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Mycosphere Doi 10.5943/mycosphere/3/2/5
157
Towards incorporating anamorphic fungi in a natural classification – checklist
and notes for 2011
Wijayawardene DNN1,2, McKenzie EHC3 and Hyde KD1,2
1Institute of Excellence in Fungal Research, Mae Fah Luang University, Chiang Rai 57100, Thailand
2School of Science, Mae Fah Luang University, Chiang Rai 57100, Thailand
3Manaaki Whenua Landcare Research, Private Bag 92170, Auckland, New Zealand
Wijayawardene DNN, McKenzie EHC, Hyde KD 2012 – Towards incorporating anamorphic fungi
in a natural classification – checklist and notes for 2011. Mycosphere 3(2), 157-228, Doi 10.5943
/mycosphere/3/2/5
A complilation of anamorphic names for both Ascomycota and Basidiomycota is provided which
comprises 2895 genera. The genera are listed against a backbone of teleomorphic relationships
where known. The study reveals that 73 genera and 95 anamorph-like genera are linked to
teleomorphic genera names, 447 genera (three anamorph-like genera) are linked to teleomorph
families, orders or classes, while for more than 1592 (55%) genera no teleomorph link is known.
The links are based on the literature and often because fungi were found in association, but have not
been proven by molecular data. Many are based on links with species other than the generic type
and thus must be considered questionable for the genus. A considerable effort is needed to establish
whether these links are correct.
Key words – Asexual fungi – life cycle – sexual fungi – taxonomy
Article Information
Received 24 March 2012
Accepted 29 March 2012
Published online 10 April 2012
Corresponding author: KD Hyde – e-mail – kdhyde3@gmail.com
Introduction
The purpose of the present paper is to
collate the knowledge of asexual fungi in
relationship to their sexual states. It attempts to
classify the asexual genera in a natural
biological framework for the Ascomycota
based on the Outline of Ascomycota 2009
(Lumbsch & Huhndorf 2010) and for the
Basidiomycota based on Kirk et al. (2008). The
present paper has used data from the previous
outline of Hyde et al. (2010) as well as
literature published in 2011 and updated with
opinions in Seifert et al. (2011).
There is now a mandate in the syste-
matics community to place one name on a
holomorphic fungus species as opposed to the
previous system (especially in Ascomycota) of
naming both sexual and asexual morphs
(Samuels et al. 2009, Hawksworth 2011;
Hawksworth et al. 2011, Taylor 2011; Wingf-
ield et al. 2011). Also, as more and more
strains of asexual genera are sequenced
(Shenoy et al. 2010, Põldmaa 2011, Reblova &
Seifert 2011), these genera will be classified in
families, orders and classes of Ascomycota and
Basidiomycota. This will result in more and
more asexual states being described and illustr-
ated without being given formal names and
therefore it is important to have a forum where
asexual data can be brought to the attention of
researchers. It is hoped that this compilation
will facilitate the move towards one fungus one
name (Shenoy et al. 2007, 2010, Hawksworth
et al. 2011). By placing all asexual genera
against the backbone classification of the sexu-
al genera it will be possible to establish, 1)
Mycosphere Doi 10.5943/mycosphere/3/2/5
158
what are the sexual states of various species,
genera,families or orders, 2) which genera have
no information, 3) which are the older or more
commonly used names for each genus, and 4)
illustrate the redundancy of the dual classif-
ication system.
This compilation also provides a single
place where asexual connections can be access-
ed and critically scrutinized so that a stable and
reliable system can be developed based on pub-
lished data. We do not claim that this compi-
lation is complete or definitive; however, it
provides a forum for displaying data and can
be added to or changed as more data becomes
available or errors are discovered. In future
issues we will explore various connections and
provide more data.
In the long term it will be desirable if
Myconet (http://www.fieldmuseum.org/myco-
net) is merged with this document, however,
this should occur at some time in the future,
when more data is available.
Material and methods
The overall scheme for the classification
of the sexual fungi follows the Outline of
Ascomycota 2009 (Lumbsch & Huhndorf
2010), with additional data on anamorphic
Basidiomycota from Kirk et al. (2008). This
scheme is chosen as the schemes in the
Dictionary of the Fungi, Index Fungorum, and
MycoBank differ to a greater or lesser extent
and it was felt wise to follow what the authors
consider is an admirable attempt to keep
abreast of the current literarure as is routinely
carried out in the Outline of Ascomycota. This
compilation also annotates changes made in the
form of notes on asexual fungi (marked with an
asterisk after the entry, with notes in the later
“Notes” section) so that changes can easily be
followed; this is not always true of the
available databases.
Asexual genera or ‘-like’ genera are
listed in bold under the appropriate fungal
classification and the link to a teleomorph
genus given where found. Dubious genera are
excluded but can be added if they are later
found to be good genera. Fossil fungi are also
excluded as it would be impossible with
present techniques available to provide a na-
tural classification for them. A question mark
before the entry means that its placement in
that family or order is not fully established and
generally follows Lumbsch & Huhndorf
(2010). A question mark before anamorph
means it is not clear that this is the anamorph
of this genus, although there are many cases
without a question mark where the link has not
equivocally been proven. An asterisk following
the names indicates an entry in the “Notes”
section.
In many genus records in Index Fungo-
rum the species is linked to more than one
anamorphic genus. Such records can be located
by searching the genus throughout the list for
repeat entries. The synonyms of genera are not
listed as they can be easily found in Index
Fungorum.
We encourage colleagues to inform us of
any missing entries, new data or misunder-
standings, which will be updated in future
compilations. We would also invite experts of
specific groups to contribute to future compi-
lations on their specific group(s) of interest and
encourage reviews or illustrated accounts of
families, orders or other groups relating to
sexual and asexual links. Reviews of asexual
genera with sections or series, such as
Aspergillus, Penicillium and Verticillium, the
powdery mildews, many of the yeasts and
yeast-like genera and the asexual smuts and
rusts are particularly needed. Similarly the
lichenized ascomycete asexual states are poorly
documented.
As there is mandate towards using one
name for both the sexual and asexual states of a
fungus we also encourage authors to submit
articles or opinions concerning the use of either
name. This might be in the form of a general
critic, arguments for or against using one name,
or an article that specifically addresses an
individual group or genus.
In the present outline we have added
(H) = hyphomycete, (C) = coelomycete, (Y) =
yeast and (R) = Rust against entries where
known. Although this grouping is artificial we
do this to facilitate finding data on either as the
groups are generally dealt with in different text.
For example, data for the hyphomycetes have
recently been complied and updated by Seifert
et al. (2011) and the new data for hyphomycet-
es included here have been extracted from this
tome. Data for coelomycetes were extracted
from Sutton (1980), Nag Raj (1993), http://
Mycosphere Doi 10.5943/mycosphere/3/2/5
159
www. Cybertr uffle.org.uk/cybernome/eng/ind-
ex.htm (Accesion date 18.03.2011) and http://-
www.landcarer esearch .co.nz/html /data asp?
ID = &NAMEKey = 26885 (Accesion date
6.12.2011).
Index to Ascomycota
Phylum ASCOMYCOTA Caval-Sm.
Subphylum TAPHRINOMYCOTINA O.E.
Erikss. & Winka
Class Taphrinomycetes O.E. Erikss. & Winka
Taphrinales Gäum. & C.W. Dodge
Protomycetaceae Gray
Saitoella Goto, Sugiy., Hamam. & Komag.
1987 (Y)
Taphrinaceae Gäum.
Lalaria R.T. Moore 1990, anamorphic
Taphrina Fr. 1815 (Y)
Subphylum SACHAROMYCOTINA O.E.
Erikss. & Winka
Class Saccharomycetes O.E. Erikss. & Winka
Saccharomycetales Kudrjanzev
Dipodascaceae Engl. & E. Gilg
Blastobotrys Klopotek 1967, anamorphic
Trichomonascus H.S. Jacks. 1948 (H)
?Candida Berkhout 1923, anamorphic
Sporopachydermia Rodr. Mir. 1978 (Y)
Geotrichum Link 1809, anamorphic
Galactomyces Redhead & Malloch
1977 and Dipodascus Lagerh. 1892 (H)
Geotrichum-like, anamorphic Basidioas-
cus Matsush. 2003 (H)
Saprochaete Coker & Shanor 1939, ana-
morphic Magnusiomyces Zender 1926
(H)
Endomycetaceae J. Schröt.
Fusidium Link 1809, possibly anamorphic
Ascocephalophora K. Matsush. &
Matsush. 1995 (H)
Lipomycetaceae E.K. Novák & Zsolt
Myxozyma Van der Walt, Weijman & Arx
1981 (Y)
Metschnikowiaceae T. Kamienski
Candida Berkhout 1923, anamorphic
Clavispora Rodr. Mir. 1979 and
Metschnikowia T. Kamienski 1899 (Y)
Pichiaceae Zender
Brettanomyces N.H. Claussen ex Custers
1940, anamorphic Dekkera Van der
Walt 1964 (Y)
Eeniella M.T. Sm., Bat. Vegte & Scheffers
1981, anamorphic Dekkera Van der
Walt (Y)
Enantiothamnus Pinoy 1911 (Y)
Hyphopichia Arx & Van der Walt 1976
(Y)
Saccharomycetaceae G. Winter
?Candida Berkhout 1923, anamorphic
Ogataea Y. Yamada, K. Maeda &
Mikata 1994, Lodderomyces Van der
Walt 1966 and Spathaspora N.H.
Nguyen, S.O. Suh & M. Blackw. 2006
(Y)
Saccharomycodaceae Kudrjanzev
Candida Berkhout 1923, anamorphic
Lodderomyces Van der Walt 1966 and
Spathaspora N.H. Nguyen, S.O. Suh &
M. Blackw. 2006 (Y)
Kloeckera Janke 1923, anamorphic Han-
seniaspora Zikes 1911 (Y)
Trichomonascaceae Kurtzman & Robnett
?Candida Berkhout 1923, anamorphic
Zygoascus M.T. Sm. 1986 (Y)
Saccharomycetales, genera incertae sedis
Aciculoconidium D.S. King & S.C. Jong
1976 (Y)
Botryozyma Shann & M.T. Sm. 1992,
anamorphic Ascobotryozyma J. Kerri-
gan, M.T. Sm. & J.D. Rogers 2001 (Y)
Candida Berkhout 1923, anamorphic
Starmerella C.A. Rosa & Lachance
1998 (Y)
Cicadomyces Šulc 1911 (Y)
Macrorhabdus Tomasz., Logan, Snowden,
Kurtzman & Phalen 2003 (Y)
Oosporidium Stautz 1931 (Y)
Pseudomycoderma H. Will 1916 (Y)
Schizoblastosporion Cif. 1930 (Y)
Trigonopsis Schachner 1929 (Y)
Saccharomycetes, genera incertae sedis
Selenotila Lagerh. 1892 (Y)
Mycosphere Doi 10.5943/mycosphere/3/2/5
160
Selenozyma Yarrow 1977 (Y)
Subphylum PEZIZOMYCOTINA O.E.
Erikss. & Winka
Class Arthoniomycetes O.E. Erikss. & Winka
Arthoniales Henssen ex D. Hawksw. & O.E.
Erikss.
Arthoniaceae Reichenb. ex Reichenb.
Helicobolomyces Matzer 1995, anamor-
phic Arthonia Ach. 1806 (C)
?Septocyta Petr. 1927, anamorphic Artho-
nia Ach. 1806 (C)
Subhysteropycnis Wedin & Hafellner
1998, anamorphic Arthonia Ach. 1806
(C)
Roccellaceae Chevall.
Sporhaplus H.B.P. Upadhyay 1964, ana-
morphic Mazosia A. Massal 1854 (C)
Class Dothideomycetes sensu O.E. Erikss. &
Winka
Subclass Dothideomycetidae P.M. Kirk, P.F.
Cannon, J.C. David & J.A. Stalpers ex C.L.
Schoch, Spatafora, Crous & Shoemaker
Capnodiales Woron.
Antennulariellaceae Woron.
Capnodendron S. Hughes 1976, anamor-
phic Antennulariella Woron. 1915 (H)
Capnofrasera S. Hughes 2003 (H)
Heteroconium Petr. 1949, anamorphic
Antennulariella Woron. 1915 (H)
Capnodiaceae (Sacc.) Höhn. ex Theiss.
Acanthorus Bat. & Cavalc. 1967 (C)
?Antennariella Bat. & Cif. 1963 (C)*
Apiosporium Kunze 1817 (C)
Conidiocarpus Woron. 1927, anamorphic
Phragmocapnias Theiss. & Syd. 1918
and Scorias Fr. 1825 (C)*
Fumagospora G. Arnaud 1911, anam-
orphic Capnodium Mont. 1848 (C)
?Fumiglobus D.R. Reynolds & G.S.
Gilbert 2006 (C)*
Microxiphium (Harv. ex Berk. & Desm.)
Thüm. 1879 (as Microxyphium) (H)*
?Mycogelidium W.Y. Zhuang 2007 (C)
Phaeoxyphiella Bat. & Cif. 1963, ana-
morphic Capnodium Mont. 1848 (C)*
?Polychaetella Speg. 1918, anamorphic
Capnodium Mont. 1848 (C)*
Polychaeton (Pers.) Lev. 1846 (C)*
Scolecoxyphium Cif. & Bat. 1956,
anamorphic Capnodium Mont. 1848
and Scorias Fr. 1825 (C)*
Tripospermum Speg. 1918 anamorphic
Triposporiopsis W.Yamam 1955 (H)*
Coccodiniaceae Höhn. ex O.E. Erikss.
Bisbyopeltis Bat. & A.F. Vital 1957 (C)
Microxiphium (Harv. ex Berk. & Desm.)
Thüm. 1879, anamorphic Dennisiella
Bat. & Cif. 1962 (H)*
Davidiellaceae C.L. Schoch, Spatafora, Crous
& Shoemaker
Acroconidiella J.C. Lindq. & Alippi 1964,
anamorphic Davidiella Crous & U.
Braun 2003 (H)
Cladosporium Link 1816, anamorphic
Davidiella Crous & U. Braun 2003 (H)
Dichocladosporium K. Schub., U. Braun
& Crous 2007 (C)
Graphiopsis Trail 1889, anamorphic
Davidiella Crous & U. Braun 2003 (H)
Hoornsmania Crous 2007 (H)
Rachicladosporium Crous, U. Braun &
C.F. Hill 2007 (H)
Toxicocladosporium Crous & U. Braun
2007 (H)
Verrucocladosporium K. Schub., Aptroot
& Crous 2007 (H)
Dissoconiaceae Crous & de Hoog
Dissoconium de Hoog, Oorschot & Hijwe-
gen 1983 (H)
Ramichloridium Stahel 1937 ex de Hoog
1977 (H)
Metacapnodiaceae S. Hughes & Corlett
Capnobotrys S. Hughes 1970, anamorphic
Metacapnodium Speg. 1918 (H)
Capnocybe S. Hughes 1966, anamorphic
Metacapnodium Speg. 1918 (H)
Capnophialophora S. Hughes 1966, ana-
morphic Metacapnodium Speg. 1918
(H)
Capnosporium S. Hughes 1976, anamor-
phic Metacapnodium Speg. 1918 (H)
Hormiokrypsis Bat. & Nascim. 1957, ana-
morphic Metacapnodium Speg. 1918
and Ophiocapnocoma Bat. & Cif. 1963
(H)
Torulopsiella Bender 1932, anamorphic
Mycosphere Doi 10.5943/mycosphere/3/2/5
161
Ophiocapnocoma Bat. & Cif. 1963 and
Metacapnodium Speg. 1918 (H)
Mycosphaerellaceae Lindau
Anguillosporella U. Braun 1995 (H)
Asperisporium Maubl. 1913 (H)
Asteromella-like, anamorphic Gillotia
Sacc. & Trotter 1913 (C)
Cercospora Fresen. 1863, anamorphic
Mycosphaerella Johanson 1884, Sphae-
rulina Sacc. and others (H)
Cercosporella Sacc. 1880, anamorphic
Mycosphaerella Johanson 1884 (H)
?Clypeispora A.W. Ramaley 1991, ana-
morphic Mycosphaerella Johanson
1884 (C)
Colletogloeum Petr. 1953, anamorphic
Mycosphaerella Johanson 1884 (C)
Coniothryium-like, anamorphic Bruneos-
phaerella Crous 2009 (C)
Deightoniella S. Hughes 1952 (H)
Didymochora Höhn. 1918, anamorphic
Euryachora Fuckel 1870 (C)
Distocercospora N. Pons & B. Sutton
1988 (H)
Dothistroma Hulbary 1941 (C)
Floricola Kohlm. & Volkm.Kohlm. 2000
(C)
Fusicladiella Höhn. 1919, anamorphic
Mycosphaerella Johanson 1884 (H)
Laocoön J.C. David 1997 (H)
Lecanosticta Syd. 1922 (C)
Microcyclospora Jana Frank, Schroers &
Crous 2010 (H)
Microcyclosporella Jana Frank, Schroers
& Crous 2010 (H)
Miuraea Hara 1948, anamorphic Myco-
sphaerella Johanson 1884 (H)
Mycopappus Redhead & G.P. White 1985,
anamorphic Mycosphaerella Johanson
1884 (H)
Mycovellosiella Rangel 1918, anamorphic
Mycosphaerella Johanson 1884 (H)
Parastenella J.C. David 1991 (H)
Passalora Fr. 1849, anamorphic Myco-
sphaerella Johanson 1884 (H)
Penidiella Crous & U. Braun 2007 (H)
Periconiella Sacc. 1885, anamorphic
Allosoma Syd. 1926 (H)
Phacellium Bonord. 1860, anamorphic
Mycosphaerella Johanson 1884 (H)
Phaeoisariopsis Ferraris 1909, anamorphic
Mycosphaerella Johanson 1884 (H)
Phaeophleospora Rangel 1916 (C)
Phaeothecoidea Crous 2007 (H)
Phloeospora Wallr. 1833, anamorphic
Mycosphaerella Johanson 1884 (C)
Placosphaeria-like, anamorphic Euryach-
ora Fuckel 1870 (C)
Polythrincium Kunze 1817, anamorphic
Cymadothea F.A. Wolf 1935 (H)
Pseudocercospora Speg. 1910 anamorphic
Mycosphaerella Johanson 1884 (H)
Pseudocercosporella Deighton 1973, ana-
morphic Mycosphaerella Johanson
1884 (H)
Pseudocercosporidium Deighton 1973 (H)
Quasiphloeospora B. Sutton, Crous &
Shamoun 1996 (H)
Ramichloridium Stahel ex de Hoog 1977
(H)
Ramichloridium-like, anamorphic Mycosp-
haerella Johanson 1884 (H)
Ramularia Unger 1833, anamorphic Myc-
osphaerella Johanson 1884 and
Melanodothis R.H. Arnold 1972 (H)
Ramulariopsis Speg. 1910, anamorphic
Mycosphaerella Johanson 1884 (H)
Ramulispora Miura 1920, anamorphic
Mycosphaerella Johanson 1884 (H)
Rhabdospora (Durieu & Mont. ex Sacc.)
Sacc. 1884 (C)
Rhexocercosporidium U. Braun 1994 (H)
Semipseudocercospora J.M. Yen 1983 (H)
Septoria Sacc. 1884 (C)
Septoria-like, anamorphic Mycosphaerella
Johanson 1884 (C)
Sirosporium Bubák & Serebrian. 1912 (H)
Sonderhenia H.J. Swart & J. Walker 1988
(C)
Stenella Syd. 1930, anamorphic Myco-
sphaerella Johanson 1884 (H)
Stenellopsis B. Huguenin 1966 (H)
Stigmina Sacc. 1980, anamorphic Myco-
sphaerella Johanson 1884 (H)
Thedgonia B. Sutton 1973, anamorphic
Mycosphaerella Johanson 1884 (H)
Thedgonia-like (H)
Trochophora R.T. Moore 1955 (H)
Verrucisporota D.E. Shaw & Alcorn 1993
(H)
Xenostigmina Crous 1998, anamorphic
Mycosphaerella Johanson 1884 or
Didymella Sacc. ex D. Sacc. 1880 (H)
Mycosphere Doi 10.5943/mycosphere/3/2/5
162
Zasmidium Fr. 1849, anamorphic Mycosp-
haerella-like and Rasutoria M.E. Barr
1987 (H)
Zymoseptoria Quaedvlieg & Crous 2011
(C)*
Capnodiales, genera incertae sedis
Capnocheirides J.L. Crane & S. Hughes
1982 (H)
Heptaster Cif., Bat. & Nascim. 1956 (H)
Hormonema-like, anamorphic Xenomeris
Syd. 1924 (H)
Houjia G.Y. Sun & Crous 2010 (H)
Micropustulomyces R.W. Barreto 1995 (C)
Phaeotheca Sigler, Tsuneda & J.W.
Carmich. 1981 (H)
Phaeothecoidiella Batzer & Crous 2010
(H)
Pseudovirgaria H.D. Shin, U. Braun,
Arzanlou & Crous 2007 (H)
Rachicladosporium Crous, U. Braun &
C.F. Hill 2007 (H)
Recurvomyces Selbmann & de Hoog 2008
(H)
Scolecostigmina U. Braun 1999 (H)
Sporidesmajora Batzer & Crous 2010 (H)
Verrucocladosporium K. Schub., Aptroot
& Crous 2007 (H)
Dothideales Lindau
Dothideaceae Chevall.
Lecanosticta Syd. 1922, anamorphic Sci-
rrhia Nitschke ex Fuckel 1870 (C)
Podoplaconema Petr. 1921, anamorphic
Omphalospora Theiss. & Syd. 1915 (C)
Dothioraceae Theiss. & H. Syd.
Aureobasidium Viala & G. Boyer 1891,
anamorphic Columnosphaeria Munk
1953 and Saccothecium Fr. 1836 (H)
Dothichiza Lib. ex Roum. 1880, ana-
morphic Dothiora Fr. 1849 and Sydo-
wia Bres. 1895 (C)
Hormonema Lagerb. & Melin 1927, ana-
morphic Dothiora Fr. 1849, Prings-
heimia Schulzer 1866 and Sydowia
Bres. 1895 (H)
Japonia Höhn. 1909, anamorphic Yoshina-
gaia Henn. 1904 (C)
Kabatina R. Schneid. & Arx 1966 (C)
?Rhizosphaera L. Mangin & Har. 1907,
anamorphic Phaeocryptopus Naumov
1915 and Dimerina Theiss. 1912 (C)
Sclerophoma Höhn. 1909, anamorphic
Sydowia Bres. 1895 (C)
Teratosphaeriaceae Crous & U. Braun
Acidomyces B.J. Baker, M.A. Lutz, S.C.
Dawson, P.L. Bond & Banfield ex
Selbmann, de Hoog & De Leo 2008 (H)
Batcheloromyces Marasas, P.S. van Wyk
& Knox-Dav. 1975 (H)
Baudoinia J.A. Scott & Unter. 2007 (H)
Capnobotryella Sugiy. 1987 (H)
Catenulostroma Crous & U. Braun 2007,
anamorphic Teratosphaeria Syd. & P.
Syd. 1912 (H)
Cibiessia Crous 2007 (H)
Colletogloeopsis Crous & M.J. Wingf.
1997, anamorphic Teratosphaeria Syd.
& P. Syd. 1912 (C)
Davisoniella H.J. Swart 1988 (C)
Devriesia Seifert & N.L. Nick. 2004 (H)
Elasticomyces Zucconi & Selbmann 2008
(H)
Friedmanniomyces Onofri 1999 (H)
Hispidoconidioma Tsuneda & Davey 2010
(H)
Hobsoniopsis D. Hawksw. 2001 (H)
Hortaea Nishim. & Miyaji 1984 (H)
Kirramyces J. Walker, B. Sutton & Pascoe
1992, anamorphic Teratosphaeria Syd.
& P. Syd. 1912 (C)
Nothostrasseria Nag Raj 1983 (C)
Passalora Fr. 1849 (H)
Penidiella Crous & U. Braun 2007 (H)
Phacellium Bonord. 1860 (H)
Phaeothecoidea Crous 2007 (H)
Pseudocercosporella Deighton 1973 (H)
Pseudoramichloridium Cheewangkoon &
Crous 2010 (H)
Pseudotaeniolina J.L. Crane & Schokn.
1986 (H)
Ramichloridium Stahel ex de Hoog 1977
(H)
Recurvomyces Selbmann & de Hoog 2008
(H)
Readeriella Syd. & P. Syd. 1908 (C)
Sporidesmium Link 1809 (H)
Staninwardia B. Sutton 1971 (C)
Stenella Syd. 1930 (H)
Trimmatostroma-like, anamorphic Terato-
sphaeria Syd. & P. Syd. 1912 (H)
Tripospermum Speg. 1918 (H)
Mycosphere Doi 10.5943/mycosphere/3/2/5
163
Myriangiales Starbäck
Elsinoaceae Höhn. ex Sacc. & Trotter
Endosporium Tsuneda 2008 (H)
Sphaceloma de Bary 1874 anamorphic
Elsinoë Racib. 1900 (C)
Xenodiella Syd. 1935, anamorphic Xeno-
dium Syd. 1935 (H)
Myriangiales, genera incertae sedis
Phaeosclera Sigler, Tsuneda & J.W. Car-
mich. 1981 (H)
Sarcinomyces Lindner 1898 (H)
Zopfiaceae G. Arnaud ex D. Hawksw. 1992
Cladosporium-like, anamorphic Zopfia Rab-
enh. 1874 (H)
Subclass Pleosporomycetidae C.L. Schoch,
Spatafora, Crous & Shoemaker
Pleosporales Luttrell ex M.E. Barr
Amniculicolaceae Yin. Zhang, C.L. Schoch, J.
Fourn., Crous & K.D. Hyde
Anguillospora Ingold 1942 (H)
Spirosphaera Beverw. 1953 (H)
Cucurbitariaceae G. Winter
Camarosporium Schulzer 1870, anamor-
phic Cucurbitaria Gray 1821 (C)
Coniothyrium-like, anamorphic Curreya
Sacc. 1883 (C)
Diplodia-like, anamorphic Cucurbitaria
Gray 1821 (C)
?Megaloseptoria Naumov 1925, anamor-
phic Cucurbitaria Gray 1821 (C)
?Phaeoseptoria Speg. 1908, anamorphic
Rhytidiella Zalasky 1968 (C)
Phialophorophoma Linder 1944 (C)
Phoma Sacc. 1880 (C)
Pleurophoma Höhn. 1914 (C)
Pleurostromella Petr. 1922, anamorphic
Cucurbitaria Gray 1821 and Gibberi-
dea Fuckel 1870 (C)
Pyrenochaeta De Not. 1849, anamorphic
Cucurbitaria Gray 1821 (C)
Pyrenochaetopsis Gruyter, Aveskamp, &
Verkley, 2010 (C)
Syntholus A.W. Ramaley & M.E. Barr
1997, anamorphic Syncarpella Theiss.
& Syd. 1915 (C)
Diademaceae Shoemaker & C.E. Babc.
Alternaria-like, anamorphic Comoclathris
Clem. 1909 (H)*
Fusicladiella Höhn. 1919, anamorphic
Didymosphaeria Fuckel 1870 (H)
Didymellaceae Gruyter, Aveskamp & Verkley
Ascochyta Lib. 1830, anamorphic Didy-
mella Sacc. ex D. Sacc. 1880 (C)
Ampelomyces Ces. ex Schltdl. 1852 (C)
Boeremia Aveskamp, Gruyter & Verkley
2010 (C)
Chaetasbolisia Speg. 1918 (C)
Dactuliochaeta G.L. Hartm. & J.B. Sincl-
air 1988, anamorphic Didymella Sacc.
1880 (C)
Microsphaeropsis Höhn. 1917 (C)
Peyronellaea Gold. ex Togliani 1952 (C)
Phoma Sacc. 1880, anamorphic Didymella
Sacc. ex D. Sacc. 1880 (C)
Piggotia Berk. & Broome 1851, possibly
anamorphic Platychora Petr. 1925 (C)
Pithomyces Berk. & Broome 1875, ana-
morphic Leptosphaerulina McAlpine
1902 (H)
Stagonosporopsis Died. 1912, anamorphic
Didymella Sacc. ex D. Sacc. 1880 (C)
(=Ascochyta Lib. 1830)
Didymosphaeriaceae Munk
Cytoplea Bizz. & Sacc. 1885, anamorphic
Roussoëlla Sacc. 1888 (C)
Dothidotthiaceae Crous & A.J.L. Phillips
?Dothiorella Sacc. 1880, anamorphic Dot-
hidotthia Höhn. 1918 (C)
Thyrostroma Höhn. 1911, anamorphic
Dothidotthia Höhn. 1918 (H)
Hypsostromataceae Huhndorf
Pleurophomopsis-like, anamorphic Hyp-
sostroma Huhndorf (C)
Lentitheciaceae Yin. Zhang, C.L. Schoch, J.
Fourn., Crous & K.D. Hyde
Stagonospora (Sacc.) Sacc. 1884 (C)
Leptosphaeriaceae M.E. Barr
Camposporium Harkn. 1884, anamorphic
Leptosphaeria Ces. & De Not. 1863
(H)
Coniothyrium Corda 1840, anamorphic
Leptosphaeria Ces. & De Not. 1863 (C)
Coniothyrium-like, anamorphic Neophaeo-
Mycosphere Doi 10.5943/mycosphere/3/2/5
164
sphaeria M.P.S. Câmara, M.E. Palm &
A.W. Ramaley 2003 (C)
Phoma Sacc. 1880, anamorphic Leptosph-
aeria Ces. & De Not. 1863 (C)
Plenodomus Preuss 1851 (C)
Pyrenochaeta De Not. 1849, possibly
anamorphic Leptosphaeria Ces. & De
Not. 1863 (C)
Lindgomycetaceae K. Hiray., Kaz. Tanaka &
Shearer
Taeniolella S. Hughes 1958 (H)
Lolia Abdel-Aziz & Abdel-Whab 2010
(C)*
Lophiostomataceae Sacc.
Anguillospora Ingold 1942, anamorphic
Massarina Sacc. 1883 (H)
Ascochyta Lib. 1830, anamorphic Tricho-
metasphaeria Munk 1953 (C)
Periconia Tode 1791 anamorphic Lophio-
stoma Ces. & De Not. 1863 (H)
Pleuorphomopsis-like, anamorphic Lophi-
ostoma Ces. & De Not. 1863 (C)
Pyricularia Sacc. 1880, anamorphic Mass-
arina Sacc. 1883 (H)
Tumularia Descals & Marvanová 1987(H)
Massariaceae Nitschke
?Aplosporella-like, anamorphic Dubitatio
Speg. 1882 (C)
Myxocyclus Riess 1852, anamorphic Mass-
aria De Not. 1844 (C)
Torula Pers. 1794 (H)
Massarinaceae Munk
Aquaticheirospora Kodsueb & W.H. Ho
2007 (H)
Ceratophoma Höhn. 1917, anamorphic
Massarina Sacc. 1883 (C)
Cheirosporium L. Cai & K.D. Hyde 2008
(H)
Clavariopsis De Wild. 1895 (H)
Helminthosporium Link 1809, anamorphic
Splanchnonema Corda 1829 (H)
Neottiosporina Subram. 1961 (C)
Pseudodictyosporium Matsush. 1971 (H)
Melanommataceae G. Winter
Aposphaeria Sacc. 1880 (C)
Beverwykella Tubaki 1975 (H)
Exosporiella P. Karst. 1892, anamorphic
Anomalemma Sivan. 1983 (H)
Nigrolentilocus R.F. Castañeda & Heredia
2001, anamorphic Melanomma Nitsch-
ke ex Fuckel 1870 (H)
Monodictys S. Hughes 1958, anamorphic
Ohleria Fuckel 1868 (H)
Monotosporella S. Hughes 1958 (H)
Pyrenochaeta De Not. 1849, anamorphic
Byssosphaeria Cooke 1879 (C)
Sporidesmiella P.M. Kirk 1982 (H)
Montagnulaceae M.E. Barr
Aschersonia Mont. 1848, anamorphic
Montagnula Berl. 1896 (C)
Microdiplodia Allesch. 1901, possibly
anamorphic Karstenula Speg. 1879 (C)
Microsphaeropsis Höhn. 1917, anamorph-
ic Paraphaeosphaeria O.E. Erikss.
1967 (C)
Paraconiothyrium Verkley 2004, possibly
anamorphic Paraphaeosphaeria O.E.
Erikss. 1967 (C)
Phaeosphaeriaceae M.E. Barr
Amarenographium O.E. Erikss. 1982, ana-
morphic Amarenomyces O.E. Erikss.
1981 (recently synonymised with
Phaeosphaeria I. Miyake 1909) (C)
Ampelomyces Ces. ex Schltdl. 1852 (C)
Chaetosphaeronema Moesz 1915 (C)
Coniothyrium Corda 1840, anamorphic
Ophiobolus Riess 1854 (C)
Harpophora W. Gams 2000, anamorphic
anamorphic Ophiobolus Riess 1854 (H)
Hendersonia-like, anamorphic Austropleo-
spora R.G. Shivas & L. Morin 2010
and Phaeosphaeria I. Miyake 1909 (C)
Mauginiella Cavara 1925 (H)
Neosetophoma Gruyter, Aveskamp &
Verkley 2010 (C)
?Parahendersonia A.W. Ramaley 1995,
anamorphic Chaetoplea (Sacc.) Clem.
1931 (C)
Paraphoma Morgan-Jones & J.F. White
1983 (C)
Phaeoseptoria Speg. 1908, anamorphic
Phaeosphaeria I. Miyake 1909 (C)
Phaeostagonospora A.W. Ramaley 1997,
anamorphic Phaeosphaeriopsis M.P.S.
Câmara, M.E. Palm & A.W. Ramaley
2003 (C)
Phoma Sacc. 1880, anamorphic Ophio-
Mycosphere Doi 10.5943/mycosphere/3/2/5
165
bolus Riess 1854 (C)
Rhabdospora (Durieu & Mont. ex Sacc.)
Sacc. 1884, anamorphic Ophiobolus
Riess 1854 (C)
Scolecosporiella Petr. 1921, anamorphic
Ophiosphaerella Speg. and Phaeo-
sphaeria I. Miyake 1909 (C)
Setophoma Gruyter, Aveskamp & Verkley
2010 (C)
?Sphaerellopsis Cooke 1883, anamorphic
Eudarluca Speg. 1908 (C)
Stagonospora (Sacc.) Sacc. 1884, anamor-
phic Phaeosphaeria I. Miyake 1909 (C)
Tiarospora Sacc. & Marchal 1885 (C)
Pleomassariaceae M.E. Barr
Ceuthodiplospora Died. 1912, anamorphic
Splanchnonema Corda (C)
Corynespora Güssow 1906, anamorphic
Pleomassaria Speg. 1880 (H)
Myxocyclus Riess 1852, anamorphic
Splanchnonema Corda 1829 (C)
?Prosthemium Kunze 1817, anamorphic
Pleomassaria Speg. 1880 (C)
Scolicosporium Lib. ex Roum. 1880, ana-
morphic Asteromassaria Höhn. 1917
(C)
Shearia Petr. 1924, anamorphic Pleomas-
saria Speg. 1880 (C)
Stegonsporium Corda 1827, anamorphic
Splanchnonema Corda 1829 (C)
Pleosporaceae Nitschke
Alternaria Nees 1816, anamorphic Lewia
M.E. Barr & E.G. Simmons 1986 (H)*
Alternaria-like anamorphic Clathrospora
Rabenh. 1857 (H)
Alternariaster E.G. Simmons 2007 (H)
Brachycladium Corda 1838, anamorphic
Crivellia Shoemaker & Inderbitzin
2006 (H)
Bipolaris Shoemaker 1959, anamorphic
Cochliobolus Drechsler 193 (H)
Chalastospora E.G. Simmons 2007 (H)
Corynespora Güssow 1906, anamorphic
Pyrenophora Fr. 1849 (H)
Curvularia Boedijn 1933, anamorphic
Cochliobolus Drechsler 1934 (H)
Dactuliophora C.L. Leakey 1964 (H)
Dendryphiella Bubák & Ranoj. 1914,
anamorphic Pleospora Rabenh. ex Ces.
& De Not. 1863 (H)
Dendryphion Wallr. 1833, anamorphic
Pleospora Rabenh. ex Ces. & De Not.
1863 (H)
Dictyosporium Corda 1836 (H)
Digitodesmium P.M. Kirk 1981 (H)
Drechslera S. Ito 1930, anamorphic Pyre-
nophora Fr. 1849 (H)
Edenia M.C. González, Anaya, Glenn,
Saucedo & Hanlin 2007 (H)
Embellisia E.G. Simmons 1971, anamor-
phic Lewia M.E. Barr & E.G. Simmons
(H)
Exserohilum K.J. Leonard & Suggs 1974,
anamorphic Setosphaeria K.J. Leonard
& Suggs 1974 (H)
Gibbago E.G. Simmons 1986 (H)
Lemonniera De Wild. 1894 (H)
Marielliottia Shoemaker 1999, anamor-
phic Pyrenophora Fr. 1849 (H)
Nimbya E.G. Simmons 1989, anamorphic
Macrospora Fuckel 1870 (H)
Prathoda Subram. 1956 (H)
Sinomyces Yong Wang bis & X.G. Zhang
2011 (H)
Stemphylium Wallr. 1833, anamorphic Ple-
ospora Rabenh. ex Ces. & De Not.
1863 (H)
Teretispora E.G. Simmons 2007 (H)
Ulocladium Preuss 1851, anamorphic Las-
iobotrys Kunze 1823 (H)
Undifilum Pryor, Creamer, Shoemaker,
McLain-Romero & Hambl. 2009 (H)
Sporormiaceae Munk
?Amorosia Mantle & D. Hawksw. 2006
(H)
Phoma Sacc. 1880 (C)
Phoma-like, anamorphic Westerdykella
Stolk 1955 (C)
Teichosporaceae M.E. Barr
Coniothyrium-like, anamorphic Immothia
M.E. Barr (C)
Tetraplosphaeriaceae Kaz. Tanaka & K. Hiray
Pseudotetraploa Kaz. Tanaka & K.
Hirayama 2009 (H)
Quadricrura Kaz. Tanaka, K. Hirayama &
Sat. Hatak. 2009 (H)
Tetraploa Berk. & Broome 1850, ana-
morphic Tetraplosphaeria Kaz. Tanaka
& K. Hirayama 2009 (H)
Mycosphere Doi 10.5943/mycosphere/3/2/5
166
Tetraploa-like, anamorphic Triplosphaeria
Kaz. Tanaka & K. Hirayama 2009 and
Polyplosphaeria Kaz. Tanaka & K.
Hirayama 2009 (H)
Trematosphaeriaceae K.D. Hyde, Yin. Zhang,
Suetrong & E.B.G. Jones
Hyphopodia-like, anamorphic Tremato-
sphaeria Fuckel, Jb. nassau. Ver.
Naturk (C)*
Pleosporales, genera incertae sedis
Berkleasmium Zobel 1854 (H)
Briansuttonia R.F. Castañeda, Minter &
Saikawa 2004 (H)
Cerebella Ces. 1851 (H)
Cheiromoniliophora Tzean & J.L. Chen
1990 (H)
Diplococcium Grove 1885 (H)
Ellisembia Subram. 1992 (H)
Epicoccum Link 1815 (H)
Fusculina Crous & Summerell 2006 (C)
Mycocentrospora Deighton 1972 (H)
Mycopappus Redhead & G.P. White 1985
anamorphic Mycodidymella C.Z. Wei,
Y. Harada & Katum. 1998 (H)
Noosia Crous, R.G. Shivas & McTaggart
2011 (H)
Ochrocladosporium Crous & U. Braun
2007 (H)
Phoma Sacc. 1880 (C)
Periconia Tode 1791(H)
Pleurophomopsis Petr. 1924, anamorphic
Astrosphaeriella Syd. & P. Syd. 1913
(C)
Polyschema H.P. Upadhyay 1966 (H)
Pseudochaetosphaeronema Punith. 1979
(C)
Pseudopassalora Crous 2011 (H)
Repetophragma Subram. 1992 (H)*
?Scleroramularia Batzer & Crous 2011
(H)
Sclerostagonospora Höhn. 1917 (C)
Sirodesmium De Not. 1849 (C)
Spiroplana Voglmayr, M.J. Park & H.D.
Shin 2011 (H)
Sporidesmium Link 1809 (C)
Versicolorisporium Sat. Hatak., Kaz.
Tanaka & Y. Harada 2008 (C)
Ybotromyces Rulamort 1986 (H)
Dothideales genera incertae sedis
Atramixtia Tsuneda, M.L. Davey &
Currah 2011 (H)
Celosporium Tsuneda & Davey 2010 (H)
Diplococcium Grove 1885, anamorphic
Otthia Nitschke ex Fuckel 1870 (H)
Pleosporomycetidae, genera incertae sedis
Acrogenospora M.B. Ellis 1971, anamor-
phic Farlowiella Sacc. 1891(H)
Dothideomycetes, orders incertae sedis
Acrospermales Minter, Peredo & A.T. Watson
Acrospermaceae Fuckel
Dactylaria Sacc. 1880, anamorphic Acro-
spermum Tode 1783 (H)
Gonatophragmium Deighton 1969, ana-
morphic Acrospermum Tode 1790 (H)
Botryosphaeriales C.L. Schoch, Crous &
Shoemaker
Botryosphaeriaceae Theiss. & H. Syd.
Alysidiella Crous 2006 (H)
Aplosporella Speg. 1880 (C)
Arthrinium Kunze 1817, anamorphic Lept-
oguignardia E. Müll. (H)
Bahusutrabeeja Subram. & Bhat 1977 (H)
Barriopsis A.J.L. Phillips, A. Alves &
Crous 2008 (C)
Dichomera Cooke 1878 (C)
Diplodia Fr. 1834 (C)
Diplodia-like, anamorphic Neodeightonia
C. Booth 1970 and Saccharata Denman
& Crous 2004 (C)
Dothichiza-like, anamorphic Leptoguig-
nardia E. Müll. 1955 (C)
Dothiorella Sacc. 1880, anamorphic Spen-
cermartinsia A.J.L. Phillips, A. Alves
& Crous 2008 (C)
Fusicoccum Corda 1829, anamorphic
Botryosphaeria Ces. & De Not. 1863
(C)
Fusicoccum-like, anamorphic Melanops
Nitschke ex Fuckel 1870 and Saccha-
rata Denman & Crous 2004 (C)
Hormonema Lagerb. & Melin 1927,
anamorphic Guignardia Viala & Ravaz
1892 (H)
Lasiodiplodia Ellis & Everh. 1896 (C)
Leptodothiorella Höhn. 1918, anamorphic
Guignardia Viala & Ravaz 1892 (C)
Macrophomina Petr. 1923 (C)
Microdiplodia Allesch. 1901 (C)
Mycosphere Doi 10.5943/mycosphere/3/2/5
167
Neofusicoccum Crous, Slippers & A.J.L.
Phillips 2006, with Botryosphaeria-like
teleomorphs (C)
Neoscytalidium Crous & Slippers 2006
(H)
Phyllosticta Pers. 1818, anamorphic Guig-
nardia Viala & Ravaz 1892 (C)
Pseudofusicoccum Mohali, Slippers &
M.J. Wingf. 2006 (C)
Sphaeropsis Sacc. 1880 (C)
Thyrostroma Höhn. 1911, anamorphic
Dothidotthia Höhn. 1918 (H)
Botryosphaeriales, genera incertae sedis
Camarosporium Schulzer 1870 (C)
Dichomera Cooke 1878 (C)
Hendersonula Speg. 1880 (C)
Hysteriales Lindau
Hysteriaceae Chevall.
Aposphaeria-like, anamorphic Gloniopsis
De Not. 1847 and Hysterobrevium E.
Boehm & C.L. Schoch 2010 (C)
Coniosporium Link 1809, anamorphic
Hysterium Pers. 1797 (H)
Plenodomus Preuss 1851, anamorphic
Psiloglonium Höhn. 1918 (C)
Septonema Corda 1837, anamorphic Oedo-
hysterium E. Boehm & C.L. Schoch
2009 (H)
Sporidesmium Link 1809, anamorphic
Psiloglonium Höhn. 1918 (H)
Jahnulales Pang, Abdel-Wahab, El-
Sharouney, E.B.G. Jones & Sivichai
Aliquandostipitaceae Inderbitzin
Brachiosphaera Nawawi 1976 (H)
Mytilinidiales Boehm, C.L. Schoch &
Spatafora
Gloniaceae (Corda) Boehm, C.L. Schoch &
Spatafora
Cenococcum Moug. & Fr. 1829 (C)
Cleistonium Speer 1986, anamorphic
Glonium Mühl. 1813 (C)
Mytilinidiaceae Kirschst.
Camaroglobulus Speer 1986, anamorphic
Mytilinidion Duby 1862 (C)
Chalara (Corda) Rabenh. 1844, anam-
orphic Quasiconcha M.E. Barr & M.
Blackw. 1981 (H)*
?Monotosporella S. Hughes 1958, anam-
orphic Actidium Fr. 1815 (H)
Papulospora Preuss 1851, anamorphic
Lophium Fr. 1818 (H)
Septonema Corda 1837, anamorphic
Mytilinidion Duby 1862 (H)
Taeniolella S. Hughes 1958, anamorphic
Mytilinidion Duby 1861 (H)
Jahnulales genera incertae sedis
Xylomyces Goos, R.D. Brooks & Lamore
1977 (H)
Patellariales D. Hawksw. & O.E. Erikss.
Patellariaceae Corda
Aposphaeria-like, anamorphic Rhytidhy-
steron Speg. 1881 (C)
Diplodia-like, anamorphic Rhytidhysteron
Speg. 1881 (C)
Venturiales Yin. Zhang, C.L. Schoch & K.D.
Hyde
Venturiaceae E. Müll. & Arx ex M.E. Barr
Acroconidiellina (Berk. & Broome) M.B.
Ellis 1971, anamorphic Acantharia
Theiss. & Syd. (H)*
Bactrodesmium Cooke 1883, anamorphic
Phragmogibbera Samuels & Rog-
erson 1990 (H)
Dictyodochium Sivan. 1984, anamorphic
Gibbera Fr. 1825 (H)
?Didymochora Hohn. 1918, anamorphic
Atopospora Petr. 1925 (C)
Cladosporium-like, anamorphic Apiosp-
orina Höhn. 1910 (H)
Fusicladium Bonord. 1851, anamorphic
Apiosporina Höhn. 1910, Caprov-
enturia Braun 1998 and Venturia Sacc.
1882 (H)
Fusicladium-like, anamorphic Acantharia
Theiss. & Syd 1918 and Protove-
nturia Berl. & Sacc. 1887 (H)
Fusicladosporium Partr. & Morgan-Jones
2003, anamorphic Venturia Sacc.
1882 (H)
Helicodendron Peyronel 1918 anamorphic
Tyrannosorus Unter. & Malloch 1995
(H)
Pithosira Petr. 1949 (H)
Spilocaea Fr. 1819, anamorphic Venturia
Sacc. 1882 (C)
Spilodochium Syd. 1927 anamorphic Pseu-
Mycosphere Doi 10.5943/mycosphere/3/2/5
168
doparodiella F. Stevens 1927 (H)
Stigmina-like, Sacc. 1980 anamorphic
Aca-ntharia Theiss. & Syd 1918 and
Gib-bera Fr. 1825 (H)
Sympoventuriaceae Yin. Zhang, C.L. Schoch
& K.D. Hyde
Fusicladium-like (H)
Sympodiella-like, anamorphic Sympoven-
turia Crous & Seifert (H)*
Veronaeopsis Arzanlou & Crous 2007
(H)*
Dothideomycetes, families incertae sedis
Ascoporiaceae Kutorga & D. Hawksw.
Plectophomella-like, anamorphic Pseudo-
solidum Lloyd 1923 (H)
Asterinaceae Hansf.
?Acarella Syd. 1927, anamorphic More-
noina Theiss. 1913 (C)
Asterostomella Speg. 1886, anamorphic
Asterina Lév. 1845 (C)
Asterostromina Bat. & A.F. Vital 1957,
anamorphic Asterodothis Theiss. (C)
Asterostomula Theiss. 1916, anamorphic
Echidnodes Theiss. & Syd. 1918 (C)
Bahusakala Subram. 1958 (H)
?Clasterosporium Schwein. 1832, anamo-
rphic Eupelte Syd. 1924 (H)
Clasterosporium Schwein. 1832, anam-
orphic Asterina Lév. 1845, Asterodothis
Theiss.1912, Yamamotoa Bat. 1960 and
Lembosia Lév. 1845 (H)
?Elachopeltis Syd. 1927, anamorphic Aph-
anopeltis Syd. 1927 (C)
Eriothyrium Speg. 1888, anamorphic
Aster-inema Bat. & Gayão 1953 (C)*
Leprieurina G. Arnaud 1918, anamorphic
Prillieuxina G. Arnaud 1918 (C)
Mahanteshamomyces Hosag. & C.K. Biju
2004 (C)
?Peltasterella Bat. & H. Maia 1959, ana-
morphic Yamamotoa Bat. 1960 (C)
Pirozynskia Subram. 1972, anamorphic
Eupelte Syd. 1924 (H)
Septoidium G. Arnaud 1921, anamorphic
Eupelte Syd. 1924 (H)
Septothyrella Höhn. 1911, anamorphic
Uleothyrium Petr. 1929 (C)
Shivamyces Hosag. 2004 (H)
Sirothyriella Höhn. 1910, anamorphic
Morenoina Thesis. 1913 (C)
?Thyrinula Petr. & Syd. 1924, anamorphic
Aulographina Arx & E. Müll. 1960 (C)
Triposporium Corda 1837, anamorphic
Batistinula Arx 1960 (H)
Corynesporascaceae Sivan.
Corynespora Güssow 1906, anamorphic
Corynesporasca Sivan. (H)
Dacampiaceae Körb.
Cyclothyrium Petr. 1923, anamorphic Pol-
ycoccum Saut. ex Körb. 1865 (C)
Lecythophora-like, anamorphic Munkoval-
saria Aptroot 1995 (H)
Englerulaceae Henn.
Capnodiastrum Speg. 1886, anamorphic
Englerula Henn. 1904 and Rhyti-
denglerula Höhn. 1918 (C)
Digitosarcinella S. Hughes 1984, anamo-
rphic Schiffnerula Höhn. 1909 (H)
Mitteriella Syd. 1933, anamorphic Schiff-
nerula Höhn. 1909 (H)
Questieriella G. Arnaud ex S. Hughes
1983, anamorphic Schiffnerula Höhn.
1909 (H)
Sarcinella Sacc. 1880, anamorphic Schiff-
nerula Höhn. 1909 (H)*
Eremomycetaceae Malloch & Cain
Arthrographis G. Cochet ex Sigler & J.W.
Carmich. 1976, anamorphic Eremomy-
ces Malloch & Cain 1971and Pitho-
ascina Valmaseda, A.T. Martínez &
Barrasa 1987 (H)
Trichosporiella Kamyschko 1960, anam-
orphic Rhexothecium Samson &
Mouch. 1975 and Eremomyces Malloch
& Cain 1971 (H)
Euantennariaceae Hughes & Corlett
Antennatula Fr. ex F. Strauss 1850, anam-
orphic Euantennaria Speg. and Strig-
opodia Bat. 1957 (H)
Capnokyma S. Hughes 1975 (H)
Capnophialophora S. Hughes 1966, ana-
morphic Strigopodia Bat. 1957 (H)
Hormisciomyces Bat. & Nascim. 1957,
anamorphic Euantennaria Speg. (H)
Mycosphere Doi 10.5943/mycosphere/3/2/5
169
Hyphosoma Syd. 1924, anamorphic Strig-
opodia Bat. 1957 (H)
Plokamidomyces Bat., C.A.A. Costa &
Cif. 1957, anamorphic Trichopeltheca
Bat., C.A.A. Costa & Cif. 1958 (H)
Racodium Fr. 1829, anamorphic Strigo-
podia Bat. 1957 (H)
Trichothallus F. Stevens 1925, anamorphic
Trichopeltheca Bat., C.A.A. Costa &
Cif. 1958 (H)
Fenestellaceae M.E. Barr
Pleurostromella Petr. 1922, anamorphic
Fenestella Tul. & C. Tul. 1863 (C)
Leptopeltidaceae Höhn. ex Trotter
?Idriella P.E. Nelson & S. Wilh. 1956,
anamorphic Dothiopeltis E. Mull. 1957
(H)
Leptothyrium-like, anamorphic Leptopeltis
Höhn. 1917 (C)
Meliolinaceae S. Hughes
Briania D.R. Reynolds 1989, anamorphic
Meliolina Syd. & P. Syd. 1914 (H)
Micropeltidaceae Clem. & Shear
Cyclopeltella Petr. 1953, anamorphic Cycl-
opeltis Petr. 1953 (C)
Heliocephala V.G. Rao, K.A. Reddy & de
Hoog 1984 (H)*
Holubovaniella R.F. Castañeda 1985 (H)*
Sirothyriella Höhn. 1910, anamorphic
Stomiopeltis Theiss. 1914 (C)
Sporidesmium-like, anamorphic Stomio-
peltis Theiss. 1914 (as Akaropeltella
M.L. Farr, 1972) (H)
Pycnothyriaceous taxa associated with spe-
cies of Stomiopeltis anamorphic Micro-
peltis Mont.
Microthyriaceae Sacc.
Asterostomula Theiss. 1916, anamorphic
Asterinella Theiss. 1912 (C)
Asteromella-like, anamorphic Asterinella
Theiss. 1912 (C)
Hansfordiella S. Hughes 1951, anamor-
phic Trichothyrium Speg. 1889 (H)
Isthmospora F. Stevens 1918, anamorphic
Trichothyrium Speg. 1889 (H)
Leptothyrium Kunze 1823, anamorphic
Microthyrium Desm. 1841 (C)
Xenogliocladiopsis Crous & W.B. Kendr.
1994, anamorphic Arnaudiella Petr.
1927 (H)
Zalerion R.T. Moore & Meyers 196, ana-
morphic Microthyrium Desm. 1841 (H)
Parmulariaceae E. Müll. & Arx ex M.E. Barr
Excipulariopsis P.M. Kirk & Spooner
1982, anamorphic Kentingia Sivan. &
W.H. Hsieh 1989 (H)
?Melanoplaca Syd. & P. Syd. 1917,
anamorphic Dothidasteroma Höhn.
1909 (C)
Placomelan Cif. 1962, anamorphic Doth-
idasteroma Höhn. 1909 (C)
Parodiellaceae Theiss. & H. Syd. ex M.E. Barr
?Ascochytopsis Henn. 1905, anamorphic
Parodiella Speg. 1880 (C)
Parodiopsidaceae Toro
?Chuppia Deighton 1965, anamorphic
Neoparodia Petr. & Cif. 1932 (H)
Cicinnobella Henn. 1904, anamorphic
Perisporiopsis Henn. 1904 (as Phaeo-
dimeriella Speg. 1908) and Dimerium
(Sacc. & P. Syd.) Sacc. & D. Sacc.
1905 (C)
Clasterosporium Schwein. 1832, anamor-
phic Balladyna Racib. 1900 (H)
Heterosporiopsis Petr. 1950, anamorphic
‘Dimerium’ (Sacc. & P. Syd.) Sacc. &
D. Sacc. 1905, Exosporinella Bender
1932, possibly anamorphic Parodiellina
Henn. ex G. Arnaud 1918 (H)
?Ophiotrichum Kunze 1849 anamorphic
Leptomeliola Höhn. 1919 (H)
Sarcinella-like, anamorphic Neoparodia
Petr. & Cif. 1932 (H)
Septoidium G. Arnaud 1921, anamorphic
Alina Racib. 1909, Perisporiopsis
Henn. 1904, Parodiellina Henn. ex G.
Arnaud 1918, Chevalieropsis G.
Arnaud 1923, Ophioparodia Petr. &
Cif. 1932 and Pilgeriella Henn. 1900
(H)
Shivamyces Hosag. 2004, anamorphic
Alina Racib. 1909 (H)
Tretospora M.B. Ellis 1976, anamorphic
Balladyna Racib. and Balladynopsis
Mycosphere Doi 10.5943/mycosphere/3/2/5
170
Theiss. & Syd. (H)
Planistromellaceae M.E. Barr
Aposphaeria-like, anamorphic Microcycl-
us Sacc., Syd. & P. Syd. 1904 (C)
Fusicladium Bonord. 1851, anamorphic
Microcyclus Sacc., Syd. & P. Syd. 1904
(H)
Hyphospora A.W. Ramaley 1996, ana-
morphic Comminutispora A.W. Rama-
ley 1996 (H)
Kellermania Ellis & Everh. 1885, ana-
morphic Planistroma A.W. Ramaley
1991 (C)
Lecanosticta Syd. 1922, anamorphic Erup-
tio M.E. Barr 1996 (C)
Pazschkeella Syd. & P. Syd. 1901, anam-
orphic Microcyclus Sacc., Syd. & P.
Syd. 1904 (C)
Piptarthron Mont. ex Höhn. 1918, anam-
orphic Planistroma A.W. Ramaley
1991 (C)
Polystomellaceae Theiss. & H. Syd.
Asteromella Pass. & Thüm. 1880 (as
Stictochorella Höhn. 1917), anamorphic
Dothidella Speg. 1880 (C)
Lasmenia Speg. 1886, anamorphic Mun-
kiella Speg. 1885 (C)
Pseudoperisporiaceae Toro
Chaetosticta Petr. & Syd. 1925, anamor-
phic Lasiostemma Theiss., Syd. & P.
Syd. 1917 and ?Nematostoma Syd. & P.
Syd. 1914 (C)
Schizothyriaceae Höhn. ex Trotter, Sacc., D.
Sacc. & Traverso
Plenotrichaius Bat. & Valle 1961, anamor-
phic Mycerema Bat., J.L. Bezerra &
Cavalc 1963 (C)
Zygophiala E.W. Mason 1945, anamorphic
Schizothyrium Desm. 1849 (H)
Tubeufiaceae M.E. Barr
?Annellospermosporella P.R. Johnst. 1999,
anamorphic Malacaria Syd. 1930 (H)
Aquaphila Goh, K.D. Hyde & W.H. Ho
1998, anamorphic Tubeufia Penz. &
Sacc. 1898 (H)*
Araneomyces Höhn. 1909, anamorphic
Paranectriella (P. Henn. & Sacc.)
Höhn. (H)
Chionomyces Deighton & Piroz. 1972,
anamorphic Melioliphila Speg. 1924
(H)
Corynespora Güssow 1906, anamorphic
Chaetosphaerulina I. Hino 1938 (H)
Eriomycopsis Speg. 1910, anamorphic
Melioliphila Speg. 1924 and Byssocallis
Syd. 1927 (H)
Guelichia Speg. 1886, anamorphic Putte-
mansia Henn. 1902 (H)
Helicoma Corda 1837, anamorphic Tubeu-
fia Penz. & Sacc. 1898, Thaxteriella
Petr. 1924 and Chlamydotubeufia Boo-
nmee & K.D. Hyde 2011 (H)*
Helicoön Morgan 1892 (H)*
Helicomyces Link 1809, anamorphic
Acanthostigma De Not. (1863) (H)*
Helicosporium Nees 1816, anamorphic
Acanthostigma De Not. (1863) and
Tubeufia Penz. & Sacc. 1898 (H)
Kamalomyces R.K. Verma, N. Sharma &
Soni 2008 (C)
Mirandina G. Arnaud ex Matsush. 1975,
anamorphic Taphrophila Scheuer 1988
(H)*
Monodictys S. Hughes 1958, anamorphic
Tubeufia Penz. & Sacc. 1898 (H)
Monodictys-like, anamorphic Tubeufia
Penz. & Sacc. 1898 (H)
Moorella P. Rag. Rao & D. Rao 1964,
anamorphic Chaetosphaerulina I. Hino
1938 (H)
Pendulispora M.B. Ellis 1961, anamorphic
Tubeufia Penz. & Sacc. 1898 (H)
Peziotrichum (Sacc.) Lindau 1900, ana-
morphic Podonectria Petch 1921 (H)
Spermatoloncha Speg. 1908 (H)
Tetracrium Henn. 1902, anamorphic Putt-
emansia Henn. 1902 and Podonectria
Petch 1921 (H)
Titaea Sacc. 1876, anamorphic Paranec-
triella (Henn. ex Sacc. & D. Sacc.)
Höhn. 1910 and Puttemansia Henn.
1902 (H)
Xenosporium Penz. & Sacc. 1901, anamor-
phic Acanthostigmella Höhn. 1905 and
Chaetosphaerulina I. Hino 1938 (as
Mycosphere Doi 10.5943/mycosphere/3/2/5
171
Thaxteriellopsis Sivan., Panwar & S.J.
Kaur 1977) (H)
Vizellaceae H.J. Swart
Chrysogloeum Petr. 1959, anamorphic
Blasdalea Sacc. & P. Syd. 1902 (C)
Manginula G. Arnaud 1918, anamorphic
Vizella Sacc. 1883 (C)
Dothideomycetes, genera incertae sedis
Acrocalymma Alcorn & J.A.G. Irwin
(1987), anamorphic Massarina, how-
ever Massarina has been shown to be
polyphyletic and its placement is there-
fore presently unclear (C)
Ampullifera Deighton 1960, anamorphic
Teratoschaeta Bat. & O.M. Fonseca
(H)
Ascochyta-like, anamorphic Gilletiella
Sacc. & P. Syd. 1899 (C)
Asteromella Pass. & Thüm. 1880 (C)
Bactrodesmium Cooke 1883, anamorphic
Stuartella Fabre 1879 (H)
Brachyconidiella R.F. Castañeda & W.B.
Kendr. 1990 (H)
Camarosporula Petr. 1954, anamorphic
Anthracostroma Petr. 1954 (C)
Chaetasbolisia Speg. 1918, anamorphic
Pseudomorfea Punith. 1981 (C)
Cladoriella Crous 2006 (H)
Coniosporium Link 1809 (H)
Coronospora M.B. Ellis 1971, anamorphic
Ascocoronospora Matsush. 2003 (H)
Cryomyces Selbmann, de Hoog, Mazzagl-
ia, Friedmann & Onofri 2005 (H)
?Cyclothyrium Petr. 1923, anamorphic
Thyridaria Sacc. 1875 (C)
Dendryphiopsis S. Hughes 1953, anamorp-
hic Kirschsteiniothelia D. Hawksw.
1985 (H)
Diederichia D. Hawksw. 2003 (C)
Dilophospora Desm. 1840, anamorphic
Lidophia J. Walker & B. Sutton 1974
(C)
Diplodia-like, anamorphic Otthia Nitschke
ex Fuckel 1870 (C)
Disculina Höhn. 1916 (C)
Funbolia Crous & Seifert 2011 (H)
Hansfordiellopsis Deighton 1960, anamor-
phic Koordersiella Höhn. 1909 (H)
Helicodendron Peyronel 1918, anamorphic
Tyrannosorus Unter. & Malloch 1995
(H)
Hiospira R.T. Moore 1962, anamorphic
Brooksia Hansf. 1956 (H)
Iledon Samuels & J.D. Rogers 1986, ana-
morphic Botryohypoxylon Samuels &
J.D. Rogers 1986 (C)
Megaloseptoria Naumov 1925 (C)
Mirandina G. Arnaud ex Matsush. 1975,
anamorphic Chaetonectrioides Matsu-
sh. 1996 (H)
Monodictys S. Hughes 1958 (H)
Peltaster Syd. & P. Syd. 1917 (C)
Phaeothecoidiella Batzer & Crous 2010
(H)
Phanerococculus Cif. 1954, anamorphic
Koordersiella Höhn. 1909 (C)
Phialophora-like, anamorphic Tirisporella
E.B.G. Jones, K.D. Hyde & Alias 1996
(H)
Phoma-like, anamorphic Tremateia Koh-
lm., Volkm.-Kohlm. & O.E. Erikss.
1995 (C)
Placosphaeria (De Not.) Sacc. 1880 (C)
Plectopycnis Bat. & A.F. Vital 1959 (C)
Pleurostromella Petr. 1922, anamorphic
Gibberidea Fuckel 1870 (C)
Pseudorobillarda M. Morelet 1968 (C)
Pycnopleiospora C.Z. Wei, Y. Harada &
Katum. 1997, anamorphic Pseudodidy-
mella C.Z. Wei, Y. Harada & Katum.
1997 (H)
Rhizopycnis D.F. Farr 1998 (C)
Scleroconidioma Tsuneda, Currah & Thor-
mann 2000 (H)
Septoriella Oudem. 1889 (C)
Troposporella P. Karst. 1892 (H)
Class Eurotiomycetes Tehler ex O.E. Eriksson
& K. Winka
Subclass Chaetothyriomycetidae Doweld
Chaetothyriales M.E. Barr
Chaetothyriaceae Hansf. ex M.E. Barr
?Ciferrioxyphium Bat. & H. Maia 1963,
anamorphic Aithaloderma P. Syd. (H)*
Cyphellophora G.A. de Vries 1962 (H)
Leptoxyphium Speg. 1918, anamorphic Ai-
thaloderma P. Syd. 1913 (H)*
Merismella Syd. 1927, anamorphic Chaet-
Mycosphere Doi 10.5943/mycosphere/3/2/5
172
othyrium Speg. 1888 (C)
Stanhughesia Constant. 1989, anamorphic
Ceramothyrium Bat. & H. Maia 1956
(H)
Tripospermum Speg. 1918, anamorphic
Phragmocapnias Theiss. & Syd. 1918,
Trichomerium Speg. 1918 and Tripo-
sporiopsis W. Yamam. 1955 (H)*
Vonarxia Bat. 1960 (H)
Herpotrichiellaceae Munk
Brycekendrickomyces Crous & M.J.
Wingf. 2009 (H)
Cladophialophora Borelli 1980, anamor-
phic Capronia Sacc. 1883 (H)
Exophiala J.W. Carmich. 1966, anamor-
phic Capronia Sacc. 1883 (H)
Fonsecaea Negroni 1936, anamorphic
Capronia Sacc. 1883 (H)
Metulocladosporiella Crous, Schroers, J.Z.
Groenew., U. Braun & K. Schub. 2006
(H)
Nadsoniella Issatsch. 1914 (H)
Phaeococcomyces de Hoog 1979 (H)
Phaeoramularia Munt.-Cvetk. 1960, ana-
morphic Capronia Sacc.1883 (H)
Phialophora Medlar 1915 (H)
Racodium Pers. 1794, anamorphic Capro-
nia Sacc. 1883 (H)
Rhinocladiella Nannf. 1934, anamorphic
Capronia Sacc. 1883 (H)
Sorocybe Fr. 1849 (H)
Thysanorea Arzanlou, W. Gams & Crous
2007 (H)
Veronaea Cif. & Montemart. 1957 (H)
Chaetothyriales M.E. Barr, genera incertae
sedis
Coniosporium Link 1809, the type species
is considered as Sirodesmium De Not.
1849 (H)
Phaeoannellomyces McGinnis & Schell
1985 (H)(= Hortaea)
Staninwardia B. Sutton 1971 (C)
Strelitziana Arzanlou & Crous 2006 (H)
Pyrenulales, genera incertae sedis
Lyromma Bat. & H. Maia 1965 (C)
Chaetothyriomycetidae, families incertae sedis
Rhynchostomataceae Winka & O. E. Erikss.
Arthropycnis Constant. 1992, anamorphic
Rhynchostoma P. Karst. (C)
Strigulaceae Zahlbr.
Discosiella Syd. & P. Syd. 1912, anamor-
phic Strigula Fr. 1823 (C)
Chaetothyriomycetidae, genera incertae sedis
Peyronelia Cif. & Gonz. Frag. 1927, ana-
morphic Glyphium Nitschke ex F.
Lehm. 1886 (H)
Phaeomoniella Crous & W. Gams 2000
(H)
Rhabdospora-like, anamorphic Dolabra C.
Booth & W.P. Ting 1964 (C)
Taeniolella S. Hughes 1958, anamorphic
Glyphium Nitschke ex F. Lehm. 1886
(H)
Subclass Eurotiomycetidae Geiser & Lutzoni
Coryneliales Seaver & Chardon
Coryneliaceae Sacc. ex Berl. & Voglino
Exophiala J.W. Carmich. 1966, anamor-
phic Bicornispora Checa, Barrasa,
M.N. Blanco & A.T. Martínez 1996 (H)
Eurotiales G.W. Martin ex Benny & Kimbr.
Monascaceae J. Schröt.
Basipetospora G.T. Cole & W.B. Kendr.
1968, anamorphic Monascus Tiegh.
1884 (H)
Fraseriella Cif. & A.M. Corte 1957, ana-
morphic Xeromyces Fraser (H)
Trichocomaceae E. Fisch.
Aspergillus P. Micheli ex Link 1809,
anamorphic Dichlaena Durieu & Mont.
1849, Emericella Berk. & Broome
1857, Eurotium Link 1809, Fennellia
B.J. Wiley & E.G. Simmons 1973,
Hemicarpenteles A.K. Sarbhoy &
Elphick 1968, Neopetromyces Frisvad
& Samson 2000, Neosartorya Malloch
& Cain 1973, Petromyces Malloch &
Cain 1973 and Warcupiella Subram.
1972 (H)*
Geosmithia J.Pitt 1979, anamorphic, Chro-
mocleista Yaguchi & Udagawa 1993
and Talaromyces C.R. Benj. 1955 (H)
Isaria Pers. 1794, anamorphic Byssochla-
mys Westling (H)
Mycosphere Doi 10.5943/mycosphere/3/2/5
173
Merimbla Pitt 1979, anamorphic Talar-
omyces C.R. Benj. 1955 (H)
Paecilomyces Bainier 1907, anamorphic
Byssochlamys Westling 1909 and Tala-
romyces C.R. Benj. 1955 (H)
Paecilomyces-like, anamorphic Cooneme-
ria Mouch. 1997 (H)
Penicillium Link 1809, anamorphic Eupe-
nicillium F. Ludw. 1892, Talaromyces
C.R. Benj. 1955, Dichlaena Durieu &
Mont. 1849 and Trichocoma Jungh.
1838 (H)
Phialosimplex Sigler, Deanna A. Sutton,
Gibas, Summerb. & Iwen 2010 (H)
Polypaecilum G. Sm. 1961, anamorphic
Dactylomyces Sopp 1912, Dichotomo-
myces Saito ex D.B.Scott 1970 and
Thermoascus Miehe 1907 (H)
Pseudocordyceps Hauman 1936, anamor-
phic Penicilliopsis Solms 1887 (H)
Sagenomella W. Gams 1978, anamorphic
Sagenoma Stolk & G.F. Orr 1974 and
Talaromyces C.R. Benj. 1955 (H)
Sarophorum Syd. & P. Syd. 1916, ana-
morphic Penicilliopsis Solms 1887 (H)
Stilbodendron Syd. & P. Syd. 1916, ana-
morphic Penicilliopsis Solms 1887 (H)
Stilbothamnium Henn. 1896 (H)
Thermomyces Tsikl. 1899 (H)
Thysanophora W.B. Kendr. 1961 (H)
Torulomyces Delitsch 1943, anamorphic
Eupenicillium F. Ludw. 1892 (H)
Eurotiomycetidae genera incertae sedis
Penicillium Link 1809, anamorphic Para-
talaromyces Matsush. 2003 (H)
Onygenales Cif. ex Benny & Kimbr.
Ajellomycetaceae Untereiner, J.A. Scott &
Sigler
Blastomyces Gilchrist & W.R. Stokes
1898, anamorphic Ajellomyces McDo-
nough & A.L. Lewis 1968 (H)
Chrysosporium Corda 1833, anamorphic
Ajellomyces McDonough & A.L. Lewis
1968 (H)
Emmonsia Cif. & Montemart. 1959, ana-
morphic Ajellomyces McDonough &
A.L. Lewis 1968 (H)
Histoplasma Darling 1906, anamorphic
Ajellomyces McDonough & A.L. Lewis
1968 (H)
Lacazia Taborda, V.A. Taborda & Mc-
Ginnis 1999 (H)
Paracoccidioides F.P. Almeida 1930 (H)
Arachnomycetaceae Gibas, Sigler & Currah
Onychocola Sigler 1990, anamorphic
Arachnomyces Massee & E.S. Salmon
1902 (H)
Gymnoascaceae Baran.
Chrysosporium-like, anamorphic Orromy-
ces B. Sur & G.R. Ghosh 1987 (H)
Malbranchea Sacc. 1882, anamorphic
Byssoonygena Guarro, Punsola & Cano
1987 and Gymnoascus Baran. 1872 (H)
Oncocladium Wallr. 1833, anamorphic
Gymnoascus Baran. 1872 (H)
Onygenaceae Berk.
?Arthrographis G. Cochet ex Sigler &
J.W. Carmich. 1976, anamorphic Ara-
chnotheca Arx 1971 (H)
Chrysosporium Corda 1833 anamorphic
Amauroascus J. Schröt., Aphanoascus
Zukal 1890, Apinisia La Touche 1968,
Arachnotheca Arx 1971, Nannizziopsis
Currah 1985, ?Polytolypa J.A. Scott &
Malloch 1993, Pectinotrichum Varsav-
sky & G.F. Orr 1971, ?Renispora Sigler
& J.W. Carmich. 1979 (H)
Coccidioides G.W. Stiles 1896 (H)
Malbranchea Sacc. 1882, anamorphic Au-
xarthron G.F. Orr & Kuehn 1963,
Byssoonygena Guarro, Punsola & Cano
1987 and Uncinocarpus Sigler & G.F.
Orr 1976 (H)
Myriodontium Samson & Polon. 1978,
anamorphic Neoarachnotheca Ulfig,
Cano & Guarro 1997 (H)
Sporendonema Desm. 1827, anamorphic
Onygena Pers. 1800 (H)
Arthrodermataceae Currah
Chrysosporium Corda 1833, anamorphic
Shanorella R.K. Benj. 1956, Arthr-
oderma Curr. 1860 and Ctenomyces
Eidam 1880 (H)
Chrysosporium Corda 1833, anamorphic
Epidermophyton Sabour. 1907 (H)
Microsporum Gruby 1843, anamorphic
Nannizzia Stockdale 1961 and Arthrod-
erma Curr. 1860 (H)
Mycosphere Doi 10.5943/mycosphere/3/2/5
174
Myceliophthora Costantin 1892, anamo-
rphic Arthroderma Curr. 1860
Trichophyton Malmsten 1848, anamorphic
Arthroderma Curr. 1860 (H)
Ascosphaeraceae L.S. Olive & Spiltoir
Chrysosporium-like, anamorphic Asco-
sphaera L.S. Olive & Spiltoir 1955 and
Bettsia Skou 1972 (H)
Onygenales, genera incertae sedis
Arthropsis Sigler, M.T. Dunn & J.W.
Carmich. 1982 (H)
Lacazia Taborda, V.A. Taborda & McG-
innis 1999 (H)
Subclass Mycocaliciomycetidae Tibell
Mycocaliciales Tibell & Wedin
Mycocaliciaceae A.F.W. Schmid
Asterophoma D. Hawksw. 1981, anamor-
phic Chaenothecopsis Vain. 1927 (C)
Catenomycopsis Tibell & Constant. 1991,
anamorphic Chaenothecopsis Vain.
1927 (H)
Phialophora-like, anamorphic Chaenothe-
copsis Vain. 1927 and Mycocalicium
Vain. 1890 (H)
Class Laboulbeniomycetes Engler
Laboulbeniales Engler
Coreomycetopsis Thaxt. 1920 (H)
Muiaria Thaxt. 1914 (H)
Tetrameronycha Speg. ex W. Rossi & M.
Blackw. 1990 (H)
Pyxidiophorales P.F. Cannon
Pyxidiophoraceae Arnold
Chalara (Corda) Rabenh. 1844 anam-
orphic Pyxidiophora Bref. & Tavel
1891 (H)*
Gliocephalis Matr. 1899 (H)
Pleurocatena G. Arnaud ex Aramb.,
Gamundí, W. Gams & G.R.W. Arnold
2007, anamorphic Pyxidiophora Bref.
& Tavel 1891 (H)
Thaxteriola Speg. 1918, anamorphic Pyxi-
diophora Bref. & Tavel 1891 (H)
Class Lecanoromycetes O.E. Erikss. & Winka
Subclass Ostropomycetidae Reeb, Lutzoni &
Cl. Roux
Agyriales Clem. & Shear
Trapeliaceae M. Choisy ex Hertel
Epithyrium (Sacc.) Trotter 1931, anamo-
rphic Sarea Fr. 1825 (C)
Pycnidiella Höhn. 1915, anamorphic Sarea
Fr. 1825 (C)
Ostropales Nannf.
Odontotremataceae D. Hawksw. & Sherwood
Pycnidial anamorph associated with
Claviradulomyces P.R. Johnst., D.C.
Park, H.C. Evans, R.W. Barreto & D.J.
Soares 2010
Stictidaceae Fr.
?Coleophoma-like, anamorphic Stictopha-
cidium Rehm 1888 (C)
?Ebollia Minter & Caine 1980, anamor-
phic Stictophacidium Rehm 1888 (C)
Phacidiella-like, anamorphic Acaros-
porina Sherwood 1977 (C)
Stictospora Cif. 1957, anamorphic Stictis
Pers. 1800 (C)
Ostropales, genera incertae sedis
Glomerobolus Kohlm. & Volkm.-Kohlm.
1996 (H)
Subclass Lecanoromycetidae P.M. Kirk, P.F.
Cannon, J.C. David & Stalpers ex Miadl.,
Lutzoni & Lumbsch
Lecanorales Nannf.
Pilocarpaceae Zahlbr.
Pyriomyces Bat. & H. Maia 1965,
anamorphic Byssoloma Trevis. 1853
(C)
Szczawinskia A. Funk 1984 (C)
Stereocaulaceae Chevall.
Lepraria Ach. 1803 (C)
Lecanorales, genera incertae sedis
Leprocaulon Nyl. 1878 (C)
Vouauxiomyces Dyko & D. Hawks. 1979,
anamorphic Abrothallus De Not. 1845
(C)
Class Leotiomycetes O.E. Erikss. & Winka
Cyttariales Luttr. ex Gamundı´
Cyttariaceae Speg.
Cytariella Palm 1932, anamorphic Cytta-
Mycosphere Doi 10.5943/mycosphere/3/2/5
175
ria Berk. 1842 (C)
Erysiphales Gwynne-Vaughan
Erysiphaceae Tul. & C. Tul.
Oidiopsis Scalia 1902, anamorphic Leveil-
lula G. Arnaud 1921 (H)
Oidium Link 1824, anamorphic Arthro-
cladiella Vassilkov 1960, Blumeria
Golovin ex Speer 1975, Brasiliomyces
Viégas 1944, Cystotheca Berk. & M.A.
Curtis 1860, Erysiphe R. Hedw. ex DC.
1805, Golovinomyces (U. Braun) V.P.
Gelyuta 1988, Medusosphaera Golovin
& Gamalizk. 1962, Microsphaera Lév.
1851, U. Braun 1999, Podosphaera
Kunze 1823, Sawadaea Miyabe 1914,
Sphaerotheca Lév. 1851, Uncinula Lév.
1851 and Uncinuliella R.Y. Zheng &
G.Q. Chen 1979 (H)
Ovulariopsis Pat. & Har. 1900, ana-
morphic Phyllactinia Lév. 1851 (H)
Streptopodium R.Y. Zheng & G.Q. Chen
1978, anamorphic Pleochaeta Sacc. &
Speg. 1881 and Phyllactinia Lév. 1851
(H)
Helotiales Nannf.
Bulgariaceae Fr.
Crinula Fr. 1821, anamorphic Holwaya
Sacc. 1889 (H)
Endomelaconium Petr. 1940, anamorphic
Bulgaria Fr. 1822 (C)
Dermateaceae Fr.
Anguillospora Ingold 1942, anamorphic
Mollisia (Fr.) P. Karst. 1871 (H)
Cadophora Lagerb. & Melin 1927, anam-
orphic Mollisia (Fr.) P. Karst. 1871,
Pyrenopeziza Fuckel 1870 and Pseud-
opezicula Korf 1975 (H)
Cadophora-like, anamorphic Phaeomol-
lisia T.N. Sieber & Grünig 2009 (H)
Casaresia Gonz. Frag. 1920, anamorphic
Mollisia (Fr.) P. Karst. 1871 (H)
Chondropodium-like, anamorphic Walto-
nia Saho 1970 (C)
Corniculariella P. Karst. 1884, anamor-
phic Dermea Fr. 1825 and Durandiella
Seaver 1932 (C)
Cryptocline Petr. 1924, anamorphic
Trochila Fr. 1849 (C)
Cryptosporiopsis Bubák & Kabát 1912,
anamorphic Neofabraea H.S. Jacks.
1913, Pezicula Tul. & C. Tul. 1865 and
?Ocellaria (Tul. & C. Tul.) P. Karst.
1871 (C)
Cryptosympodula Verkley 1999, anamor-
phic Scleropezicula Verkley 1999 (C)
Cylindrocolla Bonord. 1851, anamorphic
Calloria Fr. 1836 (H)
Cylindrosporium Grev. 1822, anamorphic
Pyrenopeziza Fuckel 1870 (C)
Cystodendron Bubák 1914, anamorphic
Mollisia (Fr.) P. Karst. 1871 Belonopsis
(Sacc.) Rehm 1891 and Niptera Fr.
1849 (H)
Diplosporonema Höhn. 1917, anamorphic
Pyrenopeziza Fuckel 1917 (C)
Discogloeum Petr. 1923, anamorphic
Spilopodia Boud. 1885 (C)
Entomosporium Lév. 1856, anamorphic
Diplocarpon F.A. Wolf 1912 (C)
Eriosporella Höhn. 1916, possibly ana-
morphic Laetinaevia Nannf. 1932 (C)
Foveostroma DiCosmo 1978, anamorphic
Dermea Fr. 1825 (C)
Gelatinosporium Peck 1873, anamorphic
Dermea Fr. 1825 and Durandiella
Seaver 1932 (C)
Glarea Bills & Paláez 1999 (H)
Gloeosporidiella Petr. 1921, anamorphic
Drepanopeziza (Kleb.) Höhn. 1917 and
Pseudopeziza Fuckel 1870 (C)
Hainesia Ellis & Sacc. 1884, anamorphic
Discohainesia Nannf. 1932 (C)
Helicodendron Peyronel 1918, anamorphic
Mollisia (Fr.) P. Karst. 1871 (H)
Laetinaevia Nannf. 1932, anamorphic
Naeviopsis B. Hein 1976 (C)
Marssonina Magnus 1906, anamorphic
Diplocarpon F.A. Wolf 1912 &
Drepanopeziza (Kleb.) Höhn. 1917 (C)
Melanodiscus Höhn. 1918, anamorphic
Spilopodia Boud. 1885 (H)
Microgloeum Petr. 1922, anamorphic
Blumeriella Arx 1961 (C)
Monostichella Höhn. 1916, anamorphic
Drepanopeziza (Kleb.) Höhn. 1917 (C)
Myrioconium Syd. & P. Syd. 1912, ana-
morphic Dibeloniella Nannf. 1932 (H)
Phacidiella P. Karst. 1884, anamorphic
Pyrenopeziza Fuckel 1870 (C)
Phialocephala W.B. Kendr. 1961
anamorphic Phaeomollisia T.N. Sieber
& Grünig 2009 (H)
Mycosphere Doi 10.5943/mycosphere/3/2/5
176
Phialocephala-like, anamorphic Phaeo-
mollisia T.N. Sieber & Grünig 2009
(H)
Phialophora-like, anamorphic Dibeloniel-
la Nannf. 1932 (H)
Phloeosporella Höhn. 1924, anamorphic
Blumeriella Arx 1961 (C)
Phlyctema Desm. 1847, anamorphic Neo-
fabraea H.S. Jacks. 1913 (C)
Pilidium Kunze 1823, anamorphic Disco-
hainesia Nannf. 1932 (C)
Rhynchosporium Heinsen ex A.B. Frank
1897 (H)*
?Sporonema Desm. 1847, anamorphic
Leptotrochila P. Karst. 1871 (C)
Trichosporiella Kamyschko 1960, ana-
morphic Laetinaevia Nannf. 1932 (H)
Trimmatostroma Corda 1837 (H)
Xenochalara M.J. Wingf. & Crous 2000
(H)
Helotiaceae Rehm
Alatospora Ingold 1942 (H)
Anguillospora Ingold 1942, anamorphic
Hymenoscyphus Gray 1821 (H)
Articulospora Ingold 1942, anamorphic
Hymenoscyphus Gray 1821(H)
Ascoconidium Seaver 1942, anamorphic
Sageria A. Funk 1975 (H)
Aureohyphozyma Hosoya & Y. Otani 1995
anamorph of Gelatinipulvinella Hosoya
& Y. Otani (Hosoay Y Otani 1995) (H)
Bloxamia Berk. & Broome 1854, anamor-
phic Bisporella Sacc. 1884 (H)
Bothrodiscus Shear 1907, anamorphic
Ascocalyx Naumov 1926 (C)
Brunchorstia Erikss. 1891, anamorphic
Gremmeniella M. Morelet 1969 (C)
?Bryophytomyces Cif. 1953, anamorphic
Hymenoscyphus Gray 1821 (H)
Cercosporella Sacc. 1880, anamorphic
Heterosphaeria Grev. 1824 (H)
Coryne Nees 1816, anamorphic Asco-
coryne J.W. Groves & D.E. Wilson
1967 (H)
Cystodendron Bubák 1914, anamorphic
Bisporella Sacc. 1884 (H)
?Cystotricha Berk. & Broome 1850, ana-
morphic Xylogramma Wallr. 1833 (C)
Deltosperma W.Y. Zhuang 1988, ana-
morphic Unguiculariopsis Rehm 1909
(C)
Dendrostilbella Höhn. 1905, anamorphic
Claussenomyces Kirschst. 1923 (H)
Dichaenopsella Petr. 1952, type is
anamorphic Godroniopsis Diehl & E.K.
Cash 1929 (C)
Digitosporium Gremmen 1953, anam-
orphic Crumenulopsis J.W. Groves
1969 (C)
Dimorphospora Tubaki 1958, anamorphic
Hymenoscyphus Gray 1821 (H)
Dothiorina Höhn. 1911, anamorphic Chlo-
rociboria Seaver ex Ramamurthi, Korf
& L.R. Batra 1958 (C)
Endomelaconium Petr. 1940, anamorphic
Austrocenangium Gamundí 1997 (C)
Eriomycopsis Speg. 1910, anamorphic
Calloriopsis Syd. & P. Syd. 1917 (H)
Eustilbum Rabenh. 1864, anamorphic Bis-
porella Sacc. 1884 (H)
Fuckelia Bonord. 1864, anamorphic God-
ronia Moug. & Lév. 1846 (C)
Geniculospora Sv. Nilsson ex Marvanová
& Sv. Nilsson 1971, anamorphic Hyme-
noscyphus Gray 1821 (H)
Gliomastix Guég. 1905 anamorphic
Ascoclavulina Y. Otani 1974 (H)*
Heteropatella Fuckel 1874, anamorphic
Heterosphaeria Grev. 1824 (C)
Hormonema-like, anamorphic Pragmopo-
ra A. Massal 1855 (H)
Idriella P.E. Nelson & S. Wilh. 1956,
anamorphic Hymenoscyphus Gray 1821
(H)
Linodochium Höhn. 1909, anamorphic
Pseudohelotium Fuckel 1870 (H)
Monodictys S. Hughes 1958 anamorphic
Hymenoscyphus Gray 1821 (H)
Periperidium Darker 1963, anamorphic
Micraspis Darker 1963 (C)
Phialophora-like, anamorphic Pseudopezi-
cula Korf 1975 (H)
Pragmopycnis B. Sutton & A. Funk 1975,
anamorphic Pragmopora A. Massal
1855 (C)
?Pseudaegerita J.L. Crane & Schokn.
1981, anamorphic Claussenomyces Kir-
schst. 1923 (H)
Pseudospiropes M.B. Ellis 1971, ana-
morphic Strossmayeria Schulzer 1881
(H)
Mycosphere Doi 10.5943/mycosphere/3/2/5
177
Rhinocladiella-like, anamorphic Bioscy-
pha Syd. 1927 (H)
Rhizothyrium Naumov 1915, anamorphic
Rhizocalyx Petr. 1928 (C)
Scytalidium Pesante 1957 anamorphic
Hymenoscyphus Gray 1821 (H)
Sirodothis Clem. 1909, anamorphic Tym-
panis Tode 1790 (C)
Sporonema Desm. 1847, anamorphic God-
ronia Moug. & Lév. 1846 (C)
Symphyosirella Seifert 2009, anamorphic
Symphyosirinia E.A. Ellis 1956 (H)
Titaeospora Bubák 1916, anamorphic
Stamnaria Fuckel 1870 (C)
Topospora Fr. 1836, anamorphic Godr-
onia Moug. & Lév. 1846 (C)
Tricladium Ingold 1942, anamorphic
Cudoniella Sacc. 1889 and Hymeno-
scyphus Gray 1821 (H)
Varicosporium W. Kegel 1906, anamor-
phic Hymenoscyphus Gray 1821 (H)
Hemiphacidiaceae Korf
Meria Vuill. 1896, anamorphic Rhabdo-
cline Syd. 1922 (H)
Rhabdogloeum Syd. 1922, anamorphic
Rhabdocline Syd. 1922 (C)
Rhabdogloeopsis Petr. 1925, anamorphic
Sarcotrochila Höhn. 1917 (C)
Hyaloscyphaceae Nannf.
Acleistia Bayl. Ell. (1917) anamorphic
Calycina Nees ex Gray 1821 (C)
Brefeldochium Verkley 2005, anamorphic
Polydesmia Boud. 1885 (H)
Chaetochalara B. Sutton & Piroz. 1965
(H)
Chalara (Corda) Rabenh. 1844, anam-
orphic Calycellina Höhn., Phaeos-
cypha Spooner 1984 and Tapesina
Lambotte 1887 (H)*
Cheiromycella Höhn. 1910, anamorphic
Hyaloscypha Boud. 1885 (H)
Clathrosphaerina Beverw. 1951, anamor-
phic Hyaloscypha Boud. 1885 (H)
Haplographium Berk. & Broome 1859,
anamorphic Dematioscypha Svrček
1977 (H)
Mycoarthris Marvanová & P.J. Fisher
2002 (H)
Naemospora Roth ex Kuntze 1898,
possibly anamorphic Lachnellula P.
Karst. 1884 (C)
Phialophora-like, anamorphic Cistella
Quél. and Hyalopeziza Fuckel 1870 (H)
Pseudaegerita J.L. Crane & Schokn. 1981,
anamorphic Hyaloscypha Boud. 1885
(H)
Septonema-like, anamorphic Ciliolarina
Svrcek 1977 (H)
Tricladium Ingold 1942, anamorphic
Hydrocina Scheuer 1991 (H)
Leotiaceae Corda
Anguillospora Ingold 1942, anamorphic
Pezoloma Clem. 1909 (H)
Aureohyphozyma Hosoya & Y. Otani
1995, anamorphic Gelatinipulvinella
Hosoya & Y. Otani 1995 (H)
Halenospora E.B.G. Jones 2009 (H)
Myrioconium-like, anamorphic Neobulga-
ria Petr. 1921 (H)
Phialophora-like, anamorphic Neobulga-
ria Petr. 1921 (H)
Loramycetaceae Dennis ex Digby & Goos
Anguillospora Ingold 1942, anamorphic
Loramyces W. Weston 1929 (H)
Phacidiaceae Fr.
Allantophomopsis Petr. 1925, anamorphic
Phacidium Fr. 1815 (C)
Apostrasseria Nag Raj 1983, anamorphic
Lophophacidium Lagerb. 1949 and
Phacidium Fr. 1815 (C)
Blennoria Moug. & Fr. 1825, anamorphic
Phacidium Fr. 1815 (C)
Ceuthospora Grev. 1826, anamorphic
Phacidium Fr. 1815 (C)
?Coma Nag Raj & W.B. Kendr. 1972,
anamorphic Ascocoma H.J. Swart (C)
Rutstroemiaceae Holst-Jensen, L.M. Kohn &
T. Schumacher
Helicodendron Peyronel 1918, anamorphic
Lambertella Höhn. 1918 (H)
Myrioconium Syd. & P. Syd. 1912, ?ana-
morphic Lanzia Sacc. 1884, Rutstro-
emia P. Karst. 1871, Scleromitrula
Mycosphere Doi 10.5943/mycosphere/3/2/5
178
S.Imai 1941, Lambertella Höhn. 1918
and Poculum Velen. 1934 (H)
Phialophora-like, anamorphic Rutstroemia
P. Karst. 1871 (H)
Sclerotiniaceae Whetzel ex Whetzel
Acarosporium Bubák & Vleugel ex Bubák
1911, anamorphic Pycnopeziza W.L.
White & Whetzel 1938 (C)
Amerosporium Speg. 1882, anamorphic
Zoellneria Velen. 1934 (C)
Amphobotrys Hennebert 1973, anamorphic
Botryotinia Whetzel 1945 (H)
Botrytis P. Micheli ex Pers. 1794,
anamorphic Botryotinia Whetzel 1945
(H)
Cristulariella Höhn. 1916, anamorphic
Nervostroma Narumi & Y. Harada
2006 (H)
Haradamyces Masuya, Kusunoki, Kosaka
& Aikawa 2009 (H)
Hinomyces Narumi & Y. Harada 2006,
anamorphic Grovesinia M.N. Cline,
J.L. Crane & S.D. Cline 1983 (H)
Monilia Bonord. 1851, anamorphic Moni-
linia Honey 1928 and Phaeosclerotinia
Hori 1916 (H)
Mycopappus Redhead & G.P. White 1985,
anamorphic Redheadia Y. Suto &
Suyama 2005 (H)
Myrioconium Syd. & P. Syd. 1912,
anamorphic Ciboria Fuckel 1870,
Ciborinia Whetzel 1945, (Fr.) P. Karst.
1871, Martininia Dumont & Korf 1970,
Myriosclerotinia N.F. Buchw. 1947,
Sclerotinia Fuckel 1870 Stromatinia
(Boud.) Boud. 1907 and Monilinia
Honey 1928 (H)
Ovulitis N.F. Buchw. 1970, anamorphic
Ovulinia F.A. Weiss 1940 (H)
Sclerotium Tode 1790, anamorphic Myrio-
sclerotinia N.F. Buchw. 1947, Sclero-
tinia Fuckel 1870 and Stromatinia
(Boud.) Boud. 1907 (H)
Septotis N.F. Buchw. ex Arx 1970,
anamorphic Septotinia Whetzel ex J.W.
Groves & M.E. Elliott 1961 (H)
Streptobotrys Hennebert 1973, anamorphic
Streptotinia Whetzel 1945 (H)
Valdensia Peyronel 1923, anamorphic
Valdensinia Peyronel 1953 (H)
Verrucobotrys Hennebert 1973, anamor-
phic Seaverinia Whetzel 1945 (H)
Vibrisseaceae Korf
Acephala Grünig & T.N. Sieber 2005 (H)
Anavirga B. Sutton 1975, anamorphic
Vibrissea Fr. 1822 (H)
Phialocephala W.B. Kendr. 1961, some
possibly anamorphic Vibrissea Fr. 1822
(H)
Phialophora-like, anamorphic Chlorovi-
brissea L.M. Kohn 1989 (H)
Helotiales, genera incertae sedis
Catenulifera Hosoya 2002, anamorphic
Hyphodiscus Kirschst. 1906 (H)
Chalara (Corda) Rabenh. 1844 (H)*
Clathrosporium Nawawi & Kuthub. 1987
(H)
Crucellisporium M.L. Farr 1968 (C)
Cystotricha Berk. & Broome 1850 (C)
Dactylaria Sacc. 1880 (H)
Diplococcium Grove 1885 (H)
Endoconidium Prill. & Delacr. 1891, ana-
morphic Gloeotinia M. Wilson, Noble
& E.G. Gray 1954 (H)
Fontanospora Dyko 1978 (H)
Fulvoflamma Crous 2006 (H)
Glarea Bills & Paláez 1999 (H)
Helgardia Crous & W. Gams 2003, ana-
morphic Oculimacula Crous & W.
Gams 2003 (H)
Humicolopsis Cabral & S. Marchand 1976
(H)
Hyalodendriella Crous 2007 (H)
Hysteropezizella Höhn. 1917 (C)
Lemonniera De Wild. 1894 (H)
Leptodontidium de Hoog 1979 (H)
Libartania Nag Raj 1979 (C)
Margaritispora Ingold 1942 (H)
Meliniomyces Hambl. & Sigler 2005 (H)
Mirandina G. Arnaud ex Matsush. 1975
(H)
Monochaetiellopsis B. Sutton & DiCosmo
1977, anamorphic Hypnotheca Tom-
merup 1970 (C)
Mycochaetophora Hara & Ogawa 1931
(H)
Neozythia Petr. 1958 (C)
Rhexocercosporidium U. Braun 1994 (H)
Rhizocladosporium Crous & U. Braun
Mycosphere Doi 10.5943/mycosphere/3/2/5
179
2007 (H)
Rhodesia Grove 1937 (C)
Scytalidium Pesante 1957 (H)
Sphaerographium Sacc. 1884 (C)
Spirosphaera Beverw. 1953 (H)
Tetracladium De Wild. 1893 (H)
Thedgonia B. Sutton 1973 (H)
Trimmatostroma Corda 1837 (H)
Variocladium Descals & Marvanová 1999
(H)
Xenopolyscytalum Crous 2011(‘2010’) (H)
Rhytismatales M.E. Barr ex Minter
Ascodichaenaceae D. Hawksw. & Sherwood
Dichaenopsella Petr. 1952, anamorphic
Ascodichaena Butin 1977 (C)
Macroallantina Speer 1987, anamorphic
Delpinoina Kuntze, 1891 (C)
Polymorphum Chevall. 1822, anamorphic
Ascodichaena Butin 1977 (C)
Cryptomycetaceae Höhn.
?Phacidiopycnis Potebnia 1912, anamor-
phic Potebniamyces Smerlis 1962 (C)
Rhytismataceae Chevall.
Conostroma Moesz 1921, anamorphic
Colpoma Wallr. 1833 (C)
Crandallia Ellis & Sacc. 1897, ana-
morphic Bifusella Höhn. 1917 and
Duplicaria Fuckel 1870 (C)
Cryocaligula Minter 1986, anamorphic
Ploioderma Darker. 1967 (C)
Hysterodiscula Petr. 1942, anamorphic
Duplicaria Fuckel 1870 (C)
Leptostroma Fr. 1815, anamorphic Lopho-
dermium Chevall. 1826 (C)
Melasmia Lév. 1846, anamorphic Rhytis-
ma Fr. 1818 (C)
Tryblidiopycnis Höhn. 1918, anamorphic
Tryblidiopsis P. Karst. 1871 (C)
Rhytismatales, genera incertae sedis
Fuligomyces Morgan-Jones & Kamal 1984
(H)
Uyucamyces H.C. Evans & Minter 1985,
anamorphic Ocotomyces H.C. Evans &
Minter 1985 (C)
Leotiomycetes, families incertae sedis
Myxotrichaceae Currah
Geomyces Traaen 1914, anamorphic
Pseudogymnoascus Raillo 1929 (H)
?Gymnostellatospora Udagawa, Uchiy. &
Kamiya 1993, anamorphic Pseudo-
gymnoascus Raillo 1929 (C)
?Malbranchea Sacc. 1882, anamorphic
Myxotrichum Kunze 1823 (H)
Oidiodendron Robak 1932, anamorphic
Byssoascus Arx 1971 and Myxotrichum
Kunze 1823 (H)
Leotiomycetes, genera incertae sedis
Brachyalara Réblová & W. Gams 2011
(H)
Chaetomella Fuckel 1870 (C)
Collophora Damm & Crous 2010 (C)
Eleutheromyces Fuckel 1870 (C)
Geniculospora Sv. Nilsson ex Marvanová
& Sv. Nilsson 1971 (H)
Hyphozyma de Hoog & M.T. Sm. 1981
(H)
Infundichalara Réblová & W. Gams 2011
(H)
Leohumicola N.L. Nick. 2005 (H)
Meliniomyces Hambl. & Sigler 2005 (H)
Scytalidium Pesante 1957, anamorphic
Xylogone Arx & T. Nilsson 1969 (H)
Tetrachaetum Ingold 1942 (H)
Tiarosporella Höhn. 1924, anamorphic
Darkera H.S. Whitney, J. Reid & Piroz.
1975 (C)
Class Orbiliomycetes O.E. Erikss. & Baral
Orbiliales Baral, O.E. Erikss., G. Marson & E.
Weber
Orbiliaceae Nannf.
Anguillospora Ingold 1942, anamorphic
anamorphic Orbilia Fr. 1836 (H)
Arthrobotrys Corda 1939, anamorphic
Orbilia Fr. 1836 (H)
Brachyphoris Juan Chen, L.L. Xu, B. Liu
& Xing Z. Liu 2007, anamorphic
Hyalorbilia Baral & G. Marson 2000
(H)
Dactylaria Sacc. 1880, anamorphic Or-
bilia Fr. 1836 (H)
Dactylella Grove 1884, anamorphic Orbi-
lia Fr. 1836 (H)
Dactylellina M. Morelet 1968 (H)
Dicranidion Harkn. 1885, anamorphic Or-
bilia Fr. 1836 (H)
Mycosphere Doi 10.5943/mycosphere/3/2/5
180
Drechslerella Subram. 1964, anamorphic
Orbilia Fr. 1836 (H)
Dwayaangam Subram. 1978, anamorphic
Orbilia Fr. 1836 (H)
Gamsylella M. Scholler, Hagedorn & A.
Rubner 1999, anamorphic Orbilia Fr.
1836 (H)
Helicoön Morgan 1892, anamorphic
Orbilia Fr. 1836 (H)
Idriella P.E. Nelson & S. Wilh. 1956,
anamorphic Orbilia Fr. 1836 (H)
Monacrosporium Oudem. 1885, anamor-
phic Orbilia Fr. 1836 (H)
Pseudotripoconidium Z.F. Yu & K.Q.
Zhang 2011, anamorphic Orbilia Fr.
1836 (H)*
Tricellula Beverw. 1954, anamorphic
Orbilia Fr. 1836 (H)
Tridentaria Preuss 1852 (H)
Trinacrium Riess 1852, anamorphic Orbi-
lia Fr. 1836 (H)
Class Pezizomycetes sensu O.E. Erikss. &
Winka
Pezizales J. Schröt.
Ascobolaceae Boud. ex Sacc.
Papulospora Preuss 1851, anamorphic
Ascobolus Pers. 1796 (H)
Papulospora-like, anamorphic Cubonia
Sacc. 1889 (H)
Rhizostilbella Wolk 1914, anamorphic
Ascobolus Pers. 1792 (H)
Caloscyphaceae Harmaja
Geniculodendron G.A. Salt 1974, anamor-
phic Caloscypha Boud. 1885 (H)
Chorioactidaceae Pfister
Kumanasamuha P.Rag. Rao & D. Rao
1964, anamorphic Chorioactis Kupfer
ex Eckblad 1968 (H)
Verticicladium Preuss 1851, anamorphic
Desmazierella Lib. 1829 (H)
Morchellaceae Reichenb.
Costantinella Matr. 1892, anamorphic
Morchella Dill. ex Pers. 1794 (H)
Pezizaceae Dumort.
Chromelosporium Corda 1833, anamor-
phic Muciturbo P.H.B. Talbot 1989,
Peziza Dill. ex Fr. 1822 and Plicaria
Fuckel 1870 (H)
Glischroderma Fuckel 1870 (C)
Oedocephalum Preuss 1851, anamorphic
Iodophanus Korf 1967 and Peziza Dill.
ex Fr. 1822 (H)
Ostracoderma Fr. 1825, anamorphic Pezi-
za Dill. ex Fr. 1822 (C)
Pyronemataceae Corda
Actinosporella Descals, Marvanová & J.
Webster 1999, anamorphic Miladina
Svrček 1972 (H)
Ascorhizoctonia Chin S. Yang & Korf
1985, anamorphic Tricharina Eckblad
1968 (H)
Basifimbria Subram. & Lodha 1968,
anamorphic Geopyxis (Pers.) Sacc. (H)
Complexipes C. Walker 1979, anamorphic
Tricharina Eckblad 1968 and Wilcoxina
Chin S. Yang & Korf 1985 (H)
?Dichobotrys Hennebert 1973, anamorphic
Pyropyxis Egger 1984 and Trichophaea
Boud. 1885 (H)
Micronematobotrys Xiang Sun & L.D.
Guo 2010 (H)
?Molliardiomyces Paden 1984, anam-
orphic Stephensia Tul. & C. Tul. 1851
(H)
?Scytalidium Pesante 1957, anamorph of
Anthracobia Boud. 1885 (H)
Rhizinaceae Bonord.
Phymatotrichopsis Hennebert 1973 (H)
Sarcoscyphaceae LeGal ex Eckblad
Molliardiomyces Paden 1984, anamorphic
Nanoscypha Denison 1972, Phillipsia
Berk. 1881, Pithya Fuckel 1870 and
Sarcoscypha (Fr.) Boud. 1885 (H)
Verticicladium Preuss 1851, anamorphic
Desmazierella Lib. 1829 (H)
Sarcosomataceae Kobayasi
Conoplea Pers. 1797, anamorphic Urnula
Fr. 1849 and Plectania Fuckel 1870 (H)
Strumella Fr. 1849, anamorphic Urnula Fr.
1849 (H)
Verticicladium Preuss 1851, anamorphic
Mycosphere Doi 10.5943/mycosphere/3/2/5
181
Sarcosoma Casp. 1891 (H)
Pezizales, genera incertae sedis
Cephaliophora Thaxt. 1903 (H)
Class Sordariomycetes sensu O.E. Erikss. &
Winka
Subclass Hypocreomycetidae O.E. Erikss. &
Winka
Coronophorales Nannf.
Chaetosphaerellaceae Huhndorf, A.N. Mill. &
F.A. Fernández
Oedemium Link 1824, anamorphic Chae-
tosphaerella E. Müll. & C. Booth 1972
(H)
Veramycina Subram. 1995, anamorphic
Chaetosphaerella E. Müll. & C. Booth
1972 (H)
Nitschkiaceae (Fitzp.) Nannf.
Acremonium-like, anamorphic Acantho-
nitschkea Speg. (H)
Glomerellales Chadef. ex Réblová, W. Gams
& Seifert
Australiascaceae Réblová & W. Gams
Monilochaetes Halst. ex Harter 1916,
anamorphic Australiasca Sivan. &
Alcorn 2002 (H)*
Glomerellaceae Locq. ex Seifert & W. Gams
Colletotrichum Corda 1831, anamorphic
Glomerella Spauld. & Schrenk 1903
(C)
Reticulascaceae Réblová & W. Gams
Cylindrotrichum Bonord.1851, anamorph-
ic Reticulascus Réblová & W. Gams
2011 (H)*
Hypocreales Lindau
Bionectriaceae Samuels & Rossman
Acremonium-like, anamorphic Dimero-
sporiella Speg. 1908, Ijuhya Starbäck
1899, Hydropisphaera Dumont 1822,
Lasionectria (Sacc.) Cooke 1884, My-
coarachis Malloch & Cain 1970,
Mycocitrus Moller 1901, Nectriella
Nitschke ex Fuckel 1870, Nectriopsis
Maire 1911, Nigrosabulum Malloch &
Cain 1970, Protocreopsis Yoshim. Doi
1977, Selinia P. Karst. 1876, and
Stilbocrea Pat. 1900 and Verrucostoma
Hirooka, Tak. Kobay. & Chaverri 2010
(H)
Albosynnema E.F. Morris 1967 (H)
Clonostachys Corda 1839, anamorphic
Bionectria Speg. 1919 (H)
Dendrodochium-like, anamorphic Nec-
triella Nitschke ex Fuckel 1870 (C)
Didymostilbe Henn. 1902, anamorphic
Peethambara Subram. & Bhat 1978 (H)
Geosmithia J.Pitt 1979 (H)*
Gliocladium-like, anamorphic Roumegue-
riella Speg. 1880 (H)
Gliomastix Guég. 1905 (H)*
Gracilistilbella Seifert 2000, anamorphic
Stilbocrea Pat. 1900 (H)
Kutilakesa Subram. 1956, anamorphic
Nectriella Nitschke ex Fuckel 1870 (H)
Rhopalocladium Schroers, Samuels & W.
Gams 1999, anamorphic Nectriopsis
Maire 1911 (H)
Sarocladium W. Gams & D. Hawksw.
1976 (H)
Spicellum Nicot & Roquebert 1976 (H)
Stanjemonium W. Gams, O'Donnell,
Schroers & M. Chr. 1999 (H)
Stilbella Lindau 1900 anamorphic
Stilbocrea Pat. 1900 (H)
Trichothecium-like, anamorphic Heleoc-
occum C.A. Jørg. 1922 (H)
Tubercularia-like, anamorphic Selinia P.
Karst. 1876 (H)
Verticillium-like, anamorphic Nectriopsis
Maire 1911 (H)
Vesicladiella Crous & M.J. Wingf. 1994
(H)
Virgatospora Finley 1967, anamorphic
Peethambara Subram. & Bhat 1978 (H)
Clavicipitaceae (Lindau) Earle ex Rogerson
Acremonium-like, anamorphic Neobarya
Lowen 1986 (H)
Albomyces I. Miyake 1908, anamorphic
Aciculosporium I. Miyake (C)
Amphichordia Fr., 1825 (H)
Aschersonia Mont. 1848, anamorphic
Hypocrella Sacc. and Moelleriella Bres.
1897 (C)
Blistum B. Sutton 1973, anamorphic
Berkelella (Sacc.) Sacc. 1891 (H)
Calcarisporium Preuss 1851, anamorphic
Neobarya Lowen (H)
Corallocytostroma Y.N. Yu & Z.Y. Zhang
Mycosphere Doi 10.5943/mycosphere/3/2/5
182
1980 (C)
Diploöspora Grove 1916, anamorphic
Neobarya Lowen 1986 (H)
Drechmeria W. Gams & H.-B. Jansson
1985 (H)
Ephelis Fr. 1849, anamorphic Atkinsonella
Diehl, Balansia Speg., 1885, Epichloë
(Fr.) Tul. & C. Tul., 1865, Dothichloë
G.F. Atk. 1894, Heteroepichloë E.
Tanaka, C. Tanaka, Gafur & Tsuda
2002, Nigrocornus Ryley & Langdon
2003 and Myriogenospora G.F. Atk.
1894 (H)
Harposporium Lohde 1874, anamorphic
Podocrella Seaver 1928 (H)
Hirsutella-like, anamorphic Orbiocrella D.
Johnson, G.H. Sung, Hywel-Jones &
Spatafora 2009 and Moelleriella Bres.
1897 (H)
Lecanicillium W. Gams & Zare 2001,
anamorphic Hypocrella Sacc. 1878and
Neobarya Lowen 1986 (H)
Metarhizium Sorokīn 1879, anamorphic
Metacordyceps G.H. Sung, J.M. Sung,
Hywel-Jones & Spatafora 2007 (H)
?Munkia Speg. 1886, anamorphic Myc-
omalus A. Møller 1901 (H)
Neomunkia Petr. 1947 (H)
Neotyphodium Glenn, C.W. Bacon &
Hanlin 1996, anamorphic Epichloë (Fr.)
Tul. & C. Tul. 1865 (H)
Nomuraea Maubl. 1903 (H)
Pleurodesmospora Samson, W. Gams &
H.C. Evans 1979 (H)
Pochonia Bat. & O.M. Fonseca 1965 (H)
Polycephalomyces Kobayasi 1941, ana-
morphic Berkelella (Sacc.) Sacc. 1891
(H)
Pseudogibellula Samson & H.C. Evans
1973, anamorphic Torrubiella Boud.
1885 (H)
Pseudomeria G.L. Barron 1980 (H)
Rotiferophthora G.L. Barron 1991 (H)
Sphacelia Lév. 1827, anamorphic Atkin-
sonella Diehl 1950 and Claviceps Tul.
1853 (H)
Ustilaginoidea Bref. 1895, anamorphic
Villosiclava E. Tanaka & C. Tanaka
2009 (H)
Cordycipitaceae Kreisel ex G.M. Sung, J.M.
Sung, Hywel-Jones & Spatafora
Akanthomyces Lebert 1858, anamorphic
Cordyceps Fr. 1824 and Torrubiella
Boud. 1885 (H)
Beauveria Vuill. 1912, anamorphic Cordy-
ceps Fr. 1824 (H)*
Engyodontium de Hoog 1978 (H)
Gibellula Cavara 1894, anamorphic
Cordyceps Fr. 1824 and Torrubiella
Boud. 1885 (H)
Granulomanus de Hoog & Samson 1978,
anamorphic Torrubiella Boud. 1885
(H)
Hirsutella Pat. 1892, anamorphic
Cordyceps Fr. 1824 and Cordyce-
pioideus Stifler 1941 (H)
Isaria Pers. 1794, anamorphic Cordyceps
Fr. 1824 and Torrubiella Boud. 1885
(H)
Lecanicillium W. Gams & Zare 2001,
anamorphic Cordyceps Fr. 1824 and
Torrubiella Boud. 1885 (H)
Microhilum H.Y. Yip & A.C. Rath 1989
(H)
Pseudogibellula Samson & H.C. Evans
1973 (H)
Rotiferophthora G.L. Barron 1991 (H)
Septofusidium W. Gams 1971, anamorphic
Cordyceps Fr. 1824 (H)
Simplicillium W. Gams & Zare 2001,
possibly anamorphic Torrubiella Boud.
1885 (H)
Tolypocladium W. Gams 1971, anamo-
rphic Cordyceps Fr. 1824 (H)
Hypocreaceae De Not.
Acremonium-like, anamorphic Protocrea
Petch 1937 and Pseudohypocrea
Yoshim. Doi 1972 (H)
Arachnocrea Z. Moravec 1956, anamor-
phic Protocrea Petch 1937 and
Sarawakus Lloyd 1924 (C)
Cladobotryum Nees 1816, anamorphic
Hypomyces (Fr.) Tul. & C. Tul. 1860
(H)*
Gliocladium Corda 1840, anamorphic
Hypocrea Fr. 1825 and Sphaerostil-
bella (Henn.) Sacc. & D. Sacc. 1905
(H)
Gliocladium-like, anamorphic Sarawakus
Lloyd 1924 (H)
Mycogone Link 1809, anamorphic Hypo-
myces (Fr.) Tul. & C. Tul. 1860 (H)
Mycosphere Doi 10.5943/mycosphere/3/2/5
183
Papulaspora-like, anamorphic Hypomyces
(Fr.) Tul. & C. Tul. 1860 (H)
Sepedonium Link 1809, anamorphic Hypo-
myces (Fr.) Tul. & C. Tul. 1860 (H)
Sibirina G.R.W. Arnold 1970, anamorphic
Hypomyces (Fr.) Tul. & C. Tul. 1860
(H)
Stephanoma Wallr. 1833, anamorphic
Hypomyces (Fr.) Tul. & C. Tul. 1860
(H)
Stilbella Lindau 1900, anamorphic
Hypocrea Fr. 1825 (H)
Stromatocrea W.B. Cooke 1952, anamor-
phic Hypocreopsis P. Karst. 1873 (H)
Trichoderma Pers. 1794, anamorphic
Hypocrea Fr. 1825 and Podostroma P.
Karst. 1892 (H)*
Trichoderma-like, anamorphic Sarawakus
Lloyd 1924 (H)
Trichothecium-like, anamorphic Hypom-
yces (Fr.) Tul. & C. Tul. 1860 (H)
Verticillium-like, anamorphic Aphysiostro-
ma Barrasa, A.T. Martínez & G.
Moreno 1986, Sphaerostilbella Henn.
1901 and Coniochaeta (Sacc.) Cooke
1887 (H)
Verticimonosporium Matsush. 1971 (H)
Nectriaceae Tul. & C. Tul.
Acremonium-like, anamorphic Cosmospo-
ra Rabenh. 1862, Nectria (Fr.) Fr. 1849
and Neocosmospora E.F. Sm. 1899 (H)
Actinostilbe Petch 1925, anamorphic La-
natonectria Samuels & Rossman 1999
(H)
Antipodium Piroz. 1974, anamorphic
Ophionectria Sacc. 1878 (H)
Aphanocladium W. Gams 1971 (H)
Atractium Link 1809 (H)
Calostilbella Höhn. 1919, anamorphic
Calostilbe Sacc. & Syd. 1902 (H)
Campylocarpon Halleen, Schroers &
Crous 2005 (H)
Chaetodochium Höhn. 1932, anamorphic
Pseudonectria Seaver 1909 (H)
Chaetopsina Rambelli 1956, anamorphic
Chaetopsinectria J. Luo & W.Y.
Zhuang and Cosmospora Rabenh. 1862
(H)
Curvicladiella Decock & Crous 2006 (H)
Curvicladium Decock & Crous 1998 (H)
Cyanochyta Höhn. 1915, anamorphic
Gibberella Sacc. 1877 (C)
Cyanophomella Höhn. 1918, anamorphic
Gibberella Sacc. 1877 (C)
Cylindrocarpon Wollenw. 1913, anamor-
phic Neonectria Wollenw. 1917,
Rugonectria P.Chaverri & Samuels
2011, Thelonectria P. Chaverri & C.G.
Salgado 2011, Ilyonectria P.Chaverri &
Samuels 2011 (H)*
Cylindrocladiella Boesew. 1982, anamor-
phic Cosmospora Rabenh. 1862 and
Nectricladiella Crous & C.L. Schoch
2000 (H)
Cylindrocladium Morgan 1892, anamor-
phic Calonectria De Not. 1867 (H)
Dacryoma Samuels 1988, anamorphic
Nectria (Fr.) Fr. 1849 (H)
Dematiocladium Allegr., Aramb., Cazau &
Crous 2005 (H)
Dichomera Cooke 1878, anamorphic
Gibberella Sacc. 1877 (C)
Flagellospora Ingold 1942, anamorphic
Nectria (Fr.) Fr. 1849 (H)
Fusarium Link 1809, anamorphic Albo-
nectria Rossman & Samuels 1999, Cya-
nonectria Samuels & Chaverri 2009,
Gibberella Sacc. 1877, Corallomyce-
tella Henn. 1904 and Haematonectria
Samuels & Nirenberg 1999 and
unknown teleomorphs (H)*
Fusarium-like, anamorphic, Geejayessia
Schroers, Gräfenhan & Seifert 2011 and
Cyanonectria Samuels & Chaverri 2009
(H)*
Fusicolla Bonord. 1851 (H)
Gliocephalotrichum J.J. Ellis & Hesselt.
1962, anamorphic Leuconectria Ross-
man, Samuels & Lowen 1993 (H)
Gliocladiopsis S.B. Saksena 1954, ana-
morphic Glionectria Crous & S.L.
Schoch 2000 (H)
Gyrostroma Naumov 1914, anamorphic
Nectria (Fr.) Fr. 1849 (C)
Heliscus Sacc. 1880, anamorphic Neone-
ctria Wollenw. 1917 . 1849 (H)
Macroconia (Wollenw.) Gräfenhan,
Seifert & Schroers 2011 (H)
Mariannaea G. Arnaud ex Samson 1974
anamorphic ‘Nectria’ (Fr.) Fr. 1849 and
Cordyceps Fr. 1833 (H)*
Mycosphere Doi 10.5943/mycosphere/3/2/5
184
Microcera Desm. 1848, anamorphic
Gibberella Sacc. 1877 (H)*
Mirandina G. Arnaud ex Matsush. 1975,
anamorphic Chaetonectrioides Mats-
ush. (H)
Nalanthamala Subram. 1956, anamorphic
Rubrinectria Rossman & Samuels 1999
(H)
Penicillifer Emden 1968, anamorphic Viri-
dispora Samuels & Rossman 1999 (H)
Pleurocolla Petr. 1924 (H)
Rhizostilbela Wolk 1914, anamorphic
Corallomycetella Henn. 1904 (H)
Septofusidium W. Gams 1971 (H)
Stachybotryna Tubaki & T. Yokoy. 1971,
anamorphic ‘Nectria’ (Fr.) Fr. 1849 (H)
Stilbella Lindau 1900, anamorphic Cosmo-
spora Rabenh. 1862 and Nectria (Fr.)
Fr. 1849 (H)*
Tilachlidium Preuss 1851, anamorphic
‘Pseudonectria’ Seaver 1909 (H)
Tubercularia Tode 1790, anamorphic
Nectria (Fr.) Fr. 1849 (H)*
Verticillium-like, anamorphic Cosmospora
Rabenh. 1862 (H)
Volutella Fr. 1832, anamorphic Cosmo-
spora Rabenh. 1862 and Pseudonectria
Seaver 1909 (H)
Xenocylindrocladium Decock, Hennebert
& Crous 1997, anamorphic Xenocalo-
nectria Crous & C.L. Schoch 2000 (H)
Zythiostroma Höhn. ex Falck 1923,
anamorphic Nectria (Fr.) Fr. 1849 (C)
Niessliaceae Kirschst.
Acremonium-like, anamorphic Tricho-
sphaerella E. Bommer, M. Rousseau &
Sacc. 1891 and Valetoniellopsis Samu-
els & M.E. Barr 1998 (H)
Custingophora-like, anamorphic Melanop-
samma Niessl 1876 (H)
Monocillium S.B. Saksena 1955, ana-
morphic Hyaloseta A.W. Ramaley 2001
and Niesslia Auersw. 1869 (H)
Ophiocordycipitaceae G.H. Sung, J.M. Sung,
Hywel-Jones & Spatafora
Chaunopycnis W. Gams 1979 (H)
Haptocillium W. Gams & Zare 2001
anamorphic Ophiocordyceps Petch
1931 (H)
Hirsutella Pat. 1892, anamorphic Ophio-
cordyceps Petch 1931 (H)
Hymenostilbe Petch 1931, anamorphic
Ophiocordyceps Petch 1931 (H)
Paraisaria Samson & B.L. Brady 1983,
anamorphic Ophiocordyceps Petch
1931 (H)
Purpureocillium Luangsa-ard, Hywel-
Jones, Houbraken & Samson 2011 (H)
Syngliocladium Petch 1932, anamorphic
Ophiocordyceps Petch 1931(H)
Tolypocladium W. Gams 1971, anamor-
phic Elaphocordyceps G.H. Sung &
Spatafora 2007 (H)
Verticillium-like, anamorphic Elaphocor-
dyceps G.H. Sung & Spatafora 2007
(H)
Hypocreales, genera incertae sedis
Acremonium Link 1809 (H)*
Acremonium-like, anamorphic Emeri-
cellopsis J.F.H. Beyma 1940 and Leu-
cosphaerina Arx 1987 (H)
Ascochytopsis-like, anamorphic Euca-
sphaeria Crous 2007 (C)
Cancellidium Tubaki 1975 (H)
Didymostilbe Henn. 1902, anamorphic
Peethambara Subram. & Bhat 1978 (H)
Escovopsis J.J. Muchovej & Della Lucia
1990 (H)
Hapsidospora Malloch & Cain 1970 (C)
Haptospora G.L. Barron 1991 (H)
Illosporiopsis D. Hawksw. 2001 (H)
Illosporium Mart. 1817 (H)
Imicles Shoemaker & Hambl. 2001 (H)
Memnoniella Höhn. 1923 (H)
Myrothecium Tode 1790 (H)
Parasarcopodium Melnik, S.J. Lee &
Crous 2004 (H)
Pseudomicrodochium B. Sutton 1975 (H)
Sarocladium W. Gams & D. Hawksw.
1976 (H)
Septomyrothecium Matsush. 1971, anam-
orphic Peethambara Subram. & Bhat
1978 (H)
Sorosporella Sorokīn 1888 (H)
Spicellum Nicot & Roquebert 1976 (H)
Sporothrix Hektoen & C.F. Perkins 1901,
anamorphic Leucosphaerina Arx 1987
(H)
Stachybotrys Corda 1837 (H)
Mycosphere Doi 10.5943/mycosphere/3/2/5
185
Stilbella Lindau 1900, anamorphic Eme-
ricellopsis J.F.H. Beyma 1940 and
Valsonectria Speg. 1881 (H)
Tilachlidium Preuss 1851 (H)
Trichothecium Link 1809 (H)*
Lulworthiales Kohlm., Spatafora & Volkm.-
Kohlm. 2000
Lulworthiaceae Kohlm., Spatafora & Volkm.-
Kohlm. 2000
Anguillospora Ingold 1942, anamorphic
Lindra I.M. Wilson 1956 (H)
Melanosporales N. Zhang & M. Blackw.
Ceratostomataceae G. Winter
Acrospeira Berk. & Broome 1857 (H)
Gonatobotrys Corda 1839, anamorphic
Melanospora Corda 1837 (H)
Harzia Costantin 1888, anamorphic Mela-
nospora Corda 1837 (H)
Papulaspora Preuss 1851, anamorphic
Melanospora Corda 1837 and Micr-
othecium Corda 1842 (H)
Proteophiala Cif. 1958, anamorphic Mela-
nospora Corda 1837, Persiciospora
P.F. Cannon & D. Hawksw. 1982,
Sphaerodes Clem. 1909, Pteridio-
sperma J.C. Krug & Jeng 1979 (H)
[Sphaerodes Clem. 1909 is a teleomorph
which was reported to produce ampul-
liform phialides on irregularly branched
conidiophores diectrly on ascomata or
on surrounding hyphae (Vujanovic &
Goh, 2009)]
Microascales Luttr. ex Benny & Kimbr.
Chadefaudiellaceae Faurel & Schotter ex
Benny & Kimbr.
Arthrographis-like, anamorphic Faurelina
Locq.-Lin. 1975 (H)
Gondwanamycetaceae Réblová, W. Gams &
Seifert
Custingophora Stolk, Hennebert &
Klopotek 1968, anamorphic Gondwa-
namyces G.J. Marais & M.J. Wingf. 1998
(H)*
Halosphaeriaceae E. Müll. & Arx ex Kohlm.
Cirrenalia Meyers & R.T. Moore 1960
(H)
Clavariopsis-like, anamorphic Corollo-
spora Werderm. 1922 (H)
Clavatospora Sv. Nilsson ex Marvanová
& Sv. Nilsson 1971, anamorphic Corol-
lospora Werderm. 1922 (H)
Culcitalna Meyers & R.T. Moore 1960,
anamorphic Halosarpheia Kohlm. & E.
Kohlm. 1977 (H)
Halosigmoidea Nakagiri, K.L. Pang &
E.B.G. Jones 2009, anamorphic Corol-
lospora Werderm. 1922 (H)
Humicola Traaen 1914, anamorphic Pseu-
dolignincola Chatmala & E.B.G. Jones
2006 and Lignincola Höhnk 1955 (H)
Monodictys S. Hughes 1958 anamorphic
Nereiospora E.B.G. Jones, R.G. Johns-
on & S.T. Moss 1983 (H)
Periconia Tode 1791, anamorphic Halo-
sphaeria Linder 1944 (H)
Sigmoidea J.L. Crane 1968, anamorphic
Corollospora Werderm. 1922 (H)
Trichocladium Harz 1871, anamorphic
Halosphaeria Linder 1944 (H)
Varicosporina Meyers & Kohlm. 1965,
anamorphic Corollospora Werderm.
1922 (H)
Microascaceae Luttr. ex Malloch
Atrhrographis G. Cochet ex Sigler & J.W.
Carmich. 1976, anamorphic Canar-
iomyces Arx (H)
Brachyconidiellopsis Decock, R.F. Casta-
ñeda & Adhikari 2004 (H)
Cephalotrichum Link 1809, anamorphic
Microascus Zukal 1885 (H)
Echinobotryum Corda 1831 (H)
Graphium Corda 1837, anamorphic Micro-
ascus Zukal 1885, Petriella Curzi 1930
and Pseudallescheria Negr. & I. Fisch.
1944 (H)
Humicola Traaen 1914, anamorphic Loph-
otrichus R.K. Benj. 1949 (H)
Parascedosporium Gilgado, Gené, Cano &
Guarro 2007 (H)
Scedosporium Sacc. ex Castell. & Chalm.
1919, anamorphic Pseudallescheria
Negr. & I. Fisch. 1944 and possibly
Petriella Curzi 1930 (H)
Scopulariopsis Bainier 1907, anamorphic
Microascus Zukal 1885 and ?Petriella
Curzi 1930 and Pithoascus Arx 1973
(H)
Mycosphere Doi 10.5943/mycosphere/3/2/5
186
Trichurus Clem. 1896 (H)
Wardomyces F.T. Brooks & Hansf. 1923,
anamorphic Microascus Zukal 188 (H)
Wardomycopsis Udagawa & Furuya 1978,
anamorphic Microascus Zukal 1885
(H)
Microascales, genera incertae sedis
Bisporostilbella Brandsb. & E.F. Morris
1971 (H)
Chalara-like, anamorphic Ceratocystis
Ellis & Halst. 1890 and Cornuvesica
C.D. Viljoen, M.J. Wingf. & K. Jacobs
2000 (H)
Gabarnaudia Samson & W. Gams 1974,
anamorphic Sphaeronaemella P. Karst.
1884 (H)
Sporendocladia G. Arnaud ex Nag Raj &
W.B. Kendr. 1975 (H)
Thielaviopsis Went 1893, anamorphic
Ceratocystis Ellis & Halst. 1890 (H)
Hypocreomycetidae, families incertae sedis
Plectosphaerellaceae W. Gams, Summerbell
& Zare
Acremonium Link 1809 (H)*
Acrostalagmus Corda 1838 (H)*
Gibellulopsis Bat. & H. Maia 1959 (H)
Musicillium Zare & W. Gams 2007 (H)
Plectosporium M.E. Palm, W. Gams &
Nirenberg 1995, anamorphic Plecto-
sphaerella Kleb. 1929 (H)
Stachylidium Link 1809 (H)*
Verticillium Nees 1816 (H)
Hypocreomycetidae, genera incertae sedis
Canalisporium Nawawi & Kuthub. 1989,
anamorphic Ascothailandia Sriindra-
sutdhi, Boonyuen, Sivichai & E.B.G.
Jones 2010 (H)
Dictyochaeta Speg. 1923, anamorphic
Ascocodinaea Samuels, Candoussau &
Magni 1997 (H)
?Glomerulispora Abdel-Wahab, Abdel-
Aziz & Nagahama 2010, anamorphic
Torpedospora Meyers 1957 (H)
Moheitospora Abdel-Wahab, Abdel-Aziz
& Nagahama 2010, anamorphic Junci-
gena Kohlm., Volkm.-Kohlm & O.E.
Erikss. 1997 (H)
Sporoschismopsis Hol.-Jech. & Hennebert
1972, anamorphic Porosphaerellopsis
Samuels & E. Müll. 1982 (H)
Subclass Sordariomycetidae O.E. Erikss. &
Winka
Boliniales P.F. Cannon 2001
Boliniaceae Rick 1931
Phaeoisaria Höhn. 1909 anamorphic
Lentomitella Höhn. 1906 (H)
Calosphaeriales M.E. Barr
Calosphaeriaceae Munk
Calosphaeriophora Réblová, L. Mostert,
W. Gams & Crous 2004, anamorphic
Calosphaeria Tul. & C. Tul. (H)
Lecythophora Nannf. 1934, anamorphic
Sulcatistroma A.W. Ramaley 2005 (H)
Phaeocrella Réblová, L. Mostert, W.
Gams & Crous 2004, anamorphic Tog-
niniella Réblová, L. Mostert, W. Gams
Crous 2004 (H)
Phialophora-like, anamorphic Jattaea
Berl. 1900 (H)
Stachybotrys-like, anamorphic Jattaea
Berl. 1900 (H)
Pleurostomataceae Réblová, L. Mostert, W.
Gams & Crous
Pleurostomophora Vijaykr., L. Mostert,
Jeewon, W. Gams, K.D. Hyde & Crous
2004, anamorphic Pleurostoma Tul. &
C. Tul. 1863 (H)
Chaetosphaeriales Huhndorf, A.N. Mill. &
F.A. Fernández
Chaetosphaeriaceae Réblová, M.E. Barr &
Samuels
Cacumisporium Preuss 1851, anamorphic
Chaetosphaeria Tul. & C. Tul. 1863
(H)
Catenularia Grove 1886, anamorphic
Chaetosphaeria Tul. & C. Tul. 1863
(H)
Chalara (Corda) Rabenh. 1844, anam-
orphic Ascochalara Réblová 1999 and
Chaetosphaeria Tul. & C. Tul. 1863
(H)*
Chloridium Link 1809, anamorphic Chae-
tosphaeria Tul. & C. Tul. 1863 and
Mycosphere Doi 10.5943/mycosphere/3/2/5
187
Melanopsammella Höhn. 1920 (H)
Codinaea Maire 1937, anamorphic Chae-
tosphaeria Tul. & C. Tul. 1863 (H)
?Cordana Preuss 1851, anamorphic Poro-
sphaerella E. Müll. & Samuels 1982
(H)
Craspedodidymum Hol.-Jech. 1972, ana-
morphic Chaetosphaeria Tul. & C. Tul.
1863 (H)
Cryptophiale Piroz. 1968, anamorphic
Chaetosphaeria Tul. & C. Tul. 1863
(H)
Custingophora Stolk, Hennebert & Klop-
otek 1968 anamorphic Chaetosphaeria
Tul. & C. Tul. 1863 (H)
Dictyochaeta Speg. 1923, anamorphic
Chaetosphaeria Tul. & C. Tul. 1863
and Striatosphaeria Samuels & E. Müll.
1979 (H)
Exserticlava S. Hughes 1978, anamorphic
Chaetosphaeria Tul. & C. Tul. 1863
(H)
Fusichalara S. Hughes & Nag Raj 1973,
anamorphic Chaetosphaeria Tul. & C.
Tul. 1863 (H)
Gonytrichum Nees & T. Nees 1818, ana-
morphic Chaetosphaeria Tul. & C. Tul.
1863 (H)
Harpophora W. Gams 2000, anamorphic
Chaetosphaeria Tul. & C. Tul. 1863
(H)
Hemicorynespora M.B. Ellis 1972, ana-
morphic Chaetosphaeria Tul. & C. Tul.
1863 (H)
Kylindria DiCosmo, S.M. Berch & W.B.
Kendr. 1983, anamorphic Chaetos-
phaeria Tul. & C. Tul. 1863 (H)
Menispora Pers. 1822, anamorphic Chae-
tosphaeria Tul. & C. Tul. 1863 (H)
Monilochaetes Halst. ex Harter 1916,
anamorphic Australiasca Sivan. &
Alcorn 2002 (H)*
Nawawia Marvanová 1980 (H)
Paliphora Sivan. & B. Sutton 1985 (H)
Phaeostalagmus W. Gams 1976, ana-
morphic Chaetosphaeria Tul. & C. Tul.
1863 (H)
Phialogeniculata Matsush. 1971 (H)
Phialophora-like, anamorphic Chaeto-
sphaeria Tul. & C. Tul. 1863 (H)
Pseudobotrytis Krzemien. & Badura 1954,
anamorphic Porosphaerella E. Müll. &
Samuels 1982 (H)
Pyrigemmula D. Magyar & R. Shoemaker
2011 (H)
Ramichloridium-like, anamorphic Chaeto-
sphaerides Matsush. 2003 (H)
Rattania Prabhugaonkar & Bhat 2009 (H)
Sporidesmium Link 1809, anamorphic
Umbrinosphaeria Réblová 1999, Miyo-
shiella Kawam. 1929 and Lecy-
thothecium Réblová & Winka 2001 (H)
Sporoschisma Berk. & Broome 1847, ana-
morphic Melanochaeta E. Müll., Harr
& Sulmont 1969 and Chaetosphaeria
Tul. & C. Tul. 1863 (H)
Stachybotrys Corda 1837, anamorphic
Melanopsamma Niessl 1876 (H)
Thozetella Kuntze 1891 (H)
Zanclospora S. Hughes & W.B. Kendr.
1965, anamorphic Chaetosphaeria Tul.
& C. Tul. 1863 (H)
Coniochaetales Huhndorf, A.N. Mill. & F.A.
Fernández
Coniochaetaceae Malloch & Cain
Cladobotryum-like, anamorphic Conioch-
aeta (Sacc.) Cooke 1887 (H)
?Dactylaria Sacc. 1880, anamorphic Syna-
ptospora Cain 1957 (H)
Lecythophora Nannf. 1934, anamorphic
Coniochaeta (Sacc.) Cooke 1887 (H)
Phialophora-like, anamorphic Barrina
A.W. Ramaley 1997 (H)
Diaporthales Nannf.
Cryphonectriaceae Gryzenh. & M.J. Wingf.
Aurapex Gryzenh. & M.J. Wingf. 2006
(C)
Chrysoporthella Gryzenh. & M.J. Wingf.
2004, anamorphic Chrysoporthe Gry-
zenh. & M.J. Wingf. 2004 (C)
Endothiella Sacc. 1906, anamorphic Cry-
phonectria (Sacc.) Sacc. & D. Sacc.
1905 (C)
Foliocryphia Cheewangkoon & Crous
2010 (C)
Prosopidicola Crous & C.L. Lennox 2004
(C)
Ursicollum Gryzenh. & M.J. Wingf. 2006
(C)
Diaporthaceae Höhn. ex Wehm.
Mazzantiella Höhn. 1925, anamorphic
Mycosphere Doi 10.5943/mycosphere/3/2/5
188
Mazzantia Mont. 1855 (C)
Phomopsis (Sacc.) Bubák 1905, anamor-
phic Diaporthe Nitschke 1870 (C)
Gnomoniaceae G. Winter
Asteroma DC. 1815, anamorphic Gnomo-
niella Sacc. 1881 and Plagiostoma
Fuckel 1870 and Pleuroceras Riess.
The type of the genus is considered to
be Montagnellina stellaris (Pers.)
Theiss. & Syd. 1915 (C)
?Depazea Fr. 1818, anamorphic Pleuro-
ceras Riess 1854 (C)
Diplodina Westend. 1857, anamorphic
Plagiostoma Fuckel 1870 (C)
Discosporium Sacc. & P. Syd. 1902 ana-
morphic Cryptodiaporthe Petr. 1921 C.
Tul. (C)
Discula Sacc. 1884, anamorphic Apiogno-
monia Höhn. 1917, Gnomonia Ces. &
De Not. 1863 and Amphiporthe Petr.
1971 (C)
Gloeosporidina Petr. 1921, anamorphic
Apiognomonia Höhn. 1917 (C)
Mazzantiella Höhn. 1925, anamorphic
Mamianiella Höhn. 1917 (C)
Millerburtonia Cif. 1951, anamorphic Pla-
giostoma Fuckel 1870 (C)
Phaeocytostroma Petr. 1921, anamorphic
Clypeoporthe Höhn 1919 (C)
?Septogloeum Sacc. 1880, anamorphic
Pleuroceras Riess 1854 (C)
Uniseta Ciccar. 1948, anamorphic Pla-
giostoma Fuckel 1870 (C)
?Zythia Fr. 1849 (C)
Melanconidaceae G. Winter
Dumortieria Westend. 1857 (as Cytospo-
rina Sacc. 1884), anamorphic Mela-
namphora Lafl. 1976 and Melogramma
Fr. 1849 (C)
Harknessia Cooke 1881, anamorphic Wue-
stneia Auersw. ex Fuckel 1863 (C)
Hendersonula Speg. 1880, anamorphic
Pseudovalsella Höhn. 1918 and Dictyo-
porthe Petr. 1955 (C)
Mastigosporella Höhn. 1914, anamorphic
Wuestneiopsis J. Reid & Dowsett 1990
(C)
Melanconiopsis Ellis & Everh. 1900, ana-
morphic Massariovalsa Sacc. 1882 (C)
Melanconium Link 1809, anamorphic
Melanconis Tul. & C. Tul. 1863 (C)
?Stilbospora Pers. 1794, anamorphic Pros-
thecium Fresen. 1852 (C)
Tubakia B. Sutton 1973, anamorphic
Dicarpella Syd. & P. Syd. 1921 (C)
Pseudoplagiostomaceae Cheewangkoon, M.J.
Wingf. & Crous
Cryptosporiopsis-like, anamorphic Pseu-
doplagiostoma Cheewangkoon, M.J.
Wingf. & Crous 2010 (C)
Pseudovalsaceae M.E. Barr
Coryneum Nees 1816, anamorphic Pseu-
dovalsa Ces. & De Not. 1863 (C)
Schizoparmeaceae Rossman
Coniella Höhn. 1918 (C)
Pilidiella Petr. & Syd. 1927, anamorphic
Schizoparme Shear 1923 (C)
Sydowiellaceae Lar. N. Vassiljeva
Cylindrosporella Höhn. 1916, anamorphic
Stegophora Syd. & P. Syd. 1916 (C)
Phoma-like, anamorphic Hapalocystis
Auersw. ex Fuckel 1863 (C)
Togniniaceae Réblová, L. Mostert, W. Gams
& Crous
Phaeoacremonium W. Gams, Crous &
M.J. Wingf. 1996, anamorphic
Togninia Berl. 1900 (H)
Valsaceae Tul. & C. Tul.
Amphicytostroma Petr. 1921, anamorphic
Amphiporthe Petr. 1971 (C)
Cytospora Ehrenb. 1818, anamorphic
Leucostoma (Nitschke) Höhn. 1917,
Valsa Fr. 1849, Valsella Fuckel 1870
and Valseutypella Höhn. 1919 (C)
Harpostroma Höhn. 1928, anamorphic
Leptosillia Höhn. 1928 (C)
?Lasmenia Speg. 1886, anamorphic Chad-
efaudiomyces Kamat, Rao, Patil &
Ullasa 1974 (C)
Phomopsis (Sacc.) Bubák 1905, anamor-
phic Clypeoporthella Petr. 1924 (C)
Diaporthales, genera incertae sedis
Craspedodidymum Hol.-Jech. 1972, ana-
morphic Thailandiomyces Pinruan,
Sakay., K.D. Hyde & E.B.G. Jones
Mycosphere Doi 10.5943/mycosphere/3/2/5
189
2008 (H)
Apoharknessia Crous & S.J. Lee 2004 (C)
Apomelasmia Grove 1937 (C)
Asterosporium Kunze 1819 (C)
Craspedodidymum Hol.-Jech. 1972, ana-
morphic Thailandiomyces Pinruan, Sa-
kay., K.D. Hyde & E.B.G. Jones 2008
(H)
Cytospora-like, anamorphic Pachytrype
Berl. ex M.E. Barr, J.D. Rogers & Y.M.
Ju 1993 (C)
Dwiroopa Subram. & Muthumary
?Greeneria Scribn. & Viala 1887 (C)
Hormonema Lagerb. & Melin 1927, ana-
morphic Valsaria Ces. & De Not. 1863
(H)
Hypodermina Höhn. 1916 (C)
Macrohilum H.J. Swart 1988 (C)
Natarajania Pratibha & Bhat 2006 (H)
Phomopsis-like, anamorphic Caudospora
Starbäck 1889 (C)
Rabenhorstia Fr. 1849, anamorphic Her-
cospora Fr. 1825 (C)
Sirococcus Preuss 1855 (C)
Stenocarpella Syd. & P. Syd. 1917 (C)
Synchaetomella Decock & Seifert 2005
(H)
Magnaporthales Thongk., Vijakr. & K.D.
Hyde
Magnaporthaceae P.F. Cannon
Clasterosporium Schwein. 1832, anamor-
phic Clasterosphaeria Sivan. 1984 (H)
Didymobotryum Sacc. 1886, anamorphic
Ceratosphaerella Huhndorf, Greif,
Mugambi & A.N. Mill. 2008 (H)
Harpophora W. Gams 2000, anamorphic
Gaeumannomyces Arx & D.L. Olivier
1952 and Magnaporthe R.A. Krause &
R.K. Webster 1972 (H)
Mycoleptodiscus Ostaz. 1968, anamorphic
Omnidemptus P.F. Cannon & Alcorn
1994 (H)
Nakataea Hara 1939, anamorphic Mag-
naporthe R.A. Krause & R.K. Webster
1972 (H)
Pyricularia Sacc. 1880, anamorphic Mag-
naporthe R.A. Krause & R.K. Webster
1972 (H)
Trichocladium-like, anamorphic Gaeuma-
nnomyces Arx & D.L. Olivier 1952 (H)
Ophiostomatales Benny & Kimbr.
Kathistaceae Malloch & M. Blackw.
Mattirolella S. Colla 1929 (C)
Termitaria Thaxt. 1920 (H)
Termitariopsis M. Blackw., Samson &
Kimbr. 1980 (H)
Ophiostomataceae Nannf.
Acremonium-like, anamorphic Frago-
sphaeria Shear 1923 (H)
Ambrosiella Brader ex Arx & Hennebert
1965, anamorphic Grosmannia Goid.
1936 and Ophiostoma Syd. & P. Syd.
1919 (H)
Graphium Corda 1837, anamorphic Op-
hiostoma Syd. & P. Syd. 1919 (H)
Hyalobelemnospora Matsush. 1993, ana-
morphic Subbaromyces Hesselt. 1953
(H)
Hyalorhinocladiella H.P. Upadhyay &
W.B. Kendr. 1975, anamorphic Cerato-
cystiopsis H.P. Upadhyay & W.B.
Kendr. 1975, Klasterskya Petr. 1940
and Ophiostoma Syd. & P. Syd. 1919
(H)
Hyalorhinocladiella-like, anamorphic Kla-
sterskya Petr. 1940 (H)
Leptographium Lagerb. & Melin 1927,
anamorphic Grosmannia Gold. 1936
(H)*
Pesotum J.L. Crane & Schokn. 1973,
anamorphic Ophiostoma Syd. & P. Syd.
1919 (H)
Pesotum-like, anamorphic Grosmannia
Gold. 1936 (H)
Phialographium H.P. Upadhyay & W.B.
Kendr. 1974, anamorphic Ophiostoma
Syd. & P. Syd. 1919 (C)
Raffaelea Arx & Hennebert 1965, anamor-
phic Grosmannia Goid. 1936 and
Ophiostoma Syd. & P. Syd. 1919 (H)
Sporothrix Hektoen & C.F. Perkins 1901,
anamorphic Ophiostoma Syd. & P. Syd.
1919 (H)
Sordariales Chad. ex D. Hawksw. & O.E.
Erikss.
Chaetomiaceae G. Winter
Acrophialophora Edward 1961 (H)
Botryotrichum Sacc. & Marchal 1885, ana-
morphic Chaetomium Kunze 1817 and
Farrowia D. Hawksw. (H)
Mycosphere Doi 10.5943/mycosphere/3/2/5
190
Humicola Traaen 1914, anamorphic Chae-
tomium Kunze 1817 (H)
Madurella Brumpt 1905 (H)
Myceliophthora Costantin 1892, anamorp-
hic Corynascus Arx 1973 and Thielavia
Zopf 1876 (H)
Papulaspora-like, Preuss 1851, anamorp-
hic Chaetomium Kunze 1817 (H)
Phialophora-like, anamorphic Chaetomid-
ium (Zopf) Sacc. 1882 (H)
Retroconis de Hoog & Bat. Vegte 1989
(H)
Taifanglania Z.Q. Liang, Y.F. Han, H.L.
Chu & R.T.V. Fox 2009, anamorphic
Thielavia Zopf 1876 (H)
Trichocladium Harz 1871, anamorphic
Chaetomium Kunze 1817 (H)
Trichosporiella Kamyschko 1960, ana-
morphic Thielavia Zopf 1876 (H)
Lasiosphaeriaceae Nannf.
Angulimaya Subram. & Lodha 1964, ana-
morphic Bombardioidea C. Moreau ex
N. Lundqv. 1972 (H)
Bagadiella Cheewangkoon & Crous 2009
(H)
Cladorrhinum Sacc. & Marchal 1885,
anamorphic Apiosordaria Arx & W.
Gams 1967, Apodospora Cain & J.H.
Mirza 1970, Arnium Nitschke ex G.
Winter 1873, Cercophora Fuckel 1870
and Podospora Ces. 1856 (H)
Mammaria Ces. ex Rabenh. 1854, ana-
morphic Pseudocercophora Subram. &
Sekar 1986 and Cercophora Fuckel
1870 (H)
Phialophora-like, anamorphic Cercophora
Fuckel 1870, Eosphaeria Höhn. 1917,
Lasiosphaeria Ces. & De Not. 1863 and
Podospora Ces. 1856 (H)
Sordariaceae G. Winter
Chrysonilia Arx 1981, anamorphic Neuro-
spora Shear & B.O. Dodge 1927 (H)
Trichosporiella Kamyschko 1960, anam-
orphic Diplogelasinospora Cain 1961
(H)
Virgariella S. Hughes 1953, anamorphic
Effetia Bartoli, Maggi & Persiani 1984
(H)
Sordariales, genera incertae sedis
Kionochaeta P.M. Kirk & B. Sutton 1986
(H)
Lasiadelphia Réblová & W. Gams 2011
(H)
Lunulospora Ingold 1942 (H)
Madurella Brumpt 1905 (H)
Monodictys S. Hughes 1958 anamorphic
Ascotaiwania Sivan. & H.S. Chang
1992 (H)
Monotosporella S. Hughes 1958, anam-
orphic Ascotaiwania Sivan. & H.S.
Chang 1992 (H)
Papulospora Preuss 1851, anamorphic
Sordaria Ces. & De Not. 1863 (H)
Ramophialophora M. Calduch, Stchigel,
Gené & Guarro 2004 (H)
Trichocladium-like, anamorphic Asco-
lacicola Ranghoo & K.D. Hyde 1998,
Ascotaiwania Sivan. & H.S. Chang
1992 (H)
Ypsilonia Lév. 1846, anamorphic Acantho-
theciella Höhn. 1911 (H)
Sordariomycetidae, families incertae sedis
Amplistromataceae Huhndorf, A.N. Mill., M.
Greif & Samuels
Acrodontium-like, anamorphic Amplistro-
ma Huhndorf, A.N. Mill., M. Greif &
Samuels 2009 (H)
Pseudogliomastix W. Gams 1985, ana-
morphic Wallrothiella Sacc. 1882 (H)
Annulatascaceae S.W. Wong, K.D. Hyde &
E.B.G. Jones
Phaeoisaria Höhn. 1909, anamorphic
Rhamphoria Niessl 1876 (H)
Phialophora-like, anamorphic Teraco-
sphaeria Réblová & Seifert 2007 (H)
Cephalothecaceae Höhn.
Acremonium-like, anamorphic Alberti-
niella Kirschst. 1936 (H)
Chalara (Corda) Rabenh. 1844, anamor-
phic Cryptendoxyla Malloch & Cain
1970 (H)*
Phialemonium W. Gams & McGinnis
1983 (H)
Tritirachium Limber 1940, anamorphic
Cephalotheca Fuckel 1871 (H)
Helminthosphaeriaceae Samuels, Candoussau
& Magní
Diplococcium Grove 1885, anamorphic
Mycosphere Doi 10.5943/mycosphere/3/2/5
191
Helminthosphaeria Fuckel 1870 (H)
Endophragmiella B. Sutton 1973, anamor-
phic Echinosphaeria A.N. Mill. &
Huhndorf 2004 (H)
Selenosporella-like, anamorphic Ruzenia
O. Hilber 2002 (H)
Spadicoides S. Hughes 1958, anamorphic
Tengiomyces Réblová 1999 (H)
Vermiculariopsiella Bender 1932, anamor-
phic Echinosphaeria A.N. Mill. &
Huhndorf 2004 (H)
Sordariomycetidae, genera incertae sedis
Sterigmatobotrys Oudem. 1886 (H)*
Brachysporiella Bat. 1952, anamorphic
Ascotaiwania Sivan. & H.S. Chang
1992 (H)
Conioscypha Höhn. 1904, anamorphic
Conioscyphascus Réblová & Seifert
2004 (H)
Endophragmiella B. Sutton 1973, ana-
morphic Phaeotrichosphaeria Sivan.
1983 (H)
Menisporopsis S. Hughes 1952, anamor-
phic Menisporopascus Matsush. 2003
(H)
Phialophora-like, anamorphic Linocarpon
Syd. & P. Syd. 1917 (H)
Pleurothecium Höhn. 1919, anamorphic
Carpoligna F.A. Fernández & Huhn-
dorf 1999 (H)
Ramichloridium-like, anamorphic Barba-
tosphaeria Reblova 2007 (H)
Rhodoveronaea Arzanlou, W. Gams &
Crous 2007 (H)
Spadicoides S. Hughes 1958 (H)
Sporothrix-like, anamorphic Barbatosph-
aeria Reblova 2008 (H)
Subclass Xylariomycetidae sensu O.E. Erikss.
& Winka
Xylariales Nannf.
Amphisphaeriaceae G. Winter
Bartalinia Tassi 1900 (C)
?Beltraniella Subram. 1952, anamorphic
Leiosphaerella Höhn. 1919 (H)
Bleptosporium Steyaert 1961, anamorphic
Amphisphaeria Ces. & De Not. 1863
(C)
Ceratosporium Schwein. 1832, anamor-
phic Iodosphaeria Samuels, E. Müll. &
O. Petrini 1987 (H)
?Chaetoconis Clem. 1909, anamorphic
Ceriospora Niessl 1876 (C)
Discosia Lib. 1837, anamorphic Adisciso
Kaz. Tanaka, Okane & Hosoya 2011
(C)*
Doliomyces Steyaert 1961 (C)
?Hyalotiopsis Punith. 1970, anamorphic
Ellurema Nag Raj & W.B. Kendr. 1986
(C)
Immersidiscosia Kaz. Tanaka, Okane &
Hosoya 2011 (C)*
Labridella Brenckle 1929, anamorphic
Griphosphaerioma Höhn. 1918 (C)
Labridium Vestergr. 1897, anamorphic
Discostroma Clem. 1909 (C)
Microdochium Syd. 1924, anamorphic
Monographella Petr. 1924 (H)
Monochaetia (Sacc.) Allesch. 1902 (C)
Monochaetinula Muthumary, Abbas & B.
Sutton 1986 (C)
Monochaetiopsis L. Cai, Jeewon & K.D.
Hyde 2003, anamorphic Dyrithiopsis L.
Cai, R. Jeewon & K.D. Hyde 2003 (C)
Morinia Berl. & Bres. 1889 (C)
?Pestalotia De Not. 1841, anamorphic
Broomella Sacc. 1883 (C)
Pestalotiopsis Steyaert 1949, anamorphic
Neobroomella Petr. 1947 and Pestalo-
sphaeria M.E. Barr 1975 (C)
Phlogicylindrium Crous, Summerb. &
Summerell 2006 (H)
Sarcostroma Cooke 1871, anamorphic
Griphosphaerioma Höhn. 1918 (C)
Seimatosporium Corda 1833, anamorphic
anamorphic Discostroma Clem. 1909
(C)
Seiridium Nees 1816, anamorphic Blogia-
scospora Shoemaker, E. Müll. &
Morgan-Jones 1966 and Lepteutypa
Petr. 1923 (C)
Selenosporella G. Arnaud ex MacGarvie
1969, anamorphic Oxydothis Penz. &
Sacc. 1898 and Iodosphaeria Samuels,
E. Müll. & O. Petrini 1987 (H)
Sporocadus Corda 1839 (C)
Synnemapestaloides T. Handa & Y.
Harada 2004 (H)
?Truncatella Steyaert 1949 anamorphic
Broomella Sacc. 1883 (C)
Vermisporium H.J. Swart & M.A. Will.
1983 (C)
Zetiasplozna Nag Raj 1993 (C)
Mycosphere Doi 10.5943/mycosphere/3/2/5
192
Diatrypaceae Nitschke
Dumortieria Westend. 1857 (some as
Cytosporina Sacc. 1884), anamorphic
Cryptosphaeria Ces. & De Not. 1863,
Diatrype Fr. 1849, Eutypa Tul. & C.
Tul. 1863 and Eutypella (Nitschke)
Sacc. 1875 (C)
Libertella Desm. 1830, anamorphic
Diatrype Fr. 1849, Diatrypella (Ces. &
De. Not.) De. Not. 1863, Eutypa Tul. &
C. Tul. 1863 and Eutypella (Nitschke)
Sacc. 1875 (C)
Diatrypaceae Nitschke 1869
Phaeoisaria Höhn. 1909 anamorphic
Eutypella (Nitschke) Sacc. 1875 (H)
Hyponectriaceae Petr.
Arthrinium Kunze 1817, anamorphic
Physalospora Niessl (H)
Aureobasidium Viala & G. Boyer 1891,
anamorphic Discosphaerina Höhn.
1917 (H)
Beltraniella Subram. 1952, anamorphic
Pseudomassaria Jacz. 1894 (H)
Hormonema-like, anamorphic Discosp-
haerina Hohn 1917 (H)
Kabatia Bubák 1904, anamorphic Disco-
sphaerina Höhn. 1917 (C)
Kabatiella Bubák 1907, anamorphic Dis-
cosphaerina Höhn. 1917 (H)
Sarcophoma Höhn. 1916, anamorphic Dis-
cosphaerina Hohn 1917 (C)
Selenophoma Maire 1907, anamorphic
Discosphaerina Hohn 1917 (C)
Graphostromataceae M.E. Barr, J.D. Rogers
& Y.M. Ju
Nodulisporium-like, anamorphic Grapho-
stroma Piroz. 1974 (H)
Xylariaceae Tul. & C. Tul.
Acanthodochium Samuels, J.D. Rogers &
Nagas., anamorphic Astrocystis Berk. &
Broome and Collodiscula I. Hino &
Katum. 1955 (H)
Arthroxylaria Seifert & W. Gams 2002
Dematophora R. Hartig 1883, anamorphic
Rosellinia De Not. 1844 (H)
?Dicyma Boulanger 1897, anamorphic
Ascotricha Berk. 1838 (H)
Geniculisynnema Okane & Nakagiri 2007,
anamorphic Nemania Gray 1821 (H)
Geniculosporium Chesters & Greenh.
1964, anamorphic Entoleuca Syd. 1922,
Leprieuria Laessøe, J.D. Rogers &
Whalley 1989, Nemania Gray 1821,
Phylacia Lév. 1845, Podosordaria Ellis
& Holw. 1897, Rosellinia De Not. 1844
and Xylaria Hill ex Schrank 1789 (H)
Geniculosporium-like, anamorphic Halo-
rosellinia Whalley, E.B.G. Jones, K.D.
Hyde & Læssøe 2000, Jumillera J.D.
Rogers, Y.M. Ju & San Martín 1997
and Kretzschmaria Fr. 1849 (H)
Hypocreodendron Henn. 1897, anamor-
phic Discoxylaria J.C. Lindq. & J.E.
Wright 1964 (H)
Libertella Desm. 1830 (C)
Libertella-like, anamorphic Barrmaelia
Rappaz 1995, Creosphaeria Theiss.
1910, Jumillera J.D. Rogers, Y.M. Ju &
San Martín 1997 and Lopadostoma
(Nitschke) Traverso 1906 (C)
Lindquistia Subram. & Chandrash. 1977,
anamorphic Poronia Willd. 1787 (H)
Masoniomyces J.L. Crane & Dumont
1975, anamorphic Camillea Fr. 1849
(H)
Mirandina-like, anamorphic Kretzschma-
riella Viégas 1944 (H)
Moelleroclavus Henn. 1902, anamorphic
Xylaria Hill ex Schrank 1789 (H)
Muscodor Worapong, Strobel & W.M.
Hess 2001 (H)
Nodulisporium Preuss 1849, anamorphic
Annulohypoxylon Y.M. Ju, J.D. Rogers
& H.M. Hsieh 2005, Biscogniauxia
Kuntze 1891, Daldinia Ces. & De Not.
1863, Entonaema Möller 1901, Hypo-
xylon Bull. 1791, Induratia Samuels, E.
Müll. & O. Petrini, Pyrenomyxa Mor-
gan 1895 (as Pulveria Malloch &
Rogerson 1977), Thamnomyces Ehrenb.
1820, Theissenia Maubl. 1914, Thue-
menella Penz. & Sacc. 1898 and
Xylaria Hill ex Schrank (H)
Nodulosporium-like, anamorphic Rhopalo-
stroma D. Hawksw. 1977 and Ruwen-
zoria J. Fornier, M. Stadler, Læssøe &
C. Decock 2010 (H)
Padixonia Subram. 1972, anamorphic
Xylaria Hill ex Schrank (H)
Mycosphere Doi 10.5943/mycosphere/3/2/5
193
Rhinocladiella-like, anamorphic Obolari-
na Pouzar 1986 (H)
Sporothrix-like, anamorphic Rostrohypo-
xylon J. Fourn. & M. Stadler 2010 and
Ruwenzoria J. Fornier, M. Stadler,
Læssøe & C. Decock 2010 (H)
Triplicaria P. Karst. 1889, anamorphic
Hypoxylon Bull. 1791 (H)
Virgaria Nees 1816, anamorphic Ascovir-
garia J.D. Rogers & Y.M. Ju 2002 (H)
Virgariella-like, anamorphic Hypoxylon
Bull. 1791 (H)
Vivantia J.D. Rogers, Y.M. Ju & Cand.
1996 (H)
Xylocladium P. Syd. ex Lindau 1900,
anamorphic Camillea Fr. 1849 (H)
Xylocoremium J.D. Rogers 1984, anamor-
phic Xylaria Hill ex Schrank 1789 (H)
Xylariales, genera incertae sedis
Chaetochalara B. Sutton & Piroz. 1965
(H)
Chalara (Corda) Rabenh. 1844 (H)*
Hadrotrichum Fuckel 1865 (H)
Humicola-like, anamorphic Ascotrichella
Valldos. & Guarro 1988 (H)
Idriella P.E. Nelson & S. Wilh. 1956,
anamorphic Phomatospora Sacc. 1875
(H)
Melanographium Sacc. 1913, possibly
anamorphic Lasiobertia Sivan. 1978
(H)
Nodulisporium-like, anamorphic Coniola-
riella D. Garcıá, Stchigel & Guarro (H)
Rhinocladiella-like, anamorphic Coniola-
riella D. Garcıá, Stchigel & Guarro (H)
Selenosporella G. Arnaud ex MacGarvie
1969, anamorphic Diatrypasimilis J.
Zhou, & Kohlm. 2010 (H)
Sporidesmina Subram. & Bhat 1989 (H)
Subramaniomyces Varghese & V.G. Rao
1980 (H)
Sordariomycetes, orders incertae sedis
Lulworthiales Kohlm., Spatafora & Volkm.-
Kohlm.
Lulworthiaceae Kohlm., Spatafora & Volkm.-
Kohlm.
Anguillospora Ingold 1942, anamorphic
Lindra I.M. Wilson 1956 (H)
Cumulospora I. Schmidt 1985, anamor-
phic Lulwoana Kohlm., Volkm.-
Kohlm., J. Campb., Spatafora & Grä-
fenhan 2005 and Lulworthia G.K.
Sutherl. 1916 (H)
Halazoon Abdel-Aziz, Abdel-Wahab &
Nagahama 2010, anamorphic Lulwort-
hia G.K. Sutherl. 1916 (C)
Hydea K.L. Pang & E.B.G. Jones 2010
(H)
Matsusporium E.B.G. Jones & K.L. Pang
2010 (H)
Moleospora Abdel-Wahab, Abdel-Aziz &
Nagahama 2010 (H)
Moromyces Abdel-Wahab, K.L. Pang,
Nagahama, Abdel-Aziz & E.B.G. Jones
2010 (H)
Zalerion R.T. Moore & Meyers 1962,
anamorphic Lulwoana Kohlm., Volkm.-
Kohlm, J. Campb., Spatafora & Gräf-
enhan 2005 (H)
Lulworthiales, genera incertae sedis
Halazoön Abdel-Aziz, Abdel Wahab &
Nagah. 2011 (H)
Phyllachorales M.E. Barr
Phyllachoraceae Theiss. & H. Syd.
Acerviclypeatus Hanlin 1990, anamorphic
Ophiodothella Boyd 1934 (C)
Baeumleria Petr. & Syd. 1927, anamo-
rphic Phyllachora Nitschke ex Fuckel
1870 (as Trabutia Sacc. & Roum. 1881)
(C)
Colletogloeum-like, anamorphic Cocco-
diella Hara 1909 (C)
Cyclodomus Höhn. 1909, anamorphic
Maculatifrons K.D. Hyde 1996 (C)
Diachorella Höhn. 1918, anamorphic
Diachora Müll. 1893 (C)
Hysterodiscula Petr. 1942, anamorphic
Orphnodactylis Malloch & A. Mallik
1998 (C)
Linochora Höhn. 1910, anamorphic Phyl-
lachora Nitschke ex Fuckel 1870 (C)
Mycohypallage B. Sutton 1963, anamor-
phic Deshpandiella Kamat & Ullasa
1973 (C)
Oswaldina Rangel 1921, anamorphic Api-
osphaeria Höhn. 1909 (C)
Polystigmina Sacc. 1884, anamorphic Pol-
ystigma DC. 1815 (C)
Pseudothiopsella Petr. 1928, anamorphic
Pseudothiella Petr. 1928 (C)
Mycosphere Doi 10.5943/mycosphere/3/2/5
194
Rhodosticta Woron. 1911, anamorphic
Polystigma DC. 1815 and Stigmatula
(Sacc.) Syd. & P. Syd. 1901 (C)
Trichosphaeriales M.E. Barr 1983
Trichosphaeriaceae G. Winter 1885
Brachysporium Sacc. 1886, anamorphic
Cryptadelphia Réblová & Seifert 2004
(H)
Koorchaloma Subram. 1953, anamorphic
Kananascus Nag Raj 1984 (H)
Stanjehughesia Subram. 1992, anamorphic
Miyoshiella Kawam. 1929 and Umbri-
nosphaeria Réblová 1999 (H)
Stromatographium Höhn. 1907, anamor-
phic Fluviostroma Samuels & E. Müll.
1980 (H)
Sordariomycetes, family incertae sedis
Apiosporaceae K.D. Hyde, J. Fröhlich, Joanne
E. Taylor & M.E. Barr
Arthrinium Kunze 1817, anamorphic Apio-
spora Sacc. 1875 (H)
Cordella Speg. 1886, anamorphic Apio-
spora Sacc. 1875 (H)
Pteroconium Sacc. ex Grove 1914, anamo-
rphic Apiospora Sacc. 1875 (H)
Scyphospora L.A. Kantsch. 1928, anamor-
phic Apiospora Sacc. 1875 (C)
Thyridiaceae O.E. Erikss. & J.Z. Yue
Pleurocytospora Petr. 1923, anamorphic
Thyridium Nitschke 1867 (C)
Sordariomycetes, genera incertae sedis
Cryptomycella Höhn. 1925, anamorphic
Cryptomycina Höhn. 1977 (C)
?Custingophora Stolk, Hennebert & Klo-
potek 1968 (H)
Didymostilbe-like, anamorphic Ornatispo-
ra K.D. Hyde, Goh, Joanne E. Taylor,
J. Fröhl. 1999 (H)
Dinemasporium Lév. 1846, anamorphic
Phomatospora Sacc. 1875 (C)
Ellisembia Subram. 1992 (H)
Linkosia A. Hern. Gut. & B. Sutton 1997
(H)
Myrmecridium Arzanlou, W. Gams &
Crous 2007 (H)
Nigrospora Zimm. 1902, anamorphic
Khuskia H.J. Huds. 1963 (H)
Penicillifer Emden 1968, anamorphic
Stellosetifera Matsush.1996 (H)
Phomatosporella Tak. Kobay. & K. Sasaki
1982, anamorphic Phomatospora Sacc.
1875 (C)
Ramichloridium-like, anamorphic Tectoni-
dula Réblová 2009 (H)
Sporothrix-like, anamorphic Phomato-
spora Sacc. 1875 and Tectonidula Réb-
lová 2009 (H)
Subphylum PEZIZOMYCOTINA, orders
incertae sedis
Lahmiales O. E. Erikss.
Lahmiaceae O. E. Erikss.
Cyphellophora G.A. de Vries 1962, ana-
morphic Lahmia Körb. 1861 (H)
ASCOMYCOTA, families incertae sedis
Families and genera that cannot be
placed in any of the classes and orders accepted
in the present classification with a high degree
of probability are listed below.
Amorphothecaceae Parbery
Hormoconis Arx & G.A. de Vries 1973,
anamorphic Amorphotheca Parbery
1969 (H)
Batistiaceae Samuels & K.F. Rodrigues
Acrostroma Seifert 1987, anamorphic
Batistia Cif. 1958 (H)
Pseudeurotiaceae Malloch & Cain
Teberdinia Sogonov, W. Gams, Summerb.
& Schroers 2005, anamorphic Pseudeu-
rotium J.F.H. Beyma 1937 (H)
Seuratiaceae Vuill. ex M.E. Barr
Atichia Flot. 1850, anamorphic Seuratia
Pat. 1904 (H)
ASCOMYCOTA, genera incertae sedis
Abropelta B. Sutton 1986 (C)
Acarellina Bat. & H. Maia 1960 (C)
Acaroconium Kocourk. & D. Hawksw.
2008 (C)
Acarocybe Syd. 1937 (H)
Acarocybella M.B. Ellis 1960 (H)
Acarocybellina Subram. 1992 (H)
Acarocybiopsis J. Mena, A. Hern. Gut. &
Mercado 1999 (H)
Acaropeltis Petr. 1937 (C)
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195
Achoropeltis Syd. 1929 (C)
Aciculariella G. Arnaud 1954 (H)
Acontium Morgan 1902 (H)
Acremoniula G. Arnaud 1954 (H)
Acrodictyella W.A. Baker & Partr. 2001
(H)
Acrodictyopsis P.M. Kirk 1983 (H)
Acrodictys M.B. Ellis 1961 (H)
Acrodontiella U. Braun & Scheuer 1995
(H)
Acrophragmis Kiffer & Reisinger 1970
(H)
Acrostaurus Deighton & Piroz. 1972 (H)
Actinocladium Ehrenb. 1819 (H)
Actinotexis Arx 1960 (C)
Actinothecium Ces. 1854 (H)
Actinothyrium Kunze 1823 (C)
Acumispora Matsush. 1980 (H)
Aegeritella Bałazy & J. Wiśn. 1974 (H)
Aenigmatospora R.F. Castañeda, Saikawa,
Guarro & Calduch 1999 (?H)
Agaricodochium X.J. Liu 1981 (H)
Agarwalomyces R.K. Verma & Kamal
1987 (H)
Agrabeeja Subram. 1995 (H)
Agyriella Sacc. 1884 (H)
Agyriellopsis Höhn. 1903 (C)
Ahmadia Syd. 1939 (C)
Ajrekarella Kamat & Kalani 1964 (C)
Alatosessilispora K. Ando & Tubaki 1984
(H)
Alciphila Harmaja 2002 (H)
Algonquinia R.F. Castañeda & W.B.
Kendr. 1991 (H)
Alloneottiosporina Nag Raj 1993 (C)
Allothyriella Bat., Cif. & Nascim. 1959
(C)
Allothyrina Bat. & J.L. Bezerra 1964 (C)
Allothyriopsis Bat., Cif. & H. Maia 1959
(C)
Alpakesa Subram. & K. Ramakr. 1954 (C)
Alpakesiopsis Abbas, B. Sutton, Ghaffar &
A. Abbas 2003 (C)
Alveophoma Alcalde 1952 (C)
Alysidiopsis B. Sutton 1973 (H)
Amallospora Penz. 1897 (H)
Amblyosporium Fresen. 1863 (H)
Amerodiscosiella M.L. Farr 1961 (C)
Amerodiscosiellina Bat. & Cavalc. 1966
(C)
Amerosporiopsis Petr. 1941 (C)
Amerosympodula Matsush. 1996 (H)
Amoenodochium Peláez & R.F. Castañeda
1996 (H)
Amoenomyces R.F. Castañeda, Saikawa &
Hennebert 1996 (H)
Amphichaetella Höhn. 1916 (H)
Amphophialis R.F. Castañeda, W.B.
Kendr. & Guarro 1998 (H)
Amphoropycnium Bat. 1963 (C)
Ampullicephala R.F. Castañeda, Minter &
M. Stadler 2009 (H)
Ampulliferina B. Sutton 1969 (H)
Anaphysmene Bubák 1906 (C)
Anarhyma M.H. Pei & Z.W. Yuan 1986
(C)
Anaselenosporella Heredia, R.F. Casta-
ñeda & R.M. Arias 2010 (H)
Ancoraspora Mig. Rodr. 1982 (H)
Ancorasporella J. Mena, Mercado &
Heredia 1998 (H)
Angiopomopsis Höhn. 1912 (C)
Angulospora Sv. Nilsson 1962 (H)
Annellodentimyces Matsush. 1985 (H)
Annellodochium Deighton 1969 (H)
Annellolacinia B. Sutton 1964 (C)
Annellophora S. Hughes 1952 (H)
Annellophorella Subram. 1962 (H)
Annellophragmia Subram. 1963 (H)
Annellosympodia McTaggart, R.G. Shivas
& U. Braun 2007 (H)
Antennopsis R. Heim 1952 (H)
Anthopsis Fil. March., A. Fontana & Luppi
Mosca 1977 (H)
Anthracoderma Speg. 1888 (C)
Antromyces Fresen. 1850 (H)
Anulohypha Cif. 1962 (H)
Anungitea B. Sutton 1973 (H)
Anungitopsis R.F. Castañeda & W.B.
Kendr. 1990 (H)
Aoria Cif. 1962 (C)
Aphanofalx B. Sutton 1986 (C)
Apiocarpella Syd. & P. Syd. 1919 (C)
Apogloeum Petr. 1954 (C)
Aporellula B. Sutton 1986 (C)
Aposporella Thaxt. 1920 (H)
Arachnophora Hennebert 1963 (H)
Arachnospora R.F. Castañeda, Minter &
Camino 2003 (H)
Arborillus Munt.-Cvetk. & Gómez-Bolea
1998 (H)
Arborispora K. Ando 1986 (H)
Arbusculina Marvanová & Descals 1987
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196
(H)
Arcuadendron Sigler & J.W. Carmich.
1976 (H)
Ardhachandra Subram. & Sudha 1978 (H)
Argopericonia B. Sutton & Pascoe 1987
(H)
Aristastoma Tehon 1933 (C)
Arthrobotryum Ces. 1854 (C)
Arthrocladium Papendorf 1969 (H)
Arthrocristula Sigler, M.T. Dunn & J.W.
Carmich. 1982 (H)
Arthrosporium Sacc. 1880 (H)
Arthrowallemia R.F. Castañeda, Dania
García & Guarro 1998 (H)
Articulophora C.J.K. Wang & B. Sutton
1982 (H)
Artocarpomyces Subram. 1996 (H)
Arxiella Papendorf 1967 (H)
Asbolisiomyces Bat. & H. Maia 1961 (C)
Ascochytopsis Henn. 1905 (C)
Ascochytulina Petr. 1922 (C)
Ascospora Mont. 1849 (C)
Ashtaangam Subram. 1995 (H)
Aspilaima Bat. & H. Maia 1961 (C)
Astelechia Cif. 1962 (H)
Asterinothyriella Bat. & Cif. 1959 (C)
Asterinothyrium Bat., Cif. & H. Maia 1959
(C)
Asteroconium Syd. & P. Syd. 1903 (C)
Asteromyces F. Moreau & V. Moreau, in
Hennenert 1962 (H)
Asterostomopora Bat. & H. Maia 1960 (C)
Asterostomopsis Bat., Cif. & H. Maia 1959
(C)
Asterostomulina Bat., J.L. Bezerra & H.
Maia 1964 (C)
Astronatelia Bat. & H. Maia 1962 (C)
Atractilina Dearn. & Barthol. 1924 (H)
Atrosetiphiale Matsush. 1995 (H)
Aurantiosacculus Dyko & B. Sutton 1979
(C)
Aurosphaeria Sun J. Lee, Strobel, Eisen-
man, Geary, Vargas & S.A. Strobel
2009 (C)
Avesicladiella W.P. Wu, B. Sutton &
Gange 1997 (H)
Avettaea Petr. & Syd. 1927 (C)
Bachmanniomyces D. Hawksw. 1981 (C)
Bacillopeltis Bat. 1957 (C)
Bactridiopsis Henn. 1904 (H)
Bactridium Kunze 1817 (H)
Bactrodesmiastrum Hol.-Jech 1984 (H)
Bactrodesmiella M.B. Ellis 1959 (H)
Badarisama Kunwar, J.B. Manandhar &
J.B. Sinclair 1986 (H)
Bahuchashaka Subram. 1978 (H)
Bahugada K.A. Reddy & Vasant Rao
1984 (H)
Bahukalasa Subram. & Chandrash. 1979
(H)
Bahusandhika Subram. 1956 (H)
Balaniopsis P.M. Kirk 1985 (H)
Balanium Wallr. 1833 (H)
Barbarosporina Ķirulis 1942 (C)
Barnettella D. Rao & P.Rag. Rao 1964 (C)
Basauxia Subram. 1995 (H)
Batistina Peres 1961 (C)
Beauveriphora Matsush 1975 (H)
Beccopycnidium F. Stevens 1930 (C)
Beejadwaya Subram. 1978 (H)
Belemnospora P.M. Kirk 1981 (H)
Bellulicauda B. Sutton 1967 (C)
Beltrania Penz. 1882 (H)
Beltraniomyces Monohar, D.K. Agarwal &
Rao 2003 (H)
Beltraniopsis Bat. & J.L. Bezerra 1960 (H)
Beniowskia Racib. 1900 (H)
Benjaminia S. Ahmad 1967 (C)
Benjpalia Subram. & Bhat 1989 (H)
Bharatheeya D'Souza & Bhat 2002 (H)
Bhatia W.A. Baker & Morgan-Jones 2009
(H)
Bibanasiella R.F. Castañeda & W.B.
Kendr. 1991 (H)
Biflagellospora Matsush. 1975 (H)
Biflagellosporella Matsush. 1993 (H)
Bimeris Petr. 1949 (C)
Bioconiosporium Bat. & J.L. Bezerra 1964
(H)
Biophomopsis Petr. 1931 (C)
Bispora Corda 1837 (H)
Bisseomyces R.F. Castañeda 1985 (H)
Blarneya D. Hawksw., Coppins & P.
James 1980 (H)
Blastacervulus H.J. Swart 1988 (C)
Blastocatena Subram. & Bhat 1989 (H)
Blastodictys M.B. Ellis 1976 (H)
Blastofusarioides Matsush. 1996 (H)
Blastophorella Boedijn 1937 (H)
Blastophorum Matsush. 1971 (H)
Blastophragma Subram. 1995 (H)
Blennoria Moug. & Fr. 1825 (C)
Blennoriopsis Petr. 1919 (C)
Blodgettia Harv. 1858 (H)
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197
Bomplandiella Speg. 1886 (H)
Bostrichonema Ces. 1867 (H)
Botryoderma Papendorf & H.P. Upadhyay
1969 (H)
Botryodiplodina Dias & Sousa da Câmara
1954 (C)
Botryomonilia Goos & Piroz. 1975 (H)
Botryosporium Corda 1831 (H)
Brachydesmiella G. Arnaud ex S. Hughes
1961 (H)
Brachysporiellina Subram. & Bhat 1989
(H)
Brachysporiopsis Yanna, W.H. Ho & K.D.
Hyde 2004 (H)
Brefeldiopycnis Petr. & Cif. 1932 (H)
Brevicatenospora R.F. Castañeda, Minter
& Saikawa 2006 (H)
Briosia Cavara 1888 (H)
Brycekendrickia Nag Raj 1973 (C)
Bulbilopycnis Matsush. 1996 (C)
Bulbocatenospora R.F. Castañeda & Iturr.
2000 (H)
Bullaserpens Bat., J.L. Bezerra & Cavalc.
1965 (C)
Calcarispora Marvanová & Marvan 1963
(H)
Calcarisporiella de Hoog 1974 (H)
Calceispora Matsush. 1975 (H)
Callistospora Petr. 1955 (C)
Calocline Syd. 1939 (C)
?Calongia D. Hawksw. & Etayo 2010 (H)
Camarographium Bubák 1916 (C)
Camaropycnis E.K. Cash 1945 (C)
Camarosporellum Tassi 1902 (C)
Camarosporiopsis Abbas, B. Sutton &
Ghaffar 2000 (C)
Camposporidium Nawawi & Kuthub. 1988
(H)
Camptomeris Syd. 1927 (H)
Campylospora Ranzoni 1953 (H)
Candelabrum Beverw. 1951 (H)
Candelosynnema K.D. Hyde & Seifert
1992 (H)