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Addition of Pumice Affects Physical Properties of Soil Used for Container Grown Plants

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Aeration of horticultural media is oft en reported to be a problem. The air filled porosity is generally considered as the quality determining factor for media and is generally determining the shape of the moisture characteristic curve. The objective of this study was to determine affects of different size and rate of pumice mixed with soil on the pore size distribution and bulk density of a mixture medium. Pumice of different size (<4 mm, <8 mm, and 4-8 mm) and rate on volumetric basis (10, 20, 30, 40, 50 % v/v) was mixed with two types of loamy soils passed through an 8 mm sieve. The moisture characteristic curve and bulk density of each mixture media was determined. From the moisture characteristic curve pore size distribution was obtained. Aeration, water conductivity, and water retention of media were evaluated. The results indicated that total porosity increased with increasing rate and size of pumice. The amount of pores important for drainage and aeration increased, but bulk density decreased significantly (p<0.01) with the increase in the rates of pumice mixed with soil. At 50 % pumice application, the increase in macropores (>100 μm diameter) were 98.2 % and 70.3 %, and the decreases in bulk density were 24.8 % and 21.0 % for soil I and II, respectively. While the mesopores (100-30 μm diameter), which are important for water conductivity, decreased significantly with pumice application in soil I, no significant changes were obtained for soil II. The amount of water held at 0.01 MPa – 0.10 MPa decreased significantly with 30 % or more pumice application in both soils. The amount of water held at 0.10 MPa or greater suctions decreased with pumice application. Significant changes in physical properties of soil-pumice mixture media were obtained for different sizes of pumice application.
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59
ORIGINAL SCIENTIFIC PAPER
Summary
Aeration of horticultural media is o en reported to be a problem.  e air
lled porosity is generally considered as the quality determining factor for
media and is generally determining the shape of the moisture characteristic
curve.  e objective of this study was to determine a ects of di erent size
and rate of pumice mixed with soil on the pore size distribution and bulk
density of a mixture medium. Pumice of di erent size (<4 mm, <8 mm, and
4-8 mm) and rate on volumetric basis (10, 20, 30, 40, 50 % v/v) was mixed
with two types of loamy soils passed through an 8 mm sieve.  e moisture
characteristic curve and bulk density of each mixture media was determined.
From the moisture characteristic curve pore size distribution was obtained.
Aeration, water conductivity, and water retention of media were evaluated.  e
results indicated that total porosity increased with increasing rate and size of
pumice.  e amount of pores important for drainage and aeration increased,
but bulk density decreased signi cantly (p<0.01) with the increase in the
rates of pumice mixed with soil. At 50 % pumice application, the increase in
macropores (>100 m diameter) were 98.2 % and 70.3 %, and the decreases in
bulk density were 24.8 % and 21.0 % for soil I and II, respectively. While the
mesopores (100-30 m diameter), which are important for water conductivity,
decreased signi cantly with pumice application in soil I, no signi cant changes
were obtained for soil II.  e amount of water held at 0.01 MPa – 0.10 MPa
decreased signi cantly with 30 % or more pumice application in both soils.  e
amount of water held at 0.10 MPa or greater suctions decreased with pumice
application. Signi cant changes in physical properties of soil-pumice mixture
media were obtained for di erent sizes of pumice application.
Key words
bulk density; pore size distribution; soil-pumice mixture media
Agriculturae Conspectus Scienti cus, Vol. 71 (2006) No. 2 (59-64)
Addition of Pumice A ects Physical
Properties of Soil Used for Container
Grown Plants
Ustun SAHIN ( )
Omer ANAPALI
Ataturk University, Faculty of Agriculture,
Department of Agricultural Structures and Irrigation
25240, Erzurum, Turkey
e-mail: ussahin@atauni.edu.tr
Received: May 18, 2006 | Accepted: June 21, 2006
Agric. conspec. sci. Vol. 71 (2006) No. 2
60 Ustun SAHIN, Omer ANAPALI
Introduction
Soil and some soilless media are used as plant-growth
media in horticulture. Liquid fertilizers are essential for
soilless plant-growth media. In soilless cultures, con-
tinuous controlling is required for soil reaction (pH),
salinity and plant nutrient equilibrium because of low
bu ering capacities of these media. On the other hand,
although plant-growth media with soil may have sev-
eral disadvantages due to not providing desirable soil at
all times, heavy texture, and need for sterilization, plant
nutrition in soil media is easier because the nutrients are
retained by soil. In addition, minor element de ciencies
are rare because the elements are on the soil exchange
sites (Holcomb 2002).
In a medium with soil, the provision of aeration and
the improvement of drainage conditions are required.
For this reason, pumice is a good amendment for aera-
tion and drainage. Pumice mixed with soil in speci c
amounts improves soil air and water conductivity, and
reduces negative e ects of crusting, cracking,  ooding,
and shrink swelling. It is also used for long periods be-
cause of its stable physical and chemical properties (Gur
et al. 1997). In general, pumice could be provided easily
since there are many pumice deposits distributed around
the world (Tuncer 1997). Pumice used only a er sieving
has a high water retention capacity, and very low bulk
density value compared to soil (Sahin et al. 2004).
e most important physical properties of a medium
for suitability are good aeration and drainage, optimum
water retention, high water conductivity and low bulk
density (Cabrera 2003). Within the physical proper-
ties, the air-water ratio is the most important (Bruckner
1997; Caron and Nkongolo 1999).  e air-water ratio can
partly be determined by the granulometry and poros-
ity (Orozco et al. 1997).  e relationship between water
energy status and water content of the medium is a re ec-
tion of the pore size distribution of the medium (Milks
et al. 1989). Pore sizes have traditionally been divided
into macropores, mesopores, micropores and ultrami-
cropores (Sahin et al. 2002).  e macropores (>100 m
diameter) supply drainage and aeration, the mesopores
(100-30 m diameter) supply water conductivity, and
the micropores (30-3 m diameter) supply water reten-
tion (Gemalmaz 1993).  e water retained in ultrami-
cropores (<3 m diameter) is unavailable for plant use
(Drzal et al. 1999).
The objective of this study was to determine the
effects of different size and rates of pumice mixed
with soil on the pore size distribution and bulk
density of the mixture medium.
Materials and methods
Two soils formed under di erent climates with 450
mm (Erzurum-Turkey) and 2200 mm (Rize-Turkey)
annual rainfalls were used. Some physical and chemical
properties of these soils and pumice are given in Table
1.  e soils used in this study sampled from Ap hori-
zon (0-20 cm depth) and origin of pumice was Ercis,
Van-Turke y.
Pumice is a very light, porous igneous rock that is
formed during volcanic eruptions. It is made up of very
tiny crystals, since they cool so quickly above ground.
e texture of pumice is rough and has many hollows
and cavities. Pumice has been used to a large extent as a
plant growing media and it lightens the soil, makes tillage
easier, improves soil aeration and holds water. Pumice
mixed with soil in speci c amounts improves soil’s air
and water conductivity, and reduces negative e ects of
crusting, cracking,  ooding, and shrink-swelling. It
can also be used for a long periods because of its stable
Properties Soil I Soil II Pumice
Soil taxonomy Ustent Aquoll -
Dominant clay type Montmorillonite Kaolinite -
Clay (%) 23.7 24.6 -
Silt (%) 39.8 27.7 -
Sand (%) 36.5 47.7 -
Bulk density (g cm-3) 1.05 1.00 0.65 (for < 4 mm)
0.63 (for < 8 mm)
0.40 (for 4-8 mm)
Particle density (g cm-3) 2.64 2.61 2.24
Wet Aggregate stability (%) 28.5 50.7 -
pH 8.2 6.7 8.4
Electrical cond. (dS m-1) 0.93 0.19 0.11
CEC (cmol(+) kg-1) 33.6 26.0 6.9
Carbonates (%) 2.7 0.1 0.8
Organic matter (%) 2.9 3.8 -
Table 1.
Some physical and chemical properties of soils and pumice used
Agric. conspec. sci. Vol. 71 (2006) No. 2
61
Addition of Pumice Affects Physical Properties of Soil Used for Container Grown Plants
physical and chemical properties and it can be provided
easily since there are many pumice deposits distributed
around the world. Pumice used only a er sieving has a
high water retention capacity, and very low bulk density
value compared to soil (Sahin et al. 2005).
Soil samples were passed through a sieve with open-
ings of 8 mm before mixing them with pumice.  e rates
of size fractions (<2 mm, 2-4 mm, and 4-8 mm) were
58.6, 26.1 and 15.3 % for Soil I and 44.2, 29.7, and 26.1
for Soil II, respectively, following sieving.
Soil and pumice mixtures were prepared as 9:1, 8:2,
7:3, 6:4, and 5:5 ratio on volumetric basis, resulting in
10 % (M1), 20 % (M2), 30 % (M3), 40 % (M4), and 50
% (M5) of pumice in the mixture media. Similar mix-
ture media were prepared for three di erent sizes of
pumice as S1: <4 mm (50 % 2-4 mm, 25 % 1-2 mm, and
25 % < 1mm), S2: <8 mm (33.3 % 4-8 mm, 33.3 % 2-4
mm, 16.7 % 1-2 mm, and 16.7 % <1 mm), S3: 4-8 mm.
Sample with no pumice mixture was used as the con-
trol medium (C).
e soil water characteristic curve (pF curve) was
determined using pressures plates (Klute 1986), and
was used as the basis for the calculation of the pore
size distribution. Water held at 0.001 MPa, 0.01 MPa,
0.03 MPa, 0.10 Mpa and 1.5 MPa was obtained when
water output stopped for a given suction.  e samples
for pF curves were packed by dropping the cylinders.
Porosity was estimated by calculation (Danielson and
Sutherland, 1986). Bulk density was determined by the
cylinder method (Blake and Hartge 1986a) on samples
packed by dropping the sample cylinders from a height
of 10 cm for 20 times.
e percentage of water-stable aggregates was deter-
mined by a wet sieving procedure (Kemper and Rosenau
1986). Particle density was determined by the pycnometer
method (Blake and Hartge 1986b), pH by a pH-meter in
saturation extract (Mc Lean 1982), Electric conductiv-
ity (EC) by an EC-meter in saturation extract (Rhoades
1982a), cation exchange capacity (CEC) by the sodium
acetate method (Rhoades 1982b), carbonates by the cal-
cimeter method (Nelson 1982) and organic matter by the
wet combustion method (Nelson and Sommers 1982).
e experimental design was factorial (soils (2),
mixture rates (5), and pumice sizes (3)) with three rep-
lications. Analysis of variance (ANOVA) for the data of
macropores, mesopores, micropores, ultramicropores
and bulk density was performed, and Duncans multiple
range tests were used for important treatments.
Results and Discussion
Total porosity increased with increasing rates and
sizes of pumice in both soils. e increasing rates in
total porosity were higher for Soil I than Soil II.  e ini-
tial total porosity was 60.4 for Soil I and 62.1 for Soil II.
Since both soils have similar texture, it may be that the
di erences in total porosity may be related to the degree
of soil structure.  erefore, pumice application to Soil
I gave better results in improving total porosity which
reached up to 12.2 % (M5) and 10.0 % (S3) as compared
to the initial porosity value.
e amount of macropores supplies aeration and
drainage increased signi cantly (p<0.01) with increas-
ing rate of pumice in the medium. Pumice mixture to
soil medium with a rate of 50 % increased the amount
of macropores by about 98.2 % in soil I and 70.3 %
in soil II compared with the control medium (Fig. 1).
Similar results were obtained in all pumice size groups.
Maximum increase was obtained for S2 treatment in
soil I, and S1 treatment in soil II (Fig. 1).  e macro-
pores increased with increasing rates of pumice in the
medium in all pumice sizes.  e maximum macropo-
res values were obtained in S2 treatment (Fig. 2). More
LSD =1.24
0.01
0
5
10
15
20
25
30
35
40
C 10 20 30 40 50 <4 <8 4-8
Soil I
Soil II
Rate of pumice (%v/v) Pumice size (mm)
(over means of S1, S2, and S3) (over means of M1,
M2, M3, M4, and M5)
Macroporosity (%)
LSD =0.96
0.01
1
0
5
10
15
20
25
30
35
40
10 20 30 40 50
<4mm
Rate of pumice (%v/v)
Macroporosity (%)
LSD =1.51
0.01
<8mm
4-8 mm
1
Figure 1.
Changes in macropores with pumice size and rate
Figure 2.
Changes in macropores with pumice rate in the different
pumice size
Agric. conspec. sci. Vol. 71 (2006) No. 2
62 Ustun SAHIN, Omer ANAPALI
Figure 7.
Changes in bulk density with pumice size and rate
Figure 8.
Changes in bulk density with pumice rate in the different
pumice size
Figure 3.
Changes in mesopores with pumice size and rate
Figure 4.
Changes in micropores with pumice size and rate
Figure 5.
Changes in ultramicropores with pumice size and rate
Figure 6.
Changes in ultramicropores with pumice rates in the
different pumice size
0
0,2
0,4
0,6
0,8
1
1,2
10 20 30 40 50
Bulk density(gm )
–3
<8mm
LSD =0.02
0.01
<4mm 4-8 mm
Rate of pumice (%v/v)
0
2
4
6
8
10
12
C10 20 30 40 50 <4 <8 4-8
Microporosity (%)
Rate of pumice (%v/v) Pumice size (mm)
(over means of S1, S2, and S3) (over means of M1,
M2, M3, M4, and M5)
Soil I
Soil II
LSD =1.21
0.01 LSD =0.94
0.01
1
0
1
2
3
4
5
6
7
8
C10 20 30 40 50 <4 <8 4-8
Mesoporosity (%)
Rate of pumice (%v/v) Pumice size (mm)
(over means of S1, S2, and S3) (over means of M1,
M2, M3, M4, and M5)
Soil I
Soil II
LSD =0.86
0.01 LSD =0.67
0.01
1
0
5
10
15
20
25
30
10 20 30 40 50
Rate of pumice (%v/v)
Ultramicroporosity (%)
<8mm
LSD =0.82
0.01
<4mm 4-8 mm
0
5
10
15
20
25
30
C 10 20 30 40 50 <4 <8 4-8
Ultramicroporosity
(
%
)
Rate of pumice (%v/v) Pumice size (mm)
(over means of S1, S2, and S3) (over means of M1,
M2, M3, M4, and M5)
Soil I
Soil IIns ns
1
1
C10 20 30 40 50 <4 <8 4-8
Rate of pumice (%v/v) Pumice size (mm)
(over means of S1, S2, and S3) (over means of M1,
M2, M3, M4, and M5)
0,5
0,6
0,7
0,8
0,9
1
1,1
Bulk density (g m )
–3
Soil I
Soil II
LSD =0.02
0.01 LSD =0.01
0.01
suitable air volume conditions were reached; due to the
macropores they were greater than 20 % in all treat-
ments (Verdonck 1984).
Pumice caused signi cant reduction of the amount
of mesopores, which e ect water conductivity, in soil
I, but no signi cant changes were determined for soil
II (Fig.3).  is may be related to di erences in soil ag-
gregation (Table 1).  e e ects of mixed ratio with all
pumice sizes on the amount of mesopores were not sig-
ni cant.
Agric. conspec. sci. Vol. 71 (2006) No. 2
63
Addition of Pumice Affects Physical Properties of Soil Used for Container Grown Plants
e amounts of micropores, which a ect water re-
tention, also decreased signi cantly (p<0.01) by higher
rates of pumice application. Signi cant decreases of
the amount of micropores occurred with M3-M5 treat-
ments in soil I and M4-M5 treatments in soil II. In soil
I, a small increase (8.1 %) in the amount of micropores
with M1 treatment was obtained, but it was statistically
insigni cant (Fig. 4). It means that pumice application
to soil medium in small rates has no signi cant e ect on
the amount of water retention. On the other hand, dif-
ferent sizes of pumice caused reduction in the amount of
micropores.  e most signi cant reduction was obtained
with S2 treatment in soil I (25.6 %) and S1 and S2 treat-
ments in soil II (13.2 and 14.3 %, respectively) (Fig. 4).
e e ects of mixed ratio with all pumice sizes on the
amount of micropores were not signi cant.
Pumice application also reduced the amount of ultra-
micropores compared to the controls (Fig. 5, and 6), but
this may not be important because water in these pores
is not available for plant use (Drzal et al. 1999).
Bulk density of the medium decreased signi cantly
(p<0.01) with the rate of pumice applied to soil medium
in both soils. In soil II, no signi cant e ect of pumice
size on bulk density was obtained, but it became more
important with pumice size in soil I (Fig.7). e bulk
density decreased signi cantly (p<0.01) with increasing
rates of pumice in the medium in all pumice sizes (Fig. 8).
In greenhouse culture, a lower bulk density is desirable
due to easier handling and less root losses during pick
up and transportation (Sahin et al. 2004).
e results of this study indicated that the optimum
air- lled porosity conditions were reached by pumice ap-
plications. While the amount of macropores increased
signi cantly (p<0.01) with increasing rate of pumice
in the medium, the amount of mesopores in one of the
soils decreased.  e amounts of micropores decreased
signi cantly (p<0.01) by higher rates of pumice applica-
tion. Pumice application reduced the amount of ultrami-
cropores compared to the controls. Bulk density of the
medium decreased signi cantly (p<0.01) with increas-
ing rate of pumice applied to soil medium. It was clearly
determined that pumice may be e ectively used in spe-
ci c amounts for improving aeration and bulk density
conditions of poorly structured soils.
Conclusions
Pumice application to soil increased the amount of
macropores and decreased the bulk density. It was clearly
determined that pumice may be e ectively used in spe-
ci c amounts for improving aeration and bulk density
conditions of poorly structured soils.
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Proceedings of the I. Isparta Pumice Symposium, pp -
Verdonc O. (). New developments in the use of graded
perlite in horticultural substrates. Acta Hort : -
acs71_09
... Different lowercases between freezing-thawing cycles indicate difference at the level of 0.05 Leuther and Schlüter (2021) expressed that frost resulted in a systematic reduction in pore sizes and macro-pore connectivity with re-consolidation of soil clods. The macro-pores supply drainage and aeration, while the meso-pores serve water conductivity and temporary water retention, and micro-pores aid permanent water retention (Sahin and Anapali, 2006). Mangalassery et al. (2013) expressed that soil aggregates were often considered a good indicator and component of soil structure as the basic factor influencing soil pore characteristics. ...
... Considering the significant negative correlation between macro-pores and field capacity (Fig. 3), an increase in the hydraulic conductivity could have incurred with the impact of a possible increase in meso-pores. Moreover, the increase in macro-pores which provides drainage and aeration was found valuable for contributing to the increase in soil aeration (Sahin and Anapali, 2006). ...
... För att åtgärda problemet med dräneringen tillsätter samma plantskola grovsand till substratet. Alternativt kan också andra grova mineraliska material, såsom perlit och pimpsten, (Sahin & Anapali 2006;Kuepper 2010) eller organiska substratkomponenter, som t.ex. kokos-eller träflis, användas för detta ändamål (Carlile et al. 2019). ...
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Peatlands cover ~3% of the earth´s land surface and have since the last ice age functioned as an important carbon sink, taking up carbon dioxide (CO2) from the atmosphere. About 10% of the global peatlands have been drained, mainly for forestry and agriculture, and to some extent for peat extraction. Drained peatlands not only lose their function as carbon sinks, but instead give rise to an emission of almost 2 billion tonnes of CO2 per year, which corresponds to ~5% of the human-caused CO2 emissions. Because rewetting of peatlands can reverse this process, many countries have launched environmental programs aimed at protecting and restoring peatlands. In the United Kingdom (UK), the political discussion about peat has led to the decision that the use of peat has to be phased out within the entire horticultural industry by 2030. Although the implementation of this goal is voluntary, there are several nurseries in the UK that already produce their container-grown plants completely without peat. As part of this study, these nurseries were contacted with a survey to take part in their experiences with peat-free cultivation systems. The main objectives of this were to investigate which growing media these peat-free nurseries use to replace peat and to identify the benefits and challenges that arise in peat-free nursery production. The results show that coir, composted bark, wood fibre, green compost and composted wool are the substrate components most used in peat-free growing media for container nursery production in the UK. The adaptation of the cultivation systems to peat-free growing media based on these components is perceived by the majority as manageable. However, the results also showed that the optimisation of plant nutrition and irrigation routines can be perceived as challenging and thus it is recommended that these aspects should be controlled carefully. In addition, the partly limited availability and varying quality of peat-free growing media is perceived as problematic. As main advantage the nurseries experience that the label “peat-free” is good for marketing and can thus be used to attract both, private and corporate customers, who want to buy sustainably produced plants. Overall, the results of this study show that it is possible to produce a wide range of different plant species completely without peat, both on small and large scale, and that a peat-free nursery can be a profitable business model.
... Treatment (T2) soil + coir pith + vermicompost + pumice stone (1:1:1:0.5) can be used to establish the above ornamental plants in plastic pot container of vertical garden systems fabricated in iron frames and found as best growing media composition [10][11]. ...
Article
An experiment was conducted on the topic “A study on vertical gardens in urban areas’’ during rainy season 2021-22 at the Research Field of Department of Horticulture, Sam Higginbottom University of Agriculture, Technology and Sciences, Prayagraj. The observations were recorded on various growth parameter. Vertical gardens are becoming a common component in contemporary garden designs at urban living spaces because of shrinking land spaces. Successful growing of plants in vertical garden systems depends up on growing container, plant chosen, growing media etc. Hence a study was carried out in the Department of Horticulture, Sam Higginbottom university of Agriculture, Technology and Sciences, Prayagraj, India with the objective to study about suitable media composition along with vermicompost, perlite, coir pith, pumice stone and peat moss on growth and performance of ornamentals plant for establishment of vertical garden and to study the performance of ornamental plants viz., Duranta erecta, Hemelia patens, Iresine lindeni, Tabernamontana divaricata, Hemelia patens, in plastic pot for establishment of vertical garden. The experiment was laid out in Completely Randomised Design in plastic pot with five treatment combination of various growing media mixtures comprising soil, river sand as basic components in combination with organic manures (vermicompost, coir pith, perlite, peat moss, pumice stone). The plant growth parameters and ornamental morphological characters observed. Among the five ornamentals plants used Golden duranta and Mini Chandani performed better as ornamental plants in vertical garden system with the growing media of soil:cocopeat:vermicompost:perlite (1:1:1:0.5).
... e other soil ameliorants were pumice and mycorrhizae. Pumice and mycorrhizae are good soil ameliorants, suitable for comparison [30,31]. e pumice use is also based on the abundant pumice availability in Indonesia due to many compositetype volcanoes. ...
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Vertical gardens are becoming a common component in contemporary garden designs at urban living space because of shrinking land spaces. Successful growing of plants in vertical garden systems depends up on growing container, plants chosen, growing media, etc. Hence a study was carried out in the Department of Horticulture, Annamalai University, during the year 2013, with the objectives to study the influence of Coir pith, Stockosorb and Geohumus as components of growing media along with FYM, Vermicompost and Leaf mould compost on growth and performance of ornamental plants for establishment of vertical garden and to study the performance of ornamental plants Viz., Philodendron erubescens Cv. ‘Gold’, Chlorophytum comosum Cv. ‘Variegatum’ and Polyscias fruticosa plants in wooden containers for establishment of vertical garden. The experiment was laid out in Completely Randomized Design in wooden containers, with ten treatment combinations of various growing media mixtures comprising red soil, river sand as basic components in combination with organic manures (FYM, vermicompost, leaf mould compost, coir pith) and hydrogels (Stocksorb and Geohumus). The plant growth characters and ornamental value index were observed. Among the three ornamental plants used, Polyscias fruticosa and Philodendron erubescens are performed better as ornamental plants in vertical garden system with the growing media of Red soil : River sand : Vermicompost @ 1:1:1/2 + Stockosorb(25g).
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Moisture retention data were collected for five porous materials: soil, phenolic foam, and three combinations of commonly used media components. Two mathematical functions were evaluated for their ability to describe the water content–soil moisture relationship. A cubic polynomial function with linear parameters previously used on container media was compared to a closed-form nonlinear parameter model developed to describe water conductivity in mineral soils. In most tests for precision, adequacy, accuracy, and validation, the nonlinear function was superior to the simpler power series. The nonlinear function provides an excellent tool for describing the water content for media with widely varying physical properties.
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Aeration in soilless mixes is often reported to be a problem. After the partial saturation and the complete drainage of potted media, a very small perched water table occurs at the bottom of the pot, resulting in media equilibrated at very high water potentials. The air-filled porosity of soilless mixes may be very low at these high water potentials, despite a very high total porosity. Air-filled porosity is therefore used as a standard index to guide substrate manufacturing, and is generally measured on disturbed media packed into cylinders and equilibrated at an arbitrary potential. Simulation studies have shown the importance of characterizing air-filled porosity at more than one water potential since air-filled porosity is dependent on container geometry for a given substrate. Latest developments also suggest that the air exchange properties within the substrate be considered to obtain more reliable indexes of aeration. Indeed, in three recent studies in which the substrate physical properties were varied, air exchange properties (gas relative diffusivity) were more closely correlated to plant growth than was the air-filled porosity. Recent studies have also focused on the in situ characterization of artificial mixes, providing new ways to measure gas relative diffusivity, air-filled porosity and water desorption curves directly in the pot, with or without actively growing plants.