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Redescription of Lagria (Apteronympha) tenenbaumi Pic, 1929, with a checklist of the Western Palaearctic species of the genus Lagria F. (Coleoptera: Tenebrionidae: Lagriini)

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Lagria tenenbaumi Pic, 1929 is redescribed and transferred to the subgenus Apteronympha Seidlitz, 1898. Lagria hirta (Linnaeus, 1758) is recorded from Israel for the first time. Type species of Lachna Billberg, 1820 (= Lagria Fabricius, 1775) is designated. A checklist of the Western Palaearctic species of Lagria and a key to the species of the subgenus Apteronympha are given.
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Труды Русского энтомологического общества. С.-Петербург, 2006. Т. 77: 219–225.
Proceedings of the Russian Entomological Society. St. Petersburg, 2006. Vol. 77: 219–225.
Redescription of Lagria (Apteronympha) tenenbaumi Pic, 1929,
with a checklist of the Western Palaearctic species of the genus
Lagria F. (Coleoptera: Tenebrionidae: Lagriini)
O. Merkl
Переописание Lagria (Apteronympha) tenenbaumi Pic, 1929
и конспект западнопалеарктических видов рода
Lagria F. (Coleoptera: Tenebrionidae: Lagriini)*
О. Меркл
This paper is dedicated to Dr Gleb Sergeevich Medvedev,
leader of the Coleoptera Department and Head of the Labo-
ratory of Insect Systematics, Zoological Institute, Russian
Academy of Sciences (St. Petersburg), President of the Rus-
sian Entomological Society, an outstanding expert of Tene-
brionidae, on the occasion of his 75th birthday.
Hungarian Natural History Museum, Baross utca 13, H-1088 Budapest, Hungary. E-mail: merkl@zoo.zoo.nhmus.hu
Abstract. Lagria tenenbaumi Pic, 1929 is redescribed and transferred to the subgenus Apteronympha
Seidlitz, 1898. Lagria hirta (Linnaeus, 1758) is recorded from Israel for the first time. Type species of
Lachna Billberg, 1820 (= Lagria Fabricius, 1775) is designated. A checklist of the Western Palaearctic
species of Lagria and a key to the species of the subgenus Apteronympha are given.
Key words. Tenebrionidae, Lagriini, Lagria tenenbaumi, Apteronympha, Lachna, type species designa-
tion, checklist, Israel.
Резюме. Приведено переописание Lagria tenenbaumi Pic, 1929, этот вид переведен в подрод Apte-
ronympha Seidlitz, 1898. Lagria hirta (Linnaeus, 1758) впервые указан из Израиля. Обозначен
типовой вид рода Lachna Billberg, 1820 (= Lagria Fabricius, 1775). Даны аннотированный список за-
паднопалеарктических видов рода Lagria и таблица для определения видов подрода Apteronympha.
Ключевые слова. Tenebrionidae, Lagriini, Lagria tenenbaumi, Apteronympha, Lachna, обозначение
типового вида, список видов, Израиль.
15th contribution to the knowledge of Lagriini. 14th contribution: Acta Zoologica Academiae Scientiarum
Hungaricae 50: 283–305.
220
Introduction
Lagria Fabricius, 1775 is a large collective genus of the primitive tenebrionoid subtribe Lagriina
with more than 150 species in the temperate, tropical and subtropical areas of Europe, Africa (including
Madagascar), Asia, New Guinea and northern parts of Australia (Northern Territory and Queensland). The
genus contains species with very different general appearance and without any synapomorphy which
would separate them from members of other genera. Virtually the absence of salient modifications in
the antennae, pronotum, elytra and legs seems to be the only reason to put them together in this genus.
Seidlitz (1898) erected the subgenus Apteronympha for two Western Mediterranean species on
the basis of flightlessness of females, although females with reduced hind wings occur in other (non-
European) species of Lagriina (even in Lagria itself) as well. Males resemble Lagria hirta (Linnaeus,
1758), the type species of the genus, in all fundamental features. There is another feature that separates
members of Apteronympha from other Western Palaearctic species of Lagria. This is the rugose (coarse
and dense) punctation of pronotum; the other species have finely and sparsely punctate (sometimes nearly
impunctate) pronotum. Although female flightlessness and difference in pronotal punctation are rather
weak traits to support subgeneric distinction, the subgenus is retained for now as a separate group, since
a thorough revision of the genus is a task of the future. Borchmann (1936) transferred Lagria pygmaea
Brancsik, 1892 to Apteronympha, but this Madagascan species is unknown to the author, and it is quite
possible that it is not closely related to the species from the Western Palaearctic.
Borchmann apparently overlooked Lagria tenenbaumi Pic, 1929, a species described from Pales-
tine, because it is absent from his monograph of World “Lagriidae” (Borchmann, 1936). Only males were
available when Pic published the very short description typical of him, so subgeneric assignment was
impossible. A redescription of the species, including the description of the female, is given below. On the
basis of the rugosely punctate pronotum and wingless female it is unequivocal that Lagria tenenbaumi
should be assigned to the subgenus Apteronympha.
Abbreviations of depositories: BMNH – Natural History Museum (formerly British Museum, Natu-
ral History), London, United Kingdom; HNHM – Hungarian Natural History Museum, Budapest, Hun-
gary; NMPR – National Museum, Prague, Czech Republic; TAUI – Tel Aviv University, Tel Aviv, Israel.
Transliteration of locality names is according to the Touring Map of Israel (1 : 250 000) + List of
settlements, antiquity sites and road distances, (2001) Survey of Israel, Cartography Department, Tel Aviv.
Lagria tenenbaumi Pic, 1929 (Figs 1–4)
Lagria tenenbaumi Pic, 1929: 2; Bodenheimer, 1937: 135.
Original description: “Lagria Tenenbaumi nov. (? var. de tristis Bon). Sat elongatus, parum nitidus, sat longe gri-
seo pubescens, fortiter et pro parte dense ruguloso-punctatus, niger, elytris postice diverse pallidioribus. Long. 6,7 mill.
Palestine: Gebatha (Tenenbaum, in coll. Tenenbaum et Pic).
En l’absence du sexe , je ne puis me prononcer catégoriquement au sujet de cette nouveauté, mais j’incline à y voir
simplement une variété, à coloration générale foncée de L. tristis Bon., les antennes ayant le dernier article environ de la
longueur des quatre précédents réunis, le prothorax aussi large que long, densément ponctué.”
Redescription. M a l e (Fig. 1). Body winged, elongate, slender, almost parallel-sided, dark brownish black (teneral
specimens paler), weakly shining; vestiture consisting of short, semierect greyish hairs leaving dorsal surface visible. Body
length 4.8–6.1 mm (n = 42).
Head short, coarsely and densely punctate, interspaces much narrower than a puncture diameter; frons shallowly im-
pressed; frontoclypeal suture deep; clypeus with anterior margin slightly concave; temples short, converging posteriorly; eyes
large, bulging, interocular distance shorter than eye diameter. Pubescence sparse, with a few long hairs on temples. Maxilla
with apical palpomere narrowly securiform. Labium with apical palpomere spindle-shaped, apex not emarginate. Antennae
when posteriorly extended surpassing posterior margin of pronotum by last two antennomeres; antennomere 1 subglobose, 2
subequal in length and width, 3 twice as long as 2, 4 slightly shorter than 3, 5 and 6 slightly shorter than 4, 7–9 as long as 5
and 6 but slightly broader, 10 subequal in length and width, 11 as long as four preceding antennomeres combined.
Prothorax nearly as long as wide, constricted just posterior to middle, widest just anterior to middle; pronotum with
dorsal surface slightly convex, with barely visible mesal impression immediately before posterior margin; punctation coarse
and dense, interspaces narrower than a puncture diameter; pubescence partly proclinate. Prothoracic hypomeron and proster-
num with coarse and sparse punctures and fine transverse wrinkles. Scutellum small, rounded triangular, coarsely punctate.
Elytra about 3.5 times as long as wide and 4 times as long as pronotum; sides parallel, barely diverging posteriorly;
punctation relatively coarse and dense, without trace of longitudinal rows; punctures separated by interspaces of 0.5 to
221
1 puncture diameter; interspaces without microreticulation. Pubescence reclinate. Mesoventrite finely punctate; mesoventral
process about 1/5 times as wide as mesocoxa; mesepisternum, metepimeron, and metepisternum coarsely and densely punc-
tate, metaventrite very finely and sparsely punctate.
Legs long and slender; femora subequal in length to tibiae, and practically not clavate; length of hind femur about
twice width of elytron at level of hind coxae; tibiae nearly straight, subequal in width; femora and tibiae without denticles,
angulations or any other modifications; tarsi definitely shorter than tibiae.
Abdominal ventrites finely and sparsely punctate, punctation becoming denser toward lateral portion; last ventrite
rounded posteriorly. Aedeagus with basale about 8 times as long as apicale, latter narrowly triangular (Figs 3, 4).
F e m a l e (Fig. 2). Body wingless, shorter and broader. Body length 6.8–7.0 mm (n = 2). Eyes small, transverse; in-
terocular distance 2.5 times eye diameter; antennae short, when posteriorly extended barely surpassing posterior margin
of pronotum; antennomere 11 twice as long as 10; prothorax slightly transverse, disc without midlongitudinal impression;
elytra distinctly widened posteriorly, with traces of longitudinal elevations in posterior half.
Material. I s r a e l . ‘Adullam, 17 V 2002, V. Mandelik & V. Chikatunov (1 , 1 ; TAUI); Amnun, light trap, 15
III 2003, V. Kravchenko & V. Chikatunov (1 ; TAUI); Amnun, light trap, 15 V 2003, V. Kravchenko & V. Chikatunov
(3 ; TAUI); Been Mashichim [= Be`er Mash`abim], 7 XII 1996, R. Hoffman (1 ; TAUI); Esdraelon, Mount Gilboa [=
Yizre’`el Valley, Hare Gilboa’], unknown date and collector [probably Q. Argaman] (1 ; TAUI); Gan Shomeron, 27 IX
1957, Hebr[ew] Univ[ersity] (1 ; TAUI); Golan, Senir, 23 IX 1997, V. Chikatunov (2 ; TAUI); Haifa, Checkpost, 20 IV
1998, V. Chikatunov & T. Pavlíček (1 ; TAUI); Hammat Gader, 15 VI 2002, light trap, V. Kravchenko (1 ; TAUI); Kok-
hav HaYarden, light trap, 15 VI 2002, V. Kravchenko (1 ; TAUI); Lod, 8 X 1997, R. Hoffman (1 ; TAUI); Majdal Shams,
2 X 2002, V. Kravchenko (1 ; TAUI); Majdal Shams, 20 V 2001, V. Kravchenko (2 , HNHM; 3 , TAUI); Merom
Figs 1, 2. Lagria (Apteronympha) tenenbaumi Pic, 1926, male and female. Scale = 2 mm.
222
Figs 3, 4. Lagria (Apteronympha) tenen-
baumi Pic, 1926, aedeagus, ventral and lateral
views. Scale = 0.5 mm.
Golan, 12 VI 2000, V. Chikatunov (1 ; TAUI); Mi-
khmoret, unknown date and collector [probably Q. Ar-
gaman] (2 ; TAUI); Nahal Dishon, Rt. 886, light trap,
22 V 1998, A. Freidberg (1 , BMNH; 2 , TAUI);
Nahal Oren, Mt. Carmel, SFS [= south-facing slope],
light trap, 30 X 1997, T. Pavlíček (2 , NMPR; 1 ,
HNHM); Nahal Oren, S1, 5 X 1997, V. Chikatunov &
T. Pavlíček (1 ; TAUI); Nahal Oren, SFS1 [= south-
facing slope], 30 X 1997, V. Chikatunov & T. Pavlíček
(1 ; TAUI); Nahal Oren, SFS2 [= south-facing slope],
light, 21 V 1998, V. Chikatunov & T. Pavlíček (1 ;
TAUI); Nahal Sansan, 17 V 2002, V. Mandelik &
V. Chikatunov (1 ; TAUI); Ramallah, 600 m, 10 IX
1958, J. Klapperich (1 ; HNHM); Ramot Avishov, 17
V 2002, V. Mandelik & V. Chikatunov (1 ; HNHM);
Samaria, Wadi el Faria [= Nahal Tirza], unknown date
and collector [probably Q. Argaman] (2 ; TAUI);
Segula, Qiryat Gat, 5 V 1996, V. Chikatunov (1 ;
TAUI); Tel Dan, 15 VI 2002, light trap, V. Kravchenko
(1 ; TAUI); Tel Dan, 5 X 2002, light trap, V.
Kravchenko & V. Chikatunov (1 , BMNH; 1 ,
TAUI); Zekharya, 17 V 2002, V. Mandelik &
V. Chikatunov (1 ; TAUI); Zur Hadassa, light trap, 29
IV 2001, V. Kravchenko (1 ; TAUI).
Distribution. All known localities of this
species are in Israel.
Diagnostic characters of L. tenenbaumi
see in the key below (with data on the distribu-
tion of the species in the Iberian Peninsula after
Español, 1964 and Viñolas, Cartagena, 2005).
Key to species of the subgenus Lagria (Apteronympha)
1a. Body unicoloured dark brownish black, at most tibiae slightly paler. Female without midlongitudinal
impression on pronotum. – Israel .............................................................. Lagria tenenbaumi Pic, 1929
1b. Body not unicoloured: at least elytra light reddish brown. Female with more or less distinct midlongi-
tudinal impression on pronotum. – Western Mediterranean .................................................................. 2
2a. Male with interocular distance more than half of eye diameter. Legs, antennae, head and pronotum
entirely black. Elytral pubescence shorter. Female with shallow midlongitudinal impression restricted
to posterior half of pronotum. – Iberian Peninsula ...................................... Lagria rubida Graells, 1855
2b. Male with interocular distance less than half of eye diameter. Legs and antennae (frequently also
pronotum and head) at least partly reddish brown. Elytral pubescence somewhat longer. Female with
deep midlongitudinal impression almost reaching anterior margin of pronotum. – Iberian Peninsula,
southern France, northern Italy, southern Switzerland; ?Morocco .. Lagria rugosula Rosenhauer, 1856
Lagria hirta (Linnaeus, 1758)
This is the second species of Lagria occurring in Israel. It has winged females, larger and bicol-
oured body (the elytra are reddish brown, the rest is black) and finely punctate pronotum.
Material. Jerusalem, 8 IV 1966, unknown collector (1 ; TAUI); N kziv [= Nahal Keziv], 4 VI 1999, M. Finkel (1
; TAUI); Nahal Keziv, 28 IX 1999, M. Finkel & V. Kravchenko (1 ; TAUI); Mt. Meiron [= Har Meron], 18 IX 1976,
A. Freidberg (1 ; TAUI), and 12 VII 2002, light trap, V. Kravchenko (1 ; TAUI).
3 4
223
Checklist of the Western Palaearctic species of the genus Lagria
Much of the synonymy is based on Borchmann (1936). A few names were synonymised by Merkl
(2004). L. brevipilis Desbr. and L. pici Normand are unknown to the author; it is not impossible that these
names are synonyms of other, well known species of Lagria occurring in North Africa.
Subgenus Apteronympha Seidlitz, 1898: 336
Type species Lagria rubida Graells, 1855, designated by Merkl, 2004: 284.
Lagria rubida Graells, 1855: 74. Spain
= Lagria parvula Perris, 1864: 291
= Lagria elliptica Fairmaire, 1880: 242
Lagria rugosula Rosenhauer, 1856: 223. Europe: France, Italy, Portugal, Spain, Switzerland;
North Africa: Morocco
= Lagria tristis Bonelli, 1812: 172, not Fabricius, 1801: 70
= Lagria glabrata Olivier, 1792: 446, not Fabricius, 1775: 125
= Lagria pubescens Latreille, 1804: 351, not Linnaeus, 1767: 603
= Lagria melichari Reitter, 1890: 173
Lagria tenenbaumi Pic, 1929: 1. Asia: Israel
Subgenus Lagria Fabricius, 1775: 124
Type species Chrysomela hirta Linnaeus, 1758, designated by Borchmann, 1916: 59.
= Lachna Billberg, 1820: 35. Type species Chrysomela hirta Linnaeus, 1758, here designated.
Lagria atripes Mulsant et Guillebeau, 1855: 74. Europe: north to Belgium, Germany and Poland;
Asia: Iran, Turkmenistan, Turkey
= Lagria atripes var. puncticollis Seidlitz, 1898: 337
= Lagria atripes var. tenuicollis Seidlitz, 1898: 337, not Fairmaire, 1895: 450
Lagria brevipilis Desbrochers des Loges, 1881: 143. North Africa: Algeria
Lagria grenieri Brisout de Barneville, 1867: 181. Europe: France, Portugal, Spain
Lagria hirta (Linnaeus, 1758: 377) (Chrysomela). Europe: north to Fennoscandia; North Africa:
Algeria, Morocco; Asia: Russia (Western and Eastern Siberia), Israel, Cyprus, Turkey, Iran, Iraq, Kazakh-
stan, Turkmenistan, Uzbekistan, Tajikistan
= Cantharis spadicea Scopoli, 1763: 43
= Chrysomela pubescens Linnaeus, 1767: 603
= Lagria glabrata Fabricius, 1775: 125
= Tenebrio villosus De Geer, 1775: 44
= Cantharis flava Geoffroy, 1785: 155
= Lagria lurida Krynicki, 1832: 137
= Lagria nudipennis Mulsant, 1856: 20
= Lagria depilis Mulsant, 1856: 21
= Lagria caucasica Motschulsky, 1860: 144
= Lagria fuscata Motschulsky, 1860: 144
= Lagria pontica Motschulsky, 1860: 144
= Lagria hirta var. limbata Desbrochers des Loges, 1881: 142
= Lagria seminuda Reitter, 1889: 259
Lagria lata Fabricius, 1801: 70. Europe: Spain; North Africa: Algeria, Morocco
= Lagria rugata Desbrochers des Loges, 1881: 145
Lagria pici Normand, 1936: 161. North Africa: Tunisia
224
Acknowledgements
Thanks are due to Dr Vladimir Chikatunov and Mr Ariel-Leib-Leonid Friedman (TAUI), Dr Tomáš
Pavlíček (Institute of Evolution, University of Haifa, Israel) and Dr Vladimir Švihla (NMPR) for sending
material investigated. Mr Friedman helped to rectify transliteration of the locality names. Dr Anthony
Davies (Canadian National Collection of Insects, Arachnids and Nematodes, Agriculture and Agri-Food
Canada, Ottawa, Canada) kindly provided information about Billberg (1820). The drawings were made by
Mr János Pál (HNHM).
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Many studies have found that the correlation between species richness (SR) and morphological diversity (MD) is positive, but the correlation degree of these parameters is not always consistent due to differences in categories and various ecological factors in the living environment. Based on this, related studies have revealed the good performance of using higher taxa in biodiversity research, not only by shifting the testing group scale from local communities to worldwide datasets but also by adding different taxonomic levels, such as the genus level. However, it remains unclear whether this positive correlation can also be applied to other categories or groups. Here, we evaluated the applicability of higher taxa in the biodiversity study of darkling beetles by using 3407 species (9 subfamilies, 89 tribes, and 678 genera), based on the correlation between taxa richness and morphological diversity in the tribe/genus/species. In addition, the continuous features prevalent in the tenebrionids, pronotum and elytron, were selected, and the morphological diversity of various groups was obtained by the geometric morphometric approach to quantify the morphologic information of features. This study found that genus/species richness in subfamilies Pimelinae and Stenochiinae was positively correlated with the change trend of MD, and the correlation between the MD of elytron and taxa richness gradually decreased from the tribe-level to the genus-level to the species-level test. The results confirm the stable morphology and simple function of the elytron and the applicability of tribe level in biodiversity research.
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A review of genus-group names for darkling beetles in the family Tenebrionidae (Insecta: Coleoptera) is presented. A catalogue of 4122 nomenclaturally available genus-group names, representing 2307 valid genera (33 of which are extinct) and 761 valid subgenera, is given. For each name the author, date, page number, gender, type species, type fixation, current status, and first synonymy (when the name is a synonym) are provided. Genus-group names in this family are also recorded in a classification framework, along with data on the distribution of valid genera and subgenera within major biogeographical realms. A list of 535 unavailable genus-group names (e.g., incorrect subsequent spellings) is included. Notes on the date of publication of references cited herein are given, when known. The following genera and subgenera are made available for the first time: Anemiadena Bouchard & Bousquet, subgen. nov. (in Cheirodes Gené, 1839), Armigena Bouchard & Bousquet, subgen. nov. (in Nesogena Mäklin, 1863), Debeauxiella Bouchard & Bousquet, subgen. nov. (in Hyperops Eschscholtz, 1831), Hyperopsis Bouchard & Bousquet, subgen. nov. (in Hyperops Eschscholtz, 1831), Linio Bouchard & Bousquet, subgen. nov. (in Nilio Latreille, 1802), Matthewsotys Bouchard & Bousquet, gen. nov. , Neosolenopistoma Bouchard & Bousquet, subgen. nov. (in Eurynotus W. Kirby, 1819), Paragena Bouchard & Bousquet, subgen. nov. (in Nesogena Mäklin, 1863), Paulianaria Bouchard & Bousquet, gen. nov. , Phyllechus Bouchard & Bousquet, gen. nov. , Prorhytinota Bouchard & Bousquet, subgen. nov. (in Rhytinota Eschscholtz, 1831), Pseudorozonia Bouchard & Bousquet, subgen. nov. (in Rozonia Fairmaire, 1888), Pseudothinobatis Bouchard & Bousquet, gen. nov. , Rhytinopsis Bouchard & Bousquet, subgen. nov. (in Thalpophilodes Strand, 1942), Rhytistena Bouchard & Bousquet, subgen. nov. (in Rhytinota Eschscholtz, 1831), Spinosdara Bouchard & Bousquet, subgen. nov. (in Osdara Walker, 1858), Spongesmia Bouchard & Bousquet, subgen. nov. (in Adesmia Fischer, 1822), and Zambesmia Bouchard & Bousquet, subgen. nov. (in Adesmia Fischer, 1822). The names Adeps Gistel, 1857 and Adepsion Strand, 1917 syn. nov. [= Tetraphyllus Laporte & Brullé, 1831], Asyrmatus Canzoneri, 1959 syn. nov. [= Pystelops Gozis, 1910], Euzadenos Koch, 1956 syn. nov. [= Selenepistoma Dejean, 1834], Gondwanodilamus Kaszab, 1969 syn. nov. [= Conibius J.L. LeConte, 1851], Gyrinodes Fauvel, 1897 syn. nov. [= Nesotes Allard, 1876], Helopondrus Reitter, 1922 syn. nov. [= Horistelops Gozis, 1910], Hybonotus Dejean, 1834 syn. nov. [= Damatris Laporte, 1840], Iphthimera Reitter, 1916 syn. nov. [= Metriopus Solier, 1835], Lagriomima Pic, 1950 syn. nov. [= Neogria Borchmann, 1911], Orphelops Gozis, 1910 syn. nov. [= Nalassus Mulsant, 1854], Phymatium Billberg, 1820 syn. nov. [= Cryptochile Latreille, 1828], Prosoblapsia Skopin & Kaszab, 1978 syn. nov. [= Genoblaps Bauer, 1921], and Pseudopimelia Gebler, 1859 syn. nov. [= Lasiostola Dejean, 1834] are established as new synonyms (valid names in square brackets). Anachayus Bouchard & Bousquet, nom. nov. is proposed as a replacement name for Chatanayus Ardoin, 1957, Genateropa Bouchard & Bousquet, nom. nov. as a replacement name for Apterogena Ardoin, 1962, Hemipristula Bouchard & Bousquet, nom. nov. as a replacement name for Hemipristis Kolbe, 1903, Kochotella Bouchard & Bousquet, nom. nov. as a replacement name for Millotella Koch, 1962, Medvedevoblaps Bouchard & Bousquet, nom. nov. as a replacement name for Protoblaps G.S. Medvedev, 1998, and Subpterocoma Bouchard & Bousquet, nom. nov. is proposed as a replacement name for Pseudopimelia Motschulsky, 1860. Neoeutrapela Bousquet & Bouchard, 2013 is downgraded to a subgenus ( stat. nov. ) of Impressosora Pic, 1952. Anchomma J.L. LeConte, 1858 is placed in Stenosini: Dichillina (previously in Pimeliinae: Anepsiini); Entypodera Gerstaecker, 1871, Impressosora Pic, 1952 and Xanthalia Fairmaire, 1894 are placed in Lagriinae: Lagriini: Statirina (previously in Lagriinae: Lagriini: Lagriina); Loxostethus Triplehorn, 1962 is placed in Diaperinae: Diaperini: Diaperina (previously in Diaperinae: Diaperini: Adelinina); Periphanodes Gebien, 1943 is placed in Stenochiinae: Cnodalonini (previously in Tenebrioninae: Helopini); Zadenos Laporte, 1840 is downgraded to a subgenus ( stat. nov. ) of the older name Selenepistoma Dejean, 1834. The type species [placed in square brackets] of the following available genus-group names are designated for the first time: Allostrongylium Kolbe, 1896 [ Allostrongylium silvestre Kolbe, 1896], Auristira Borchmann, 1916 [ Auristira octocostata Borchmann, 1916], Blapidocampsia Pic, 1919 [ Campsia pallidipes Pic, 1918], Cerostena Solier, 1836 [ Cerostena deplanata Solier, 1836], Coracostira Fairmaire, 1899 [ Coracostira armipes Fairmaire, 1899], Dischidus Kolbe, 1886 [ Helops sinuatus Fabricius, 1801], Eccoptostoma Gebien, 1913 [ Taraxides ruficrus Fairmaire, 1894], Ellaemus Pascoe, 1866 [ Emcephalus submaculatus Brême, 1842], Epeurycaulus Kolbe, 1902 [ Epeurycaulus aldabricus Kolbe, 1902], Euschatia Solier, 1851 [ Euschatia proxima Solier, 1851], Heliocaes Bedel, 1906 [ Blaps emarginata Fabricius, 1792], Hemipristis Kolbe, 1903 [ Hemipristis ukamia Kolbe, 1903], Iphthimera Reitter, 1916 [ Stenocara ruficornis Solier, 1835], Isopedus Stein, 1877 [ Helops tenebrioides Germar, 1813], Malacova Fairmaire, 1898 [ Malacova bicolor Fairmaire, 1898], Modicodisema Pic, 1917 [ Disema subopaca Pic, 1912], Peltadesmia Kuntzen, 1916 [ Metriopus platynotus Gerstaecker, 1854], Phymatium Billberg, 1820 [ Pimelia maculata Fabricius, 1781], Podoces Péringuey, 1886 [ Podoces granosula Péringuey, 1886], Pseuduroplatopsis Pic, 1913 [ Borchmannia javana Pic, 1913], Pteraulus Solier, 1848 [ Pteraulus sulcatipennis Solier, 1848], Sciaca Solier, 1835 [ Hylithus disctinctus Solier, 1835], Sterces Champion, 1891 [ Sterces violaceipennis Champion, 1891] and Teremenes Carter, 1914 [ Tenebrio longipennis Hope, 1843]. Evidence suggests that some type species were misidentified. In these instances, information on the misidentification is provided and, in the following cases, the taxonomic species actually involved is fixed as the type species [placed in square brackets] following requirements in Article 70.3 of the International Code of Zoological Nomenclature: Accanthopus Dejean, 1821 [ Tenebrio velikensis Piller & Mitterpacher, 1783], Becvaramarygmus Masumoto, 1999 [ Dietysus nodicornis Gravely, 1915], Heterophaga Dejean, 1834 [ Opatrum laevigatum Fabricius, 1781], Laena Dejean, 1821, [ Scaurus viennensis Sturm, 1807], Margus Dejean, 1834 [ Colydium castaneum Herbst, 1797], Pachycera Eschscholtz, 1831 [ Tenebrio buprestoides Fabricius, 1781], Saragus Erichson, 1842 [ Celibe costata Solier, 1848], Stene Stephens, 1829 [ Colydium castaneum Herbst, 1797], Stenosis Herbst, 1799 [ Tagenia intermedia Solier, 1838] and Tentyriopsis Gebien, 1928 [ Tentyriopsis pertyi Gebien, 1940]. The following First Reviser actions are proposed to fix the precedence of names or nomenclatural acts (rejected name or act in square brackets): Stenosis ciliaris Gebien, 1920 as the type species for Afronosis G.S. Medvedev, 1995 [ Stenosis leontjevi G.S. Medvedev, 1995], Alienoplonyx Bremer, 2019 [ Alienolonyx ], Amblypteraca Mas-Peinado, Buckley, Ruiz & García-París, 2018 [ Amplypteraca ], Caenocrypticoides Kaszab, 1969 [ Caenocripticoides ], Deriles Motschulsky, 1872 [ Derilis ], Eccoptostira Borchmann, 1936 [ Ecoptostira ], † Eodromus Haupt, 1950 [† Edromus ], Eutelus Solier, 1843 [ Lutelus ], Euthriptera Reitter, 1893 [ Enthriptera ], Meglyphus Motschulsky, 1872 [ Megliphus ], Microtelopsis Koch, 1940 [ Extetranosis Koch, 1940, Hypermicrotelopsis Koch, 1940], Neandrosus Pic, 1921 [ Neoandrosus ], Nodosogylium Pic, 1951 [ Nodosogilium ], Notiolesthus Motschulsky, 1872 [ Notiolosthus ], Pseudeucyrtus Pic, 1916 [ Pseudocyrtus ], Pseudotrichoplatyscelis Kaszab, 1960 [ Pseudotrichoplatynoscelis and Pseudotrichoplatycelis ], Rhydimorpha Koch, 1943 [ Rhytimorpha ], Rhophobas Motschulsky, 1872 [ Rophobas ], Rhyssochiton Gray, 1831 [ Ryssocheton and Ryssochiton ], Sphaerotidius Kaszab, 1941 [ Spaerotidius ], Stira Agassiz, 1846 (Mollusca) [ Stira Agassiz, 1846 (Coleoptera)], Sulpiusoma Ferrer, 2006 [ Sulpiosoma ] and Taenobates Motschulsky, 1872 [ Taeniobates ]. Supporting evidence is provided for the conservation of usage of Cyphaleus Westwood, 1841 nomen protectum over Chrysobalus Boisduval, 1835 nomen oblitum.
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On the occurence of Lagria rugosula Rosenhauer, 1856 in Switzerland (Coleoptera: Tenebrionidae, Lagriinae). An overview on the occurrence of Lagria rugosula Rosenhauer, 1856 in Switzerland is given. The 76 investigated specimens are mainly from localities of the cantons Valais and Ticino, but also from Vaud and Grisons. Most records are from light traps.
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In this 7th report of the Project Entomofauna of Turkey, totally 1720 species and 490 genera recorded in Turkey are listed, with their synonyms. Provincial codes, if known, are added to the species. An attempt on evaluating of some dipterists are made for the first time. A selected bibliography are also given.
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In this 9th report of the Project Entomofauna of Turkey, development of the Entomofauna concept, research conditions and similar problems are discussed to some degree. Number of the pterygot species of Turkey based upon the info-system of the Cesa are explained. According to this, 41000 taxa (family-, genus-, and species-groups) are included to the Infosystem of the Cesa (with the exception of the Lepidoptera). Total number of the pterygot species of Turkey based upon the info-system of the Cesa increased to 14244. Concerning the present information of the database of the Cesa on the Entomofauna of Turkey, totally 13146 taxa (except Lepidoptera), including (789 family-, 2631 genus-, and 9056 species-group taxa) are evaluated. Among them, 3185 coleopteran species of Turkey belong to 739 genera of the 249 family group taxa are mentioned here. In the temporary lists submitted below, all the scientific names of the species and the genera are arranged alphabetically with their families. Besides the scientific names of the valid family group taxa (superfamilies, families, subfamilies, tribes, and subtribes) of the Coleoptera fauna of Turkey are listed. Chronologically arranged synonymous names are added to each taxa, if available. Distributional information are given as codes of the provinces of Turkey. References are partly published in the previous issue of this journal. Totally 103 coleopteran images of 17 families are used here for the first time. Illustrations are based upon field or laboratory studies of the authors. A selected bibliography is also given.
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New and interesting records of Lagriinae (Coleoptera: Tenebrionidae) for the province of Huelva, Andalusia, Southwest of Spain La familia Tenebrionidae esta representada en la Península Ibérica por siete subfamilias, entre las cuales, Lagriinae esta compuesta por tres tribus: Lagriini, Cossyphini y Belopini, en las que se integran tres géneros y seis subgénero. Tras la aparición de la monografía de VIÑOLAS & CARTA-GENA (2005), se aprecia una ausencia de datos para la provincia andaluza más occidental, respecto a otras provincias mucho mejor prospectadas. En esta nota se pretende ampliar el registro corológico onubense y andaluz, aportando los datos procedentes de la colección del autor. El material que se relaciona ha sido colectado por socios de la aso-ciación entomológica Insecol de Huelva y por el autor, en la provincia de Huelva en los últimos veintisiete años, utilizando los métodos habituales de captura según los distintos géneros de esta subfamilia de coleópteros: mangueo en flores, trampas de luz, inspección de materiales en el suelo y trampas de caída (pitfall). En un caso se proporciona una localidad de la vecina provincia de Cádiz. Todo el material está depositado en la colección de referencia del autor (JJLP) y todas las medidas corresponden a la longi-tud total del cuerpo. En esta nota se presentan cinco especies de Lagriinae, de las cuales, todas son novedosas para la provincia de Huelva, según la bibliografía consultada, reforzando y ampliando con estas diez citas (nueve para Huelva y una para Cádiz), la corología de los mismos, destacando la presencia de Lagria lata (Fabricius, 1801), que amplía notablemente su distribución. Se señalan además algunas de las diferencias encontradas entre los ejemplares del género Lagria de la península Ibérica con los existente en la región báltica. Para la identificación del material recolectado se han utilizado los siguientes trabajos: BOUCHARD et al.
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New data about distribution of three species of the genus Lagria Fabricius, 1755 (L. hirta (Linnaeus, 1758), L. atripes Mulsant et Guillebeau, 1855 and L. laticollis Motschulsky, 1860) in the European part of Russia (Belgorod and Lipetsk regions) and adjacent territories of Ukraine (Lugansk region) are given. A key to the three species is given. Images of L. laticollis are published for the first time.
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Un index alphabétique des noms d'espèces est proposé pour le catalogue systématique et atlas des Coléoptères Tenebrionidae de France publié par l'un des auteurs il y a quatre ans. Quelques corrections sont apportées. Abstract-A species-names alphabetical index is proposed for the systematic catalogue and atlas of French Coleoptera Tenebrionidae published by one of the authors four years ago. Some corrections are effected.
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Type species is designated for Apteronympha SEIDLITZ, 1898. The following genus-group synonyms are proposed: Diversogria PIC, 1954, Heterogria FAIRMAIRE, 1896, Lagriocera FAIRMAIRE, 1896, Lagriodes BORCHMANN, 1930, Pachylagria BORCHMANN, 1912 and Wallardilagria PIC, 1910 are junior subjective synonyms of Xanthalia FAIRMAIRE, 1894. Trachelolagria PIC, 1941 is transferred to the family Cleridae. New species-group synonyms are pro-posed in Anisostira (11), Cerogria (2), Chlorophila (6), Lagria (9), and Xanthalia (3). New combinations are proposed in Arthromacra (1), Casnonidea (1), Donaciolagria (2), and Xanthalia (61). Xanthalia borchmanni Merkl, 2004 is proposed as a replacement name for Xanthalia pilosa (BORCHMANN, 1943), secondary homonym of Xanthalia pilosa (BORCHMANN, 1930). Lectotypes are designated for Chlorophila portschinskii (SEMENOV, 1891), Lagria caucasica MOTSCHULSKY, 1860, Lagria fuscata MOTSCHULSKY, 1860, Lagria laticollis MOTSCHULSKY, 1860, Lagria subtilipunctata SEIDLITZ, 1898 and Clerus villosus THUNBERG, 1821 (= Lopholagria villosa). New country and Chinese province records are given for five species. Arthromacra chifengi sp. n. is described from Taiwan.
Contribution au Catalogue des Coléoptères de la Tunisie (9e fascicule)
N o r m a n d H 1936. Contribution au Catalogue des Coléoptères de la Tunisie (9e fascicule). Bull. Soc. Hist. Nat. Afr. Nord. 27(5): 144–164.
Entomologia Carniolica exhibens Insecta Carnioliae indigena et distributa in ordines, genera , species, varietates methodo Linnaeana. I. Vindobonae: N. Trattner
  • S C O P O L I
S c o p o l i J. A. 1763. Entomologia Carniolica exhibens Insecta Carnioliae indigena et distributa in ordines, genera, species, varietates methodo Linnaeana. I. Vindobonae: N. Trattner. 420 pp.
Description de quelques espèces nouvelles de Coléoptères et notes diverses
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P e r r i s É. 1864. Description de quelques espèces nouvelles de Coléoptères et notes diverses. Ann. Soc. Entomol. France. Quatrième série. 4: 275–303.