregoviTs & a. kun, slide Nos LGN 819, LGN 801 (W 8384), LGN 802 (W 8373); 1 w, 10 km E of Mutan, 400
m, 5.vii.1997, leg. b. HerCzig & s. kováCs, slide No. LGN 804 (W 8383); 1 x, from the same locality, 7-8.iv.1997,
leg. G. CsoRba & L. Ronkay; 1 x, from the same site, 12.vi.1997, leg. b. HeRCzig & L. Ronkay; 2 ww, 10 km SE of
Mutan, 470 m, 5.xii.1997, leg. gy. Fábián (coll. MWM, G. Ronkay and HNHM).
Diagnosis. The new species resembles externally Meganola calligrapha but easily distinguishable by its slightly
darker, more brownish forewing ground colour, the absence of the darkened longitudinal band of the forewing
(being typical of M. calligrapha) and the presence of a rather conspicuous, slightly elongate blackish patch in the
lower third of the forewing in the xx which is absent from M. calligrapha.
The differences between the x genitalia of the two species are rather large. The valva of M. zolotuhini is some-
what longer, the basal plate of the harpe is much more elongated, the erect part of the harpe is somewhat longer,
narrower at base but thicker apically; in addition, the sacculus lacks the long, nger-like process which is present
in M. calligrapha. The aedeagus of the new species is somewhat shorter, medially more curved than that of M.
calligrapha; the apex of aedeagus is pointed in M. zolotuhini, while truncate in M. calligrapha. The armature of the
vesica is also different: the new species has a single, basally rather broad, apically elongated and acute cornutus,
while the cornutus of M. calligrapha is much shorter and more robust.
The w genitalia of M. zolotuhini differ from those of M. calligrapha by the following features: apophyses anteriores
are slightly longer, apically pointed, the lobes of the 8th sternite are slightly shorter and narrower, the funnel-like
ostium narrower, less sclerotized, and the ductus bursae is much longer and entirely membraneous. The anterior
apophyses of M. calligrapha are apically rounded, the funnel-like ostium is much broader, strongly sclerotized,
and the ductus bursae is strongly sclerotized in its distal two-thirds. The corpus bursae of the new species is more
elongated and the two signa are more remote from each other than in M. calligrapha.
Distribution. The new spoecies is distributed in the northern mountainous areas of Thailand, Laos and Vietnam
and the southern, more subtropical and rather low regions of Taiwan.
Meganola postmediana spec. nov.
(Pl. 7, g. 6; gen. g. 44)
Holotype. x, “N. Thailand, Chiang Mai Prov., between Chiang Dao and Kariang, 900 m, 98°48’E, 19°25’N, 26.x.2002, leg. B.
Herczig & G. Ronkay” slide No. LGN 986 (W 8372) (coll. MWM).
Paratype. 1 w, NE Thailand, 750 m, Chaiyaphum, Phu KhieoWS, km 14 on road to HQ, 6.xi.1991, leg. i.J. kitCHing & a.m. Cotton
BM 1992-9, slide No. LGN 1294 (BM Arct. 6114)) (coll. BMNH).
Diagnosis. The closest relative of the new species, despite the conspicuous external differences, is M. rucostata.
Meganola postmediana is somewhat smaller in size (wingspan and length of forewing 17 and 8 mm, those of M.
rucostata 18-25 mm and 9-12 mm, respectively), having dark grey costal area in the basal half of forewing, and
the crosslines are more indistinct, broader and less sinuous. The forewing costal area of M. rucostata is dark
red-brown and crosslines are more sharply dened, much ner and more sinuous.
The x genitalia of the two species show remarkable differences but the basic structure is the same, displaying
their close relationship. Comparing their genitalia, M. postmediana has narrower, apically rounded uncus, longer,
medially narrower, apically dilated and broadly rounded valva, much longer, slightly narrower and more arched
harpe, without dentation on its caudal margin, the aedeagus is somewhat more arched, and the vesica is armed
with a large, elongated cornutus. The uncus of M. rucostata is more robust at base, apically acutely pointed,
the valva is shorter, gradually tapering, apically narrowly rounded, the harpe has well-discernible dentation on its
caudal margin, and the vesica lacks the cornutus.
In the w genitalia, the ovipositor, the ostium and the ductus bursae are in general similar, the specic differences
49
can be found in the shape and size of the corpus bursae and the signa: the new species has much smaller corpus
bursae and a pair of more robust, rather triangular signa. The proximal margin of the 8th sternite is also somewhat
different in the two species: it is slightly wavy in M. postmediana while is straight in M. rucostata.
Distribution. Northern Thailand.
Meganola triangulalis (LeeCH, 1890)
(Pl. 7, gs 8-9)
Nola triangulalis LeeCH, 1890, Proceedings of the Zoological Society of London 1888: 608, pl. 31, g. 12. Type-locality: Japan,
Satsuma. Holotype: x, in coll. BMNH.
Material examined:
Thailand. Prov. Nan: 1 specimen, 6 km N of Bo Luang, 1050 m, 19.viii.1999, leg. T. Csővári & l. mikus; 1 specimen, 5 km N of
Bo Luang, 1000 m, 10.ii.2000, leg. m. HRebLay, y. sHeRpani & a. szabó; 2 specimens, 25 km N of Bo Luang, 1150 m, 11.xi.1999,
leg. a. szabó & z. CzeRe; 1 specimen, from the same site, 3.ii.2000, leg. m. HRebLay, y. sHeRpani & a. szabó; 1 x, from the same
locality, 17.xi.1998, leg. T. Csővári & l. mikus, slide No. LGN 1057 (W 8371); 1 specimen, 30 km E of Pua, 1700 m, 20.i.2004,
leg. p. HentsCHeL & a. szabó. Prov. Chiang Mai: 2 specimens, 7 km W of Pa Pae, 1230 m, 21. and 27.xi.1998, leg. T. Csővári & l.
mikus; 4 specimens, between Chiang Dao and Kariang, 900 m, 98º48’E, 19º25’N, 26.x.2002; 3 specimens, from the same locality,
8.xi.2002, leg. b. HeRCzig & g. Ronkay; 4 specimens, 4 km SE of Pang Faen, 1100 m, 13.i.2004, leg. p. HentsCHeL & a. szabó; 4
specimens, from the same site, 31.i.2004, leg. a. szabó. Prov. Mae Hong Song: 1 specimen, between Pa Pae and Khun Sa, 1250
m, 98º39’E, 19º08’N, leg. b. HeRCzig & g. Ronkay (coll. G. Ronkay and MWM).
Meganola punctilineata (WiLeman & soutH, 1919)
(Pl. 7, gs 10-11)
Pisara punctilinea WiLeman & soutH, 1919, Entomologist 52: 270. Type-locality: Philippines, Mindanao, Lanao, Kolambugan.
Holotype: w, in coll. BMNH.
Material examined:
Thailand. Prov. Chiang Mai: 4 specimens, 4 km SE of Pang Faen, 1100 m, 13.i.2004, 26.i.2004 and 31.i.2004, slide No. LGN 999
(W 8370), leg. p. HentsCHeL & a. szabó; 1 w, from the same site, 27.i.2004, slide LGN 1000 (W 8381) leg. a. szabó. Prov. Chiang
Rai: 1 w, 1 km SE of Khun-Kon Waterfall, 600 m, 15.xi.1998, leg. T. Csővári & l. mikus, slide No. LGN 501 (W 8380). Prov. Nan:
1 specimen, 5 km N of Ban Luang, 350 m, between Pi Nai and Pi Tai, 100º27’E, 18º56’N, 4.xi.2002, leg. b. HeRCzig & g. Ronkay
(coll. G. Ronkay and MWM).
Meganola tenebrosa (Hampson, 1896)
(Pl. 7, gs 12-13)
Nola tenebrosa Hampson, 1896, Fauna of British India, Moths 4: 504. Type-locality: Bhutan. Holotype: w, in coll. BMNH.
Material examined:
Thailand. Prov. Nan: 1 x, 5 km N of Ban Luang, 350 m, between Pi Nai and Pi Tai, 100º27’E, 18º56’N, 4.xi.2002, leg. b. HeRCzig
& g. Ronkay, slide No. LGN 1058; 1 w, 25 km N of Bo Luang, 1150 m, 17.ii.1998, leg. m. HRebLay & Cs. szabóky, slide No. LGN
1060 (W 8429). Prov. Chiang Mai: 1 x, Mok Fa Garden Resort, 450 m, 98º48’E, 19º06’N, 1.xi.2002, leg. b. HeRCzig & g. Ronkay,
slide No. LGN 1061 (W 8419) (coll. G. Ronkay and MWM).
50
Meganola suffusata (WiLeman & West, 1929)
(Pl. 7, gs 14-15)
Nola suffusata WiLeman & West, 1929, Annals and Magazine of Natural History 10(3): 189. Type-locality: [Taiwan] Formosa, Kan-
shirei. Holotype: w, in coll. BMNH.
Material examined:
Thailand. Prov. Nan: 1 specimen, Doi Phuka NP, between Pua and Bo Luang, 1350 m, 101º05’E, 19º12’N, 3.xi.2002, leg. b.
HeRCzig & g. Ronkay; 1 w, 5 km N of Bo Luang, 1000 m, 4.ii.2000, leg. m. HRebLay, y. sHeRpani & a. szabó, slide No. LGN 495 (W
8418). Prov. Chiang Mai: 1 specimen, between Chiang Dao and Kariang, 900 m, 98º48’E, 19º25’N, 26.x.2002, leg. b. HeRCzig &
g. Ronkay (coll. G. Ronkay and MWM).
Meganola major (Hampson, 1891)
(Pl. 7, gs 16-17)
Nola major Hampson, 1891, Illustrated Typical Specimens of the Lepidoptera Heterocera in the Collection of the British Museum 8:
4, 48, pl. 139, g. 13. Type-locality: India [Tamil Nadu], Nilgiri district. Syntypes: in coll. BMNH.
Material examined:
Thailand. Prov. Nan: 1 specimen, 30 km E of Pua, 1700 m, 1.iii.1998; 5 specimens, from the same site, 18., 20. and 27.ii.1998,
leg. m. HRebLay & Cs. szabóky; 1 specimen, 5 km E of Bo Luang, 610 m, 23.xi.1998, leg. T. Csővári & l. mikus; 1 specimen, 22
km N of Bo Luang, 1120 m, 24.i.1999; 1 x, 25 km N of Bo Luang, 1150 m, 11.xi.1999, leg. a. szabó & z. CzeRe. Prov. Mae Hong
Song: 6 specimens, 25 km NE of Pai, 1560 m, 2.iii.1998, leg. m. HRebLay & Cs. szabóky; 2 specimens, 10 km NE of Pai, 1560 m,
28.i.1999, leg. a. szabó & z. CzeRe; 1 specimen, from the same locality, 3.xii.1998, leg. m. HRebLay, i. soós & y. sHeRpani; 1 speci-
men, 21 km NW of Pai, 1360 m, 7.ii.1998; 1 specimen, 1 km S of Bahundanda, 1000 m, 6.ii.1998, leg. m. HRebLay & Cs. szabóky;
Prov. Chiang Mai: 1 specimen, 23 km NW of Sop Kha, 1 km E of Kop Dong, 1650 m, 23.ii.1998; 2 spec. 15 km SW of Wiang
Haeng, 1400 m, 3.iii.1998, leg. m. HRebLay & Cs. szabóky; 1 specimen, 4 km W of Pa Pae, 1050 m, 28.xi.1998, leg. T. Csővári &
L. mikus; 1 specimen, 18 km NW of Fang, 2100 m, 25.ii.1998, leg. m. HRebLay & Cs. szabóky; 1 specimen, Mt. Doi Phahompok,
20 km NW of Fang, 2450 m, 24.i.2004, leg. a. szabó; 9 specimens, 6 km SE of Pang Faen, 1100 m, 16.ix.1999, leg. a. szabó & z.
CzeRe; 1 specimen, same site, 29.xi.1998, leg. T. Csővári & l. mikus; 12 specimens, 4 km SE of Pang Faen, 1100 m, 13, 18., and
26.i.2004, leg. p. HentsCHeL & a. szabó; 17 specimens, from the same site, 27. and 31.i.2004, leg. a. szabó; 1 specimen, from the
same locality, 14.xi.1999, leg. m. HRebLay & i. soós; 1 specimen, 12 km NW of Chiang Dao, 750 m, 12.xi.1998, leg. T. Csővári & l.
mikus; 4 specimens, between Chiang Dao and Kariang, 900 m, 98º48’E, 19º25’N, 26.x.2002 and 8.xi.2002; 1 specimen, between
Fang and Nor Lae, 1600 m, 99º06’E, 20º02’N, 7.xi.2002; 1 specimen, Mae Suai, 500 m, Charin Garden Resort, 99º35’E, 19º47’N,
2.xi.2002, leg. b. HeRCzig & g. Ronkay; 1 specimen, Doi Inthanon, NP, 2300 m, 21.i.1999, leg. a. szabó & z. CzeRe; 1 specimen, 2
km S of Kop Dong, 1800 m, 24.iii.1998, leg. T. Csővári & P. sTéger. Prov. Chiang Rai: 1 specimen, 1 km SE of Khun-Kon Waterfall,
600 m, 15.xi.1998, leg. T. Csővári & l. mikus. Prov. Phayao: 1 specimen, 15 km W of Huai Fuang, 740 m, 9.viii.1999, leg. T. Csővári
& L. mikus (coll. G. Ronkay and MWM).
Meganola argentescens (Hampson, 1895)
(Pl. 7, gs 18-19)
Pisara argentescens Hampson, 1895, Catalogue of the Lepidoptera Phalaenae in the British Museum 5: 296. Type-locality: [India]
Sikkim. Holotype: w, in coll. BMNH.
Material examined:
Thailand. Prov. Nan: 1 specimen, Doi Phuka NP, between Pua and Bo Luang, 1350 m, 101º05’E, 19º12’N, 3.xi.2002, leg. b. HeRCzig
& g. Ronkay; 2 xx, 30 km E of Pua, 1700 m, 20-21.vi.1998, leg. i. soós & a. szabó, slide Nos LGN 150 (W 8428), LGN 180 (W
8417). Prov. Chiang Mai: 1 specimen, between Fang and Nor Lae, 99º06’E, 20º02’N, 28.x.2002; 4 specimen, between Chiang Dao
and Kariang, 900 m, 98º48’E, 19º25’N, 26.x.2002, leg. b. HeRCzig & g. Ronkay; Prov. Mae Hong Song: 2 specimens, between Pa
Pae and Khun Sa, 1250 m, 98º39’E, 19º08’N, leg. b. HeRCzig & g. Ronkay (coll. G. Ronkay and MWM).
51
Meganola brunellus (Hampson, 1893)
(Pl. 7, gs 20-21)
Roeselia brunellus Hampson, 1893, Illustrated typical Specimens of the Lepidoptera Heterocera in the Collection of the British
Museum 9: 15, 89, pl. 158, g. 31. Type-locality: [Sri Lanka] Ceylon, Pundaloya. Syntypes: in coll. BMNH.
Meganola pseudohypena inoue, 1982, in inoue et al., Moths of Japan 1: 667, pl. 154, gs 26-27, pl. 351, g. 1, pl. 353, g. 9. Type-
locality: Japan, Ishigaki Island, Mt. Banna. Holotype: w, in coll. NSM Tokyo.
Celamoides corticella eeCke, 1926, Zoologische Mededeelingen, Leiden 9: 39, pl. 2, g. 10. Type-locality: [Indonesia] Sumatra,
Fort de Kock. Syntypes: 9 specimens, in coll. RNH Leiden.
Material examined:
Thailand. Prov. Chiang Mai: 7 specimens, between Fang and Nor Lae, 1600 m, 99º06’E, 20º02’N, 28.x.2002; 2 specimens, from
the same site, 12.xi.2002; 2 specimens, 4 km S of Kop Dong, 99º03’E, 19º52’N, 1800 m, 6.xi.2002, leg. b. HeRCzig & g. Ronkay;
1 specimen, 23 km NW of Sop Kha, 1 km E of Kop Dong, 1650 m, 14.i.2004, leg. p. HentsCHeL & a. szabó (coll. G. Ronkay and
MWM).
„Meganola” igorkostjuki spec. nov.
(Pl. 7, g. 7; gen. g. 45)
Holotype. x, „North Thailand, Prov. Chiang Mai, 1100 m, 4 km SE of Pang Faen, 13.i.2004, leg. A. szabó”, slide No. LGN 1062
(W 8382) (coll. MWM).
Diagnosis. The new species belongs to a remote phyletic line within Meganola, representing most probably a
distinct genus. As the species is known only by the x sex, the more detailed analysis of its taxonomic position is
postponed until the discovery of the w.
The differences between “Meganola” igorkostjuki and the other Meganola species are rather conspicuous. The
new species has a very characteristic and rather unique external appearance due to its very dark brown forewing
ground colour, the prominent yellowish apical streak, the slightly arcuate (but not sinuous), rather broad, yellowish
postmedial line and the distinctly marked black orbicular stigma.
The x genitalia display numerous autapomorphic features, like the long and slender, slightly curved and acutely
pointed uncus, the absence of subscaphial lobes, the very narrow, simple, apically rounded valva, the rather short,
bifurcate, more or less V-shaped harpe and the large, long V-shaped vinculum. The aedeagus is simple, almost
straight, the vesica is armed by a straight, relatively short, but strong cornutus. The w is unknown.
Distribution. The new species is known from the type locality only.
Nola ochrolopha (Hampson, 1911)
(Pl. 8, gs 1-2)
Celama ochrolopha Hampson, 1911, Annals and Magazine of Natural History 8(8): 397. Type-locality: [India] Sikkim. Holotype: w,
in coll. BMNH.
Material examined:
Thailand. Prov. Chiang Mai: 2 specimens, 20 km NW of Sop Kha, 2 km S of Kop Dong, 1800 m, 7.i.1999, leg. a. szabó & z. CzeRe;
1 x, from the same locality, 5.xii.1998, leg. m. HRebLay, i. soós & y. sHeRpani, slide No. LGN 1071 (W 8426); 1 specimen, 23 km
NW of Sop Kha, 1 km E of Kop Dong, 1650 m, 14.i.2004, leg. p. HentsCHeL & a. szabó; 5 specimens, Mt. Doi Phahompok, 20 km
NW of Fang, 2450 m, 24.i.2004; 4 specimens, Mt. Doi Phahompok, 20 km NW of Fang, 2150 m, 22-25. and 28.i.2004, leg. a.
szabó; 2 specimens, 18 km NW of Fang, 2100 m, 18.i.1999 and 20.i.1999, leg. a. szabó & z. CzeRe; 1 specimen, from the same
locality, 5.xi.1999, leg. m. HRebLay & i. soós; 1 specimen, Mt. Doi Phahompok, 16 km NW of Fang, 2000 m, 10.i.1999, leg. a.
szabó & z. CzeRe; 1 specimen, from the same locality, 27.iii.1998, leg. T. Csővári & P. sTéger; 1 specimen, 20 km NW of Mae Ai,
1650 m, 26.i.1999, leg. a. szabó & z. CzeRe; 6 specimens, between Fang and Nor Lae, 1600 m, 99º06’E, 20º02’N, 28.x., 7.xi. and
52
12.xi.2002, leg. b. HeRCzig & g. Ronkay. Prov. Nan: 2 specimens, 30 km E of Pua, 1700 m, 22-23.i.1999 and 25.i.1999, slide No.
LGN 1069 (W 8427); 1 x, 25 km N of Bo Luang, 1150 m, 14.i.1999, leg. a. szabó & z. CzeRe, slide No. LGN 1070 (W 8416) (coll.
G. Ronkay and MWM).
Nola defreina spec. nov.
(Pl. 8, gs 3-4; gen. g. 48)
Holotype. x, „Thailand, Prov. Nan, 30 km E of Pua, 1700 m, 20.i.2004, leg. p. Hentschel & A. Szabó”, slide No. LGN 1019 (W
8415) (coll. MWM).
Paratypes. Thailand. Prov. Mae Hong Song: 1 x, 1 km S of Bahundanda, 1000 m, 6.ii.1998; 1 x, 21 km NW of Pai, 1360 m,
7.ii.1998, leg. m. HRebLay & Cs. szabóky. Prov. Chiang Mai: 1 x, 20 km NW of Sop Kha, 2 km S of Kop Dong,
1800 m, 7.i.1999, leg. a. szabó & z. CzeRe, slide No. LGN 1018 (W 8414); 1 x, 15 km SW of Wiang Haeng, 1400
m, 9.ii.1998, leg. m. HRebLay & Cs. szabóky; 1 w, Mt. Doi Phahompok, 16 km NW of Fang, 2000 m, 4.iv.1998,
leg. T. Csővári & P. sTéger, slide No. LGN 1017 (W 8425); 1 x, between Fang and Nor Lae, 1600 m, 99º06’E,
20º02’N, 12.xi.2002, leg. b. HeRCzig & g. Ronkay; 1 x, from the same site, 18.ii.1998, leg. m. HRebLay & Cs.
szabóky. Prov. Nan: 1 x, 30 km E of Pua, 1700 m, 24.xi.1998, leg. T. Csővári & l. mikus; 4 xx, 1 w, from the
same locality, 20.i.2004, leg. p. HentsCHeL & a. szabó; 1 w, from the same locality, 5.ii.2000, leg. m. HRebLay,
y. sHeRpani & a. szabó (coll. G. Ronkay & MWM). Khasis.1 x, Oct. 1890, Nat. Coll., RotHsCHiLD Bequest B.M.
1939-1, 1 x, March 1894, Nat. Coll., RotHsCHiLD Bequest B.M. 1939-1 (coll. BMNH). Nepal. Annapurna Himal: 1
x, 2 km N Landrung, 1540 m, 83º49’E, 28º23’N, 8.iv.1995, leg. gy. m. LászLó & g. Ronkay (coll. MWM). China.
Guangxi: 2 xx, 2 ww, Dayao Shan, Jingxiu, 100 km SE of Liuzhou, 23º45’N, 109º45’E, 1200 m, 15-30.iii.2005,
leg. sinjaev & his team (coll. MWM).
Diagnosis. The new species is somewhat similar to Nola ochrolopha by its pale greyish white forewing ground
colour and the sharply dened medial line, but it is easily distinguished by its much narrower and more strongly
curved dark medial suffusion, the less sharply dened subterminal line, and the more greyish, and less shining
forewing. Nola defreina resembles also Nola sindhulica but the new species has somewhat paler greyish forewing
ground colour, more distinct double medial line the inner line of which is rather pale and less sinuous, this inner
line is sharper and conspicuously dentate in N. sindhulica.
The x genitalia of the related species show also clearly recognisable differences. N. defreina has much deeply cleft
valva compared to N. sindhulica and especially to N. ochrolopha (latter species has rather shallow medial incision
in the valva), much shorter and more robust harpe than in N. ochrolopha (the shape of harpe of the new species
is rather similar to that of N. sindhulica). The aedeagus of the new species is slightly slenderer than that of N.
sindhulica but much shorter than in N. ochrolopha, having a small, but conspicuous carinal tooth which is absent
in both related species. The vesica of N. defreina is armed by a short but rather robust, cornutus, which is angled
medially in 90°; the cornutus of N. ochrolopha is conspicuously longer and thinner, medially strongly reclinate, while
the cornutus of N. sindhulica is only slightly longer and medially less strongly bent than that of N. defreina.
The w genitalia of N. defreina display also characteristic differences compared with that of N. ochrolopha: the 8th
segment is shorter, the ostium and ductus bursae are narrower and less strongly sclerotized, the proximal sec-
tion of ductus bursae has conspicuous, elongated sclerotization (a unique feature of the species within the genus
Nola), and the presence of a pair of strongly sclerotized, more or less crater-shaped signa (N. ochrolopha has a
single, rounded signum).
Distribution. The new species has a relatively long area in the eastern Himalayan region, extending from Central
Nepal (Annapurna Himal) throughout Assam and northern Indochina to SE China (Guangxi).
Nola sindhulica inoue, 1998
(Pl. 8, g. 5)
Nola sindhulica inoue, 1998, Tinea 15 (Supplement 1): 91, pl. 140, g. 15, text g. 882. Type-locality: Nepal, Janakpur, Bijayach-
hap. Holotype: x, in coll. inoue.
Material examined:
53
Thailand. Prov. Chiang Mai: 1 x, 20 km NW of Sop Kha, 2 km S of Kop Dong, 1800 m, 7.i.1999, leg. a. szabó & z. CzeRe, slide
No. LGN 1072 (W 8424) (coll. MWM); 1 x, Doi Suthep-Pui NP. 4.ii.1989; 1 w, Doi Suthep-Pui NP. 29.xi.1989, leg. a.m. Cotton &
i.J. kitCHing BM 1989-57 (coll. BMNH).
Nola atrocincta inoue, 1998
(Pl. 8, gs 6-7)
Nola atrocincta inoue, 1998, Tinea 15 (Supplement 1): 91, pl. 140, g. 14, text g. 880. Type-locality: Nepal, Solukhumbu, Mnid-
ingna. Holotype: x, in coll. NSMT.
Material examined:
Thailand. Prov. Chiang Mai: 2 specimens, Mt. Doi Phahompok, 16 km NW of Fang, 2000 m, 15.ii.1998; 1 specimen, from the same
site, 24.ii.1998, leg. m. HRebLay & Cs. szabóky; 1 x, Mt. Doi Phahompok, 20 km NW of Fang, 2150 m 28.i.2004, slide No. LGN
978 (W 8413); 2 specimens, from the same site, 22.i.2004, slide No. LGN 979 (W 8423) (w), leg. a. szabó; 1 specimen, Mt. Doi
Phahompok, 20 km NW of Fang, 2000 m, 15.i.2004, leg. p. HentsCHeL & a. szabó. Prov. Nan: 1 w, Doi Phuka NP, between Pua and
Bo Luang, 1350 m, 101º05’E, 19º12’N, 3.xi.2002, leg. b. HeRCzig & g. Ronkay, slide No. LGN 1254 (coll. G. Ronkay and MWM).
Nola sikkima (mooRe, 1888) stat. rev.
Roeselia sikkima mooRe, 1888, Description of new Indian Lepidopterous Insects from the Collection of the late Mr. W. S. Atkinson
1888: 287. Type-locality: [India] [Sikkim] Darjeeling. Lectotype: w, in coll. BMNH, here designated.
Remarks. Nola sikkima (mooRe, 1888) has been treated by Hampson (1900) as a synonym of Celama exuosa
pouJaDe, 1886. Though the type of Celama exuosa has not been examined by the authors, inoue (1996) suppos-
edly examined it when transferred exuosa to Rhynchopalpus Hampson, 1893. The examination of the w genitalia
of one of the syntypes of N. sikkima undoubtedly shows its correct placement in the genus Nola LeaCH, 1830; it
cannot be combined with Rhynchopalpus (which is a synonym of Meganola DyaR, 1898) or Manoba WaLkeR, 1863.
Thus, Nola sikkima cannot be conspecic with Meganola exuosa. On the other hand, the syntype series of Nola
sikkima contains very probably two species: the dissected w syntype deposited in the BMNH and the w specimen
of the two syntypes deposited in ZMHU are identical, representing Nola sikkima. The third syntype in ZMHU the
abdomen of which is lost are supposedly not conspecic with the other syntypes and represent a new species, N.
wernerthomasi. This latter new species is described below, but the third syntype of N. sikkima cannot be identied
satisfactorily without the examination of its genitalia, therefore it is not added into the type-series. To clarify this
rather confused situation, the designation of the lectotype of N. sikkima is essential.
Lectotype of Roeselia sikkima mooRe, 1888, here designated: w, „red ring type label”, „Darjeeling”, „Roeselia sikkima w type Moore”,
„Moore Coll. 94-106”. Slide No. BM Arct. 891 in coll. BMNH.
Remarks. The species has not yet been recorded from Thailand.
Nola wernerthomasi spec. nov.
(Pl. 8, gs 8-9; gen. g. 49)
Holotype. x, „Thailand, Changwat Chiang Mai, Mt. Doi Phahompok, 18 km NW of Fang, 2100m, 7.ii.2000, leg. Hreblay & Szabó”,
slide No. LGN 1011 (W 8412) (coll. MWM).
Paratypes. Thailand. Prov. Chiang Mai: 1 x, Mt. Doi Phahompok, 18 km NW of Fang, 2100 m, 20.i.1999, slide LGN 1129 (W
8411), leg. a. szabó & z. CzeRe; 3 ww, 14 km NW of Fang, 2000 m, 14.ii.2000 leg. m. HRebLay, slide Nos LGN
1012 (W 8421), LGN 1246 (W 8420), LGN 1248 (W 8439); 1 w, Mt. Doi Inthanon, NP, 2300 m, 7-8.ix.1999 leg.
a. szabó & z. CzeRe, slide No. LGN 1245 (W 8410). Prov. Nan: 1 x, 30 km E of Pua, 1700 m, 10.xi.1999, leg. m.
HRebLay, slide No. LGN 1075 (W 8422) (coll. MWM). Vietnam. Prov. Lao Cai: 3 xx, 1 w, 3 km NW of Cat Cat,
2000 m, 1.xii.1997, leg. g. kósa, l. PeregoviTs & l. ronkay; 2 xx, Fan-si-pan Mts, 14 km NW Sa Pa, 2000 m,
103º46,06’E, 22º20,9’N, 4-6.xii.1997, leg. l. PeregoviTs & l. ronkay; 2 xx, Fan-si-pan Mts, 16 km NW Sa Pa,
16-17.iii.1998, leg. l. PeregoviTs & T. vásárHelyi (coll. HNHM). Nepal. Ganesh Himal: 1 w, 2 km W Gholjong,
2420 m, 85º18’E, 28º11’N, 12.x.1995, leg. l. PeregoviTs & l. ronkay (coll. HNHM). Sikkim. 1 x, Sikkim, Kurseong,
R.p. bRetanDeau, 1894; 1 x, Sikkim, 7000, July 1895, J.g. piLCHeR 97-31, slide No. BM Arct. 894; 2 xx, with the
54
same data; 2 xx, 1 w, same site, but collected in August 1895; 1 w, same site, but collected in May 1895, slide
No. BM Arct. 895; 1 x, Sikhim, VIII.1909, F. moLLeR, 1910-140; 1 x, Khasis, Sept. 1895, Nat. Coll; 3 xx, 2 ww,
Khasis, Nat. Coll. (coll. BMNH).
Diagnosis. The new species is closely related to Nola sikkima but clearly separable by its much darker, more
distinctly marked forewing median area and more sharply dened medial and postmedial lines. As Nola sikkima
is available only by w specimens, only the w genitalia of the two species can be compared. N. wernerthomasi has
less elongate papillae anales, somewhat thicker and longer more strongly sclerotized ostium bursae, conspicu-
ously shorter and thicker ductus bursae and considerably larger, more rounded and more strongly sclerotized
signum bursae. The x genitalia of the new species are characterized by their medium deeply cleft valvae with
broadly rounded apex, the very slightly arcuate, medium-long, slightly tapering, apically rounded harpe, the rather
elongate, apically pointed vinculum, and the relatively short, medium thick aedeagus armed with a medium-long,
falcate, pointed, single cornutus.
Distribution. An eastern Himalayan species, all known localities lie in the southern side of the Himalayan main chain,
from Central Nepal and Sikkim to the northern areas of Indochina (Northern Thailand, North Vietnam).
Nola peguense (Hampson, 1894)
(Pl. 8, gs 10-11)
Deltapterum peguense Hampson, 1894, Fauna of British India, Moths 2: 138, g. 93. Type-locality: [Myanmar] Burma, E. Pegu.
Holotype: x, in coll. BMNH.
Material examined:
Thailand. Prov. Nan: 7 specimens, 30 km E of Pua, 1700 m, 10. and 13.xi.1999, slide No. LGN 968 (W 8448), leg. m. HRebLay; 4
specimens, from the same locality, 1700 m, 12-13.i.1999, 22-23.i.1999 and 25.i.1999, slide No. LGN 969 (W 8437), leg. a. szabó
& z. CzeRe; 1 specimen, from the same site, 16.xi.1998, leg. T. Csővári & l. mikus; 6 specimens, from the same locality, 18.ii.1998
and 27.ii.1998, slide No. LGN 1056 (W 8438), leg. m. HRebLay & Cs. szabóky; 7 specimens, from the same locality, 20-21.i.2004,
leg. a. szabó & p. HentsCHeL; 2 specimens, Doi Phuka NP, between Pua and Bo Luang, 1350 m, 101º05’E, 19º12’N, 3.xi.2002,
leg. b. HeRCzig & g. Ronkay; 1 specimen, 25 km N of Bo Luang, 1150 m, 11.xi.1999, leg. a. szabó & z. CzeRe; 2 xx, 5 km N of Bo
Luang, 1000 m, 4.ii.2000, leg. m. HRebLay, y. sHeRpani & a. szabó, slide Nos LGN 972 (W 8446), LGN 955 (W 8449). Prov. Chiang
Mai: 1 specimen, Mt. Doi Phahompok, 16 km NW of Fang, 2000 m, 27.iii.1998, leg. T. Csővári & P. sTéger; 3 specimens, Mt. Doi
Phahompok, 18 km NW of Fang, 2100 m, 15.xi.1999, leg. m. HRebLay; 1 specimen, Mt. Doi Phahompok, 20 km NW of Fang, 2450
m, 24.i.2004, leg. a. szabó; 15 specimens, between Fang and Nor Lae, 1600 m, 99º06’E, 20º02’N, 12.xi.2002; 5 specimens, from
the same locality, 28.x.2002 and 7.xi.2002, slide No. LGN 952 (x), leg. b. HeRCzig & g. Ronkay; 5 specimens, 20 km NW of Mae
Ai, 1650 m, 26.i.1999, slide No. (LGN 970 (W 8447), leg. a. szabó & z. CzeRe; 2 specimens, 15 km SW of Wiang Haeng, 1400 m,
9.ii.1998, slide No. LGN 973 (W 8436) (w), leg. m. HRebLay & Cs. szabóky. Prov. Mae Hong Song: 1 specimen, 21 km NW of Pai,
1360 m, 7.ii.1998, leg. m. HRebLay & Cs. szabóky (coll. G. Ronkay and MWM).
Nola pinratana spec. nov.
(Pl. 8, gs 12-13; gen. g. 50)
Holotype. x, „North Thailand, Prov. Chiang Mai, 1600 m, between Fang and Nor Lae, 99°06’E, 20°02’N, 12.xi.2002, leg. B.
HeRCzig & g. Ronkay” (coll. MWM).
Paratypes. Thailand. Prov. Chiang Mai: 1 w, 2 km S of Ban Kum, 1700 m, 13-14.vi.1998, leg. i. soós & a. szabó; 1 x, between
Fang and Nor Lae, 1600 m, 99º06’E, 20º02’N, 12.xi.2002, leg. b. HeRCzig & g. Ronkay; 1 x, 15 km SW of Wiang
Haeng, 1400 m, 9.ii.1998, leg. m. HRebLay & Cs. szabóky, slide No. LGN 872 (W 8435); 2 xx, 20 km NW of
Sop Kha, 2 km S of Kop Dong, 1800 m, 7.i.1999; 1 w, from the same locality, 27.i.1999, slide No. LGN 1064 (W
8445), leg. a. szabó & z. CzeRe. Prov. Nan: 1 w, 30 km E of Pua, 1700 m, 25.i.1999, leg. a. szabó & z. CzeRe
(coll. G. Ronkay and MWM).
Diagnosis. The new species cannot be confused with the other congeners. Its external appearance is rather unique
having almost uniformly dark graphite greyish forewings with somewhat darker, small costal spots and diffuse,
shadow-like crosslines. The conguration of the x genitalia display closer relationship with Nola wernerthomasi,
the main differences are as follows: the new species has slightly longer but more robust, medially curved, apically
pointed harpe, much shorter aedeagus and slenderer, ner, more curved cornutus in the vesica.
55
The harpe of N. wernerthomasi is almost straight and apically rounded, the aedeagus is considerably longer, and
the cornutus of the vesica is thicker, less falcate than in N. pinratana. The w genitalia of the two species show
similarly conspicuous differences: the new species has considerably longer 8th segment, much shorter sclerotized
part of ostium bursae, and different shape of signum bursae which is elongate, crest-like in N. pinratana, and
rounded in N. wernerthomasi.
Distribution. Northern Thailand.
Nola duplicilinea (Hampson, 1900)
(Pl. 8, gs 14-15)
Celama duplicilinea Hampson, 1900, Catalogue of the Lepidoptera Phalaenae in the British Museum 2: 12, pl. 18, g. 8. Type-local-
ity: [India] Sikkim. Holotype: x, in coll. BMNH.
Celamoides pseudastigma eeCke, 1920, Zoologische Mededeelingen Leiden 5: 120, gs 8-9. Type-locality: [Indonesia] Java,
Preanger; Malabar, Tjinjiroean. Syntypes: one x and eight ww, in coll. RNH Leiden.
Celama indenita Roepke, 1948, Tijdschrift voor Entomologie 89: 210, pl. 14, g. 7. Type-locality: [Indonesia] Sumatra, Mt. Tang-
gamus. Holotype: w, in coll. RNH Leiden.
Material examined:
Thailand. Prov. Nan: 9 specimens, 30 km E of Pua, 1700 m, 12-13. and 25.i.1999, slide No. LGN 1114 (W 8444), leg. a. szabó &
z. CzeRe; 3 specimens, from the same site, 2.ii.2000, leg. m. HRebLay, y. sHeRpani & a. szabó; 1 specimen, from the same locality,
20.ii.1998, leg. T. Csővári & l. mikus; 5 specimens, from the same locality, 20-21.i.2004, slide No. LGN 1027 (W 8443) (w), leg. a.
szabó & p. HentsCHeL. Prov. Chiang Mai: 1 specimen, Mt. Doi Phahompok, 16 km NW of Fang, 2000 m, 4.iv.1998, leg. T. Csővári
& p. stégeR; 1 specimen, from the same locality, 15.ii.1998, leg. m. HRebLay & Cs. szabóky; 4 specimens, Mt. Doi Phahompok,
18 km NW of Fang, 2100 m, 9. and 20.i.1999, slide No LGN 1110 (W 8433) (x), leg. a. szabó & z. CzeRe; 1 specimen, from the
same locality, 25.ii.1998, leg. m. HRebLay & Cs. szabóky; 1 specimen, from the same site, 7.ii.2000, leg. M. HRebLay & a. szabó; 6
specimens, Mt. Doi Phahompok, 20 km NW of Fang, 2450 m, 24.i.2004; 2 specimens, same site, 22-25.i.2004, leg. a. szabó; 6
specimens, 20 km NW of Sop Kha, 2 km S of Kop Dong, 1800 m, 8.i.1999, slide No. LGN 1116 (W 8434) (w); 3 specimens, from
the same site, 7. and 27.i.1999, leg a. szabó & z. CzeRe; 4 specimens, same site, 21.ii.1998, leg. m. HRebLay & Cs. szabóky; 3
specimens, between Fang and Nor Lae, 1600 m, 99º06’E, 20º02’N, 28.x.2002; 5 specimens, from the same locality, 7.xi.2002 and
12.xi.2002, slide Nos LGN 1055, LGN 1080 (xx); 2 specimens, 4 km S of Kop Dong, 99º03’E, 19º52’N, 1800 m, 6.xi.2002, leg.
b. HeRCzig & g. Ronkay; 1 specimen, Mt. Doi Inthanon, 2300 m, 14.xii.1998, leg. m. HRebLay, i. soós & y. sHeRpani; 1 specimen,
from the same locality, 21.i.1999, leg. a. szabó & z. CzeRe. Prov. Mae Hong Song: 1 specimen, 1 km S of Bahundanda, 1000 m,
6.ii.1998, leg. m. HRebLay & Cs. szabóky (coll. G. Ronkay and MWM).
Nola fraterna (mooRe, 1888)
(Pl. 8, gs 16-17)
Roeselia fraterna mooRe, 1888, Proceedings of the Zoological Society of London 1888: 393. Type-locality: [India] [Himachal
Pradesh], Kangra. Holotype: w, in coll. BMNH.
Material examined:
Thailand. Prov. Chiang Mai: 7 specimens, between Fang and Nor Lae, 1600 m, 99º06’E, 20º02’N, 7.xi.2002; 10 specimens, from
the same site, 28.x.2002 and 12.xi.2002, leg. b. HeRCzig & g. Ronkay; 1 specimen, 4 km SE of Pang Faen, 1100 m, 31.i.2004,
leg. a. szabó; 1 specimen, 20 km NW of Sop Kha, 2 km S of Kop Dong, 1800 m, 27.i.1999, leg a. szabó & z. CzeRe; 1 specimen,
15 km SW of Wiang Haeng, 1400 m, 9.ii.1998, leg. m. HRebLay & Cs. szabóky. Prov. Nan: 1 specimen, 4 km W of Pha Lak,
100º34’E, 19º21’N, 5.xi.2002, leg. b. HeRCzig & g. Ronkay; 1 specimen, 22 km N of Bo Luang, 1100 m, 18-19.vi.1998, leg. i. soós
& a. szabó; 1 specimen, 5 km N of Bo Luang, 1000 m, 4.ii.2000, leg. m. HRebLay, y. sHeRpani & a. szabó. Prov. Mae Hong Song: 5
specimens, 21 km NW of Pai, 1360 m, 7.ii.1998; 2 specimens, 1 km S of Bahundanda, 1000 m, 6.ii.1998, leg. m. HRebLay & Cs.
szabóky (coll. G. Ronkay and MWM).
Nola bifascialis (WaLkeR, 1865)
(Pl. 8, gs 18-19)
Pisara bifascialis WaLkeR, 1865, List of the Specimens of Lepidopterous Insects in the Collection of the British Museum 31: 244.
56
Type-locality: Borneo, Sarawak. Type(s): in coll. UM Oxford.
Material examined:
Thailand. Prov. Chiang Mai: 1 x, 1 w, 4 km SE of Pang Faen, 1100 m, 31.i.2004, leg. a. szabó, slide Nos LGN 1154 (W 8432) (x),
LGN 1155 (W 8442) (w); 2 specimens, from the same site, 26.i.2004, leg. p. HentsCHeL & a. szabó; 1 specimen, Mok Fa Garden
Resort, 450 m, 98º48’E, 19º06’N, 1.xi.2002, leg. b. HeRCzig & g. Ronkay. Prov. Nan: 1 specimen, 5 km N of Ban Luang, 350 m,
between Pi Nai and Pi Tai, 100º27’E, 18º56’N, 4.xi.2002, leg. b. HeRCzig & g. Ronkay (coll. MWM).
Nola mesotherma (Hampson, 1909)
(Pl. 8, gs 20-21)
Celama mesotherma Hampson, 1909, Annals and Magazine of Natural History 8(4): 349. Type-locality: [Sri Lanka] Ceylon. Holotype:
w, in coll. BMNH.
Material examined:
Thailand. Prov. Chiang Mai: 1 w, between Chiang Dao and Kariang, 900 m, 98º48’E, 19º25’N, 26.x.2002, leg. b. HeRCzig & g.
Ronkay, slide No. LGN 1159 (W 8431) (coll. MWM).
Nola nepalpumila inoue, 1998
(Pl. 9, gs 1-2)
Nola nepalpumila inoue, 1998, Tinea 15 (Supplement 1): 89, pl. 140, g. 6, text g. 877. Type-locality: India [Sikkim], W Bengal,
Darjeeling. Holotype: x, in coll. NSMT.
Material examined:
Thailand. Prov. Chiang Mai: 1 x, 1 w, between Fang and Nor Lae, 99º06’E, 20º02’N, 28.x.2002, slide Nos LGN 1086 (x), LGN
1085 (w); 2 specimens, from the same site, 12.xi.2002; 1 x, 4 km S of Kop Dong, 99º03’E, 19º52’N, 1800 m, 6.xi.2002, slide No.
LGN 1038 (W 8430), leg. b. HeRCzig & g. Ronkay; 1 specimen, Doi Inthanon, NP, 2300 m, 14.xii.1998, leg. m. HRebLay, i. soós &
y. sHeRpani; 1 specimen, 20 km NW of Mae Ai, 1650 m, 19.xii.1998, leg. m. HRebLay, i. soós & y. sHeRpani. Prov. Nan: 1 fem, Doi
Phuka NP, between Pua and Bo Luang, 1350 m, 101º05’E, 19º12’N, 3.xi.2002, leg. b. HeRCzig & g. Ronkay, slide No. LGN 1037
(W 8441); 1 specimen, 30 km E of Pua, 1700 m, 20.ii.1998, leg. T. Csővári & l. mikus; 1 specimen, from the same site, 8.iv.1998,
leg. T. Csővári & P. sTéger; 2 specimens, from the same locality, 25.i.1999, leg. a. szabó & z. CzeRe; 1 specimen, 25 km N of Bo
Luang, 1150 m, 3.ii.2000, leg. m. HRebLay, y. sHeRpani & a. szabó (coll. G. Ronkay and MWM).
Nola kiisigna spec. nov.
(Pl. 9, g. 3; gen. g. 51)
Holotype. x, „N. Thailand, Prov. Chiang Mai, 900m, between Chiang Dao and Kariang, 98º48’E, 19º25’N, 08.11.2002 leg. B.
HeRCzig & g. Ronkay”, slide No. LGN 1097 (coll. G. Ronkay).
Diagnosis. Nola kiisigna, N. ineffectalis WaLkeR, 1864 and N. tincta WiLeman & soutH, 1919, described from the
Philippines, form a common, specialised phyletic line, according to their unique and characteristic signa: it is very
narrow and long, ribbon-like situated crosswise to the main (longitudinal) axis.
The new species is closely related to N. ineffectalis WaLkeR, 1864 (see the Fig. 51) according to their similarly pale
grey forewing ground colour with somewhat darker reversed triangular costal patch. The two taxa are, however
easily distinguishable by the following features: the new species has considerably narrower, more elongate forew-
ing with more greyish ground colour, more sharply dened costal patch and crosslines, while the forewing ground
colour of N. ineffectalis is more brownish, with paler costal patch and crosslines.
57
The w genitalia of N. kiisigna differ from those of N. ineffectalis by their much shorter papillae anales, much longer,
more strongly sclerotized 8th segment and much shorter apophyses posteriores; the ostium bursae of the new
species is narrower than that of N. ineffectalis. Last but not least, the signum is slightly curved in N. kiisigna while
straight in N. ineffectalis. The x is unknown.
Distribution. The new species is known from the type locality only.
Nola canioralis (WaLkeR, 1863)
(Pl. 9, g. 6)
Necla canioralis WaLkeR, 1863, List of the Specimens of Lepidopterous Insects in the Collection of the British Museum 27: 100.
Type locality: Borneo, Sarawak. Holotype, w in coll. BMNH.
Material examined:
Thailand. Prov. Chiang Mai: 1 w, 6 km SE of Pang Faen, 1100 m, 16.ix.1999 leg. a. szabó & z. CzeRe, slide No. LGN 172 (W
14990) (coll. MWM).
Nola opalina (WaLkeR, 1862)
(Pl. 9, g. 5)
Pisara opalina WaLkeR, 1862, Journal of the Proceedings of the Linnean Society of London (Zoology) 6: 118, by monotypy. Type-
locality: Borneo, Sarawak. Type(s) in coll. UM Oxford.
Material examined:
Thailand. Prov. Nan: 2 specimens, 4 km W of Pha Lak, 100º34’E, 19º21’N, 5.xi.2002, leg. b. HeRCzig & g. Ronkay; Prov. Chiang
Mai: 2 specimens, 4 km SE of Pang Faen, 1100 m, 13.i.2004, leg. p. HentsCHeL & a. szabó; Prov. Mae Hong Song: 1 w, 21 km NW
of Pai, 1360 m, 7.ii.1998, leg. m. HRebLay & Cs. szabóky, slide No. LGN 1063 (W 8440) (coll. G. Ronkay and MWM).
Remarks. Nola canioralis and N. opalina are sister species, being externally very similar to each other. As only the w
holotype of N. canioralis has been dissected, and the genitalia of the types of N. opalina are unknown, only the ww
of N. canioralis can be properly identied yet. The further revision of the species complex would be desirable.
Nola ceylonica Hampson, 1893
(Pl. 9, gs 7-8)
Nola ceylonica Hampson, 1893, Illustrations of Typical Specimens of Lepidoptera Heterocera in the Collection of the British Museum
9: 15, 88, pl. 158, g. 13. Type-locality: [Sri Lanka] Ceylon, Pundaloya. Holotype: x, in coll. BMNH.
Nola kimurai yazaki, 1995, Japan Heterocerists’ Journal 182: 104. Type-locality: Japan, Okinawa Island.
Material examined:
Thailand. Prov. Chiang Mai: 1 x, between Chiang Dao and Kariang, 900 m, 98º48’E, 19º25’N, 26.x.2002, leg. b. HeRCzig & g. Ronkay,
slide No. LGN 1084; 1 x, between Fang and Nor Lae, 1600 m, 99º06’E, 20º02’N, 12.xi.2002, slide No. LGN 1141 (W 14992); 1 w,
between Chiang Dao and Kariang, 900 m, 98º48’E, 19º25’N, 26.x.2002, slide No. LGN 1150 (W 14994), leg. b. HeRCzig & g. Ronkay;
1 x, 4 km SE of Pang Faen, 1100 m, 31.i.2004, leg. a. szabó, slide No. LGN 1149 (W 14993) (coll. G. Ronkay and MWM).
Nola euryzonata (Hampson, 1900)
(Pl. 9, gs 9-10)
Celama euryzonata Hampson, 1900, Catalogue of the Lepidoptera Phalaenae in the British Museum 2: 16, pl. 18, g. 11. Type-
58
locality: [Indonesia] Sumatra. Holotype: w, in coll. BMNH.
Material examined:
Thailand. Prov. Nan: 1 w, 25 km N of Bo Luang, 1150 m, 17.xi.1998, leg. T. Csővári & l. mikus, slide No. LGN 1158 (W 14995)
(coll. MWM).
Nola spec. close to Nola euryzonata
(Pl. 9, g. 11)
Material examined:
Thailand. Prov. Nan: 1 x, 750 m, 4 km W of Pha Lak, 100º34’E, 19º21’N, 5.xi.2002, leg. b. HeRCzig & g. Ronkay, slide No. LGN
1092 (coll. G. Ronkay).
Remarks. The old sense „Celama” species (type: „Celama” liparisalis WaLkeR, 1865) are presently under revision.
The preliminary results indicate that the species group is far more complex than the actual knowledge shows. The
exact identication of the species, therefore, requires the full revision of the lineage.
Nola lucidalis (WaLkeR, 1865)
(Pl. 9, gs 12-13)
Pisara lucidalis WaLkeR, 1865, List of the Specimens of Lepidopterous Insects in the Collection of the British Museum 31: 245.
Type-locality: Borneo, Sarawak. Type: x, in coll. UM Oxford.
Thailand. Prov. Nan: 1 x, 750 m, 4 km W of Pha Lak, 100º34’E, 19º21’N, 5.xi.2002, leg. b. HeRCzig & g. Ronkay, slide No. LGN
1156 (coll. G. Ronkay). 1 x, Chaiyaphum, 650 m, Ohu Khieo WS, km 7, Sala Prom checkpoint, 4/9.xi.1991, leg. i.J. kitCHing &
a.m. Cotton BM 1992-9 (coll. BMNH).
Nola erythrostigmata Hampson, 1894
(Pl. 9, gs 14-15)
Nola erythrostigmata Hampson, 1894, Fauna of British India, Moths 2: 140. Type-locality: [India] [Nagaland] Naga Hills. Holotype:
x, in coll. BMNH.
Material examined:
Thailand. Prov. Chiang Mai: 4 specimens, 4 km SE of Pang Faen, 1100 m, 13.i.2004, 18.i.2004, and 26.i.2004, slide No. LGN
1144 (W 14999), leg. p. HentsCHeL & a. szabó; 3 specimen, from the same site, 27.i.2004 and 31.i.2004, leg. a. szabó; 1 speci-
men, between Chiang Dao and Kariang, 900 m, 98º48’E, 19º25’N, 26.x.2002, leg. b. HeRCzig & g. Ronkay. Prov. Mae Hong Son:
1 specimen, 10 km NE of Pai, 1560 m, 28.i.1999, leg. a. szabó & z. CzeRe; 3 specimens, 21 km NW of Pai, 1360 m, 7.ii.1998, leg.
m. HRebLay & Cs. szabóky, slide Nos LGN 1147 (W 14996) (x), LGN 1145 (W 14997) (w). Prov. Nan: 1 x, 5 km N of Bo Luang,
1000 m, 4.ii.2000, leg. M. HRebLay & a. szabó, slide No. LGN 1146 (W 14998) (coll. G. Ronkay and MWM).
Nola spec. close to Nola erythrostigmata No. 1.
(Pl. 9, g. 16)
Material examined:
Thailand. Prov. Mae Hong Song: 1 x, 21 km NW of Pai, 1360 m, 7.ii.1998, slide No. LGN 1142 (W 15000); 1 w, 1 km S of Bahun-
danda, 1000 m, 6.ii.1998, leg. m. HRebLay & Cs. szabóky, slide No. LGN 1157 (W 15001) (coll. MWM).
59
Nola spec. close to Nola erythrostigmata No. 2.
(Pl. 9, g. 17)
Material examined:
Thailand. Prov. Chiang Mai: 1 x, 15 km SW Wiang Haeng, 1400 m, 9.ii.1998, leg. m. HRebLay & Cs. szabóky, slide No. LGN 1143
(W 15002) (coll. MWM).
Remarks. During the study on the Thai Nolinae material two closely related species of N. erythrostigmata were
distinguished which display certain external and/or genital differences. The exact identication of these two species
requires the revision of the old sense „Celama” species group.
Nola thyrophora (Hampson, 1914)
(Pl. 9, gs 18-19)
Pisara thyrophora Hampson, 1914, Catalogue of the Amatidae and Arctiidae(Nolinae and Lithosiinae) in the Collection of the British
Museum 1914: 390, pl. 22, g. 9. Type-locality: [Taiwan] Formosa, Kanshirei. Holotype: x, in coll. BMNH.
Material examined:
Thailand. Prov. Chiang Mai: 2 specimens, 4 km SE of Pang Faen, 1100 m, 31.i.2004, leg. a. szabó; 2 specimens, from the same
site, 26.i.2004, leg. p. HentsCHeL & a. szabó; 1 specimen, between Chiang Dao and Kariang, 900 m, 98º48’E, 19º25’N, 26.x.2002,
leg. b. HeRCzig & g. Ronkay. Prov. Mae Hong Song: 1 specimen, 1 km S of Bahundanda, 1000 m, 6.ii.1998, leg. m. HRebLay & Cs.
szabóky. Prov. Nan: 1 x, 1 w, 5 km N of Bo Luang, 1000 m, 4.ii.2000, leg. m. HRebLay, y. sHeRpani & a. szabó, slide Nos LGN 1117
(W 15003), LGN 1140 (W 15004) (coll. MWM).
Nola endotherma (Hampson, 1918)
(Pl. 9, g. 20)
Celama endotherma Hampson, 1918, Novitates Zoologicae 25: 95. Type-locality: [India] [Meghalaya] Khasis. Holotype: w, in coll.
BMNH.
Material examined:
Thailand. Prov. Nan: 1 w, 30 km E of Pua, 1700 m, 20.i.2004, leg. p. HentsCHeL & a. szabó, slide No. LGN 993 (W 15005) (coll.
MWM).
Nola mediolineata spec. nov.
(Pl. 9, g. 21; gen. g. 52)
Holotype. x, „Thailand, Prov. Nan, 1700m, 30 km E of Pua, 20.i.2004, leg. P. Hentschel & A. Szabó”, slide No. LGN 991 (W 7316)
(coll. MWM).
Paratypes. Thailand. Prov. Nan: 1 x, Doi Phu Kha, Pua, 1680 m, 16-20.ii.1993, leg. W. speiDeL (coll. speiDeL). 1 w, Khao Yai NP,
900 m, 3.ii.1987, Col. m.g. aLLen, BM 1996-300, slide No. LGN 1332 (BM Arct. 6211) (coll. BMNH). 1 x, Khao
Yai, 20-24.viii.1990, leg. k.t. paRk, slide No. LGN 1337 (coll. HNHM).
Diagnosis. The new species resembles externally Inouenola diversalis but the similarity of the two species is
only supercial, the two similar species belong to two different genera according to their genitalia features. Nola
mediolineata has, in comparison with I. diversalis, much paler grey forewing ground colour, somewhat narrower,
apically less pointed forewing and conspicuous, sharply dened, very slightly arched medial line. The antemedial
line of I. diversalis is sharply dened, almost straight, and the medial line is absent.
The conguration of x genitalia of the new species clearly show the generic features of Nola, characterized by
60
the following features: valva medially deeply incised, valval arms relatively broad, apically broadly rounded; harpe
conspicuously short, robust, thorn-like, vinculum medium long, rather elongated, apically sharply pointed. Aedeagus
short, medially slightly curved, vesica with a single, large cornutus.
The typical features of the w genitalia are the conspicuously long apophyses, the very simple, membranous ostium
bursae, the somewhat swollen, membranous distal-, and the strongly sclerotized, wrinkled proximal section of
ductus bursae. The cervix bursae of the new species is well-developed, relatively broad at base; the corpus bursae
is slightly ovoidal, and the signum is small, crescent-like.
Distribution. Northern Thailand.
Nola serrativalva spec. nov.
(Pl. 10, g. 1; gen. g. 53)
Holotype. x, „N. Thailand, Chiang Mai Prov., between Chiang Dao and Kariang, 900m, 98º48’E, 19º25’N, 26.x.2002 leg. B. HeRCzig
& g. Ronkay”, slide No. LGN 1093 (coll. G. Ronkay).
Diagnosis. The new species, together with the next species, N. picurka, are the smallest known members of the
entire subfamily: wingspan 8-9 mm, length of forewing 3-4 mm. The most typical external feature is the presence
of a pair of characteristic, ne but distinct, elongate-rounded red-brownish spots along the costal margin of the
forewing, these spots are more prominent than in N. picurka.
The conguration of the x genitalia of N. serrativalva is rather unusual within the genus Nola: the valval shape is
typical of the genus with the deep medial incision and narrow, apically rounded arms, but the ventral arm has a
heavily sclerotized, sharply dentate crest along the whole length of the saccular margin of valva. The vinculum is
broad, triangular, apically pointed, the aedeagus is medium-long, rather straight, the vesica is armed by a single,
well-developed cornutus. The w is unknown. The genitalia of N. serrativalva and N. picurka are rather dissimilar
(see the Figs 53 and 54), a more detailed comparison is given under the diagnosis of the latter species.
Distribution. The species is known by its unique holotype only.
Nola picurka spec. nov.
(Pl. 11, g. 21; gen. g. 54)
Holotype. x, „NW Thailand: 500m, Fang, Hortic. Res. Sta. 8-10.xi.1988, Coll. J.D. Bradley, Angoon Lewvanich & D.S. Fletcher”,
slide No. LGN 1354 (BM Arct. 6212) (coll. BMNH).
Paratypes. Thailand. 1 x, with the same data as the holotype; 1 x, Khao Yai NP, Park HQ, 720m, 2-4.xi.1988, Coll. J.D. bRaDLey,
angoon lewvaniCH & D.s. fleTCHer, slide No. LGN 1351 (BM Arct. 6213); 1 x, Ramkhamhaeng NP, 27.x.1990,
leg. j.D. braDley & angoon lewvaniCH, Brit. Mus. 1990-221, slide No. LGN 1352 (BM Arct. 6214). Sabah. 1 x,
Crocker Range NP, 5º45’N, 116º19’E, lower montane forest, 7-12.viii.1991, k.R. tuCk, op. RaLeigH BM 1991-136,
slide No. LGN 1353 (BM Arct. 6215) (coll. BMNH). Vietnam. Ca 90 x specimens, Prov. Nghe An, Pu Huong
Nature Reserve, 19°20`12N, 205°01`18E, 380 m, 6-8.x.2008, leg. G. CsoRba (coll. HNHM); Cambodia. Prov.
Mondolkiri: 4 xx, Seima Biodiversity Conservation Area, between Seima and O’Rang, 12º12’12N, 107º01’09E,
300 m, 30.i.2006, leg. g. CsoRba & g. Ronkay (coll. MWM).
Diagnosis. Nola picurka is similar in size and wingpattern to the previously described N. serrativalva, having
somewhat paler forewing ground colour and less sharply dened elongate-rounded costal spots. Despite their
rather similar external appearance, the x genitalia of the two species display conspicuous differences. The most
prominent difference is the presence or absence of the saccular crest, besides this key feature, there are several
smaller ones which are as follows. Nola picurka has, in comparison with N. serrativalva, somewhat broader valval
arms, and a bilobate harpe the proximal lobe of which is swollen, rounded, more or less ball-like with very ne
scobination on its distal surface while the distal lobe is a short, but relatively robust and slightly curved hook. The
new species has much longer and narrower vinculum, with a short, but distinct, slightly nger-like process, the
aedeagus is somewhat longer, more slender, apically more pointed, and the vesica has no cornutus (N. serrativalva
has a well-developed cornutus in the vesica). The w is unknown.
61
Distribution. The new species occurs in the northern parts of Thailand, Vietnam and Cambodia.
Nola fasciata (WaLkeR, 1866)
(Pl. 10, gs 2-3)
Minnagara fasciata WaLkeR, 1866, List of the Specimens of Lepidopterous Insects in the Collection of the British Museum 35:
1903. Type-locality: [Moluccas] Sula [Island]. Lectotype: w, designated by Swinhoe, (as type) 1892, Catalogue east. and Australian
Lepidoptera Heterocera 1: 99, in coll. UM, Oxford.
Nola nigrifascia Hampson, 1891, Illustrations of Typical Specimens of Lepidoptera Heterocera in the Collection of the British Museum
8: 5, 48, pl. 139, g. 15. Type-locality: India [Tamil Nadu], Nilgiri district. Holotype: x, in coll. BMNH.
Material examined:
Thailand. Prov. Nan: 3 specimens, 5 km E of Bo Luang, 610 m, 23.xi.1998, leg. T. Csővári & l. mikus; 1 specimen, 25 km N of Bo
Luang, 1150 m, 11.xi.1999, leg. a. szabó & z. CzeRe; 2 specimens, 22 km N of Bo Luang, 1100 m, 29-30.vi.1998; 2 specimens, from
the same site, 18-19.vi.1998, leg. I. soós & a. szabó; 2 specimens, 4 km W of Pha Lak, 100º34’E, 19º21’N, 5.xi.2002; 2 specimens,
5 km N of Ban Luang, 350 m, between Pi Nai and Pi Tai, 100º27’E, 18º56’N, 4.xi.2002; 1 specimen, Doi Phuka NP, between Pua
and Bo Luang, 1350 m, 101º05’E, 19º12’N, 3.xi.2002 leg. b. HeRCzig & g. Ronkay. Prov. Phayao: 1 specimen, 15 km SE of Chiang
Muan, 640 m, 26.xi.1998, leg. T. Csővári & l. mikus; 1 specimen, Phayao, 15 km W of Huai Fuang, 740 m, 9.viii.1999, leg. T. Csővári
& L. mikus. Prov. Chiang Mai: 1 specimen, 23 km NW of Sop Kha, 1 km E of Kop Dong, 1650 m, 13.xi.1998; 7 specimens, 12 km
NW of Chiang Dao, 750 m, 12.xi.1998, leg. T. Csővári & l. mikus; 6 specimens, between Chiang Dao and Kariang, 900 m, 98º48’E,
19º25’N, 26.x.2002, slide No. LGN 1091; 10 specimens, from the same site, 8.xi.2002, leg. b. HeRCzig & g. Ronkay; 1 specimen,
20 km NW of Mae Ai, 1650 m, 6.xii.1998, leg. m. HRebLay, i. soós & y. sHeRpani; 2 specimens, 6 km SE of Pang Faen, 1100 m,
29.i.1999, leg. a. szabó & z. CzeRe; 1 specimen, from the same locality, 21.viii.1999, leg. T. Csővári & l. mikus; 4 specimen, 4 km
SE of Pang Faen, 1100 m, 13.i.2004; 5 specimens, from the same site, 18.i.2004 and 26.i.2004, leg. p. HentsCHeL & a. szabó; 8
specimens, from the same locality, 27.i.2004 and 31.i.2004, leg. a. szabó; 1 specimen, Mt. Doi Phahompok, 16 km NW of Fang,
2000 m, 15.ii.1998, leg. m. HRebLay & Cs. szabóky; 1 specimen, Doi Inthanon, 2300 m, 23 km NW Sop Kha, 1650 m, 14.i.2004,
leg. a. szabó & p. HentsCHeL. Prov. Chiang Rai: 2 specimens, 1 km SE of Khun-Kon Waterfall, 600 m, 15.xi.1998, leg. T. Csővári
& L. mikus. Prov. Mae Hong Song: 2 specimen, between Pa Pae and Khun Sa, 1250 m, 98º39’E, 19º08’N, leg. b. HeRCzig & g.
Ronkay; 2 specimens, 10 km NE of Pai, 1560 m, 3.xii.1998, leg. m. HRebLay, i. soós & y. sHeRpani; 1 specimen, 21 km NW of Pai,
1360 m, 7.ii.1998, leg. m. HRebLay & Cs. szabóky (coll. G. Ronkay and MWM).
„Nola” quadrimaculata species group (2 distinct species)
(Pl. 10, gs 6-7)
Material examined:
Thailand. Prov. Chiang Mai: 1 x, 10 km NW of Fang, 550 m, 14.xi.1998, slide No. LGN 1136 (W 15007); 1 specimen, 12 km NW
of Chiang Dao, 750 m, 12.xi.1998, leg. T. Csővári & l. mikus; Prov. Nan: 1 fem, 5 km N of Ban Luang, 350 m, between Pi Nai
and Pi Tai, 100º27’E, 18º56’N, 4.xi.2002, slide No. LGN 1151 (W 15008); 1 specimen, between Chiang Dao and Kariang, 900
m, 98º48’E, 19º25’N, 26.x.2002, leg. b. HeRCzig & g. Ronkay; 1 x, 5 km N of Bo Luang, 1000 m, 4.ii.2000, leg. m. HRebLay, y.
sHeRpani & a. szabó, slide No. LGN 1135 (W 15009); 1 specimen, 5 km E of Bo Luang, 610 m, 23.xi.1998, leg. T. Csővári & l.
mikus (coll. MWM).
Remarks. There are two species occurring in Thailand which belong to the Nola quadrimaculata species-group.
The members of this group are externally almost indistinguishable, the exact identication requires the examina-
tion of the genitalia which display rather distinct specic features. As the species-group is currently under revision,
the two Thai species are yet unidentied.
There are, however, some difculties which make the revision of the group rather problematic. Unfortunately, the
abdomen of the w type of N. internella is lost and, the type of N. quadrimaculata HeyLaeRts has not been examined
by the authors and HoLLoWay (pers. comm.) as the type was not found in RMS, Leiden. Thus, the designation of the
neotypes of the two taxa would be indispensable for the monographic treatment of the N. internella species-group.
The revision of the species-group is intended to publish in a separate article.
62
„Nola” infralba inoue, 1976
(Pl. 10, g. 8)
Nola infralba inoue, 1976 Bulletin of the Faculty of Domestic Sciences, of Otsuma Woman’s University 12: 165. Type-locality: Japan,
Chiba Prefecture, Awa-gun, near Kamogawa-machi, Kami. Holotype: in coll inoue.
Material examined:
Thailand. Prov. Nan: 1 x, 1700 m, 30 km E of Pua, 20.i.2004, leg. p. HentsCHeL & a. szabó, slide No. LGN 1016 (W 15010) (coll.
MWM).
Tshodanola gen. nov.
Type species: Tshodanola gabriella spec. nov.
Species content:
Tshodanola gabriella spec. nov.
Diagnosis. The only known species of the new genus has a rather unusual external appearance, it differs from
the taxa of all other Noline genera by the following features: the forewing ground colour is shining golden brown,
the costal margin is decorated with a row of very prominent, elongated dark red-brown spots appearing as basal
dots of the basal, antemedial, medial, postmedial and praeterminal lines. The crosslines are slightly arcuate,
antemedial and medial lines relatively broad and diffuse, dark grey, postmedial line rather narrow, pale yellowish
brown, praeterminal line interrupted, consisting of a row of conspicuous dark red-brown patches, terminal line
continuous, pale grey.
The x genitalia of the new genus indicate the close relationship between Tshodanola and Nola: the clasping ap-
paratus is characterized by the medium deeply incised valvae with apically slightly dilated arms (a feature typical
of Nola), and the short, robust, slightly arched, apically pointed harpe. The very narrow and rather elongated, acute
vinculum of the new genus is characteristic for Tshodanola, such vinculum is unusual and very rare in Nola.
The main generic feature of the x genitalia is the presence of a huge cornuti eld in the vesica, consisting of nu-
merous small, but elongated and acute cornuti; no such cornuti eld with so large number of cornuti is known in
Nola. The only exception is the „Nola” marginata-shakishimana-umbrata species complex, which represent most
probably a distinct genus (see also the remarks under N. marginata), this group has a large cornuti eld in the
vesica, but the clasping apparatus completely different.
The w genitalia of the new genus is characterized by the rather short papillae anales, the relatively long apophyses
posteriores, the short, simple 8th segment with short apophyses anteriores, the simple, rounded ostium bursae,
the sclerotized, gradually dilated ductus bursae, the relatively small cervix bursae, the elongated-ovoidal corpus
bursae and by the small, but conspicuous, strongly sclerotized, spine-like signum.
Tshodanola gabriella spec. nov.
(Pl. 10, gs 15-16; gen. g. 57)
Holotype. x: „China, Guangxi, Dayao Shan, Jingxiu, 100 km SE of Liuzhou, 23º45’N, 109º45’E, 1200m, 15-30.iii.2005, leg. siniaev
& his team”, slide No. LGN 1161 (W 15017) (coll. MWM).
Paratypes. China. Guangxi: 4 ww, with the same data as the holotype, slide Nos LGN 1049 (W 15018), LGN 1050 (W 15019)
(coll. MWM). Thailand. Prov. Nan: 1 w, Doi Phuka N.P., between Pua and Bo Luang, 1350 m, 101˚05’E, 19˚12’N,
3.xi.2002, leg. b. HeRCzig & g. Ronkay, slide No. LGN 781 (G. Ronkay). Vietnam. Prov. Lao Cai: 3 ww, Fan-si-
pan Mts, 1920 m, 4 km SW Cat Cat, 14.v.1998, leg. Frontier Vietnam (coll. HNHM); 1 w, Mt. Fan-si-pan, Sa Pa,
Nebelwald, 2400 m, 22º15’N, 103º45’E, 2-4.iii.1995, leg. DR. R. bReCHLin (coll. MWM).
Diagnosis. The diagnostic features of the new species are given above, under the diagnosis of the genus Tshod-
anola.
63
Distribution. The new species is distributed in the northern parts of Thailand and Vietnam and the neighbouring
province in south-eastern China (Guangxi).
„Nola” shakishimana inoue, 2001
(Pl. 10, g. 19)
Nola sakishimana inoue, 2001, Japan Heterocerists’ Journal 212: 226, gs 5-6, 18-20. Type-locality: Japan, Maesato, Ishigakijima.
Holotype: w, in coll. BMNH.
Material examined:
Thailand. Prov. Mae Hong Song: 1 x, 21 km NW of Pai, 1360 m, 7.ii.1998, leg. m. HRebLay & Cs. szabóky, slide No. LGN 1065
(W 15023) (coll. MWM).
„Nola” spec. close to Nola marginata
(Pl. 10, gs 17-18)
Material examined:
Thailand. Prov. Chiang Mai: 2 ww, 4 km SE of Pang Faen, 1100 m, 31.i.2004, leg. a. szabó, slide No. LGN 1160 (W 15020); 1 x,
between Chiang Dao and Kariang, 900 m, 98º48’E, 19º25’N, 26.x.2002, slide No. LGN 1260; 1 x, 4 km S of Kop Dong, 1800 m,
99º03’E, 19º52’N, 6.xi.2002, slide No. LGN 1261, leg. b. HeRCzig & g. Ronkay. Prov. Nan: 1 x, 30 km E of Pua, 1700 m, 5.ii.2000,
slide No. LGN 520 (W 15021), leg. m. HRebLay, y. sHeRpani & a. szabó; 1 x, Doi Phuka NP, between Pua and Bo Luang, 1350 m,
101º05’E, 19º12’N, 3.xi.2002, leg. b. HeRCzig & g. Ronkay. Prov. Mae Hong Song: 1 x, between Pa Pae and Khun Sa, 1250 m,
98º39’E, 19º08’N, leg. b. HeRCzig & g. Ronkay, slide No. LGN 1259 (W 15022) (coll. G. Ronkay and MWM).
Remarks. The „Nola” marginata-shakishimana-umbrata species-complex is a rather distinct phyletic group display-
ing certain unique morphological features which differ signicantly from the important generic features of Nola.
The species of the complex is presently under revision; the results, containing the description of a new genus for
the lineage, will be published in a separate paper.
„Nola” cretacea (Hampson, 1901)
(Pl. 10, gs 9-11)
Celama cretacea Hampson, 1901, Annals and Magazine of Natural History 7: 177. Type-locality: [India][Maharashtra] [Mumbai]
Bombay. Syntypes: four ww, in coll. BMNH.
Nola calcicola HoLLoWay, 2003, The Moths of Borneo 18: 55, pl. 3, g. 111. Type-locality: Borneo, Sarawak. Holotype: x, in coll.
BMNH; syn. n.
Type material examined:
Syntype w of Celama cretacea Hampson, 1901: „TYPE” (Red ring type label), „Bombay”, slide No. BM Arct. 1801 (coll. BMNH).
Holotype of Nola calcicola HoLLoWay, 2003: „Borneo, Sarawak, Gunong Mulu National Park, R.G.S. exped., 1977-8 (g.D. Hol-
loway et al.) Site 26, April, G. Api, Pinnacle, 1200 m, 428545, open scrub, slide No. BM Noct. 17097” (coll. BMNH).
Additional material examined:
Thailand. Prov. Chiang Mai: 1 w, between Fang and Nor Lae, 99º06’E, 20º02’N, 28.x.2002, slide No. LGN 1096; 1 w, 4 km W of
Pha Lak, 100º34’E, 19º21’N, 5.xi.2002, slide No. LGN 1095, leg. b. HeRCzig & g. Ronkay; 1 x, 2 km S of Ban Kum, 1700 m, 23-
24.vi.1998, leg. I. soós & a. szabó, slide No. LGN 1148 (W 15011); 1 w, 4 km SE of Pang Faen, 1100 m, 31.i.2004, leg. a. szabó,
slide No. LGN 1139 (W 15012). Prov. Nan: 1 x, 30 km E of Pua, 1700 m, 20.i.2004, leg. p. HentsCHeL & a. szabó, slide No. LGN
1138 (W 15013) (coll. G. Ronkay and MWM).
64
Remarks. The syntype series of N. cretacea consists of only w specimens. HoLLoWay (2003) failed to examine the
w syntypes of N. cretacea and described N. calcicola based on the x holotype. The genitalia of all x specimens
of N. cretacea from Thailand and Nepal are matching well with those of the holotype of N. calcicola. Thus, the
species described from Borneo represent the x sex of the more widespread N. cretacea, and is synonymised
here with N. cretacea.
Neonola mesosticta Hampson, 1900
(Pl. 10, gs 4-5)
Neonola mesosticta Hampson, 1900, Catalogue of the Lepidoptera Phalaenae in the British Museum 2: 4, g. 2. Type-locality:
[Borneo] [Laut Island] Pulo Laut. Holotype: x, in coll. BMNH.
Material examined:
Thailand. Prov. Chiang Mai: 1 specimen, between Chiang Dao and Kariang, 900 m, 98º48’E, 19º25’N, 26.x.2002; 1 specimen,
between Fang and Nor Lae, 99º06’E, 20º02’N, 28.x.2002, leg. b. HeRCzig & g. Ronkay. Prov. Nan: 1 x, 30 km E of Pua, 1700 m,
20.i.2004, leg. p. HentsCHeL & a. szabó, slide No. LGN 994 (W 15006) (coll. G. Ronkay and MWM).
Spininola gen. nov.
Type species: Nola loxoscia Hampson, 1900
Species content:
S. loxoscia (Hampson, 1900) comb. nov.
S. vesiculalis (eeCke, 1926) comb. nov.
S. trilinea (maRumo, 1923) comb. nov.
S. denticulata (mooRe, 1888) comb. nov.
S. fuscibasalis (Hampson, 1896) comb. nov.
S. armata spec. nov.
Diagnosis. The new genus represents a rather compact phyletic unit being closely related to Nola. The external
appearance of the species of Spininola can be rather variable, no common diagnostic feature is found. The sepa-
ration of the new genus is based on the conguration of the x genitalia. The new genus is characterized by the
following features of the x genitalia: Spininola has a ne, usually rather short uncus, while the species of Nola
always lack the uncus. The valvae of Spininola are very deeply incised medially, the dorsal arm of the valva is
narrow at basal third, then dilated distally and rounded apically. The ventral arm of the valva has slightly curved
ventral and more or less straight dorsal margin, the sacculus is narrow but strongly sclerotized. The apex (and
sometimes the dorsal margin of the ventral arm) of the valva is armed with strong, long and acute, easily remov-
able spines of different numbers; Nola lacks such spines from the valvae. The harpe of Spininola is conspicuously
long, elongated and more or less S-shaped, it is slender but strongly sclerotized and apically pointed; the harpe
of Nola is always shorter and differently shaped. The vinculum of the new genus is simple, shortly triangular and
apically pointed; the aedeagus is medium-long, simple, slightly arched, the vesica is without cornuti, but sometimes
with a scobinated section.
The w genitalia of the new genus have relatively short, conical papillae anales, long apophyses posteriores, very
short 8th tergite and rather broad 8th sternite, with strongly sclerotized lobes at the base of the rather short and
thin apophyses anteriores. Ostium bursae relatively broad, more or less cup-shaped; sclerotized part of ductus
bursae short; ductus bursae relatively long and narrow, cervix bursae weakly developed. Corpus bursae ovoid,
signum bursae consists of a longitudinal, scobinate band and a pair of rather long, strong, nger-like processi with
pointed apices.
Spininola loxoscia (Hampson, 1900)
(Pl. 10, gs 12-13; gen. g. 55)
Nola loxoscia Hampson, 1900, Catalogue of the Lepidoptera Phalaenae in the British Museum 2: 33, pl. 18, g. 33. Type-locality:
65
[India] Sikkim. Holotype: x, in coll. BMNH.
Material examined:
Thailand. Prov. Nan: 2 specimens, Doi Phuka NP, between Pua and Bo Luang, 1350 m, 101º05’E, 19º12’N, 3.xi.2002, leg. b. HeRCzig
& g. Ronkay; 1 x, 25 km N of Bo Luang, 1150 m, 11.xi.1999, leg. a. szabó & z. CzeRe. Prov. Chiang Mai: 1 x, Mt. Doi Inthanon,
NP, 2300 m, 16.i.2004, leg. p. HentsCHeL & a. szabó, slide No. LGN 1001 (W 15014) (coll. MWM).
Spininola armata spec. nov.
(Pl. 10, g. 14; gen. g. 56)
Holotype. x, „Thailand, Changwat Chiang Mai, 4 km SE of Pang Faen, 1100m, 31.i.2004., Leg.: Attila Szabó”, slide No. LGN 982
(W 15015) (coll. MWM).
Paratypes. Thailand, Prov. Nan: 3 xx, 25 km N of Bo Luang, 1150 m, 03.ii.2000 leg. m. HRebLay & a. szabó, slide No. LGN 1257
(W 15016). Prov. Chiang Mai: 1 x, 4 km S of Kop Dong, 1800 m, 99º03’E, 19º52’N, 6.xi.2002, leg. b. HeRCzig &
g. Ronkay (coll. MWM). 1 x, Prov. Chiang Mai, 1600 m, between Fang and Nor Lae, 99º06’E, 20º02’N, 12.XI.2002
leg. B. HeRCzig & g. Ronkay (coll. G. Ronkay).
Diagnosis. The new species is similar externally to Spininola vesiculalis described from Sumatra by its greyish
forewing ground colour and the ne darker grey streaks and spots on the forewing, but easily distinguishable by
the following features: S. armata has more greyish forewing ground colour, while S. vesiculalis is rather brownish
grey. In the x genitalia, the new species have much shorter uncus, somewhat shorter and more slender dorsal
arm of valva, slightly shorter and much broader ventral arm of valva, and conspicuously shorter, somewhat thin-
ner harpe compared with those of S. vesiculalis. The shape of the aedeagus of the two species is rather similar:
medium-long and narrow, medially slightly curved, but S. armata has a scobinated section in the vesica which is
absent in S. vesiculalis. The w is unknown.
Distribution. Northern Thailand.
Njalkanola gen. nov.
Type species: Celamoides bimaculata eeCke, 1920
Species content:
Njalkanola bimaculata (eeCke, 1920)
Diagnosis. The only known species of the new genus has numerous apomorphic features in the genitalia of both
sexes. The x genitalia display the relationship with Nola by the absence of uncus and the bilobate valva, all other
features and the general structure of the copulatory organ are diagnostic for Njalkanola. The valva is shallowly
incised, the dorsal arm of valva is very broad, weakly sclerotized, tapering apically, with short and rounded apical
process; the ventral arm of the valva is narrower and more strongly sclerotized than the dorsal one, having an
elongated, digitiform apical process. A very characteristical feature of Njalkanola is the extraordinarily long, thin,
almost straight, apically curved and pointed harpe, this structure is unique within the whole subfamily. The acutely
pointed apex of the aedeagus and the rather extensive scobination of the vesica are also distinctive features of
the genus.
In the w genitalia, Njalkanola has very narrow, elongated papillae anales with very long, basally rather thick apo-
physes posteriores, relatively long 8th segment with medium-long apophyses anteriores. The most characteristic
feature of the new genus is the extremely long, strongly sclerotized and partly scobinate, wrinkled and folded
ductus bursae surrounded by a rather thick, mucous membrane. The corpus bursae of Njalkanola is conspicuously
small, rounded, without signum.
66
Njalkanola bimaculata (eeCke, 1920) comb. nov.
(Pl. 10, gs 20-21; gen. g. 58)
Celamoides bimaculata eeCke, 1920, Zoologische Mededeelingen Leiden 5: 121, g. 10. Type-locality: [Indonesia] Java, Preanger.
Syntypes: one x and three ww, in coll. RNH Leiden.
Material examined:
Thailand. Prov. Chiang Mai: 2 specimens, between Fang and Nor Lae, 1600 m, 99º06’E, 20º02’N, 7.xi.2002; 5 specimens, from
the same site, 28.x.2002, slide No. LGN 983 (W 15024) (x); 2 specimens, from the same site, 12.xi.2002, slide No. LGN 984 (W
15025) (w); 8 specimens, between Chiang Dao and Kariang, 900 m, 98º48’E, 19º25’N, 26.x.2002; 1 specimen, from the same
site, 8.xi.2002; 1 specimen, 4 km S of Kop Dong, 99º03’E, 19º52’N, 1800 m, 6.xi.2002, leg. b. HeRCzig & g. Ronkay. Prov. Mae
Hong Song: 1 specimen, between Pa Pae and Khun Sa, 1250 m, 98º39’E, 19º08’N, 31.x.2002, leg. b. HeRCzig & g. Ronkay. Prov.
Nan: 1 specimen, Doi Phuka NP, between Pua and Bo Luang, 1350 m, 101º05’E, 19º12’N, 3.xi.2002, leg. b. HeRCzig & g. Ronkay
(coll. G. Ronkay and MWM).
Acknowledgements
The authors are indebted to Mr maRtin R. Honey, Mr geoFF maRtin and Dr JeRemy HoLLoWay (BMNH London) for the opportunity to
check the type material in their museum collections. We are also grateful to Dr miCHaeL FibigeR (Sorø) and Dr WoLFgang speiDeL
(Olching) for the kind loan of materials for study. Our sincere thanks to Dr LászLó Ronkay (HNHM Budapest) for his kind help in
the study of the Nolidae material of the Hungarian Natural History Museum and the useful advices during the preparation of this
paper.
This research received support from the SYNTHESYS Project http://www.synthesys.info/ which is nanced by European Com-
munity Research Infrastructure Action under the FP6 „Structuring the European Research Area” Programme.
67
References
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