Redescription of Lagria (Apteronympha) tenenbaumi Pic, 1929, with a checklist of the Western Palaearctic species of the genus Lagria F. (Coleoptera: Tenebrionidae: Lagriini)

Abstract

Lagria tenenbaumi Pic, 1929 is redescribed and transferred to the subgenus Apteronympha Seidlitz, 1898. Lagria hirta (Linnaeus, 1758) is recorded from Israel for the first time. Type species of Lachna Billberg, 1820 (= Lagria Fabricius, 1775) is designated. A checklist of the Western Palaearctic species of Lagria and a key to the species of the subgenus Apteronympha are given.

Full text

219
Труды Русского энтомологического общества. С.-Петербург, 2006. Т. 77: 219–225.
Proceedings of the Russian Entomological Society. St. Petersburg, 2006. Vol. 77: 219–225.
Redescription of Lagria (Apteronympha) tenenbaumi Pic, 1929,
with a checklist of the Western Palaearctic species of the genus
Lagria F. (Coleoptera: Tenebrionidae: Lagriini)∗
O. Merkl
Переописание Lagria (Apteronympha) tenenbaumi Pic, 1929
и конспект западнопалеарктических видов рода
Lagria F. (Coleoptera: Tenebrionidae: Lagriini)*
О. Меркл
This paper is dedicated to Dr Gleb Sergeevich Medvedev,
leader of the Coleoptera Department and Head of the Labo-
ratory of Insect Systematics, Zoological Institute, Russian
Academy of Sciences (St. Petersburg), President of the Rus-
sian Entomological Society, an outstanding expert of Tene-
brionidae, on the occasion of his 75th birthday.
Hungarian Natural History Museum, Baross utca 13, H-1088 Budapest, Hungary. E-mail: merkl@zoo.zoo.nhmus.hu
Abstract. Lagria tenenbaumi Pic, 1929 is redescribed and transferred to the subgenus Apteronympha
Seidlitz, 1898. Lagria hirta (Linnaeus, 1758) is recorded from Israel for the first time. Type species of
Lachna Billberg, 1820 (= Lagria Fabricius, 1775) is designated. A checklist of the Western Palaearctic
species of Lagria and a key to the species of the subgenus Apteronympha are given.
Key words. Tenebrionidae, Lagriini, Lagria tenenbaumi, Apteronympha, Lachna, type species designa-
tion, checklist, Israel.
Резюме. Приведено переописание Lagria tenenbaumi Pic, 1929, этот вид переведен в подрод Apte-
ronympha Seidlitz, 1898. Lagria hirta (Linnaeus, 1758) впервые указан из Израиля. Обозначен
типовой вид рода Lachna Billberg, 1820 (= Lagria Fabricius, 1775). Даны аннотированный список за-
паднопалеарктических видов рода Lagria и таблица для определения видов подрода Apteronympha.
Ключевые слова. Tenebrionidae, Lagriini, Lagria tenenbaumi, Apteronympha, Lachna, обозначение
типового вида, список видов, Израиль.
∗ 15th contribution to the knowledge of Lagriini. 14th contribution: Acta Zoologica Academiae Scientiarum
Hungaricae 50: 283–305.

220
Introduction
Lagria Fabricius, 1775 is a large collective genus of the primitive tenebrionoid subtribe Lagriina
with more than 150 species in the temperate, tropical and subtropical areas of Europe, Africa (including
Madagascar), Asia, New Guinea and northern parts of Australia (Northern Territory and Queensland). The
genus contains species with very different general appearance and without any synapomorphy which
would separate them from members of other genera. Virtually the absence of salient modifications in
the antennae, pronotum, elytra and legs seems to be the only reason to put them together in this genus.
Seidlitz (1898) erected the subgenus Apteronympha for two Western Mediterranean species on
the basis of flightlessness of females, although females with reduced hind wings occur in other (non-
European) species of Lagriina (even in Lagria itself) as well. Males resemble Lagria hirta (Linnaeus,
1758), the type species of the genus, in all fundamental features. There is another feature that separates
members of Apteronympha from other Western Palaearctic species of Lagria. This is the rugose (coarse
and dense) punctation of pronotum; the other species have finely and sparsely punctate (sometimes nearly
impunctate) pronotum. Although female flightlessness and difference in pronotal punctation are rather
weak traits to support subgeneric distinction, the subgenus is retained for now as a separate group, since
a thorough revision of the genus is a task of the future. Borchmann (1936) transferred Lagria pygmaea
Brancsik, 1892 to Apteronympha, but this Madagascan species is unknown to the author, and it is quite
possible that it is not closely related to the species from the Western Palaearctic.
Borchmann apparently overlooked Lagria tenenbaumi Pic, 1929, a species described from Pales-
tine, because it is absent from his monograph of World “Lagriidae” (Borchmann, 1936). Only males were
available when Pic published the very short description typical of him, so subgeneric assignment was
impossible. A redescription of the species, including the description of the female, is given below. On the
basis of the rugosely punctate pronotum and wingless female it is unequivocal that Lagria tenenbaumi
should be assigned to the subgenus Apteronympha.
Abbreviations of depositories: BMNH – Natural History Museum (formerly British Museum, Natu-
ral History), London, United Kingdom; HNHM – Hungarian Natural History Museum, Budapest, Hun-
gary; NMPR – National Museum, Prague, Czech Republic; TAUI – Tel Aviv University, Tel Aviv, Israel.
Transliteration of locality names is according to the Touring Map of Israel (1 : 250 000) + List of
settlements, antiquity sites and road distances, (2001) Survey of Israel, Cartography Department, Tel Aviv.
Lagria tenenbaumi Pic, 1929 (Figs 1–4)
Lagria tenenbaumi Pic, 1929: 2; Bodenheimer, 1937: 135.
Original description: “Lagria Tenenbaumi nov. (? var. de tristis Bon). ♂ Sat elongatus, parum nitidus, sat longe gri-
seo pubescens, fortiter et pro parte dense ruguloso-punctatus, niger, elytris postice diverse pallidioribus. Long. 6,7 mill.
Palestine: Gebatha (Tenenbaum, in coll. Tenenbaum et Pic).
En l’absence du sexe ♀, je ne puis me prononcer catégoriquement au sujet de cette nouveauté, mais j’incline à y voir
simplement une variété, à coloration générale foncée de L. tristis Bon., les antennes ayant le dernier article environ de la
longueur des quatre précédents réunis, le prothorax aussi large que long, densément ponctué.”
Redescription. M a l e (Fig. 1). Body winged, elongate, slender, almost parallel-sided, dark brownish black (teneral
specimens paler), weakly shining; vestiture consisting of short, semierect greyish hairs leaving dorsal surface visible. Body
length 4.8–6.1 mm (n = 42).
Head short, coarsely and densely punctate, interspaces much narrower than a puncture diameter; frons shallowly im-
pressed; frontoclypeal suture deep; clypeus with anterior margin slightly concave; temples short, converging posteriorly; eyes
large, bulging, interocular distance shorter than eye diameter. Pubescence sparse, with a few long hairs on temples. Maxilla
with apical palpomere narrowly securiform. Labium with apical palpomere spindle-shaped, apex not emarginate. Antennae
when posteriorly extended surpassing posterior margin of pronotum by last two antennomeres; antennomere 1 subglobose, 2
subequal in length and width, 3 twice as long as 2, 4 slightly shorter than 3, 5 and 6 slightly shorter than 4, 7–9 as long as 5
and 6 but slightly broader, 10 subequal in length and width, 11 as long as four preceding antennomeres combined.
Prothorax nearly as long as wide, constricted just posterior to middle, widest just anterior to middle; pronotum with
dorsal surface slightly convex, with barely visible mesal impression immediately before posterior margin; punctation coarse
and dense, interspaces narrower than a puncture diameter; pubescence partly proclinate. Prothoracic hypomeron and proster-
num with coarse and sparse punctures and fine transverse wrinkles. Scutellum small, rounded triangular, coarsely punctate.
Elytra about 3.5 times as long as wide and 4 times as long as pronotum; sides parallel, barely diverging posteriorly;
punctation relatively coarse and dense, without trace of longitudinal rows; punctures separated by interspaces of 0.5 to

221
1 puncture diameter; interspaces without microreticulation. Pubescence reclinate. Mesoventrite finely punctate; mesoventral
process about 1/5 times as wide as mesocoxa; mesepisternum, metepimeron, and metepisternum coarsely and densely punc-
tate, metaventrite very finely and sparsely punctate.
Legs long and slender; femora subequal in length to tibiae, and practically not clavate; length of hind femur about
twice width of elytron at level of hind coxae; tibiae nearly straight, subequal in width; femora and tibiae without denticles,
angulations or any other modifications; tarsi definitely shorter than tibiae.
Abdominal ventrites finely and sparsely punctate, punctation becoming denser toward lateral portion; last ventrite
rounded posteriorly. Aedeagus with basale about 8 times as long as apicale, latter narrowly triangular (Figs 3, 4).
F e m a l e (Fig. 2). Body wingless, shorter and broader. Body length 6.8–7.0 mm (n = 2). Eyes small, transverse; in-
terocular distance 2.5 times eye diameter; antennae short, when posteriorly extended barely surpassing posterior margin
of pronotum; antennomere 11 twice as long as 10; prothorax slightly transverse, disc without midlongitudinal impression;
elytra distinctly widened posteriorly, with traces of longitudinal elevations in posterior half.
Material. I s r a e l . ‘Adullam, 17 V 2002, V. Mandelik & V. Chikatunov (1 ♂, 1 ♀; TAUI); Amnun, light trap, 15
III 2003, V. Kravchenko & V. Chikatunov (1 ♂; TAUI); Amnun, light trap, 15 V 2003, V. Kravchenko & V. Chikatunov
(3 ♂; TAUI); Been Mashichim [= Be`er Mash`abim], 7 XII 1996, R. Hoffman (1 ♂; TAUI); Esdraelon, Mount Gilboa [=
Yizre’`el Valley, Hare Gilboa’], unknown date and collector [probably Q. Argaman] (1 ♂; TAUI); Gan Shomeron, 27 IX
1957, Hebr[ew] Univ[ersity] (1 ♂; TAUI); Golan, Senir, 23 IX 1997, V. Chikatunov (2 ♂; TAUI); Haifa, Checkpost, 20 IV
1998, V. Chikatunov & T. Pavlíček (1 ♂; TAUI); Hammat Gader, 15 VI 2002, light trap, V. Kravchenko (1 ♂; TAUI); Kok-
hav HaYarden, light trap, 15 VI 2002, V. Kravchenko (1 ♂; TAUI); Lod, 8 X 1997, R. Hoffman (1 ♂; TAUI); Majdal Shams,
2 X 2002, V. Kravchenko (1 ♂; TAUI); Majdal Shams, 20 V 2001, V. Kravchenko (2 ♂, HNHM; 3 ♂, TAUI); Merom
Figs 1, 2. Lagria (Apteronympha) tenenbaumi Pic, 1926, male and female. Scale = 2 mm.

222
Figs 3, 4. Lagria (Apteronympha) tenen-
baumi Pic, 1926, aedeagus, ventral and lateral
views. Scale = 0.5 mm.
Golan, 12 VI 2000, V. Chikatunov (1 ♂; TAUI); Mi-
khmoret, unknown date and collector [probably Q. Ar-
gaman] (2 ♂; TAUI); Nahal Dishon, Rt. 886, light trap,
22 V 1998, A. Freidberg (1 ♂, BMNH; 2 ♂, TAUI);
Nahal Oren, Mt. Carmel, SFS [= south-facing slope],
light trap, 30 X 1997, T. Pavlíček (2 ♂, NMPR; 1 ♂,
HNHM); Nahal Oren, S1, 5 X 1997, V. Chikatunov &
T. Pavlíček (1 ♂; TAUI); Nahal Oren, SFS1 [= south-
facing slope], 30 X 1997, V. Chikatunov & T. Pavlíček
(1 ♂; TAUI); Nahal Oren, SFS2 [= south-facing slope],
light, 21 V 1998, V. Chikatunov & T. Pavlíček (1 ♂;
TAUI); Nahal Sansan, 17 V 2002, V. Mandelik &
V. Chikatunov (1 ♂; TAUI); Ramallah, 600 m, 10 IX
1958, J. Klapperich (1 ♂; HNHM); Ramot Avishov, 17
V 2002, V. Mandelik & V. Chikatunov (1 ♀; HNHM);
Samaria, Wadi el Faria [= Nahal Tirza], unknown date
and collector [probably Q. Argaman] (2 ♂; TAUI);
Segula, Qiryat Gat, 5 V 1996, V. Chikatunov (1 ♂;
TAUI); Tel Dan, 15 VI 2002, light trap, V. Kravchenko
(1 ♂; TAUI); Tel Dan, 5 X 2002, light trap, V.
Kravchenko & V. Chikatunov (1 ♂, BMNH; 1 ♂,
TAUI); Zekharya, 17 V 2002, V. Mandelik &
V. Chikatunov (1 ♂; TAUI); Zur Hadassa, light trap, 29
IV 2001, V. Kravchenko (1 ♂; TAUI).
Distribution. All known localities of this
species are in Israel.
Diagnostic characters of L. tenenbaumi
see in the key below (with data on the distribu-
tion of the species in the Iberian Peninsula after
Español, 1964 and Viñolas, Cartagena, 2005).
Key to species of the subgenus Lagria (Apteronympha)
1a. Body unicoloured dark brownish black, at most tibiae slightly paler. Female without midlongitudinal
impression on pronotum. – Israel .............................................................. Lagria tenenbaumi Pic, 1929
1b. Body not unicoloured: at least elytra light reddish brown. Female with more or less distinct midlongi-
tudinal impression on pronotum. – Western Mediterranean .................................................................. 2
2a. Male with interocular distance more than half of eye diameter. Legs, antennae, head and pronotum
entirely black. Elytral pubescence shorter. Female with shallow midlongitudinal impression restricted
to posterior half of pronotum. – Iberian Peninsula ...................................... Lagria rubida Graells, 1855
2b. Male with interocular distance less than half of eye diameter. Legs and antennae (frequently also
pronotum and head) at least partly reddish brown. Elytral pubescence somewhat longer. Female with
deep midlongitudinal impression almost reaching anterior margin of pronotum. – Iberian Peninsula,
southern France, northern Italy, southern Switzerland; ?Morocco .. Lagria rugosula Rosenhauer, 1856
Lagria hirta (Linnaeus, 1758)
This is the second species of Lagria occurring in Israel. It has winged females, larger and bicol-
oured body (the elytra are reddish brown, the rest is black) and finely punctate pronotum.
Material. Jerusalem, 8 IV 1966, unknown collector (1 ♂; TAUI); N kziv [= Nahal Keziv], 4 VI 1999, M. Finkel (1
♂; TAUI); Nahal Keziv, 28 IX 1999, M. Finkel & V. Kravchenko (1 ♂; TAUI); Mt. Meiron [= Har Meron], 18 IX 1976,
A. Freidberg (1 ♂; TAUI), and 12 VII 2002, light trap, V. Kravchenko (1 ♀; TAUI).
3 4

223
Checklist of the Western Palaearctic species of the genus Lagria
Much of the synonymy is based on Borchmann (1936). A few names were synonymised by Merkl
(2004). L. brevipilis Desbr. and L. pici Normand are unknown to the author; it is not impossible that these
names are synonyms of other, well known species of Lagria occurring in North Africa.
Subgenus Apteronympha Seidlitz, 1898: 336
Type species Lagria rubida Graells, 1855, designated by Merkl, 2004: 284.
Lagria rubida Graells, 1855: 74. Spain
= Lagria parvula Perris, 1864: 291
= Lagria elliptica Fairmaire, 1880: 242
Lagria rugosula Rosenhauer, 1856: 223. Europe: France, Italy, Portugal, Spain, Switzerland;
North Africa: Morocco
= Lagria tristis Bonelli, 1812: 172, not Fabricius, 1801: 70
= Lagria glabrata Olivier, 1792: 446, not Fabricius, 1775: 125
= Lagria pubescens Latreille, 1804: 351, not Linnaeus, 1767: 603
= Lagria melichari Reitter, 1890: 173
Lagria tenenbaumi Pic, 1929: 1. Asia: Israel
Subgenus Lagria Fabricius, 1775: 124
Type species Chrysomela hirta Linnaeus, 1758, designated by Borchmann, 1916: 59.
= Lachna Billberg, 1820: 35. Type species Chrysomela hirta Linnaeus, 1758, here designated.
Lagria atripes Mulsant et Guillebeau, 1855: 74. Europe: north to Belgium, Germany and Poland;
Asia: Iran, Turkmenistan, Turkey
= Lagria atripes var. puncticollis Seidlitz, 1898: 337
= Lagria atripes var. tenuicollis Seidlitz, 1898: 337, not Fairmaire, 1895: 450
Lagria brevipilis Desbrochers des Loges, 1881: 143. North Africa: Algeria
Lagria grenieri Brisout de Barneville, 1867: 181. Europe: France, Portugal, Spain
Lagria hirta (Linnaeus, 1758: 377) (Chrysomela). Europe: north to Fennoscandia; North Africa:
Algeria, Morocco; Asia: Russia (Western and Eastern Siberia), Israel, Cyprus, Turkey, Iran, Iraq, Kazakh-
stan, Turkmenistan, Uzbekistan, Tajikistan
= Cantharis spadicea Scopoli, 1763: 43
= Chrysomela pubescens Linnaeus, 1767: 603
= Lagria glabrata Fabricius, 1775: 125
= Tenebrio villosus De Geer, 1775: 44
= Cantharis flava Geoffroy, 1785: 155
= Lagria lurida Krynicki, 1832: 137
= Lagria nudipennis Mulsant, 1856: 20
= Lagria depilis Mulsant, 1856: 21
= Lagria caucasica Motschulsky, 1860: 144
= Lagria fuscata Motschulsky, 1860: 144
= Lagria pontica Motschulsky, 1860: 144
= Lagria hirta var. limbata Desbrochers des Loges, 1881: 142
= Lagria seminuda Reitter, 1889: 259
Lagria lata Fabricius, 1801: 70. Europe: Spain; North Africa: Algeria, Morocco
= Lagria rugata Desbrochers des Loges, 1881: 145
Lagria pici Normand, 1936: 161. North Africa: Tunisia

224
Acknowledgements
Thanks are due to Dr Vladimir Chikatunov and Mr Ariel-Leib-Leonid Friedman (TAUI), Dr Tomáš
Pavlíček (Institute of Evolution, University of Haifa, Israel) and Dr Vladimir Švihla (NMPR) for sending
material investigated. Mr Friedman helped to rectify transliteration of the locality names. Dr Anthony
Davies (Canadian National Collection of Insects, Arachnids and Nematodes, Agriculture and Agri-Food
Canada, Ottawa, Canada) kindly provided information about Billberg (1820). The drawings were made by
Mr János Pál (HNHM).
References
Billberg G.J. 1820. Enumeratio insectorum in museo Gust. Joh. Billberg. Stockholm: Gadelianis. [4] + 138 pp.
B o d e n h e i m e r F . S . 1937. Prodromus faunae Palestinae. Essai sur les éléments zoogéographiques et historiques
du sud-ouest du sous-règne Paléarctique. Mém. Inst. Egypte. 33: 1–286.
Bonelli F.A. 1812. Specimen faunae subalpinae sistens Insecta Pedemontii hucusque inedita, aut rariora, aut ea
quae commodi damnive gratia quod inferunt, prudentis agricolae magis interest cognoscere. Per genera naturalia
et species digesta. Fasc. I. Coleoptera plerumque inedita comprehendens. Mem. R. Soc. Agric. Torino. 9: 149–183.
B o r c h m a n n F . 1916. Die Lagriinae (Unterfamilie der Lagriidae). Arch. Naturgeschichte. 81A [1915]: 46–188.
B o r c h m a n n F . 1936. Coleoptera Heteromera Fam. Lagriidae. Genera Insectorum. Fasc. 204. Bruxelles: Louis
Desmet-Verteneuil. 561 pp.
B r i s o u t d e B a r n e v i l l e C . 1867. Lagria grenieri. Matériaux pour servir à la faune des Coléoptères de
France. Paris: A. Grenier: 181–182.
D e G e e r C . 1775. Mémoires pour servir a l’Histoires des Insectes. T. 5. Stockholm: Pierre Hesselberg. 448 pp.
Desbrochers des Loges M.
1881. Insectes Coléoptères du nord de l’Afrique nouveaux ou peu connus 1.
Mémoire. Tenebrionides. Bull. Acad. Hippone. 16: 51–168.
Español F . 1964. Las Lagria de España. Graellsia. 20: 281–286.
F a b r i c i u s J . C . F . 1775. Systema entomologiae, sistens insectorum classes, ordines, genera, species, adiectis
synonymis, locis, descriptionibus, observationibus. Flensburgi et Lipsiae: Libraria Kortii. [32] + 832 pp.
F a b r i c i u s J . C . F . 1801. Systema eleutheratorum secundum ordines, genera, species adiectis synonimis, locis,
observationibus, descriptionibus. Kiliae: Bibliopolii Academici. T. 1. xxiv + 506 pp.; T. 2. 687 pp.
F a i r m a i r e L . 1880. Descriptions de Coléoptères d’Espagne et de Turquie. Ann. Soc. Entomol. France. Cinquième
série. 10: 237–244.
F a i r m a i r e L . 1895. Description de Coléoptères de Madagascar et îles voisines. Ann. Soc. Entomol. Belgique. 39:
443–454.
G e o f f r o y E . L . 1785. In: Fourcroy S. F.: Entomologia parisiensis; sive Catalogus Insectorum quae in Agro
Parisiensi reperiuntur; secundum methodum Geoffraeanam in sectiones, genera & species distributus: cui addita
sunt nomina trivalia & fere trecentae novae species. 1. Parisiis: Serpentineis. vii + 231 pp.
G r a e l l s D . M . P . 1855. Memorias de la Comision del Mapa Geológico de España, año de 1855. Parte zoo-
lógica. Madrid: Imprenta Nacional. 111 pp.
K r y n i c k i I . 1832. Enumeratio Coleopterorum Rossiae meridionalis et praecipue in Universitatis Caesareae
Charkoviensis circulo obvenientium, que annorum 1827–1831 spatio observavit. Bull. Soc. Natur. Mosc. 5: 65–179.
Latreille P.A. 1804. Histoire naturelle, générale et particulière, des crustacés et des insectes. Ouvrage faisant
suite aux oeuvres de Leclerc de Buffon, et partie du cours complet d'histoire naturelle rédigé par C.S. Sonnini,
membre de plusieurs sociétés savantes. Tome dixième. Paris: F. Dufart. 445 pp.
L i n n a e u s C . 1758. Systema naturae per regna tria naturae, secundum classes, ordines, genera species, cum
characteribus, differentiis, synonymis, locis. Editio decima, reformata. Tomus I. Holmiae: Laurentii Salvii. [4] +
823 + [1] pp.
M e r k l O . 2004. On taxonomy, nomenclature, and distribution of some Palaearctic Lagriini, with description of
a new species from Taiwan (Coleoptera: Tenebrionidae). Acta Zool. Acad. Sci. Hung. 50: 283–305.
M o t s c h u l s k y V . d e . 1860. Coléoptères de la Sibérie orientale et en particulier des rives de l’Amur par MM.
Schrenck, Maack, Ditmar, Voznessenski etc. Dr. L. v. Schrenk's Reisen und Forschungen im Amur-Lande. Band
II. Zweite Lieferung. Coleopteren. St.-Pétersbourg: Kaiserl. Akad. Wissenschaften. 79–257 + [1] pp.

225
Mulsant E., Guillebeau F. 1855. Notes pour servir a l’histoire des Lagries (Insectes Coléopteres Hétéro-
mères). Ann. Soc. Linn. Lyon. 2: 65–74.
M u l s a n t E . 1856. Histoire Naturelle des Coléoptères de France. Latipennes. L. Paris: Maison. [4] + 44 + [5] pp.
N o r m a n d H 1936. Contribution au Catalogue des Coléoptères de la Tunisie (9e fascicule). Bull. Soc. Hist. Nat.
Afr. Nord. 27(5): 144–164.
O l i v i e r A . G . 1792 [1791]. Encyclopédie Méthodique. Histoire naturelle. Insectes. Tome sixième. (1). Paris:
Panckoucke: 369–704.
P e r r i s É . 1864. Description de quelques espèces nouvelles de Coléoptères et notes diverses. Ann. Soc. Entomol.
France. Quatrième série. 4: 275–303.
P i c M . 1929. Notes diverses, nouveautés. Échange. 45: 1–2.
Reitter E. 1889. Berichte über die von E. v. Oertzen im Jahre 1887 in Griechenland u. Klein-Asien gesammelten
Coleopteren. IX. Neue Arten aus verschiedenen Familien. Deutsche Entomol. Zeitschr.: 251–259.
Reitter E. 1890. Neue Coleopteren aus Europa, der angrenzenden Ländern und Sibirien, mit Bemerkungen über
bekannte Arten. Deutsche Entomol. Zeitschr.: 165–176.
R o s e n h a u e r W . G . 1856. Die Thiere Andalusiens nach dem Resultate einer Reise zusammengestellt, nebst den
Beschreibungen von 249 neuen oder bis jetzt unbeschriebenen Gattungen und Arten. Erlangen: Theodor Blaesing.
VIII + 429 pp.
S c o p o l i J . A . 1763. Entomologia Carniolica exhibens Insecta Carnioliae indigena et distributa in ordines, gen-
era, species, varietates methodo Linnaeana. I. Vindobonae: N. Trattner. 420 pp.
Seidlitz G. von. 1898. Lagriidae. Naturgeschichte der Insecten Deutschlands. Erste Abtheilung. Coleoptera.
Fünfter Band. Zweite Hälfte. Berlin: Nicolaische Verlags-Buchhandlung: 306–364.
V i ño l a s A., C a r t a g e n a M. C. 2005. Fauna de Tenebrionidae de la Península Ibérica y Baleares. Vol. I. La-
griinae y Pimeliinae. Barcelona: Argania editio. 428 pp.