(PDF) Towards incorporating anamorphic fungi in a natural classification

Mycosphere Doi 10.5943/mycosphere/3/2/5

157

Towards incorporating anamorphic fungi in a natural classification – checklist

and notes for 2011

Wijayawardene DNN1,2, McKenzie EHC3 and Hyde KD1,2

1Institute of Excellence in Fungal Research, Mae Fah Luang University, Chiang Rai 57100, Thailand

2School of Science, Mae Fah Luang University, Chiang Rai 57100, Thailand

3Manaaki Whenua Landcare Research, Private Bag 92170, Auckland, New Zealand

Wijayawardene DNN, McKenzie EHC, Hyde KD 2012 – Towards incorporating anamorphic fungi

in a natural classification – checklist and notes for 2011. Mycosphere 3(2), 157-228, Doi 10.5943

/mycosphere/3/2/5

A complilation of anamorphic names for both Ascomycota and Basidiomycota is provided which

comprises 2895 genera. The genera are listed against a backbone of teleomorphic relationships

where known. The study reveals that 73 genera and 95 anamorph-like genera are linked to

teleomorphic genera names, 447 genera (three anamorph-like genera) are linked to teleomorph

families, orders or classes, while for more than 1592 (55%) genera no teleomorph link is known.

The links are based on the literature and often because fungi were found in association, but have not

been proven by molecular data. Many are based on links with species other than the generic type

and thus must be considered questionable for the genus. A considerable effort is needed to establish

whether these links are correct.

Key words – Asexual fungi – life cycle – sexual fungi – taxonomy

Article Information

Received 24 March 2012

Accepted 29 March 2012

Published online 10 April 2012

Corresponding author: KD Hyde – e-mail – kdhyde3@gmail.com

Introduction

The purpose of the present paper is to

collate the knowledge of asexual fungi in

relationship to their sexual states. It attempts to

classify the asexual genera in a natural

biological framework for the Ascomycota

based on the Outline of Ascomycota 2009

(Lumbsch & Huhndorf 2010) and for the

Basidiomycota based on Kirk et al. (2008). The

present paper has used data from the previous

outline of Hyde et al. (2010) as well as

literature published in 2011 and updated with

opinions in Seifert et al. (2011).

There is now a mandate in the syste-

matics community to place one name on a

holomorphic fungus species as opposed to the

previous system (especially in Ascomycota) of

naming both sexual and asexual morphs

(Samuels et al. 2009, Hawksworth 2011;

Hawksworth et al. 2011, Taylor 2011; Wingf-

ield et al. 2011). Also, as more and more

strains of asexual genera are sequenced

(Shenoy et al. 2010, Põldmaa 2011, Reblova &

Seifert 2011), these genera will be classified in

families, orders and classes of Ascomycota and

Basidiomycota. This will result in more and

more asexual states being described and illustr-

ated without being given formal names and

therefore it is important to have a forum where

asexual data can be brought to the attention of

researchers. It is hoped that this compilation

will facilitate the move towards one fungus one

name (Shenoy et al. 2007, 2010, Hawksworth

et al. 2011). By placing all asexual genera

against the backbone classification of the sexu-

al genera it will be possible to establish, 1)

Mycosphere Doi 10.5943/mycosphere/3/2/5

158

what are the sexual states of various species,

genera,families or orders, 2) which genera have

no information, 3) which are the older or more

commonly used names for each genus, and 4)

illustrate the redundancy of the dual classif-

ication system.

This compilation also provides a single

place where asexual connections can be access-

ed and critically scrutinized so that a stable and

reliable system can be developed based on pub-

lished data. We do not claim that this compi-

lation is complete or definitive; however, it

provides a forum for displaying data and can

be added to or changed as more data becomes

available or errors are discovered. In future

issues we will explore various connections and

provide more data.

In the long term it will be desirable if

Myconet (http://www.fieldmuseum.org/myco-

net) is merged with this document, however,

this should occur at some time in the future,

when more data is available.

Material and methods

The overall scheme for the classification

of the sexual fungi follows the Outline of

Ascomycota 2009 (Lumbsch & Huhndorf

2010), with additional data on anamorphic

Basidiomycota from Kirk et al. (2008). This

scheme is chosen as the schemes in the

Dictionary of the Fungi, Index Fungorum, and

MycoBank differ to a greater or lesser extent

and it was felt wise to follow what the authors

consider is an admirable attempt to keep

abreast of the current literarure as is routinely

carried out in the Outline of Ascomycota. This

compilation also annotates changes made in the

form of notes on asexual fungi (marked with an

asterisk after the entry, with notes in the later

“Notes” section) so that changes can easily be

followed; this is not always true of the

available databases.

Asexual genera or ‘-like’ genera are

listed in bold under the appropriate fungal

classification and the link to a teleomorph

genus given where found. Dubious genera are

excluded but can be added if they are later

found to be good genera. Fossil fungi are also

excluded as it would be impossible with

present techniques available to provide a na-

tural classification for them. A question mark

before the entry means that its placement in

that family or order is not fully established and

generally follows Lumbsch & Huhndorf

(2010). A question mark before anamorph

means it is not clear that this is the anamorph

of this genus, although there are many cases

without a question mark where the link has not

equivocally been proven. An asterisk following

the names indicates an entry in the “Notes”

section.

In many genus records in Index Fungo-

rum the species is linked to more than one

anamorphic genus. Such records can be located

by searching the genus throughout the list for

repeat entries. The synonyms of genera are not

listed as they can be easily found in Index

Fungorum.

We encourage colleagues to inform us of

any missing entries, new data or misunder-

standings, which will be updated in future

compilations. We would also invite experts of

specific groups to contribute to future compi-

lations on their specific group(s) of interest and

encourage reviews or illustrated accounts of

families, orders or other groups relating to

sexual and asexual links. Reviews of asexual

genera with sections or series, such as

Aspergillus, Penicillium and Verticillium, the

powdery mildews, many of the yeasts and

yeast-like genera and the asexual smuts and

rusts are particularly needed. Similarly the

lichenized ascomycete asexual states are poorly

documented.

As there is mandate towards using one

name for both the sexual and asexual states of a

fungus we also encourage authors to submit

articles or opinions concerning the use of either

name. This might be in the form of a general

critic, arguments for or against using one name,

or an article that specifically addresses an

individual group or genus.

In the present outline we have added

(H) = hyphomycete, (C) = coelomycete, (Y) =

yeast and (R) = Rust against entries where

known. Although this grouping is artificial we

do this to facilitate finding data on either as the

groups are generally dealt with in different text.

For example, data for the hyphomycetes have

recently been complied and updated by Seifert

et al. (2011) and the new data for hyphomycet-

es included here have been extracted from this

tome. Data for coelomycetes were extracted

from Sutton (1980), Nag Raj (1993), http://

Mycosphere Doi 10.5943/mycosphere/3/2/5

159

www. Cybertr uffle.org.uk/cybernome/eng/ind-

ex.htm (Accesion date 18.03.2011) and http://-

www.landcarer esearch .co.nz/html /data asp?

ID = &NAMEKey = 26885 (Accesion date

6.12.2011).

Index to Ascomycota

Phylum ASCOMYCOTA Caval-Sm.

Subphylum TAPHRINOMYCOTINA O.E.

Erikss. & Winka

Class Taphrinomycetes O.E. Erikss. & Winka

Taphrinales Gäum. & C.W. Dodge

Protomycetaceae Gray

Saitoella Goto, Sugiy., Hamam. & Komag.

1987 (Y)

Taphrinaceae Gäum.

Lalaria R.T. Moore 1990, anamorphic

Taphrina Fr. 1815 (Y)

Subphylum SACHAROMYCOTINA O.E.

Erikss. & Winka

Class Saccharomycetes O.E. Erikss. & Winka

Saccharomycetales Kudrjanzev

Dipodascaceae Engl. & E. Gilg

Blastobotrys Klopotek 1967, anamorphic

Trichomonascus H.S. Jacks. 1948 (H)

?Candida Berkhout 1923, anamorphic

Sporopachydermia Rodr. Mir. 1978 (Y)

Geotrichum Link 1809, anamorphic

Galactomyces Redhead & Malloch

1977 and Dipodascus Lagerh. 1892 (H)

Geotrichum-like, anamorphic Basidioas-

cus Matsush. 2003 (H)

Saprochaete Coker & Shanor 1939, ana-

morphic Magnusiomyces Zender 1926

(H)

Endomycetaceae J. Schröt.

Fusidium Link 1809, possibly anamorphic

Ascocephalophora K. Matsush. &

Matsush. 1995 (H)

Lipomycetaceae E.K. Novák & Zsolt

Myxozyma Van der Walt, Weijman & Arx

1981 (Y)

Metschnikowiaceae T. Kamienski

Candida Berkhout 1923, anamorphic

Clavispora Rodr. Mir. 1979 and

Metschnikowia T. Kamienski 1899 (Y)

Pichiaceae Zender

Brettanomyces N.H. Claussen ex Custers

1940, anamorphic Dekkera Van der

Walt 1964 (Y)

Eeniella M.T. Sm., Bat. Vegte & Scheffers

1981, anamorphic Dekkera Van der

Walt (Y)

Enantiothamnus Pinoy 1911 (Y)

Hyphopichia Arx & Van der Walt 1976

(Y)

Saccharomycetaceae G. Winter

?Candida Berkhout 1923, anamorphic

Ogataea Y. Yamada, K. Maeda &

Mikata 1994, Lodderomyces Van der

Walt 1966 and Spathaspora N.H.

Nguyen, S.O. Suh & M. Blackw. 2006

(Y)

Saccharomycodaceae Kudrjanzev

Candida Berkhout 1923, anamorphic

Lodderomyces Van der Walt 1966 and

Spathaspora N.H. Nguyen, S.O. Suh &

M. Blackw. 2006 (Y)

Kloeckera Janke 1923, anamorphic Han-

seniaspora Zikes 1911 (Y)

Trichomonascaceae Kurtzman & Robnett

?Candida Berkhout 1923, anamorphic

Zygoascus M.T. Sm. 1986 (Y)

Saccharomycetales, genera incertae sedis

Aciculoconidium D.S. King & S.C. Jong

1976 (Y)

Botryozyma Shann & M.T. Sm. 1992,

anamorphic Ascobotryozyma J. Kerri-

gan, M.T. Sm. & J.D. Rogers 2001 (Y)

Candida Berkhout 1923, anamorphic

Starmerella C.A. Rosa & Lachance

1998 (Y)

Cicadomyces Šulc 1911 (Y)

Macrorhabdus Tomasz., Logan, Snowden,

Kurtzman & Phalen 2003 (Y)

Oosporidium Stautz 1931 (Y)

Pseudomycoderma H. Will 1916 (Y)

Schizoblastosporion Cif. 1930 (Y)

Trigonopsis Schachner 1929 (Y)

Saccharomycetes, genera incertae sedis

Selenotila Lagerh. 1892 (Y)

Mycosphere Doi 10.5943/mycosphere/3/2/5

160

Selenozyma Yarrow 1977 (Y)

Subphylum PEZIZOMYCOTINA O.E.

Erikss. & Winka

Class Arthoniomycetes O.E. Erikss. & Winka

Arthoniales Henssen ex D. Hawksw. & O.E.

Erikss.

Arthoniaceae Reichenb. ex Reichenb.

Helicobolomyces Matzer 1995, anamor-

phic Arthonia Ach. 1806 (C)

?Septocyta Petr. 1927, anamorphic Artho-

nia Ach. 1806 (C)

Subhysteropycnis Wedin & Hafellner

1998, anamorphic Arthonia Ach. 1806

(C)

Roccellaceae Chevall.

Sporhaplus H.B.P. Upadhyay 1964, ana-

morphic Mazosia A. Massal 1854 (C)

Class Dothideomycetes sensu O.E. Erikss. &

Winka

Subclass Dothideomycetidae P.M. Kirk, P.F.

Cannon, J.C. David & J.A. Stalpers ex C.L.

Schoch, Spatafora, Crous & Shoemaker

Capnodiales Woron.

Antennulariellaceae Woron.

Capnodendron S. Hughes 1976, anamor-

phic Antennulariella Woron. 1915 (H)

Capnofrasera S. Hughes 2003 (H)

Heteroconium Petr. 1949, anamorphic

Antennulariella Woron. 1915 (H)

Capnodiaceae (Sacc.) Höhn. ex Theiss.

Acanthorus Bat. & Cavalc. 1967 (C)

?Antennariella Bat. & Cif. 1963 (C)*

Apiosporium Kunze 1817 (C)

Conidiocarpus Woron. 1927, anamorphic

Phragmocapnias Theiss. & Syd. 1918

and Scorias Fr. 1825 (C)*

Fumagospora G. Arnaud 1911, anam-

orphic Capnodium Mont. 1848 (C)

?Fumiglobus D.R. Reynolds & G.S.

Gilbert 2006 (C)*

Microxiphium (Harv. ex Berk. & Desm.)

Thüm. 1879 (as Microxyphium) (H)*

?Mycogelidium W.Y. Zhuang 2007 (C)

Phaeoxyphiella Bat. & Cif. 1963, ana-

morphic Capnodium Mont. 1848 (C)*

?Polychaetella Speg. 1918, anamorphic

Capnodium Mont. 1848 (C)*

Polychaeton (Pers.) Lev. 1846 (C)*

Scolecoxyphium Cif. & Bat. 1956,

anamorphic Capnodium Mont. 1848

and Scorias Fr. 1825 (C)*

Tripospermum Speg. 1918 anamorphic

Triposporiopsis W.Yamam 1955 (H)*

Coccodiniaceae Höhn. ex O.E. Erikss.

Bisbyopeltis Bat. & A.F. Vital 1957 (C)

Microxiphium (Harv. ex Berk. & Desm.)

Thüm. 1879, anamorphic Dennisiella

Bat. & Cif. 1962 (H)*

Davidiellaceae C.L. Schoch, Spatafora, Crous

& Shoemaker

Acroconidiella J.C. Lindq. & Alippi 1964,

anamorphic Davidiella Crous & U.

Braun 2003 (H)

Cladosporium Link 1816, anamorphic

Davidiella Crous & U. Braun 2003 (H)

Dichocladosporium K. Schub., U. Braun

& Crous 2007 (C)

Graphiopsis Trail 1889, anamorphic

Davidiella Crous & U. Braun 2003 (H)

Hoornsmania Crous 2007 (H)

Rachicladosporium Crous, U. Braun &

C.F. Hill 2007 (H)

Toxicocladosporium Crous & U. Braun

2007 (H)

Verrucocladosporium K. Schub., Aptroot

& Crous 2007 (H)

Dissoconiaceae Crous & de Hoog

Dissoconium de Hoog, Oorschot & Hijwe-

gen 1983 (H)

Ramichloridium Stahel 1937 ex de Hoog

1977 (H)

Metacapnodiaceae S. Hughes & Corlett

Capnobotrys S. Hughes 1970, anamorphic

Metacapnodium Speg. 1918 (H)

Capnocybe S. Hughes 1966, anamorphic

Metacapnodium Speg. 1918 (H)

Capnophialophora S. Hughes 1966, ana-

morphic Metacapnodium Speg. 1918

(H)

Capnosporium S. Hughes 1976, anamor-

phic Metacapnodium Speg. 1918 (H)

Hormiokrypsis Bat. & Nascim. 1957, ana-

morphic Metacapnodium Speg. 1918

and Ophiocapnocoma Bat. & Cif. 1963

(H)

Torulopsiella Bender 1932, anamorphic

Mycosphere Doi 10.5943/mycosphere/3/2/5

161

Ophiocapnocoma Bat. & Cif. 1963 and

Metacapnodium Speg. 1918 (H)

Mycosphaerellaceae Lindau

Anguillosporella U. Braun 1995 (H)

Asperisporium Maubl. 1913 (H)

Asteromella-like, anamorphic Gillotia

Sacc. & Trotter 1913 (C)

Cercospora Fresen. 1863, anamorphic

Mycosphaerella Johanson 1884, Sphae-

rulina Sacc. and others (H)

Cercosporella Sacc. 1880, anamorphic

Mycosphaerella Johanson 1884 (H)

?Clypeispora A.W. Ramaley 1991, ana-

morphic Mycosphaerella Johanson

1884 (C)

Colletogloeum Petr. 1953, anamorphic

Mycosphaerella Johanson 1884 (C)

Coniothryium-like, anamorphic Bruneos-

phaerella Crous 2009 (C)

Deightoniella S. Hughes 1952 (H)

Didymochora Höhn. 1918, anamorphic

Euryachora Fuckel 1870 (C)

Distocercospora N. Pons & B. Sutton

1988 (H)

Dothistroma Hulbary 1941 (C)

Floricola Kohlm. & Volkm.Kohlm. 2000

(C)

Fusicladiella Höhn. 1919, anamorphic

Mycosphaerella Johanson 1884 (H)

Laocoön J.C. David 1997 (H)

Lecanosticta Syd. 1922 (C)

Microcyclospora Jana Frank, Schroers &

Crous 2010 (H)

Microcyclosporella Jana Frank, Schroers

& Crous 2010 (H)

Miuraea Hara 1948, anamorphic Myco-

sphaerella Johanson 1884 (H)

Mycopappus Redhead & G.P. White 1985,

anamorphic Mycosphaerella Johanson

1884 (H)

Mycovellosiella Rangel 1918, anamorphic

Mycosphaerella Johanson 1884 (H)

Parastenella J.C. David 1991 (H)

Passalora Fr. 1849, anamorphic Myco-

sphaerella Johanson 1884 (H)

Penidiella Crous & U. Braun 2007 (H)

Periconiella Sacc. 1885, anamorphic

Allosoma Syd. 1926 (H)

Phacellium Bonord. 1860, anamorphic

Mycosphaerella Johanson 1884 (H)

Phaeoisariopsis Ferraris 1909, anamorphic

Mycosphaerella Johanson 1884 (H)

Phaeophleospora Rangel 1916 (C)

Phaeothecoidea Crous 2007 (H)

Phloeospora Wallr. 1833, anamorphic

Mycosphaerella Johanson 1884 (C)

Placosphaeria-like, anamorphic Euryach-

ora Fuckel 1870 (C)

Polythrincium Kunze 1817, anamorphic

Cymadothea F.A. Wolf 1935 (H)

Pseudocercospora Speg. 1910 anamorphic

Mycosphaerella Johanson 1884 (H)

Pseudocercosporella Deighton 1973, ana-

morphic Mycosphaerella Johanson

1884 (H)

Pseudocercosporidium Deighton 1973 (H)

Quasiphloeospora B. Sutton, Crous &

Shamoun 1996 (H)

Ramichloridium Stahel ex de Hoog 1977

(H)

Ramichloridium-like, anamorphic Mycosp-

haerella Johanson 1884 (H)

Ramularia Unger 1833, anamorphic Myc-

osphaerella Johanson 1884 and

Melanodothis R.H. Arnold 1972 (H)

Ramulariopsis Speg. 1910, anamorphic

Mycosphaerella Johanson 1884 (H)

Ramulispora Miura 1920, anamorphic

Mycosphaerella Johanson 1884 (H)

Rhabdospora (Durieu & Mont. ex Sacc.)

Sacc. 1884 (C)

Rhexocercosporidium U. Braun 1994 (H)

Semipseudocercospora J.M. Yen 1983 (H)

Septoria Sacc. 1884 (C)

Septoria-like, anamorphic Mycosphaerella

Johanson 1884 (C)

Sirosporium Bubák & Serebrian. 1912 (H)

Sonderhenia H.J. Swart & J. Walker 1988

(C)

Stenella Syd. 1930, anamorphic Myco-

sphaerella Johanson 1884 (H)

Stenellopsis B. Huguenin 1966 (H)

Stigmina Sacc. 1980, anamorphic Myco-

sphaerella Johanson 1884 (H)

Thedgonia B. Sutton 1973, anamorphic

Mycosphaerella Johanson 1884 (H)

Thedgonia-like (H)

Trochophora R.T. Moore 1955 (H)

Verrucisporota D.E. Shaw & Alcorn 1993

(H)

Xenostigmina Crous 1998, anamorphic

Mycosphaerella Johanson 1884 or

Didymella Sacc. ex D. Sacc. 1880 (H)

Mycosphere Doi 10.5943/mycosphere/3/2/5

162

Zasmidium Fr. 1849, anamorphic Mycosp-

haerella-like and Rasutoria M.E. Barr

1987 (H)

Zymoseptoria Quaedvlieg & Crous 2011

(C)*

Capnodiales, genera incertae sedis

Capnocheirides J.L. Crane & S. Hughes

1982 (H)

Heptaster Cif., Bat. & Nascim. 1956 (H)

Hormonema-like, anamorphic Xenomeris

Syd. 1924 (H)

Houjia G.Y. Sun & Crous 2010 (H)

Micropustulomyces R.W. Barreto 1995 (C)

Phaeotheca Sigler, Tsuneda & J.W.

Carmich. 1981 (H)

Phaeothecoidiella Batzer & Crous 2010

(H)

Pseudovirgaria H.D. Shin, U. Braun,

Arzanlou & Crous 2007 (H)

Rachicladosporium Crous, U. Braun &

C.F. Hill 2007 (H)

Recurvomyces Selbmann & de Hoog 2008

(H)

Scolecostigmina U. Braun 1999 (H)

Sporidesmajora Batzer & Crous 2010 (H)

Verrucocladosporium K. Schub., Aptroot

& Crous 2007 (H)

Dothideales Lindau

Dothideaceae Chevall.

Lecanosticta Syd. 1922, anamorphic Sci-

rrhia Nitschke ex Fuckel 1870 (C)

Podoplaconema Petr. 1921, anamorphic

Omphalospora Theiss. & Syd. 1915 (C)

Dothioraceae Theiss. & H. Syd.

Aureobasidium Viala & G. Boyer 1891,

anamorphic Columnosphaeria Munk

1953 and Saccothecium Fr. 1836 (H)

Dothichiza Lib. ex Roum. 1880, ana-

morphic Dothiora Fr. 1849 and Sydo-

wia Bres. 1895 (C)

Hormonema Lagerb. & Melin 1927, ana-

morphic Dothiora Fr. 1849, Prings-

heimia Schulzer 1866 and Sydowia

Bres. 1895 (H)

Japonia Höhn. 1909, anamorphic Yoshina-

gaia Henn. 1904 (C)

Kabatina R. Schneid. & Arx 1966 (C)

?Rhizosphaera L. Mangin & Har. 1907,

anamorphic Phaeocryptopus Naumov

1915 and Dimerina Theiss. 1912 (C)

Sclerophoma Höhn. 1909, anamorphic

Sydowia Bres. 1895 (C)

Teratosphaeriaceae Crous & U. Braun

Acidomyces B.J. Baker, M.A. Lutz, S.C.

Dawson, P.L. Bond & Banfield ex

Selbmann, de Hoog & De Leo 2008 (H)

Batcheloromyces Marasas, P.S. van Wyk

& Knox-Dav. 1975 (H)

Baudoinia J.A. Scott & Unter. 2007 (H)

Capnobotryella Sugiy. 1987 (H)

Catenulostroma Crous & U. Braun 2007,

anamorphic Teratosphaeria Syd. & P.

Syd. 1912 (H)

Cibiessia Crous 2007 (H)

Colletogloeopsis Crous & M.J. Wingf.

1997, anamorphic Teratosphaeria Syd.

& P. Syd. 1912 (C)

Davisoniella H.J. Swart 1988 (C)

Devriesia Seifert & N.L. Nick. 2004 (H)

Elasticomyces Zucconi & Selbmann 2008

(H)

Friedmanniomyces Onofri 1999 (H)

Hispidoconidioma Tsuneda & Davey 2010

(H)

Hobsoniopsis D. Hawksw. 2001 (H)

Hortaea Nishim. & Miyaji 1984 (H)

Kirramyces J. Walker, B. Sutton & Pascoe

1992, anamorphic Teratosphaeria Syd.

& P. Syd. 1912 (C)

Nothostrasseria Nag Raj 1983 (C)

Passalora Fr. 1849 (H)

Penidiella Crous & U. Braun 2007 (H)

Phacellium Bonord. 1860 (H)

Phaeothecoidea Crous 2007 (H)

Pseudocercosporella Deighton 1973 (H)

Pseudoramichloridium Cheewangkoon &

Crous 2010 (H)

Pseudotaeniolina J.L. Crane & Schokn.

1986 (H)

Ramichloridium Stahel ex de Hoog 1977

(H)

Recurvomyces Selbmann & de Hoog 2008

(H)

Readeriella Syd. & P. Syd. 1908 (C)

Sporidesmium Link 1809 (H)

Staninwardia B. Sutton 1971 (C)

Stenella Syd. 1930 (H)

Trimmatostroma-like, anamorphic Terato-

sphaeria Syd. & P. Syd. 1912 (H)

Tripospermum Speg. 1918 (H)

Mycosphere Doi 10.5943/mycosphere/3/2/5

163

Myriangiales Starbäck

Elsinoaceae Höhn. ex Sacc. & Trotter

Endosporium Tsuneda 2008 (H)

Sphaceloma de Bary 1874 anamorphic

Elsinoë Racib. 1900 (C)

Xenodiella Syd. 1935, anamorphic Xeno-

dium Syd. 1935 (H)

Myriangiales, genera incertae sedis

Phaeosclera Sigler, Tsuneda & J.W. Car-

mich. 1981 (H)

Sarcinomyces Lindner 1898 (H)

Zopfiaceae G. Arnaud ex D. Hawksw. 1992

Cladosporium-like, anamorphic Zopfia Rab-

enh. 1874 (H)

Subclass Pleosporomycetidae C.L. Schoch,

Spatafora, Crous & Shoemaker

Pleosporales Luttrell ex M.E. Barr

Amniculicolaceae Yin. Zhang, C.L. Schoch, J.

Fourn., Crous & K.D. Hyde

Anguillospora Ingold 1942 (H)

Spirosphaera Beverw. 1953 (H)

Cucurbitariaceae G. Winter

Camarosporium Schulzer 1870, anamor-

phic Cucurbitaria Gray 1821 (C)

Coniothyrium-like, anamorphic Curreya

Sacc. 1883 (C)

Diplodia-like, anamorphic Cucurbitaria

Gray 1821 (C)

?Megaloseptoria Naumov 1925, anamor-

phic Cucurbitaria Gray 1821 (C)

?Phaeoseptoria Speg. 1908, anamorphic

Rhytidiella Zalasky 1968 (C)

Phialophorophoma Linder 1944 (C)

Phoma Sacc. 1880 (C)

Pleurophoma Höhn. 1914 (C)

Pleurostromella Petr. 1922, anamorphic

Cucurbitaria Gray 1821 and Gibberi-

dea Fuckel 1870 (C)

Pyrenochaeta De Not. 1849, anamorphic

Cucurbitaria Gray 1821 (C)

Pyrenochaetopsis Gruyter, Aveskamp, &

Verkley, 2010 (C)

Syntholus A.W. Ramaley & M.E. Barr

1997, anamorphic Syncarpella Theiss.

& Syd. 1915 (C)

Diademaceae Shoemaker & C.E. Babc.

Alternaria-like, anamorphic Comoclathris

Clem. 1909 (H)*

Fusicladiella Höhn. 1919, anamorphic

Didymosphaeria Fuckel 1870 (H)

Didymellaceae Gruyter, Aveskamp & Verkley

Ascochyta Lib. 1830, anamorphic Didy-

mella Sacc. ex D. Sacc. 1880 (C)

Ampelomyces Ces. ex Schltdl. 1852 (C)

Boeremia Aveskamp, Gruyter & Verkley

2010 (C)

Chaetasbolisia Speg. 1918 (C)

Dactuliochaeta G.L. Hartm. & J.B. Sincl-

air 1988, anamorphic Didymella Sacc.

1880 (C)

Microsphaeropsis Höhn. 1917 (C)

Peyronellaea Gold. ex Togliani 1952 (C)

Phoma Sacc. 1880, anamorphic Didymella

Sacc. ex D. Sacc. 1880 (C)

Piggotia Berk. & Broome 1851, possibly

anamorphic Platychora Petr. 1925 (C)

Pithomyces Berk. & Broome 1875, ana-

morphic Leptosphaerulina McAlpine

1902 (H)

Stagonosporopsis Died. 1912, anamorphic

Didymella Sacc. ex D. Sacc. 1880 (C)

(=Ascochyta Lib. 1830)

Didymosphaeriaceae Munk

Cytoplea Bizz. & Sacc. 1885, anamorphic

Roussoëlla Sacc. 1888 (C)

Dothidotthiaceae Crous & A.J.L. Phillips

?Dothiorella Sacc. 1880, anamorphic Dot-

hidotthia Höhn. 1918 (C)

Thyrostroma Höhn. 1911, anamorphic

Dothidotthia Höhn. 1918 (H)

Hypsostromataceae Huhndorf

Pleurophomopsis-like, anamorphic Hyp-

sostroma Huhndorf (C)

Lentitheciaceae Yin. Zhang, C.L. Schoch, J.

Fourn., Crous & K.D. Hyde

Stagonospora (Sacc.) Sacc. 1884 (C)

Leptosphaeriaceae M.E. Barr

Camposporium Harkn. 1884, anamorphic

Leptosphaeria Ces. & De Not. 1863

(H)

Coniothyrium Corda 1840, anamorphic

Leptosphaeria Ces. & De Not. 1863 (C)

Coniothyrium-like, anamorphic Neophaeo-

Mycosphere Doi 10.5943/mycosphere/3/2/5

164

sphaeria M.P.S. Câmara, M.E. Palm &

A.W. Ramaley 2003 (C)

Phoma Sacc. 1880, anamorphic Leptosph-

aeria Ces. & De Not. 1863 (C)

Plenodomus Preuss 1851 (C)

Pyrenochaeta De Not. 1849, possibly

anamorphic Leptosphaeria Ces. & De

Not. 1863 (C)

Lindgomycetaceae K. Hiray., Kaz. Tanaka &

Shearer

Taeniolella S. Hughes 1958 (H)

Lolia Abdel-Aziz & Abdel-Whab 2010

(C)*

Lophiostomataceae Sacc.

Anguillospora Ingold 1942, anamorphic

Massarina Sacc. 1883 (H)

Ascochyta Lib. 1830, anamorphic Tricho-

metasphaeria Munk 1953 (C)

Periconia Tode 1791 anamorphic Lophio-

stoma Ces. & De Not. 1863 (H)

Pleuorphomopsis-like, anamorphic Lophi-

ostoma Ces. & De Not. 1863 (C)

Pyricularia Sacc. 1880, anamorphic Mass-

arina Sacc. 1883 (H)

Tumularia Descals & Marvanová 1987(H)

Massariaceae Nitschke

?Aplosporella-like, anamorphic Dubitatio

Speg. 1882 (C)

Myxocyclus Riess 1852, anamorphic Mass-

aria De Not. 1844 (C)

Torula Pers. 1794 (H)

Massarinaceae Munk

Aquaticheirospora Kodsueb & W.H. Ho

2007 (H)

Ceratophoma Höhn. 1917, anamorphic

Massarina Sacc. 1883 (C)

Cheirosporium L. Cai & K.D. Hyde 2008

(H)

Clavariopsis De Wild. 1895 (H)

Helminthosporium Link 1809, anamorphic

Splanchnonema Corda 1829 (H)

Neottiosporina Subram. 1961 (C)

Pseudodictyosporium Matsush. 1971 (H)

Melanommataceae G. Winter

Aposphaeria Sacc. 1880 (C)

Beverwykella Tubaki 1975 (H)

Exosporiella P. Karst. 1892, anamorphic

Anomalemma Sivan. 1983 (H)

Nigrolentilocus R.F. Castañeda & Heredia

2001, anamorphic Melanomma Nitsch-

ke ex Fuckel 1870 (H)

Monodictys S. Hughes 1958, anamorphic

Ohleria Fuckel 1868 (H)

Monotosporella S. Hughes 1958 (H)

Pyrenochaeta De Not. 1849, anamorphic

Byssosphaeria Cooke 1879 (C)

Sporidesmiella P.M. Kirk 1982 (H)

Montagnulaceae M.E. Barr

Aschersonia Mont. 1848, anamorphic

Montagnula Berl. 1896 (C)

Microdiplodia Allesch. 1901, possibly

anamorphic Karstenula Speg. 1879 (C)

Microsphaeropsis Höhn. 1917, anamorph-

ic Paraphaeosphaeria O.E. Erikss.

1967 (C)

Paraconiothyrium Verkley 2004, possibly

anamorphic Paraphaeosphaeria O.E.

Erikss. 1967 (C)

Phaeosphaeriaceae M.E. Barr

Amarenographium O.E. Erikss. 1982, ana-

morphic Amarenomyces O.E. Erikss.

1981 (recently synonymised with

Phaeosphaeria I. Miyake 1909) (C)

Ampelomyces Ces. ex Schltdl. 1852 (C)

Chaetosphaeronema Moesz 1915 (C)

Coniothyrium Corda 1840, anamorphic

Ophiobolus Riess 1854 (C)

Harpophora W. Gams 2000, anamorphic

anamorphic Ophiobolus Riess 1854 (H)

Hendersonia-like, anamorphic Austropleo-

spora R.G. Shivas & L. Morin 2010

and Phaeosphaeria I. Miyake 1909 (C)

Mauginiella Cavara 1925 (H)

Neosetophoma Gruyter, Aveskamp &

Verkley 2010 (C)

?Parahendersonia A.W. Ramaley 1995,

anamorphic Chaetoplea (Sacc.) Clem.

1931 (C)

Paraphoma Morgan-Jones & J.F. White

1983 (C)

Phaeoseptoria Speg. 1908, anamorphic

Phaeosphaeria I. Miyake 1909 (C)

Phaeostagonospora A.W. Ramaley 1997,

anamorphic Phaeosphaeriopsis M.P.S.

Câmara, M.E. Palm & A.W. Ramaley

2003 (C)

Phoma Sacc. 1880, anamorphic Ophio-

Mycosphere Doi 10.5943/mycosphere/3/2/5

165

bolus Riess 1854 (C)

Rhabdospora (Durieu & Mont. ex Sacc.)

Sacc. 1884, anamorphic Ophiobolus

Riess 1854 (C)

Scolecosporiella Petr. 1921, anamorphic

Ophiosphaerella Speg. and Phaeo-

sphaeria I. Miyake 1909 (C)

Setophoma Gruyter, Aveskamp & Verkley

2010 (C)

?Sphaerellopsis Cooke 1883, anamorphic

Eudarluca Speg. 1908 (C)

Stagonospora (Sacc.) Sacc. 1884, anamor-

phic Phaeosphaeria I. Miyake 1909 (C)

Tiarospora Sacc. & Marchal 1885 (C)

Pleomassariaceae M.E. Barr

Ceuthodiplospora Died. 1912, anamorphic

Splanchnonema Corda (C)

Corynespora Güssow 1906, anamorphic

Pleomassaria Speg. 1880 (H)

Myxocyclus Riess 1852, anamorphic

Splanchnonema Corda 1829 (C)

?Prosthemium Kunze 1817, anamorphic

Pleomassaria Speg. 1880 (C)

Scolicosporium Lib. ex Roum. 1880, ana-

morphic Asteromassaria Höhn. 1917

(C)

Shearia Petr. 1924, anamorphic Pleomas-

saria Speg. 1880 (C)

Stegonsporium Corda 1827, anamorphic

Splanchnonema Corda 1829 (C)

Pleosporaceae Nitschke

Alternaria Nees 1816, anamorphic Lewia

M.E. Barr & E.G. Simmons 1986 (H)*

Alternaria-like anamorphic Clathrospora

Rabenh. 1857 (H)

Alternariaster E.G. Simmons 2007 (H)

Brachycladium Corda 1838, anamorphic

Crivellia Shoemaker & Inderbitzin

2006 (H)

Bipolaris Shoemaker 1959, anamorphic

Cochliobolus Drechsler 193 (H)

Chalastospora E.G. Simmons 2007 (H)

Corynespora Güssow 1906, anamorphic

Pyrenophora Fr. 1849 (H)

Curvularia Boedijn 1933, anamorphic

Cochliobolus Drechsler 1934 (H)

Dactuliophora C.L. Leakey 1964 (H)

Dendryphiella Bubák & Ranoj. 1914,

anamorphic Pleospora Rabenh. ex Ces.

& De Not. 1863 (H)

Dendryphion Wallr. 1833, anamorphic

Pleospora Rabenh. ex Ces. & De Not.

1863 (H)

Dictyosporium Corda 1836 (H)

Digitodesmium P.M. Kirk 1981 (H)

Drechslera S. Ito 1930, anamorphic Pyre-

nophora Fr. 1849 (H)

Edenia M.C. González, Anaya, Glenn,

Saucedo & Hanlin 2007 (H)

Embellisia E.G. Simmons 1971, anamor-

phic Lewia M.E. Barr & E.G. Simmons

(H)

Exserohilum K.J. Leonard & Suggs 1974,

anamorphic Setosphaeria K.J. Leonard

& Suggs 1974 (H)

Gibbago E.G. Simmons 1986 (H)

Lemonniera De Wild. 1894 (H)

Marielliottia Shoemaker 1999, anamor-

phic Pyrenophora Fr. 1849 (H)

Nimbya E.G. Simmons 1989, anamorphic

Macrospora Fuckel 1870 (H)

Prathoda Subram. 1956 (H)

Sinomyces Yong Wang bis & X.G. Zhang

2011 (H)

Stemphylium Wallr. 1833, anamorphic Ple-

ospora Rabenh. ex Ces. & De Not.

1863 (H)

Teretispora E.G. Simmons 2007 (H)

Ulocladium Preuss 1851, anamorphic Las-

iobotrys Kunze 1823 (H)

Undifilum Pryor, Creamer, Shoemaker,

McLain-Romero & Hambl. 2009 (H)

Sporormiaceae Munk

?Amorosia Mantle & D. Hawksw. 2006

(H)

Phoma Sacc. 1880 (C)

Phoma-like, anamorphic Westerdykella

Stolk 1955 (C)

Teichosporaceae M.E. Barr

Coniothyrium-like, anamorphic Immothia

M.E. Barr (C)

Tetraplosphaeriaceae Kaz. Tanaka & K. Hiray

Pseudotetraploa Kaz. Tanaka & K.

Hirayama 2009 (H)

Quadricrura Kaz. Tanaka, K. Hirayama &

Sat. Hatak. 2009 (H)

Tetraploa Berk. & Broome 1850, ana-

morphic Tetraplosphaeria Kaz. Tanaka

& K. Hirayama 2009 (H)

Mycosphere Doi 10.5943/mycosphere/3/2/5

166

Tetraploa-like, anamorphic Triplosphaeria

Kaz. Tanaka & K. Hirayama 2009 and

Polyplosphaeria Kaz. Tanaka & K.

Hirayama 2009 (H)

Trematosphaeriaceae K.D. Hyde, Yin. Zhang,

Suetrong & E.B.G. Jones

Hyphopodia-like, anamorphic Tremato-

sphaeria Fuckel, Jb. nassau. Ver.

Naturk (C)*

Pleosporales, genera incertae sedis

Berkleasmium Zobel 1854 (H)

Briansuttonia R.F. Castañeda, Minter &

Saikawa 2004 (H)

Cerebella Ces. 1851 (H)

Cheiromoniliophora Tzean & J.L. Chen

1990 (H)

Diplococcium Grove 1885 (H)

Ellisembia Subram. 1992 (H)

Epicoccum Link 1815 (H)

Fusculina Crous & Summerell 2006 (C)

Mycocentrospora Deighton 1972 (H)

Mycopappus Redhead & G.P. White 1985

anamorphic Mycodidymella C.Z. Wei,

Y. Harada & Katum. 1998 (H)

Noosia Crous, R.G. Shivas & McTaggart

2011 (H)

Ochrocladosporium Crous & U. Braun

2007 (H)

Phoma Sacc. 1880 (C)

Periconia Tode 1791(H)

Pleurophomopsis Petr. 1924, anamorphic

Astrosphaeriella Syd. & P. Syd. 1913

(C)

Polyschema H.P. Upadhyay 1966 (H)

Pseudochaetosphaeronema Punith. 1979

(C)

Pseudopassalora Crous 2011 (H)

Repetophragma Subram. 1992 (H)*

?Scleroramularia Batzer & Crous 2011

(H)

Sclerostagonospora Höhn. 1917 (C)

Sirodesmium De Not. 1849 (C)

Spiroplana Voglmayr, M.J. Park & H.D.

Shin 2011 (H)

Sporidesmium Link 1809 (C)

Versicolorisporium Sat. Hatak., Kaz.

Tanaka & Y. Harada 2008 (C)

Ybotromyces Rulamort 1986 (H)

Dothideales genera incertae sedis

Atramixtia Tsuneda, M.L. Davey &

Currah 2011 (H)

Celosporium Tsuneda & Davey 2010 (H)

Diplococcium Grove 1885, anamorphic

Otthia Nitschke ex Fuckel 1870 (H)

Pleosporomycetidae, genera incertae sedis

Acrogenospora M.B. Ellis 1971, anamor-

phic Farlowiella Sacc. 1891(H)

Dothideomycetes, orders incertae sedis

Acrospermales Minter, Peredo & A.T. Watson

Acrospermaceae Fuckel

Dactylaria Sacc. 1880, anamorphic Acro-

spermum Tode 1783 (H)

Gonatophragmium Deighton 1969, ana-

morphic Acrospermum Tode 1790 (H)

Botryosphaeriales C.L. Schoch, Crous &

Shoemaker

Botryosphaeriaceae Theiss. & H. Syd.

Alysidiella Crous 2006 (H)

Aplosporella Speg. 1880 (C)

Arthrinium Kunze 1817, anamorphic Lept-

oguignardia E. Müll. (H)

Bahusutrabeeja Subram. & Bhat 1977 (H)

Barriopsis A.J.L. Phillips, A. Alves &

Crous 2008 (C)

Dichomera Cooke 1878 (C)

Diplodia Fr. 1834 (C)

Diplodia-like, anamorphic Neodeightonia

C. Booth 1970 and Saccharata Denman

& Crous 2004 (C)

Dothichiza-like, anamorphic Leptoguig-

nardia E. Müll. 1955 (C)

Dothiorella Sacc. 1880, anamorphic Spen-

cermartinsia A.J.L. Phillips, A. Alves

& Crous 2008 (C)

Fusicoccum Corda 1829, anamorphic

Botryosphaeria Ces. & De Not. 1863

(C)

Fusicoccum-like, anamorphic Melanops

Nitschke ex Fuckel 1870 and Saccha-

rata Denman & Crous 2004 (C)

Hormonema Lagerb. & Melin 1927,

anamorphic Guignardia Viala & Ravaz

1892 (H)

Lasiodiplodia Ellis & Everh. 1896 (C)

Leptodothiorella Höhn. 1918, anamorphic

Guignardia Viala & Ravaz 1892 (C)

Macrophomina Petr. 1923 (C)

Microdiplodia Allesch. 1901 (C)

Mycosphere Doi 10.5943/mycosphere/3/2/5

167

Neofusicoccum Crous, Slippers & A.J.L.

Phillips 2006, with Botryosphaeria-like

teleomorphs (C)

Neoscytalidium Crous & Slippers 2006

(H)

Phyllosticta Pers. 1818, anamorphic Guig-

nardia Viala & Ravaz 1892 (C)

Pseudofusicoccum Mohali, Slippers &

M.J. Wingf. 2006 (C)

Sphaeropsis Sacc. 1880 (C)

Thyrostroma Höhn. 1911, anamorphic

Dothidotthia Höhn. 1918 (H)

Botryosphaeriales, genera incertae sedis

Camarosporium Schulzer 1870 (C)

Dichomera Cooke 1878 (C)

Hendersonula Speg. 1880 (C)

Hysteriales Lindau

Hysteriaceae Chevall.

Aposphaeria-like, anamorphic Gloniopsis

De Not. 1847 and Hysterobrevium E.

Boehm & C.L. Schoch 2010 (C)

Coniosporium Link 1809, anamorphic

Hysterium Pers. 1797 (H)

Plenodomus Preuss 1851, anamorphic

Psiloglonium Höhn. 1918 (C)

Septonema Corda 1837, anamorphic Oedo-

hysterium E. Boehm & C.L. Schoch

2009 (H)

Sporidesmium Link 1809, anamorphic

Psiloglonium Höhn. 1918 (H)

Jahnulales Pang, Abdel-Wahab, El-

Sharouney, E.B.G. Jones & Sivichai

Aliquandostipitaceae Inderbitzin

Brachiosphaera Nawawi 1976 (H)

Mytilinidiales Boehm, C.L. Schoch &

Spatafora

Gloniaceae (Corda) Boehm, C.L. Schoch &

Spatafora

Cenococcum Moug. & Fr. 1829 (C)

Cleistonium Speer 1986, anamorphic

Glonium Mühl. 1813 (C)

Mytilinidiaceae Kirschst.

Camaroglobulus Speer 1986, anamorphic

Mytilinidion Duby 1862 (C)

Chalara (Corda) Rabenh. 1844, anam-

orphic Quasiconcha M.E. Barr & M.

Blackw. 1981 (H)*

?Monotosporella S. Hughes 1958, anam-

orphic Actidium Fr. 1815 (H)

Papulospora Preuss 1851, anamorphic

Lophium Fr. 1818 (H)

Septonema Corda 1837, anamorphic

Mytilinidion Duby 1862 (H)

Taeniolella S. Hughes 1958, anamorphic

Mytilinidion Duby 1861 (H)

Jahnulales genera incertae sedis

Xylomyces Goos, R.D. Brooks & Lamore

1977 (H)

Patellariales D. Hawksw. & O.E. Erikss.

Patellariaceae Corda

Aposphaeria-like, anamorphic Rhytidhy-

steron Speg. 1881 (C)

Diplodia-like, anamorphic Rhytidhysteron

Speg. 1881 (C)

Venturiales Yin. Zhang, C.L. Schoch & K.D.

Hyde

Venturiaceae E. Müll. & Arx ex M.E. Barr

Acroconidiellina (Berk. & Broome) M.B.

Ellis 1971, anamorphic Acantharia

Theiss. & Syd. (H)*

Bactrodesmium Cooke 1883, anamorphic

Phragmogibbera Samuels & Rog-

erson 1990 (H)

Dictyodochium Sivan. 1984, anamorphic

Gibbera Fr. 1825 (H)

?Didymochora Hohn. 1918, anamorphic

Atopospora Petr. 1925 (C)

Cladosporium-like, anamorphic Apiosp-

orina Höhn. 1910 (H)

Fusicladium Bonord. 1851, anamorphic

Apiosporina Höhn. 1910, Caprov-

enturia Braun 1998 and Venturia Sacc.

1882 (H)

Fusicladium-like, anamorphic Acantharia

Theiss. & Syd 1918 and Protove-

nturia Berl. & Sacc. 1887 (H)

Fusicladosporium Partr. & Morgan-Jones

2003, anamorphic Venturia Sacc.

1882 (H)

Helicodendron Peyronel 1918 anamorphic

Tyrannosorus Unter. & Malloch 1995

(H)

Pithosira Petr. 1949 (H)

Spilocaea Fr. 1819, anamorphic Venturia

Sacc. 1882 (C)

Spilodochium Syd. 1927 anamorphic Pseu-

Mycosphere Doi 10.5943/mycosphere/3/2/5

168

doparodiella F. Stevens 1927 (H)

Stigmina-like, Sacc. 1980 anamorphic

Aca-ntharia Theiss. & Syd 1918 and

Gib-bera Fr. 1825 (H)

Sympoventuriaceae Yin. Zhang, C.L. Schoch

& K.D. Hyde

Fusicladium-like (H)

Sympodiella-like, anamorphic Sympoven-

turia Crous & Seifert (H)*

Veronaeopsis Arzanlou & Crous 2007

(H)*

Dothideomycetes, families incertae sedis

Ascoporiaceae Kutorga & D. Hawksw.

Plectophomella-like, anamorphic Pseudo-

solidum Lloyd 1923 (H)

Asterinaceae Hansf.

?Acarella Syd. 1927, anamorphic More-

noina Theiss. 1913 (C)

Asterostomella Speg. 1886, anamorphic

Asterina Lév. 1845 (C)

Asterostromina Bat. & A.F. Vital 1957,

anamorphic Asterodothis Theiss. (C)

Asterostomula Theiss. 1916, anamorphic

Echidnodes Theiss. & Syd. 1918 (C)

Bahusakala Subram. 1958 (H)

?Clasterosporium Schwein. 1832, anamo-

rphic Eupelte Syd. 1924 (H)

Clasterosporium Schwein. 1832, anam-

orphic Asterina Lév. 1845, Asterodothis

Theiss.1912, Yamamotoa Bat. 1960 and

Lembosia Lév. 1845 (H)

?Elachopeltis Syd. 1927, anamorphic Aph-

anopeltis Syd. 1927 (C)

Eriothyrium Speg. 1888, anamorphic

Aster-inema Bat. & Gayão 1953 (C)*

Leprieurina G. Arnaud 1918, anamorphic

Prillieuxina G. Arnaud 1918 (C)

Mahanteshamomyces Hosag. & C.K. Biju

2004 (C)

?Peltasterella Bat. & H. Maia 1959, ana-

morphic Yamamotoa Bat. 1960 (C)

Pirozynskia Subram. 1972, anamorphic

Eupelte Syd. 1924 (H)

Septoidium G. Arnaud 1921, anamorphic

Eupelte Syd. 1924 (H)

Septothyrella Höhn. 1911, anamorphic

Uleothyrium Petr. 1929 (C)

Shivamyces Hosag. 2004 (H)

Sirothyriella Höhn. 1910, anamorphic

Morenoina Thesis. 1913 (C)

?Thyrinula Petr. & Syd. 1924, anamorphic

Aulographina Arx & E. Müll. 1960 (C)

Triposporium Corda 1837, anamorphic

Batistinula Arx 1960 (H)

Corynesporascaceae Sivan.

Corynespora Güssow 1906, anamorphic

Corynesporasca Sivan. (H)

Dacampiaceae Körb.

Cyclothyrium Petr. 1923, anamorphic Pol-

ycoccum Saut. ex Körb. 1865 (C)

Lecythophora-like, anamorphic Munkoval-

saria Aptroot 1995 (H)

Englerulaceae Henn.

Capnodiastrum Speg. 1886, anamorphic

Englerula Henn. 1904 and Rhyti-

denglerula Höhn. 1918 (C)

Digitosarcinella S. Hughes 1984, anamo-

rphic Schiffnerula Höhn. 1909 (H)

Mitteriella Syd. 1933, anamorphic Schiff-

nerula Höhn. 1909 (H)

Questieriella G. Arnaud ex S. Hughes

1983, anamorphic Schiffnerula Höhn.

1909 (H)

Sarcinella Sacc. 1880, anamorphic Schiff-

nerula Höhn. 1909 (H)*

Eremomycetaceae Malloch & Cain

Arthrographis G. Cochet ex Sigler & J.W.

Carmich. 1976, anamorphic Eremomy-

ces Malloch & Cain 1971and Pitho-

ascina Valmaseda, A.T. Martínez &

Barrasa 1987 (H)

Trichosporiella Kamyschko 1960, anam-

orphic Rhexothecium Samson &

Mouch. 1975 and Eremomyces Malloch

& Cain 1971 (H)

Euantennariaceae Hughes & Corlett

Antennatula Fr. ex F. Strauss 1850, anam-

orphic Euantennaria Speg. and Strig-

opodia Bat. 1957 (H)

Capnokyma S. Hughes 1975 (H)

Capnophialophora S. Hughes 1966, ana-

morphic Strigopodia Bat. 1957 (H)

Hormisciomyces Bat. & Nascim. 1957,

anamorphic Euantennaria Speg. (H)

Mycosphere Doi 10.5943/mycosphere/3/2/5

169

Hyphosoma Syd. 1924, anamorphic Strig-

opodia Bat. 1957 (H)

Plokamidomyces Bat., C.A.A. Costa &

Cif. 1957, anamorphic Trichopeltheca

Bat., C.A.A. Costa & Cif. 1958 (H)

Racodium Fr. 1829, anamorphic Strigo-

podia Bat. 1957 (H)

Trichothallus F. Stevens 1925, anamorphic

Trichopeltheca Bat., C.A.A. Costa &

Cif. 1958 (H)

Fenestellaceae M.E. Barr

Pleurostromella Petr. 1922, anamorphic

Fenestella Tul. & C. Tul. 1863 (C)

Leptopeltidaceae Höhn. ex Trotter

?Idriella P.E. Nelson & S. Wilh. 1956,

anamorphic Dothiopeltis E. Mull. 1957

(H)

Leptothyrium-like, anamorphic Leptopeltis

Höhn. 1917 (C)

Meliolinaceae S. Hughes

Briania D.R. Reynolds 1989, anamorphic

Meliolina Syd. & P. Syd. 1914 (H)

Micropeltidaceae Clem. & Shear

Cyclopeltella Petr. 1953, anamorphic Cycl-

opeltis Petr. 1953 (C)

Heliocephala V.G. Rao, K.A. Reddy & de

Hoog 1984 (H)*

Holubovaniella R.F. Castañeda 1985 (H)*

Sirothyriella Höhn. 1910, anamorphic

Stomiopeltis Theiss. 1914 (C)

Sporidesmium-like, anamorphic Stomio-

peltis Theiss. 1914 (as Akaropeltella

M.L. Farr, 1972) (H)

Pycnothyriaceous taxa associated with spe-

cies of Stomiopeltis anamorphic Micro-

peltis Mont.

Microthyriaceae Sacc.

Asterostomula Theiss. 1916, anamorphic

Asterinella Theiss. 1912 (C)

Asteromella-like, anamorphic Asterinella

Theiss. 1912 (C)

Hansfordiella S. Hughes 1951, anamor-

phic Trichothyrium Speg. 1889 (H)

Isthmospora F. Stevens 1918, anamorphic

Trichothyrium Speg. 1889 (H)

Leptothyrium Kunze 1823, anamorphic

Microthyrium Desm. 1841 (C)

Xenogliocladiopsis Crous & W.B. Kendr.

1994, anamorphic Arnaudiella Petr.

1927 (H)

Zalerion R.T. Moore & Meyers 196, ana-

morphic Microthyrium Desm. 1841 (H)

Parmulariaceae E. Müll. & Arx ex M.E. Barr

Excipulariopsis P.M. Kirk & Spooner

1982, anamorphic Kentingia Sivan. &

W.H. Hsieh 1989 (H)

?Melanoplaca Syd. & P. Syd. 1917,

anamorphic Dothidasteroma Höhn.

1909 (C)

Placomelan Cif. 1962, anamorphic Doth-

idasteroma Höhn. 1909 (C)

Parodiellaceae Theiss. & H. Syd. ex M.E. Barr

?Ascochytopsis Henn. 1905, anamorphic

Parodiella Speg. 1880 (C)

Parodiopsidaceae Toro

?Chuppia Deighton 1965, anamorphic

Neoparodia Petr. & Cif. 1932 (H)

Cicinnobella Henn. 1904, anamorphic

Perisporiopsis Henn. 1904 (as Phaeo-

dimeriella Speg. 1908) and Dimerium

(Sacc. & P. Syd.) Sacc. & D. Sacc.

1905 (C)

Clasterosporium Schwein. 1832, anamor-

phic Balladyna Racib. 1900 (H)

Heterosporiopsis Petr. 1950, anamorphic

‘Dimerium’ (Sacc. & P. Syd.) Sacc. &

D. Sacc. 1905, Exosporinella Bender

1932, possibly anamorphic Parodiellina

Henn. ex G. Arnaud 1918 (H)

?Ophiotrichum Kunze 1849 anamorphic

Leptomeliola Höhn. 1919 (H)

Sarcinella-like, anamorphic Neoparodia

Petr. & Cif. 1932 (H)

Septoidium G. Arnaud 1921, anamorphic

Alina Racib. 1909, Perisporiopsis

Henn. 1904, Parodiellina Henn. ex G.

Arnaud 1918, Chevalieropsis G.

Arnaud 1923, Ophioparodia Petr. &

Cif. 1932 and Pilgeriella Henn. 1900

(H)

Shivamyces Hosag. 2004, anamorphic

Alina Racib. 1909 (H)

Tretospora M.B. Ellis 1976, anamorphic

Balladyna Racib. and Balladynopsis

Mycosphere Doi 10.5943/mycosphere/3/2/5

170

Theiss. & Syd. (H)

Planistromellaceae M.E. Barr

Aposphaeria-like, anamorphic Microcycl-

us Sacc., Syd. & P. Syd. 1904 (C)

Fusicladium Bonord. 1851, anamorphic

Microcyclus Sacc., Syd. & P. Syd. 1904

(H)

Hyphospora A.W. Ramaley 1996, ana-

morphic Comminutispora A.W. Rama-

ley 1996 (H)

Kellermania Ellis & Everh. 1885, ana-

morphic Planistroma A.W. Ramaley

1991 (C)

Lecanosticta Syd. 1922, anamorphic Erup-

tio M.E. Barr 1996 (C)

Pazschkeella Syd. & P. Syd. 1901, anam-

orphic Microcyclus Sacc., Syd. & P.

Syd. 1904 (C)

Piptarthron Mont. ex Höhn. 1918, anam-

orphic Planistroma A.W. Ramaley

1991 (C)

Polystomellaceae Theiss. & H. Syd.

Asteromella Pass. & Thüm. 1880 (as

Stictochorella Höhn. 1917), anamorphic

Dothidella Speg. 1880 (C)

Lasmenia Speg. 1886, anamorphic Mun-

kiella Speg. 1885 (C)

Pseudoperisporiaceae Toro

Chaetosticta Petr. & Syd. 1925, anamor-

phic Lasiostemma Theiss., Syd. & P.

Syd. 1917 and ?Nematostoma Syd. & P.

Syd. 1914 (C)

Schizothyriaceae Höhn. ex Trotter, Sacc., D.

Sacc. & Traverso

Plenotrichaius Bat. & Valle 1961, anamor-

phic Mycerema Bat., J.L. Bezerra &

Cavalc 1963 (C)

Zygophiala E.W. Mason 1945, anamorphic

Schizothyrium Desm. 1849 (H)

Tubeufiaceae M.E. Barr

?Annellospermosporella P.R. Johnst. 1999,

anamorphic Malacaria Syd. 1930 (H)

Aquaphila Goh, K.D. Hyde & W.H. Ho

1998, anamorphic Tubeufia Penz. &

Sacc. 1898 (H)*

Araneomyces Höhn. 1909, anamorphic

Paranectriella (P. Henn. & Sacc.)

Höhn. (H)

Chionomyces Deighton & Piroz. 1972,

anamorphic Melioliphila Speg. 1924

(H)

Corynespora Güssow 1906, anamorphic

Chaetosphaerulina I. Hino 1938 (H)

Eriomycopsis Speg. 1910, anamorphic

Melioliphila Speg. 1924 and Byssocallis

Syd. 1927 (H)

Guelichia Speg. 1886, anamorphic Putte-

mansia Henn. 1902 (H)

Helicoma Corda 1837, anamorphic Tubeu-

fia Penz. & Sacc. 1898, Thaxteriella

Petr. 1924 and Chlamydotubeufia Boo-

nmee & K.D. Hyde 2011 (H)*

Helicoön Morgan 1892 (H)*

Helicomyces Link 1809, anamorphic

Acanthostigma De Not. (1863) (H)*

Helicosporium Nees 1816, anamorphic

Acanthostigma De Not. (1863) and

Tubeufia Penz. & Sacc. 1898 (H)

Kamalomyces R.K. Verma, N. Sharma &

Soni 2008 (C)

Mirandina G. Arnaud ex Matsush. 1975,

anamorphic Taphrophila Scheuer 1988

(H)*

Monodictys S. Hughes 1958, anamorphic

Tubeufia Penz. & Sacc. 1898 (H)

Monodictys-like, anamorphic Tubeufia

Penz. & Sacc. 1898 (H)

Moorella P. Rag. Rao & D. Rao 1964,

anamorphic Chaetosphaerulina I. Hino

1938 (H)

Pendulispora M.B. Ellis 1961, anamorphic

Tubeufia Penz. & Sacc. 1898 (H)

Peziotrichum (Sacc.) Lindau 1900, ana-

morphic Podonectria Petch 1921 (H)

Spermatoloncha Speg. 1908 (H)

Tetracrium Henn. 1902, anamorphic Putt-

emansia Henn. 1902 and Podonectria

Petch 1921 (H)

Titaea Sacc. 1876, anamorphic Paranec-

triella (Henn. ex Sacc. & D. Sacc.)

Höhn. 1910 and Puttemansia Henn.

1902 (H)

Xenosporium Penz. & Sacc. 1901, anamor-

phic Acanthostigmella Höhn. 1905 and

Chaetosphaerulina I. Hino 1938 (as

Mycosphere Doi 10.5943/mycosphere/3/2/5

171

Thaxteriellopsis Sivan., Panwar & S.J.

Kaur 1977) (H)

Vizellaceae H.J. Swart

Chrysogloeum Petr. 1959, anamorphic

Blasdalea Sacc. & P. Syd. 1902 (C)

Manginula G. Arnaud 1918, anamorphic

Vizella Sacc. 1883 (C)

Dothideomycetes, genera incertae sedis

Acrocalymma Alcorn & J.A.G. Irwin

(1987), anamorphic Massarina, how-

ever Massarina has been shown to be

polyphyletic and its placement is there-

fore presently unclear (C)

Ampullifera Deighton 1960, anamorphic

Teratoschaeta Bat. & O.M. Fonseca

(H)

Ascochyta-like, anamorphic Gilletiella

Sacc. & P. Syd. 1899 (C)

Asteromella Pass. & Thüm. 1880 (C)

Bactrodesmium Cooke 1883, anamorphic

Stuartella Fabre 1879 (H)

Brachyconidiella R.F. Castañeda & W.B.

Kendr. 1990 (H)

Camarosporula Petr. 1954, anamorphic

Anthracostroma Petr. 1954 (C)

Chaetasbolisia Speg. 1918, anamorphic

Pseudomorfea Punith. 1981 (C)

Cladoriella Crous 2006 (H)

Coniosporium Link 1809 (H)

Coronospora M.B. Ellis 1971, anamorphic

Ascocoronospora Matsush. 2003 (H)

Cryomyces Selbmann, de Hoog, Mazzagl-

ia, Friedmann & Onofri 2005 (H)

?Cyclothyrium Petr. 1923, anamorphic

Thyridaria Sacc. 1875 (C)

Dendryphiopsis S. Hughes 1953, anamorp-

hic Kirschsteiniothelia D. Hawksw.

1985 (H)

Diederichia D. Hawksw. 2003 (C)

Dilophospora Desm. 1840, anamorphic

Lidophia J. Walker & B. Sutton 1974

(C)

Diplodia-like, anamorphic Otthia Nitschke

ex Fuckel 1870 (C)

Disculina Höhn. 1916 (C)

Funbolia Crous & Seifert 2011 (H)

Hansfordiellopsis Deighton 1960, anamor-

phic Koordersiella Höhn. 1909 (H)

Helicodendron Peyronel 1918, anamorphic

Tyrannosorus Unter. & Malloch 1995

(H)

Hiospira R.T. Moore 1962, anamorphic

Brooksia Hansf. 1956 (H)

Iledon Samuels & J.D. Rogers 1986, ana-

morphic Botryohypoxylon Samuels &

J.D. Rogers 1986 (C)

Megaloseptoria Naumov 1925 (C)

Mirandina G. Arnaud ex Matsush. 1975,

anamorphic Chaetonectrioides Matsu-

sh. 1996 (H)

Monodictys S. Hughes 1958 (H)

Peltaster Syd. & P. Syd. 1917 (C)

Phaeothecoidiella Batzer & Crous 2010

(H)

Phanerococculus Cif. 1954, anamorphic

Koordersiella Höhn. 1909 (C)

Phialophora-like, anamorphic Tirisporella

E.B.G. Jones, K.D. Hyde & Alias 1996

(H)

Phoma-like, anamorphic Tremateia Koh-

lm., Volkm.-Kohlm. & O.E. Erikss.

1995 (C)

Placosphaeria (De Not.) Sacc. 1880 (C)

Plectopycnis Bat. & A.F. Vital 1959 (C)

Pleurostromella Petr. 1922, anamorphic

Gibberidea Fuckel 1870 (C)

Pseudorobillarda M. Morelet 1968 (C)

Pycnopleiospora C.Z. Wei, Y. Harada &

Katum. 1997, anamorphic Pseudodidy-

mella C.Z. Wei, Y. Harada & Katum.

1997 (H)

Rhizopycnis D.F. Farr 1998 (C)

Scleroconidioma Tsuneda, Currah & Thor-

mann 2000 (H)

Septoriella Oudem. 1889 (C)

Troposporella P. Karst. 1892 (H)

Class Eurotiomycetes Tehler ex O.E. Eriksson

& K. Winka

Subclass Chaetothyriomycetidae Doweld

Chaetothyriales M.E. Barr

Chaetothyriaceae Hansf. ex M.E. Barr

?Ciferrioxyphium Bat. & H. Maia 1963,

anamorphic Aithaloderma P. Syd. (H)*

Cyphellophora G.A. de Vries 1962 (H)

Leptoxyphium Speg. 1918, anamorphic Ai-

thaloderma P. Syd. 1913 (H)*

Merismella Syd. 1927, anamorphic Chaet-

Mycosphere Doi 10.5943/mycosphere/3/2/5

172

othyrium Speg. 1888 (C)

Stanhughesia Constant. 1989, anamorphic

Ceramothyrium Bat. & H. Maia 1956

(H)

Tripospermum Speg. 1918, anamorphic

Phragmocapnias Theiss. & Syd. 1918,

Trichomerium Speg. 1918 and Tripo-

sporiopsis W. Yamam. 1955 (H)*

Vonarxia Bat. 1960 (H)

Herpotrichiellaceae Munk

Brycekendrickomyces Crous & M.J.

Wingf. 2009 (H)

Cladophialophora Borelli 1980, anamor-

phic Capronia Sacc. 1883 (H)

Exophiala J.W. Carmich. 1966, anamor-

phic Capronia Sacc. 1883 (H)

Fonsecaea Negroni 1936, anamorphic

Capronia Sacc. 1883 (H)

Metulocladosporiella Crous, Schroers, J.Z.

Groenew., U. Braun & K. Schub. 2006

(H)

Nadsoniella Issatsch. 1914 (H)

Phaeococcomyces de Hoog 1979 (H)

Phaeoramularia Munt.-Cvetk. 1960, ana-

morphic Capronia Sacc.1883 (H)

Phialophora Medlar 1915 (H)

Racodium Pers. 1794, anamorphic Capro-

nia Sacc. 1883 (H)

Rhinocladiella Nannf. 1934, anamorphic

Capronia Sacc. 1883 (H)

Sorocybe Fr. 1849 (H)

Thysanorea Arzanlou, W. Gams & Crous

2007 (H)

Veronaea Cif. & Montemart. 1957 (H)

Chaetothyriales M.E. Barr, genera incertae

sedis

Coniosporium Link 1809, the type species

is considered as Sirodesmium De Not.

1849 (H)

Phaeoannellomyces McGinnis & Schell

1985 (H)(= Hortaea)

Staninwardia B. Sutton 1971 (C)

Strelitziana Arzanlou & Crous 2006 (H)

Pyrenulales, genera incertae sedis

Lyromma Bat. & H. Maia 1965 (C)

Chaetothyriomycetidae, families incertae sedis

Rhynchostomataceae Winka & O. E. Erikss.

Arthropycnis Constant. 1992, anamorphic

Rhynchostoma P. Karst. (C)

Strigulaceae Zahlbr.

Discosiella Syd. & P. Syd. 1912, anamor-

phic Strigula Fr. 1823 (C)

Chaetothyriomycetidae, genera incertae sedis

Peyronelia Cif. & Gonz. Frag. 1927, ana-

morphic Glyphium Nitschke ex F.

Lehm. 1886 (H)

Phaeomoniella Crous & W. Gams 2000

(H)

Rhabdospora-like, anamorphic Dolabra C.

Booth & W.P. Ting 1964 (C)

Taeniolella S. Hughes 1958, anamorphic

Glyphium Nitschke ex F. Lehm. 1886

(H)

Subclass Eurotiomycetidae Geiser & Lutzoni

Coryneliales Seaver & Chardon

Coryneliaceae Sacc. ex Berl. & Voglino

Exophiala J.W. Carmich. 1966, anamor-

phic Bicornispora Checa, Barrasa,

M.N. Blanco & A.T. Martínez 1996 (H)

Eurotiales G.W. Martin ex Benny & Kimbr.

Monascaceae J. Schröt.

Basipetospora G.T. Cole & W.B. Kendr.

1968, anamorphic Monascus Tiegh.

1884 (H)

Fraseriella Cif. & A.M. Corte 1957, ana-

morphic Xeromyces Fraser (H)

Trichocomaceae E. Fisch.

Aspergillus P. Micheli ex Link 1809,

anamorphic Dichlaena Durieu & Mont.

1849, Emericella Berk. & Broome

1857, Eurotium Link 1809, Fennellia

B.J. Wiley & E.G. Simmons 1973,

Hemicarpenteles A.K. Sarbhoy &

Elphick 1968, Neopetromyces Frisvad

& Samson 2000, Neosartorya Malloch

& Cain 1973, Petromyces Malloch &

Cain 1973 and Warcupiella Subram.

1972 (H)*

Geosmithia J.Pitt 1979, anamorphic, Chro-

mocleista Yaguchi & Udagawa 1993

and Talaromyces C.R. Benj. 1955 (H)

Isaria Pers. 1794, anamorphic Byssochla-

mys Westling (H)

Mycosphere Doi 10.5943/mycosphere/3/2/5

173

Merimbla Pitt 1979, anamorphic Talar-

omyces C.R. Benj. 1955 (H)

Paecilomyces Bainier 1907, anamorphic

Byssochlamys Westling 1909 and Tala-

romyces C.R. Benj. 1955 (H)

Paecilomyces-like, anamorphic Cooneme-

ria Mouch. 1997 (H)

Penicillium Link 1809, anamorphic Eupe-

nicillium F. Ludw. 1892, Talaromyces

C.R. Benj. 1955, Dichlaena Durieu &

Mont. 1849 and Trichocoma Jungh.

1838 (H)

Phialosimplex Sigler, Deanna A. Sutton,

Gibas, Summerb. & Iwen 2010 (H)

Polypaecilum G. Sm. 1961, anamorphic

Dactylomyces Sopp 1912, Dichotomo-

myces Saito ex D.B.Scott 1970 and

Thermoascus Miehe 1907 (H)

Pseudocordyceps Hauman 1936, anamor-

phic Penicilliopsis Solms 1887 (H)

Sagenomella W. Gams 1978, anamorphic

Sagenoma Stolk & G.F. Orr 1974 and

Talaromyces C.R. Benj. 1955 (H)

Sarophorum Syd. & P. Syd. 1916, ana-

morphic Penicilliopsis Solms 1887 (H)

Stilbodendron Syd. & P. Syd. 1916, ana-

morphic Penicilliopsis Solms 1887 (H)

Stilbothamnium Henn. 1896 (H)

Thermomyces Tsikl. 1899 (H)

Thysanophora W.B. Kendr. 1961 (H)

Torulomyces Delitsch 1943, anamorphic

Eupenicillium F. Ludw. 1892 (H)

Eurotiomycetidae genera incertae sedis

Penicillium Link 1809, anamorphic Para-

talaromyces Matsush. 2003 (H)

Onygenales Cif. ex Benny & Kimbr.

Ajellomycetaceae Untereiner, J.A. Scott &

Sigler

Blastomyces Gilchrist & W.R. Stokes

1898, anamorphic Ajellomyces McDo-

nough & A.L. Lewis 1968 (H)

Chrysosporium Corda 1833, anamorphic

Ajellomyces McDonough & A.L. Lewis

1968 (H)

Emmonsia Cif. & Montemart. 1959, ana-

morphic Ajellomyces McDonough &

A.L. Lewis 1968 (H)

Histoplasma Darling 1906, anamorphic

Ajellomyces McDonough & A.L. Lewis

1968 (H)

Lacazia Taborda, V.A. Taborda & Mc-

Ginnis 1999 (H)

Paracoccidioides F.P. Almeida 1930 (H)

Arachnomycetaceae Gibas, Sigler & Currah

Onychocola Sigler 1990, anamorphic

Arachnomyces Massee & E.S. Salmon

1902 (H)

Gymnoascaceae Baran.

Chrysosporium-like, anamorphic Orromy-

ces B. Sur & G.R. Ghosh 1987 (H)

Malbranchea Sacc. 1882, anamorphic

Byssoonygena Guarro, Punsola & Cano

1987 and Gymnoascus Baran. 1872 (H)

Oncocladium Wallr. 1833, anamorphic

Gymnoascus Baran. 1872 (H)

Onygenaceae Berk.

?Arthrographis G. Cochet ex Sigler &

J.W. Carmich. 1976, anamorphic Ara-

chnotheca Arx 1971 (H)

Chrysosporium Corda 1833 anamorphic

Amauroascus J. Schröt., Aphanoascus

Zukal 1890, Apinisia La Touche 1968,

Arachnotheca Arx 1971, Nannizziopsis

Currah 1985, ?Polytolypa J.A. Scott &

Malloch 1993, Pectinotrichum Varsav-

sky & G.F. Orr 1971, ?Renispora Sigler

& J.W. Carmich. 1979 (H)

Coccidioides G.W. Stiles 1896 (H)

Malbranchea Sacc. 1882, anamorphic Au-

xarthron G.F. Orr & Kuehn 1963,

Byssoonygena Guarro, Punsola & Cano

1987 and Uncinocarpus Sigler & G.F.

Orr 1976 (H)

Myriodontium Samson & Polon. 1978,

anamorphic Neoarachnotheca Ulfig,

Cano & Guarro 1997 (H)

Sporendonema Desm. 1827, anamorphic

Onygena Pers. 1800 (H)

Arthrodermataceae Currah

Chrysosporium Corda 1833, anamorphic

Shanorella R.K. Benj. 1956, Arthr-

oderma Curr. 1860 and Ctenomyces

Eidam 1880 (H)

Chrysosporium Corda 1833, anamorphic

Epidermophyton Sabour. 1907 (H)

Microsporum Gruby 1843, anamorphic

Nannizzia Stockdale 1961 and Arthrod-

erma Curr. 1860 (H)

Mycosphere Doi 10.5943/mycosphere/3/2/5

174

Myceliophthora Costantin 1892, anamo-

rphic Arthroderma Curr. 1860

Trichophyton Malmsten 1848, anamorphic

Arthroderma Curr. 1860 (H)

Ascosphaeraceae L.S. Olive & Spiltoir

Chrysosporium-like, anamorphic Asco-

sphaera L.S. Olive & Spiltoir 1955 and

Bettsia Skou 1972 (H)

Onygenales, genera incertae sedis

Arthropsis Sigler, M.T. Dunn & J.W.

Carmich. 1982 (H)

Lacazia Taborda, V.A. Taborda & McG-

innis 1999 (H)

Subclass Mycocaliciomycetidae Tibell

Mycocaliciales Tibell & Wedin

Mycocaliciaceae A.F.W. Schmid

Asterophoma D. Hawksw. 1981, anamor-

phic Chaenothecopsis Vain. 1927 (C)

Catenomycopsis Tibell & Constant. 1991,

anamorphic Chaenothecopsis Vain.

1927 (H)

Phialophora-like, anamorphic Chaenothe-

copsis Vain. 1927 and Mycocalicium

Vain. 1890 (H)

Class Laboulbeniomycetes Engler

Laboulbeniales Engler

Coreomycetopsis Thaxt. 1920 (H)

Muiaria Thaxt. 1914 (H)

Tetrameronycha Speg. ex W. Rossi & M.

Blackw. 1990 (H)

Pyxidiophorales P.F. Cannon

Pyxidiophoraceae Arnold

Chalara (Corda) Rabenh. 1844 anam-

orphic Pyxidiophora Bref. & Tavel

1891 (H)*

Gliocephalis Matr. 1899 (H)

Pleurocatena G. Arnaud ex Aramb.,

Gamundí, W. Gams & G.R.W. Arnold

2007, anamorphic Pyxidiophora Bref.

& Tavel 1891 (H)

Thaxteriola Speg. 1918, anamorphic Pyxi-

diophora Bref. & Tavel 1891 (H)

Class Lecanoromycetes O.E. Erikss. & Winka

Subclass Ostropomycetidae Reeb, Lutzoni &

Cl. Roux

Agyriales Clem. & Shear

Trapeliaceae M. Choisy ex Hertel

Epithyrium (Sacc.) Trotter 1931, anamo-

rphic Sarea Fr. 1825 (C)

Pycnidiella Höhn. 1915, anamorphic Sarea

Fr. 1825 (C)

Ostropales Nannf.

Odontotremataceae D. Hawksw. & Sherwood

Pycnidial anamorph associated with

Claviradulomyces P.R. Johnst., D.C.

Park, H.C. Evans, R.W. Barreto & D.J.

Soares 2010

Stictidaceae Fr.

?Coleophoma-like, anamorphic Stictopha-

cidium Rehm 1888 (C)

?Ebollia Minter & Caine 1980, anamor-

phic Stictophacidium Rehm 1888 (C)

Phacidiella-like, anamorphic Acaros-

porina Sherwood 1977 (C)

Stictospora Cif. 1957, anamorphic Stictis

Pers. 1800 (C)

Ostropales, genera incertae sedis

Glomerobolus Kohlm. & Volkm.-Kohlm.

1996 (H)

Subclass Lecanoromycetidae P.M. Kirk, P.F.

Cannon, J.C. David & Stalpers ex Miadl.,

Lutzoni & Lumbsch

Lecanorales Nannf.

Pilocarpaceae Zahlbr.

Pyriomyces Bat. & H. Maia 1965,

anamorphic Byssoloma Trevis. 1853

(C)

Szczawinskia A. Funk 1984 (C)

Stereocaulaceae Chevall.

Lepraria Ach. 1803 (C)

Lecanorales, genera incertae sedis

Leprocaulon Nyl. 1878 (C)

Vouauxiomyces Dyko & D. Hawks. 1979,

anamorphic Abrothallus De Not. 1845

(C)

Class Leotiomycetes O.E. Erikss. & Winka

Cyttariales Luttr. ex Gamundı´

Cyttariaceae Speg.

Cytariella Palm 1932, anamorphic Cytta-

Mycosphere Doi 10.5943/mycosphere/3/2/5

175

ria Berk. 1842 (C)

Erysiphales Gwynne-Vaughan

Erysiphaceae Tul. & C. Tul.

Oidiopsis Scalia 1902, anamorphic Leveil-

lula G. Arnaud 1921 (H)

Oidium Link 1824, anamorphic Arthro-

cladiella Vassilkov 1960, Blumeria

Golovin ex Speer 1975, Brasiliomyces

Viégas 1944, Cystotheca Berk. & M.A.

Curtis 1860, Erysiphe R. Hedw. ex DC.

1805, Golovinomyces (U. Braun) V.P.

Gelyuta 1988, Medusosphaera Golovin

& Gamalizk. 1962, Microsphaera Lév.

1851, U. Braun 1999, Podosphaera

Kunze 1823, Sawadaea Miyabe 1914,

Sphaerotheca Lév. 1851, Uncinula Lév.

1851 and Uncinuliella R.Y. Zheng &

G.Q. Chen 1979 (H)

Ovulariopsis Pat. & Har. 1900, ana-

morphic Phyllactinia Lév. 1851 (H)

Streptopodium R.Y. Zheng & G.Q. Chen

1978, anamorphic Pleochaeta Sacc. &

Speg. 1881 and Phyllactinia Lév. 1851

(H)

Helotiales Nannf.

Bulgariaceae Fr.

Crinula Fr. 1821, anamorphic Holwaya

Sacc. 1889 (H)

Endomelaconium Petr. 1940, anamorphic

Bulgaria Fr. 1822 (C)

Dermateaceae Fr.

Anguillospora Ingold 1942, anamorphic

Mollisia (Fr.) P. Karst. 1871 (H)

Cadophora Lagerb. & Melin 1927, anam-

orphic Mollisia (Fr.) P. Karst. 1871,

Pyrenopeziza Fuckel 1870 and Pseud-

opezicula Korf 1975 (H)

Cadophora-like, anamorphic Phaeomol-

lisia T.N. Sieber & Grünig 2009 (H)

Casaresia Gonz. Frag. 1920, anamorphic

Mollisia (Fr.) P. Karst. 1871 (H)

Chondropodium-like, anamorphic Walto-

nia Saho 1970 (C)

Corniculariella P. Karst. 1884, anamor-

phic Dermea Fr. 1825 and Durandiella

Seaver 1932 (C)

Cryptocline Petr. 1924, anamorphic

Trochila Fr. 1849 (C)

Cryptosporiopsis Bubák & Kabát 1912,

anamorphic Neofabraea H.S. Jacks.

1913, Pezicula Tul. & C. Tul. 1865 and

?Ocellaria (Tul. & C. Tul.) P. Karst.

1871 (C)

Cryptosympodula Verkley 1999, anamor-

phic Scleropezicula Verkley 1999 (C)

Cylindrocolla Bonord. 1851, anamorphic

Calloria Fr. 1836 (H)

Cylindrosporium Grev. 1822, anamorphic

Pyrenopeziza Fuckel 1870 (C)

Cystodendron Bubák 1914, anamorphic

Mollisia (Fr.) P. Karst. 1871 Belonopsis

(Sacc.) Rehm 1891 and Niptera Fr.

1849 (H)

Diplosporonema Höhn. 1917, anamorphic

Pyrenopeziza Fuckel 1917 (C)

Discogloeum Petr. 1923, anamorphic

Spilopodia Boud. 1885 (C)

Entomosporium Lév. 1856, anamorphic

Diplocarpon F.A. Wolf 1912 (C)

Eriosporella Höhn. 1916, possibly ana-

morphic Laetinaevia Nannf. 1932 (C)

Foveostroma DiCosmo 1978, anamorphic

Dermea Fr. 1825 (C)

Gelatinosporium Peck 1873, anamorphic

Dermea Fr. 1825 and Durandiella

Seaver 1932 (C)

Glarea Bills & Paláez 1999 (H)

Gloeosporidiella Petr. 1921, anamorphic

Drepanopeziza (Kleb.) Höhn. 1917 and

Pseudopeziza Fuckel 1870 (C)

Hainesia Ellis & Sacc. 1884, anamorphic

Discohainesia Nannf. 1932 (C)

Helicodendron Peyronel 1918, anamorphic

Mollisia (Fr.) P. Karst. 1871 (H)

Laetinaevia Nannf. 1932, anamorphic

Naeviopsis B. Hein 1976 (C)

Marssonina Magnus 1906, anamorphic

Diplocarpon F.A. Wolf 1912 &

Drepanopeziza (Kleb.) Höhn. 1917 (C)

Melanodiscus Höhn. 1918, anamorphic

Spilopodia Boud. 1885 (H)

Microgloeum Petr. 1922, anamorphic

Blumeriella Arx 1961 (C)

Monostichella Höhn. 1916, anamorphic

Drepanopeziza (Kleb.) Höhn. 1917 (C)

Myrioconium Syd. & P. Syd. 1912, ana-

morphic Dibeloniella Nannf. 1932 (H)

Phacidiella P. Karst. 1884, anamorphic

Pyrenopeziza Fuckel 1870 (C)

Phialocephala W.B. Kendr. 1961

anamorphic Phaeomollisia T.N. Sieber

& Grünig 2009 (H)

Mycosphere Doi 10.5943/mycosphere/3/2/5

176

Phialocephala-like, anamorphic Phaeo-

mollisia T.N. Sieber & Grünig 2009

(H)

Phialophora-like, anamorphic Dibeloniel-

la Nannf. 1932 (H)

Phloeosporella Höhn. 1924, anamorphic

Blumeriella Arx 1961 (C)

Phlyctema Desm. 1847, anamorphic Neo-

fabraea H.S. Jacks. 1913 (C)

Pilidium Kunze 1823, anamorphic Disco-

hainesia Nannf. 1932 (C)

Rhynchosporium Heinsen ex A.B. Frank

1897 (H)*

?Sporonema Desm. 1847, anamorphic

Leptotrochila P. Karst. 1871 (C)

Trichosporiella Kamyschko 1960, ana-

morphic Laetinaevia Nannf. 1932 (H)

Trimmatostroma Corda 1837 (H)

Xenochalara M.J. Wingf. & Crous 2000

(H)

Helotiaceae Rehm

Alatospora Ingold 1942 (H)

Anguillospora Ingold 1942, anamorphic

Hymenoscyphus Gray 1821 (H)

Articulospora Ingold 1942, anamorphic

Hymenoscyphus Gray 1821(H)

Ascoconidium Seaver 1942, anamorphic

Sageria A. Funk 1975 (H)

Aureohyphozyma Hosoya & Y. Otani 1995

anamorph of Gelatinipulvinella Hosoya

& Y. Otani (Hosoay Y Otani 1995) (H)

Bloxamia Berk. & Broome 1854, anamor-

phic Bisporella Sacc. 1884 (H)

Bothrodiscus Shear 1907, anamorphic

Ascocalyx Naumov 1926 (C)

Brunchorstia Erikss. 1891, anamorphic

Gremmeniella M. Morelet 1969 (C)

?Bryophytomyces Cif. 1953, anamorphic

Hymenoscyphus Gray 1821 (H)

Cercosporella Sacc. 1880, anamorphic

Heterosphaeria Grev. 1824 (H)

Coryne Nees 1816, anamorphic Asco-

coryne J.W. Groves & D.E. Wilson

1967 (H)

Cystodendron Bubák 1914, anamorphic

Bisporella Sacc. 1884 (H)

?Cystotricha Berk. & Broome 1850, ana-

morphic Xylogramma Wallr. 1833 (C)

Deltosperma W.Y. Zhuang 1988, ana-

morphic Unguiculariopsis Rehm 1909

(C)

Dendrostilbella Höhn. 1905, anamorphic

Claussenomyces Kirschst. 1923 (H)

Dichaenopsella Petr. 1952, type is

anamorphic Godroniopsis Diehl & E.K.

Cash 1929 (C)

Digitosporium Gremmen 1953, anam-

orphic Crumenulopsis J.W. Groves

1969 (C)

Dimorphospora Tubaki 1958, anamorphic

Hymenoscyphus Gray 1821 (H)

Dothiorina Höhn. 1911, anamorphic Chlo-

rociboria Seaver ex Ramamurthi, Korf

& L.R. Batra 1958 (C)

Endomelaconium Petr. 1940, anamorphic

Austrocenangium Gamundí 1997 (C)

Eriomycopsis Speg. 1910, anamorphic

Calloriopsis Syd. & P. Syd. 1917 (H)

Eustilbum Rabenh. 1864, anamorphic Bis-

porella Sacc. 1884 (H)

Fuckelia Bonord. 1864, anamorphic God-

ronia Moug. & Lév. 1846 (C)

Geniculospora Sv. Nilsson ex Marvanová

& Sv. Nilsson 1971, anamorphic Hyme-

noscyphus Gray 1821 (H)

Gliomastix Guég. 1905 anamorphic

Ascoclavulina Y. Otani 1974 (H)*

Heteropatella Fuckel 1874, anamorphic

Heterosphaeria Grev. 1824 (C)

Hormonema-like, anamorphic Pragmopo-

ra A. Massal 1855 (H)

Idriella P.E. Nelson & S. Wilh. 1956,

anamorphic Hymenoscyphus Gray 1821

(H)

Linodochium Höhn. 1909, anamorphic

Pseudohelotium Fuckel 1870 (H)

Monodictys S. Hughes 1958 anamorphic

Hymenoscyphus Gray 1821 (H)

Periperidium Darker 1963, anamorphic

Micraspis Darker 1963 (C)

Phialophora-like, anamorphic Pseudopezi-

cula Korf 1975 (H)

Pragmopycnis B. Sutton & A. Funk 1975,

anamorphic Pragmopora A. Massal

1855 (C)

?Pseudaegerita J.L. Crane & Schokn.

1981, anamorphic Claussenomyces Kir-

schst. 1923 (H)

Pseudospiropes M.B. Ellis 1971, ana-

morphic Strossmayeria Schulzer 1881

(H)

Mycosphere Doi 10.5943/mycosphere/3/2/5

177

Rhinocladiella-like, anamorphic Bioscy-

pha Syd. 1927 (H)

Rhizothyrium Naumov 1915, anamorphic

Rhizocalyx Petr. 1928 (C)

Scytalidium Pesante 1957 anamorphic

Hymenoscyphus Gray 1821 (H)

Sirodothis Clem. 1909, anamorphic Tym-

panis Tode 1790 (C)

Sporonema Desm. 1847, anamorphic God-

ronia Moug. & Lév. 1846 (C)

Symphyosirella Seifert 2009, anamorphic

Symphyosirinia E.A. Ellis 1956 (H)

Titaeospora Bubák 1916, anamorphic

Stamnaria Fuckel 1870 (C)

Topospora Fr. 1836, anamorphic Godr-

onia Moug. & Lév. 1846 (C)

Tricladium Ingold 1942, anamorphic

Cudoniella Sacc. 1889 and Hymeno-

scyphus Gray 1821 (H)

Varicosporium W. Kegel 1906, anamor-

phic Hymenoscyphus Gray 1821 (H)

Hemiphacidiaceae Korf

Meria Vuill. 1896, anamorphic Rhabdo-

cline Syd. 1922 (H)

Rhabdogloeum Syd. 1922, anamorphic

Rhabdocline Syd. 1922 (C)

Rhabdogloeopsis Petr. 1925, anamorphic

Sarcotrochila Höhn. 1917 (C)

Hyaloscyphaceae Nannf.

Acleistia Bayl. Ell. (1917) anamorphic

Calycina Nees ex Gray 1821 (C)

Brefeldochium Verkley 2005, anamorphic

Polydesmia Boud. 1885 (H)

Chaetochalara B. Sutton & Piroz. 1965

(H)

Chalara (Corda) Rabenh. 1844, anam-

orphic Calycellina Höhn., Phaeos-

cypha Spooner 1984 and Tapesina

Lambotte 1887 (H)*

Cheiromycella Höhn. 1910, anamorphic

Hyaloscypha Boud. 1885 (H)

Clathrosphaerina Beverw. 1951, anamor-

phic Hyaloscypha Boud. 1885 (H)

Haplographium Berk. & Broome 1859,

anamorphic Dematioscypha Svrček

1977 (H)

Mycoarthris Marvanová & P.J. Fisher

2002 (H)

Naemospora Roth ex Kuntze 1898,

possibly anamorphic Lachnellula P.

Karst. 1884 (C)

Phialophora-like, anamorphic Cistella

Quél. and Hyalopeziza Fuckel 1870 (H)

Pseudaegerita J.L. Crane & Schokn. 1981,

anamorphic Hyaloscypha Boud. 1885

(H)

Septonema-like, anamorphic Ciliolarina

Svrcek 1977 (H)

Tricladium Ingold 1942, anamorphic

Hydrocina Scheuer 1991 (H)

Leotiaceae Corda

Anguillospora Ingold 1942, anamorphic

Pezoloma Clem. 1909 (H)

Aureohyphozyma Hosoya & Y. Otani

1995, anamorphic Gelatinipulvinella

Hosoya & Y. Otani 1995 (H)

Halenospora E.B.G. Jones 2009 (H)

Myrioconium-like, anamorphic Neobulga-

ria Petr. 1921 (H)

Phialophora-like, anamorphic Neobulga-

ria Petr. 1921 (H)

Loramycetaceae Dennis ex Digby & Goos

Anguillospora Ingold 1942, anamorphic

Loramyces W. Weston 1929 (H)

Phacidiaceae Fr.

Allantophomopsis Petr. 1925, anamorphic

Phacidium Fr. 1815 (C)

Apostrasseria Nag Raj 1983, anamorphic

Lophophacidium Lagerb. 1949 and

Phacidium Fr. 1815 (C)

Blennoria Moug. & Fr. 1825, anamorphic

Phacidium Fr. 1815 (C)

Ceuthospora Grev. 1826, anamorphic

Phacidium Fr. 1815 (C)

?Coma Nag Raj & W.B. Kendr. 1972,

anamorphic Ascocoma H.J. Swart (C)

Rutstroemiaceae Holst-Jensen, L.M. Kohn &

T. Schumacher

Helicodendron Peyronel 1918, anamorphic

Lambertella Höhn. 1918 (H)

Myrioconium Syd. & P. Syd. 1912, ?ana-

morphic Lanzia Sacc. 1884, Rutstro-

emia P. Karst. 1871, Scleromitrula

Mycosphere Doi 10.5943/mycosphere/3/2/5

178

S.Imai 1941, Lambertella Höhn. 1918

and Poculum Velen. 1934 (H)

Phialophora-like, anamorphic Rutstroemia

P. Karst. 1871 (H)

Sclerotiniaceae Whetzel ex Whetzel

Acarosporium Bubák & Vleugel ex Bubák

1911, anamorphic Pycnopeziza W.L.

White & Whetzel 1938 (C)

Amerosporium Speg. 1882, anamorphic

Zoellneria Velen. 1934 (C)

Amphobotrys Hennebert 1973, anamorphic

Botryotinia Whetzel 1945 (H)

Botrytis P. Micheli ex Pers. 1794,

anamorphic Botryotinia Whetzel 1945

(H)

Cristulariella Höhn. 1916, anamorphic

Nervostroma Narumi & Y. Harada

2006 (H)

Haradamyces Masuya, Kusunoki, Kosaka

& Aikawa 2009 (H)

Hinomyces Narumi & Y. Harada 2006,

anamorphic Grovesinia M.N. Cline,

J.L. Crane & S.D. Cline 1983 (H)

Monilia Bonord. 1851, anamorphic Moni-

linia Honey 1928 and Phaeosclerotinia

Hori 1916 (H)

Mycopappus Redhead & G.P. White 1985,

anamorphic Redheadia Y. Suto &

Suyama 2005 (H)

Myrioconium Syd. & P. Syd. 1912,

anamorphic Ciboria Fuckel 1870,

Ciborinia Whetzel 1945, (Fr.) P. Karst.

1871, Martininia Dumont & Korf 1970,

Myriosclerotinia N.F. Buchw. 1947,

Sclerotinia Fuckel 1870 Stromatinia

(Boud.) Boud. 1907 and Monilinia

Honey 1928 (H)

Ovulitis N.F. Buchw. 1970, anamorphic

Ovulinia F.A. Weiss 1940 (H)

Sclerotium Tode 1790, anamorphic Myrio-

sclerotinia N.F. Buchw. 1947, Sclero-

tinia Fuckel 1870 and Stromatinia

(Boud.) Boud. 1907 (H)

Septotis N.F. Buchw. ex Arx 1970,

anamorphic Septotinia Whetzel ex J.W.

Groves & M.E. Elliott 1961 (H)

Streptobotrys Hennebert 1973, anamorphic

Streptotinia Whetzel 1945 (H)

Valdensia Peyronel 1923, anamorphic

Valdensinia Peyronel 1953 (H)

Verrucobotrys Hennebert 1973, anamor-

phic Seaverinia Whetzel 1945 (H)

Vibrisseaceae Korf

Acephala Grünig & T.N. Sieber 2005 (H)

Anavirga B. Sutton 1975, anamorphic

Vibrissea Fr. 1822 (H)

Phialocephala W.B. Kendr. 1961, some

possibly anamorphic Vibrissea Fr. 1822

(H)

Phialophora-like, anamorphic Chlorovi-

brissea L.M. Kohn 1989 (H)

Helotiales, genera incertae sedis

Catenulifera Hosoya 2002, anamorphic

Hyphodiscus Kirschst. 1906 (H)

Chalara (Corda) Rabenh. 1844 (H)*

Clathrosporium Nawawi & Kuthub. 1987

(H)

Crucellisporium M.L. Farr 1968 (C)

Cystotricha Berk. & Broome 1850 (C)

Dactylaria Sacc. 1880 (H)

Diplococcium Grove 1885 (H)

Endoconidium Prill. & Delacr. 1891, ana-

morphic Gloeotinia M. Wilson, Noble

& E.G. Gray 1954 (H)

Fontanospora Dyko 1978 (H)

Fulvoflamma Crous 2006 (H)

Glarea Bills & Paláez 1999 (H)

Helgardia Crous & W. Gams 2003, ana-

morphic Oculimacula Crous & W.

Gams 2003 (H)

Humicolopsis Cabral & S. Marchand 1976

(H)

Hyalodendriella Crous 2007 (H)

Hysteropezizella Höhn. 1917 (C)

Lemonniera De Wild. 1894 (H)

Leptodontidium de Hoog 1979 (H)

Libartania Nag Raj 1979 (C)

Margaritispora Ingold 1942 (H)

Meliniomyces Hambl. & Sigler 2005 (H)

Mirandina G. Arnaud ex Matsush. 1975

(H)

Monochaetiellopsis B. Sutton & DiCosmo

1977, anamorphic Hypnotheca Tom-

merup 1970 (C)

Mycochaetophora Hara & Ogawa 1931

(H)

Neozythia Petr. 1958 (C)

Rhexocercosporidium U. Braun 1994 (H)

Rhizocladosporium Crous & U. Braun

Mycosphere Doi 10.5943/mycosphere/3/2/5

179

2007 (H)

Rhodesia Grove 1937 (C)

Scytalidium Pesante 1957 (H)

Sphaerographium Sacc. 1884 (C)

Spirosphaera Beverw. 1953 (H)

Tetracladium De Wild. 1893 (H)

Thedgonia B. Sutton 1973 (H)

Trimmatostroma Corda 1837 (H)

Variocladium Descals & Marvanová 1999

(H)

Xenopolyscytalum Crous 2011(‘2010’) (H)

Rhytismatales M.E. Barr ex Minter

Ascodichaenaceae D. Hawksw. & Sherwood

Dichaenopsella Petr. 1952, anamorphic

Ascodichaena Butin 1977 (C)

Macroallantina Speer 1987, anamorphic

Delpinoina Kuntze, 1891 (C)

Polymorphum Chevall. 1822, anamorphic

Ascodichaena Butin 1977 (C)

Cryptomycetaceae Höhn.

?Phacidiopycnis Potebnia 1912, anamor-

phic Potebniamyces Smerlis 1962 (C)

Rhytismataceae Chevall.

Conostroma Moesz 1921, anamorphic

Colpoma Wallr. 1833 (C)

Crandallia Ellis & Sacc. 1897, ana-

morphic Bifusella Höhn. 1917 and

Duplicaria Fuckel 1870 (C)

Cryocaligula Minter 1986, anamorphic

Ploioderma Darker. 1967 (C)

Hysterodiscula Petr. 1942, anamorphic

Duplicaria Fuckel 1870 (C)

Leptostroma Fr. 1815, anamorphic Lopho-

dermium Chevall. 1826 (C)

Melasmia Lév. 1846, anamorphic Rhytis-

ma Fr. 1818 (C)

Tryblidiopycnis Höhn. 1918, anamorphic

Tryblidiopsis P. Karst. 1871 (C)

Rhytismatales, genera incertae sedis

Fuligomyces Morgan-Jones & Kamal 1984

(H)

Uyucamyces H.C. Evans & Minter 1985,

anamorphic Ocotomyces H.C. Evans &

Minter 1985 (C)

Leotiomycetes, families incertae sedis

Myxotrichaceae Currah

Geomyces Traaen 1914, anamorphic

Pseudogymnoascus Raillo 1929 (H)

?Gymnostellatospora Udagawa, Uchiy. &

Kamiya 1993, anamorphic Pseudo-

gymnoascus Raillo 1929 (C)

?Malbranchea Sacc. 1882, anamorphic

Myxotrichum Kunze 1823 (H)

Oidiodendron Robak 1932, anamorphic

Byssoascus Arx 1971 and Myxotrichum

Kunze 1823 (H)

Leotiomycetes, genera incertae sedis

Brachyalara Réblová & W. Gams 2011

(H)

Chaetomella Fuckel 1870 (C)

Collophora Damm & Crous 2010 (C)

Eleutheromyces Fuckel 1870 (C)

Geniculospora Sv. Nilsson ex Marvanová

& Sv. Nilsson 1971 (H)

Hyphozyma de Hoog & M.T. Sm. 1981

(H)

Infundichalara Réblová & W. Gams 2011

(H)

Leohumicola N.L. Nick. 2005 (H)

Meliniomyces Hambl. & Sigler 2005 (H)

Scytalidium Pesante 1957, anamorphic

Xylogone Arx & T. Nilsson 1969 (H)

Tetrachaetum Ingold 1942 (H)

Tiarosporella Höhn. 1924, anamorphic

Darkera H.S. Whitney, J. Reid & Piroz.

1975 (C)

Class Orbiliomycetes O.E. Erikss. & Baral

Orbiliales Baral, O.E. Erikss., G. Marson & E.

Weber

Orbiliaceae Nannf.

Anguillospora Ingold 1942, anamorphic

anamorphic Orbilia Fr. 1836 (H)

Arthrobotrys Corda 1939, anamorphic

Orbilia Fr. 1836 (H)

Brachyphoris Juan Chen, L.L. Xu, B. Liu

& Xing Z. Liu 2007, anamorphic

Hyalorbilia Baral & G. Marson 2000

(H)

Dactylaria Sacc. 1880, anamorphic Or-

bilia Fr. 1836 (H)

Dactylella Grove 1884, anamorphic Orbi-

lia Fr. 1836 (H)

Dactylellina M. Morelet 1968 (H)

Dicranidion Harkn. 1885, anamorphic Or-

bilia Fr. 1836 (H)

Mycosphere Doi 10.5943/mycosphere/3/2/5

180

Drechslerella Subram. 1964, anamorphic

Orbilia Fr. 1836 (H)

Dwayaangam Subram. 1978, anamorphic

Orbilia Fr. 1836 (H)

Gamsylella M. Scholler, Hagedorn & A.

Rubner 1999, anamorphic Orbilia Fr.

1836 (H)

Helicoön Morgan 1892, anamorphic

Orbilia Fr. 1836 (H)

Idriella P.E. Nelson & S. Wilh. 1956,

anamorphic Orbilia Fr. 1836 (H)

Monacrosporium Oudem. 1885, anamor-

phic Orbilia Fr. 1836 (H)

Pseudotripoconidium Z.F. Yu & K.Q.

Zhang 2011, anamorphic Orbilia Fr.

1836 (H)*

Tricellula Beverw. 1954, anamorphic

Orbilia Fr. 1836 (H)

Tridentaria Preuss 1852 (H)

Trinacrium Riess 1852, anamorphic Orbi-

lia Fr. 1836 (H)

Class Pezizomycetes sensu O.E. Erikss. &

Winka

Pezizales J. Schröt.

Ascobolaceae Boud. ex Sacc.

Papulospora Preuss 1851, anamorphic

Ascobolus Pers. 1796 (H)

Papulospora-like, anamorphic Cubonia

Sacc. 1889 (H)

Rhizostilbella Wolk 1914, anamorphic

Ascobolus Pers. 1792 (H)

Caloscyphaceae Harmaja

Geniculodendron G.A. Salt 1974, anamor-

phic Caloscypha Boud. 1885 (H)

Chorioactidaceae Pfister

Kumanasamuha P.Rag. Rao & D. Rao

1964, anamorphic Chorioactis Kupfer

ex Eckblad 1968 (H)

Verticicladium Preuss 1851, anamorphic

Desmazierella Lib. 1829 (H)

Morchellaceae Reichenb.

Costantinella Matr. 1892, anamorphic

Morchella Dill. ex Pers. 1794 (H)

Pezizaceae Dumort.

Chromelosporium Corda 1833, anamor-

phic Muciturbo P.H.B. Talbot 1989,

Peziza Dill. ex Fr. 1822 and Plicaria

Fuckel 1870 (H)

Glischroderma Fuckel 1870 (C)

Oedocephalum Preuss 1851, anamorphic

Iodophanus Korf 1967 and Peziza Dill.

ex Fr. 1822 (H)

Ostracoderma Fr. 1825, anamorphic Pezi-

za Dill. ex Fr. 1822 (C)

Pyronemataceae Corda

Actinosporella Descals, Marvanová & J.

Webster 1999, anamorphic Miladina

Svrček 1972 (H)

Ascorhizoctonia Chin S. Yang & Korf

1985, anamorphic Tricharina Eckblad

1968 (H)

Basifimbria Subram. & Lodha 1968,

anamorphic Geopyxis (Pers.) Sacc. (H)

Complexipes C. Walker 1979, anamorphic

Tricharina Eckblad 1968 and Wilcoxina

Chin S. Yang & Korf 1985 (H)

?Dichobotrys Hennebert 1973, anamorphic

Pyropyxis Egger 1984 and Trichophaea

Boud. 1885 (H)

Micronematobotrys Xiang Sun & L.D.

Guo 2010 (H)

?Molliardiomyces Paden 1984, anam-

orphic Stephensia Tul. & C. Tul. 1851

(H)

?Scytalidium Pesante 1957, anamorph of

Anthracobia Boud. 1885 (H)

Rhizinaceae Bonord.

Phymatotrichopsis Hennebert 1973 (H)

Sarcoscyphaceae LeGal ex Eckblad

Molliardiomyces Paden 1984, anamorphic

Nanoscypha Denison 1972, Phillipsia

Berk. 1881, Pithya Fuckel 1870 and

Sarcoscypha (Fr.) Boud. 1885 (H)

Verticicladium Preuss 1851, anamorphic

Desmazierella Lib. 1829 (H)

Sarcosomataceae Kobayasi

Conoplea Pers. 1797, anamorphic Urnula

Fr. 1849 and Plectania Fuckel 1870 (H)

Strumella Fr. 1849, anamorphic Urnula Fr.

1849 (H)

Verticicladium Preuss 1851, anamorphic

Mycosphere Doi 10.5943/mycosphere/3/2/5

181

Sarcosoma Casp. 1891 (H)

Pezizales, genera incertae sedis

Cephaliophora Thaxt. 1903 (H)

Class Sordariomycetes sensu O.E. Erikss. &

Winka

Subclass Hypocreomycetidae O.E. Erikss. &

Winka

Coronophorales Nannf.

Chaetosphaerellaceae Huhndorf, A.N. Mill. &

F.A. Fernández

Oedemium Link 1824, anamorphic Chae-

tosphaerella E. Müll. & C. Booth 1972

(H)

Veramycina Subram. 1995, anamorphic

Chaetosphaerella E. Müll. & C. Booth

1972 (H)

Nitschkiaceae (Fitzp.) Nannf.

Acremonium-like, anamorphic Acantho-

nitschkea Speg. (H)

Glomerellales Chadef. ex Réblová, W. Gams

& Seifert

Australiascaceae Réblová & W. Gams

Monilochaetes Halst. ex Harter 1916,

anamorphic Australiasca Sivan. &

Alcorn 2002 (H)*

Glomerellaceae Locq. ex Seifert & W. Gams

Colletotrichum Corda 1831, anamorphic

Glomerella Spauld. & Schrenk 1903

(C)

Reticulascaceae Réblová & W. Gams

Cylindrotrichum Bonord.1851, anamorph-

ic Reticulascus Réblová & W. Gams

2011 (H)*

Hypocreales Lindau

Bionectriaceae Samuels & Rossman

Acremonium-like, anamorphic Dimero-

sporiella Speg. 1908, Ijuhya Starbäck

1899, Hydropisphaera Dumont 1822,

Lasionectria (Sacc.) Cooke 1884, My-

coarachis Malloch & Cain 1970,

Mycocitrus Moller 1901, Nectriella

Nitschke ex Fuckel 1870, Nectriopsis

Maire 1911, Nigrosabulum Malloch &

Cain 1970, Protocreopsis Yoshim. Doi

1977, Selinia P. Karst. 1876, and

Stilbocrea Pat. 1900 and Verrucostoma

Hirooka, Tak. Kobay. & Chaverri 2010

(H)

Albosynnema E.F. Morris 1967 (H)

Clonostachys Corda 1839, anamorphic

Bionectria Speg. 1919 (H)

Dendrodochium-like, anamorphic Nec-

triella Nitschke ex Fuckel 1870 (C)

Didymostilbe Henn. 1902, anamorphic

Peethambara Subram. & Bhat 1978 (H)

Geosmithia J.Pitt 1979 (H)*

Gliocladium-like, anamorphic Roumegue-

riella Speg. 1880 (H)

Gliomastix Guég. 1905 (H)*

Gracilistilbella Seifert 2000, anamorphic

Stilbocrea Pat. 1900 (H)

Kutilakesa Subram. 1956, anamorphic

Nectriella Nitschke ex Fuckel 1870 (H)

Rhopalocladium Schroers, Samuels & W.

Gams 1999, anamorphic Nectriopsis

Maire 1911 (H)

Sarocladium W. Gams & D. Hawksw.

1976 (H)

Spicellum Nicot & Roquebert 1976 (H)

Stanjemonium W. Gams, O'Donnell,

Schroers & M. Chr. 1999 (H)

Stilbella Lindau 1900 anamorphic

Stilbocrea Pat. 1900 (H)

Trichothecium-like, anamorphic Heleoc-

occum C.A. Jørg. 1922 (H)

Tubercularia-like, anamorphic Selinia P.

Karst. 1876 (H)

Verticillium-like, anamorphic Nectriopsis

Maire 1911 (H)

Vesicladiella Crous & M.J. Wingf. 1994

(H)

Virgatospora Finley 1967, anamorphic

Peethambara Subram. & Bhat 1978 (H)

Clavicipitaceae (Lindau) Earle ex Rogerson

Acremonium-like, anamorphic Neobarya

Lowen 1986 (H)

Albomyces I. Miyake 1908, anamorphic

Aciculosporium I. Miyake (C)

Amphichordia Fr., 1825 (H)

Aschersonia Mont. 1848, anamorphic

Hypocrella Sacc. and Moelleriella Bres.

1897 (C)

Blistum B. Sutton 1973, anamorphic

Berkelella (Sacc.) Sacc. 1891 (H)

Calcarisporium Preuss 1851, anamorphic

Neobarya Lowen (H)

Corallocytostroma Y.N. Yu & Z.Y. Zhang

Mycosphere Doi 10.5943/mycosphere/3/2/5

182

1980 (C)

Diploöspora Grove 1916, anamorphic

Neobarya Lowen 1986 (H)

Drechmeria W. Gams & H.-B. Jansson

1985 (H)

Ephelis Fr. 1849, anamorphic Atkinsonella

Diehl, Balansia Speg., 1885, Epichloë

(Fr.) Tul. & C. Tul., 1865, Dothichloë

G.F. Atk. 1894, Heteroepichloë E.

Tanaka, C. Tanaka, Gafur & Tsuda

2002, Nigrocornus Ryley & Langdon

2003 and Myriogenospora G.F. Atk.

1894 (H)

Harposporium Lohde 1874, anamorphic

Podocrella Seaver 1928 (H)

Hirsutella-like, anamorphic Orbiocrella D.

Johnson, G.H. Sung, Hywel-Jones &

Spatafora 2009 and Moelleriella Bres.

1897 (H)

Lecanicillium W. Gams & Zare 2001,

anamorphic Hypocrella Sacc. 1878and

Neobarya Lowen 1986 (H)

Metarhizium Sorokīn 1879, anamorphic

Metacordyceps G.H. Sung, J.M. Sung,

Hywel-Jones & Spatafora 2007 (H)

?Munkia Speg. 1886, anamorphic Myc-

omalus A. Møller 1901 (H)

Neomunkia Petr. 1947 (H)

Neotyphodium Glenn, C.W. Bacon &

Hanlin 1996, anamorphic Epichloë (Fr.)

Tul. & C. Tul. 1865 (H)

Nomuraea Maubl. 1903 (H)

Pleurodesmospora Samson, W. Gams &

H.C. Evans 1979 (H)

Pochonia Bat. & O.M. Fonseca 1965 (H)

Polycephalomyces Kobayasi 1941, ana-

morphic Berkelella (Sacc.) Sacc. 1891

(H)

Pseudogibellula Samson & H.C. Evans

1973, anamorphic Torrubiella Boud.

1885 (H)

Pseudomeria G.L. Barron 1980 (H)

Rotiferophthora G.L. Barron 1991 (H)

Sphacelia Lév. 1827, anamorphic Atkin-

sonella Diehl 1950 and Claviceps Tul.

1853 (H)

Ustilaginoidea Bref. 1895, anamorphic

Villosiclava E. Tanaka & C. Tanaka

2009 (H)

Cordycipitaceae Kreisel ex G.M. Sung, J.M.

Sung, Hywel-Jones & Spatafora

Akanthomyces Lebert 1858, anamorphic

Cordyceps Fr. 1824 and Torrubiella

Boud. 1885 (H)

Beauveria Vuill. 1912, anamorphic Cordy-

ceps Fr. 1824 (H)*

Engyodontium de Hoog 1978 (H)

Gibellula Cavara 1894, anamorphic

Cordyceps Fr. 1824 and Torrubiella

Boud. 1885 (H)

Granulomanus de Hoog & Samson 1978,

anamorphic Torrubiella Boud. 1885

(H)

Hirsutella Pat. 1892, anamorphic

Cordyceps Fr. 1824 and Cordyce-

pioideus Stifler 1941 (H)

Isaria Pers. 1794, anamorphic Cordyceps

Fr. 1824 and Torrubiella Boud. 1885

(H)

Lecanicillium W. Gams & Zare 2001,

anamorphic Cordyceps Fr. 1824 and

Torrubiella Boud. 1885 (H)

Microhilum H.Y. Yip & A.C. Rath 1989

(H)

Pseudogibellula Samson & H.C. Evans

1973 (H)

Rotiferophthora G.L. Barron 1991 (H)

Septofusidium W. Gams 1971, anamorphic

Cordyceps Fr. 1824 (H)

Simplicillium W. Gams & Zare 2001,

possibly anamorphic Torrubiella Boud.

1885 (H)

Tolypocladium W. Gams 1971, anamo-

rphic Cordyceps Fr. 1824 (H)

Hypocreaceae De Not.

Acremonium-like, anamorphic Protocrea

Petch 1937 and Pseudohypocrea

Yoshim. Doi 1972 (H)

Arachnocrea Z. Moravec 1956, anamor-

phic Protocrea Petch 1937 and

Sarawakus Lloyd 1924 (C)

Cladobotryum Nees 1816, anamorphic

Hypomyces (Fr.) Tul. & C. Tul. 1860

(H)*

Gliocladium Corda 1840, anamorphic

Hypocrea Fr. 1825 and Sphaerostil-

bella (Henn.) Sacc. & D. Sacc. 1905

(H)

Gliocladium-like, anamorphic Sarawakus

Lloyd 1924 (H)

Mycogone Link 1809, anamorphic Hypo-

myces (Fr.) Tul. & C. Tul. 1860 (H)

Mycosphere Doi 10.5943/mycosphere/3/2/5

183

Papulaspora-like, anamorphic Hypomyces

(Fr.) Tul. & C. Tul. 1860 (H)

Sepedonium Link 1809, anamorphic Hypo-

myces (Fr.) Tul. & C. Tul. 1860 (H)

Sibirina G.R.W. Arnold 1970, anamorphic

Hypomyces (Fr.) Tul. & C. Tul. 1860

(H)

Stephanoma Wallr. 1833, anamorphic

Hypomyces (Fr.) Tul. & C. Tul. 1860

(H)

Stilbella Lindau 1900, anamorphic

Hypocrea Fr. 1825 (H)

Stromatocrea W.B. Cooke 1952, anamor-

phic Hypocreopsis P. Karst. 1873 (H)

Trichoderma Pers. 1794, anamorphic

Hypocrea Fr. 1825 and Podostroma P.

Karst. 1892 (H)*

Trichoderma-like, anamorphic Sarawakus

Lloyd 1924 (H)

Trichothecium-like, anamorphic Hypom-

yces (Fr.) Tul. & C. Tul. 1860 (H)

Verticillium-like, anamorphic Aphysiostro-

ma Barrasa, A.T. Martínez & G.

Moreno 1986, Sphaerostilbella Henn.

1901 and Coniochaeta (Sacc.) Cooke

1887 (H)

Verticimonosporium Matsush. 1971 (H)

Nectriaceae Tul. & C. Tul.

Acremonium-like, anamorphic Cosmospo-

ra Rabenh. 1862, Nectria (Fr.) Fr. 1849

and Neocosmospora E.F. Sm. 1899 (H)

Actinostilbe Petch 1925, anamorphic La-

natonectria Samuels & Rossman 1999

(H)

Antipodium Piroz. 1974, anamorphic

Ophionectria Sacc. 1878 (H)

Aphanocladium W. Gams 1971 (H)

Atractium Link 1809 (H)

Calostilbella Höhn. 1919, anamorphic

Calostilbe Sacc. & Syd. 1902 (H)

Campylocarpon Halleen, Schroers &

Crous 2005 (H)

Chaetodochium Höhn. 1932, anamorphic

Pseudonectria Seaver 1909 (H)

Chaetopsina Rambelli 1956, anamorphic

Chaetopsinectria J. Luo & W.Y.

Zhuang and Cosmospora Rabenh. 1862

(H)

Curvicladiella Decock & Crous 2006 (H)

Curvicladium Decock & Crous 1998 (H)

Cyanochyta Höhn. 1915, anamorphic

Gibberella Sacc. 1877 (C)

Cyanophomella Höhn. 1918, anamorphic

Gibberella Sacc. 1877 (C)

Cylindrocarpon Wollenw. 1913, anamor-

phic Neonectria Wollenw. 1917,

Rugonectria P.Chaverri & Samuels

2011, Thelonectria P. Chaverri & C.G.

Salgado 2011, Ilyonectria P.Chaverri &

Samuels 2011 (H)*

Cylindrocladiella Boesew. 1982, anamor-

phic Cosmospora Rabenh. 1862 and

Nectricladiella Crous & C.L. Schoch

2000 (H)

Cylindrocladium Morgan 1892, anamor-

phic Calonectria De Not. 1867 (H)

Dacryoma Samuels 1988, anamorphic

Nectria (Fr.) Fr. 1849 (H)

Dematiocladium Allegr., Aramb., Cazau &

Crous 2005 (H)

Dichomera Cooke 1878, anamorphic

Gibberella Sacc. 1877 (C)

Flagellospora Ingold 1942, anamorphic

Nectria (Fr.) Fr. 1849 (H)

Fusarium Link 1809, anamorphic Albo-

nectria Rossman & Samuels 1999, Cya-

nonectria Samuels & Chaverri 2009,

Gibberella Sacc. 1877, Corallomyce-

tella Henn. 1904 and Haematonectria

Samuels & Nirenberg 1999 and

unknown teleomorphs (H)*

Fusarium-like, anamorphic, Geejayessia

Schroers, Gräfenhan & Seifert 2011 and

Cyanonectria Samuels & Chaverri 2009

(H)*

Fusicolla Bonord. 1851 (H)

Gliocephalotrichum J.J. Ellis & Hesselt.

1962, anamorphic Leuconectria Ross-

man, Samuels & Lowen 1993 (H)

Gliocladiopsis S.B. Saksena 1954, ana-

morphic Glionectria Crous & S.L.

Schoch 2000 (H)

Gyrostroma Naumov 1914, anamorphic

Nectria (Fr.) Fr. 1849 (C)

Heliscus Sacc. 1880, anamorphic Neone-

ctria Wollenw. 1917 . 1849 (H)

Macroconia (Wollenw.) Gräfenhan,

Seifert & Schroers 2011 (H)

Mariannaea G. Arnaud ex Samson 1974

anamorphic ‘Nectria’ (Fr.) Fr. 1849 and

Cordyceps Fr. 1833 (H)*

Mycosphere Doi 10.5943/mycosphere/3/2/5

184

Microcera Desm. 1848, anamorphic

Gibberella Sacc. 1877 (H)*

Mirandina G. Arnaud ex Matsush. 1975,

anamorphic Chaetonectrioides Mats-

ush. (H)

Nalanthamala Subram. 1956, anamorphic

Rubrinectria Rossman & Samuels 1999

(H)

Penicillifer Emden 1968, anamorphic Viri-

dispora Samuels & Rossman 1999 (H)

Pleurocolla Petr. 1924 (H)

Rhizostilbela Wolk 1914, anamorphic

Corallomycetella Henn. 1904 (H)

Septofusidium W. Gams 1971 (H)

Stachybotryna Tubaki & T. Yokoy. 1971,

anamorphic ‘Nectria’ (Fr.) Fr. 1849 (H)

Stilbella Lindau 1900, anamorphic Cosmo-

spora Rabenh. 1862 and Nectria (Fr.)

Fr. 1849 (H)*

Tilachlidium Preuss 1851, anamorphic

‘Pseudonectria’ Seaver 1909 (H)

Tubercularia Tode 1790, anamorphic

Nectria (Fr.) Fr. 1849 (H)*

Verticillium-like, anamorphic Cosmospora

Rabenh. 1862 (H)

Volutella Fr. 1832, anamorphic Cosmo-

spora Rabenh. 1862 and Pseudonectria

Seaver 1909 (H)

Xenocylindrocladium Decock, Hennebert

& Crous 1997, anamorphic Xenocalo-

nectria Crous & C.L. Schoch 2000 (H)

Zythiostroma Höhn. ex Falck 1923,

anamorphic Nectria (Fr.) Fr. 1849 (C)

Niessliaceae Kirschst.

Acremonium-like, anamorphic Tricho-

sphaerella E. Bommer, M. Rousseau &

Sacc. 1891 and Valetoniellopsis Samu-

els & M.E. Barr 1998 (H)

Custingophora-like, anamorphic Melanop-

samma Niessl 1876 (H)

Monocillium S.B. Saksena 1955, ana-

morphic Hyaloseta A.W. Ramaley 2001

and Niesslia Auersw. 1869 (H)

Ophiocordycipitaceae G.H. Sung, J.M. Sung,

Hywel-Jones & Spatafora

Chaunopycnis W. Gams 1979 (H)

Haptocillium W. Gams & Zare 2001

anamorphic Ophiocordyceps Petch

1931 (H)

Hirsutella Pat. 1892, anamorphic Ophio-

cordyceps Petch 1931 (H)

Hymenostilbe Petch 1931, anamorphic

Ophiocordyceps Petch 1931 (H)

Paraisaria Samson & B.L. Brady 1983,

anamorphic Ophiocordyceps Petch

1931 (H)

Purpureocillium Luangsa-ard, Hywel-

Jones, Houbraken & Samson 2011 (H)

Syngliocladium Petch 1932, anamorphic

Ophiocordyceps Petch 1931(H)

Tolypocladium W. Gams 1971, anamor-

phic Elaphocordyceps G.H. Sung &

Spatafora 2007 (H)

Verticillium-like, anamorphic Elaphocor-

dyceps G.H. Sung & Spatafora 2007

(H)

Hypocreales, genera incertae sedis

Acremonium Link 1809 (H)*

Acremonium-like, anamorphic Emeri-

cellopsis J.F.H. Beyma 1940 and Leu-

cosphaerina Arx 1987 (H)

Ascochytopsis-like, anamorphic Euca-

sphaeria Crous 2007 (C)

Cancellidium Tubaki 1975 (H)

Didymostilbe Henn. 1902, anamorphic

Peethambara Subram. & Bhat 1978 (H)

Escovopsis J.J. Muchovej & Della Lucia

1990 (H)

Hapsidospora Malloch & Cain 1970 (C)

Haptospora G.L. Barron 1991 (H)

Illosporiopsis D. Hawksw. 2001 (H)

Illosporium Mart. 1817 (H)

Imicles Shoemaker & Hambl. 2001 (H)

Memnoniella Höhn. 1923 (H)

Myrothecium Tode 1790 (H)

Parasarcopodium Melnik, S.J. Lee &

Crous 2004 (H)

Pseudomicrodochium B. Sutton 1975 (H)

Sarocladium W. Gams & D. Hawksw.

1976 (H)

Septomyrothecium Matsush. 1971, anam-

orphic Peethambara Subram. & Bhat

1978 (H)

Sorosporella Sorokīn 1888 (H)

Spicellum Nicot & Roquebert 1976 (H)

Sporothrix Hektoen & C.F. Perkins 1901,

anamorphic Leucosphaerina Arx 1987

(H)

Stachybotrys Corda 1837 (H)

Mycosphere Doi 10.5943/mycosphere/3/2/5

185

Stilbella Lindau 1900, anamorphic Eme-

ricellopsis J.F.H. Beyma 1940 and

Valsonectria Speg. 1881 (H)

Tilachlidium Preuss 1851 (H)

Trichothecium Link 1809 (H)*

Lulworthiales Kohlm., Spatafora & Volkm.-

Kohlm. 2000

Lulworthiaceae Kohlm., Spatafora & Volkm.-

Kohlm. 2000

Anguillospora Ingold 1942, anamorphic

Lindra I.M. Wilson 1956 (H)

Melanosporales N. Zhang & M. Blackw.

Ceratostomataceae G. Winter

Acrospeira Berk. & Broome 1857 (H)

Gonatobotrys Corda 1839, anamorphic

Melanospora Corda 1837 (H)

Harzia Costantin 1888, anamorphic Mela-

nospora Corda 1837 (H)

Papulaspora Preuss 1851, anamorphic

Melanospora Corda 1837 and Micr-

othecium Corda 1842 (H)

Proteophiala Cif. 1958, anamorphic Mela-

nospora Corda 1837, Persiciospora

P.F. Cannon & D. Hawksw. 1982,

Sphaerodes Clem. 1909, Pteridio-

sperma J.C. Krug & Jeng 1979 (H)

[Sphaerodes Clem. 1909 is a teleomorph

which was reported to produce ampul-

liform phialides on irregularly branched

conidiophores diectrly on ascomata or

on surrounding hyphae (Vujanovic &

Goh, 2009)]

Microascales Luttr. ex Benny & Kimbr.

Chadefaudiellaceae Faurel & Schotter ex

Benny & Kimbr.

Arthrographis-like, anamorphic Faurelina

Locq.-Lin. 1975 (H)

Gondwanamycetaceae Réblová, W. Gams &

Seifert

Custingophora Stolk, Hennebert &

Klopotek 1968, anamorphic Gondwa-

namyces G.J. Marais & M.J. Wingf. 1998

(H)*

Halosphaeriaceae E. Müll. & Arx ex Kohlm.

Cirrenalia Meyers & R.T. Moore 1960

(H)

Clavariopsis-like, anamorphic Corollo-

spora Werderm. 1922 (H)

Clavatospora Sv. Nilsson ex Marvanová

& Sv. Nilsson 1971, anamorphic Corol-

lospora Werderm. 1922 (H)

Culcitalna Meyers & R.T. Moore 1960,

anamorphic Halosarpheia Kohlm. & E.

Kohlm. 1977 (H)

Halosigmoidea Nakagiri, K.L. Pang &

E.B.G. Jones 2009, anamorphic Corol-

lospora Werderm. 1922 (H)

Humicola Traaen 1914, anamorphic Pseu-

dolignincola Chatmala & E.B.G. Jones

2006 and Lignincola Höhnk 1955 (H)

Monodictys S. Hughes 1958 anamorphic

Nereiospora E.B.G. Jones, R.G. Johns-

on & S.T. Moss 1983 (H)

Periconia Tode 1791, anamorphic Halo-

sphaeria Linder 1944 (H)

Sigmoidea J.L. Crane 1968, anamorphic

Corollospora Werderm. 1922 (H)

Trichocladium Harz 1871, anamorphic

Halosphaeria Linder 1944 (H)

Varicosporina Meyers & Kohlm. 1965,

anamorphic Corollospora Werderm.

1922 (H)

Microascaceae Luttr. ex Malloch

Atrhrographis G. Cochet ex Sigler & J.W.

Carmich. 1976, anamorphic Canar-

iomyces Arx (H)

Brachyconidiellopsis Decock, R.F. Casta-

ñeda & Adhikari 2004 (H)

Cephalotrichum Link 1809, anamorphic

Microascus Zukal 1885 (H)

Echinobotryum Corda 1831 (H)

Graphium Corda 1837, anamorphic Micro-

ascus Zukal 1885, Petriella Curzi 1930

and Pseudallescheria Negr. & I. Fisch.

1944 (H)

Humicola Traaen 1914, anamorphic Loph-

otrichus R.K. Benj. 1949 (H)

Parascedosporium Gilgado, Gené, Cano &

Guarro 2007 (H)

Scedosporium Sacc. ex Castell. & Chalm.

1919, anamorphic Pseudallescheria

Negr. & I. Fisch. 1944 and possibly

Petriella Curzi 1930 (H)

Scopulariopsis Bainier 1907, anamorphic

Microascus Zukal 1885 and ?Petriella

Curzi 1930 and Pithoascus Arx 1973

(H)

Mycosphere Doi 10.5943/mycosphere/3/2/5

186

Trichurus Clem. 1896 (H)

Wardomyces F.T. Brooks & Hansf. 1923,

anamorphic Microascus Zukal 188 (H)

Wardomycopsis Udagawa & Furuya 1978,

anamorphic Microascus Zukal 1885

(H)

Microascales, genera incertae sedis

Bisporostilbella Brandsb. & E.F. Morris

1971 (H)

Chalara-like, anamorphic Ceratocystis

Ellis & Halst. 1890 and Cornuvesica

C.D. Viljoen, M.J. Wingf. & K. Jacobs

2000 (H)

Gabarnaudia Samson & W. Gams 1974,

anamorphic Sphaeronaemella P. Karst.

1884 (H)

Sporendocladia G. Arnaud ex Nag Raj &

W.B. Kendr. 1975 (H)

Thielaviopsis Went 1893, anamorphic

Ceratocystis Ellis & Halst. 1890 (H)

Hypocreomycetidae, families incertae sedis

Plectosphaerellaceae W. Gams, Summerbell

& Zare

Acremonium Link 1809 (H)*

Acrostalagmus Corda 1838 (H)*

Gibellulopsis Bat. & H. Maia 1959 (H)

Musicillium Zare & W. Gams 2007 (H)

Plectosporium M.E. Palm, W. Gams &

Nirenberg 1995, anamorphic Plecto-

sphaerella Kleb. 1929 (H)

Stachylidium Link 1809 (H)*

Verticillium Nees 1816 (H)

Hypocreomycetidae, genera incertae sedis

Canalisporium Nawawi & Kuthub. 1989,

anamorphic Ascothailandia Sriindra-

sutdhi, Boonyuen, Sivichai & E.B.G.

Jones 2010 (H)

Dictyochaeta Speg. 1923, anamorphic

Ascocodinaea Samuels, Candoussau &

Magni 1997 (H)

?Glomerulispora Abdel-Wahab, Abdel-

Aziz & Nagahama 2010, anamorphic

Torpedospora Meyers 1957 (H)

Moheitospora Abdel-Wahab, Abdel-Aziz

& Nagahama 2010, anamorphic Junci-

gena Kohlm., Volkm.-Kohlm & O.E.

Erikss. 1997 (H)

Sporoschismopsis Hol.-Jech. & Hennebert

1972, anamorphic Porosphaerellopsis

Samuels & E. Müll. 1982 (H)

Subclass Sordariomycetidae O.E. Erikss. &

Winka

Boliniales P.F. Cannon 2001

Boliniaceae Rick 1931

Phaeoisaria Höhn. 1909 anamorphic

Lentomitella Höhn. 1906 (H)

Calosphaeriales M.E. Barr

Calosphaeriaceae Munk

Calosphaeriophora Réblová, L. Mostert,

W. Gams & Crous 2004, anamorphic

Calosphaeria Tul. & C. Tul. (H)

Lecythophora Nannf. 1934, anamorphic

Sulcatistroma A.W. Ramaley 2005 (H)

Phaeocrella Réblová, L. Mostert, W.

Gams & Crous 2004, anamorphic Tog-

niniella Réblová, L. Mostert, W. Gams

Crous 2004 (H)

Phialophora-like, anamorphic Jattaea

Berl. 1900 (H)

Stachybotrys-like, anamorphic Jattaea

Berl. 1900 (H)

Pleurostomataceae Réblová, L. Mostert, W.

Gams & Crous

Pleurostomophora Vijaykr., L. Mostert,

Jeewon, W. Gams, K.D. Hyde & Crous

2004, anamorphic Pleurostoma Tul. &

C. Tul. 1863 (H)

Chaetosphaeriales Huhndorf, A.N. Mill. &

F.A. Fernández

Chaetosphaeriaceae Réblová, M.E. Barr &

Samuels

Cacumisporium Preuss 1851, anamorphic

Chaetosphaeria Tul. & C. Tul. 1863

(H)

Catenularia Grove 1886, anamorphic

Chaetosphaeria Tul. & C. Tul. 1863

(H)

Chalara (Corda) Rabenh. 1844, anam-

orphic Ascochalara Réblová 1999 and

Chaetosphaeria Tul. & C. Tul. 1863

(H)*

Chloridium Link 1809, anamorphic Chae-

tosphaeria Tul. & C. Tul. 1863 and

Mycosphere Doi 10.5943/mycosphere/3/2/5

187

Melanopsammella Höhn. 1920 (H)

Codinaea Maire 1937, anamorphic Chae-

tosphaeria Tul. & C. Tul. 1863 (H)

?Cordana Preuss 1851, anamorphic Poro-

sphaerella E. Müll. & Samuels 1982

(H)

Craspedodidymum Hol.-Jech. 1972, ana-

morphic Chaetosphaeria Tul. & C. Tul.

1863 (H)

Cryptophiale Piroz. 1968, anamorphic

Chaetosphaeria Tul. & C. Tul. 1863

(H)

Custingophora Stolk, Hennebert & Klop-

otek 1968 anamorphic Chaetosphaeria

Tul. & C. Tul. 1863 (H)

Dictyochaeta Speg. 1923, anamorphic

Chaetosphaeria Tul. & C. Tul. 1863

and Striatosphaeria Samuels & E. Müll.

1979 (H)

Exserticlava S. Hughes 1978, anamorphic

Chaetosphaeria Tul. & C. Tul. 1863

(H)

Fusichalara S. Hughes & Nag Raj 1973,

anamorphic Chaetosphaeria Tul. & C.

Tul. 1863 (H)

Gonytrichum Nees & T. Nees 1818, ana-

morphic Chaetosphaeria Tul. & C. Tul.

1863 (H)

Harpophora W. Gams 2000, anamorphic

Chaetosphaeria Tul. & C. Tul. 1863

(H)

Hemicorynespora M.B. Ellis 1972, ana-

morphic Chaetosphaeria Tul. & C. Tul.

1863 (H)

Kylindria DiCosmo, S.M. Berch & W.B.

Kendr. 1983, anamorphic Chaetos-

phaeria Tul. & C. Tul. 1863 (H)

Menispora Pers. 1822, anamorphic Chae-

tosphaeria Tul. & C. Tul. 1863 (H)

Monilochaetes Halst. ex Harter 1916,

anamorphic Australiasca Sivan. &

Alcorn 2002 (H)*

Nawawia Marvanová 1980 (H)

Paliphora Sivan. & B. Sutton 1985 (H)

Phaeostalagmus W. Gams 1976, ana-

morphic Chaetosphaeria Tul. & C. Tul.

1863 (H)

Phialogeniculata Matsush. 1971 (H)

Phialophora-like, anamorphic Chaeto-

sphaeria Tul. & C. Tul. 1863 (H)

Pseudobotrytis Krzemien. & Badura 1954,

anamorphic Porosphaerella E. Müll. &

Samuels 1982 (H)

Pyrigemmula D. Magyar & R. Shoemaker

2011 (H)

Ramichloridium-like, anamorphic Chaeto-

sphaerides Matsush. 2003 (H)

Rattania Prabhugaonkar & Bhat 2009 (H)

Sporidesmium Link 1809, anamorphic

Umbrinosphaeria Réblová 1999, Miyo-

shiella Kawam. 1929 and Lecy-

thothecium Réblová & Winka 2001 (H)

Sporoschisma Berk. & Broome 1847, ana-

morphic Melanochaeta E. Müll., Harr

& Sulmont 1969 and Chaetosphaeria

Tul. & C. Tul. 1863 (H)

Stachybotrys Corda 1837, anamorphic

Melanopsamma Niessl 1876 (H)

Thozetella Kuntze 1891 (H)

Zanclospora S. Hughes & W.B. Kendr.

1965, anamorphic Chaetosphaeria Tul.

& C. Tul. 1863 (H)

Coniochaetales Huhndorf, A.N. Mill. & F.A.

Fernández

Coniochaetaceae Malloch & Cain

Cladobotryum-like, anamorphic Conioch-

aeta (Sacc.) Cooke 1887 (H)

?Dactylaria Sacc. 1880, anamorphic Syna-

ptospora Cain 1957 (H)

Lecythophora Nannf. 1934, anamorphic

Coniochaeta (Sacc.) Cooke 1887 (H)

Phialophora-like, anamorphic Barrina

A.W. Ramaley 1997 (H)

Diaporthales Nannf.

Cryphonectriaceae Gryzenh. & M.J. Wingf.

Aurapex Gryzenh. & M.J. Wingf. 2006

(C)

Chrysoporthella Gryzenh. & M.J. Wingf.

2004, anamorphic Chrysoporthe Gry-

zenh. & M.J. Wingf. 2004 (C)

Endothiella Sacc. 1906, anamorphic Cry-

phonectria (Sacc.) Sacc. & D. Sacc.

1905 (C)

Foliocryphia Cheewangkoon & Crous

2010 (C)

Prosopidicola Crous & C.L. Lennox 2004

(C)

Ursicollum Gryzenh. & M.J. Wingf. 2006

(C)

Diaporthaceae Höhn. ex Wehm.

Mazzantiella Höhn. 1925, anamorphic

Mycosphere Doi 10.5943/mycosphere/3/2/5

188

Mazzantia Mont. 1855 (C)

Phomopsis (Sacc.) Bubák 1905, anamor-

phic Diaporthe Nitschke 1870 (C)

Gnomoniaceae G. Winter

Asteroma DC. 1815, anamorphic Gnomo-

niella Sacc. 1881 and Plagiostoma

Fuckel 1870 and Pleuroceras Riess.

The type of the genus is considered to

be Montagnellina stellaris (Pers.)

Theiss. & Syd. 1915 (C)

?Depazea Fr. 1818, anamorphic Pleuro-

ceras Riess 1854 (C)

Diplodina Westend. 1857, anamorphic

Plagiostoma Fuckel 1870 (C)

Discosporium Sacc. & P. Syd. 1902 ana-

morphic Cryptodiaporthe Petr. 1921 C.

Tul. (C)

Discula Sacc. 1884, anamorphic Apiogno-

monia Höhn. 1917, Gnomonia Ces. &

De Not. 1863 and Amphiporthe Petr.

1971 (C)

Gloeosporidina Petr. 1921, anamorphic

Apiognomonia Höhn. 1917 (C)

Mazzantiella Höhn. 1925, anamorphic

Mamianiella Höhn. 1917 (C)

Millerburtonia Cif. 1951, anamorphic Pla-

giostoma Fuckel 1870 (C)

Phaeocytostroma Petr. 1921, anamorphic

Clypeoporthe Höhn 1919 (C)

?Septogloeum Sacc. 1880, anamorphic

Pleuroceras Riess 1854 (C)

Uniseta Ciccar. 1948, anamorphic Pla-

giostoma Fuckel 1870 (C)

?Zythia Fr. 1849 (C)

Melanconidaceae G. Winter

Dumortieria Westend. 1857 (as Cytospo-

rina Sacc. 1884), anamorphic Mela-

namphora Lafl. 1976 and Melogramma

Fr. 1849 (C)

Harknessia Cooke 1881, anamorphic Wue-

stneia Auersw. ex Fuckel 1863 (C)

Hendersonula Speg. 1880, anamorphic

Pseudovalsella Höhn. 1918 and Dictyo-

porthe Petr. 1955 (C)

Mastigosporella Höhn. 1914, anamorphic

Wuestneiopsis J. Reid & Dowsett 1990

(C)

Melanconiopsis Ellis & Everh. 1900, ana-

morphic Massariovalsa Sacc. 1882 (C)

Melanconium Link 1809, anamorphic

Melanconis Tul. & C. Tul. 1863 (C)

?Stilbospora Pers. 1794, anamorphic Pros-

thecium Fresen. 1852 (C)

Tubakia B. Sutton 1973, anamorphic

Dicarpella Syd. & P. Syd. 1921 (C)

Pseudoplagiostomaceae Cheewangkoon, M.J.

Wingf. & Crous

Cryptosporiopsis-like, anamorphic Pseu-

doplagiostoma Cheewangkoon, M.J.

Wingf. & Crous 2010 (C)

Pseudovalsaceae M.E. Barr

Coryneum Nees 1816, anamorphic Pseu-

dovalsa Ces. & De Not. 1863 (C)

Schizoparmeaceae Rossman

Coniella Höhn. 1918 (C)

Pilidiella Petr. & Syd. 1927, anamorphic

Schizoparme Shear 1923 (C)

Sydowiellaceae Lar. N. Vassiljeva

Cylindrosporella Höhn. 1916, anamorphic

Stegophora Syd. & P. Syd. 1916 (C)

Phoma-like, anamorphic Hapalocystis

Auersw. ex Fuckel 1863 (C)

Togniniaceae Réblová, L. Mostert, W. Gams

& Crous

Phaeoacremonium W. Gams, Crous &

M.J. Wingf. 1996, anamorphic

Togninia Berl. 1900 (H)

Valsaceae Tul. & C. Tul.

Amphicytostroma Petr. 1921, anamorphic

Amphiporthe Petr. 1971 (C)

Cytospora Ehrenb. 1818, anamorphic

Leucostoma (Nitschke) Höhn. 1917,

Valsa Fr. 1849, Valsella Fuckel 1870

and Valseutypella Höhn. 1919 (C)

Harpostroma Höhn. 1928, anamorphic

Leptosillia Höhn. 1928 (C)

?Lasmenia Speg. 1886, anamorphic Chad-

efaudiomyces Kamat, Rao, Patil &

Ullasa 1974 (C)

Phomopsis (Sacc.) Bubák 1905, anamor-

phic Clypeoporthella Petr. 1924 (C)

Diaporthales, genera incertae sedis

Craspedodidymum Hol.-Jech. 1972, ana-

morphic Thailandiomyces Pinruan,

Sakay., K.D. Hyde & E.B.G. Jones

Mycosphere Doi 10.5943/mycosphere/3/2/5

189

2008 (H)

Apoharknessia Crous & S.J. Lee 2004 (C)

Apomelasmia Grove 1937 (C)

Asterosporium Kunze 1819 (C)

Craspedodidymum Hol.-Jech. 1972, ana-

morphic Thailandiomyces Pinruan, Sa-

kay., K.D. Hyde & E.B.G. Jones 2008

(H)

Cytospora-like, anamorphic Pachytrype

Berl. ex M.E. Barr, J.D. Rogers & Y.M.

Ju 1993 (C)

Dwiroopa Subram. & Muthumary

?Greeneria Scribn. & Viala 1887 (C)

Hormonema Lagerb. & Melin 1927, ana-

morphic Valsaria Ces. & De Not. 1863

(H)

Hypodermina Höhn. 1916 (C)

Macrohilum H.J. Swart 1988 (C)

Natarajania Pratibha & Bhat 2006 (H)

Phomopsis-like, anamorphic Caudospora

Starbäck 1889 (C)

Rabenhorstia Fr. 1849, anamorphic Her-

cospora Fr. 1825 (C)

Sirococcus Preuss 1855 (C)

Stenocarpella Syd. & P. Syd. 1917 (C)

Synchaetomella Decock & Seifert 2005

(H)

Magnaporthales Thongk., Vijakr. & K.D.

Hyde

Magnaporthaceae P.F. Cannon

Clasterosporium Schwein. 1832, anamor-

phic Clasterosphaeria Sivan. 1984 (H)

Didymobotryum Sacc. 1886, anamorphic

Ceratosphaerella Huhndorf, Greif,

Mugambi & A.N. Mill. 2008 (H)

Harpophora W. Gams 2000, anamorphic

Gaeumannomyces Arx & D.L. Olivier

1952 and Magnaporthe R.A. Krause &

R.K. Webster 1972 (H)

Mycoleptodiscus Ostaz. 1968, anamorphic

Omnidemptus P.F. Cannon & Alcorn

1994 (H)

Nakataea Hara 1939, anamorphic Mag-

naporthe R.A. Krause & R.K. Webster

1972 (H)

Pyricularia Sacc. 1880, anamorphic Mag-

naporthe R.A. Krause & R.K. Webster

1972 (H)

Trichocladium-like, anamorphic Gaeuma-

nnomyces Arx & D.L. Olivier 1952 (H)

Ophiostomatales Benny & Kimbr.

Kathistaceae Malloch & M. Blackw.

Mattirolella S. Colla 1929 (C)

Termitaria Thaxt. 1920 (H)

Termitariopsis M. Blackw., Samson &

Kimbr. 1980 (H)

Ophiostomataceae Nannf.

Acremonium-like, anamorphic Frago-

sphaeria Shear 1923 (H)

Ambrosiella Brader ex Arx & Hennebert

1965, anamorphic Grosmannia Goid.

1936 and Ophiostoma Syd. & P. Syd.

1919 (H)

Graphium Corda 1837, anamorphic Op-

hiostoma Syd. & P. Syd. 1919 (H)

Hyalobelemnospora Matsush. 1993, ana-

morphic Subbaromyces Hesselt. 1953

(H)

Hyalorhinocladiella H.P. Upadhyay &

W.B. Kendr. 1975, anamorphic Cerato-

cystiopsis H.P. Upadhyay & W.B.

Kendr. 1975, Klasterskya Petr. 1940

and Ophiostoma Syd. & P. Syd. 1919

(H)

Hyalorhinocladiella-like, anamorphic Kla-

sterskya Petr. 1940 (H)

Leptographium Lagerb. & Melin 1927,

anamorphic Grosmannia Gold. 1936

(H)*

Pesotum J.L. Crane & Schokn. 1973,

anamorphic Ophiostoma Syd. & P. Syd.

1919 (H)

Pesotum-like, anamorphic Grosmannia

Gold. 1936 (H)

Phialographium H.P. Upadhyay & W.B.

Kendr. 1974, anamorphic Ophiostoma

Syd. & P. Syd. 1919 (C)

Raffaelea Arx & Hennebert 1965, anamor-

phic Grosmannia Goid. 1936 and

Ophiostoma Syd. & P. Syd. 1919 (H)

Sporothrix Hektoen & C.F. Perkins 1901,

anamorphic Ophiostoma Syd. & P. Syd.

1919 (H)

Sordariales Chad. ex D. Hawksw. & O.E.

Erikss.

Chaetomiaceae G. Winter

Acrophialophora Edward 1961 (H)

Botryotrichum Sacc. & Marchal 1885, ana-

morphic Chaetomium Kunze 1817 and

Farrowia D. Hawksw. (H)

Mycosphere Doi 10.5943/mycosphere/3/2/5

190

Humicola Traaen 1914, anamorphic Chae-

tomium Kunze 1817 (H)

Madurella Brumpt 1905 (H)

Myceliophthora Costantin 1892, anamorp-

hic Corynascus Arx 1973 and Thielavia

Zopf 1876 (H)

Papulaspora-like, Preuss 1851, anamorp-

hic Chaetomium Kunze 1817 (H)

Phialophora-like, anamorphic Chaetomid-

ium (Zopf) Sacc. 1882 (H)

Retroconis de Hoog & Bat. Vegte 1989

(H)

Taifanglania Z.Q. Liang, Y.F. Han, H.L.

Chu & R.T.V. Fox 2009, anamorphic

Thielavia Zopf 1876 (H)

Trichocladium Harz 1871, anamorphic

Chaetomium Kunze 1817 (H)

Trichosporiella Kamyschko 1960, ana-

morphic Thielavia Zopf 1876 (H)

Lasiosphaeriaceae Nannf.

Angulimaya Subram. & Lodha 1964, ana-

morphic Bombardioidea C. Moreau ex

N. Lundqv. 1972 (H)

Bagadiella Cheewangkoon & Crous 2009

(H)

Cladorrhinum Sacc. & Marchal 1885,

anamorphic Apiosordaria Arx & W.

Gams 1967, Apodospora Cain & J.H.

Mirza 1970, Arnium Nitschke ex G.

Winter 1873, Cercophora Fuckel 1870

and Podospora Ces. 1856 (H)

Mammaria Ces. ex Rabenh. 1854, ana-

morphic Pseudocercophora Subram. &

Sekar 1986 and Cercophora Fuckel

1870 (H)

Phialophora-like, anamorphic Cercophora

Fuckel 1870, Eosphaeria Höhn. 1917,

Lasiosphaeria Ces. & De Not. 1863 and

Podospora Ces. 1856 (H)

Sordariaceae G. Winter

Chrysonilia Arx 1981, anamorphic Neuro-

spora Shear & B.O. Dodge 1927 (H)

Trichosporiella Kamyschko 1960, anam-

orphic Diplogelasinospora Cain 1961

(H)

Virgariella S. Hughes 1953, anamorphic

Effetia Bartoli, Maggi & Persiani 1984

(H)

Sordariales, genera incertae sedis

Kionochaeta P.M. Kirk & B. Sutton 1986

(H)

Lasiadelphia Réblová & W. Gams 2011

(H)

Lunulospora Ingold 1942 (H)

Madurella Brumpt 1905 (H)

Monodictys S. Hughes 1958 anamorphic

Ascotaiwania Sivan. & H.S. Chang

1992 (H)

Monotosporella S. Hughes 1958, anam-

orphic Ascotaiwania Sivan. & H.S.

Chang 1992 (H)

Papulospora Preuss 1851, anamorphic

Sordaria Ces. & De Not. 1863 (H)

Ramophialophora M. Calduch, Stchigel,

Gené & Guarro 2004 (H)

Trichocladium-like, anamorphic Asco-

lacicola Ranghoo & K.D. Hyde 1998,

Ascotaiwania Sivan. & H.S. Chang

1992 (H)

Ypsilonia Lév. 1846, anamorphic Acantho-

theciella Höhn. 1911 (H)

Sordariomycetidae, families incertae sedis

Amplistromataceae Huhndorf, A.N. Mill., M.

Greif & Samuels

Acrodontium-like, anamorphic Amplistro-

ma Huhndorf, A.N. Mill., M. Greif &

Samuels 2009 (H)

Pseudogliomastix W. Gams 1985, ana-

morphic Wallrothiella Sacc. 1882 (H)

Annulatascaceae S.W. Wong, K.D. Hyde &

E.B.G. Jones

Phaeoisaria Höhn. 1909, anamorphic

Rhamphoria Niessl 1876 (H)

Phialophora-like, anamorphic Teraco-

sphaeria Réblová & Seifert 2007 (H)

Cephalothecaceae Höhn.

Acremonium-like, anamorphic Alberti-

niella Kirschst. 1936 (H)

Chalara (Corda) Rabenh. 1844, anamor-

phic Cryptendoxyla Malloch & Cain

1970 (H)*

Phialemonium W. Gams & McGinnis

1983 (H)

Tritirachium Limber 1940, anamorphic

Cephalotheca Fuckel 1871 (H)

Helminthosphaeriaceae Samuels, Candoussau

& Magní

Diplococcium Grove 1885, anamorphic

Mycosphere Doi 10.5943/mycosphere/3/2/5

191

Helminthosphaeria Fuckel 1870 (H)

Endophragmiella B. Sutton 1973, anamor-

phic Echinosphaeria A.N. Mill. &

Huhndorf 2004 (H)

Selenosporella-like, anamorphic Ruzenia

O. Hilber 2002 (H)

Spadicoides S. Hughes 1958, anamorphic

Tengiomyces Réblová 1999 (H)

Vermiculariopsiella Bender 1932, anamor-

phic Echinosphaeria A.N. Mill. &

Huhndorf 2004 (H)

Sordariomycetidae, genera incertae sedis

Sterigmatobotrys Oudem. 1886 (H)*

Brachysporiella Bat. 1952, anamorphic

Ascotaiwania Sivan. & H.S. Chang

1992 (H)

Conioscypha Höhn. 1904, anamorphic

Conioscyphascus Réblová & Seifert

2004 (H)

Endophragmiella B. Sutton 1973, ana-

morphic Phaeotrichosphaeria Sivan.

1983 (H)

Menisporopsis S. Hughes 1952, anamor-

phic Menisporopascus Matsush. 2003

(H)

Phialophora-like, anamorphic Linocarpon

Syd. & P. Syd. 1917 (H)

Pleurothecium Höhn. 1919, anamorphic

Carpoligna F.A. Fernández & Huhn-

dorf 1999 (H)

Ramichloridium-like, anamorphic Barba-

tosphaeria Reblova 2007 (H)

Rhodoveronaea Arzanlou, W. Gams &

Crous 2007 (H)

Spadicoides S. Hughes 1958 (H)

Sporothrix-like, anamorphic Barbatosph-

aeria Reblova 2008 (H)

Subclass Xylariomycetidae sensu O.E. Erikss.

& Winka

Xylariales Nannf.

Amphisphaeriaceae G. Winter

Bartalinia Tassi 1900 (C)

?Beltraniella Subram. 1952, anamorphic

Leiosphaerella Höhn. 1919 (H)

Bleptosporium Steyaert 1961, anamorphic

Amphisphaeria Ces. & De Not. 1863

(C)

Ceratosporium Schwein. 1832, anamor-

phic Iodosphaeria Samuels, E. Müll. &

O. Petrini 1987 (H)

?Chaetoconis Clem. 1909, anamorphic

Ceriospora Niessl 1876 (C)

Discosia Lib. 1837, anamorphic Adisciso

Kaz. Tanaka, Okane & Hosoya 2011

(C)*

Doliomyces Steyaert 1961 (C)

?Hyalotiopsis Punith. 1970, anamorphic

Ellurema Nag Raj & W.B. Kendr. 1986

(C)

Immersidiscosia Kaz. Tanaka, Okane &

Hosoya 2011 (C)*

Labridella Brenckle 1929, anamorphic

Griphosphaerioma Höhn. 1918 (C)

Labridium Vestergr. 1897, anamorphic

Discostroma Clem. 1909 (C)

Microdochium Syd. 1924, anamorphic

Monographella Petr. 1924 (H)

Monochaetia (Sacc.) Allesch. 1902 (C)

Monochaetinula Muthumary, Abbas & B.

Sutton 1986 (C)

Monochaetiopsis L. Cai, Jeewon & K.D.

Hyde 2003, anamorphic Dyrithiopsis L.

Cai, R. Jeewon & K.D. Hyde 2003 (C)

Morinia Berl. & Bres. 1889 (C)

?Pestalotia De Not. 1841, anamorphic

Broomella Sacc. 1883 (C)

Pestalotiopsis Steyaert 1949, anamorphic

Neobroomella Petr. 1947 and Pestalo-

sphaeria M.E. Barr 1975 (C)

Phlogicylindrium Crous, Summerb. &

Summerell 2006 (H)

Sarcostroma Cooke 1871, anamorphic

Griphosphaerioma Höhn. 1918 (C)

Seimatosporium Corda 1833, anamorphic

anamorphic Discostroma Clem. 1909

(C)

Seiridium Nees 1816, anamorphic Blogia-

scospora Shoemaker, E. Müll. &

Morgan-Jones 1966 and Lepteutypa

Petr. 1923 (C)

Selenosporella G. Arnaud ex MacGarvie

1969, anamorphic Oxydothis Penz. &

Sacc. 1898 and Iodosphaeria Samuels,

E. Müll. & O. Petrini 1987 (H)

Sporocadus Corda 1839 (C)

Synnemapestaloides T. Handa & Y.

Harada 2004 (H)

?Truncatella Steyaert 1949 anamorphic

Broomella Sacc. 1883 (C)

Vermisporium H.J. Swart & M.A. Will.

1983 (C)

Zetiasplozna Nag Raj 1993 (C)

Mycosphere Doi 10.5943/mycosphere/3/2/5

192

Diatrypaceae Nitschke

Dumortieria Westend. 1857 (some as

Cytosporina Sacc. 1884), anamorphic

Cryptosphaeria Ces. & De Not. 1863,

Diatrype Fr. 1849, Eutypa Tul. & C.

Tul. 1863 and Eutypella (Nitschke)

Sacc. 1875 (C)

Libertella Desm. 1830, anamorphic

Diatrype Fr. 1849, Diatrypella (Ces. &

De. Not.) De. Not. 1863, Eutypa Tul. &

C. Tul. 1863 and Eutypella (Nitschke)

Sacc. 1875 (C)

Diatrypaceae Nitschke 1869

Phaeoisaria Höhn. 1909 anamorphic

Eutypella (Nitschke) Sacc. 1875 (H)

Hyponectriaceae Petr.

Arthrinium Kunze 1817, anamorphic

Physalospora Niessl (H)

Aureobasidium Viala & G. Boyer 1891,

anamorphic Discosphaerina Höhn.

1917 (H)

Beltraniella Subram. 1952, anamorphic

Pseudomassaria Jacz. 1894 (H)

Hormonema-like, anamorphic Discosp-

haerina Hohn 1917 (H)

Kabatia Bubák 1904, anamorphic Disco-

sphaerina Höhn. 1917 (C)

Kabatiella Bubák 1907, anamorphic Dis-

cosphaerina Höhn. 1917 (H)

Sarcophoma Höhn. 1916, anamorphic Dis-

cosphaerina Hohn 1917 (C)

Selenophoma Maire 1907, anamorphic

Discosphaerina Hohn 1917 (C)

Graphostromataceae M.E. Barr, J.D. Rogers

& Y.M. Ju

Nodulisporium-like, anamorphic Grapho-

stroma Piroz. 1974 (H)

Xylariaceae Tul. & C. Tul.

Acanthodochium Samuels, J.D. Rogers &

Nagas., anamorphic Astrocystis Berk. &

Broome and Collodiscula I. Hino &

Katum. 1955 (H)

Arthroxylaria Seifert & W. Gams 2002

Dematophora R. Hartig 1883, anamorphic

Rosellinia De Not. 1844 (H)

?Dicyma Boulanger 1897, anamorphic

Ascotricha Berk. 1838 (H)

Geniculisynnema Okane & Nakagiri 2007,

anamorphic Nemania Gray 1821 (H)

Geniculosporium Chesters & Greenh.

1964, anamorphic Entoleuca Syd. 1922,

Leprieuria Laessøe, J.D. Rogers &

Whalley 1989, Nemania Gray 1821,

Phylacia Lév. 1845, Podosordaria Ellis

& Holw. 1897, Rosellinia De Not. 1844

and Xylaria Hill ex Schrank 1789 (H)

Geniculosporium-like, anamorphic Halo-

rosellinia Whalley, E.B.G. Jones, K.D.

Hyde & Læssøe 2000, Jumillera J.D.

Rogers, Y.M. Ju & San Martín 1997

and Kretzschmaria Fr. 1849 (H)

Hypocreodendron Henn. 1897, anamor-

phic Discoxylaria J.C. Lindq. & J.E.

Wright 1964 (H)

Libertella Desm. 1830 (C)

Libertella-like, anamorphic Barrmaelia

Rappaz 1995, Creosphaeria Theiss.

1910, Jumillera J.D. Rogers, Y.M. Ju &

San Martín 1997 and Lopadostoma

(Nitschke) Traverso 1906 (C)

Lindquistia Subram. & Chandrash. 1977,

anamorphic Poronia Willd. 1787 (H)

Masoniomyces J.L. Crane & Dumont

1975, anamorphic Camillea Fr. 1849

(H)

Mirandina-like, anamorphic Kretzschma-

riella Viégas 1944 (H)

Moelleroclavus Henn. 1902, anamorphic

Xylaria Hill ex Schrank 1789 (H)

Muscodor Worapong, Strobel & W.M.

Hess 2001 (H)

Nodulisporium Preuss 1849, anamorphic

Annulohypoxylon Y.M. Ju, J.D. Rogers

& H.M. Hsieh 2005, Biscogniauxia

Kuntze 1891, Daldinia Ces. & De Not.

1863, Entonaema Möller 1901, Hypo-

xylon Bull. 1791, Induratia Samuels, E.

Müll. & O. Petrini, Pyrenomyxa Mor-

gan 1895 (as Pulveria Malloch &

Rogerson 1977), Thamnomyces Ehrenb.

1820, Theissenia Maubl. 1914, Thue-

menella Penz. & Sacc. 1898 and

Xylaria Hill ex Schrank (H)

Nodulosporium-like, anamorphic Rhopalo-

stroma D. Hawksw. 1977 and Ruwen-

zoria J. Fornier, M. Stadler, Læssøe &

C. Decock 2010 (H)

Padixonia Subram. 1972, anamorphic

Xylaria Hill ex Schrank (H)

Mycosphere Doi 10.5943/mycosphere/3/2/5

193

Rhinocladiella-like, anamorphic Obolari-

na Pouzar 1986 (H)

Sporothrix-like, anamorphic Rostrohypo-

xylon J. Fourn. & M. Stadler 2010 and

Ruwenzoria J. Fornier, M. Stadler,

Læssøe & C. Decock 2010 (H)

Triplicaria P. Karst. 1889, anamorphic

Hypoxylon Bull. 1791 (H)

Virgaria Nees 1816, anamorphic Ascovir-

garia J.D. Rogers & Y.M. Ju 2002 (H)

Virgariella-like, anamorphic Hypoxylon

Bull. 1791 (H)

Vivantia J.D. Rogers, Y.M. Ju & Cand.

1996 (H)

Xylocladium P. Syd. ex Lindau 1900,

anamorphic Camillea Fr. 1849 (H)

Xylocoremium J.D. Rogers 1984, anamor-

phic Xylaria Hill ex Schrank 1789 (H)

Xylariales, genera incertae sedis

Chaetochalara B. Sutton & Piroz. 1965

(H)

Chalara (Corda) Rabenh. 1844 (H)*

Hadrotrichum Fuckel 1865 (H)

Humicola-like, anamorphic Ascotrichella

Valldos. & Guarro 1988 (H)

Idriella P.E. Nelson & S. Wilh. 1956,

anamorphic Phomatospora Sacc. 1875

(H)

Melanographium Sacc. 1913, possibly

anamorphic Lasiobertia Sivan. 1978

(H)

Nodulisporium-like, anamorphic Coniola-

riella D. Garcıá, Stchigel & Guarro (H)

Rhinocladiella-like, anamorphic Coniola-

riella D. Garcıá, Stchigel & Guarro (H)

Selenosporella G. Arnaud ex MacGarvie

1969, anamorphic Diatrypasimilis J.

Zhou, & Kohlm. 2010 (H)

Sporidesmina Subram. & Bhat 1989 (H)

Subramaniomyces Varghese & V.G. Rao

1980 (H)

Sordariomycetes, orders incertae sedis

Lulworthiales Kohlm., Spatafora & Volkm.-

Kohlm.

Lulworthiaceae Kohlm., Spatafora & Volkm.-

Kohlm.

Anguillospora Ingold 1942, anamorphic

Lindra I.M. Wilson 1956 (H)

Cumulospora I. Schmidt 1985, anamor-

phic Lulwoana Kohlm., Volkm.-

Kohlm., J. Campb., Spatafora & Grä-

fenhan 2005 and Lulworthia G.K.

Sutherl. 1916 (H)

Halazoon Abdel-Aziz, Abdel-Wahab &

Nagahama 2010, anamorphic Lulwort-

hia G.K. Sutherl. 1916 (C)

Hydea K.L. Pang & E.B.G. Jones 2010

(H)

Matsusporium E.B.G. Jones & K.L. Pang

2010 (H)

Moleospora Abdel-Wahab, Abdel-Aziz &

Nagahama 2010 (H)

Moromyces Abdel-Wahab, K.L. Pang,

Nagahama, Abdel-Aziz & E.B.G. Jones

2010 (H)

Zalerion R.T. Moore & Meyers 1962,

anamorphic Lulwoana Kohlm., Volkm.-

Kohlm, J. Campb., Spatafora & Gräf-

enhan 2005 (H)

Lulworthiales, genera incertae sedis

Halazoön Abdel-Aziz, Abdel Wahab &

Nagah. 2011 (H)

Phyllachorales M.E. Barr

Phyllachoraceae Theiss. & H. Syd.

Acerviclypeatus Hanlin 1990, anamorphic

Ophiodothella Boyd 1934 (C)

Baeumleria Petr. & Syd. 1927, anamo-

rphic Phyllachora Nitschke ex Fuckel

1870 (as Trabutia Sacc. & Roum. 1881)

(C)

Colletogloeum-like, anamorphic Cocco-

diella Hara 1909 (C)

Cyclodomus Höhn. 1909, anamorphic

Maculatifrons K.D. Hyde 1996 (C)

Diachorella Höhn. 1918, anamorphic

Diachora Müll. 1893 (C)

Hysterodiscula Petr. 1942, anamorphic

Orphnodactylis Malloch & A. Mallik

1998 (C)

Linochora Höhn. 1910, anamorphic Phyl-

lachora Nitschke ex Fuckel 1870 (C)

Mycohypallage B. Sutton 1963, anamor-

phic Deshpandiella Kamat & Ullasa

1973 (C)

Oswaldina Rangel 1921, anamorphic Api-

osphaeria Höhn. 1909 (C)

Polystigmina Sacc. 1884, anamorphic Pol-

ystigma DC. 1815 (C)

Pseudothiopsella Petr. 1928, anamorphic

Pseudothiella Petr. 1928 (C)

Mycosphere Doi 10.5943/mycosphere/3/2/5

194

Rhodosticta Woron. 1911, anamorphic

Polystigma DC. 1815 and Stigmatula

(Sacc.) Syd. & P. Syd. 1901 (C)

Trichosphaeriales M.E. Barr 1983

Trichosphaeriaceae G. Winter 1885

Brachysporium Sacc. 1886, anamorphic

Cryptadelphia Réblová & Seifert 2004

(H)

Koorchaloma Subram. 1953, anamorphic

Kananascus Nag Raj 1984 (H)

Stanjehughesia Subram. 1992, anamorphic

Miyoshiella Kawam. 1929 and Umbri-

nosphaeria Réblová 1999 (H)

Stromatographium Höhn. 1907, anamor-

phic Fluviostroma Samuels & E. Müll.

1980 (H)

Sordariomycetes, family incertae sedis

Apiosporaceae K.D. Hyde, J. Fröhlich, Joanne

E. Taylor & M.E. Barr

Arthrinium Kunze 1817, anamorphic Apio-

spora Sacc. 1875 (H)

Cordella Speg. 1886, anamorphic Apio-

spora Sacc. 1875 (H)

Pteroconium Sacc. ex Grove 1914, anamo-

rphic Apiospora Sacc. 1875 (H)

Scyphospora L.A. Kantsch. 1928, anamor-

phic Apiospora Sacc. 1875 (C)

Thyridiaceae O.E. Erikss. & J.Z. Yue

Pleurocytospora Petr. 1923, anamorphic

Thyridium Nitschke 1867 (C)

Sordariomycetes, genera incertae sedis

Cryptomycella Höhn. 1925, anamorphic

Cryptomycina Höhn. 1977 (C)

?Custingophora Stolk, Hennebert & Klo-

potek 1968 (H)

Didymostilbe-like, anamorphic Ornatispo-

ra K.D. Hyde, Goh, Joanne E. Taylor,

J. Fröhl. 1999 (H)

Dinemasporium Lév. 1846, anamorphic

Phomatospora Sacc. 1875 (C)

Ellisembia Subram. 1992 (H)

Linkosia A. Hern. Gut. & B. Sutton 1997

(H)

Myrmecridium Arzanlou, W. Gams &

Crous 2007 (H)

Nigrospora Zimm. 1902, anamorphic

Khuskia H.J. Huds. 1963 (H)

Penicillifer Emden 1968, anamorphic

Stellosetifera Matsush.1996 (H)

Phomatosporella Tak. Kobay. & K. Sasaki

1982, anamorphic Phomatospora Sacc.

1875 (C)

Ramichloridium-like, anamorphic Tectoni-

dula Réblová 2009 (H)

Sporothrix-like, anamorphic Phomato-

spora Sacc. 1875 and Tectonidula Réb-

lová 2009 (H)

Subphylum PEZIZOMYCOTINA, orders

incertae sedis

Lahmiales O. E. Erikss.

Lahmiaceae O. E. Erikss.

Cyphellophora G.A. de Vries 1962, ana-

morphic Lahmia Körb. 1861 (H)

ASCOMYCOTA, families incertae sedis

Families and genera that cannot be

placed in any of the classes and orders accepted

in the present classification with a high degree

of probability are listed below.

Amorphothecaceae Parbery

Hormoconis Arx & G.A. de Vries 1973,

anamorphic Amorphotheca Parbery

1969 (H)

Batistiaceae Samuels & K.F. Rodrigues

Acrostroma Seifert 1987, anamorphic

Batistia Cif. 1958 (H)

Pseudeurotiaceae Malloch & Cain

Teberdinia Sogonov, W. Gams, Summerb.

& Schroers 2005, anamorphic Pseudeu-

rotium J.F.H. Beyma 1937 (H)

Seuratiaceae Vuill. ex M.E. Barr

Atichia Flot. 1850, anamorphic Seuratia

Pat. 1904 (H)

ASCOMYCOTA, genera incertae sedis

Abropelta B. Sutton 1986 (C)

Acarellina Bat. & H. Maia 1960 (C)

Acaroconium Kocourk. & D. Hawksw.

2008 (C)

Acarocybe Syd. 1937 (H)

Acarocybella M.B. Ellis 1960 (H)

Acarocybellina Subram. 1992 (H)

Acarocybiopsis J. Mena, A. Hern. Gut. &

Mercado 1999 (H)

Acaropeltis Petr. 1937 (C)

Mycosphere Doi 10.5943/mycosphere/3/2/5

195

Achoropeltis Syd. 1929 (C)

Aciculariella G. Arnaud 1954 (H)

Acontium Morgan 1902 (H)

Acremoniula G. Arnaud 1954 (H)

Acrodictyella W.A. Baker & Partr. 2001

(H)

Acrodictyopsis P.M. Kirk 1983 (H)

Acrodictys M.B. Ellis 1961 (H)

Acrodontiella U. Braun & Scheuer 1995

(H)

Acrophragmis Kiffer & Reisinger 1970

(H)

Acrostaurus Deighton & Piroz. 1972 (H)

Actinocladium Ehrenb. 1819 (H)

Actinotexis Arx 1960 (C)

Actinothecium Ces. 1854 (H)

Actinothyrium Kunze 1823 (C)

Acumispora Matsush. 1980 (H)

Aegeritella Bałazy & J. Wiśn. 1974 (H)

Aenigmatospora R.F. Castañeda, Saikawa,

Guarro & Calduch 1999 (?H)

Agaricodochium X.J. Liu 1981 (H)

Agarwalomyces R.K. Verma & Kamal

1987 (H)

Agrabeeja Subram. 1995 (H)

Agyriella Sacc. 1884 (H)

Agyriellopsis Höhn. 1903 (C)

Ahmadia Syd. 1939 (C)

Ajrekarella Kamat & Kalani 1964 (C)

Alatosessilispora K. Ando & Tubaki 1984

(H)

Alciphila Harmaja 2002 (H)

Algonquinia R.F. Castañeda & W.B.

Kendr. 1991 (H)

Alloneottiosporina Nag Raj 1993 (C)

Allothyriella Bat., Cif. & Nascim. 1959

(C)

Allothyrina Bat. & J.L. Bezerra 1964 (C)

Allothyriopsis Bat., Cif. & H. Maia 1959

(C)

Alpakesa Subram. & K. Ramakr. 1954 (C)

Alpakesiopsis Abbas, B. Sutton, Ghaffar &

A. Abbas 2003 (C)

Alveophoma Alcalde 1952 (C)

Alysidiopsis B. Sutton 1973 (H)

Amallospora Penz. 1897 (H)

Amblyosporium Fresen. 1863 (H)

Amerodiscosiella M.L. Farr 1961 (C)

Amerodiscosiellina Bat. & Cavalc. 1966

(C)

Amerosporiopsis Petr. 1941 (C)

Amerosympodula Matsush. 1996 (H)

Amoenodochium Peláez & R.F. Castañeda

1996 (H)

Amoenomyces R.F. Castañeda, Saikawa &

Hennebert 1996 (H)

Amphichaetella Höhn. 1916 (H)

Amphophialis R.F. Castañeda, W.B.

Kendr. & Guarro 1998 (H)

Amphoropycnium Bat. 1963 (C)

Ampullicephala R.F. Castañeda, Minter &

M. Stadler 2009 (H)

Ampulliferina B. Sutton 1969 (H)

Anaphysmene Bubák 1906 (C)

Anarhyma M.H. Pei & Z.W. Yuan 1986

(C)

Anaselenosporella Heredia, R.F. Casta-

ñeda & R.M. Arias 2010 (H)

Ancoraspora Mig. Rodr. 1982 (H)

Ancorasporella J. Mena, Mercado &

Heredia 1998 (H)

Angiopomopsis Höhn. 1912 (C)

Angulospora Sv. Nilsson 1962 (H)

Annellodentimyces Matsush. 1985 (H)

Annellodochium Deighton 1969 (H)

Annellolacinia B. Sutton 1964 (C)

Annellophora S. Hughes 1952 (H)

Annellophorella Subram. 1962 (H)

Annellophragmia Subram. 1963 (H)

Annellosympodia McTaggart, R.G. Shivas

& U. Braun 2007 (H)

Antennopsis R. Heim 1952 (H)

Anthopsis Fil. March., A. Fontana & Luppi

Mosca 1977 (H)

Anthracoderma Speg. 1888 (C)

Antromyces Fresen. 1850 (H)

Anulohypha Cif. 1962 (H)

Anungitea B. Sutton 1973 (H)

Anungitopsis R.F. Castañeda & W.B.

Kendr. 1990 (H)

Aoria Cif. 1962 (C)

Aphanofalx B. Sutton 1986 (C)

Apiocarpella Syd. & P. Syd. 1919 (C)

Apogloeum Petr. 1954 (C)

Aporellula B. Sutton 1986 (C)

Aposporella Thaxt. 1920 (H)

Arachnophora Hennebert 1963 (H)

Arachnospora R.F. Castañeda, Minter &

Camino 2003 (H)

Arborillus Munt.-Cvetk. & Gómez-Bolea

1998 (H)

Arborispora K. Ando 1986 (H)

Arbusculina Marvanová & Descals 1987

Mycosphere Doi 10.5943/mycosphere/3/2/5

196

(H)

Arcuadendron Sigler & J.W. Carmich.

1976 (H)

Ardhachandra Subram. & Sudha 1978 (H)

Argopericonia B. Sutton & Pascoe 1987

(H)

Aristastoma Tehon 1933 (C)

Arthrobotryum Ces. 1854 (C)

Arthrocladium Papendorf 1969 (H)

Arthrocristula Sigler, M.T. Dunn & J.W.

Carmich. 1982 (H)

Arthrosporium Sacc. 1880 (H)

Arthrowallemia R.F. Castañeda, Dania

García & Guarro 1998 (H)

Articulophora C.J.K. Wang & B. Sutton

1982 (H)

Artocarpomyces Subram. 1996 (H)

Arxiella Papendorf 1967 (H)

Asbolisiomyces Bat. & H. Maia 1961 (C)

Ascochytopsis Henn. 1905 (C)

Ascochytulina Petr. 1922 (C)

Ascospora Mont. 1849 (C)

Ashtaangam Subram. 1995 (H)

Aspilaima Bat. & H. Maia 1961 (C)

Astelechia Cif. 1962 (H)

Asterinothyriella Bat. & Cif. 1959 (C)

Asterinothyrium Bat., Cif. & H. Maia 1959

(C)

Asteroconium Syd. & P. Syd. 1903 (C)

Asteromyces F. Moreau & V. Moreau, in

Hennenert 1962 (H)

Asterostomopora Bat. & H. Maia 1960 (C)

Asterostomopsis Bat., Cif. & H. Maia 1959

(C)

Asterostomulina Bat., J.L. Bezerra & H.

Maia 1964 (C)

Astronatelia Bat. & H. Maia 1962 (C)

Atractilina Dearn. & Barthol. 1924 (H)

Atrosetiphiale Matsush. 1995 (H)

Aurantiosacculus Dyko & B. Sutton 1979

(C)

Aurosphaeria Sun J. Lee, Strobel, Eisen-

man, Geary, Vargas & S.A. Strobel

2009 (C)

Avesicladiella W.P. Wu, B. Sutton &

Gange 1997 (H)

Avettaea Petr. & Syd. 1927 (C)

Bachmanniomyces D. Hawksw. 1981 (C)

Bacillopeltis Bat. 1957 (C)

Bactridiopsis Henn. 1904 (H)

Bactridium Kunze 1817 (H)

Bactrodesmiastrum Hol.-Jech 1984 (H)

Bactrodesmiella M.B. Ellis 1959 (H)

Badarisama Kunwar, J.B. Manandhar &

J.B. Sinclair 1986 (H)

Bahuchashaka Subram. 1978 (H)

Bahugada K.A. Reddy & Vasant Rao

1984 (H)

Bahukalasa Subram. & Chandrash. 1979

(H)

Bahusandhika Subram. 1956 (H)

Balaniopsis P.M. Kirk 1985 (H)

Balanium Wallr. 1833 (H)

Barbarosporina Ķirulis 1942 (C)

Barnettella D. Rao & P.Rag. Rao 1964 (C)

Basauxia Subram. 1995 (H)

Batistina Peres 1961 (C)

Beauveriphora Matsush 1975 (H)

Beccopycnidium F. Stevens 1930 (C)

Beejadwaya Subram. 1978 (H)

Belemnospora P.M. Kirk 1981 (H)

Bellulicauda B. Sutton 1967 (C)

Beltrania Penz. 1882 (H)

Beltraniomyces Monohar, D.K. Agarwal &

Rao 2003 (H)

Beltraniopsis Bat. & J.L. Bezerra 1960 (H)

Beniowskia Racib. 1900 (H)

Benjaminia S. Ahmad 1967 (C)

Benjpalia Subram. & Bhat 1989 (H)

Bharatheeya D'Souza & Bhat 2002 (H)

Bhatia W.A. Baker & Morgan-Jones 2009

(H)

Bibanasiella R.F. Castañeda & W.B.

Kendr. 1991 (H)

Biflagellospora Matsush. 1975 (H)

Biflagellosporella Matsush. 1993 (H)

Bimeris Petr. 1949 (C)

Bioconiosporium Bat. & J.L. Bezerra 1964

(H)

Biophomopsis Petr. 1931 (C)

Bispora Corda 1837 (H)

Bisseomyces R.F. Castañeda 1985 (H)

Blarneya D. Hawksw., Coppins & P.

James 1980 (H)

Blastacervulus H.J. Swart 1988 (C)

Blastocatena Subram. & Bhat 1989 (H)

Blastodictys M.B. Ellis 1976 (H)

Blastofusarioides Matsush. 1996 (H)

Blastophorella Boedijn 1937 (H)

Blastophorum Matsush. 1971 (H)

Blastophragma Subram. 1995 (H)

Blennoria Moug. & Fr. 1825 (C)

Blennoriopsis Petr. 1919 (C)

Blodgettia Harv. 1858 (H)

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197

Bomplandiella Speg. 1886 (H)

Bostrichonema Ces. 1867 (H)

Botryoderma Papendorf & H.P. Upadhyay

1969 (H)

Botryodiplodina Dias & Sousa da Câmara

1954 (C)

Botryomonilia Goos & Piroz. 1975 (H)

Botryosporium Corda 1831 (H)

Brachydesmiella G. Arnaud ex S. Hughes

1961 (H)

Brachysporiellina Subram. & Bhat 1989

(H)

Brachysporiopsis Yanna, W.H. Ho & K.D.

Hyde 2004 (H)

Brefeldiopycnis Petr. & Cif. 1932 (H)

Brevicatenospora R.F. Castañeda, Minter

& Saikawa 2006 (H)

Briosia Cavara 1888 (H)

Brycekendrickia Nag Raj 1973 (C)

Bulbilopycnis Matsush. 1996 (C)

Bulbocatenospora R.F. Castañeda & Iturr.

2000 (H)

Bullaserpens Bat., J.L. Bezerra & Cavalc.

1965 (C)

Calcarispora Marvanová & Marvan 1963

(H)

Calcarisporiella de Hoog 1974 (H)

Calceispora Matsush. 1975 (H)

Callistospora Petr. 1955 (C)

Calocline Syd. 1939 (C)

?Calongia D. Hawksw. & Etayo 2010 (H)

Camarographium Bubák 1916 (C)

Camaropycnis E.K. Cash 1945 (C)

Camarosporellum Tassi 1902 (C)

Camarosporiopsis Abbas, B. Sutton &

Ghaffar 2000 (C)

Camposporidium Nawawi & Kuthub. 1988

(H)

Camptomeris Syd. 1927 (H)

Campylospora Ranzoni 1953 (H)

Candelabrum Beverw. 1951 (H)

Candelosynnema K.D. Hyde & Seifert

1992 (H)

Towards incorporating anamorphic fungi in a natural classification - Checklist and notes for 2011

Abstract