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New Species of the Tenebrionid Beetle Genus Cylindronotus Fald. (Coleoptera, Tenebrionidae) from the North Caucasus

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  • Daghestan Federal Research Centre of the Russian Academy of Sciences

Abstract and Figures

Two new species of the genus Cylindronotus Fald. from the North Caucasus are described: C. dom-baicus sp. n. from Karachai-Cherkess Republic and C. alanicus sp. n. from North Ossetia. A new subgenus, Cau-casonotus subgen. n. (type species Cylindronotus dombaicus sp. n.), is erected to include the two new species in addition to C. diteras All. and C. adriani Rtt. The paper presents descriptions of two new species of the genus Cylindronotus: C. dombaicus sp. n. and C. alanicus sp. n. from Karachai-Cherkess Republic and North Ossetia. The species are placed in the new subgenus Caucasonotus subgen. n., in addition to C. alanicus Rtt. and C. diteras All. previously assigned by Reitter (1922) to the subgenus Nalassus. The holotypes and part of paratypes are deposited at the Zoological Institute, Russian Academy of Sciences, St. Petersburg; the rest of types, in the Zoological Mu-seum, Faculty of Zoology, Rostov State University, Rostov-on-Don.
Content may be subject to copyright.
Entomological Review, Vol. 80, No. 1, 2000, pp. 50–53. Translated from Entomologicheskoe Obozrenie, Vol. 79, No. 1, 2000, pp. 107–111.
Original Russian Text Copyright © 2000 by Nabozhenko.
English Translation Copyright © 2000 by MAIK “Nauka/Interperiodica” (Russia).
50
New Species of the Tenebrionid Beetle Genus Cylindronotus
Fald. (Coleoptera, Tenebrionidae) from the North Caucasus
M. V. Nabozhenko
Rostov State University, Rostov-on-Don, Russia
Received February 25, 1999
Abstract—Two new species of the genus Cylindronotus Fald. from the North Caucasus are described: C. dom-
baicus sp. n. from Karachai-Cherkess Republic and C. alanicus sp. n. from North Ossetia. A new subgenus, Cau-
casonotus subgen. n. (type species Cylindronotus dombaicus sp. n.), is erected to include the two new species in
addition to C. diteras All. and C. adriani Rtt.
The paper presents descriptions of two new species
of the genus Cylindronotus: C. dombaicus sp. n. and
C. alanicus sp. n. from Karachai-Cherkess Republic
and North Ossetia. The species are placed in the new
subgenus Caucasonotus subgen. n., in addition to C.
alanicus Rtt. and C. diteras All. previously assigned
by Reitter (1922) to the subgenus Nalassus. The
holotypes and part of paratypes are deposited at the
Zoological Institute, Russian Academy of Sciences, St.
Petersburg; the rest of types, in the Zoological Mu-
seum, Faculty of Zoology, Rostov State University,
Rostov-on-Don.
GENUS CYLINDRONOTUS FALD.
Subgenus CAUCASONOTUS Nabozhenko, subgen. n.
Type species C. dombaicus sp. n.
Description. Eyes small, widely separate and very
weakly convex (in dorsal view). Lateral margins of
pronotum flattened along entire length or, at least, at
posterior and anterior angles. Anterior pronotal angles
acute, often widely rounded, noticeably protruding
forward. All antennal segments rounded in cross-
section, except slightly compressed 9–11th. First visi-
ble sternite of male abdomen with spot of long coarse
hairs. Surface of sternite under the spot strongly shin-
ing, with sparser punctation; punctures large and elon-
gate. Aedeagus compressed and keel-shaped produced
apically.
Differential diagnosis. This subgenus belongs to
the group of subgenera with apically compressed
aedeagus (Fig. 2). In the Caucasus, this group is also
represented by the subgenera Nalassus and Helopo-
cerodes. The subgenus Caucasonotus is most closely
related to Nalassus; clearly differing from this and all
other subgenera in the not convex and widely sepa-
rated eyes (Fig. 1a) and punctation of the first visible
sternite of male abdomen under the hairy spot; from
Nalassus, it additionally differs in the protruding ante-
rior angles of pronotum. Along with the species de-
scribed here, the subgenus Caucasonotus also includes
C. adriani Rtt. and C. diteras All. previously placed in
Nalassus by Reitter (1922). All known species of the
subgenus Caucasonotus occur in the alpine and subal-
pine belts, being distributed within the Caucasus;
whereas species of Nalassus are mainly European
forest inhabitants, with the only species, C. (Nalassus)
brevicollis Kryn., occurring in foothills of the W Cau-
casus.
Cylindronotus (Caucasonotus) dombaicus
Nabozhenko, sp. n.
Material. Holotype: b-
lic, Mt. Semenov-Bashi (W of Dombai), 2800 m,
21.VI.1997 (M.V. Nabozhenko). Paratypes: 10
23
D.A. Dubovikov); 2
25.VI.1997 (M.V. Nabozhenko); 1
Turye (Dzhuguturlukel), 20.VI.1997 (M.V. Nabo-
zhenko).
Description. Male. Body black, dull. Head widest
at the level of eyes. Outer margin of clypeus very
weakly arcuate. Genae before eyes gently rounded,
protruding. Lateral margin of head with hardly notice-
able emargination between apex of genal angle and
outer angles of clypeus. Eyes weakly convex. Ratio of
head width to distance between eyes 1.33. Temples
roundly converging to neck constriction. Head punc-
NEW SPECIES OF THE TENEBRIONID BEETLE GENUS CYLINDRONOTUS FALD.
ENTOMOLOGICAL REVIEW Vol. 80 No. 1 2000
5
1
tation coarse and dense, punctures often merged (es-
pecially on sides). Antennae widening to apex, ex-
tending as far as elytral base. Second antennal segment
slightly elongate (1.3);1 3rd strongly elongate (2.5),
1.8 times as long as 2nd and 1.4 times, as 4th; 46th
segments elongate (1.5); 79th slightly elongate (1.2
1.3); 10th rounded, as long as wide; 11th elongate
(1.6), with blunt conical apex. All antennal segments
covered with pale hairs denser on apical segments.
Pronotum transverse (length 1.3 times width) [defi-
nitely 1.3 times as wide as long.Ed.]; widest before
the middle; weakly rounded laterally; narrowly flat-
tened at posterior and anterior angles and, often, also
along lateral margin. Pronotal sides not emarginate
before posterior angles, weakly arcuate or linearly
converging to base. Posterior pronotal angles well
defined, slightly obtuse; anterior angles acute, widely
rounded, and extending as far as posterior eye margin.
Pronotum emarginate anteriorly, less deeply so at
base; base weakly emarginate; sides and base with
wide distinct, anterior margin, with very narrow mar-
gination. Pronotal disc uniformly strongly convex,
glabrous, coarsely and rather densely punctate (inter-
vals between punctures subequal to puncture diame-
_____________
1 Numbers in parentheses give length to width ratio for antennal
segments.
ter). Propleura coarsely rugose. Elytra oblong (1.5
times as long as wide). Elytral intervals flattened;
striae narrow but distinct, with punctures merged into
a single entire row. Surface of intervals with fine
regular punctation (34 punctures across interval) and
small but deep transverse rugae. Elytral epipleura with
deep longitudinal and transverse rugae and deep basal
depression. Body ventrally glabrous; mesepisterna
with dense coarse punctation; metepisterna weakly
punctate, shining. Abdominal sternites with moder-
ately dense punctation; laterally, punctures elongate,
often fused. First visible sternite with V-shaped spot
of dense golden-rufous hairs; surface under the spot
shining, with large elongate punctures. Fore tarsus
rather narrow; its 2nd4th segments cordate, weakly
transverse; 2nd and 3rd segments 0.8 times as wide as
long; 4th 0.9 times as wide as long; 5th 2.8 times as
long as wide. Middle and hind tarsi narrow. Body
length 8 mm.
Male genitalia. Aedeagal apex keel-shaped, pro-
duced. The keel obtuse-angled (in lateral view),
sharply and linearly turning into apex. Sides of genita-
lia sparsely punctate (Fig. 2b).
Female. Usually larger than male. Antennae slight-
ly narrower and shorter. Pronotum occasionally with
two noticeable admedian depression. First visible ab-
dominal sternite without hairy spot.
Fig. 1. Cylindronotus Fald.: (a, b) head; (c, d) pronotum. (a) C. diteras All.; (b) C. brevicollis Kryn.; (c) C. dombaicus sp. n.;
(d) C. alanicus sp. n.
NABOZHENKO
ENTOMOLOGICAL REVIEW Vol. 80 No. 1 2000
52
Ecology. Occurs in the alpine belt, usually on dry
areas or in organic debris between stones.
Comparative notes. The species is similar to Cyl-
indronotus alanicus sp. n.; differing from it in the
sides of pronotum, not emarginate at base; presence of
transverse rugae on the elytra; punctures in elytral
striae, merged into entire rows; rounded genae; and
presence of a hairy spot only on the first visible ab-
dominal sternite, whereas C. alanicus sp. n. has such
spots on the 1st and 2nd sternites.
Cylindronotus (Caucasonotus) alanicus
Nabozhenko, sp. n.
Material. Holotype: Khuma
ratkhokh, 25.VII.1997 (D.G. Kasatkin). Paratypes:
2 t-
kin); 3 Kariukhokh, 5.VII.1997 (D.G.
Kasatkin); 2
(M.V. Nabozhenko, I.V. Shokhin).
Description. Male. Body dark fuscous, weakly
shining; tibiae, antennae, and mouthparts dark rufous.
Anterior margin of clypeus straight. Outer margins of
genae before eyes linearly converging to clypeus. Lat-
eral margin of head with small emargination between
apex of genal angle and outer angle of clypeus. Eyes
weakly convex. Ratio of head width at the level of
eyes to distance between eyes 1.4. Temples roundly
converging to neck constriction. Head punctation
rather dense; punctures not merged. Antennae widen-
ing to apex. Second antennal segment rounded (nearly
as long as wide); 3rd strongly elongate (2.5), 2.3 times
as long as 2nd and 1.4 times, as 4th; 4th elongate (2);
Fig. 2. Cylindronotus Fald., parameres, dorsal and lateral view: (a) C. alanicus sp. n.; (b) C. dombaicus sp. n.; (c) C. diteras All.;
(d) C. adriani Rtt.
NEW SPECIES OF THE TENEBRIONID BEETLE GENUS CYLINDRONOTUS FALD.
ENTOMOLOGICAL REVIEW Vol. 80 No. 1 2000
5
3
58th moderately elongate (1.5); 9th 1.4 times as long
as wide; 10th weakly elongate (1.2); 11th elongate
(1.5). All antennal segments covered with rufous hairs.
Pronotum transverse (length 1.4 times width) [defi-
nitely, 1.4 times as wide as longEd.]; widest in the
middle; its sides noticeably arcuate, emarginate at
base; base weakly emarginate (Fig. 1d); anterior mar-
gin noticeably emarginate; margination on sides well
defined, wider near middle and at base; margination of
anterior margin medially hardly visible, smoothed.
Pronotal disc evenly strongly convex, flattened only at
posterior and anterior angles, glabrous, rather densely
punctate (intervals between punctures equal to punc-
ture diameter); punctures large, elongate on sides of
disc. Propleura with rather small but distinct rugae.
Elytra elongate (1.5 times as long as wide). Elytral
intervals almost flat; striae consisting of rounded
punctures not merged into entire rows. Surface of ely-
tral intervals without transverse rugae, uniformly cov-
ered with fine punctation. Body ventrally glabrous,
shining. Epipleura shining, rugose, with deep basal
depression. Abdominal sternites with uniform puncta-
tion; laterally, often with fine rugae at bases. First and
second visible sternites with spots of dark rufous hairs.
Spot on 1st sternite large, V-shaped; that on 2nd ster-
nite rather small, transverse. Surface of sternites under
the spots shining, with large elongate punctures.
Fore tarsus as in the preceding species. Body length
7.5 mm.
Male genitalia. Parameres weakly hastate, keel (in
lateral view) with more acute apex.
Female. Differs from male in more strongly arcuate
anterior margin of pronotum and abdominal sternites
lacking hairy spots.
Comparative notes. The species is similar to the
preceding one (C. dombaicus sp. n.); differing from it
in the pronotal sides emarginate at base; genae linearly
converging to clypeus; absence of transverse rugae on
elytra; and elytral punctures not merged into entire
striae. In addition, C. alanicus sp. n. has two hairy
spots on the 1st and 2nd visible abdominal sternites.
Key to Species of the Subgenus Caucasonotus
subgen. n. of the Genus Cylindronotus
1(2). Anterior angles of pronotum with acute apices,
not rounded; lateral margination of pronotum
wide at anterior angles, separated from disc by
rather deep sulcus. Body widely oval, strongly
convex, and shining; 1st and 2nd visible sternites
of male abdomen with V-shaped spots of long
coarse hairs, surface under the spots slightly de-
pressed. 68 mm. Caucasus, subalpine belt .........
....................................................... C. diteras All.
2(1) Anterior angles of pronotum rounded apically.
3(4) All segments of male antennae strongly thick-
ened; 2nd6th segments the thickest. Pronotum
narrowly flattened along entire length of lateral
margin. Body strongly elongate, parallel-sided;
elytra not convex. Dark fuscous. 910 mm.
Northern Abkhazia (Klukhor Pass) ......................
...................................................... C. adriani Rtt.
4(3) Antennal segments in male not thickened, gradu-
ally widening to apex; 811 segments the thick-
est. Elytra convex, not parallel-sided.
5(6) Sides of pronotum not emarginate at base. Pro-
notal disc narrowly flattened along entire length
of lateral margin (Fig. 1c). Punctation on pro-
notum coarse. Outer margins of genae gently
rounded along entire length. Punctures in elytral
striae fused into entire rows. Elytral intervals
covered with distinct transverse rugae. First visi-
ble sternite of male abdomen with hairy spot.
Black, dull. 7.59 mm. Karachai-Cherkess Re-
public ................................... C. dombaicus sp. n.
6(5) Outer lateral margins of pronotum shallowly but
distinctly emarginate at base. Pronotal disc flat-
tened at posterior and anterior angles (Fig. 1a).
Outer margins of genae linearly converging to
clypeus. Punctures in elytral striae not fused into
entire rows. Elytral intervals without transverse
rugae. First and second visible sternites of male
abdomen with hairy spots. Dark fuscous, weakly
shining. 79 mm. North Ossetia ...........................
.................................................. C. alanicus sp. n.
REFERENCES
1. Allard, E., Revision des Helopides vrais de Lacordare,
LAbeille, 1876, vol. 14, pp. 180.
2. Allard, E., Revision des Helopides vrais, Mitt. Schweiz.
Ent. Ges., 1880, pp. 12268.
3. Ardoin, P., Contribution a letude des Helopinae de
France, Ann. Soc. Ent. France, 1958, pp. 949.
4. Reitter, E., Bestimmungstabelle der palaearktischen
Helopina: II. Teil, Wien. Ent. Ztg., 1922, vol. 39,
pp. 113171.
5. Seidlitz, G., Naturgeschichte der Insecten Deutsch-
lands. Coleoptera, Vol. V, 1898.
... Species of the genus Nalassus are widespread in the Palaearctic from the Atlantic to the Pacific. Most diversity of the genus is observed in the Western Palaearctic, especially in the Mediterranean (Antoine 1949, Ardoin 1958, Español 1961, Nabozhenko and Löbl 2008, on the Caucasus (Nabozhenko 2000, 2001, 2008a, 2013b, Nabozhenko and Dzhambazishvili 2001, Nabozhenko and Abdurakhmanov 2007, Abdurakhmanov and Nabozhenko 2011, in Iran and Turkmenistan (Medvedev 1987, 1999, Nabozhenko 2006, 2010, 2014. Isolated generic enclave is located in the South-Eastern Kazakhstan and Western China (Tarbagatai) (Medvedev 1987, Nabozhenko 2012, Pacific region (Russian Far East, Korean Peninsula, Eastern China and Japan (Nabozhenko 2012, Nabozhenko andIvanov 2015). ...
... The genus Nalassus included 4 subgenera: nominotypical, Helopondrus, Caucasonotus Nabozhenko, 2000 and Helopocerodes. The first two subgenera are distributed in Europe (to Western Kazakhstan on east), Eastern Anatolia, Iran, on the Caucasus and locally in North Africa. ...
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... The genus Nalassus Mulsant, 1854 is well studied on the Caucasus [Nabozhenko, 2000[Nabozhenko, , 2001[Nabozhenko, , 2005Nabozhenko, Dzhambazishvili, 2001;Nabozhenko, Abdurakhmanov, 2007;Abdurakhmanov, Nabozhenko, 2011]. Twenty two species of four subgenera are known from the region to the present time. ...
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A review of genus-group names for darkling beetles in the family Tenebrionidae (Insecta: Coleoptera) is presented. A catalogue of 4122 nomenclaturally available genus-group names, representing 2307 valid genera (33 of which are extinct) and 761 valid subgenera, is given. For each name the author, date, page number, gender, type species, type fixation, current status, and first synonymy (when the name is a synonym) are provided. Genus-group names in this family are also recorded in a classification framework, along with data on the distribution of valid genera and subgenera within major biogeographical realms. A list of 535 unavailable genus-group names (e.g., incorrect subsequent spellings) is included. Notes on the date of publication of references cited herein are given, when known. The following genera and subgenera are made available for the first time: Anemiadena Bouchard & Bousquet, subgen. nov. (in Cheirodes Gené, 1839), Armigena Bouchard & Bousquet, subgen. nov. (in Nesogena Mäklin, 1863), Debeauxiella Bouchard & Bousquet, subgen. nov. (in Hyperops Eschscholtz, 1831), Hyperopsis Bouchard & Bousquet, subgen. nov. (in Hyperops Eschscholtz, 1831), Linio Bouchard & Bousquet, subgen. nov. (in Nilio Latreille, 1802), Matthewsotys Bouchard & Bousquet, gen. nov. , Neosolenopistoma Bouchard & Bousquet, subgen. nov. (in Eurynotus W. Kirby, 1819), Paragena Bouchard & Bousquet, subgen. nov. (in Nesogena Mäklin, 1863), Paulianaria Bouchard & Bousquet, gen. nov. , Phyllechus Bouchard & Bousquet, gen. nov. , Prorhytinota Bouchard & Bousquet, subgen. nov. (in Rhytinota Eschscholtz, 1831), Pseudorozonia Bouchard & Bousquet, subgen. nov. (in Rozonia Fairmaire, 1888), Pseudothinobatis Bouchard & Bousquet, gen. nov. , Rhytinopsis Bouchard & Bousquet, subgen. nov. (in Thalpophilodes Strand, 1942), Rhytistena Bouchard & Bousquet, subgen. nov. (in Rhytinota Eschscholtz, 1831), Spinosdara Bouchard & Bousquet, subgen. nov. (in Osdara Walker, 1858), Spongesmia Bouchard & Bousquet, subgen. nov. (in Adesmia Fischer, 1822), and Zambesmia Bouchard & Bousquet, subgen. nov. (in Adesmia Fischer, 1822). The names Adeps Gistel, 1857 and Adepsion Strand, 1917 syn. nov. [= Tetraphyllus Laporte & Brullé, 1831], Asyrmatus Canzoneri, 1959 syn. nov. [= Pystelops Gozis, 1910], Euzadenos Koch, 1956 syn. nov. [= Selenepistoma Dejean, 1834], Gondwanodilamus Kaszab, 1969 syn. nov. [= Conibius J.L. LeConte, 1851], Gyrinodes Fauvel, 1897 syn. nov. [= Nesotes Allard, 1876], Helopondrus Reitter, 1922 syn. nov. [= Horistelops Gozis, 1910], Hybonotus Dejean, 1834 syn. nov. [= Damatris Laporte, 1840], Iphthimera Reitter, 1916 syn. nov. [= Metriopus Solier, 1835], Lagriomima Pic, 1950 syn. nov. [= Neogria Borchmann, 1911], Orphelops Gozis, 1910 syn. nov. [= Nalassus Mulsant, 1854], Phymatium Billberg, 1820 syn. nov. [= Cryptochile Latreille, 1828], Prosoblapsia Skopin & Kaszab, 1978 syn. nov. [= Genoblaps Bauer, 1921], and Pseudopimelia Gebler, 1859 syn. nov. [= Lasiostola Dejean, 1834] are established as new synonyms (valid names in square brackets). Anachayus Bouchard & Bousquet, nom. nov. is proposed as a replacement name for Chatanayus Ardoin, 1957, Genateropa Bouchard & Bousquet, nom. nov. as a replacement name for Apterogena Ardoin, 1962, Hemipristula Bouchard & Bousquet, nom. nov. as a replacement name for Hemipristis Kolbe, 1903, Kochotella Bouchard & Bousquet, nom. nov. as a replacement name for Millotella Koch, 1962, Medvedevoblaps Bouchard & Bousquet, nom. nov. as a replacement name for Protoblaps G.S. Medvedev, 1998, and Subpterocoma Bouchard & Bousquet, nom. nov. is proposed as a replacement name for Pseudopimelia Motschulsky, 1860. Neoeutrapela Bousquet & Bouchard, 2013 is downgraded to a subgenus ( stat. nov. ) of Impressosora Pic, 1952. Anchomma J.L. LeConte, 1858 is placed in Stenosini: Dichillina (previously in Pimeliinae: Anepsiini); Entypodera Gerstaecker, 1871, Impressosora Pic, 1952 and Xanthalia Fairmaire, 1894 are placed in Lagriinae: Lagriini: Statirina (previously in Lagriinae: Lagriini: Lagriina); Loxostethus Triplehorn, 1962 is placed in Diaperinae: Diaperini: Diaperina (previously in Diaperinae: Diaperini: Adelinina); Periphanodes Gebien, 1943 is placed in Stenochiinae: Cnodalonini (previously in Tenebrioninae: Helopini); Zadenos Laporte, 1840 is downgraded to a subgenus ( stat. nov. ) of the older name Selenepistoma Dejean, 1834. The type species [placed in square brackets] of the following available genus-group names are designated for the first time: Allostrongylium Kolbe, 1896 [ Allostrongylium silvestre Kolbe, 1896], Auristira Borchmann, 1916 [ Auristira octocostata Borchmann, 1916], Blapidocampsia Pic, 1919 [ Campsia pallidipes Pic, 1918], Cerostena Solier, 1836 [ Cerostena deplanata Solier, 1836], Coracostira Fairmaire, 1899 [ Coracostira armipes Fairmaire, 1899], Dischidus Kolbe, 1886 [ Helops sinuatus Fabricius, 1801], Eccoptostoma Gebien, 1913 [ Taraxides ruficrus Fairmaire, 1894], Ellaemus Pascoe, 1866 [ Emcephalus submaculatus Brême, 1842], Epeurycaulus Kolbe, 1902 [ Epeurycaulus aldabricus Kolbe, 1902], Euschatia Solier, 1851 [ Euschatia proxima Solier, 1851], Heliocaes Bedel, 1906 [ Blaps emarginata Fabricius, 1792], Hemipristis Kolbe, 1903 [ Hemipristis ukamia Kolbe, 1903], Iphthimera Reitter, 1916 [ Stenocara ruficornis Solier, 1835], Isopedus Stein, 1877 [ Helops tenebrioides Germar, 1813], Malacova Fairmaire, 1898 [ Malacova bicolor Fairmaire, 1898], Modicodisema Pic, 1917 [ Disema subopaca Pic, 1912], Peltadesmia Kuntzen, 1916 [ Metriopus platynotus Gerstaecker, 1854], Phymatium Billberg, 1820 [ Pimelia maculata Fabricius, 1781], Podoces Péringuey, 1886 [ Podoces granosula Péringuey, 1886], Pseuduroplatopsis Pic, 1913 [ Borchmannia javana Pic, 1913], Pteraulus Solier, 1848 [ Pteraulus sulcatipennis Solier, 1848], Sciaca Solier, 1835 [ Hylithus disctinctus Solier, 1835], Sterces Champion, 1891 [ Sterces violaceipennis Champion, 1891] and Teremenes Carter, 1914 [ Tenebrio longipennis Hope, 1843]. Evidence suggests that some type species were misidentified. In these instances, information on the misidentification is provided and, in the following cases, the taxonomic species actually involved is fixed as the type species [placed in square brackets] following requirements in Article 70.3 of the International Code of Zoological Nomenclature: Accanthopus Dejean, 1821 [ Tenebrio velikensis Piller & Mitterpacher, 1783], Becvaramarygmus Masumoto, 1999 [ Dietysus nodicornis Gravely, 1915], Heterophaga Dejean, 1834 [ Opatrum laevigatum Fabricius, 1781], Laena Dejean, 1821, [ Scaurus viennensis Sturm, 1807], Margus Dejean, 1834 [ Colydium castaneum Herbst, 1797], Pachycera Eschscholtz, 1831 [ Tenebrio buprestoides Fabricius, 1781], Saragus Erichson, 1842 [ Celibe costata Solier, 1848], Stene Stephens, 1829 [ Colydium castaneum Herbst, 1797], Stenosis Herbst, 1799 [ Tagenia intermedia Solier, 1838] and Tentyriopsis Gebien, 1928 [ Tentyriopsis pertyi Gebien, 1940]. The following First Reviser actions are proposed to fix the precedence of names or nomenclatural acts (rejected name or act in square brackets): Stenosis ciliaris Gebien, 1920 as the type species for Afronosis G.S. Medvedev, 1995 [ Stenosis leontjevi G.S. Medvedev, 1995], Alienoplonyx Bremer, 2019 [ Alienolonyx ], Amblypteraca Mas-Peinado, Buckley, Ruiz & García-París, 2018 [ Amplypteraca ], Caenocrypticoides Kaszab, 1969 [ Caenocripticoides ], Deriles Motschulsky, 1872 [ Derilis ], Eccoptostira Borchmann, 1936 [ Ecoptostira ], † Eodromus Haupt, 1950 [† Edromus ], Eutelus Solier, 1843 [ Lutelus ], Euthriptera Reitter, 1893 [ Enthriptera ], Meglyphus Motschulsky, 1872 [ Megliphus ], Microtelopsis Koch, 1940 [ Extetranosis Koch, 1940, Hypermicrotelopsis Koch, 1940], Neandrosus Pic, 1921 [ Neoandrosus ], Nodosogylium Pic, 1951 [ Nodosogilium ], Notiolesthus Motschulsky, 1872 [ Notiolosthus ], Pseudeucyrtus Pic, 1916 [ Pseudocyrtus ], Pseudotrichoplatyscelis Kaszab, 1960 [ Pseudotrichoplatynoscelis and Pseudotrichoplatycelis ], Rhydimorpha Koch, 1943 [ Rhytimorpha ], Rhophobas Motschulsky, 1872 [ Rophobas ], Rhyssochiton Gray, 1831 [ Ryssocheton and Ryssochiton ], Sphaerotidius Kaszab, 1941 [ Spaerotidius ], Stira Agassiz, 1846 (Mollusca) [ Stira Agassiz, 1846 (Coleoptera)], Sulpiusoma Ferrer, 2006 [ Sulpiosoma ] and Taenobates Motschulsky, 1872 [ Taeniobates ]. Supporting evidence is provided for the conservation of usage of Cyphaleus Westwood, 1841 nomen protectum over Chrysobalus Boisduval, 1835 nomen oblitum.
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Two new species of the tenebrionid beetle genus Nalassus, N. (Caucasonotus) svaneticus sp. n. and N. (Helopondrus) adzharicus sp. n., are described from the Caucasus (Georgia). One of the species of the genus Nalassus, described in the present paper, belongs to the endemic Caucasian subgenus Caucasonotus (Nabozhenko, 2000), whose representatives are distributed, with some exception, within the Great Caucasus. The other species belongs to the subgenus Helopondrus Reitter, whose range covers the Caucasus, N Iran, and the European part of the CIS; several species are known from Europe. Types of the described species are deposited at the Zoolo-gical Institute, Russian Academy of Sciences (St. Pe-tersburg), and part of type specimens, at the Institute of Zoology, Georgian Academy of Sciences (Tbilisi).
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