BookPDF Available

Pojoce kobilice Slovenije/Singing Orthoptera of Slovenia.

August 2014

Edition: 1
Publisher: EGEA, Institution for Nature
ISBN: 978-961-93692-0-3

Authors:

EGEA, Institution for Nature

The book of singing Orthoptera of Slovenia represents 97 species and two subspecies of bush-crickets and grasshoppers in sound and photographs. The book is richly illustrated with photographs as it is meant for the general public in order to make it easy to recognise species of Othoptera, based on images as well as sound. In order to facilitate the identification of species, each species is represented by four photos and an oscillogram of the song. Each species is also represented by basic data including the Slovenian and English common name, body size, audiblehearing distance from where each species can be heard, data on species habitats and known distribution in Slovenia. In addition to this book there is also included the first up to date checklist of currently known Orthoptera species in Slovenia. Several species in the book actually represent the first finds of species in Slovenia. Most species in the checklist are also indicated by their Slovenian common names. Based on all this information the book brings much new information on Orthoptera fauna in Slovenia. This will not only be useful just for the general public but also for the professionals who will find many novelties on Orthoptera fauna in Slovenia. In researching the checklist of Slovenian Orthoptera, we found out that there are currently 157 species present in Slovenia, 5 were deleted from the list, some species were updated to their current valid or revised names. Certainly, this checklist will now be much faster updated and supplemented with new species, as it represents a long-awaited review of Slovenian Orthoptera.

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POJOČE KOBILICE SLOVENIJE

*****

SINGING ORTHOPTERA OF SLOVENIA

STANISLAV GOMBOC

BLAŽ ŠEGULA

Ljubljana, 2014

Knjiga pojoče kobilice Slovenije predstavlja 97 vrst in dve podvrsti kobilic v zvoku in sliki. Knjiga je

slikovno bogato opremljena, saj je namenjena širši javnosti, za čim enostavnejše spoznavanje vrst

kobilic, tako po slikah kot tudi po zvoku. Za lažjo določitev je vsaka vrsta predstavljena s 4 fotograﬁ-

jami in oscilogramom napeva. Pri vsaki vrsti so predstavljeni osnovni podatki o vrsti, vključno s

slovenskim in angleškim poimenovanjem vrste, velikostjo telesa, slišno razdaljo, s katere lahko

slišimo posamezno vrsto, podatku o habitatih vrste in razširjenosti vrste v Sloveniji. Poleg slikovnega

dela, je knjigi dodan prvi ažuren popis vseh vrst kobilic, ki jih trenutno najdemo v Sloveniji. Nekaj vrst

v knjigi dejansko predstavlja tudi prve najdbe vrst pri nas. Prav tako je večina vrst dobila še poljudna

slovenska imena. Knjiga tako prinaša precej novosti na področju poznavanja kobilic v Sloveniji. Tako

ne bo uporabna le za širšo javnost ampak tudi za strokovnjake, ki bodo v njej našli precej novosti na

področju poznavanja kobilic Slovenije. V popisu vrst slovenske favne ugotavljamo, da je trenutno v

Sloveniji prisotnih 157 vrst kobilic, 5 smo črtali iz seznama, nekaj vrst smo ažurirali na trenutno

veljavna imena oz. njihov revidiran status pri nas. Gotovo se bo ta seznam sedaj veliko hitreje

ažuriral in dopolnjeval z novimi vrstami, saj gre za dolgo pričakovani pregled vrst kobilic pri nas.

The book of singing Orthoptera of Slovenia represents 97 species and two subspecies of

bush-crickets and grasshoppers in sound and photographs. The book is richly illustrated with

photographs as it is meant for the general public in order to make it easy to recognise species of

Othoptera, based on images as well as sound. In order to facilitate the identiﬁcation of species, each

species is represented by four photos and an oscillogram of the song. Each species is also

represented by basic data including the Slovenian and English common name, body size, audible-

hearing distance from where each species can be heard, data on species habitats and known

distribution in Slovenia. In addition to this book there is also included the ﬁrst up to date checklist of

currently known Orthoptera species in Slovenia. Several species in the book actually represent the

ﬁrst ﬁnds of species in Slovenia. Most species in the checklist are also indicated by their Slovenian

common names. Based on all this information the book brings much new information on Orthoptera

fauna in Slovenia. This will not only be useful just for the general public but also for the professionals

who will ﬁnd many novelties on Orthoptera fauna in Slovenia. In researching the checklist of

Slovenian Orthoptera, we found out that there are currently 157 species present in Slovenia, 5 were

deleted from the list, some species were updated to their current valid or revised names. Certainly,

this checklist will now be much faster updated and supplemented with new species, as it represents

a long-awaited review of Slovenian Orthoptera.

CMY

Pojoče kobilice Slovenije/Singing Orthoptera of Slovenia

Priročnik za določanje pojočih vrst kobilic po napevih in slikah/Manual for identiﬁcation

of singing Orthoptera based on songs and images

Besedilo/Text:

Zvoki/Songs:

Fotograﬁje/Photographs:

Lektura/Proofreaders:

Izdajatelj/Publisher: EGEA, Zavod za naravo/EGEA, Institution for Nature, Tacenska cesta

135A, SI-1113 Ljubljana, Slovenija

Naslovnica/Cover page: Akvarel, Blaž Šegula

Oblikovanje/Design: Emi international d.o.o., Lan Treven, Blaž Šegula,

Stanislav Gomboc

Tehnična priprava/Technical editing: Emi international d.o.o., Lan Treven,

Branka Foršček

Tisk/Print: Trajanus d.o.o., Kranj

Leto izdaje/Year of publication: 2014

Število izvodov/Copies: 250

CIP – Kataložni zapis o publikaciji

Narodna in univerzitetna knjižnica, Ljubljana

595.727/.728(497.4)

591.582:595.727/.728

GOMBOC, Stanislav

Pojoče kobilice Slovenije: priročnik za določanje pojočih vrst kobilic po napevih in slikah

= Singing Orthoptera of Slovenia: manual for identiﬁcation of singing Orthoptera based

on songs and images / [besedilo] Stanislav Gomboc, Blaž Šegula; [fotograﬁje Blaž

Šegula ... et al.]. - Ljubljana: Egea, zavod za naravo, 2014

Citiranje\Citation:

Book: Gomboc S., Šegula B. 2014: Pojoče kobilice Slovenije/Singing Orthoptera of

Slovenia. EGEA, Zavod za naravo, Ljubljana: 1-240.

CD: Šegula B., Gomboc S. 2014: Pojoče kobilice Slovenije/Singing Orthoptera of

Slovenia. Napevi/Songs. EGEA, Zavod za naravo, Ljubljana: CD.

ISBN 978-961-93692-0-3

1. Šegula, Blaž

274536960

Stanislav Gomboc, Blaž Šegula

Blaž Šegula

Blaž Šegula, Stanislav Gomboc,

Matjaž Bedjanič, Nina Šegula

Julia Ellis Burnet, Maja Žitnik, Gregor Torkar

UVOD

Knjiga in priložena zgoščenka obrav-

navata le pojoče vrste kobilic Sloveni-

je, ki smo jih uspeli posneti in fotogra-

ﬁrati pri večletnem proučevanju kobilic

na Slovenskem. V knjigi in na priloženi

zgoščenki je predstavljenih 99 taksonov

kobilic, od tega 97 vrst in 2 podvrsti.

Knjiga je slikovni dodatek k zgoščenki

s posnetimi napevi naših kobilic, ki se

oglašajo na nam slišen način. Za laž-

jo orientacijo po fotograﬁjah v knjigi in

zvokih, so vse vrste v knjigi in na zgo-

ščenki označene z enotno, zaporedno

številko taksona. Na ta način uporab-

nik lažje poišče napev na zgoščenki in

v knjigi pogleda, kateri vrsti pripada ter

obratno. Knjiga in zgoščenka skupaj sta

namenjeni ljubiteljem narave, ki želijo

spoznati slovenske vrste pojočih kobilic.

Sodobni mobilni telefoni domala že vsi

omogočajo predvajanje napevov kobi-

lic, ki so na zgoščenki v formatu MPEG

3, zato sta knjiga in mobilni telefon ti-

sto, kar potrebujemo za prepoznavanje

zvokov in kobilic v naravi. Zvoke prene-

semo na mobilni telefon, knjigo priroč-

nega formata pa spravimo v žep in že

smo pripravljeni na opazovanje kobilic.

Namen knjige in zgoščenke je spodbu-

diti dodatno terensko delo, saj na tem

področju manjka poljudna literatura, ki

je strokovno dovolj zanesljiva za lažjo

določitev sorodnih vrst. S knjigo želimo

tudi opozoriti na veliko biotsko raznovr-

stnost pojočih kobilic v Sloveniji.

V kompletu so predstavljene le vrste ko-

bilic, ki proizvajajo nam slišne napeve,

ostalih 60 vrst pa tu ni zajetih. Od sku-

paj 157 za Slovenijo znanih vrst kobilic

(objavljenih in neobjavljenih najdb), jih

INTRODUCTION

The book and the enclosed CD present

only the singing species of Orthoptera in

Slovenia, which we managed to record

and to photograph after studying them

for several years. They represent 99 taxa

of the Slovenian Orthoptera, namely 97

species and 2 subspecies. The book is a

photographic supplement to the CD with

sound recordings of Orthopteran son-

gs which are audible to our ears. For a

better orientation on species in the book

and on the CD, photos and sounds are

labeled with the same serial number.

Thus, the user can easily ﬁnd the song

on the CD and the corresponding spe-

cies in the book or vice versa. The book

and the CD together are intended for na-

ture lovers, who will ﬁnd it an easier way

to expand their knowledge on Slovenian

species of singing Orthoptera. Currently,

almost every mobile phone can be used

to play back songs of Orthoptera, which

are on the CD in MPEG 3 format; so that

the book and a cell phone are all we need

to identify sounds and Orthoptera in their

natural environment. With sounds trans-

ferred to a mobile phone and the book in

your pocket - you are ready to observe

the Orthoptera in the wild. In addition,

the book and the CD are geared towards

encouraging ﬁeldwork, as there is no po-

pular literature covering this domain, and

it is sufﬁciently professionally reliable to

facilitate the identiﬁcation of closely rela-

ted species. It should be noted, however,

that the book and the CD present only

species that produce songs which are

audible to the human ear; the remaining

60 species are not included in this set.

Out of a total of 157 known species of

je v knjigi predstavljenih malo manj kot

dve tretjini. Od pojočih, pri nas prisotnih

vrst, nam ni uspelo posneti nekaterih

redkih ali za Slovenijo vprašljivih vrst

kobilic. Nekaterih kljub starim naved-

bam nismo več našli, ali pa so njihove

navedbe za Slovenijo vprašljive, zaradi

napačne identiﬁkacije oz. takratne ravni

literature, ki ni razlikovala toliko vrst kot

jih poznamo danes. Nekatere, nekdaj

prisotne vrste pri nas, so zaradi spre-

memb rabe prostora, kot kaže, pri nas

izginile. Vrste, ki nam jih v zadnjih letih

ni uspelo najti so: Stenobothrus ﬁscheri

(Eversmann, 1848), Bucephaloptera bu-

cephala (Brunner von Wattenwyl, 1882),

Poecilimon thoracicus (Fieber, 1853),

Pachytrachis frater (Brunner von Watten-

wyl, 1882), Nemobius sylvestris (Bosc,

179 2), Arcyptera microptera (Fischer de

Waldheim, 1833), Arcyptera brevipennis

(Brunner von Wattenwyl, 1861), Chorthi-

ppus dichrous (Eversmann, 1859), Pse-

udochorthippus montanus (Charpenti-

er, 1825), Chorthippus pullus (Philippi,

1830), Myrmeleotettix maculatus (Thun-

berg, 1815), Celes variabilis (Pallas,

1771). Prav tako nam ni uspelo posneti

nekaterih vrst, ki smo jih pri nas sicer

potrdili, jih pa zaradi pomanjkanja ma-

teriala in časa doslej še nismo posne-

li: Euchorthippus pulvinatus (Fischer de

Waldheim, 1846), Andreiniimon nuptialis

(Karny, 1918), nekaj vrst iz rodu Isophya

in Platycleis s.l. Druge vrste kobilic se ne

oglašajo na način, da bi jih lahko zaznali

z našim sluhom ali drugimi čutili, čeprav

vemo, da se med seboj sporazumevajo

tudi z osciliranjem, ki ga proizvedejo z

zadkom, tipalkami, čeljustmi ali nogami,

navadno na zelo kratke razdalje, tudi z

drgnjenjem samčka ob samičko. Naša

čutila teh oscilacij ne zaznavajo, ker

Orthoptera in Slovenia (published and un-

published ﬁndings), a little less than two-

-thirds of them are portrayed here. From

the singing species living in Slovenia, we

have failed to capture some rare species;

in Slovenia the existence of some among

these is currently questionable. Despite

quotes in older sources, we could no lon-

ger ﬁnd some of them: this is either due

to misidentiﬁcation or to the level of old

sources that did not differentiate as many

species as are known today. Some spe-

cies formerly present in Slovenia seem

to have disappeared due to changes in

land use. Species that we could not ﬁnd

in recent years are: Stenobothrus ﬁscheri

(Eversmann, 1848), Bucephaloptera bu-

cephala (Brunner von Wattenwyl, 1882),

Poecilimon thoracicus (Fieber, 1853), Pa-

chytrachis frater (Brunner von Wattenwyl,

1882), Nemobius sylvestris (Bosc, 1792),

Arcyptera microptera (Fischer de Wald-

heim, 1833), Arcyptera brevipennis (Brun-

ner von Wattenwyl, 1861), Chorthippus

dichrous (Eversmann, 1859), Chorthi-

ppus montanus (Charpentier, 1825),

Chorthippus pullus (Philippi, 1830), Myr-

meleotettix maculatus (Thunberg, 1815),

Celes variabilis (Pallas, 1771). Owing to a

lack of material and time, we also have

not been successful in recording some

of the species, whose presence in Slove-

nia has recently been conﬁrmed. These

are: Euchorthippus pulvinatus (Fischer

de Waldheim, 1846), Andreiniimon nup-

tialis (Karny, 1918), some species from

Genus Isophya and Platycleis s.l. Other

Orthoptera species do not produce audi-

ble songs in a manner that could be per-

ceived by our senses - although we know

that they communicate with one another

in other ways, including short distance

oscillations produced by the abdomen,

Ime/Common name temna krivocerka Dark Bush-cricket

Velikost/Size 18-25 mm 18-25 mm

Slišna razdalja/Hearing distance 0,6 m, vabilni napev 0.6 m, Calling song

Habitat Grmovje, drevesa in višja robna

vegetacija, pogosto na robidi in

jesenu.

Shrubs, trees and higher edge

vegetation, mainly on Rubus and

Fraxinus.

Razširjenost/Distribution Od obale do Kraškega roba. From the seaside to the Kraški

rob (Karst edge).

Ensifera - dolgotipalčnice

Phaneropteridae - srparice

7. Barbitistes ocskayi Charpentier, 1850

Foto: Stanislav Gomboc

Foto: Blaž Šegula

Foto: Stanislav Gomboc

170

Ime/Common name dolgokrila škrebetulja Large Banded Grasshopper

Velikost/Size 22-42 mm 22-42 mm

Slišna razdalja/Hearing distance 15 m, vabilni napev 15 m, Calling song

Habitat Sredogorska do gorska bujna

travišča, svetel gozdni rob s

podrastjo.

Middle mountain to mountain

high grasslands, light forest edge

with undergrowth.

Razširjenost/Distribution Primorsk, Notranjska, južna

pobočja Gorenjske.

Primorska, Notranjska, southern

slopes of Gorenjska.

Caelifera - kratkotipalčnice

Acrididae - ščebetulje

74. Arcyptera fusca (Pallas, 1773)

Foto: Stanislav Gomboc

171

Foto: Stanislav Gomboc

Foto: Blaž Šegula

222

Seznam v Sloveniji najdenih vrst kobilic, s pregledom posnetih zvokov/

Checlist of Slovenian Orthoptera, with overwiev of recorded species

Spec.Nr. CD.TrackNr. Family, Species

Phaneropteridae - srparice

001 1Phaneroptera falcata (Poda, 1761) - navadna srparica

002 2Phaneroptera nana Fieber, 1853 - primorska srparica

003 3Tylopsis lilifolia (Fabricius, 1793) - lilijevka

004 4Acrometopa macropoda (Burmeister, 1838) - dolgonoga zelenolistka

005 5Isophya brevicauda Ramme, 1931 - kratkolegla nežnica

006

Isophya camptoxypha (Fieber, 1853) - vzhodna nežnica

syn. = Isophya brevipennis Brunner von Wattenwyl, 1878,

Few audible observations in Eastern Slovenia from Gergely Szövényi and

Anton Koschuh without collected specimens. Conﬁrmed in 2014 from

Stanislav Gomboc by sound recording and specimens. In publication.

007 Isophya kraussii Brunner von Wattenwyl, 1878 - kraussova nežnica

not = Isophya pyrenaea (Serville, 1839) as used in (Us 1992)

008 Isophya modesta (Frivaldskyi, 1867) - skromna nežnica

009 6Isophya modestior Brunner von Wattenwyl, 1882 - velika nežnica

010 7Barbitistes ocskayi Charpentier, 1850 - temna krivocerka

011 8Barbitistes serricauda (Fabricius, 1798) - celinska krivocerka

012 9Barbitistes yersini Brunner von Wattenwyl, 1878 - yersinova krivocerka

013 Andreiniimon nuptialis (Karny, 1918)

014 10 Leptophyes albovittata (Kollar, 1833) - beloproga steblikovka

015 11 Leptophyes boscii Fieber, 1853 - črnoroba steblikovka

016 12 Leptophyes laticauda (Frivaldsky, 1867) - dolgolegla steblikovka

017 13 Leptophyes punctatissima (Bosc, 1792) - pikčasta steblikovka

018 14 Poecilimon ampliatus Brunner von Wattenwyl, 1878 - cvetna lepotka

019 15 Poecilimon elegans Brunner von Wattenwyl, 1878 - postavna lepotka

020 16 Poecilimon gracilis (Fieber, 1853) - vitka lepotka

021 17 Poecilimon ornatus (Schmidt, 1849) - prešerna lepotka, južna lepotka

022 18 Poecilimon schmidtii (Fieber, 1853) - šmidtova lepotka

Poecilimon thoracicus (Fieber, 1853)

Not jet conﬁrmed for Slovenia even it is mentioned in Us (1992). From

Italy known from one old record from Gorizia (Ramme 1933b), at the

Slovenian border.

023 19 Polysarcus denticauda (Charpentier, 1825) - hribska srparica

234

Index

Acheta domesticus 148, 225

Acrida ungarica 226

Acrididae 162, 164, 166, 168, 170,

172, 174, 176, 178, 180, 182,

184, 186, 188, 190, 192, 194,

196, 198, 200, 202, 204, 206,

208, 210, 212, 214, 216, 218,

220, 226, 229, 230

Acrometopa macropoda 4, 30, 222

Acrotylus patruelis 226

Aeropedellus variegatus 227

Aiolopus strepens 226

Aiolopus thalassinus 226

Alpine Bush-cricket 138

Alpine Dark Bush-cricket 112

alpska ščebetulja 206, 228

Anacridium aegyptium 226

Andreiniimon nuptialis 9, 222, 229

Antaxius difformis 138, 224

Arachnocephalus vestitus 225

Arcyptera brevipennis 9, 227

Arcyptera fusca 170, 227

Arcyptera microptera 227

Barbitistes ocskayi 36, 222

Barbitistes serricauda 38, 222

Barbitistes yersini 40, 222

beloproga popadavka 84, 223

beloproga regljalka 184, 227

beloproga steblikovka 42, 222

beloproga ščebetulja 210, 228

beloroba ščebetulja 200, 228

Bicoloriana bicolor 5, 98, 223

Bicoloriana kuntzeni 100, 223

Black Cricket 150

Black-spotted Grasshopper 186

Blue-winged Grasshopper 164, 166

Blue-winged Locust 166

Bog Bush-cricket 102

Bordeaux Cricket 152

bordojski muren 152, 225

Bow-winged Grasshopper 210

bradavičarka 82, 223

Bradyporidae 140, 142, 144, 224

bramorji 15, 160, 225

bratska zalezovalka 224

bronasta trsnica 172, 227

Brown-spotted Bush-cricket 96

Bucephaloptera bucephala 9, 224

bukova kobilica 226

bukova listnica 226

bukova olivka 226

Caelifera 10, 15, 16, 162, 164, 166,

168, 170, 172, 174, 176, 178,

180, 182, 184, 186, 188, 190,

192, 194, 196, 198, 200, 202,

204, 206, 208, 210, 212, 214,

216, 218, 220, 230, 231, 233

Calliptamus italicus 226

Celes variabilis 9, 226

celinska grmovnica 124, 224

celinska krivocerka 38, 222

centralnoalpska regljalka 227

ceperojeva trnovratka 225

Chabrier’s Bush-cricket 124

Chorthippus albomarginatus 200, 228

Chorthippus alticola 206, 228, 231

Chorthippus apricarius 208, 228, 233

Chorthippus biguttulus 210, 228

Chorthippus bornhalmi 14, 212, 228

Chorthippus brunneus 214, 228

Chorthippus dichrous 9, 228

Chorthippus dorsatus 204, 228

Chorthippus eisentrauti 228, 231

Chorthippus mollis 216, 218, 228

Chorthippus mollis ignifer 218, 228

Chorthippus montanus 9, 227

Chorthippus oschei 13, 14, 202, 228