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steppenlebensräume europas – GefährdunG, erhaltunGsmassnahmen und schutz
435
Dry grasslands of Germany – call to support an initiative
for a consistent, plot-based classication
ute jandt, thomas becker & jürGen denGler
Zusammenfassung
Abstract
We present the plans for a comprehensive, national dry grassland database for Germany, in which
vegetation-plot records from different databases will be collected. These data will particularly be
used for a plot-based, consistent classication of dry grassland vegetation in Germany. The results of
the classication project will then be published in a special feature contribution to Applied Vegeta-
tion Science and in the series Synopsis der Panzengesellschaften Deutschlands. We warmly invite
persons who have dry grassland relevés to participate in this project by contributing data and/or
helping with data analysis.
Trockenrasen in Deutschland – Aufruf zur Mithilfe bei einer Initiative für eine einheitliche,
Aufnahme-basierte Klassikation
Es werden die Pläne für eine einheitliche, nationale Datenbank der Trockenrasen Deutschlands vor-
gestellt, in der die Vegetations-Daten aus verschiedenen Datenbanken zusammengeführt werden
sollen. Hiermit soll zunächst eine Aufnahme-basierte, einheitliche Klassikation der Trockenrasen-
Vegetation in Deutschland erstellt werden. Wir beabsichtigen, die Ergebnisse des Klassikations-
Projektes in der Zeitschrift Applied Vegetation Science sowie in der Reihe Synopsis der Pan-
zengesellschaften Deutschlands zu veröffentlichen. Eigentümer von Vegetationsaufnahmen von
Trockenrasen bitten wir hiermit herzlich um Mithilfe bei der Realisierung dieses Projektes.
Dry grasslands are in the focus of nature conservation throughout Central Europe, as they are extre-
mely diverse and at the same time highly endangered. However, the conservation and protection of
dry grassland habitats in Germany is largely based on traditional subjective classication schemes
developed decades ago (e.g. krAusch 1961, 1968, korneck 1974, 1975, oBerdorfer 1978, jeckel
1984). In general, these studies were based on limited datasets both in terms of geographic coverage
and numbers of relevés, and could not rely on the numerical methods available today. More recently,
there have been a few plot-based studies with transparent numerical methods, such as jAndt (1999)
for the Harz and Kyffhäuser regions and denGler (2004a, 2004b) for Mecklenburg-Vorpommern,
but none covered the whole of Germany. Accordingly, the available country-wide classications
(Pott 1995, schuBert et al. 2001, rennwAld 2002) are basically amalgamations of different, often
incompatible concepts. There is therefore a great need for a consistent syntaxonomic classication of
dry grasslands in Germany to meet the standards of neighbouring countries (e.g. mucinA et al. 1993,
schAminée et al. 1996, chytrý 2007, jAnišoVá 2007). There is also evidence that the discreteness of
some syntaxa is overrated while other units are largely underestimated or even overlooked (e.g. the
acidophilous semi-dry grasslands of the lower mountain ranges in southern Germany; see Becker
et al. 2012).
With the initiative of the German Arbeitsgruppe Trockenrasen presented in this article, we intend to
classify the syntaxa of dry grasslands based on a comprehensive vegetation database covering the
whole territory of Germany and all dry grassland types, characterise the derived syntaxa oristically
and ecologically, and analyse their geographical distribution (for photographic examples of syntaxa
see Figures 2–17). Further, we plan to compile conservation-relevant information for each distingu-
ished syntaxon, e.g. appropriate conservation measures and information on historical development.
For the future, we envision the establishment of an internet-platform, which will enable anyone
interested in dry grassland vegetation to share their data, analytical methods and results. The Ger-
man Dry Grassland Database will be part of the GVRD (German Vegetation Reference Database;
GIVD-ID EU-DE-014, jAndt&Bruelheide 2012), which is the vegetation data research platform
of the newly established German Centre for Integrative Biodiversity Research, iDiv (http://www.
idiv-biodiversity.de/). This database initiative of the German Arbeitsgruppe Trockenrasen within the
European Dry Grassland Group (EDGG; see VrAhnAkis et al. 2013) is meant to support nature con-
servation as well as research on biodiversity patterns and other ecological topics. Its success relies on
the willingness of persons with dry grassland vegetation data to contribute them to a joint database.
1
Introduction
436 jandt et al.: Dry grasslands of Germany – call to support an initiative
The common database format of the already existing database is TURBOVEG (hennekens&schA-
minée 2001), which enables easy data exchange via the comprehensive species reference list Ger-
manSL (jAnsen & denGler 2008). Presently, databases managed by the three initiators comprise
approx. 26,000 dry grassland relevés from Germany (Table 1), mainly from the vegetation classes
Festuco-Brometea, Koelerio-Corynephoretea (incl. Sedo-Scleranthetea), Violetea calaminariae,
Trifolio-Geranietea, and Elyno-Seslerietea. However, the spatial coverage is still incomplete, es-
pecially for the Northwest German lowlands (Northrhine-Westphalia, Lower Saxony and Schles-
wig-Holstein), Saxony and South Germany except the Swabian-Franconian Alb (Fig. 1). Therefore,
further efforts are needed to include data from other (public or private) databases and to digitize
relevés from undersampled regions.
2
Data management
and methods
Table 1:
Contributing databases with dry
grassland relevés from Germany.
Database identiers and names are
given according to the Global In-
dex of Vegetation-Plot Databases
(GIVD; http://www.givd.info/;
denGler et al. 2011, 2012).
Database identier Database name Number of German dry
in GIVD in GIVD grassland plots contained
EU-00-002 Database Dry Grasslands in the Nordic ca. 3,500
and Baltic Region
EU-00-007 VIOLETEA – heavy metal grasslands ca. 600
EU-DE-001 VegMV – the vegetation database of ca. 2,000
Mecklenburg-Vorpommern
EU-DE-013 VegetWeb – the national online-repository ca. 1,000
of vegetation plots from Germany
EU-DE-014 German Vegetation Reference Database (GVRD) ca. 17,000
EU-DE-034 Dry Grassland Central Germany Database ca. 3,000
Fig. 1:
Distribution of the approx. 18.000
dry grassland vegetation plots with
geographic coordinates in the con-
tributing databases. Occurrences
were pooled to central points of
the topographical map grid of
Germany.
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437
Fig. 4:
Cirsio-Brachypodion pinnati with Adonis vernalis,
Kyffhäuser, Thuringia (Photo: U. Jandt).
Fig. 5:
Festucion valesiacae, Mainz Basin, Rhineland-Palatinate
(Photo: J. Dengler, JD125289).
Fig. 2:
Bromion erecti, Swabian Alb, Baden-Württemberg
(Photo: J. Dengler, JD080712).
Fig. 3:
Xero-Bromion erecti, Unstrut Valley, Saxony-Anhalt
(Photo: T. Becker).
Fig. 6:
Armerion elongatae, Upper Rhine Valley, Hesse
(Photo: J. Dengler, JD103044).
Fig. 7:
De-alpine grasslands with Sesleria albicans, Franconian Alb,
Bavaria (Photo: J. Dengler, JD093884)
Fig. 8:
Koelerio-Phleion phleoidis. Rheinhesse, Rhineland-Palatinate
(Photo: J. Dengler, JD125060).
Fig. 9:
Festucion pallentis. Southern Harz, Saxony-Anhalt
(Photo: U. Jandt).
438 jandt et al.: Dry grasslands of Germany – call to support an initiative
Fig. 12:
Hyperico perforati-Scleranthion perennis, Lahn-Dill High-
lands, Hesse (Photo: J. Dengler, JD102744).
Fig. 13:
Corynephorion canescentis, Elbe Valley, Lower Saxony
(Photo: J. Dengler, JD080650).
Fig. 10:
Koelerion glaucae, Mainz Basin, Rhineland-Palatinate
(Photo: T. Becker).
Fig. 11:
Alysso alyssoidis-Sedion, Unstrut Valley, Saxony-Anhalt
(Photo: T. Becker).
Fig. 14:
Armerion halleri, Harz Mts., Lower Saxony
(Photo: T. Becker).
Fig. 15.:
Veronicion dillenii, Unstrut Valley, Thuringia
(Photo: T. Becker).
Fig. 16:
Sileno conicae-Cerastion semidecandri, Upper Rhine Valley,
Hesse (Photo: J. Dengler, JD102903).
Fig. 17:
Thero-Airion, Upper Rhine Valley, Hesse
(Photo: J. Dengler, JD103011).
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439
We would like to encourage colleagues who have dry grassland relevés not yet in our databases
(check with author search on the GVRD homepage, http://www.biologie.uni-halle.de/bot/vegetati-
on_db/ or ask authors), to enter contribute to the database project. Interested researchers may also
join the classication project. Numerical methods will be applied for data classication, like cluster
analysis (e.g. roleček et al. 2009), and delity analyses (e.g. chytrý et al. 2002, tichý&chytrý
2006), and denition of species groups with the Cocktail method (Bruelheide 1995). We intend
to publish the results in a special feature contribution in Applied Vegetation Science (denGler et
al. 2013) and in the German series Synopsis der Panzengesellschaften Deutschlands (http://www.
tuexenia.de/index.php?id=58). Vegetation-plot data gathered within the German Dry Grassland Da-
tabase will be made available for further research, environment protection and teaching purposes via
iDiv’s database portal (under construction).
We thank all the people who have recorded vegetation-plot data and let us include their records into
the databases. Thanks also to the many student helpers, who have devoted time and effort to the
digitization of vegetation relevés. Gunnar Seidler (Halle/Saale) designed the map of Germany with
dry grassland coordinates (Fig. 1). Parts of our work have been funded by the DFG Priority Program
1374 “Infrastructure-Biodiversity-Exploratories” (DFG BR 1698/11-1).
Acknowledgements
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Addresses of authors
Dr. Ute Jandt
Martin Luther University Halle Wittenberg
Institute of Biology / Geobotany and Botanical Garden
Am Kirchtor 1
06108 Halle
GERMANY
E-Mail: ute.jandt@botanik.uni-halle.de
German Centre for Integrative Biodiversity Research (iDiv)
University of Leipzig
Deutscher Platz 5e
04103 Leipzig
GERMANY
Dr. Thomas Becker
Regional and Environmental Sciences / Geobotany
University of Trier
Behringstr. 21
54296 Trier
GERMANY
E-Mail beckerth@uni-trier.de
PD Dr. Jürgen Dengler
Disturbance Ecology
University of Bayreuth
Universitätsstr. 30
95447 Bayreuth
GERMANY
E-Mail juergen.dengler@uni-bayreuth.de
German Centre for Integrative Biodiversity Research (iDiv)
University of Leipzig
Deutscher Platz 5e
04103 Leipzig
GERMANY