A revised checklist of Hawaiian mosses

Abstract

A revised and updated literature-based checklist of Hawaiian mosses is presented. Geographic coverage includes the eight main Hawaiian Islands; the Northwestern Hawaiian Islands are excluded. The checklist is alphabetically ordered by scientific names; the family is noted for each genus. Synonyms and misapplied names are cross-referenced to the accepted names. A bibliography of supporting references is included.

Full text

Tropical Bryology 25: 35-69, 2004
A revised checklist of Hawaiian mosses
G. W. Staples, C. T. Imada, W. J. Hoe, and C. W. Smith
Correction for Page 38:
Hawaiian Moss Flora in Summary
Endemic Indigenous Alien Total
Families 0 41 1 42
Genera 2 126 7 135
Species 75 166 14 255
Total named taxa 90 169 14 273
Correction for Page 39:
Frahm et al. 2000, cited under Amphidium tortuosum, should correctly refer to the following:
Frahm, J.-P., T. Klöcker, R. Schmidt, and C. Schöter. 2000. Revision der Gattung Amphidium
(Musci, Dicranaceae). Tropical Bryology 18: 173-184.

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TROPICAL BRYOLOGY 25 (2004)
Revised checklist of Hawaiian mosses
Tropical Bryology 25: 35-69, 2004
A revised checklist of Hawaiian mosses1
G. W. StaplesA, C. T. ImadaA, W. J. Hoe,B and C. W. SmithC
A – Hawaii Biological Survey, Bishop Museum, 1525 Bernice St., Honolulu, Hawaii 96817
U.S.A.
B – deceased;
C – Department of Botany, The Natural History Museum, Cromwell Road, London, SW7
5BD, U.K.
Abstract. A revised and updated literature-based checklist of Hawaiian mosses is presented.
Geographic coverage includes the eight main Hawaiian Islands; the Northwestern Hawaiian Islands
are excluded. The checklist is alphabetically ordered by scientific names; the family is noted for each
genus. Synonyms and misapplied names are cross-referenced to the accepted names. A bibliography
of supporting references is included.
Introduction
The Hawaii Biological Survey (HBS) was
established as a program of Bishop Museum by
the Hawaii State Legislature in 1992, specifically
to create and maintain accurate inventories of all
species present in the Hawaiian Islands, and to
document their presence with voucher specimen
collections and bibliographies of relevant
scientific literature. In the ensuing eleven years,
comprehensive taxonomic inventories have been
compiled for many groups of organisms, both
native and alien, that comprise the Hawaiian biota.
Vascular plants and marine algae have been the
primary focus of botanical inventory efforts;
research on non-marine algae, terrestrial
cryptogamic plants, fungi, and lichens has lagged
behind.
This checklist is intended to fill one of
the gaps in our knowledge of Hawaiian terrestrial
cryptogamic plants—the mosses. Also in
preparation by HBS staff members are checklists
and bibliographies for the Hawaiian anthocerotes
(hornworts) and hepatics (liverworts). A non-
marine algal checklist for the Hawaiian Islands is
also in an advanced stage of preparation (A.
Sherwood, pers. comm.). Eventually a checklist
of Hawaiian lichens is planned. These ongoing
inventories will help the Hawaii Biological Survey
to fulfill its mission by systematically compiling
literature-based inventories for each of these
groups of organisms.
Genesis of the current checklist
This checklist of Hawaiian mosses begins
where its predecessor—the checklist compiled
by Hoe (1974)—left off, and updates that
document by revising the nomenclature, adding
new taxa and distributional records, and
providing a bibliography of literature that
supports the names in current use. The checklist
1 Contribution No. 2004–001 to the Hawaii
Biological Survey.

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TROPICAL BRYOLOGY 25 (2004)
Staples et al.
and bibliography were compiled by a
collaborative effort between HBS staff members
(Staples, Imada) working in partnership with two
bryological specialists (Hoe, Smith), who had
assembled a cache of literature citations and new
distribution records pertinent to Hawaiian
mosses. Funding from the USDA–PLANTS
database project in 1998 provided the impetus
for Staples and Imada to undertake conversion
of Hoe’s 1974 moss checklist to an electronic
format and then begin to update the taxonomy
and distribution information it contained. Hoe
and Smith generously shared their preliminary
cache of information in order to facilitate the
update process. Sadly, in May 2000, Hoe died
suddenly, a great loss to the botanical community
in Hawai‘i and to the world bryological
community. After some reorganization the project
resumed and was completed by Staples and
Imada. In the final version of the checklist,
taxonomic decisions were made by Staples, with
review and critical input from Smith and two
independent reviewers.
Coverage
It must be emphasized that this is a
literature-based taxonomic checklist, not a
specimen-based inventory. It is compiled from
literature published from the mid-1970s to the
present (2003). The starting point is the checklist
published by Hoe (1974), who summarized the
literature and reconciled the scientific names for
Hawaiian mosses in use up to that time. The
present checklist attempts to account for all
scientific names in use for Hawaiian mosses in
the latter decades of the twentieth century; it is
not a comprehensive nomenclator for all names
that have ever been published for, or applied to,
Hawaiian mosses. No nomenclatural novelties or
taxonomic changes are initiated in this checklist.
Geographically, coverage focuses on the
eight main Hawaiian Islands. There are, as yet,
no published reports for bryophytes from the
uninhabited Northwestern Hawaiian Islands,
although a few bryophyte collections were made
there in the last decades of the twentieth century.
It had been Bill Hoe’s intention to identify these
voucher specimens and write them up for
publication but the specimens remain unstudied
as this checklist goes to print. They await
inclusion in a future iteration of the Hawaiian
moss checklist.
In the last quarter of the twentieth
century there was a renaissance of bryological
research worldwide and as a result many
revisions, floristic accounts, and several
monographs have been published that impact
Hawaiian moss nomenclature. These regional and
global studies resulted in significant taxonomic
changes, as well as reporting discovery of several
taxa new to the Islands, either rare native mosses
or newly naturalized alien species. This checklist
aims to summarize all this information in a concise
format, as follows.
Format
Accepted scientific name
The main body of the checklist is
alphabetically organized by genus, species, and
infraspecific taxon. Accepted names appear in
bold italic type. Authors are included for the
genera, species, and infraspecific epithets.
Authors of moss genera have been standardized
against the classification proposed in Bryophyte
Biology (Shaw & Goffinet 2000); authors for five
genera not found in that classification have been
checked on the MOST website (cited below). All
author names are abbreviated according to
Authors of Plant Names style (Brummitt & Powell
1992). References that explain the taxonomic
concept or provide geographic distribution data
are cited after the accepted taxon name.
The first mention of a genus includes
the family assignment on the same line. Most
genera are still placed in the same families
recognized by Hoe (1974), but a few new family
placements have been made since then. In such
cases, we provide first the family assignment
used by Hoe (1974), and mention alternative
family placements afterward, with a reference to
the source.
Synonyms and misapplied names (in
italics) follow the accepted name in paragraph
style. These are limited primarily to names in use
for Hawaiian mosses in the latter part of the
twentieth century, although a few names from
other parts of the world that enter into the
nomenclature for particular taxa have been
included to enable tracking names through the
literature.

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TROPICAL BRYOLOGY 25 (2004)
Revised checklist of Hawaiian mosses
Status
Following the taxonomy is a column
indicating status of the taxon in the Hawaiian
flora: end indicates an endemic, ind indicates
indigenous, and nat signifies naturalized. A ?
following any of these signifies uncertainty about
the status of the taxon in the Hawaiian flora. While
some research has been devoted to the mode of
introduction of pteridophytes and angiosperms
to Hawai‘i, resulting in a list of species that are
likely to have been introduced by Polynesian
voyagers that settled the archipelago and another
(much larger) list of plants that arrived following
European contact, little attention has been
devoted to bryophytes in this regard. The authors
have used their best judgment in cases where
the status is not clear and appended a ? to
whatever status we assigned, to draw attention
to the species and invite investigation from
specialists.
One case worth noting is Racopilum
cuspidigerum, which is widespread in the
Southern Hemisphere but is found only in the
Hawaiian Islands in the Northern Hemisphere. It
is a prime candidate as a Polynesian introduction
to Hawai‘i, and indeed to many of the other
Polynesian islands in the South Pacific (S. Olson,
pers. comm.).
Island distribution
Following the status category is a
summary of geographic distribution on the eight
main Hawaiian Islands. The island name
abbreviations are explained at the foot of each
page. An island abbreviation appears in the row
to the right of the taxon name to signify that the
taxon is reported in the literature from that island.
Absence of the island name abbreviation
indicates the moss is not (yet) known from that
island. Given the inadequate state of moss
collecting in the Hawaiian Islands, many new
island distribution records are sure to be found
in the future.
Synonym list and Literature Consulted
Following the main body of the checklist,
an alphabetical list of all synonyms and
misapplied names is provided. This list cross-
references every synonym or misapplied name
to the main entry found under the scientific name
accepted in this checklist. The checklist
concludes with a bibliography of the references
cited in the checklist; more than 130 bryological
titles were consulted in the course of the project,
but only those providing taxonomic or
distribution data pertinent to this checklist are
included here.
Anomalies and incomplete records
A few moss names without any island
distribution are included, based on (mostly) older
literature that reports the species from the
Hawaiian (or sometimes Sandwich) Islands
without stating on which island the species was
found. Two examples in the list are Dicranoweisia
cirrata and Plagiothecium denticulatum. Their
presence in the Hawaiian Islands needs to be
confirmed and documented with contemporary
collections.
Another anomaly is Mniobryoides
degeneri, an invalid name published without a
description or indication of a type specimen by
H. Hörmann. It has been pointed out (S. Olson,
pers. comm.) that this is probably a
misidentification for some well-known species.
However, M. degeneriae has never been assessed
by a competent bryologist to ascertain its true
identity. We leave the name in the checklist to
point out that more study is required.
A note about family circumscriptions
The past 25 years has brought an
explosion in research on bryophyte systematics
and a great deal of new information has accrued,
which has led to many changes in moss
classification. Family classification in particular
has become increasingly problematic as new data
provide insights into relationships between
genera. Several new families have been erected
and other genera have been reassigned. We have
indicated the family where Hoe (1974) assigned
each genus, followed by alternative family
assignments where these were discovered. Two
principle reference works were consulted for
family assignments: first, Bryophyte Biology
(Shaw & Goffinet 2000), and in a few cases the
bryological nomenclator included in the
W3MOST database of mosses (http://
mobot.mobot.org/W3T/Search/most.html) on the
TROPICOS website.

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Staples et al.
The way forward
It is hoped that by bringing together the literature
and providing an updated checklist of names in
current use, one or more competent bryologists
might be inspired to undertake preparation of a
moss flora for the Hawaiian Islands. The last
floristic account of the Hawaiian mosses is that
by Bartram (1933, 1939). Bill Hoe intended to write
a modern flora of the Hawaiian mosses, but his
untimely death prevented that. It is our hope that
professional bryologists will take up the challenge
to complete this much needed reference work.
Those who would study Hawaiian
mosses will find type material and historic
specimens widely distributed in herbaria around
the world. In the last decades of the twentieth
century, D. Vitt and his students as well as a team
of cryptogamic botanists from the National
Museum of Tokyo collected mosses throughout
the main Hawaiian Islands. But by far the most
comprehensive late twentieth century collections
of Hawaiian mosses are those made by Hoe
between 1965 and 1995. Today the first set of
Hoe’s collections resides in two herbaria: the
Herbarium Pacificum, Bishop Museum, Honolulu
(BISH) and the Claude Philipps Herbarium,
Delaware State University, Dover (DOV), with
duplicates widely distributed in other herbaria.
Hoe donated his personal herbarium to BISH in
1983. His residual duplicates and some new
material acquired after 1983 were deposited at
DOV after the sale of Hoe’s bryological library
and herbarium. The Hoe collections in BISH and
DOV provide an excellent resource for bryological
researchers and afford a solid foundation for any
floristic account of Hawaiian mosses.
Acknowledgments
Funding for initial data compilation was
made available by the USDA-PLANTS program;
we thank Scott Petersen and Mark Skinner for
their support and assistance. Our Museum
colleagues Katie Anderson and Karin Kami
assisted with data entry and bibliographic
compilation. Bishop Museum librarian Patrice
Belcher obtained numerous critical references
through interlibrary loan services. Reviews of the
draft checklist by Storrs Olson, U.S. National
Museum, and James Shevock, U.S. Geological
Survey, brought to light several bryological
articles overlooked during the compilation as well
as many suggestions for its improvement. Bill
Buck, New York Botanical Garden, added further
literature citations and name changes when he
edited the manuscript for Tropical Bryology. The
authors gratefully acknowledge all assistance
rendered; any errors of fact or interpretation are
the sole responsibility of the authors.
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Hawaiian Moss Flora in Summary
Endemic Indigenous Alien Total
Families0411 42
Genera 2 126 7 135
Species 75 166 14 255
Total named taxa 90 169 14 273

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species. Bryologist 83(4): 405–436.
Vitt, D.H. 1981. The genus Macrocoma II.
Geographical variation in the Macrocoma
tenue—M. sullivantii species complex.
Bryologist 83(4): 437–450.
Vitt, D.H. 1983. The New Zealand species of the
pantropical genus Macromitrium
(Orthotrichaceae: Musci): taxonomy,
phylogeny and phytogeography. Journal of
the Hattori Botanical Laboratory 54: 1–94.
Vitt, D.H., and W.J. Hoe. 1980. Andreaea in
Hawaii. Bryologist 83(2): 212–214.
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Garden, St. Louis MO.
Watanabe, R., and Z. Iwatsuki. 1981. New name
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259–261.
Wichman, J.R., and H. St. John. 1990. A chronicle
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Wilczek, R., and F. Demaret. 1976. Bryum
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Zander, R.H. 1977. The tribe Pleuroweisieae
(Pottiaceae, Musci) in Middle America.
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Zander, R.H. 1978. New combinations in
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Zander, R.H. 1982. Oxystegus tenuirostris var.
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Bryologica et Lichenologica 9(4): 72–74.
Zander, R.H. 1993. Genera of Pottiaceae: mosses
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Zander, R.H., and P.M. Eckel. 1982.
Hymenostylium recurvirostrum var. insigne
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Zander, R.H., and W.J. Hoe. 1979. Geographic
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tortelloides). Bryologist 82(1): 84–87.
Zander, R.H., and D.H. Vitt. 1979. Gametophytic
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Island abbreviations: Ni Ni‘ihau; K Kaua‘i; O
O‘ahu; Mo Moloka‘i; M Maui; Ka Kahoolawe;
H Hawai‘i.

43
TROPICAL BRYOLOGY 25 (2004)
Revised checklist of Hawaiian mosses
FullName STATUS NI K O MO L M KA H
ACROPORIUM Mitt. (Sematophyllaceae)
Acroporium fuscoflavum (Paris) Broth. [Hoe 1974,
Miller et al. 1978]
Acroporium fuscoflavum (Paris) Broth. var.
baldwinii (Paris) Broth.
?Acroporium sigmatodontium misapplied
ind K O Mo L M H
AEROBRYOPSIS M.Fleisch. (Meteoriaceae)
Aerobryopsis subdivergens (Broth.) Broth. ssp. scariosa
(E.B.Bartram) Nog. [Hoe 1974, Noguchi 1976]
Aerobryopsis scariosa E.B.Bartram
Ptychomnion aciculare misapplied
ind K O Mo M
Aerobryopsis wallichii (Brid.) M.Fleisch. [Hoe 1974,
Noguchi 1976]
Aerobryopsis longissima (Dozy & Molk.) M.Fleisch.
Aerobryopsis longissima (Dozy & Molk.)
M.Fleisch. var. dozyana M.Fleisch.
ind K O Mo L M H
AMPHIDIUM Schimp. (Dicranaceae, or
Rhabdoweisiaceae [Shaw & Goffinet 2000])
Amphidium lapponicum (Hedw.) Schimp. [Hoe 1979] ind M
Amphidium tortuosum (Hornsch.) Cufod. [Frahm et al.
2000, Muñoz & Pando 2000]
Amphidium cyathicarpum (Mont.) Broth.
Grimmia haleakalae Reichardt
Grimmia haliacalae Reichardt
ind M H
ANDREAEA Hedw. (Andreaeaceae)
Andreaea acutifolia Hook. F. & Wilson ssp. acuminata
(Mitt.) Vitt [Vitt 1980a, Vitt & Hoe 1980]
Andreaea acuminata Mitt.
ind M H
Andreaea rupestris Hedw. [Hoe 1977, Hoe 1979, Vitt
1980a, Vitt & Hoe 1980]
Andreaea perpapillosa Broth.
ind M
ANOECTANGIUM Schwägr. (Pottiaceae)
Anoectangium aestivum (Hedw.) Mitt. [Hoe 1974,
Zander 1977, Zander & Vitt 1979]
Anoectangium euchloron (Schwägr.) Mitt.
Anoectangium haleakalae (Müll.Hal.) Paris var.
haleakalae
Anoectangium haleakalae (Müll.Hal.) Paris var.
laxulum E.B.Bartram
Anoectangium harttiae E.B.Bartram
ind K O Mo L M H
ANOMOBRYUM Schimp. (Bryaceae)
Anomobryum angustirete Broth. [Hoe 1974] end K L M

44
TROPICAL BRYOLOGY 25 (2004)
Staples et al.
FullName STATUS NI K O MO L M KA H
BALDWINIELLA M.Fleisch. (Neckeraceae)
Baldwiniella kealeensis (Reichardt) E.B.Bartram [Hoe
1974, Hoe 1975]
end K O Mo L M H
BARBELLOPSIS Broth. (Meteoriaceae)
Barbellopsis trichophora (Mont.) W.R.Buck [Hoe 1974,
Streimann 1993, Buck 1998]
Barbella trichophora (Mont.) M.Fleisch.
ind K O Mo M H
BARBULA Hedw. (Pottiaceae)
Barbula indica (Hook.) Spreng. [Hoe 1978, Zander 1993,
Darigo 1996]
Barbula cruegeri Sond. ex Müll.Hal.
Barbula unguiculata misapplied
nat O M
BARTRAMIA Hedw. (Bartramiaceae)
Bartramia baldwinii Müll.Hal. [Hoe 1974] end M
Bartramia halleriana Hedw. [Hoe 1974] ind M
BRACHYMENIUM Schwägr. (Bryaceae)
Brachymenium exile (Dozy & Molk.) Bosch & Sande
Lac. [Hoe 1974]
ind K O Mo M H
Brachymenium indicum (Dozy & Molk.) Bosch & Sande
Lac. [Hoe 1979]
ind O H
BRACHYTHECIUM Schimp. (Brachytheciaceae)
Brachythecium hawaiicum E.B.Bartram [Hoe 1974] end M H
Brachythecium lamprocarpum (Müll.Hal.) A.Jaeger
[Hoe 1974]
ind K M H
Brachythecium plumosum (Hedw.) B.S.G. [Hoe 1974]
Brachythecium oxyrrhynchum (Dozy & Molk.)
A.Jaeger
Cirriphyllum oxyrrhynchum (Dozy & Molk.)
M.Fleisch.
ind K O Mo M H
Brachythecium rutabulum (Hedw.) B.S.G. [Hoe 1974] ind M
BREUTELIA (Bruch & Schimp.) Schimp.
(Bartramiaceae)
Breutelia affinis (Hook.) Mitt. [Hoe 1974, Virtanen 1997]
Breutelia kilaueae (Müll.Hal.) Broth.
ind H
Breutelia arundinifolia (Duby) M.Fleisch. [Hoe 1974,
Virtanen 1997]
ind K O M H
Breutelia crassicaulis (Müll.Hal.) Paris [Virtanen 1997] ind M H
BRYOERYTHROPHYLLUM P.C.Chen (Pottiaceae)
Bryoerythrophyllum recurvirostre (Hedw.) P.C.Chen
[Hoe 1974]
Didymodon recurvirostris (Hedw.) Jenn.
ind M

45
TROPICAL BRYOLOGY 25 (2004)
Revised checklist of Hawaiian mosses
FullName STATUS NI K O MO L M KA H
BRYUM Hedw. (Bryaceae)
Bryum argenteum Hedw. [Hoe 1974, Miller et al. 1978,
Tan & Koponen 1989]
Bryum argenteum Hedw. var. lanatum (P. Beauv.)
Hampe
Bryum leucophyllum Dozy & Molk.
ind K O M H
Bryum atrovirens Brid. [Hoe 1974]
Bryum erythrocarpum Schwägr.
ind O M H
Bryum baldwinii Broth. [Hoe 1974] end M
Bryum caespiticium Hedw. [Hoe 1974] ind M H
Bryum chrysoneuron Müll.Hal. [Miller et al. 1978] ind H
Bryum conicum Hornsch. [Ochi 1980]
Bryum ceramiocarpum Müll.Hal.
ind M
Bryum flavituber R.Wilczek & Demaret [Wilczek &
Demaret 1976]
nat O
Bryum hawaiicum Hoe [Hoe 1974]
Bryum crassicostatum Broth.
end M H
Bryum klinggraeffii Schimp. [Hoe 1979] nat O
Bryum lisae De Not. var. cuspidatum (Bruch & Schimp.)
Margad. [Olson 1999a]
nat O
Bryum mildeanum Jur. [Hoe 1974, Ochi 1974, Hoe 1979,
Ignatov & Afonina 1992]
Bryum apiculatum Schwägr.
Bryum mauiensis Broth.
Bryum megalostegium Sull.
Bryum nitens Hook.
ind K O L M
Bryum pseudocapillare Besch. [Hoe 1975] nat? O
Bryum radiculosum Brid. [Olson 1999a] nat K
BUCKIELLA Ireland (Hypnaceae)
Buckiella draytonii (Sull.) Ireland [Hoe 1974, Ireland
2001]
Hypnum draytoni(i) Sull.
Hypnum eudorae Sull.
Plagiothecium draytonii (Sull.) Müll.Hal.
end K O Mo M H
CALYMPERES Sw. (Calymperaceae)
Calymperes fasciculatum Dozy & Molk. [Hoe 1974,
Reese et al. 1986]
Calymperes hawaiiense E.B.Bartram
ind O
Calymperes tenerum Müll.Hal. [Hoe 1974] ind K O Mo M H
CAMPTOCHAETE Reichardt (Lembophyllaceae)
Camptochaete pulvinata (Hook.f. & Wilson) A.Jaeger
[Hoe 1974, Tangney 1997]
ind M
CAMPYLOPODIUM (Müll.Hal.) Besch. (Dicranaceae)
Campylopodium medium (Duby) Giese & J.-P.Frahm ind K O Mo M H

46
TROPICAL BRYOLOGY 25 (2004)
Staples et al.
FullName STATUS NI K O MO L M KA H
[Hoe 1974, Giese & Frahm 1985]
Campylopodium euphorocladum (Müll.Hal.) Besch.
CAMPYLOPUS Brid. (Dicranaceae)
Campylopus exasperatus (Nees & Blume) Brid. [Frahm
1991]
ind K
Campylopus fragilis (Brid.) Bruch. & Schimp. ssp.
zollingerianus (Müll.Hal.) J.-P.Frahm [Hoe 1974,
Frahm 1991, Frahm 1992]
Campylopus boswellii (Müll.Hal.) Paris
Campylopus boswellii (Müll.Hal.) Paris var.
capitulatus E.B.Bartram
ind K O Mo M
Campylopus hawaiicus (Müll.Hal.) A.Jaeger var.
densifolius (Ångstr.) J.-P.Frahm [Frahm 1978, Frahm
1991]
Campylopus densifolius Ångstr.
Campylopus densiretis Broth.
Campylopus wawreanus Reichardt
end K O Mo M H
Campylopus hawaiicus (Müll.Hal.) A.Jaeger var.
hawaiicoflexuosus (Müll.Hal.) J.-P.Frahm [Frahm
1978, Frahm 1991]
Campylopus densifolius Ångstr. var. falcatus Broth.
Campylopus densifolius Ångstr. var.
hawaiicoflexuosus (Müll.Hal.) H.A.Mill.
Campylopus hawaiicoflexuosus (Müll.Hal.)
Müll.Hal.
Campylopus hawaiicoflexuosus (Müll.Hal.) Paris
Campylopus perangustifolius Müll.Hal.
Campylopus purpureoflavescens Hampe, in part
Dicranum hawaiicoflexuosum Müll.Hal.
end K O Mo L M H
Campylopus hawaiicus (Müll.Hal.) A.Jaeger var.
hawaiicus [Frahm 1978, Frahm 1991]
Campylopus densifolius Ångstr. var.
purpureoflavescens (Hampe ex Müll.Hal.)
H.A.Mill.
Campylopus purpureoflavescens Hampe, in part
Dicranum hawaiicum Müll.Hal.
ind K O Mo L M H
Campylopus incurvatus J.-P.Frahm & Hoe [Frahm & Hoe
1978, Frahm 1991]
end M H
Campylopus laxitextus Sande Lac. [Hoe 1974, Frahm
1991]
Campylopus fumarioli Müll.Hal.
ind K O Mo L M H
Campylopus praemorsus (Müll.Hal.) A.Jaeger [Frahm
1976, Hoe 1979, Frahm 1991]
Campylopus exasperatus (Nees & Blume) Brid.
Campylopus microcephalus (Müll.Hal.) Paris
Dicranum microcephalum Müll.Hal.
Dicranum praemorsum Müll.Hal.
end K O L M H

47
TROPICAL BRYOLOGY 25 (2004)
Revised checklist of Hawaiian mosses
FullName STATUS NI K O MO L M KA H
Campylopus schmidii (Müll.Hal.) A.Jaeger ssp.
hemitrichius (Müll.Hal.) J.-P.Frahm [Hoe 1974, Frahm
1991]
Campylopus skottsbergii Broth.
Campylopus tubulosus E.B.Bartram
Dicranum skottsbergii Broth.
ind K O M H
Campylopus schmidii (Müll.Hal.) A.Jaeger ssp. schmidii
[Hoe 1974, Hoe 1977, Frahm 1991]
Campylopus aureus Bosch & Sande Lac.
Campylopus introflexus misapplied
Campylopus introflexus (Hedw.) Brid. var.
polytrichoides misapplied
Campylopus polytrichoides misapplied
ind K O M H
Campylopus umbellatus (Arn.) Paris var. pterotoneuron
(Müll.Hal.) J.-P.Frahm [Hoe 1974, Frahm 1991]
Campylopus geniculatus Ångstr.
ind O
Campylopus umbellatus (Arn.) Paris var. umbellatus
[Hoe 1974, Frahm 1991]
?Campylopus didrichsenii (Müll.Hal.) A.Jaeger
?Campylopus pertristis (Müll.Hal.) Paris
Campylopus richardii misapplied
Campylopus sandwicensis Paris
Campylopus umbellatus (Arn.) E.B.Bartram
Dicranum didrichsenii Müll.Hal.
Dicranum pertriste Müll.Hal.
Thysanomitrion hawaiicum Müll.Hal.
Thysanomitrion umbellatum Arn.
ind K O L M H
Campylopus wheeleri (Müll.Hal.) Paris [Hoe 1974,
Frahm 1991]
Dicranoloma wheeleri (Müll.Hal.) Paris
Dicranum wheeleri Hampe ex Müll.Hal.
end K M H
CERATODON Brid. (Ditrichaceae)
Ceratodon purpureus (Hedw.) Brid. ssp. purpureus [Hoe
1974, Hoe 1977, Miller et al. 1978, Burley & Pritchard
1990]
Ceratodon purpureus (Hedw.) Brid. fma. purpureus
Ceratodon purpureus (Hedw.) Brid. fma. xanthopus
(Sull.) E.Britton ex Grout
ind K O M H
CHENIA R.H.Zander (Pottiaceae)
Chenia leptophylla (Müll.Hal.) R.H.Zander [Hoe 1974,
Zander 1993]
Tortula rhizophylla (Sakurai) Z.Iwats. & K.Saito
Tortula vectensis E.F.Warb. & Crundw.
nat O
CLAOPODIUM (Lesq. & James) Renauld & Cardot
(Thuidiaceae, or Leskeaceae [Shaw & Goffinet 2000])
Claopodium prionophyllum (Müll. Hal.) Broth. [Hoe
1974]
ind O Mo M H

48
TROPICAL BRYOLOGY 25 (2004)
Staples et al.
FullName STATUS NI K O MO L M KA H
Claopodium amblystegioides Dixon
Claopodium nervosum M.Fleisch.
Claopodium whippleanum (Sull.) Renauld & Cardot
[Hoe 1974]
Claopodium hawaiiense R.S.Williams
ind K O M
CLASTOBRYOPSIS M.Fleisch. (Sematophyllaceae)
Clastobryopsis robusta (Broth.) M.Fleisch. [Hoe 1974,
Tan & Jia 1999]
Aptychella robusta (Broth.) M.Fleisch.
ind K M
CTENIDIUM (Schimp.) Mitt. (Hypnaceae)
Ctenidium elegantulum Broth. [Hoe 1974, Nishimura
1985]
end O M H
CYRTOPUS (Brid.) Hook.f. (Cyrtopodaceae)
Cyrtopus setosus (Hedw.) Hook.f. [Hoe 1974] ind H
DALTONIA Hook. & Taylor (Hookeriaceae, or
Daltoniaceae [Shaw & Goffinet 2000])
Daltonia baldwinii Broth. [Hoe 1974] end M H
Daltonia contorta Müll.Hal. [Hoe 1974, Hoe 1979] ind K O Mo L M H
Daltonia pseudostenophylla E.B.Bartram [Hoe 1974] ind O M H
Daltonia rufescens Broth. [Hoe 1974] ind O M H
DICRANELLA (Müll.Hal.) Schimp. (Dicranaceae)
Dicranella exilis Sull. [Hoe 1974] end M
Dicranella hawaiica (Müll.Hal.) Broth. var. hawaiica
[Hoe 1974]
Microdus hawaiicus (Müll.Hal.) Paris
end K Mo M
Dicranella hawaiica (Müll.Hal.) Broth. var. tomentella
E.B.Bartram [Hoe 1974]
end O M
Dicranella hochreutineri Cardot [Hoe 1974]
Dicranella hochreuteri misapplied
ind K O L M H
Dicranella integrifolia E.B.Bartram [Hoe 1974] end O Mo M H
Dicranella rigidula E.B.Bartram [Hoe 1974] end K O H
DICRANODONTIUM Bruch & Schimp. (Dicranaceae)
Dicranodontium denudatum (Brid.) E.Britton [Hoe 1974] ind M
Dicranodontium falcatum Broth. var. atrovirens (Broth.)
E.B.Bartram [Hoe 1974]
end M H
Dicranodontium porodictyon Cardot & Thér. [Hoe 1974,
Hoe 1979, Frahm 1997]
Dicranodontium falcatum Broth. var. falcatum
Dicranodontium hawaiicum Broth.
ind K O Mo M H
DICRANOLOMA (Renauld) Renauld (Dicranaceae)
Dicranoloma gracile Broth. ex E.B.Bartram [Hoe 1974] end M

49
TROPICAL BRYOLOGY 25 (2004)
Revised checklist of Hawaiian mosses
FullName STATUS NI K O MO L M KA H
DICRANOWEISIA Milde (Dicranaceae, or
Rhabdoweisiaceae [Shaw & Goffinet 2000])
Dicranoweisia cirrata (Hedw.) Lindb. [Hoe 1974, Miller
et al. 1978]
ind H?
DICRANUM Hedw. (Dicranaceae)
Dicranum speirophyllum Mont. var. breviflagellare
(Müll.Hal.) E.B.Bartram [Hoe 1974]
end O M H
Dicranum speirophyllum Mont. var. condensatum (Sull.)
Wijk & Margad. [Hoe 1974]
end H
Dicranum speirophyllum Mont. var. elongatum (Sull.)
Wijk & Margad. [Hoe 1974]
end M
Dicranum speirophyllum Mont. var. speirophyllum [Hoe
1974]
end K O Mo M H
DIDYMODON Hedw. (Pottiaceae)
Didymodon insulanus (De Not.) M.O.Hill [Hoe 1974,
Zander 1978, Corley et al. 1981]
Barbula vinealis Brid. var. cylindrica (Taylor)
Boulay
Barbula vinealis Brid. var. flaccida B.S.G.
Didymodon vinealis (Brid.) R.H.Zander var.
flaccidus (Bruch & Schimp.) R.H.Zander
ind O M
DISTICHIUM Bruch & Schimp. (Ditrichaceae)
Distichium capillaceum (Hedw.) B.S.G. [Hoe 1974] ind H
DISTICHOPHYLLUM Dozy & Molk. (Hookeriaceae,
or Daltoniaceae [Shaw & Goffinet 2000])
Distichophyllum freycinetii (Schwägr.) Mitt. var.
crasseturgescens (Müll.Hal.) E.B.Bartram ex Hoe [Hoe
1974]
Mniadelphus freycineti (Schwägr.) Müll.Hal. var.
crasseturgescens Müll.Hal.
end
Distichophyllum freycinetii (Schwägr.) Mitt. var.
freycinetii [Hoe 1974]
end K O Mo L M H
Distichophyllum paradoxum (Mont.) Mitt. [Hoe 1974] end K O Mo L M H
ECTROPOTHECIUM Mitt. (Hypnaceae)
Ectropothecium arcuatum Mitt. [Hoe 1974] end K O M H
Ectropothecium decurrens (Sull.) Nishimura [Hoe 1974,
Nishimura 1985]
Ctenidium decurrens (Sull.) Broth.
Ctenidium pulcherrimum Broth.
Ctenidium submalacobolum Broth.
Cupressina decurrens (Sull.) Müll.Hal.
Ectropothecium rockii Broth.
Hypnum decurrens Sull.
end K O Mo M H

50
TROPICAL BRYOLOGY 25 (2004)
Staples et al.
FullName STATUS NI K O MO L M KA H
Stereodon decurrens (Sull.) Mitt.
Taxiphyllum gracile Broth.
Ectropothecium sandwichense (Hook. & Arn.) Mitt.
[Hoe 1974, Hoe 1979]
ind K O Mo L M H
Ectropothecium tutuilum (Sull.) Mitt. [Miller et al. 1978] ind H
Ectropothecium viridifolium E.B.Bartram [Hoe 1974,
Wichman & St. John 1990]
end Ni O L M
Ectropothecium zollingeri (Müll. Hal.) A.Jaeger var.
filicaule (M.Fleisch.) H.A.Mill., H.Whittier &
B.Whittier [Hoe 1974, Hoe 1975, Miller et al. 1978]
Glossadelphus zollingeri (Müll.Hal.) M.Fleisch. var.
filicaulis (M.Fleisch.) M.Fleisch.
ind K O Mo L M H
ENCALYPTA Hedw. (Encalyptaceae)
Encalypta ciliata Hedw. [Horton 1983]
Encalypta scabrata E.B.Bartram
ind M H
Encalypta rhabdocarpa Schwägr. [Hoe 1974]
Encalypta rhaptocarpa misapplied
ind H
Encalypta sandwicensis Sull. [Hoe 1974, Hoe 1977] ind M H
ENTODON Müll.Hal. (Entodontaceae)
Entodon plicatus Müll.Hal. [Hoe 1974, Buck 1990]
Entodon hillebrandii Müll.Hal.
Entodon solanderi (Ångstr.) A.Jaeger
ind K O M H
Entodon subcuspidatus (Müll.Hal.) E.B.Bartram [Hoe
1974, Hoe 1979]
ind K Mo M
ENTODONTOPSIS Broth. (Stereophyllaceae)
Entodontopsis anceps (Bosch & Sande Lac.) W.R.Buck
& Ireland [Hoe 1974, Buck & Ireland 1985]
Stereophyllum oahuense Broth.
ind O
ENTOSTHODON Schwägr. (Funariaceae)
Entosthodon subintegrus (Broth.) H.A.Mill., H.Whittier
& B.Whittier [Miller et al. 1978]
Funaria subintegra Broth.
end K O Mo L M H
FABRONIA Raddi (Fabroniaceae)
Fabronia ciliaris (Brid.) Brid. [Hoe 1977] ind M
Fabronia degeneri E.B.Bartram [Hoe 1974] end O Mo H
Fabronia nietneri Müll.Hal. [Hoe 1974, Hoe 1975] ind K O M H
FISSIDENS Hedw. (Fissidentaceae)
Fissidens aphelotaxifolius Pursell [Pursell & Hoe 1977] ind M
Fissidens bryoides Hedw. [Pursell & Hoe 1977]
Fissidens hawaiicus E.B.Bartram
Fissidens insularis E.B.Bartram
Fissidens oahuensis E.B.Bartram
ind K O Mo L M H

51
TROPICAL BRYOLOGY 25 (2004)
Revised checklist of Hawaiian mosses
FullName STATUS NI K O MO L M KA H
Fissidens delicatulus Ångstr. [Pursell & Hoe 1977] end K O L M H
Fissidens elegans Brid. [Pursell & Hoe 1977]
Fissidens baldwinii Broth.
Fissidens bishopii Hoe
ind K O Mo L M H
Fissidens hoei Pursell [Pursell & Hoe 1977] end O L
Fissidens kilaueae Hoe & H.A.Crum [Pursell & Hoe
1977]
end K O L M H
Fissidens lancifolius E.B.Bartram [Pursell & Hoe 1977] end K O Mo L M H
Fissidens lindbergii Mitt. [Pursell & Hoe 1977, Darigo
1996, Pursell 1997]
Fissidens crumii Hoe
nat O M
Fissidens nothotaxifolius Pursell & Hoe [Pursell & Hoe
1977]
end K O Mo L M H
Fissidens pacificus Ångstr. [Hoe 1974, Pursell & Hoe
1977]
Fissidens mauiensis Müll.Hal.
end K O M H
Fissidens submarginatus Bruch [Pursell & Hoe 1977,
Hoe 1979, Bruggeman-Nannenga & Pursell 1995]
Fissidens intermedius Müll.Hal.
ind K O Mo L M H
Fissidens taxifolius Hedw. [Pursell & Hoe 1977] ind O M
FLORIBUNDARIA M.Fleisch. (Meteoriaceae)
Floribundaria floribunda (Dozy & Molk.) M.Fleisch.
[Hoe 1974, Tan 1984]
Floribundaria baldwinii Broth.
ind O M H
FUNARIA Hedw. (Funariaceae)
Funaria hygrometrica Hedw. [Hoe 1974, Hoe 1975] ind O H
GAROVAGLIA Endl. (Pterobryaceae, or Garovagliaceae
[Shaw & Goffinet 2000])
Garovaglia powellii Mitt. var. haleakalae (Broth.) During
[During 1977]
Garovaglia haleakalae Broth.
ind M
GLOSSADELPHUS M.Fleisch.
1
(Sematophyllaceae)
Glossadelphus acutifolius E.B.Bartram [Hoe 1974, Tixier
1988]
end M
Glossadelphus limnobioides Broth. [Hoe 1974, Tixier
1988]
end O M H
Glossadelphus mauiensis Broth. [Hoe 1974, Tixier 1988] end M
Glossadelphus ogatae Broth. & Yasuda [Hoe 1974, Hoe
1979, Tixier 1988]
Glossadelphus chrysobasilaris Broth.
ind M H
1
Buck (1987b) reduced the type species of Glossadelphus to Phyllodon so the genus name is not usable.
The correct placement for the 4 species left here remains to be determined.

52
TROPICAL BRYOLOGY 25 (2004)
Staples et al.
FullName STATUS NI K O MO L M KA H
GRIMMIA Hedw. (Grimmiaceae)
Grimmia anodon Bruch & Schimp. [Hoe 1977, Hoe
1979, Muñoz & Pando 2000]
Grimmia apocarpa Hedw. var. pulvinata (Hedw.)
G.N.Jones
Gymnostomum pulvinatum Hedw.
ind M H
Grimmia laevigata (Brid.) Brid. [Hoe 1974, Hoe 1979,
Muñoz & Pando 2000]
ind M H
Grimmia longirostris Hook. [Hoe 1977,Muñoz 1998]
Grimmia pilifera misapplied
ind M
Grimmia mauiense Greven [Greven 2003] end M H
Grimmia maunakeaense Greven [Greven 2003] end M H
Grimmia pulvinata (Hedw.) Sm. [Muñoz & Pando 2000]
Leersia pulvinata Hedw.
ind M
Grimmia trichophylla Grev. [Hoe 1974, Muñoz & Pando
2000]
ind M H
HAGENIELLA Broth. (Sematophyllaceae)
Hageniella micans (Mitt.) B.C.Tan & Y.Jia [Hoe 1974,
Tan & Jia 1999]
Hageniella pacifica Broth.
ind K M H
HAPLOCLADIUM Müll.Hal. (Leskeaceae)
Haplocladium microphyllum (Hedw.) Broth. [Hoe 1974,
Watanabe & Iwatuski 1981, Zilstra 1990]
Bryohaplocladium microphyllum (Hedw.) R.Watan.
& Z.Iwats.
ind O
HAPLOHYMENIUM Dozy & Molk. (Thuidiaceae, or
Anomodontaceae [Shaw & Goffinet 2000])
Haplohymenium triste (Ces.) Kindb. [Hoe 1974, Hoe
1979]
ind K O M H
HETEROPHYLLIUM (Schimp.) Kindb.
(Sematophyllaceae)
Heterophyllium subauriculatum (Müll.Hal.) Broth. [Hoe
1974]
end M H
HOLOMITRIUM Brid. (Dicranaceae)
Holomitrium seticalycinum Müll.Hal. [Hoe 1974, Hoe
1979]
end K O Mo L M H
Holomitrium squarrifolium E.B.Bartram [Hoe 1974]
Holomitrium ferriei misapplied
end O H
HOMALIODENDRON M.Fleisch. (Neckeraceae)
Homaliodendron flabellatum (Sm.) M.Fleisch. [Hoe
1974]
Homalia densa Bosw.
Homalia praelonga Reichardt
ind K O Mo L M H

53
TROPICAL BRYOLOGY 25 (2004)
Revised checklist of Hawaiian mosses
FullName STATUS NI K O MO L M KA H
HOOKERIA Sm. (Hookeriaceae)
Hookeria acutifolia Hook. & Grev. [Hoe 1974, Hoe
1975]
ind O L M H
HYMENOSTYLIUM Brid. (Pottiaceae)
Hymenostylium recurvirostrum (Hedw.) Dixon var.
cylindricum (E.B.Bartram) R.H.Zander [Zander &
Eckel 1982]
Merceyopsis crassinervis (Broth.) E.B.Bartram
Molendoa crassinervis (Broth.) E.B.Bartram
ind H
Hymenostylium recurvirostrum (Hedw.) Dixon var.
recurvirostrum [Zander 1993]
Hymenostylium firmum (Müll.Hal.) Broth. ex
E.B.Bartram
ind K O M H
HYOPHILA Brid. (Pottiaceae)
Hyophila involuta (Hook.) A.Jaeger [Hoe 1974]
Hyophila dozy-molkenboeri M.Fleisch.
ind K O Mo M H
HYPNUM Hedw. (Hypnaceae)
Hypnum plumaeforme Wilson [Crum & Mueller-
Dombois 1968, Hoe 1974]
nat M H
HYPOPTERYGIUM Brid. (Hypopterygiaceae)
Hypopterygium sandwicense Broth. [Hoe 1974, Kruijer
2002]
end M H
ISOPTERYGIUM Mitt. (Hypnaceae)
Isopterygium albescens (Hook.) A.Jaeger [Hoe 1974] ind K O Mo L M H
Isopterygium vineale E.B.Bartram [Hoe 1974] end K O Mo M H
LEPTOBRYUM (Bruch & Schimp.) Wilson (Bryaceae,
or Meesiaceae [Shaw & Goffinet 2000])
Leptobryum pyriforme (Hedw.) Wilson [Hoe 1974] ind M H
LEPTODONTIUM (Müll.Hal.) Lindb. (Pottiaceae)
Leptodontium flexifolium (Dicks. ex With.) Hampe [Hoe
1974, Hoe 1977]
Leptodontium brevicaule E.B.Bartram
ind M H
LEUCOBRYUM Hampe (Leucobryaceae)
Leucobryum gracile Sull. var. gracile [Hoe 1974] end K O Mo L M H
Leucobryum gracile Sull. var. hamatum E.B.Bartram
[Hoe 1974]
end K O Mo M
Leucobryum pachyphyllum Müll.Hal. [Hoe 1974] end K O Mo L M
Leucobryum seemannii Mitt. var. fumarioli (Müll.Hal.)
H.A.Mill. [Hoe 1974]
end H

54
TROPICAL BRYOLOGY 25 (2004)
Staples et al.
FullName STATUS NI K O MO L M KA H
Leucobryum hawaiense (Reichardt) E.B.Bartram var.
fumarioli (Müll.Hal.) E.B.Bartram
Leucobryum solfatarae Müll.Hal. var. fumarioli
(Müll.Hal.) Wijk & Margad.
Leucobryum seemannii Mitt. var. seemannii [Hoe 1974,
Hoe 1975]
Leucobryum hawaiense (Reichardt) E.B.Bartram
Leucobryum solfatarae Müll.Hal. var. hawaiense
(Reichardt) Wijk & Marg.
end K O Mo L M H
Leucobryum seemannii Mitt. var. solfatarae (Müll.Hal.)
H.A.Mill. [Hoe 1974]
Leucobryum hawaiense (Reichardt) E.B.Bartram var.
solfatarae (Müll.Hal.) E.B.Bartram
Leucobryum solfatarae Müll.Hal.
end K Mo M H
LEUCOLOMA Brid. (Dicranaceae)
Leucoloma molle (Müll.Hal.) Mitt. [Hoe 1974] ind O M H
Leucoloma scaberulum E.B.Bartram [Hoe 1974] end O
LEUCOPHANES Brid. (Leucobryaceae, or
Calymperaceae [Shaw & Goffinet 2000])
Leucophanes glaucum (Schwägr.) Mitt. [Hoe 1974] ind O
LIMBELLA (Müll.Hal.) Müll.Hal. (Amblystegiaceae)
Limbella tricostata (Sull.) Müll.Hal. [Hoe 1974, Christy
1987]
Sciaromium tricostatum (Sull.) Mitt.
end K O Mo M H
MACROCOMA (Müll.Hal.) Grout (Orthotrichaceae)
Macrocoma tenue (Hook. & Grev.) Vitt ssp. sullivantii
(Müll.Hal.) Vitt [Hoe 1974, Vitt 1980b, Vitt 1981]
Macrocoma intricata (Müll.Hal.) H.A.Miller,
H.Whittier & B.Whittier
Macromitrium intricatum Müll.Hal.
ind K O Mo M H
MACROMITRIUM Brid. (Orthotrichaceae)
Macromitrium brevisetum Mitt. [Hoe 1974, Hoe 1979] end K O Mo L M H
Macromitrium emersulum Müll.Hal. [Hoe 1974, Hoe
1975]
end K O Mo L M H
Macromitrium incurvifolium (Hook. & Grev.) Schwägr.
[Hoe 1974]
Macromitrium cumingii Müll.Hal.
ind K M H
Macromitrium microstomum (Hook. & Grev.) Schwägr.
[Hoe 1974, Vitt 1983]
Macromitrium owahiense Müll.Hal.
Macromitrium reinwardtii Schwägr.
ind K O Mo L M H
Macromitrium piliferum Schwägr. [Hoe 1974] end K O Mo L M H

55
TROPICAL BRYOLOGY 25 (2004)
Revised checklist of Hawaiian mosses
FullName STATUS NI K O MO L M KA H
MACROTHAMNIUM M.Fleisch. (Hylocomiaceae)
Macrothamnium macrocarpum (Reinw. & Hornsch.)
M.Fleisch. [Hoe 1974]
ind M
MICRODUS Schimp. ex Besch. (Dicranaceae)
Microdus hillebrandii (Müll.Hal.) Paris [Hoe 1974]
Dicranella hillebrandii (Müll.Hal.) Broth.
ind O Mo
MNIOBRYOIDES Hörmann (Bryaceae)
Mniobryoides degeneriae Hörmann [Hoe 1974] end H
MNIUM Hedw. (Mniaceae)
Mnium marginatum (With.) P.Beauv. [Hoe 1974]
Mnium serratum Schrad. ex Brid.
ind M
NECKERA Hedw. (Neckeraceae)
Neckera hawaiicopennata Müll.Hal. [Hoe 1974] end M
NECKEROPSIS Reichardt (Neckeraceae)
Neckeropsis lepineana (Mont.) M.Fleisch. [Hoe 1974,
Enroth 1989]
ind K O M
Neckeropsis obtusata (Mont.) Broth. [Olson 1999b] nat K
OCTOBLEPHARUM Hedw. (Leucobryaceae, or
Calymperaceae [Shaw & Goffinet 2000])
Octoblepharum albidum Hedw. [Hoe 1974, Hoe 1975] ind K O Mo M H
ORTHODONTIUM Wilson (Bryaceae, or
Orthodontiaceae [Shaw & Goffinet 2000])
Orthodontium pellucens (Hook.) B.S.G. [Hoe 1974, Hoe
1977]
ind K M
ORTHOTRICHUM Hedw. (Orthotrichaceae)
Orthotrichum alpestre Hornsch. var. berggrenii
(E.B.Bartram) Lewinsky [Hoe 1974, Lewinksy 1993]
Orthotrichum berggrenii E.B.Bartram
end O? H
Orthotrichum diaphanum Schrad. ex Brid. [Hoe 1974,
Hoe 1977, Lewinsky 1993]
ind M H
Orthotrichum hawaiicum Müll.Hal. [Hoe 1974] end M H
Orthotrichum incrassatum Lewinsky [Lewinsky 1993] end H
Orthotrichum lyellii Hook. & Taylor [Hoe 1974,
Lewinsky 1993]
Orthotrichum verrucatum Müll.Hal.
ind
Orthotrichum rupestre Schleich. & Schwägr. [Hoe 1974,
Lewinsky 1993]
Orthotrichum hillebrandii Müll.Hal.
ind O M H

56
TROPICAL BRYOLOGY 25 (2004)
Staples et al.
FullName STATUS NI K O MO L M KA H
OXYRRHYNCHIUM (Schimp.) Warnst.
(Brachytheciaceae)
Oxyrrhynchium vagans (A.Jaeger) Ignatova & Huttunen
[Hoe 1974, Hoe 1975, Ignatov & Huttunen 2002]
Eurhynchium vagans (A.Jaeger) E.B.Bartram
Rhynchostegium vagans A.Jaeger
ind K O Mo L M H
OXYSTEGUS (Limpr.) Hilp. (Pottiaceae)
Oxystegus tenuirostris (Hook. & Taylor) A.J.E.Sm. var.
gemmiparus (Schimp.) R.H.Zander [Hoe 1977, Hoe
1979, Zander 1982]
Trichostomum tenuirostre (Hook. & Taylor) Lindb.
ind K O M
PALAMOCLADIUM Müll.Hal. (Brachytheciaceae)
Palamocladium wilkesianum (Sull.) Müll.Hal. [Hoe
1974, Hofmann 1997]
Hypnum wilkesianum Sull.
Meteorium pallidovirens Müll.Hal.
Meteorium sciuroides Müll.Hal.
Palamocladium aptychoides Müll.Hal.
Palamocladium sciurellum Müll.Hal.
Palamocladium wilkesianum (Sull.) Müll.Hal. var.
altisetum Müll.Hal.
Palamocladium wilkesianum (Sull.) Müll.Hal. var.
sciuroides (Müll.Hal.) Wijk & Margad.
Pleuropus aptychoides (Müll.Hal.) Broth.
Pleuropus pallidovirens (Müll.Hal.) Broth.
Pleuropus sciurellus (Müll.Hal.) Broth.
Pleuropus wilkesianus (Sull.) Broth.
Pleuropus wilkesianus (Sull.) Broth. var. sciuroides
(Müll.Hal.) E.B.Bartram
end K O Mo L M H
PELEKIUM Mitt. (Thuidiaceae)
Pelekium versicolor (Hornsch. ex Müll.Hal.) Touw [Hoe
1974, Touw 2001]
Thuidium crenulatum Mitt.
Thuidium nanophyllum Müll.Hal.
ind O M H
PHILONOTIS Brid. (Bartramiaceae)
Philonotis falcata (Hook.) Mitt. [Hoe 1974] ind K O Mo M
Philonotis hastata (Duby) Wijk & Margad. [Hoe 1974]
Philonotis laxissima Mitt.
ind O M H
Philonotis hawaica (Müll.Hal.) Broth. [Hoe 1974] end K O Mo L M H
Philonotis turneriana (Schwägr.) Mitt. var. sullivantii
(Müll.Hal.) E.B.Bartram [Hoe 1974]
end O M H
Philonotis turneriana (Schwägr.) Mitt. var. turneriana
[Hoe 1974]
ind K O Mo L M H

57
TROPICAL BRYOLOGY 25 (2004)
Revised checklist of Hawaiian mosses
FullName STATUS NI K O MO L M KA H
PILOTRICHELLA (Müll.Hal.) Besch. (Meteoriaceae,
or Lembophyllaceae [Shaw & Goffinet 2000])
Pilotrichella mauiensis (Sull.) A.Jaeger [Hoe 1974] end O M H
PLAGIOMNIUM T.J.Kop. (Mniaceae)
Plagiomnium rostratum (Schrad.) T.J.Kop. [Hoe 1974]
Mnium rostratum Schrad.
Plagiomnium prorepens (Müll.Hal.) T.J.Kop.
Plagiomnium rhynchosporum (Hook.) T.J.Kop.
ind K O Mo M H
PLAGIOPUS Brid. (Bartramiaceae)
Plagiopus oederiana (Sw.) H.A.Crum & L.E.Anderson
[Hoe 1974]
Plagiopus longisetus (Brid.) E.B.Bartram
Plagiopus oederi (Brid.) Limpr.
ind M
PLAGIOTHECIUM Bruch & Schimp.
(Plagiotheciaceae)
Plagiothecium cavifolium (Brid.) Z.Iwats. [Hoe 1977] ind M
Plagiothecium denticulatum (Hedw.) Schimp. [Hoe
1974]
ind
Plagiothecium mauiense Broth. [Hoe 1974] end M
PLATYHYPNIDIUM M.Fleisch. (Amblystegiaceae, or
Brachytheciaceae [Shaw & Goffinet 2000])
Platyhypnidium muelleri (A.Jaeger) M.Fleisch. [Hoe
1974]
Eurhynchium mulleri misapplied
ind K O Mo M
POGONATUM P. Beauv. (Polytrichaceae)
Pogonatum tahitense Schimp. ex Besch. [Hoe 1974]
Pogonatum baldwinii (Müll.Hal.) Paris
ind K O Mo L M H
POHLIA Hedw. (Bryaceae, or Mniaceae [Shaw &
Goffinet 2000])
Pohlia baldwinii (Broth. ex E.B.Bartram) W.Schultze-
Motel [Hoe 1974]
Webera baldwinii Broth. ex E.B.Bartram
end M
Pohlia cruda (Hedw.) Lindb. [Hoe 1974, Hoe 1979]
Webera cruda (Hedw.) Fürnr.
ind M H
Pohlia flexuosa Hook. [Hoe 1974, Gangulee 1974]
Pohlia gracilescens (E.B.Bartram) E.B.Bartram
Pohlia gracillima (Cardot) Horik. & Ochi
Pohlia leucostoma (Bosch & Sande Lac.) M.Fleisch.
Pohlia leucostomoides (Broth.) E.B.Bartram
Webera gracilescens E.B.Bartram
Webera leucostomoides Broth.
ind K O Mo M H
Pohlia mauiensis (Broth. ex E.B.Bartram) W.Schultze-
Motel [Hoe 1974]
end M H

58
TROPICAL BRYOLOGY 25 (2004)
Staples et al.
FullName STATUS NI K O MO L M KA H
POLYTRICHUM Hedw. (Polytrichaceae)
Polytrichum commune Hedw. [Hoe 1974] ind H
Polytrichum juniperinum Hedw. [Hoe 1974, Hyvönen
1986]
ind H
Polytrichum piliferum Hedw. [Hoe 1974] ind M H
PSEUDOSCLEROPODIUM (Limpr.) M.Fleisch.
(Brachytheciaceae)
Pseudoscleropodium purum (Hedw.) M.Fleisch. [Hoe
1974]
nat H
PSEUDOSYMBLEPHARIS Broth. (Pottiaceae)
Pseudosymblepharis angustata (Mitt.) Hilp. [Hoe 1974,
Hoe 1979, Norris & Koponen 1989]
Pseudosymblepharis mauiensis (Mull.Hal.) Broth.
ind K O Mo M H
PSEUDOTAXIPHYLLUM Z.Iwats. (Hypnaceae)
Pseudotaxiphyllum elegans (Brid.) Z.Iwats. [Hoe 1979,
Iwatsuki 1987]
Isopterygium elegans (Brid.) Lindb.
ind M
PTEROGONIDIUM Broth. (Sematophyllaceae)
Pterogonidium pulchellum (Hook.) Müll.Hal. [Hoe 1979] nat? O
PTYCHOMITRIUM Fürnr. (Ptychomitriaceae)
Ptychomitrium lepidomitrium (Müll.Hal.) Schimp. ex
Besch. [Hoe 1974, Cao et al. 2003]
Ptychomitrium mauiense Broth.
ind M H
PYLAISIADELPHA Broth. (Meteoriaceae)
Pylaisiadelpha opaeodon (Sull.) W.R. Buck [Hoe 1974,
Buck 1984]
Brotherella opaeodon (Sull.) Broth.
Hypnum opaeodon Sull.
ind K O Mo M H
PYRRHOBRYUM Mitt. (Rhizogoniaceae)
Pyrrhobryum spiniforme (Hedw.) Mitt. [Hoe 1974,
Koponen et al. 1986]
Rhizogonium spiniforme (Hedw.) Bruch
ind K O Mo L M H
RACOMITRIUM Brid. (Grimmiaceae)
Racomitrium crispulum (Hook.f. & Wilson) Hook.f. &
Wilson [Hoe 1974]
ind K M H
Racomitrium fasciculare (Schrad. ex Hedw.) Brid. var.
erosum Broth. [Hoe 1974]
end M
Racomitrium lanuginosum (Hedw.) Brid. var.
lanuginosum [Hoe 1974]
ind M

59
TROPICAL BRYOLOGY 25 (2004)
Revised checklist of Hawaiian mosses
FullName STATUS NI K O MO L M KA H
Racomitrium lanuginosum (Hedw.) Brid. var.
pruinosum Wilson [Hoe 1974]
Racomitrium lanuginosum (Hedw.) Brid. var.
sandvicense Reichardt
ind K O Mo M H
RACOPILUM P.Beauv. (Racopilaceae)
Racopilum cuspidigerum (Schwägr.) Ångstr. [Hoe 1974]
Rhacomitrium cuspidigerum misapplied
nat? K O Mo L M H
RADULINA W.R.Buck & B.C.Tan (Sematophyllaceae)
Radulina hamata (Dozy & Molk.) W.R.Buck & B.C.Tan
[Hoe 1974, Buck & Tan 1989]
Trichosteleum hamatum (Dozy & Molk.) A.Jaeger
ind K O Mo L M H
REMYELLA Müll.Hal. (Brachytheciaceae)
Remyella hawaica Müll.Hal. [Hoe 1974, Ignatov &
Huttunen 2002]
Rhynchostegiella hawaica (Müll.Hal.) Broth.
end O
RHABDOWEISIA Bruch & Schimp. (Dicranaceae, or
Rhabdoweisiaceae [Shaw & Goffinet 2000])
Rhabdoweisia crenulata (Mitt.) H.Jameson [Hoe 1975,
Frahm et al. 2000]
ind M
Rhabdoweisia crispata (With.) Lindb. [Hoe 1974, Hoe
1977, Hoe 1979]
Rhabdoweisia denticulata B.S.G.
ind M H
RHAPHIDOSTICHUM M.Fleisch. (Sematophyllaceae)
Rhaphidostichum pustulatum Hoe [Hoe 1974] end O
RHIZOGONIUM Brid. (Rhizogoniaceae)
Rhizogonium pungens Sull. [Hoe 1974] end K O Mo L M H
RHODOBRYUM (Schimp.) Limpr. (Bryaceae)
Rhodobryum giganteum (Schwägr.) Paris [Hoe 1974,
Crosby et al. 1983, Tan & Iwatsuki 1991]
Bryum giganteum (Schwägr.) Arn.
ind K O M
RHYNCHOSTEGIUM Bruch & Schimp.
(Brachytheciaceae)
Rhynchostegium celebicum (Sande Lac.) A.Jaeger [Hoe
1974, Hoe 1977]
Eurhynchium celebicum (Sande Lac.) E.B.Bartram
ind K O Mo M
Rhynchostegium gaudichaudii (Mont.) A.Jaeger [Hoe
1974]
ind
Rhynchostegium selaginellifolium Müll.Hal. var.
recurvirameum (Müll.Hal.) Wijk & Margad. [Hoe
1974]
Eurhynchium selaginellifolium var. recurvirameum
end K O H

60
TROPICAL BRYOLOGY 25 (2004)
Staples et al.
FullName STATUS NI K O MO L M KA H
(Müll.Hal.) E.B.Bartram
Rhynchostegium selaginellifolium Müll.Hal. var.
selaginellifolium [Hoe 1974]
Eurhynchium selaginellifolium (Müll.Hal.)
E.B.Bartram
end K O Mo M H
ROSULABRYUM J.R.Spence (Bryaceae)
Rosulabryum billardierei (Schwägr.) J.R.Spence
[Mohamed 1979, Anderson et al. 1990, Spence 1996]
Bryum andicola Hook.
Bryum decaisnei Dozy & Molk.
Bryum limbatomarginatum Müll.Hal.
Bryum truncorum misapplied
ind O M H
Rosulabryum capillare (Hedw.) J.R.Spence [Hoe 1974,
Hoe 1977, Spence 1996]
Bryum capillare Hedw.
Bryum vinoviride E.B.Bartram
ind K O M H
SAELANIA Lindb. (Ditrichaceae)
Saelania glaucescens (Hedw.) Broth. [Hoe 1974] ind M
SCHISTIDIUM Bruch & Schimp. (Grimmiaceae)
Schistidium apocarpum (Hedw.) Bruch & Schimp.
[Schultze-Motel 1963, Muñoz & Pando 2000]
Grimmia apocarpa Hedw.
ind M
Schistidium strictum (Turner) Loeske ex Mårtensson
[Hoe 1974, Cao & Churchill 1995, Muñoz & Pando
2000]
Grimmia apocarpa Hedw. var. stricta (Turner) Hook.
& Taylor
Grimmia scabrifolia Broth.
Grimmia stricta Turner
ind M
SCHIZYMENIUM Harv. (Bryaceae, or Mniaceae [Shaw
& Goffinet 2000])
Schizymenium campylocarpum (Hook. & Arn.) A.J.Shaw
var. campylocarpum [Zanten 1971, Shaw 1985]
Mielichhoferia campylocarpa (Hook. & Arn.) Mitt.
Weissia campylocarpa Hook. & Arn. in Hook.
ind H
Schizymenium nealiae (E.B.Bartram) A.J.Shaw [Hoe
1974, Shaw 1985]
Mielichhoferia nealiae E.B.Bartram
end H
Schizymenium pulvinatum (Müll.Hal.) A.J.Shaw [Hoe
1974, Shaw 1985]
Mielichhoferia pulvinata Müll.Hal.
end M H
SCOPELOPHILA (Mitt.) Lindb. (Pottiaceae)
Scopelophila infericola Hoe [Hoe 1974] end H

61
TROPICAL BRYOLOGY 25 (2004)
Revised checklist of Hawaiian mosses
FullName STATUS NI K O MO L M KA H
SEMATOPHYLLUM Mitt. (Sematophyllaceae)
Sematophyllum hawaiiense (Broth.) Broth. [Hoe 1974,
Hoe 1979]
ind K O L M H
Sematophyllum subpinnatum (Brid.) E.Britton [Hoe
1974, Hoe 1979, Buck 1983]
Sematophyllum caespitosum misapplied
ind O H
SPHAGNUM L. (Sphagnaceae)
Sphagnum palustre L. [Hoe 1978, Karlin & Andrus
1995]
Sphagnum henryense misapplied
Sphagnum lonchocladum Müll.Hal.
Sphagnum perichaetiale misapplied
Sphagnum vitjianum misapplied
ind H
Sphagnum wheeleri Müll.Hal. [Hoe 1974, Karlin &
Andrus 1995, Karlin 2001]
Sphagnum compactum misapplied
Sphagnum rigidulum Warnst.
Sphagnum vulcanicum Warnst.
end M
SPLACHNOBRYUM Müll.Hal. (Splachnobryaceae, or
Pottiaceae [Shaw & Goffinet 2000])
Splachnobryum obtusum (Brid.) Müll.Hal. [Hoe 1974,
Arts 2001]
nat O
STREPTOPOGON Mitt. (Pottiaceae)
Streptopogon erythrodontus (Taylor) Wilson [Hoe 1974] ind M
SYMPHYODON Mont. (Symphyodontaceae)
Symphyodon pygmaeus (Broth.) S.He & Snider [Hoe
1974, Hoe 1977, He & Snider 1992]
Glossadelphus abortivapicus Hoe
ind O M
SYNTRICHIA Brid. (Pottiaceae)
Syntrichia fragilis (Taylor) Ochyra [Hoe 1974, Kürschner
2000]
Tortula alpina (Brid.) Arn. var. inermis (Milde) De
Not.
Tortula fragilis Taylor
ind M H
Syntrichia princeps (De Not.) Mitt. [Hoe 1974,
Kürschner 2000]
Tortula princeps De Not.
ind M H
SYRRHOPODON Schwägr. (Calymperaceae)
Syrrhopodon armatus Mitt. [Hoe 1974, Orbán & Reese
1986]
Syrrhopodon oahuensis Broth.
ind O M H
Syrrhopodon hawaiicus Müll.Hal. [Hoe 1974] end K O Mo M H
Syrrhopodon prolifer Schwägr. [Hoe 1974, Reese et al. end O Mo M H

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FullName STATUS NI K O MO L M KA H
1986]
Syrrhopodon kilaueae Müll.Hal.
Syrrhopodon semiliber (Mitt.) Besch. [Hoe 1974, Reese
et al. 1986]
Calymperes semiliberum Mitt.
Calymperopsis semiliber (Mitt.) M.Fleisch.
ind O H
TAXIPHYLLUM M.Fleisch. (Hypnaceae)
Taxiphyllum laevifolium (Mitt.) W.R.Buck [Hoe 1974,
Buck 1987b, Tixier 1988]
Glossadelphus baldwinii Broth.
Glossadelphus laevifolius (Mitt.) E.B.Bartram
ind K O Mo M H
Taxiphyllum torrentium (Besch.) W.R.Buck [Hoe 1974,
Buck 1987b, Tixier 1988]
Glossadelphus irroratus E.B.Bartram
Glossadelphus torrentium (Besch.) M.Fleisch.
ind M
TAXITHELIUM Mitt. (Sematophyllaceae)
Taxithelium mundulum (Sull.) E.B.Bartram [Hoe 1974]
Taxithelium undulatum misapplied
ind K O Mo L M H
TAYLORIA Hook. (Splachnaceae)
Tayloria sandwicensis (Müll.Hal.) Broth. [Hoe 1974] end M
THAMNIOPSIS (Mitt.) Fleisch. (Hookeriaceae, or
Callicostaceae [Buck 1987a])
Thamniopsis utacamundiana (Mont.) W.R Buck [Hoe
1974, Buck 1987a]
Hookeria purpurea Müll.Hal.
Hookeriopsis purpurea (Müll.Hal.) Broth.
Hookeriopsis purpurea (Müll.Hal.) Broth. var.
acuminatula (Müll.Hal.) E.B.Bartram
Hookeriopsis purpurea (Müll.Hal.) Broth. var.
ligulacea (Müll.Hal.) E.B.Bartram
ind K O Mo M H
THAMNOBRYUM Nieuwl. (Neckeraceae)
Thamnobryum speciosum (Broth.) Hoe [Hoe 1974]
Thamnium speciosum Broth.
end M H
THUIDIUM Bruch & Schimp. (Thuidiaceae)
Thuidium cymbifolium (Dozy & Molk.) Dozy & Molk.
[Hoe 1974, Touw 2001]
Thuidium glabratum Broth.
?Thuidium havaiense Reichardt
Thuidium oahuense Broth.
Thuidium plicatum Mitt.
Thuidium plicatum Mitt. var. brevifolium
E.B.Bartram
ind K O Mo L M H

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FullName STATUS NI K O MO L M KA H
TIMMIA Hedw. (Timmiaceae)
Timmia megapolitana Hedw. ssp. bavarica (Hessl.)
Brassard [Brassard 1984]
Timmia bavarica Hessl.
ind H
TORTELLA (Lindb.) Limpr. (Pottiaceae)
Tortella fragilis (Hook. & Wilson) Limpr. var.
tortelloides (S.W.Greene) R.H.Zander & Hoe [Zander
& Hoe 1979]
Sarconeurum tortelloides S.W.Greene
Tortella tortelloides (S.W.Greene) H.Rob.
ind M
Tortella humilis (Hedw.) Jenn. [Hoe 1974]
Tortella caespitosa (Schwägr.) Limpr.
ind K O L M H
Tortella tortuosa (Hedw.) Limpr. [Hoe 1979] ind L
TORTULA Hedw. (Pottiaceae)
Tortula atrovirens (Sm.) Lindb. [Hoe 1974, Hoe 1979,
Corley et al. 1981, Zander 1993]
Desmatodon convolutus (Brid.) Grout
ind M H
TRACHYLOMA Brid. (Pterobryaceae, or
Trachylomataceae [Shaw & Goffinet 2000])
Trachyloma indicum Mitt. [Hoe 1974]
Trachyloma tahitense Besch.
ind M
TRACHYPODOPSIS M.Fleisch. (Trachypodaceae, or
Meteoriaceae [Shaw & Goffinet 2000])
Trachypodopsis auriculata (Mitt.) M.Fleisch. [Hoe 1974]
Trachypodopsis ornans (Reichardt) M.Fleisch.
Trachypus ornans (Reichardt) Müll.Hal. fma.
simplex Broth.
ind K O Mo M H
Trachypodopsis serrulata (P.Beauv.) M.Fleisch. var.
crispatula (Hook.) Zanten [Hoe 1979]
ind H
TRACHYPUS Reinw. & Hornsch. (Trachypodaceae, or
Meteoriaceae [Shaw & Goffinet 2000])
Trachypus bicolor Reinw. & Hornsch. [Hoe 1974, Hoe
1979]
ind K M H
Trachypus humilis Lindb. var. flagellifer (Broth. ex
Herzog) H.A.Mill. [Zanten 1959, Miller 1967, Hoe
1974]
Trachypus humilis Lindb. var. tenerrimus (Herzog)
Zanten
Trachypus mauiensis Broth.
ind K O M
Trachypus humilis Lindb. var. humilis [Hoe 1974, Hoe
1977]
ind K M
TREMATODON Michx. (Dicranaceae, or Bruchiaceae
[Shaw & Goffinet 2000])

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FullName STATUS NI K O MO L M KA H
Trematodon latinervis Müll.Hal. [Hoe 1974]
Trematodon longicollis misapplied
ind K O Mo M H
TRICHOSTOMUM Bruch (Pottiaceae)
Trichostomum brachydontium Bruch [Sollman 1984]
Trichostomum oblongifolium E.B.Bartram
ind K O Mo M H
Trichostomum crispulum Bruch [Hoe 1974, Hoe 1975,
Norris & Koponen 1989]
Trichostomum bartramii H.A.Mill.
Trichostomum mauiense Broth.
ind K O Mo L M H
ULOTA D.Mohr (Orthotrichaceae)
Ulota cervina Hoe & H.A.Crum [Hoe 1974] end M
VESICULARIA (Müll.Hal.) Müll.Hal. (Hypnaceae)
Vesicularia inflectens (Brid.) Müll.Hal. [Hoe 1979] ind M
Vesicularia perviridis (Ångstr.) Müll.Hal. [Hoe 1974]
Vesicularia graminicolor Müll.Hal.
end K O Mo L M H
WEISSIA Hedw. (Pottiaceae)
Weissia controversa Hedw. [Hoe 1974, Wichman & St.
John 1990]
Weisia viridula Hedw.
ind Ni K O H
Weissia ovalis (R.S.Williams) E.B.Bartram [Hoe 1974] end K O Mo L M H
ZYGODON Hook. & Taylor (Orthotrichaceae)
Zygodon reinwardtii (Hornsch.) A.Braun [Hoe 1974] ind M
Zygodon rubrigemmius (Hoe & H.A.Crum) R.H.Zander
& Vitt [Hoe 1974, Zander & Vitt 1979]
Anoectangium rubrigemmium Hoe & H.A.Crum
end M
Zygodon tetragonostomus A.Braun ex B.S.G. [Hoe 1974] ind K O M H

65
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CROSS-REFERENCED LIST OF SYNONYMS AND MISAPPLIED NAMES
Acroporium fuscoflavum var. baldwinii See Acroporium fuscoflavum
?Acroporium sigmatodontium misapplied See Acroporium fuscoflavum
Aerobryopsis longissima See Aerobryopsis wallichii
Aerobryopsis longissima var. dozyana See Aerobryopsis wallichii
Aerobryopsis scariosa See Aerobryopsis subdivergens ssp. scariosa
Amphidium cyathicarpum See Amphidium tortuosum
Andreaea acuminata See Andreaea acutifolia ssp. acuminata
Andreaea perpapillosa See Andreaea rupestris
Anoectangium euchloron See Anoectangium aestivum
Anoectangium haleakalae var. haleakalae See Anoectangium aestivum
Anoectangium haleakalae var. laxulum See Anoectangium aestivum
Anoectangium harttiae See Anoectangium aestivum
Anoectangium rubrigemmium See Zygodon rubrigemmius
Aptychella robusta See Clastobryopsis robusta
Barbella trichophora See Barbellopsis trichophora
Barbula cruegeri See Barbula indica
Barbula unguiculata misapplied See Barbula indica
Barbula vinealis var. cylindrica See Didymodon insulanus
Barbula vinealis var. flaccida See Didymodon insulanus
Brachythecium oxyrrhynchum See Brachythecium plumosum
Breutelia kilaueae See Breutelia affinis
Brotherella opaeodon See Pylaisiadelpha opaeodon
Bryohaplocladium microphyllum See Haplocladium microphyllum
Bryum andicola Hook. See Rosulabryum billardierei
Bryum apiculatum See Bryum mildeanum
Bryum argenteum var. lanatum See Bryum argenteum
Bryum capillare See Rosulabryum capillare
Bryum ceramiocarpum See Bryum conicum
Bryum crassicostatum See Bryum hawaiicum
Bryum decaisnei See Rosulabryum billardierei
Bryum erythrocarpum See Bryum atrovirens
Bryum giganteum See Rhodobryum giganteum
Bryum leucophyllum See Bryum argenteum
Bryum limbatomarginatum See Rosulabryum billardierei
Bryum mauiensis See Bryum mildeanum
Bryum megalostegium See Bryum mildeanum
Bryum nitens See Bryum mildeanum
Bryum truncorum misapplied See Rosulabryum billardierei
Bryum vinoviride See Rosulabryum capillare
Calymperes hawaiiense See Calymperes fasciculatum
Calymperes semiliberum See Syrrhopodon semiliber
Calymperopsis semiliber See Syrrhopodon semiliber
Campylopodium euphorocladum See Campylopodium medium
Campylopus aureus See Campylopus schmidii ssp. schmidii
Campylopus boswellii See Campylopus fragilis ssp. zollingerianus
Campylopus boswellii var. capitulatus See Campylopus fragilis ssp. zollingerianus
Campylopus densifolius var. densifolius See Campylopus hawaiicus var. densifolius
Campylopus densifolius var. falcatus See Campylopus hawaiicus var. hawaiicoflexuosus

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Campylopus densifolius var. hawaiicoflexuosus See Campylopus hawaiicus var. hawaiicoflexuosus
Campylopus densifolius var. purpureoflavescens See Campylopus hawaiicus var. hawaiicus
Campylopus densiretis See Campylopus hawaiicus var. densifolius
?Campylopus didrichsenii See Campylopus umbellatus var. umbellatus
Campylopus exasperatus See Campylopus praemorsus
Campylopus fumarioli See Campylopus laxitextus
Campylopus geniculatus See Campylopus umbellatus var. pterotoneuron
Campylopus hawaiicoflexuosus See Campylopus hawaiicus var. hawaiicoflexuosus
Campylopus introflexus misapplied See Campylopus schmidii ssp. schmidii
Campylopus introflexus var. polytrichoides misapplied See Campylopus schmidii ssp. schmidii
Campylopus microcephalus See Campylopus praemorsus
Campylopus perangustifolius See Campylopus hawaiicus var. hawaiicoflexuosus
?Campylopus pertristis See Campylopus umbellatus var. umbellatus
Campylopus polytrichoides misapplied See Campylopus schmidii ssp. schmidii
Campylopus purpureoflavescens, in part See Campylopus hawaiicus var. hawaiicoflexuosus, C.
hawaiicus var. hawaiicus
Campylopus richardii misapplied See Campylopus umbellatus var. umbellatus
Campylopus sandwicensis See Campylopus umbellatus var. umbellatus
Campylopus skottsbergii See Campylopus schmidii ssp. hemitrichius
Campylopus tubulosus See Campylopus schmidii ssp. hemitrichius
Campylopus wawreanus See Campylopus hawaiicus var. densifolius
Ceratodon purpureus fma. xanthopus See Ceratodon purpureus ssp. purpureus
Cirriphyllum oxyrrhynchum See Brachythecium plumosum
Claopodium amblystegioides See Claopodium prionophyllum
Claopodium hawaiiense See Claopodium whippleanum
Claopodium nervosum See Claopodium prionophyllum
Ctenidium decurrens See Ectropothecium decurrens
Ctenidium pulcherrimum See Ectropothecium decurrens
Ctenidium submalacobolum See Ectropothecium decurrens
Cupressina decurrens See Ectropothecium decurrens
Desmatodon convolutus See Tortula atrovirens
Dicranella hillebrandii See Microdus hillebrandii
Dicranella hochreuteri misapplied See Dicranella hochreutineri
Dicranodontium falcatum var. falcatum See Dicranodontium porodictyon
Dicranodontium hawaiicum See Dicranodontium porodictyon
Dicranoloma wheeleri See Campylopus wheeleri
Dicranum didrichsenii See Campylopus umbellatus var. umbellatus
Dicranum hawaiicoflexuosum See Campylopus hawaiicus var. hawaiicoflexuosus
Dicranum hawaiicum See Campylopus hawaiicus var. hawaiicus
Dicranum microcephalum See Campylopus praemorsus
Dicranum pertriste See Campylopus umbellatus var. umbellatus
Dicranum praemorsum See Campylopus praemorsus
Dicranum skottsbergii See Campylopus schmidii ssp. hemitrichius
Dicranum wheeleri See Campylopus wheeleri
Didymodon recurvirostris See Bryoerythrophyllum recurvirostre
Didymodon vinealis var. flaccidus See Didymodon insulanus
Ectropothecium rockii See Ectropothecium decurrens
Encalypta rhaptocarpa misapplied See Encalypta rhabdocarpa
Encalypta scabrata See Encalypta ciliata
Entodon hillebrandii See Entodon plicatus

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Entodon solanderi See Entodon plicatus
Eurhynchium celebicum See Rhynchostegium celebicum
Eurhynchium muelleri misapplied See Platyhypnidium muelleri
Eurhynchium selaginellifolium See Rhynchostegium selaginellifolium var. selaginellifolium
Eurhynchium selaginellifolium var. recurvirameum See Rhynchostegium selaginellifolium var.
recurvirameum
Eurhynchium vagans See Oxyrrhynchium vagans
Fissidens baldwinii See Fissidens elegans
Fissidens bishopii See Fissidens elegans
Fissidens crumii See Fissidens lindbergii
Fissidens hawaiicus See Fissidens bryoides
Fissidens insularis See Fissidens bryoides
Fissidens intermedius See Fissidens submarginatus
Fissidens mauiensis See Fissidens pacificus
Fissidens oahuensis See Fissidens bryoides
Floribundaria baldwinii See Floribundaria floribunda
Funaria subintegra See Entosthodon subintegrus
Garovaglia haleakalae See Garovaglia powellii var. haleakalae
Glossadelphus abortivapicus See Symphyodon pygmaeus
Glossadelphus baldwinii See Taxiphyllum laevifolium
Glossadelphus chrysobasilaris See Glossadelphus ogatae
Glossadelphus irroratus See Taxiphyllum torrentium
Glossadelphus laevifolius See Taxiphyllum laevifolium
Glossadelphus torrentium See Taxiphyllum torrentium
Glossadelphus zollingeri var. filicaulis See Ectropothecium zollingeri var. filicaule
Grimmia apocarpa See Schistidium apocarpum
Grimmia apocarpa var. pulvinata See Grimmia anodon
Grimmia apocarpa var. stricta See Schistidium strictum
Grimmia haleakalae See Amphidium tortuosum
Grimmia haliacalae See Amphidium tortuosum
Grimmia pilfera See Grimmia longirostris
Grimmia scabrifolia See Schistidium strictum
Grimmia stricta See Schistidium strictum
Gymnostoma pulvinata See Grimmia anodon
Hageniella pacifica See Hageniella micans
Holomitrium ferriei misapplied See Holomitrium squarrifolium
Homalia densa See Homaliodendron flabellatum
Homalia praelonga See Homaliodendron flabellatum
Hookeria purpurea See Thamniopsis utacamundiana
Hookeriopsis purpurea See Thamniopsis utacamundiana
Hookeriopsis purpurea var. acuminatula See Thamniopsis utacamundiana
Hookeriopsis purpurea var. ligulacea See Thamniopsis utacamundiana
Hymenostylium firmum See Hymenostylium recurvirostrum var. recurvirostrum
Hyophila dozy-molkenboeri See Hyophila involuta
Hypnum decurrens See Ectropothecium decurrens
Hypnum draytoni(i) See Buckiella draytonii
Hypnum eudorae See Buckiella draytonii
Hypnum opaeodon See Pylaisiadelpha opaeodon
Hypnum wilkesianum See Palamocladium wilkesianum
Isopterygium elegans See Pseudotaxiphyllum elegans

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TROPICAL BRYOLOGY 25 (2004)
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Leptodontium brevicaule See Leptodontium flexifolium
Leucobryum hawaiense See Leucobryum seemannii var. seemannii
Leucobryum hawaiense var. fumarioli See Leucobryum seemannii var. fumarioli
Leucobryum hawaiense var. solfatarae See Leucobryum seemannii var. solfatarae
Leucobryum solfatarae See Leucobryum seemannii var. solfatarae
Leucobryum solfatarae var. fumarioli See Leucobryum seemannii var. fumarioli
Leucobryum solfatarae var. hawaiense See Leucobryum seemannii var. seemannii
Macrocoma intricata See Macrocoma tenue ssp. sullivantii
Macromitrium cumingii See Macromitirum incurvifolium
Macromitrium intricatum See Macrocoma tenue ssp. sullivantii
Macromitrium owahiense See Macromitrium microstomum
Macromitrium reinwardtii See Macromitrium microstomum
Merceyopsis crassinervis See Hymenostylium recurvirostrum var. cylindricum
Meteorium pallidovirens See Palamocladium wilkesianum
Meteorium sciuroides See Palamocladium wilkesianum
Microdus hawaiicus See Dicranella hawaiica var. hawaiica
Mielichhoferia campylocarpa See Schizymenium campylocarpum var. campylocarpum
Mielichhoferia nealiae See Schizymenium nealiae
Mielichhoferia pulvinata See Schizymenium pulvinatum
Mniadelphus freycineti var. crasseturgescens See Distichophyllum freycinetii var.
crasseturgescens
Mnium rostratum See Plagiomnium rostratum
Mnium serratum See Mnium marginatum
Molendoa crassinervis See Hymenostylium recurvirostrum var. cylindricum
Orthotrichum berggrenii See Orthotrichum alpestre var. berggrenii
Orthotrichum hillebrandii See Orthotrichum rupestre
Orthotrichum verrucatum See Orthotrichum lyellii
Palamocladium aptychoides See Palamocladium wilkesianum
Palamocladium sciurellum See Palamocladium wilkesianum
Palamocladium wilkesianum var. altisetum See Palamocladium wilkesianum
Palamocladium wilkesianum var. sciuroides See Palamocladium wilkesianum
Philonotis laxissima See Philonotis hastata
Plagiomnium prorepens See Plagiomnium rostratum
Plagiomnium rhynchosporum See Plagiomnium rostratum
Plagiopus longisetus See Plagiopus oederiana
Plagiopus oederi See Plagiopus oederiana
Plagiothecium draytonii See Buckiella draytonii
Pleuropus aptychoides See Palamocladium wilkesianum
Pleuropus pallidovirens See Palamocladium wilkesianum
Pleuropus sciurellus See Palamocladium wilkesianum
Pleuropus wilkesianus See Palamocladium wilkesianum
Pleuropus wilkesianus var. sciuroides See Palamocladium wilkesianum
Pogonatum baldwinii See Pogonatum tahitense
Pohlia gracilescens See Pohlia flexuosa
Pohlia gracillima See Pohlia flexuosa
Pohlia leucostoma See Pohlia flexuosa
Pohlia leucostomoides See Pohlia flexuosa
Pseudosymblepharis mauiensis See Pseudosymblepharis angustata
Ptychomitrium mauiense See Ptychomitrium lepidomitrium
Ptychomnion aciculare misapplied See Aerobryopsis subdivergens ssp. scariosa

69
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Racomitrium lanuginosum var. sandvicense See Racomitrium lanuginosum var. pruinosum
Rhabdoweisia denticulata See Rhabdoweisia crispata
Rhacomitrium cuspidigerum misapplied See Racopilum cuspidigerum
Rhizogonium spiniforme See Pyrrhobryum spiniforme
Rhynchostegiella hawaica See Remyella hawaica
Rhynchostegium vagans See Oxyrrhynchium vagans
Sarconeurum tortelloides See Tortella fragilis var. tortelloides
Sciaromium tricostatum See Limbella tricostata
Sematophyllum caespitosum See Sematophyllum subpinnatum
Sphagnum compactum misapplied See Sphagnum wheeleri
Sphagnum henryense misapplied See Sphagnum palustre
Sphagnum lonchocladum See Sphagnum palustre
Sphagnum perichaetiale misapplied See Sphagnum palustre
Sphagnum rigidulum See Sphagnum wheeleri
Sphagnum vitjianum misapplied See Sphagnum palustre
Sphagnum vulcanicum See Sphagnum wheeleri
Stereodon decurrens See Ectropothecium decurrens
Stereophyllum oahuense See Entodontopsis anceps
Syrrhopodon kilaueae See Syrrhopodon prolifer
Syrrhopodon oahuensis See Syrrhopodon armatus
Taxiphyllum gracile See Ectropothecium decurrens
Taxithelium undulatum misapplied See Taxithelium mundulum
Thamnium speciosum See Thamnobryum speciosum
Thuidium crenulatum See Pelekium versicolor
Thuidium glabratum See Thuidium cymbifolium
?Thuidium havaiense See Thuidium cymbifolium
Thuidium nanophyllum See Pelekium versicolor
Thuidium oahuense See Thuidium cymbifolium
Thuidium plicatum See Thuidium cymbifolium
Thuidium plicatum var. brevifolium See Thuidium cymbifolium
Thysanomitrion hawaiicum See Campylopus umbellatus var. umbellatus
Thysanomitrion umbellatum See Campylopus umbellatus var. umbellatus
Timmia bavarica See Timmia megapolitana ssp. bavarica
Tortella caespitosa See Tortella humilis
Tortella tortelloides See Tortella fragilis var. tortelloides
Tortula alpina var. inermis See Syntrichia fragilis
Tortula fragilis See Syntrichia fragilis
Tortula princeps See Syntrichia princeps
Tortula rhizophylla See Chenia leptophylla
Tortula vectensis See Chenia leptophylla
Trachyloma tahitense See Trachyloma indicum
Trachypodopsis ornans See Trachypodopsis auriculata
Trachypus humilis var. tenerrimus See Trachypus humilis var. flagellifer
Trachypus mauiensis See Trachypus humilis var. flagellifer
Trachypus ornans fma. simplex See Trachypodopsis auriculata
Trematodon longicollis misapplied See Trematodon latinervis
Trichosteleum hamatum See Radulina hamata
Trichostomum bartramii See Trichostomum crispulum
Trichostomum mauiense See Trichostomum crispulum
Trichostomum oblongifolium See Trichostomum brachydontium

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Trichostomum tenuirostre See Oxystegus tenuirostris var. gemmiparus
Vesicularia graminicolor See Vesicularia perviridis
Webera baldwinii See Pohlia baldwinii
Webera cruda See Pohlia cruda
Webera gracilescens See Pohlia flexuosa
Webera leucostomoides See Pohlia flexuosa
Weisia viridula See Weissia controversa
Weissia campylocarpa See Schizymenium campylocarpum var. campylocarpum