Checklist of the shore fishes of Europa Island, Mozambique Channel, southwestern Indian Ocean, including 302 new records

Abstract

An annotated checklist of the fish species of Europa Island (Mozambique Channel, southwestern Indian Ocean) comprises a total of 389 species in 62 families. 302 species are recorded from Europa Island for the first time. All species are autochthonous; no introduced species have been found. The fish fauna is exclusively marine, with the Labridae, Pomacentridae, Serranidae, Gobiidae and Acanthuridae being the families with most representatives.

The fish fauna at Europa Island is typical for offshore, low islands in the southwestern Indian Ocean. Zoogeo¬graphically, the main element of the fish fauna of Europa Island consists of widespread tropical Indo-Pacific species (292 species, 75.1 % of he total occurring species). A total of 13 species (3.3 %) are found worldwide, either cir¬cumtropical or circumtropical including warm temperate zones. An additional 76 species (19.5 %) are Indian Ocean endemics, including 36 western Indian Ocean endemics (9.2 %), and 10 southwestern Indian Ocean endemics (2.6 %). No endemic fish species are reported for Europa Island.

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Stuttgarter Beiträge zur Naturkunde A, Neue Serie 6: 247–276; Stuttgart, 30.IV.2013
1 Introduction
Europa Island is situated in the southern part of the Mo-
zambique Channel, southwestern Indian Ocean at 22°22'S
40°22'E, approximately 355 km westnorthwest of Toliara
(Madagascar), and 529 km eastnortheast of Inhambane
(Mozambique). The island was named after the British ship
‘Europa’ which visited in 1774. As one of the Îles Éparses,
it has been administered by France since 1897, and is part
of the Terres australes et antarctiques françaises (TAAF)
since 2005. A small settlement existed since ca. 1860 but
was abandoned in the 1920s. Today the island is uninhab-
ited except for a small garrison hosting a detachment of the
French army, and a French gendarme. There is no harbour
available, only a boat mooring off the northwestern fring-
ing reef.
The island measures 7 by 6 km (28 km2). It is a for-
mer atoll which was uplifted to a maximum altitude of
7 m, leaving the fossil coral reefs dry. Nowadays it is sur-
rounded by a fringing reef with a narrow and shallow la-
goon, which only widens towards the north of the island,
where there is also the entrance of an extensive inland la-
goon system which is surrounded by mangroves on its in-
ner side. Several typical coastal marine habitats are shown
in Figs. 2–5.
The knowledge of the fish fauna of Europa Island was
hitherto limited to a publication by Fourmanoir (1952),
who recorded 99 fish species from the island.
Checklist of the shore fishes of Europa Island, Mozambique
Channel, southwestern Indian Ocean, including 302 new records
ronald Fricke, Patrick durville, Giacomo Bernardi, PhiliPPe Borsa, Gérard mou-tham &
Pascale chaBanet
Abstract
An annotated checklist of the fish species of Europa Island (Mozambique Channel, southwestern Indian Ocean)
comprises a total of 389 species in 62 families. 302 species are recorded from Europa Island for the first time. All
species are autochthonous; no introduced species have been found. The fish fauna is exclusively marine, with the
Labridae, Pomacentridae, Serranidae, Gobiidae and Acanthuridae being the families with most representatives.
The fish fauna at Europa Island is typical for offshore, low islands in the southwestern Indian Ocean. Zoogeo-
graphically, the main element of the fish fauna of Europa Island consists of widespread tropical Indo-Pacific species
(292 species, 75.1 % of the total occurring species). A total of 13 species (3.3 %) are found worldwide, either cir-
cumtropical or circumtropical including warm temperate zones. An additional 76 species (19.5 %) are Indian Ocean
endemics, including 36 western Indian Ocean endemics (9.2 %), and 10 southwestern Indian Ocean endemics (2.6 %).
No endemic fish species are reported for Europa Island.
K ey wo rd s: Checklist, Pisces, southwestern Indian Ocean, Europa Island, new records, zoogeography.
Zusammenfassung
Die Checkliste der Fische der Insel Europa (Kanal von Mosambik, südwestlicher Indischer Ozean) enthält 389
Arten in 62 Familien. 302 Arten werden zum ersten Mal von der Insel Europa beobachtet. Alle Fischarten sind au-
thochthon; es wurden keine durch den Menschen eingeführten oder eingeschleppten Arten gefunden. Alle Fischarten
sind marin; die artenreichsten Fischfamilien sind die Labridae, Pomacentridae, Serranidae, Gobiidae und Acanthu-
ridae.
Die Fischfauna der Insel Europa ist typisch für küstenferne, niedrige Inseln im südwestlichen Indischen Ozean.
Zoogeographisch haben 75,1 % der nachgewiesenen Fischarten eine großräumige indo-pazifische Verbreitung, 3,3 %
der Arten werden in allen tropischen Meeren gefunden, und 19,5 % der Arten sind im Indischen Ozean endemisch,
davon 9,2 % nur im westlichen Indischen Ozean und 2,6 % im südwestlichen Indischen Ozean. Es wurden keine en-
demischen Fischarten bei der Insel Europa gefunden.
Contents
1 Introduction ...........................................................................................................................................................247
2 Methods and Materials ..........................................................................................................................................248
3 Checklist................................................................................................................................................................248
4 Fish fauna of Europa Island ..................................................................................................................................273
5 References .............................................................................................................................................................274

stuttGarter BeiträGe zur naturkunde a
248 Neue Serie 6
The present paper provides an updated checklist of the
shore fishes of the island, based on 1) visual censuses dur-
ing a visit of the R/V ‘Marion Dufresne 2’ (April 2011),
where 243 fish species were observed (P. durville, P.
chaBanet, G. Bernardi, P. Borsa & G. mou-tham), and
2) the mission BioReCIE 2011 (Biodiversity, Resources
and Conservation of coral reefs at Îles Éparses; 7–13 Nov.
2011), where numerous fish species were observed and
collected (P. chaBanet, P. durville, R. Fricke).
Acknowledgments
We would like to thank the administration of the TAAF
(Terres australes et antarctiques françaises), Saint-Pierre, La
Réunion, for research permits and financial support; the ‘Forces
armées de la zone Sud de l’océan Indien’ (FAZSOI), detach-
ment of the ‘2e régiment de parachutistes d’infanterie de marine’,
Saint-Pierre, Réunion, and officers of the French ‘Gendarmerie
natio nale’, for logistic support; the captains and crews of SV
‘Antsiva’, Mahajanga, and R/V ‘Marion Dufresne 2’, Marseille
for support, excellent catering and safe transport of the two expe-
ditions to and from Europa Island. We are grateful to E. BretaGne
(French Army) who provided underwater photographs of fishes
taken at Europa Island, to C. conand (Grenoble) who identified
a sea cucumber, the host of a carapid fish, and to T. alPermann
(SMF) who provided collection equipment and catalogue num-
bers. We also gratefully acknowledge the participation of W.-J.
chen and J.-D. durand of the PHYLIP team (Expedition with
R/V Marion Dufresne, April 2011).
Fig. 1. Europa Island with stations of mission BioReCIE 2011, Nov. 2011. – Solid line represents the coast at high tide, the dotted areas
show the fringing reef.

Fricke et alii, checklist oF the shore Fishes oF euroPa island 249
Tab. 1. Station data of the mission BioReCIE 2011 to Europa Island, Nov. 2011.
Station Area Geographic
Coordinates
Date,
Time
Habitat Depth Observers
EU2 East 22°21'10.476"S
40°23'48.228"E
10 Nov. 2011
12:30–13:30 h
Coral reef slope 13 m Pascale chaBanet, Patrick
durville
EU3 West 22°22'22.8"S
40°19'29.388"E
9 Nov. 2011
09:40–10:40 h
Coral reef slope
(Fig. 4)
11–13 m Pascale chaBanet, Patrick
durville
EU4 South 22°24'14.436"S
40°22'12.936"E
9 Nov. 2011
13:40–14:40 h
Coral reef slope 12 m Pascale chaBanet, Patrick
durville
EU5 North 22°20'28.356"S
40°20'15.468"E
11 Nov. 2011
08:00–09:00 h;
13 Nov. 2011
08:00–09:00
Reef at (Fig. 2) 0–2 m Pascale chaBanet, Patrick
durville, ronald Fricke
EU6 North 22°20'26.268"S
40°20'13.776"E
8 Nov. 2011
10:10–11:10 h
Coral reef slope 8–10 m Pascale chaBanet, Patrick
durville
EU7 North 22°19'45.948"S
40°21'54.288"E
7 Nov. 2011
09:00–10:00 h
Coral reef slope 9–12 m Pascale chaBanet, Patrick
durville
B5 Southwest 22°23'02.436"S
40°20'15"E
11 Nov. 2011
14:15–15:15 h
Coral reef slope 10 m Pascale chaBanet, Patrick
durville
B8 Southeast 22°23'07.008"S
40°23'18.816"E
10 Nov. 2011
09:18–10:18 h
Coral reef slope 11 m Pascale chaBanet, Patrick
durville
B12 Northeast 22°20'07.116"S
40°23'19.968"E
12 Nov. 2011
09:00–10:00 h
Coral reef slope 10 m Pascale chaBanet, Patrick
durville
B14 West 22°21'32.436"S
40°19'46.776"E
11 Nov. 2011
09:15–10:15 h
Coral reef slope 12 m Pascale chaBanet, Patrick
durville
B23 Southeast 22°22'50.87"S
40°23'29.64"E
12 Nov. 2011 Reef at 0–1 m Patrick durville
B25 South 22°23'51.29"S
40°22'15.30"E
12 Nov. 2011 Reef at 0–1 m Patrick durville
B28 West 22°21'34.524"S
40°19'52.32"E
9 Nov. 2011
08:10–09:20 h;
11 Nov. 2011
10:15–11:45 h;
12 Nov. 2011
10:30–12:20 h
Reef at with tidal
pools (Fig. 3)
0–1 m ronald Fricke
B31 North 22°20'25.81"S
40°20'42.75"E
7 Nov. 2011
09:30–10:30 h
Reef at 0–1 m ronald Fricke
B32 North 22°20'21.04"S
40°20'42.35"E
7 Nov. 2011
10:30–11:30 h
Reef at 0–1 m ronald Fricke
B33 North 22°20'24.58"S
40°20'51.88"E
7 Nov. 2011
08:30–09:30 h
Reef at 0–1 m ronald Fricke
B34 North 22°20'16.52"S
40°20'54.51"E
8 Nov. 2011
09:30–11:30 h
Lagoon 0–1 m ronald Fricke
B37 North 22°20'07.00"S
40°21'57.50"E
10 Nov. 2011
08:50–09:50 h
Lagoon 0–1 m ronald Fricke
B49 Centre 22°20'09.19"S
40°21'13.55"E
8 Nov. 2011
11:30–12:30 h
Reef at 0–1 m ronald Fricke
B50 Centre 22°19'59.54"S
40°21'53.67"E
10 Nov. 2011
09:50–10:50 h
Reef at 0–1 m ronald Fricke
Mangroves Centre 22°22'19.29"S
40°22'22.73"E
11 Nov. 2011 Mangroves, sand
bottom (Fig. 5)
0–2 m Pascale chaBanet, Patrick
durville
Mooring North 22°20'24.15"S
40°20'11.64"E
11 Nov. 2011 Sand and patch
reef
18 m Patrick durville
Pond South 22°23'21.79"S
40°22'37.59"E
12 Nov. 2011 Sand, fossil coral 0–1 m Patrick durville

stuttGarter BeiträGe zur naturkunde a
250 Neue Serie 6
We appreciate the support of several funding agencies who
made the expeditions to Europa Island possible: the Institut
d’Écologie et Environnement du Centre National pour la Re-
cherche (INEE-CNRS), the Institut National des Sciences de
l’Univers (INSU), the Institut de Recherche pour le Développe-
ment (IRD), the Fondation pour la Recherche sur la Biodiversité
(FRB), the Agence des Aires Marines Protégées (AAMP), the
Run Sea Sciences program (Europe), and the Veolia Environment
Foundation.
2 Methods and Materials
The present paper provides a list of all shore fishes recorded
from Europa Island. For each species, the first record from Eu-
ropa Island is given. During the BioReCIE 2011 project, attempts
were made to confirm previous records of fishes from the island.
If a species is recorded from Europa Island for the first time, in-
formation on the source of the new record is presented. In ad-
dition, synonyms and misidentifications used in the literature
on Europa Island are provided, as well as English and French
vernacular names. The genus and species classification follows
eschmeyer & Fricke (2011). The species accounts include data
on the stations where the species were observed or collected. The
expedition with R/V Marion Dufresne in April 2011 is abbrevi-
ated as MD-11. Details of the station data of Europa Island (mis-
sion BioReCIE 2011) are given in Tab. 1; the positions of the sta-
tions (abbreviated “St.” in the text) are shown in Fig. 1.
While some new records are based on specimens collected, a
considerable percentage of the new records is based on observa-
tions and/or photographs, as collecting was difficult due to strong
currents. New records are only included when the identification
was unambiguous. For materials in collections, a catalogue num-
ber is provided, followed by the number of specimens, in paren-
theses.
For the distribution analysis, the following works were con-
sulted to assess the fish fauna of islands and island groups in the
Western Indian Ocean: Seychelles, smith & smith (1963) and
randall & eGmont (1994), updated using various papers in-
cluding allen (1972), dawson (1967, 1968, 1977), Fraser &
lachner (1985), Golani (1984), hoese & reader (1985), hol-
leman (2005, 2007), keith et al. (2004), kim & amaoka (2001),
lachner & karnella (1978, 1980), Polunin & luBBock (1977),
randall (1999, 2001), randall & clements (2001), randall &
heemstra (1985, 2009), randall & lourie (2009), randall et
al. (2008), schwarzhans & møller (2007), sPrinGer & williams
(1994), starck (1969), tyler (1966), uiBlein (2011); Maldives,
randall & anderson (1993) and anderson et al. (1998), updated
using various papers including allen & erdmann (2006), allen
& randall (2002), Gon & randall (2003), kuiter & deBelius
(1999), randall (2001, 2011), smith-vaniz (2010); Chagos Ar-
chipelago, winterBottom & anderson (1997), updated using
various papers including Böhlke (2000), craiG (2008), Gill &
edwards (2004), holleman (2005), randall & earle (2008),
schwarzhans et al. (2005); Réunion, Fricke et al. (2009); Mau-
ritius, Fricke (1999), updated using various papers including
Fraser & allen (2001), Fricke (2009), Gill & Fricke (2001),
holleman (2007), randall (2004, 2007, 2011), randall & Gon
(2006), randall & schultz (2008), randall & smith (2001),
schwarzhans & møller (2007), sPrinGer & Fricke (2000);
Rodrigues, heemstra et al. (2004), updated using papers in-
cluding Gill & edwards (2004) and holleman (2005). The gen-
eral distribution of fish species occurring at Europa Island was
accessed using eschmeyer & Fricke (2011), Fricke et al. (2009)
and Froese & Pauly (2011).
Abbreviation of museum collection
SMF Forschungsinstitut Senckenberg, Frankfurt/Main, Ger-
many
3 Checklist
Carcharhinidae
Carcharhinus albimarginatus (Rüppell, 1837) – Silvertip shark;
requin pointe blanche. – D is t ri bu t io n: New record from
Europa Island, based on an observation by P. durville et al.
during the expedition MD-11; also observed by P. durville &
P. chaBanet on 9–10 Nov. 2011, St. EU3, B8.
Carcharhinus amblyrhynchos (Bleeker, 1856) – Grey reef shark;
Requin dagsit. – D is t ri b u t io n : New record from Europa
Island, based on observation by P. durville & P. chaBanet
on 9–10 Nov. 2011, St. EU4, B8.
Carcharhinus melanopterus (Quoy & Gaimard, 1824) – Blacktip
reef shark; Requin à pointes noires. – D i s t r i b u t i o n :
First record from Europa Island by Fourmanoir (1952:
172, pl. 10B); record confirmed by P. durville et al. during
the expedition MD-11, an observation by P. Borsa in the
mangrove area on 7 April 2011, and by R. Fricke on 10 Nov.
2011, St. B37; also observed by P. durville & P. chaBanet,
12 Nov. 2011, St. B12, B25.
Carcharhinus obscurus (LeSueur, 1818) – Dusky shark; Requin
sombre. – D is tr i b u t i o n : First record from Europa Island
by Fourmanoir (1952: 179).
Galeocerdo cuvier (Péron & LeSueur in LeSueur, 1822) – Tiger
hark; Requin tigre. – D i s t r i b u t i o n : First record from
Europa Island by Fourmanoir (1952: 179, pl. 10A) as Ga-
leocerdo Cuvieri.
Triaenodon obesus (Rüppell, 1837) – Whitetip reef shark, blunt-
head shark; Requin corail. – D i st r ib u ti on : New record
from Europa Island, based on observation by P. durville &
P. chaBanet on 7 Nov. 2011, St. EU7.
Sphyrnidae
Sphyrna lewini (Griffith & Smith, 1834) – Scalloped hammer-
head; Requin-marteau halicorne. – D i s t r i b u t i o n : New
record from Europa Island, based on a photograph by E.
BretaGne, taken in November 2011; the species was also ob-
served by P. durville et al. (as Sphyrna sp.) during the expe-
dition MD-11; also observed by P. durville & P. chaBanet
on 9–10 Nov. 2011, St. EU3, B8.
Sphyrna mokarran (Rüppell, 1837) – Great hammerhead shark;
Grand requin-marteau. – D i s t r i b u t i o n : New record
from Europa Island, based on photograph by E. BretaGne,
taken in November 2011.
Dasyatidae
Neotrygon kuhlii (Müller & Henle, 1841) – Blue-spotted sting-
ray; Raie à points bleus. – D i s t r i b u t i o n : New record
from Europa Island, based on an observation by P. durville
et al. during the expedition MD-11.

Fricke et alii, checklist oF the shore Fishes oF euroPa island 251
Taeniura sp. – D i s t r i b u t i o n : Recorded from Europa Island
by P. durville & P. chaBanet, 12 Nov. 2011, St. B25.
Albulidae
Albula glossodonta (Forsskål in Niebuhr, 1775) – Indo-Pacific
bonefish; Banane (Réunion). – D i s t r i b u t i o n : Reported
from Europa Island as Albula vulpes by Fourmanoir (1952:
174, pl. 11A).
Moringuidae
Moringua javanica (Kaup, 1856) – Java spaghetti eel. – D i st r i-
bution: New record from Europa Island, based on speci-
mens collected by R. Fricke on 11–12 Nov. 2011, St. B28.
– M at er ia l: SMF 34766 (4), SMF 34807 (5).
Muraenidae
Echidna nebulosa (Ahl, 1789) – Snowflake moray, floral moray;
Murène étoilée. – D i s t r i b u t i o n : New record from Eu-
ropa Island, based on specimens collected by R. Fricke on
11–12 Nov. 2011, St. B28. – Ma te ri al : SMF 34767 (3),
SMF 34808 (5).
Gymnothorax breedeni McCosker & Randall, 1977 – Blackcheek
moray; Murène masquée. – D i s t r i b u t i o n : New record
from Europa Island, based on observation by P. durville &
P. chaBanet on 9 Nov. 2011, St. EU3.
Gymnothorax buroensis (Bleeker, 1857) – Vagrant moray;
Murène errante. – D i s t r i b u t i o n : New record from Eu-
ropa Island, based on specimens collected by R. Fricke on 12
Nov. 2011, St. B28. – M a t e r i a l : SMF 34809 (6).
Gymnothorax flavimarginatus (Rüppell, 1830) – Yellow-edged
moray, yellow-margined moray; Congre queue verte, murène
à queue verte (Réunion), murène à bord jaune (France). –
D i s t r i b u t i o n : First record from Europa Island as Lyco-
dontis Lemayi by Fourmanoir (1952: 188).
Gymnothorax griseus (Lacepède [ex Commerson], 1803) – Geo-
metric moray; Murène tatouée. – D i s t ri bu t i o n: New re-
cord from Europa Island, based on observations by R. Fricke
on 7–9 Nov. 2011, St. B28, B32.
Gymnothorax javanicus (Bleeker, 1859) – Giant moray; Murène
javanaise. – D i st ri bu ti on : New record from Europa Is-
land, based on an observation by P. durville & P. chaBanet
during the expedition MD-11.
Gymnothorax meleagris (Shaw in Shaw & Nodder, 1795) –
Whitemouth moray, Turkey moray; Congre pintade (Réun-
ion), murène voie lactée (France). – D i st r ib u ti on : New
record from Europa Island, based on observation by P. dur-
ville & P. chaBanet on 9–10 Nov. 2011, St. EU2, EU3.
Gymnothorax pictus (Ahl, 1789) – Peppered moray, paintspotted
moray; Congre (Réunion), murène sidérale (France) (Fig. 6).
– D i s t r i b u t i o n : First record from Europa Island as Si-
derea picta by Fourmanoir (1952: 170); record confirmed by
P. durville et al. during the expedition MD-11, a photograph
taken by P. Borsa in the mangrove area on 7 April 2011,
and specimens collected by R. Fricke on 11–12 Nov. 2011,
St. B28; also observed by P. durville & P. chaBanet,
12 Nov. 2011, St. B25. – M a te ri a l : SMF 34768 (1), SMF
34810 (2).
Gymnothorax rueppellii (McClelland, 1844) – Banded moray,
yellow-headed moray. – Di st r ib ut i on : New record from
Europa Island, based on specimens collected by R. Fricke on
11–12 Nov. 2011, St. B28. – Ma te ri al : SMF 34769 (2),
SMF 34811 (1).
Ophichthidae
Myrichthys colubrinus (Boddaert, 1781) – Ringed snake eel,
banded snake eel, harlequin snake eel. – D i s t r i b u t i o n :
First record from Europa Island by Fourmanoir 1952: 188.
Myrichthys maculosus (Cuvier, 1816) – Ocellated snake eel,
spotted snake eel, tiger snake eel; Anguille-serpent maculée.
– Distribution: New record from Europa Island, based
on specimens collected by R. Fricke on 11–12 Nov. 2011,
St. B28. – M at er ia l: SMF 34770 (1), SMF 34812 (2).
Congridae
Conger cinereus Klunzinger [ex Rüppell], 1830 – Longfin Afri-
can conger; Congre à moustache. – D i s t r i b u t i o n : New
record from Europa Island, based on specimens collected by
R. Fricke on 11–12 Nov. 2011, St. B28. – M a te ri a l : SMF
34752 (1), SMF 34795 (11).
Chanidae
Chanos chanos (Forsskål in Niebuhr, 1775) – Milkfish; Chano.
– Distribution: New record from Europa Island, based
on visual record by nicole Gravier-Bonnet on 3 June 2006
in a little pond on southeastern part of Europa Island; record
confirmed based on photograph by E. BretaGne, taken off the
reef on the north shore of Europa Island in November 2011,
and on an observation by R. Fricke on 11 Nov. 2011, St. EU5.
Clupeidae
Herklotsichthys quadrimaculatus (Rüppell, 1837) – Bluestripe
herring, blueline herring; Sardine (Réunion). – D i st r i b u -
tion: New record from Europa Island, based on a photo-
graph by P. durville taken on 11 Nov. 2011 among the man-
groves in the inner lagoon of Europa Island.
Spratelloides gracilis (Temminck & Schlegel, 1846) – Silver-
stripe round herring. – D i st r i b u t i o n : New record from
Europa Island, based on a photograph by P. durville taken
on 11 Nov. 2011 among the mangroves in the inner lagoon
of Europa Island.
Synodontidae
Synodus binotatus Schultz in Schultz, Herald, Lachner, Welander
& Woods, 1953 – Twospot lizardfish; Anoli à deux taches.
– Distribution: New record from Europa Island, based
on an observation by R. Fricke on 7–10 Nov. 2011, St. B33,
B34, B37, B50.
Synodus dermatogenys Fowler, 1912 – Sand lizardfish; Anoli
bigarré. – Distribution: New record from Europa Is-
land, based on a specimen collected by R. Fricke on 12 Nov.
2011, St. B28. – M a t e r i a l : SMF 34823 (1).
Carapidae
Encheliophis homei (Richardson, 1846) – Silver pearlfish. –
Distribution: New record from Europa Island, based
on a specimen collected and photographed by P. chaBanet
on 11 Nov. 2011, among the mangroves in the inner lagoon.
The fish was associated with Bohadschia atra Massin, Raso-

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252 Neue Serie 6
lofonirina, Conand & Samyn, 1999 (C. conand, pers. comm.,
31 Mar. 2012). – Ma te ri al : SMF 34784 (1).
Antennariidae
Antennarius commerson (Lacepède [ex Commerson] in Anony-
mus, 1798) – Big angler, giant frogfish, Commerson’s frog-
fish; Grenouille de Commerson. – D i s t r i b u t i o n : New
record from Europa Island, based on a specimen collected by
P. durville & P. chaBanet on 7 Nov. 2011, St. EU7.
Atherinidae
Atherinomorus lacunosus (Bloch & Schneider [ex Forster], 1801)
– Hardyhead silverside; Athérine têtue. – Di s tr i b u t i o n :
Reported from Europa Island as Atherina sp. by Fourmanoir
(1952: 187).
Mugilidae
Chelon macrolepis (Smith, 1846) – Largescale mullet; Mulet à
grandes écailles. – D i s t r i b u t i o n : First record from Eu-
ropa Island as Liza macrolepis by Fourmanoir (1952: 175);
record confirmed by P. durville et al. based on a observation
at St. 2 (equals St. B34 of this paper) during an expedition
with R/V Marion Dufresne on 6 April 2011, by a photograph
taken in the mangrove area by P. Borsa on 7 April 2011, and
by R. Fricke on 9 Nov. 2011, St. B28.
Valamugil sp. – D i s t r ib ut io n : Observed at Europa Island by
P. durville et al. during the expedition MD-11.
Hemiramphidae
Hemiramphus far (Forsskål in Niebuhr, 1775) – Black-barred
halfbeak; Demi-bec bagnard. – D i s t r i b u t i o n : New re-
cord from Europa Island, based on an observation by P. dur-
ville et al. during the expedition MD-11.
Hyporhamphus dussumieri (Valenciennes in Cuvier & Valenci-
ennes, 1847) – Dussumier’s halfbeak; Demi-bec de Dussu-
mier. – Di st ri b u t i o n : First record from Europa Island by
Fourmanoir (1952: 185).
Belonidae
Strongylura leiura (Bleeker, 1850) – Yellowfin needlefish; Or-
phie littorale (Fig. 7). – D i s tr i b u ti o n : New record from
Europa Island, based on an observation and photograph by P.
durville et al. during the expedition MD-11, and an observa-
tion by P. Borsa in the mangrove area on 7 April 2011.
Tylosurus crocodilus (Péron & LeSueur in LeSueur, 1821) –
Hound needlefish, crocodile needlefish; Aiguille (Réunion),
orphie crocodile (France). – D i s t r i b u t i o n : Reported
from Europa Island as Belone choram (non Rüppell, 1837)
by Fourmanoir (1952: 170), and as Tylosurus choram (non
Rüppell, 1837) by Fourmanoir (1952: 185); record of Tylo-
surus crocodilus confirmed by P. durville et al. during the
expedition MD-11.
Holocentridae
Myripristis adusta Bleeker, 1853 – Shadowfin soldier; Marignan
ombré. – Distribution: New record from Europa Island,
based on an observation by P. durville & P. chaBanet on 7
Nov. 2011, St. EU7.
Myripristis berndti Jordan & Evermann, 1903 – Blotcheye sol-
dierfish, bigscale soldierfish; Cardinal (Réunion), marignan
à oeillères (France). – D i s t r i b u t i o n : New record from
Europa Island, based on an observation by P. durville et al.
during the expedition MD-11; also observed by P. durville &
P. chaBanet on 7–12 Nov. 2011, St. B5, B8, B12, B14, EU2,
EU7; and by R. Fricke on 11 Nov. 2011, St. EU5.
Myripristis kuntee Valenciennes in Cuvier & Valenciennes, 1831
– Shoulderbar soldierfish, epaulette soldierfish, pearly sol-
dierfish; Marignan ardoisé. – D is t r i b u ti on : New record
from Europa Island, based on an observation by P. durville
et al. during the expedition MD-11.
Myripristis murdjan (Forsskål in Niebuhr, 1775) – Pinecone
soldierfish, blotcheye soldier, red soldierfish; Cardinal gros
écailles (Réunion), marignan pomme de pin (France). – D i s-
t r ib ut i on : First record from Europa Island by Fourmanoir
(1952: 184); record confirmed by R. Fricke based on an ob-
servation on 10 Nov. 2011, St. B37.
Myripristis vittata Valenciennes in Cuvier & Valenciennes, 1831
– Immaculate soldier, white-tipped soldierfish; Cardinal petit
écailles (Réunion). – D i s t r i b u t i o n : New record from
Europa Island, based on an observation by P. durville et al.
during the expedition MD-11; also observed by P. durville
& P. chaBanet on 7 Nov. 2011, St. EU7.
Neoniphon sammara (Forsskål in Niebuhr, 1775) – Bloodspot
squirrelfish; Marignan tacheté. – D i st r ib u ti o n: First re-
cord from Europa Island as Holocentrus sammara by Four-
manoir (1952: 185); record confirmed by P. durville et al.
during the expedition MD-11; also observed by P. durville
& P. chaBanet on 7 Nov. 2011, St. EU7.
Sargocentron caudimaculatum (Rüppell, 1838) – Silverspot
squirrelfish, tailspot squirrelfish; Marignan rouge et argent.
– Distribution: New record from Europa Island, based
on an observation by P. durville et al. during the expedition
MD-11; also observed by P. durville & P. chaBanet on 7–12
Nov. 2011, St. B12, B14, EU2, EU7; record confirmed by R.
Fricke based on an observation on 9–11 Nov. 2011, St. EU5.
Sargocentron diadema (Lacepède [ex Commerson], 1802) –
Crown squirrelfish; Soldat couronné. – D i s t r i b u t i o n :
First record from Europa Island as Holocentrus diadema by
Fourmanoir (1952: 174).
Sargocentron spiniferum (Forsskål in Niebuhr, 1775) – Sabre
squirrelfish, long-jawed squirrelfish; Cardinal lancette (Ré-
union), marignan sabre (France). – D i s t r i b u t i o n : First
record from Europa Island as Holocentrus spiniferus by
Fourmanoir (1952: 183); record confirmed by P. durville
et al. based on a observation at St. 1 (equals St. EU5 of this
paper) during an expedition with R/V Marion Dufresne on 5
April 2011; also observed by P. durville & P. chaBanet on
7–12 Nov. 2011, St. B8, B12, EU7.
Syngnathidae
Choeroichthys smithi Dawson, 1976 – Smith’s short-bodied pipe-
fish. – Distribution: New record from Europa Island,
based on specimens collected by R. Fricke on 11 Nov. 2011,
St. B28. – M at er ia l: SMF 34781 (5).
Scorpaenidae
Caracanthus madagascariensis (Guichenot, 1869) – Spotted
croucher. – Di st ri bu ti on : First record from Europa Is-

Fricke et alii, checklist oF the shore Fishes oF euroPa island 253
land as Caracanthus zeylonicus by Fourmanoir (1952: 187);
record confirmed by P. durville et al. during the expedition
MD-11.
Pterois miles (Bennett, 1828) – Devil firefish; Poisson armée. –
D i s t r i b u t i o n : Reported from Europa Island as Pterois
volitans (non Linnaeus, 1758) by Fourmanoir (1952: 175);
Figs. 2–7. Europa Island (2–6) and shore shes from this island (7, 8). – 2. North coast with Gendarmerie building; photograph by
P. Borsa, April 2011. 3. West coast; aerial photograph by P. Borsa, April 2011. 4. West coast; photograph by R. Fricke, 9 Nov. 2011.
5. Inner lagoon; aerial photograph by P. Borsa, April 2011. 6. Gymnothorax pictus; photograph by P. Borsa, 7 April 2011. 7. Strongy-
lura leiura; photograph by P. Borsa, 7 April 2011.

stuttGarter BeiträGe zur naturkunde a
254 Neue Serie 6
Figs. 8–13. Shore shes of Europa Island. – 8. Epinephelus tukula; photograph by P. durville, April 2011. 9. Priacanthus hamrur;
photograph by P. durville, April 2011. 10. Caranx melampygus; photograph by E. BretaGne, Oct. 2011. 11 . Gerres longirostris; pho-
tograph by P. Borsa, 7 April 2011. 12. Monotaxis grandoculis; photograph by P. Borsa, 5 April 2011. 13. Mulloidichthys vanicolensis
(foreground) and Gnathodentex aureolineatus (background); also includes Labroides dimidiatus and Gomphosus caeruleus (below),
and Naso elegans (background right); photograph by P. durville, April 2011.

Fricke et alii, checklist oF the shore Fishes oF euroPa island 255
Figs. 14–19. Shore shes of Europa Island. – 14. Chaetodon auriga; photograph by P. Borsa, 6 April 2011. 15. Chaetodon trifasciatus;
photograph by P. Borsa, 6 April 2011. 16. Chaetodon xanthocephalus; photograph by P. Borsa, 6 April 2011. 17. Pomacentrus
caeruleus; photograph by R. Fricke, 8 Nov. 2011. 18. Amblygobius semicinctus; photograph by P. durville, 11 Nov. 2011. 19. Sphy-
raena barracuda; photograph by P. Borsa, 7 April 2011.

stuttGarter BeiträGe zur naturkunde a
256 Neue Serie 6
record confirmed by R. Fricke based on an observation at
Europa on 7 Nov. 2011, St. B33.
Scorpaenodes guamensis (Quoy & Gaimard, 1824) – Guam scor-
pionfish. – Distribution: New record from Europa Is-
land, based on specimens collected by R. Fricke on 12 Nov.
2011, St. B28. – M a t e r i a l : SMF 34817 (2).
Scorpaenodes parvipinnis (Garrett, 1864) – Coral scorpionfish.
– Distribution: New record from Europa Island, based
on specimens collected by R. Fricke on 9–12 Nov. 2011,
St. B28. – M a t e r i a l : SMF 34735 (1), SMF 34777 (4),
SMF 34818 (7).
Sebastapistes strongia (Cuvier in Cuvier & Valenciennes, 1829)
– Barchin scorpionfish. – D i s tr i bu t i on : First record from
Europa Island as Scorpaena kowiensis and Cantapus oglinus
by Fourmanoir (1952: 187).
Serranidae
R e m a r k s : A record of Epinephelus guaza (non Linnaeus,
1758) by Fourmanoir (1952: 184) could not be identified to
the species level.
Aethaloperca rogaa (Forsskål in Niebuhr, 1775) – Redmouth
grouper; Vieille roga. – Di s t r ib u t i o n : New record from
Europa Island, based on an observation by P. durville et al.
during the expedition MD-11; also observed by P. durville
& P. chaBanet on 7–12 Nov. 2011, St. EU3, EU7, B12, B14.
Cephalopholis argus Bloch & Schneider, 1801 – Peacock
grouper, peacock rockcod; La prude (Réunion), mérou cé-
leste (France). – D i s t r i b u t i o n : First record from Eu-
ropa Island by Fourmanoir (1952: 183); record confirmed
by P. durville et al. based on a observation at St. 1 (equals
St. EU5 of this paper) during an expedition with R/V Marion
Dufresne on 5 April 2011; also observed by P. durville & P.
chaBanet on 7–12 Nov. 2011, St. B5, B8, B12, B14, EU2,
EU3, EU5, EU7, and by R. Fricke based on an observation
and specimens collected on 11–12 Nov. 2011, St. B28, EU5.
– M at er ia l: SMF 34778 (2), SMF 34819 (2).
Cephalopholis miniata (Forsskål in Niebuhr, 1775) – Coral
hind, coral rockcod; Ananas batard (Réunion), vieille étoilée
(France). – D i s t r i b u t i o n : First record from Europa Is-
land as Cephalopholis miniatus by Fourmanoir (1952: 183);
record confirmed by P. durville et al. during the expedition
MD-11; also observed by P. durville & P. chaBanet on 9–11
Nov. 2011, St. B14, EU3.
Cephalopholis nigripinnis (Valenciennes in Cuvier & Valenci-
ennes, 1828) – Darkfin hind; Vieille aile noire. – Di st r ib u -
tion: New record from Europa Island, based on an observa-
tion by P. durville et al. during the expedition MD-11; also
observed by P. durville & P. chaBanet on 7–12 Nov. 2011,
St. B5, B12, B14, EU2, EU7. – R e m a rk s : Closely related
to Cephalopholis urodeta (Forster in Bloch & Schneider,
1801) which is restricted to the western Pacific.
Cephalopholis polleni (Bleeker, 1868) – Harlequin hind, har-
lequin rockcod; Cabot guignan (Réunion), vieille arlequin
(France). – Distribution: New record from Europa Is-
land, based on an observation by P. durville et al. during the
expedition MD-11.
Cephalopholis sexmaculata (Rüppell, 1830) – Sixspot hind,
sixblotch rockcod; Rouge l’allié (Réunion), vieille six taches
(France). – Distribution: New record from Europa Is-
land, based on an observation by P. durville et al. during the
expedition MD-11; also observed by P. durville & P. cha-
Banet on 7–11 Nov. 2011, St. B14, EU3, EU7.
Cephalopholis sonnerati (Valenciennes in Cuvier & Valenci-
ennes, 1828) – Tomato hind, tomato rockcod; Rouge ananas,
la prude rouge (Réunion), vieille ananas (France). – D is t ri -
b u t i o n : First record from Europa Island by Fourmanoir
(1952: 183).
Epinephelus areolatus (Forsskål in Niebuhr, 1775) – Areolate
grouper. – Di st r i b ut io n: First record from Europa Island
by Fourmanoir (1952: 182).
Epinephelus coeruleopunctatus (Bloch, 1790) – White-spotted
grouper; Loche à taches blanches. – D i st r ib ut io n: New
record from Europa Island, based on a photograph by P. dur-
ville taken on 11 Nov. 2011 among the mangroves in the in-
ner lagoon of Europa Island.
Epinephelus coioides (Hamilton, 1822) – Orange-spotted
grouper; Mérou à taches oranges. – D i s t r i b u t i o n : New
record from Europa Island, based on an observation by P.
durville, 11 Nov. 2011, St. B14.
Epinephelus flavocaeruleus (Lacepède [ex Commerson], 1802)
– Blue-and-yellow grouper; Le plat, platte ailes jaunes (Ré-
union), mérou faraud (France). – D i st ri bu ti on : First re-
cord from Europa Island by Fourmanoir (1952: 183); record
confirmed by P. durville et al. during the expedition MD-11;
also observed by P. durville & P. chaBanet on 9–11 Nov.
2011, St. B5, EU4.
Epinephelus fuscoguttatus (Forsskål in Niebuhr, 1775) – Brown-
marbled grouper, blotchy rockcod; Mérou marbré. – Di s-
tribution: New record from Europa Island, based on an
observation by P. durville et al. during the expedition MD-
11.
Epinephelus hexagonatus (Bloch & Schneider [ex Forster], 1801)
– Star-spotted grouper; Maccabit, maccabit gris, risadel (Ré-
union), mérou mélifère (France). – D i s t r i b u t i o n : New
record from Europa Island, based on an observation by P.
durville et al. during the expedition MD-11; also by R.
Fricke based on an observation and a specimen collected on
11 Nov. 2011, St. B28, EU5; also observed by P. durville, 12
Nov. 2011, St. B25. – M a t e r i a l : SMF 34779 (1).
Epinephelus maculatus (Bloch, 1790) – Highfin grouper; Loche
de sable. – D i st r ib u ti o n: First record from Europa Island
by Fourmanoir (1952: 172).
Epinephelus merra Bloch, 1793 – Honeycomb grouper, dwarf
spotted grouper; Maccabit, maccabit blanc (Réunion), griffin
(Réunion, young specimens), plat gris (Réunion, adult speci-
mens), mérou gateau de cire (France). – D i s t r i b u t i o n :
First record from Europa Island by Fourmanoir (1952: 172);
record confirmed by P. durville et al. based on observations
at St. 1 and St. 2 (equals St. EU5 and B34 of this paper) dur-
ing an expedition with R/V Marion Dufresne on 5–6 April
2011, and by R. Fricke on 8–12 Nov. 2011, St. B28, B34; also
observed by P. durville, 12 Nov. 2011, St. B12. – M a t e -
r ia l: SMF 34820 (1).
Epinephelus oceanicus (Lacepède, 1802) – Indian Ocean blacktip
grouper; Rougette (Réunion), mérou oriflamine (France). –
D i st r i bu t io n : Reported from Europa Island as Epinephe-
lus fasciatus (non Forsskål 1775) by Fourmanoir (1952: 184).

Fricke et alii, checklist oF the shore Fishes oF euroPa island 257
Epinephelus ongus (Bloch, 1790) – Specklefin grouper; Loche à
taches claires. – D is tr ib ut io n: New record from Europa
Island, based on a photograph taken by P. chaBanet on 12
Nov. 2011, St. B12.
Epinephelus polyphekadion (Bleeker, 1849) – Camouflage
grouper; Loche crasseuse. – D i s t r i b u t i o n : New record
from Europa Island, based on a photograph by P. durville
taken on 11 Nov. 2011 among the mangroves in the inner la-
goon of Europa Island.
Epinephelus spilotoceps Schultz in Schultz, Herald, Lachner,
Welander & Woods, 1953 – Foursaddle grouper, foursaddle
rockcod; Mérou à quatre selles. – D is tr ib ut io n: New re-
cord from Europa Island, based on an observation by P. dur-
ville et al. at St. 1 (equals St. EU5 of this paper) during an
expedition with R/V Marion Dufresne on 5 April 2011; also
observed by P. durville & P. chaBanet on 7–11 Nov. 2011,
St. B5, EU2, EU5, EU7, and by R. Fricke on 11–12 Nov.
2011, St. EU5. – M a t e r i a l : SMF 34821 (1).
Epinephelus tauvina (Forsskål in Niebuhr, 1775) – Greasy
grouper, greasy rockcod; Vieille lutre, vieille nègre (Réun-
ion), mérou loutre (France). – D is t ri b ut io n : New record
from Europa Island, based on an observation by P. durville
et al. during the expedition MD-11; also observed by P. dur-
ville, 11 Nov. 2011, St. B5.
Epinephelus tukula Morgans, 1959 – Potato grouper, potato bass;
Mérou patate (Fig. 8). – D is tr ib u t i o n : New record from
Europa Island, based on observations by P. durville et al. at
St. 1 and St. 2 (equals St. EU5 and B34 of this paper) during
an expedition with R/V Marion Dufresne on 5–6 April 2011;
also observed by P. durville & P. chaBanet on 7–11 Nov.
2011, St. B5, B14, EU2, EU3, EU5, EU7; record confirmed
based on photograph by E. BretaGne, taken in November
2011.
Gracila albomarginata (Fowler & Bean, 1930) – Slenderspine
grouper; Vieille (Réunion), mérou bord rouge (France). –
Distribution: New record from Europa Island, based
on an observation by P. durville et al. during the expedi-
tion MD-11; also observed by P. durville, 11–12 Nov. 2011,
St. B12, B14.
Grammistes sexlineatus (Thunberg, 1792) – Sixstripe soapfish,
skunkfish; Savonette (Réunion), savon rayes d’or (France).
– D i s tr i b u ti o n : First record from Europa Island by Four-
manoir (1952: 172); record confirmed based on specimens
collected by R. Fricke on 11–12 Nov. 2011, St. B28. – Ma -
t er ia l: SMF 34780 (1), SMF 34822 (2).
Nemanthias carberryi Smith, 1954 – Threadfin anthias. – D i s -
tribution: New record from Europa Island, based on an
observation by P. durville et al. during the expedition MD-
11; also observed by P. durville & P. chaBanet on 9–12 Nov.
2011, St. EU3, B5, B8, B12, B14.
Plectropomus laevis (Lacepède [ex Commerson], 1801) – Black-
saddle coral grouper; Mérou selle. – D i st ri bu ti on : New
record from Europa Island; previously reported as Plectro-
pomus maculatus (non Bloch, 1790) by Fourmanoir (1952:
183).
Plectropomus pessuliferus (Fowler, 1904) – Roving coral
grouper; Mérou loche vagabonde. – D i st r ib u t i o n : New
record from Europa Island, based on an observation by P.
durville et al. during the expedition MD-11.
Plectropomus punctatus (Quoy & Gaimard, 1824) – Marbled
coral grouper. – Di st ri bu ti o n : New record from Europa
Island, based on an observation and a photograph by P. dur-
ville, 11–12 Nov. 2011, St. B12, and among the mangroves
in the inner lagoon.
Pseudanthias evansi (Smith, 1954) – Yellowback anthias, yel-
lowtail goldie. – D is t ri b u t io n : New record from Europa
Island, based on an observation by P. durville et al. during
the expedition MD-11; also observed by P. durville & P.
chaBanet on 7–11 Nov. 2011, St. B14, EU7.
Pseudanthias squamipinnis (Peters, 1855) – Sea goldie. – D i s-
tribution: New record from Europa Island, based on an
observation by P. durville et al. during the expedition MD-
11; also observed by P. durville & P. chaBanet on 7–12 Nov.
2011, St. B5, B8, B12, B14, EU2, EU3, EU7.
Variola louti (Forsskål in Niebuhr, 1775) – Coronation grouper;
Druide, rouge grand queue (Réunion), croissant queue jaune
(France). – D i s t r i b u t i o n : First record from Europa Is-
land by Fourmanoir (1952: pl. 11B); record confirmed by P.
durville et al. during the expedition MD-11; also observed
by P. durville & P. chaBanet on 7–12 Nov. 2011, St. B5, B8,
B12, B14, EU2, EU3, EU4, EU7.
Pseudochromidae
Chlidichthys johnvoelckeri Smith, 1953 – Cerise dottyback. –
Distribution: New record from Europa Island, based on
an observation as Pseudochromis fridmani (non Klausewitz,
1968) by P. durville et al. during the expedition MD-11, a
specimen collected by P. durville & P. chaBanet on 10 Nov.
2011, St. B23, and a photograph taken by P. durville on 7
Nov. 2011, St. EU7; also observed by P. durville & P. cha-
Banet on 11 Nov. 2011, St. B14. – Ma t e r ia l : SMF 34741
(1).
Teraponidae
Terapon jarbua (Forsskål in Niebuhr, 1775) – Jarbua, crescent-
banded grunter, thornfish; Violon jarbua. – D i s tr i bu t io n :
New record from Europa Island; previously reported as
Therapon sp. by Fourmanoir (1952: 170); record confirmed
by P. durville et al. during the expedition MD-11, a pho-
tograph taken by P. Borsa in the mangrove area on 7 April
2011, and by R. Fricke based on an observation at Europa on
7 Nov. 2011, St. B31.
Kuhliidae
Kuhlia mugil (Bloch & Schneider [ex Forster], 1801) – Barred
flagtail; Hareng (Réunion), crocro drapeau (France). – D i s -
t r i b u t i o n : Reported from Europa Island as Dules taeni-
urus by Fourmanoir (1952: 170); record confirmed based on
a specimen collected by R. Fricke on 11 Nov. 2011, St. B28;
also observed by P. durville, 12 Nov. 2011, St. B25. – Ma -
t er ia l: SMF 34758 (1).
Priacanthidae
Heteropriacanthus cruentatus (Lacepède, 1801) – Glasseye;
Beau clair du large (Réunion), beau clair de roche (France).
– Distribution: New record from Europa Island, based
on an observation by P. durville et al. during the expedition
MD-11.

stuttGarter BeiträGe zur naturkunde a
258 Neue Serie 6
Priacanthus hamrur (Forsskål in Niebuhr, 1775) – Moontail
bullseye; Cardinal, beau clair (Réunion), beau clair miroir
(France) (Fig. 9). – D i s t r i b u t i o n : New record from
Europa Island, based on an observation by P. durville et al.
during the expedition MD-11; also observed by P. durville,
10–11 Nov. 2011, St. B8, B14.
Apogonidae
R em ar ks : A record of Apogon novemfasciatus by Fourmanoir
(1952: 172) could not be identified to the species level.
Apogon semiornatus Peters, 1877 – Oblique-banded cardinalfish.
– Distribution: New record from Europa Island, based
on specimens collected by R. Fricke on 12 Nov. 2011,
St. B28. – M at er ia l: SMF 34786 (4).
Cheilodipterus artus Smith, 1961 – Wolf cardinalfish; Apogon
strié. – Distribution: New record from Europa Island,
based on an observation by P. durville et al. during the expe-
dition MD-11; also observed by P. durville & P. chaBanet
on 7 Nov. 2011, St. EU7.
Ostorhinchus apogonoides (Bleeker, 1856) – Shirt-tooth cardi-
nalfish. – Distribution: New record from Europa Island,
based on an observation by P. durville et al. during the ex-
pedition MD-11.
Ostorhinchus holotaenia (Regan, 1905) – Copperstriped cardi-
nalfish. – Distribution: New record from Europa Island,
based on a specimen collected by R. Fricke on 11 Nov. 2011,
St. B28. – M at er ia l: SMF 34743 (1).
Ostorhinchus taeniophorus (Regan, 1908) – Reef-flat cardi-
nalfish, ninestripe cardinal. – D i s t r i b u t i o n : New re-
cord from Europa Island, based on specimens collected by
R. Fricke on 9–12 Nov. 2011, St. B28; also observed and
collected by P. durville, 10–12 Nov. 2011, St. EU2, B25.
– M a t e ri al : SMF 34727 (2), SMF 34744 (2), SMF 34787
(8), SMF 34827 (2).
Pristiapogon kallopterus (Bleeker, 1856) – Iridescent cardi-
nalfish, spinyhead cardinal. – D is tr i b u t i on : New record
from Europa Island, based on an observation by P. durville
et al. during the expedition MD-11; also observed and photo-
graphed by P. durville on 7 Nov. 2011, St. EU7.
Rhabdamia gracilis (Bleeker, 1856) – Luminous cardinalfish. –
Distribution: New record from Europa Island, based
on specimens collected by R. Fricke on 11–12 Nov. 2011,
St. B28. – M at er ia l: SMF 34745 (5), SMF 34788 (2).
Malacanthidae
Malacanthus latovittatus (Lacepède [ex Commerson], 1801) –
Blue tilefish, blue blanquillo, striped blanquillo; Matajuel
bleu. – Distribution: New record from Europa Island,
based on an observation by P. durville et al. during the ex-
pedition MD-11.
Echeneidae
Echeneis naucrates Linnaeus, 1758 – Live sharksucker, shark
remora; Pilote (Réunion). – D i s t r i b u t i o n : First record
from Europa Island by Fourmanoir (1952: 187); record con-
firmed by P. durville et al. during the expedition MD-11.
Remora remora (Linnaeus, 1758) – Common remora; Pilote (Ré-
union), rémora (France). – D i s t r i b u t i o n : First record
from Europa Island by Fourmanoir (1952: 187); record con-
firmed based on photograph by E. BretaGne, taken in No-
vember 2011.
Carangidae
R e ma rk s: A record of Chorinemus sp. by Fourmanoir (1952:
186) could not identified to the species level.
Atule mate (Cuvier in Cuvier & Valenciennes, 1833) – Yellowtail
scad; Carangue maté. – Distribution: New record from
Europa Island, based on a photograph by P. durville taken
on 11 Nov. 2011 among the mangroves in the inner lagoon
of Europa Island.
Carangoides fulvoguttatus (Forsskål in Niebuhr, 1775) – Yellow-
spotted trevally, yellow-dotted trevally; Carangue pailletée.
– Distribution: New record from Europa Island, based
on an observation by P. durville et al. during the expedition
MD-11.
Carangoides orthogrammus (Jordan & Gilbert, 1882) – Island
trevally; Carangue des Îles. – D is tr ib ut io n: New record
from Europa Island, based on an observation by P. durville
et al. during the expedition MD-11; also observed by P. dur-
ville, 11 Nov. 2011, St. B14.
Caranx ignobilis (Forsskål in Niebuhr, 1775) – Giant trevally;
Carangue grosse tête (Réunion), carangue têtue (France). –
D i st ri bu ti on : First record from Europa Island by Four-
manoir (1952: 183); record confirmed by P. durville et al.
during the expedition MD-11, and based on photograph by
E. BretaGne, taken in November 2011; also observed by P.
durville, 10–11 Nov. 2011, St. B5, B8, B14, EU2.
Caranx lugubris Poey, 1860 – Black kingfish, black jack; Ca-
rangue noire (France). – D is t ri bu t io n: New record from
Europa Island, based on observation by P. durville & P. cha-
Banet on 7 Nov. 2011, St. EU7.
Caranx melampygus Cuvier in Cuvier & Valenciennes, 1833 –
Bluefin trevally, bluefin kingfish; Carangue bleu (Réunion),
carangue aile bleue (France) (Fig. 10). – Di st ri bu ti on :
First record from Europa Island as Caranx stellatus by Four-
manoir (1952: 174); record confirmed by P. durville et al.
based on observations at St. 1 and St. 2 (equals St. EU5 and
St. B34 of this paper) during an expedition with R/V Marion
Dufresne on 5–6 April 2011; also observed by P. durville &
P. chaBanet on 7–12 Nov. 2011, St. B12, B14, B25, EU3,
EU4, EU7, and by R. Fricke on 11 Nov. 2011, St. EU5.
Caranx sexfasciatus Quoy & Gaimard, 1825 – Bigeye trevally,
bigeye kingfish; Carangue (Réunion), carangue vorace
(France). – Distribution: New record from Europa Is-
land; previously reported as Caranx hippos (non Linné, 1766)
by Fourmanoir (1952: 186); record of Caranx sexfasciatus
confirmed by P. durville et al. during the expedition MD-
11; also observed and photographed by P. durville, 12 Nov.
2011, St. B12, and among the mangroves in the inner lagoon.
Caranx tille Cuvier in Cuvier & Valenciennes 1833 – Tille
trevally; Carangue tille. – D i s t r i b u t i o n : New record
from Europa Island, based on observation by P. durville &
P. chaBanet on 7–11 Nov. 2011, St. EU7, and by R. Fricke
on 8 Nov. 2011, St. B34.
Elagatis bipinnulata (Quoy & Gaimard, 1825) – Rainbow runner;
Comète saumon. – D i s t r i b u t i o n : First record from Eu-

Fricke et alii, checklist oF the shore Fishes oF euroPa island 259
ropa Island as Elagastis bipinnulatus by Fourmanoir (1952:
186); record confirmed by P. durville et al. during the expe-
dition MD-11; also observed by P. durville, 10 Nov. 2011,
St. B8.
Scomberoides lysan (Forsskål in Niebuhr, 1775) – Double-spot-
ted queenfish, leatherback; Pompre, sauteur sabre. – D i s -
tribution: New record from Europa Island, based on an
observation by P. durville et al. during the expedition MD-
11; also observed by P. durville, 12 Nov. 2011, St. B25.
Trachinotus blochii (Lacepède [ex Commerson], 1801) – Snub-
nose pompano; Pompaneau lune. – D i s t r i b u t i o n : First
record from Europa Island as Trachinotus Blochii by Four-
manoir (1952: 186); record confirmed by P. durville et al.
based on a observation at St. 2 (equals St. B34 of this paper)
during an expedition with R/V Marion Dufresne on 6 April
2011, and by R. Fricke based on an observation on 11 Nov.
2011, St. EU5; also observed by P. durville, 10–12 Nov.
2011, St. B8, B14, B25.
Lutjanidae
Aphareus furca (Lacepède [ex Commerson], 1801) – Blue small-
tooth jobfish; Lantanier noir (Réunion), vivaneau tidents
(France). – Distribution: New record from Europa Is-
land, based on an observation by P. durville et al. during
the expedition MD-11; also observed by P. durville & P.
chaBanet on 7–12 Nov. 2011, St. B5, B8, B12, B14, EU2,
EU3, EU7.
Aprion virescens Valenciennes in Cuvier & Valenciennes, 1830
– Green jobfish; Thazard blanc (Réunion), vivaneau job
(France). – Distribution: New record from Europa Is-
land, based on an observation by P. durville et al. during the
expedition MD-11; also observed by P. durville & P. cha-
Banet on 9–11 Nov. 2011, St. B5, B8, B14, EU3.
Lutjanus argentimaculatus (Forsskål in Niebuhr, 1775) – River
snapper, mangrove red snapper; Vivaneau des mangroves
(France). – D i s t r i b ut i o n : First record from Europa Island
as Lutianus argentimaculatus by Fourmanoir (1952: 178); re-
cord confirmed by P. durville et al. based on a observation at
St. 1 (equals St. EU5 of this paper) during an expedition with
R/V Marion Dufresne on 5 April 2011, and by R. Fricke on
8 Nov. 2011, St. B49; also observed by P. durville, 12 Nov.
2011, in a pond on the south side of the island.
Lutjanus bohar (Forsskål in Niebuhr, 1775) – Twinspot snapper;
Vara-vara (Réunion), vivaneau chien rouge (France). – D i s-
tribution: New record from Europa Island, based on an
observation by P. durville et al. during the expedition MD-
11; also observed by P. durville & P. chaBanet on 7–12 Nov.
2011, St. B5, B8, B12, B14, EU2, EU3, EU4, EU5, EU7.
Lutjanus fulviflamma (Forsskål in Niebuhr, 1775) – Dory snap-
per, blackspot snapper; Vivaneau gibelot (France). – D i s -
t r i b u t i o n : First record from Europa Island as Lutianus
fulviflamma by Fourmanoir (1952: 170); record confirmed
by P. durville et al. during the expedition MD-11, and a pho-
tograph taken by P. Borsa in the mangrove area on 7 April
2011.
Lutjanus fulvus (Schneider [ex Forster] in Bloch & Schneider,
1801) – Yellow striped snapper, flametail snapper; Dorée
(Réunion), vivaneau queue noire (France). – D i s t r i b u -
t io n : First record from Europa Island as Lutianus vaigiensis
by Fourmanoir (1952: 174); record confirmed by P. durville
et al. based on a observation at St. 1 (equals St. EU5 of this
paper) during an expedition with R/V Marion Dufresne on 5
April 2011, an observation by P. Borsa in the mangrove area
on 7 April 2011, and by R. Fricke based on observations at
Europa on 7–8 Nov. 2011, St. B31, B33, B34.
Lutjanus gibbus (Forsskål in Niebuhr, 1775) – Humpback snap-
per; Marie-Therèse (Réunion), vivaneau pagaie (France).
– D i s t r i b u t i o n : First record from Europa Island as Lu-
tjanus gibbus by Fourmanoir (1952: 182); record confirmed
by P. durville et al. based on a observation at St. 1 (equals
St. EU5 of this paper) during an expedition with R/V Marion
Dufresne on 5 April 2011, an observation by P. Borsa in the
mangrove area on 7 April 2011, and by R. Fricke based on an
observation on 8–10 Nov. 2011, St. B34, B50; also observed
by P. durville & P. chaBanet on 7–10 Nov. 2011, St. B8,
EU7.
Lutjanus johnii (Bloch, 1792) – John’s snapper. – D i s t r i b u -
t i o n : First record from Europa Island as Lutianus Johni
by Fourmanoir (1952: 170); record confirmed by R. Fricke
based on observations at Europa on 7–10 Nov. 2011, St. B31,
B33, B34, B49, B50.
Lutjanus kasmira (Forsskål in Niebuhr, 1775) – Blue-banded
snapper, blue-lined snapper; Le petite jaune (Réunion), vi-
vaneau à raies bleues (France). – D i st ri bu ti on : First re-
cord from Europa Island as Lutianus kasmira by Fourmanoir
(1952: 184); record confirmed by P. durville et al. based on
observations at St. 1 and St. 2 (equals St. EU5 and B34 of
this paper) during an expedition with R/V Marion Dufresne
on 5–6 April 2011, and by R. Fricke on 8 Nov. 2011, St. B49.
Lutjanus lutjanus Bloch, 1790 – Bigeye snapper; Vivaneau gros
yeux. – D is tr ib ut i o n : First record from Europa Island as
Lutianus lineolatus by Fourmanoir (1952: 186).
Lutjanus monostigma (Cuvier in Cuvier & Valenciennes, 1828) –
Onespot snapper; Vivaneau églefin. – Di s tr i b ut i on : New
record from Europa Island, based on an observation by P.
durville et al. during the expedition MD-11; also observed
by P. durville & P. chaBanet on 7–11 Nov. 2011, St. B8,
B14, EU5, EU7, and by R. Fricke on 11 Nov. 2011, St. EU5.
Lutjanus rivulatus (Cuvier in Cuvier & Valenciennes, 1828) –
Scribbled snapper, speckled snapper; Lèvre de boeuf, bon-
homme tombé (Réunion), vivaneau maori (France). – D i s-
tribution: New record from Europa Island, based on an
observation by P. durville et al. during the expedition MD-
11; also observed by P. durville, 10 Nov. 2011, St. B8.
Lutjanus sebae (Cuvier, 1816) – Emperor snapper; Vivaneau
bourgeois. – D i st r ib u ti o n: First record from Europa Is-
land as Lutianus sebae by Fourmanoir (1952: 184).
Macolor niger (Forsskål in Niebuhr, 1775) – Black snapper,
black-and-white snapper; Vivaneau plate. – Di s t r i b u -
tion: New record from Europa Island, based on an observa-
tion by P. durville et al. during the expedition MD-11; also
observed and photographed by P. durville on 7–11 Nov.
2011, St. EU7, and among the mangroves in the inner lagoon.
Pristipomoides filamentosus (Valenciennes in Cuvier & Valenci-
ennes, 1830) – Rosy jobfish, blue-spotted jobfish, crimson
jobfish; Vivaneau commun, vivaneau blanc (Réunion), colas
fil (France). – D is t ri bu t i o n: First record from Europa Is-
land by Fourmanoir (1952: 184).

stuttGarter BeiträGe zur naturkunde a
260 Neue Serie 6
Caesionidae
Caesio caerulaurea Lacepède [ex Commerson], 1801 – Scissor-
tailed fusilier; Maquereau (Réunion), caesio azuror (France).
– Distribution: New record from Europa Island, based
on an observation by P. durville et al. during the expedition
MD-11.
Caesio lunaris Cuvier [ex Ehrenberg] in Cuvier & Valenciennes,
1830 – Blue fusilier; Caesio à croissant. – D i s t r i bu t i o n :
New record from Europa Island, based on an observation by
P. durville et al. during the expedition MD-11; also observed
by P. durville, 10–12 Nov. 2011, St. B5, B12, B14, EU2.
Caesio teres Seale, 1906 – Beautiful fusilier, yellowtail fusilier;
Caesio à dos jaune et bleu. – D i s t ri b u t io n: New record
from Europa Island, based on an observation by P. durville
et al. during the expedition MD-11; also observed by P. dur-
ville & P. chaBanet on 7–12 Nov. 2011, St. B12, EU2, EU7.
Caesio xanthonota Bleeker, 1853 – Yellowback fusilier; Caesio à
dos jaune. – Distribution: New record from Europa Is-
land, based on an observation by P. durville et al. during the
expedition MD-11; also observed by P. durville & P. cha-
Banet on 7–11 Nov. 2011, St. B14, EU7; record confirmed
based on photograph by E. BretaGne, taken in November
2011.
Pterocaesio chrysozona (Cuvier [ex Kuhl & Hasselt] in Cuvier
& Valenciennes, 1830) – Goldband fusilier; Caesio ceinture
d’or. – Distribution: New record from Europa Island,
based on an observation by P. durville et al. during the ex-
pedition MD-11.
Pterocaesio pisang (Bleeker, 1853) – Banana fusilier; Caesio
rose. – Distribution: New record from Europa Island,
based on an observation by P. durville et al. during the ex-
pedition MD-11.
Ptercaesio tile (Cuvier in Cuvier & Valenciennes, 1830) – Blues-
treak fusilier, dark-banded fusilier; Fusilier à ligne olive. –
Distribution: New record from Europa Island, based
on an observation by P. durville et al. during the expedition
MD-11; also observed by P. durville & P. chaBanet on 7
Nov. 2011, St. EU7.
Gerreidae
Gerres longirostris (Lacepède, 1801) – Longtail silver-biddy;
Blanche gouvernail (Fig. 11). – Di s t r i b u t i o n : New re-
cord from Europa Island, based on a photograph taken by P.
Borsa during an expedition with R/V Marion Dufresne on 7
April 2011.
Gerres oyena (Forsskål in Niebuhr, 1775) – Common silverbiddy,
common mojarra; Blanche commune. – D i s t r i b u t i o n :
First record from Europa Island by Fourmanoir (1952: 170,
pl. 12A); record confirmed by P. durville et al. during the ex-
pedition MD-11, an observation by P. Borsa in the mangrove
area on 7 April 2011, and by R. Fricke based on observations
at Europa on 7–10 Nov. 2011, St. B28, B31, B33, B34, B37,
B50.
Haemulidae
Diagramma picta (Thunberg, 1792) – Sailfin rubberlip; dia-
gramme voilier. – Di s t r i b u t i o n : New record from Eu-
ropa Island, based on photograph by E. BretaGne, taken in
November 2011.
Plectorhinchus gaterinus (Forsskål in Niebuhr, 1775) – Black-
spotted sweetlips; Gueule pavée (Réunion), diagramme
moucheté (France). – D i s t r i b u t i o n : New record from
Europa Island, based on an observation by P. durville et al.
during the expedition MD-11; also observed by P. durville
& P. chaBanet on 7 Nov. 2011, St. EU7.
Plectorhinchus obscurus (Günther, 1872) – Giant sweetlips;
Loche casteix, lèvre en caoutchouc. – D i st r ib u t io n : New
record from Europa Island, based on an observation by P.
durville et al. during the expedition MD-11; also observed
by P. durville & P. chaBanet on 7–12 Nov. 2011, St. B8,
B12, B14, B25, EU5, EU7, and in a pond on the south side of
the island; and by R. Fricke on 11 Nov. 2011, St. EU5.
Plectorhinchus vittatus (Linnaeus, 1758) – Indian Ocean oriental
sweetlips; Diagramme à bandes horizontales. – D i st ri bu -
tion: New record from Europa Island, based on an observa-
tion as Plectorhinchus orientalis by P. durville et al. during
the expedition MD-11, and by R. Fricke on 11 Nov. 2011,
St. EU5.
Sparidae
Acanthopagrus bifasciatus (Forsskål in Niebuhr, 1775) – Twobar
seabream; Pagre double bande. – D i s t ri b u t io n : First re-
cord from Europa Island by Fourmanoir (1952: 186).
Rhabdosargus sarba (Forsskål in Niebuhr, 1775) – Natal stump-
nose, gold-lined seabream; Gueule pavée (Réunion), sargue
doré (France). – D i s t r i b u t i o n : Recorded from Europa
Island as Acanthopagrus sarba by Fourmanoir (1952: 174),
and as Rhabdosargus sarba by Fourmanoir (1952: pl. 12B).
Lethrinidae
Gnathodentex aureolineatus (Lacepède [ex Commerson], 1802)
– Yellowspot emperor, glowfish; Petit clair (Réunion), em-
pereur strié (France) (Fig. 13). – D i s t r i b u t i o n : New
record from Europa Island, based on an observation by P.
durville et al. during the expedition MD-11.
Lethrinus nebulosus (Forsskål in Niebuhr, 1775) – Spangled em-
peror; Capitaine blanc (Réunion), empereur moris (France).
– Distribution: New record from Europa Island, based
on a photograph by P. durville on 11 Nov. 2011 near the
mangroves in the inner lagoon of Europa Island.
Lethrinus obsoletus (Forsskål in Niebuhr, 1775) – Orange-striped
emperor; Bossu d’herbe. – D i s t r i b u t i o n : New record
from Europa Island, based on an observation by P. durville
et al. during the expedition MD-11, and a photograph taken
by P. durville on 12 Nov. 2011, St. EU5.
Lethrinus olivaceus Valenciennes in Cuvier & Valenciennes,
1830 – Longface emperor; Bec de cane allongé, bec de cane
malabar, lethrinus à museau long. – Di st ri bu ti on : New
record from Europa Island, based on an observation by P.
durville et al. during the expedition MD-11.
Lethrinus xanthochilus Klunzinger, 1870 – Yellowlip emperor;
Empereur bec de cane. – D i s tr i bu t i on : New record from
Europa Island, based on an observation by P. durville et al.
during the expedition MD-11; also observed by P. durville,
10 Nov. 2011, St. B8, EU2.
Monotaxis grandoculis (Forsskål in Niebuhr, 1775) – Bigeye em-
peror, bigeye barenose; Gueule pavée (Réunion), empereur

Fricke et alii, checklist oF the shore Fishes oF euroPa island 261
bossu (France) (Fig. 12). – D i s t r i b u t i o n : New record
from Europa Island, based on an observation by P. durville
et al. and photograph by P. Borsa at St. 1 (equals St. EU5 of
this paper) during an expedition with R/V Marion Dufresne
on 5 April 2011; also observed by P. durville & P. chaBanet
on 7–11 Nov. 2011, St. B8, B14, EU2, EU7.
Mullidae
Mulloidichthys ayliffe (Uiblein, 2011) – Western Indian Ocean
yellowfin goatfish, Western Indian Ocean flame goatfish;
Capucin queue jaune. – D i s t r i bu ti on : New record from
Europa Island, based on an observation by P. durville et al.
during the expedition MD-11. – R e m a r k s : This is taxo-
nomically the same species that was recorded from Réunion
by Fricke et al. (2009: 68) as Mulloidichthys auriflamma
(non Forsskål in Niebuhr, 1775).
Mulloidichthys flavolineatus (Lacepède [ex Commerson], 1801)
– Yellowstripe goatfish; Capucin (Réunion), capuciva à
bande jaune (France). – D is t r i b u t i o n : New record from
Europa Island, based on an observation by P. durville et al.
during the expedition MD-11; also photographed by P. dur-
ville on 11 Nov. 2011 at Europa Island.
Mulloidichthys vanicolensis (Valenciennes in Cuvier & Valen-
ciennes, 1831) – Yellowfin goatfish; Surmullet de Vanicolo
(France) (Fig. 13). – D is t ri b ut i on : New record from Eu-
ropa Island, based on a photograph by P. durville on 11 Nov.
2011 near the mangroves of Europa Island.
Parupeneus barberinus (Lacepède [ex Commerson], 1801) –
Dash-and-dot goatfish; Capucin sergent (Réunion), rouget-
barbet barberin (France). – D i s t r i b u t i o n : New record
from Europa Island, based on an observation by P. durville
et al. during the expedition MD-11, and on observations by
R. Fricke on 7–11 Nov. 2011, St. B31, B37, B50, EU5; also
observed by P. durville, 12 Nov. 2011, St. EU5.
Parupeneus cyclostomus (Lacepède [ex Commerson], 1801) –
Yellowsaddle goatfish; Capucin malbar (Réunion), rouget-
barbet doré (France). – D i st ri bu ti on : New record from
Europa Island, based on an observation by P. durville et al.
at St. 1 (equals St. EU5 of this paper) during an expedition
with R/V Marion Dufresne on 5 April 2011; also observed by
P. durville & P. chaBanet on 7–11 Nov. 2011, St. B5, EU3,
EU7, and by R. Fricke on 11 Nov. 2011, St. EU5.
Parupeneus macronemus (Lacepède [ex Commerson], 1801) –
Longbarbel goatfish, band-dot goatfish; Rouget-barbet ban-
deau. – D i s t r i b u t i o n : First record from Europa Island
as Pseudopeneus macronema by Fourmanoir (1952: 184);
record confirmed by P. durville et al. during the expedition
MD-11; also observed by P. durville & P. chaBanet on 9–11
Nov. 2011, St. EU3, B5, B14, EU2.
Parupeneus pleurostigma (Bennett, 1831) – Blackspot goatfish,
sidespot goatfish; Rouget-barbet pastille. – D i s tr i b u ti o n :
New record from Europa Island, based on an observation by
P. durville et al. during the expedition MD-11.
Parupeneus trifasciatus (Lacepède [ex Commerson], 1801)
– Two-barred goatfish; Capucin (Réunion), rouget-barbet
double-tache (France). – D i s t r i b u t i o n : New record
from Europa Island, based on an observation by P. durville
et al. during the expedition MD-11; also observed by P. dur-
ville & P. chaBanet on 7–12 Nov. 2011, St. B5, B8, B12,
B14, EU2, EU3, EU5, EU7, and by R. Fricke on 10–11 Nov.
2011, St. B50, EU5.
Pempheridae
Parapriacanthus ransonneti Steindachner, 1870 – Slender
sweeper, pygmy sweeper. – D i s t r i b u t i o n : New record
from Europa Island, based on an observation by P. durville
et al. during the expedition MD-11.
Pempheris sp. (aff. Pempheris nesogallica Cuvier in Cuvier &
Valenciennes, 1831). – D i s t r i b u t i o n : Observed at Eu-
ropa Island by P. durville et al. during the expedition MD-
11; also observed by P. durville & P. chaBanet on 7 Nov.
2011, St. EU7.
Pempheris schwenkii Bleeker, 1855 – Blackstripe sweeper. –
Distribution: New record from Europa Island, based
on an observation by P. durville et al. during the expedition
MD-11.
Kyphosidae
Kyphosus bigibbus Lacepède [ex Commerson], 1801 – Grey
chub, brown chub. – D i s t r i b u t i o n : New record from
Europa Island, based on a photograph taken by E. BretaGne
in Oct. 2011, a photograph by P. durville taken on 11 Nov.
2011 among the mangroves in the inner lagoon, and on ob-
servations by R. Fricke on 10–11 Nov. 2011, St. B50, EU5.
Kyphosus cinerascens (Forsskål in Niebuhr, 1775) – Blue sea-
chub, lowfin rudderfish. – D i s t r i b u t i o n : New record
from Europa Island, based on an observation by P. durville
et al. during the expedition MD-11; previously listed as Ky-
phosus sp. by Fourmanoir (1952: 170); also observed by P.
durville & P. chaBanet on 7–12 Nov. 2011, St. EU7, B25.
Monodactylidae
Monodactylus argenteus (Linnaeus, 1758) – Silver moony, mono,
Natal moony; Poisson lune argenté. – D is t r i bu t io n: First
record from Europa Island by Fourmanoir (1952: 174); re-
cord confirmed by P. durville et al. during the expedition
MD-11, and a photograph taken by P. Borsa in the mangrove
area on 7 April 2011; also observed by P. durville, 12 Nov.
2011, in a pond on the south side of the island.
Chaetodontidae
Chaetodon auriga Forsskål in Niebuhr, 1775 – Threadfin but-
terflyfish (Fig. 14). – D i s t r i b u t i o n : First record from
Europa Island by Fourmanoir (1952: 172); record confirmed
by P. durville et al. based on a observation at St. 2 (equals
St. B34 of this paper) during an expedition with R/V Marion
Dufresne on 6 April 2011, and an observation by P. Borsa
in the mangrove area on 7 April 2011; also observed by P.
durville & P. chaBanet on 7–12 Nov. 2011, St. B5, B8, B12,
B14, EU2, EU3, EU4, EU5, EU7, and by R. Fricke on 8–10
Nov. 2011, St. B34, B37, B49, B50.
Chaetodon bennetti Cuvier in Cuvier & Valenciennes, 1831 –
Bluelashed butterflyfish; Papillon de Bennett. – D is t r i b u -
tion: New record from Europa Island, based on observation
by P. durville & P. chaBanet on 7–9 Nov. 2011, St. EU3,
EU7.
Chaetodon falcula Bloch, 1795 – Blackwedged butterflyfish;
Papillon indien à deux selles. – D i s t r i b u t i o n : New re-

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262 Neue Serie 6
cord from Europa Island, based on an observation by P. dur-
ville et al. during the expedition MD-11, and by R. Fricke on
11 Nov. 2011, St. EU5; also observed by P. durville, 10 Nov.
2011, St. EU2.
Chaetodon guttatissimus Bennett, 1833 – Peppered butterflyfish,
spotted butterflyfish, gorgeous gussy; Chétodon moucheté.
– Distribution: New record from Europa Island, based
on an observation by P. durville et al. during the expedition
MD-11; also observed by P. durville & P. chaBanet on 7–12
Nov. 2011, St. B8, B12, B14, EU2, EU3, EU7.
Chaetodon interruptus Ahl, 1923 – Teardrop butterflyfish, lime-
spot butterflyfish; Chétodon à tache de citron. – D is tr ib u-
tion: New record from Europa Island, based on an observa-
tion by P. durville et al. during the expedition MD-11; also
observed by P. durville & P. chaBanet on 9–12 Nov. 2011,
St. B5, EU2, EU3, EU5.
Chaetodon kleinii Bloch, 1790 – Klein’s butterflyfish, whitespot-
ted butterflyfish; Papillon (Réunion). – D i s t r i b u t i o n :
New record from Europa Island, based on an observation by
P. durville et al. during the expedition MD-11, and by R.
Fricke on 10 Nov. 2011, St. B37.
Chaetodon lineolatus Cuvier [ex Quoy & Gaimard] in Cuvier &
Valenciennes, 1831 – Lined butterflyfish; Chétodon linéolé.
– Distribution: New record from Europa Island, based
on an observation by R. Fricke on 7–10 Nov. 2011, St. B31,
B50; also observed by P. durville, 10 Nov. 2011, St. B8.
Chaetodon lunula (Lacepède [ex Commerson], 1802) – Racoon
butterflyfish, halfmoon butterflyfish; Chétodon à croissant. –
D i st ri bu ti on : First record from Europa Island by Four-
manoir (1952: 172); record confirmed by P. durville et al.
during the expedition MD-11; and by R. Fricke based on ob-
servations and specimens collected at Europa on 7–12 Nov.
2011, St. B28, B31, B33, B34, B37, B50, EU5; also observed
by P. durville & P. chaBanet on 7–12 Nov. 2011, St. B5, B8,
B12, B14, B25, EU2, EU3, EU5, EU7. – M a te ri al : SMF
34751 (1), SMF 34794 (2).
Chaetodon madagaskariensis Ahl, 1923 – Pearly butterflyfish;
Chétodon de Madagascar. – Di st ri bu ti on : New record
from Europa Island, based on an observation by P. durville
et al. during the expedition MD-11; also observed by P. dur-
ville & P. chaBanet on 9 Nov. 2011, St. EU3.
Chaetodon melannotus Bloch & Schneider, 1801 – Black-backed
butterflyfish; Chétodon à dos noir. – Di st ri bu ti on : New
record from Europa Island, based on observation by P. dur-
ville & P. chaBanet on 9 Nov. 2011, St. EU3.
Chaetodon meyeri Schneider [ex Renard] in Bloch & Schneider,
1801 – Meyer’s butterflyfish, scrawled butterflyfish. – D i s -
tribution: New record from Europa Island, based on an
observation by P. durville et al. during the expedition MD-
11; also observed by P. durville & P. chaBanet on 7–12 Nov.
2011, St. EU3, EU4, EU7, B5, B8, B12, B14, EU2, EU5.
Chaetodon trifascialis Quoy & Gaimard, 1825 – Chevroned but-
terflyfish, rightangle butterflyfish. – D i st r ib u ti o n: New
record from Europa Island, based on an observation by P.
durville et al. during the expedition MD-11; also observed
by P. durville & P. chaBanet on 7–12 Nov. 2011, St. B12,
EU2, EU3, EU7.
Chaetodon trifasciatus Park, 1797 – Redfin butterflyfish, purple
butterflyfish, melon butterflyfish (Fig. 15). – D i s t r i b u -
tion: New record from Europa Island, based on an observa-
tion by P. durville et al. at St. 2 (equals St. B34 of this paper)
during an expedition with R/V Marion Dufresne on 6 April
2011, and on an observation by R. Fricke on 7–11 Nov. 2011,
St. B31, B49, B50, EU5; also observed by P. durville & P.
chaBanet on 7–12 Nov. 2011, St. B5, B12, B14, EU5, EU7.
Chaetodon vagabundus Linnaeus, 1758 – Vagabond butterfly-
fish; Chétodon à vagabond. – D is tr i b u t i o n : New record
from Europa Island, based on an observation by P. durville
et al. during the expedition MD-11, and by R. Fricke on 11
Nov. 2011, St. EU5; also observed by P. durville, 12 Nov.
2011, St. B12, EU5.
Chaetodon xanthocephalus Bennett, 1833 – Yellowhead butter-
flyfish; Chétodon à tête jaune (Fig. 16). – D is t ri b ut io n:
New record from Europa Island, based on an observation by
P. durville et al. at St. 2 (equals St. B34 of this paper) during
an expedition with R/V Marion Dufresne on 6 April 2011,
and by R. Fricke based on an observation at Europa on 7–11
Nov. 2011, St. B33, B50, EU5; also observed by P. durville
& P. chaBanet on 7–11 Nov. 2011, St. B5, EU2, EU3, EU7.
Chaetodon zanzibarensis Playfair in Playfair & Günther, 1867 –
Zanzibar butterflyfish; Chétodon de Zanzibar. – Di st ri bu -
tion: New record from Europa Island, based on an observa-
tion by P. durville et al. during the expedition MD-11.
Forcipiger flavissimus Jordan & McGregor in Jordan & Ever-
mann, 1898 – Long-nosed butterflyfish; Papillon longnez.
– Distribution: New record from Europa Island, based
on an observation by P. durville et al. during the expedition
MD-11; also observed by P. durville & P. chaBanet on 9–10
Nov. 2011, St. B8, EU3.
Hemitaurichthys zoster (Bennett, 1831) – Black pyramid but-
terflyfish, brushtooth butterflyfish. – D i s t ri bu t i o n: New
record from Europa Island, based on an observation by P.
durville et al. during the expedition MD-11.
Heniochus acuminatus (Linnaeus, 1758) – Coachman, longfin
bannerfish; Pavillon (Réunion). – D i s t r i b u t i o n : New
record from Europa Island, based on an observation by P.
durville et al. during the expedition MD-11; also observed
by P. durville, 11–12 Nov. 2011, St. B5, B12, B14.
Heniochus monoceros Cuvier in Cuvier & Valenciennes, 1831 –
Masked bannerfish, masked coachman; Pavillon (Réunion),
hénioche cornu (France). – D i s t r i b u t i o n : New record
from Europa Island, based on an observation by P. durville
et al. during the expedition MD-11; also observed by P. dur-
ville, 10–11 Nov. 2011, St. B5, EU2.
Pomacanthidae
Apolemichthys trimaculatus (Cuvier [ex Lacepède] in Cuvier &
Valenciennes, 1831) – Threespot angelfish, flagfin angelfish;
Poisson-ange à trois taches. – D is t r i bu ti o n : New record
from Europa Island, based on an observation by P. durville
et al. during the expedition MD-11; also observed by P. dur-
ville & P. chaBanet on 7–11 Nov. 2011, St. B5, B8, EU3,
EU4, EU7.
Centropyge acanthops (Norman, 1922) – African pygmy angel-
fish, jumping bean, orangeback angelfish. – D i s t r i b u -
tion: New record from Europa Island, based on an observa-
tion by P. durville et al. during the expedition MD-11.

Fricke et alii, checklist oF the shore Fishes oF euroPa island 263
Centropyge bispinosa (Günther, 1860) – Coral beauty; Poisson-
ange à deux épines. – D i s t r i b u t i o n : New record from
Europa Island, based on an observation by P. durville et al.
during the expedition MD-11; also observed by P. durville,
10–11 Nov. 2011, St. B8, B14, EU2.
Centropyge multispinis (Playfair in Playfair & Günther, 1867)
– Dusky angelfish, dusky cherub. – D i s t r i b u t i o n : New
record from Europa Island, based on an observation by P.
durville et al. during the expedition MD-11; also observed
by P. durville & P. chaBanet on 7–12 Nov. 2011, St. B5, B8,
B12, B14, EU2, EU3, EU4, EU7, and by R. Fricke on 10–11
Nov. 2011, St. B50, EU5.
Pomacanthus chrysurus (Cuvier in Cuvier & Valenciennes, 1831)
– Goldtail angelfish; Poisson-ange à oreille tachée. – D i s -
tribution: New record from Europa Island, based on an
observation by P. durville et al. during the expedition MD-
11; also observed by P. durville & P. chaBanet on 9 Nov.
2011, St. EU3.
Pomacanthus imperator (Bloch, 1787) – Emperor angelfish;
Poisson-ange impérial; Poisson-ange imérial. – D i st ri bu -
tion: New record from Europa Island, based on an observa-
tion by P. durville et al. during the expedition MD-11; also
observed by P. durville & P. chaBanet on 7–12 Nov. 2011,
St. B5, B14, EU3, EU5, EU7.
Pomacanthus semicirculatus (Cuvier in Cuvier & Valenciennes,
1831) – Semicircle angelfish; Poisson-ange bleu. – D i s t r i -
bution: New record from Europa Island, based on an ob-
servation by P. durville, 10–11 Nov. 2011, St. B5, B8.
Pygoplites diacanthus (Boddaert, 1772) – Regal angelfish; Pois-
son-ange à bandes bleues, poisson duc. – D i st r ib u ti o n:
New record from Europa Island, based on an observation by
P. durville et al. during the expedition MD-11; also observed
by P. durville & P. chaBanet on 7–11 Nov. 2011, St. B12,
B14, EU2, EU7.
Cirrhitidae
R em ar k s : A record of Paracirrhites sp. by Fourmanoir (1952:
183) could not be identified to the species level.
Amblycirrhitus bimacula (Jenkins, 1903) – Twospot hawkfish.
– Distribution: New record from Europa Island, based
on observation by P. durville & P. chaBanet on 9–11 Nov.
2011, St. B5, EU4.
Cirrhitichthys oxycephalus (Bleeker, 1855) – Coral hawkfish;
Épervier lutin. – D is t ri bu t io n: New record from Europa
Island, based on an observation by P. durville et al. during
the expedition MD-11; also observed and photographed by P.
durville on 7–12 Nov. 2011, St. EU3, B12, B14, EU4, EU5,
EU7, and by R. Fricke on 9 Nov. 2011, St. B28.
Cirrhitus pinnulatus (Bloch & Schneider [ex Forster], 1801) –
Stocky hawkfish; Domingue (Réunion). – D i s t ri bu ti on :
New record from Europa Island, based on an observation by
R. Fricke on 9 Nov. 2011, St. B28, and by a photograph by P.
chaBanet on the NE side of the island.
Paracirrhites arcatus (Cuvier [ex Parkinson] in Cuvier & Va-
lenciennes, 1829) – Arc-eye hawkfish, horseshoe hawkfish.
– Distribution: New record from Europa Island, based
on an observation by P. durville et al. during the expedi-
tion MD-11; also observed by P. durville & P. chaBanet on
7–12 Nov. 2011, St. B5, B8, B12, B14, EU2, EU3, EU4, EU5,
EU7, and by R. Fricke on 11 Nov. 2011, St. EU5.
Paracirrhites forsteri (Schneider in Bloch & Schneider, 1801) –
Freckled hawkfish, blackside hawkfish. – D i st ri bu ti on :
New record from Europa Island, based on an observation by
P. durville et al. during the expedition MD-11; also observed
by P. durville & P. chaBanet on 7–12 Nov. 2011, St. B5, B8,
B12, B14, EU2, EU3, EU5, EU7.
Pomacentridae
Abudefduf notatus (Day, 1870) – Yellowtail sergeant. – D i s t r i-
bution: New record from Europa Island, based on an ob-
servation by P. durville et al. during the expedition MD-11;
also observed and collected by P. durville & P. chaBanet
on 7–12 Nov. 2011, St. B25, EU7, and in a pond on the south
side of the island; and by R. Fricke based on an observation
and specimens collected on 9–11 Nov. 2011, St. B28. – Ma -
t er ia l: SMF 34732 (1), SMF 34771 (1), SMF 34835 (1).
Abudefduf septemfasciatus (Cuvier in Cuvier & Valenciennes,
1830) – Banded sergeant, sevenbar damsel; Sergent-major.
– Distribution: New record from Europa Island, based
on an observation and a photograph by P. durville, 12 Nov.
2011, St. B25.
Abudefduf sexfasciatus (Lacepède [ex Commerson], 1801) –
Scissortail sergeant, stripetail damsel. – D i s t r i b u t i o n :
New record from Europa Island, based on an observation by
P. durville et al. during the expedition MD-11.
Abudefduf sordidus (Forsskål in Niebuhr, 1775) – Blackspot ser-
geant, spot damsel; Sergent à tache noire. – D i s t ri b u t i on :
First record from Europa Island by Fourmanoir (1952: 170);
record confirmed by P. durville et al. based on a observation
at St. 2 (equals St. B34 of this paper) during an expedition
with R/V Marion Dufresne on 6 April 2011, an observation
by P. Borsa in the mangrove area on 7 April 2011, and by R.
Fricke on 9 Nov. 2011, St. B28; also observed by P. durville,
12 Nov. 2011, St. B25.
Abudefduf sparoides (Quoy & Gaimard, 1825) – False-eye ser-
geant, false-eye damsel. – D i s t r i b u t i o n : New record
from Europa Island, based on an observation by P. durville
et al. at St. 2 (equals St. B34 of this paper) during an ex-
pedition with R/V Marion Dufresne on 6 April 2011; record
confirmed by R. Fricke based on observations on 7–10 Nov.
2011, St. B28, B32, B33, B34, B37, B49.
Abudefduf vaigiensis (Quoy & Gaimard, 1825) – Indo-Pacific
sergeant. – Distribution: New record from Europa Is-
land, previously reported as A. saxatilis (non Linnaeus, 1758)
by Fourmanoir (1952: 170); record confirmed by P. durville
et al. based on a observation at St. 2 (equals St. B34 of this
paper) during an expedition with R/V Marion Dufresne on
6 April 2011, and by R. Fricke based on observations and
specimens collected at Europa on 7–12 Nov. 2011, St. B28,
B31, B32; also observed by P. durville, 12 Nov. 2011,
St. B25. – M a t e r i a l : SMF 34733 (2), SMF 34772 (1),
SMF 34813 (1).
Amphiprion akallopisos Bleeker, 1853 – Skunk clownfish; Pois-
son-clown mouflette. – D i s t r i b u t i o n : New record from
Europa Island, based on the observation of a single specimen
by P. durville et al. during the expedition MD-11. – R e -
m a r k s : This is the most common species of anemonefish
at Europa Island.

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264 Neue Serie 6
Amphiprion allardi Klausewitz, 1970 – Twobar anemonefish,
Allard’s anemonefish; Poisson-clown d’Allard. – Di s t r i -
bution: New record from Europa Island; previously listed
as A. polymnus (non Linnaeus, 1758) by Fourmanoir (1952:
186); observed at Europa Island by P. durville et al. during
the expedition MD-11.
Amphiprion latifasciatus Allen, 1972 – Madagascar anemone-
fish; Poisson-clown de Madagascar. – Di st ri bu ti on :
New record from Europa Island, based on a single specimen
by P. durville et al. during the expedition MD-11.
Chromis agilis Smith, 1960 – Reef chromis, agile chromis. –
Distribution: New record from Europa Island, based on
an observation by P. durville & P. chaBanet on 7 Nov. 2011,
St. EU7.
Chromis dimidiata (Klunzinger, 1871) – Chocolate dip; Demoi-
selle bicolore. – D i s t r i b u t i o n : New record from Eu-
ropa Island, based on an observation by P. durville et al.
during the expedition MD-11; also observed by P. durville
& P. chaBanet on 7–12 Nov. 2011, St. B5, B8, B12, B14,
EU2, EU3, EU5, EU7, and by R. Fricke on 8–11 Nov. 2011,
St. B49, B50, EU5.
Chromis nigrura Smith, 1960 – Blacktail chromis. – D is tr i bu -
tion: New record from Europa Island, based on an observa-
tion by P. durville et al. during the expedition MD-11; also
observed by P. durville & P. chaBanet on 7–12 Nov. 2011,
St. B5, B8, B12, B14, EU2, EU3, EU4, EU5, EU7.
Chromis opercularis (Günther in Playfair & Günther, 1867) –
Doublebar chromis. – D i s t r i b u t i o n : New record from
Europa Island, based on an observation by P. durville et al.
during the expedition MD-11; also observed by P. durville
& P. chaBanet on 7–12 Nov. 2011, St. B5, B8, B12, B14,
EU2, EU3, EU7.
Chromis ternatensis (Bleeker, 1856) – Golden chromis, blue-
green chromis. – D i s tr i b ut i o n : New record from Europa
Island, based on an observation by P. durville et al. during
the expedition MD-11; also observed and photographed by P.
durville on 7–12 Nov. 2011, St. B12, B14, EU7.
Chromis viridis Cuvier [ex Ehrenberg] in Cuvier & Valenciennes,
1830 – Green chromis, blue-green chromis; Petit cafre (Ré-
union), demoiselle bleue (France). – D i st ri bu ti on : New
record from Europa Island, based on an observation by P.
durville et al. during the expedition MD-11, on photographs
by P. durville and P. chaBanet taken in Nov. 2011, and on
an observation by R. Fricke on 7–8 Nov. 2011, St. B31, B34,
B49.
Chromis weberi Fowler & Bean, 1928 – Weber’s chromis. – D i s -
tribution: New record from Europa Island, based on an
observation by P. durville et al. during the expedition MD-
11; also observed and photographed by P. durville on 7 Nov.
2011, St. EU7, and by R. Fricke on 11 Nov. 2011, St. EU5.
Chrysiptera brownriggii (Bennett, 1828) – Surge demoiselle,
blueribbon demoiselle. – D i s t r i b u t i o n : New record
from Europa Island, based on observations by P. durville et
al. at St. 1 and St. 2 (equals St. EU5 and B34 of this paper)
during an expedition with R/V Marion Dufresne on 5–6 April
2011, and by R. Fricke on 7–11 Nov. 2011, St. B28, B31,
B32, B33, B34, B37, B49, B50, EU5; also observed by P.
durville, 12 Nov. 2011, St. B25, EU5; previously listed as
Abudefduf biocellatus (non Quoy & Gaimard, 1825) by Four-
manoir (1952: 172).
Chrysiptera glauca (Cuvier in Cuvier & Valenciennes, 1830)
– Grey demoiselle, blue damsel. – D i s t r i b u t i o n : New
record from Europa Island, based on observations by P.
durville et al. at St. 1 and St. 2 (equals St. EU5 and B34 of
this paper) during an expedition with R/V Marion Dufresne
on 5–6 April 2011, and on observations and specimens col-
lected by R. Fricke on 7–12 Nov. 2011, St. B28, B31, B34,
B37, B50, EU5; also observed by P. durville, 12 Nov. 2011,
St. B25. – M a t e r i a l : SMF 34734 (1), SMF 34773 (1),
SMF 34814 (6).
Chrysiptera unimaculata (Cuvier in Cuvier & Valenciennes,
1830) – Onespot demoiselle; Demoiselle à une tache. – D is -
tribution: New record from Europa Island, based on an
observation by P. durville et al. during the expedition MD-
11, and on observations and specimens collected by R. Fricke
on 7–12 Nov. 2011, St. B28, B31, B32, B33, B34, B37, B49,
B50, EU5; also observed by P. durville, 12 Nov. 2011, St.
B25, EU5. – M at er ia l: SMF 34774 (1), SMF 34815 (10).
Dascyllus aruanus (Linnaeus, 1758) – Humbug dascyllus, zebra
humbug, whitetail dascyllus; Demoiselle à queue blanche. –
D i s t r i b u t i o n : First record from Europa Island as Das-
cyllus aruanus by Fourmanoir (1952: 174), and as Dascyl-
lus amanius by Fourmanoir (1952: 186); record confirmed
by P. durville et al. based on a observation at St. 2 (equals
St. B34 of this paper) during an expedition with R/V Marion
Dufresne on 6 April 2011, and by R. Fricke based on observa-
tions at Europa on 7–10 Nov. 2011, St. B31, B32, B33, B34,
B37, B49, B50.
Dascyllus carneus Fischer, 1885 – Twobar humbug, cloudy das-
cyllus. – Distribution: New record from Europa Island,
based on a photograph by P. durville taken on 11 Nov. 2011,
on the north side of the island below the boat mooring, at
18 m depth.
Dascyllus trimaculatus (Rüppell, 1829) – Domino, threespot das-
cyllus; Demoiselle à trois taches. – D i s t r i b u t i o n : New
record from Europa Island, based on an observation by P.
durville et al. at St. 2 (equals St. B34 of this paper) during
an expedition with R/V Marion Dufresne on 6 April 2011,
and by R. Fricke on 8–10 Nov. 2011, St. B34, B37, B50; also
observed by P. durville & P. chaBanet on 7–12 Nov. 2011,
St. B12, EU7.
Lepidozygus tapeinosoma (Bleeker, 1856) – Fusilier damselfish;
Poisson demoiselle bleu à dos jaune. – D i s t r i b u t i o n :
New record from Europa Island, based on an observation by
P. durville et al. during the expedition MD-11; also observed
by P. durville & P. chaBanet on 9–12 Nov. 2011, St. B8,
B12, B14, EU2, EU3, EU4.
Neopomacentrus fuliginosus (Smith, 1960) – African demoiselle.
– Distribution: New record from Europa Island, based
on an observation by P. durville et al. during the expedition
MD-11.
Plectroglyphidodon dickii (Liénard, 1839) – Narrowbar damsel,
Dick’s damsel, blackbar devil; Demoiselle de Dick. – Di s -
tribution: New record from Europa Island, based on an
observation by P. durville et al. during the expedition MD-
11; also observed by P. durville & P. chaBanet on 7–12 Nov.

Fricke et alii, checklist oF the shore Fishes oF euroPa island 265
2011, St. B5, B8, B12, B14, EU2, EU3, EU4, EU5, EU7, and
by R. Fricke on 8–11 Nov. 2011, St. B49, EU5.
Plectroglyphidodon imparipennis (Sauvage in Vaillant & Sau-
vage, 1875) – Brighteye damsel, stop-start damsel. – D i s -
tribution: New record from Europa Island, based on an
observation by P. durville et al. during the expedition MD-
11, and on observations and specimens collected by R. Fricke
on 7–12 Nov. 2011, St. B28, B32, EU5; also observed by P.
durville, 12 Nov. 2011, St. B25, EU5. – M a t e r i a l : SMF
34775 (2), SMF 34816 (1).
Plectroglyphidodon johnstonianus Fowler & Ball, 1924 – John-
ston damsel, widebar damsel. – D i s t r i b u t i o n : New re-
cord from Europa Island, based on an observation by P. dur-
ville et al. during the expedition MD-11; also observed by
P. durville & P. chaBanet on 7–12 Nov. 2011, St. B5, B12,
B14, EU2, EU3, EU4, EU7, and by R. Fricke on 8–11 Nov.
2011, St. B49, B50, EU5.
Plectroglyphidodon lacrymatus (Quoy & Gaimard, 1825) –
White-spotted devil; Bouteur perlée. – D i s t r i b u t i o n :
New record from Europa Island, based on a photograph by P.
durville taken on 12 Nov. 2011, St. EU5.
Plectroglyphidodon phoenixensis (Schultz, 1943) – Phoenix
devil. – Distribution: New record from Europa Island,
based on an photograph by P. chaBanet taken on 10 Nov.
2011, St. B23, and an observation and a specimen collected
by R. Fricke on 11 Nov. 2011, St. B28, EU5; also observed
by P. durville, 12 Nov. 2011, St. B25. – Ma t er i al : SMF
34776 (1).
Pomacentrus agassizii Bliss, 1883 – Creole damsel. – D i st r i-
bution: New record from Europa Island; previously listed
as Pomacentrus tripunctatus (non Cuvier in Cuvier & Valen-
ciennes, 1830) by Fourmanoir (1952: 172).
Pomacentrus caeruleus Quoy & Gaimard, 1825 – Caerulean
damsel; Bouteur bleu et jaune (Fig. 17). – D is tr ib ut io n:
New record from Europa Island, based on an observation
and specimens collected by R. Fricke on 8–10 Nov. 2011,
St. B34, B37, B50; also a photograph by P. durville, taken
on 12 Nov. 2011, St. EU5.
Pomacentrus pavo (Bloch, 1787) – Sapphire damsel; Bouteur
paon. – Distribution: New record from Europa Island,
based on an observation and a photograph by P. chaBanet
on 12 Nov. 2011, in the mangrove area in the inner lagoon of
Europa Island.
Stegastes luteobrunneus (Smith, 1960) – Indian Ocean dark dam-
sel. – Distribution: New record from Europa Island,
based on an observation by P. durville et al. during the ex-
pedition MD-11, and on observations by R. Fricke on 7–11
Nov. 2011, St. B31, B32, B34, B49, EU5.
Stegastes nigricans (Lacepède [ex Commerson], 1802) – Dusky
gregory; Négrillon. – D i s t r i b u t i o n : New record from
Europa Island, based on an observation by P. durville et al.
during the expedition MD-11.
Stegastes pelicieri Allen & Emery, 1985 – Mauritian gregory. –
Distribution: New record from Europa Island, based
on an observation by P. durville et al. during the expedition
MD-11; also observed by P. durville & P. chaBanet on 7–12
Nov. 2011, St. B5, B8, B12, B14, EU2, EU5, EU7, and by R.
Fricke on 8–11 Nov. 2011, St. B37, B49, B50, EU5.
Labridae
Anampses caeruleopunctatus Rüppell, 1829 – Blue-spotted
wrasse, blue-spotted tamarin; Labre constellé (female), labre
bleu (male). – D i s t r i b u t i o n : New record from Europa
Island, based on an observation by P. durville et al. during
the expedition MD-11; also observed by P. durville, 10–12
Nov. 2011, St. B5, B8, B12, B14, EU2.
Anampses meleagrides Valenciennes in Cuvier & Valenciennes,
1840 – Yellowtail tamarin; Labre à queue jaune. – D is t ri -
bution: New record from Europa Island, based on an ob-
servation by P. durville et al. during the expedition MD-11;
also observed by P. durville, 10–12 Nov. 2011, St. B8, B12,
B14.
Bodianus anthioides (Bennett, 1832) – Lyretail hogfish; Labre
lyre. – Distribution: New record from Europa Island,
based on an observation by P. durville et al. during the expe-
dition MD-11; also observed by P. durville & P. chaBanet
on 7 Nov. 2011, St. EU7.
Bodianus axillaris (Bennett, 1832) – Axilspot hogfish, turncoat
hogfish; Labre à tache pectorale. – D i s t r i b u t i o n : New
record from Europa Island, based on an observation by P.
durville et al. during the expedition MD-11; also observed
by P. durville & P. chaBanet on 7–11 Nov. 2011, St. B5, B8,
B14, EU2, EU7.
Bodianus bilunulatus (Lacepède [ex Commerson], 1801) – Sad-
dleback hogfish; Vieille à selle noire. – Di s t r i b u t i o n :
New record from Europa Island, based on an observation
by P. durville et al. during the expedition MD-11, and on
an observation by R. Fricke on 7 Nov. 2011, St. B32; also
observed by P. durville & P. chaBanet on 9–12 Nov. 2011,
St. B12, EU3.
Bodianus diana (Lacepède [ex Commerson], 1801) – Diana’s
hogfish; Labre diane. – D i st r i b u t i o n : First record from
Europa Island as Lepidaploïs diana by Fourmanoir (1952:
184); record confirmed by P. durville et al. during the ex-
pedition MD-11; also observed and photographed by P. dur-
ville on 7–12 Nov. 2011, St. B12, B14, EU2, EU5, EU7.
Cheilinus chlorourus (Bloch, 1791) – Floral wrasse; Vieille ta-
chetée. – Distribution: New record from Europa Island,
based on juvenile specimens collected by R. Fricke on 11–12
Nov. 2011, St. B28. – M a t e r i a l : SMF 34759 (1), SMF
34801 (1).
Cheilinus fasciatus (Bloch, 1791) – Red-breasted wrasse; Vieille
rayée. – Distribution: New record from Europa Island,
based on an observation by P. durville et al. during the ex-
pedition MD-11; also observed by P. durville, 12 Nov. 2011,
St. EU5.
Cheilinus oxycephalus Bleeker, 1853 – Snooty wrasse; Vieille à
bec pointu. – D i s t r i b u t i o n : New record from Europa
Island, based on an observation by P. durville et al. during
the expedition MD-11; also observed by P. durville, 10 Nov.
2011, St. EU2.
Cheilinus trilobatus Lacepède [ex Commerson], 1801 – Tripletail
wrasse; Lachaux (Réunion), vieille triple queue, vieille tri-
lobé (France). – D is tr ib ut io n: New record from Europa
Island, based on an observation by P. durville et al. during
the expedition MD-11, and by R. Fricke on 10–11 Nov. 2011,
St. B50, EU5; also observed by P. durville, 11 Nov. 2011,
St. B5.

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266 Neue Serie 6
Cheilinus undulatus Rüppell, 1835 – Humphead wrasse;
Napoléon. – Distribution: New record from Europa Is-
land, based on an observation by P. durville et al. during the
expedition MD-11; record confirmed based on photograph by
E. BretaGne, taken in November 2011; also observed by P.
durville, 12 Nov. 2011, St. B12.
Cirrhilabrus exquisitus Smith, 1957 – Exquisite wrasse; Labre
exquis. – Distribution: New record from Europa Island,
based on an observation by P. durville et al. during the expe-
dition MD-11; also observed by P. durville & P. chaBanet
on 9–12 Nov. 2011, St. B12, B14, EU2, EU3.
Coris aygula Lacepède [ex Commerson], 1801 – Clown coris;
Loupe côte (Réunion), clarisse clown (France). – D i s t r i -
bution: New record from Europa Island, based on an ob-
servation by P. durville et al. during the expedition MD-11;
also observed by P. durville, 10 Nov. 2011, St. EU2.
Coris caudimacula (Quoy & Gaimard, 1834) – Spottail coris;
Girelle à tache caudale. – D i s t r i b u t i o n : New record
from Europa Island, based on a photograph taken by P. dur-
ville on 12 Nov. 2011, St. EU5.
Coris cuvieri (Bennett, 1831) – African coris; Clarisse africaine.
– Distribution: New record from Europa Island, based
on an observation by P. durville et al. during the expedition
MD-11, by R. Fricke on 11 Nov. 2011, St. EU5, and on a
photograph of a juvenile taken by P. durville, St. EU6; also
observed by P. durville, 10 Nov. 2011, St. B8.
Cymolutes praetextatus (Quoy & Gaimard, 1834) – Knife wrasse,
knife razorfish. – D i s t r i b u t i o n : New record from Eu-
ropa Island, based on a photograph by P. durville taken on
11 Nov. 2011 among the mangroves in the inner lagoon of
Europa Island.
Epibulus insidiator (Pallas, 1770) – Slingjaw wrasse; Epibule
gourami. – Distribution: New record from Europa Is-
land, based on an observation by P. durville et al. during
the expedition MD-11; also observed by P. durville, 11 Nov.
2011, St. B14.
Gomphosus caeruleus Lacepède [ex Commerson], 1801 – Indian
Ocean bird wrasse, birdfish; Poisson oiseau (Fig. 13). – Di s -
t r i b u t i o n: First record from Europa Island as Gomphosus
coerulens, Gomphosus varius and Gomphosus sp. by Four-
manoir (1952: 184); record confirmed by P. durville et al.
based on a observation at St. 1 (equals St. EU5 of this paper)
during an expedition with R/V Marion Dufresne on 5 April
2011, and by R. Fricke based on observations at Europa on
7–11 Nov. 2011, St. B28, B31, B32, B33, B34, B37, B49,
B50, EU5; also observed by P. durville & P. chaBanet on
9–12 Nov. 2011, St. B5, B8, B12, B14, EU2, EU3, EU4, EU5.
Halichoeres cosmetus Randall & Smith, 1982 – Adorned wrasse;
Girelle à raies d’or. – Di st ri bu ti on : New record from
Europa Island, based on an observation by P. durville et al.
during the expedition MD-11; also observed by P. durville
& P. chaBanet on 9–12 Nov. 2011, St. B5, B12, EU2, EU3.
Halichoeres hortulanus (Lacepède [ex Commerson], 1801) –
Checkerboard wrasse; Bonpartère (Réunion), lalo damier
(France). – D i s t r i b u t i o n : First record from Europa Is-
land as Halichoeres centriquadrus by Fourmanoir (1952:
184); record confirmed by P. durville et al. during the ex-
pedition MD-11, and by R. Fricke based on an observation
at Europa on 7–11 Nov. 2011, St. B28, B31, B32, B34, B37,
B49, B50, EU5; also observed by P. durville & P. chaBanet
on 7–12 Nov. 2011, St. B5, B8, B12, B14, EU2, EU5, EU7.
Halichoeres lapillus Smith, 1947 – Jewelled wrasse; Girelle à pe-
tits cailloux. – D i s t r i b u t i o n : New record from Europa
Island, based on an observation by R. Fricke on 8 Nov. 2011,
St. B34.
Halichoeres nebulosus (Valenciennes in Cuvier & Valenciennes,
1839) – Nebulous wrasse; Girelle nébuleuse. – D i st r ib u-
tion: New record from Europa Island, based on observa-
tions and specimens collected by R. Fricke on 8–11 Nov.
2011, St. B28, B34, B37, B50, EU5, and a specimen collected
by P. durville & P. chaBanet on 12 Nov. 2011, St. B25. –
M a t e r i a l : SMF 34730 (1), SMF 34760 (2), SMF 34834
(1).
Halichoeres nigrescens (Bloch & Schneider, 1801) – Bubblefin
wrasse. – Distribution: New record from Europa Is-
land, based on an observation and specimens collected by R.
Fricke on 11–12 Nov. 2011, St. B28, EU5. – Ma te ri al :
SMF 34761 (3), SMF 34802 (1).
Halichoeres scapularis (Bennett, 1832) – Zigzag wrasse; Girelle
en arc-en-ciel. – D is t r i bu ti on : New record from Europa
Island, based on an observation and a specimen collected by
R. Fricke at Europa on 7–12 Nov. 2011, St. B28, B33. – Ma -
t er ia l: SMF 34762 (1), SMF 34803 (1).
Halichoeres trispilus Randall & Smith, 1982 – Threespot wrasse;
Girelle trois points. – D i s t r i b u t i o n : New record from
Europa Island, based on observations by R. Fricke at Europa
on 7–11 Nov. 2011, St. B31, B33, B34, B37, B49, B50, EU5.
Hemigymnus fasciatus (Bloch, 1792) – Barred thicklip wrasse;
Chien noir (Réunion), tamarin à bandes noires (France). –
Distribution: New record from Europa Island, based
on an observation by P. durville et al. during the expedition
MD-11; also observed by P. durville & P. chaBanet on 7–12
Nov. 2011, St. B5, B8, B12, B14, EU2, EU3, EU7.
Hologymnosus annulatus (Lacepède [ex Commerson], 1801) –
Ring wrasse; Labre annelé. – D is t r i b u t i o n : New record
from Europa Island, based on an observation by R. Fricke at
Europa on 10 Nov. 2011, St. B50; also observed by P. dur-
ville, 11 Nov. 2011, St. B14.
Labrichthys unilineatus (Guichenot, 1847) – Tubelip wrasse. –
Distribution: New record from Europa Island, based on
an observation by P. durville, 11 Nov. 2011, St. B14.
Labroides bicolor Fowler & Bean, 1928 – Bicoloured cleaner
wrasse; Poisson nettoyeur bicolore. – D is t ri b ut io n : New
record from Europa Island, based on an observation by P.
durville et al. during the expedition MD-11; also observed
by P. durville & P. chaBanet on 7–12 Nov. 2011, St. B5,
B12, B14, EU2, EU3, EU7, and by R. Fricke on 11 Nov.
2011, St. EU5.
Labroides dimidiatus (Valenciennes in Cuvier & Valenciennes,
1839) – Bluestreak cleaner wrasse; Poisson nettoyeur com-
mun (Fig. 13). – D i s tr i b u ti on : First record from Europa
Island as Fissilabrus dimidiatus by Fourmanoir (1952: 172);
record confirmed by P. durville et al. during the expedition
MD-11, and by R. Fricke based on observations at Europa on
7–10 Nov. 2011, St. B31, B32, B33, B34, B49, B50, EU5;
also observed by P. durville & P. chaBanet on 7–12 Nov.
2011, St. B5, B8, B12, EU2, EU3, EU5, EU7.

Fricke et alii, checklist oF the shore Fishes oF euroPa island 267
Labropsis xanthonota Randall, 1981 – Wedge-tailed wrasse. –
Distribution: New record from Europa Island, based
on an observation by P. durville et al. during the expedition
MD-11; also observed by P. durville, 11 Nov. 2011, St. B14.
Macropharyngodon bipartitus Smith, 1957 – Vermiculate wrasse,
divided wrasse; Girelle diamant. – D i s t r i b u t i o n : New
record from Europa Island, based on an observation by P.
durville et al. during the expedition MD-11; also observed
by P. durville, 12 Nov. 2011, St. B12.
Oxycheilinus digrammus (Lacepède [ex Commerson], 1801) –
Cheek-lined wrasse, bandcheek wrasse; Vieille barbe noire.
– D is tr i b u t i on : First record from Europa Island as Chei-
linus diagrammus by Fourmanoir (1952: 184); record con-
firmed by P. durville, based on an observation on 7–10 Nov.
2011, St. B8, EU7.
Pseudocheilinus dispilus Randall, 1999 – Mascarene sixstripe
wrasse. – Distribution: New record from Europa Island,
based on an observation by P. durville et al. during the ex-
pedition MD-11; collected by P. durville & P. chaBanet on
8–9 Nov. 2011, St. EU3, EU4, 6; also observed by P. durville
& P. chaBanet on 10–12 Nov. 2011, St. B5, B8, B12, EU2.
Pseudocheilinus octotaenia Jenkins, 1901 – Eight-lined wrasse.
– Distribution: New record from Europa Island, based
on observation by P. durville & P. chaBanet on 7 Nov. 2011,
St. EU7.
Pseudodax moluccanus (Valenciennes in Cuvier & Valenciennes,
1840) – Chiseltooth wrasse; Vieille ciseau. – Di s t r i b u -
tion: New record from Europa Island, based on an observa-
tion by P. durville et al. during the expedition MD-11; also
observed by P. durville, 10–11 Nov. 2011, St. B8, B14, EU2.
Pteragogus flagellifer (Valenciennes in Cuvier & Valenciennes,
1839) – Cocktail wrasse. – D i s t r i b u t i o n : First record
from Europa Island as Duymaeria flagellifera by Fourma-
noir (1952: 184), and as Callyodon flagellifera by Fourma-
noir (1952: 187); record confirmed by R. Fricke based on
observations at Europa on 7–10 Nov. 2011, St. B31, B50; also
observed by P. durville, 11 Nov. 2011, St. B14. – M a t e -
r ia l: SMF 34740 (1).
Stethojulis albovittata (Bonnaterre [ex Koelreuther], 1788) –
Blue-lined wrasse. – D i s t r i b u t i o n : New record from
Europa Island, based on an observation by P. durville et al.
during the expedition MD-11, and on observations and speci-
mens collected by R. Fricke on 7–12 Nov. 2011, St. B28,
B32, B37, B50, EU5; also observed by P. durville & P. cha-
Banet on 9–12 Nov. 2011, St. B5, B25, EU3, EU4, EU5; pre-
viously listed as Stethojulis sp. by Fourmanoir (1952: 172).
– M at er ia l: SMF 34763 (4), SMF 34804 (7).
Thalassoma amblycephalum (Bleeker, 1856) – Twotone wrasse.
– Distribution: New record from Europa Island, based
on an observation by P. durville et al. during the expedition
MD-11, and by R. Fricke on 8–12 Nov. 2011, St. B28, B37,
B49, B50, EU5. – M at er ia l: SMF 34805 (2).
Thalassoma genivittatum (Valenciennes in Cuvier & Valenci-
ennes, 1839) – Redcheek wrasse. – D i s t r i b u t i o n : New
record from Europa Island, based on an observation by P.
durville et al. during the expedition MD-11; also observed
by P. durville & P. chaBanet on 7–12 Nov. 2011, St. B5,
B12, B14, B25, EU2, EU3, EU4, EU5, EU7.
Thalassoma hardwicke (Bennett, 1830) – Sixbar wrasse; Girelle
à six bandes. – D i s t r i b u t i o n : First record from Europa
Island as Thalassoma hardwicki by Fourmanoir (1952: 170),
and as Thalassoma hardwickei by Fourmanoir (1952: 187);
record confirmed by P. durville et al. during the expedition
MD-11, and by R. Fricke based on observations at Europa
on 7–11 Nov. 2011, St. B28, B31, B32, B33, B34, B49, B50,
EU5; also observed by P. durville, 12 Nov. 2011, St. EU5.
Thalassoma hebraicum (Lacepède [ex Commerson], 1801) –
Goldbar wrasse. – D i s t r i b u t i o n : First record from Eu-
ropa Island as Thalassoma hebraïcum by Fourmanoir (1952:
184); record confirmed by P. durville et al. during the ex-
pedition MD-11; also observed and a specimen collected by
P. durville & P. chaBanet on 7–12 Nov. 2011, St. B5, B8,
B25, EU2, EU3, EU4, EU5, EU7, and by R. Fricke on 11
Nov. 2011, St. EU5.
Thalassoma lunare (Linnaeus, 1758) – Moon wrasse; Girelle
verte. – D i s t r i b u t i o n : First record from Europa Island
by Fourmanoir (1952: 184); record confirmed by P. durville
et al. during the expedition MD-11, and by R. Fricke based
on observations at Europa on 7–11 Nov. 2011, St. B31, B33,
B34, B37, B49, B50, EU5; also observed by P. durville & P.
chaBanet on 7–11 Nov. 2011, St. B5, B14, EU7.
Thalassoma purpureum (Forsskål in Niebuhr, 1775) – Surge
wrasse; Girelle hublot. – D i s tr i b ut i o n : New record from
Europa Island, based on an observation by P. durville et al.
during the expedition MD-11.
Thalassoma quinquevittatum (Lay & Bennett, 1839) – Fivestripe
wrasse. – Distribution: New record from Europa Island,
based on an observation by R. Fricke based on observations
on 7–11 Nov. 2011, St. B28, B32, EU5.
Thalassoma trilobatum (Lacepède [ex Commerson], 1801) –
Ladder wrasse, Christmas wrasse; Calonnier côte (Réunion),
girelle de Noël (France). – D i s t r i b u t i o n : New record
from Europa Island, based on an observation by P. durville
et al. during the expedition MD-11, and specimens collected
by R. Fricke on 11–12 Nov. 2011, St. B28; also observed
by P. durville, 12 Nov. 2011, St. B25. – Ma t er i al : SMF
34764 (2), SMF 34806 (1).
Wetmorella nigropinnata (Seale, 1901) – Sharpnose wrasse. –
Distribution: New record from Europa Island, based on
a specimen collected by P. durville & P. chaBanet on 9 Nov.
2011, St. EU3.
Scaridae
Calotomus carolinus (Valenciennes in Cuvier & Valenciennes,
1840) – Stareye parrotfish, Christmas parrotfish, bucktooth
parrotfish; Perroquet des Carolines. – D i st r ib u ti o n: New
record from Europa Island, based on an observation by P.
durville et al. during the expedition MD-11.
Chlorurus sordidus (Forsskål in Niebuhr, 1775) – Bullethead par-
rotfish, daisy parrotfish; Perroquet marguerite. – D is t r ib u -
tion: New record from Europa Island, based on observation
by P. durville & P. chaBanet on 7–12 Nov. 2011, St. B5, B8,
B12, B14, EU2, EU3, EU4, EU7, and by R. Fricke on 8–10
Nov. 2011, St. B34, B37, B50.
Chlorurus strongylocephalus (Bleeker, 1854) – Steephead par-
rotfish; Perroquet vert (Réunion). – D i s t r i b u t i o n : New

stuttGarter BeiträGe zur naturkunde a
268 Neue Serie 6
record from Europa Island, based on an observation by P.
durville et al. at St. 1 (equals St. EU5 of this paper) dur-
ing an expedition with R/V Marion Dufresne on 5 April
2011, and on observations by R. Fricke on 7–10 Nov. 2011,
St. B31, B32, B33, B49, B50; also observed by P. durville
& P. chaBanet on 7 Nov. 2011, St. EU7, B14. – R e m ar ks :
This species is similar to Chlorurus microrhinos (Bleeker,
1854) of the western Pacific Ocean.
Scarus forsteni (Bleeker, 1861) – Forsten’s parrotfish; Perroquet
mauve. – Distribution: New record from Europa Island,
based on observation by P. durville & P. chaBanet on 7 Nov.
2011, St. EU7.
Scarus frenatus Lacepède [ex Commerson], 1802 – Bridled par-
rotfish, vermiculate parrotfish. – D i s t r i b u t i o n : New
record from Europa Island, based on an observation by R.
Fricke on 7–11 Nov. 2011, St. B31, B34, B37, B49, B50,
EU5; also observed by P. durville & P. chaBanet on 7–12
Nov. 2011, St. B8, B12, EU7.
Scarus ghobban Forsskål in Niebuhr, 1775 – Blue-barred par-
rotfish; Perroquet bleu (Réunion), perroquet à écailles jaunes
(France). – D i s t r i b u t i o n : First record from Europa Is-
land as Callyodon guttatus by Fourmanoir (1952: 172); re-
cord confirmed by R. Fricke on an observation on 8–11 Nov.
2011, St. B49, EU5; also observed by P. durville, 12 Nov.
2011, St. EU5.
Scarus niger Forsskål in Niebuhr, 1775 – Perroquet dorade. –
Distribution: New record from Europa Island, based
on an observation by P. durville et al. during the expedition
MD-11.
Scarus rubroviolaceus Bleeker, 1847 – Ember parrotfish; Perro-
quet braisé. – D i s t r i b u t i o n : New record from Europa
Island, based on an observation by P. durville et al. during
the expedition MD-11; also observed by P. durville & P.
chaBanet on 7–12 Nov. 2011, St. B5, B8, B12, B14, EU3,
EU4, EU5, EU7, and by R. Fricke on 11 Nov. 2011, St. EU5.
Scarus scaber Valenciennes in Cuvier & Valenciennes, 1840
– Dusky-capped parrotfish; Perroquet marron (Réunion),
perroquet à cinq celles (France). – D i s t r i b u t i o n : New
record from Europa Island, based on an observation by P.
durville et al. during the expedition MD-11; also observed
by P. durville & P. chaBanet on 9–12 Nov. 2011, St. B12,
B14, EU3.
Scarus tricolor Bleeker, 1847 – Tricolour parrotfish; Perroquet
tricolore. – D i s t r i b u t i o n : New record from Europa Is-
land, based on observation by P. durville & P. chaBanet on
7–12 Nov. 2011, St. B5, B8, B12, EU2, EU3, EU7.
Pinguipedidae
Parapercis hexophtalma (Cuvier [ex Ehrenberg] in Cuvier & Va-
lenciennes, 1829) – Speckled sandperch, spotted sandsmelt;
Pinge pintade. – D is t r i bu ti o n : New record from Europa
Island, based on observation by P. durville & P. chaBanet
on 10–11 Nov. 2011, St. B14, EU2.
Parapercis punctulata (Cuvier in Cuvier & Valenciennes, 1829)
– Spotted sandperch, spotted sandsmelt. – D i s t ri bu t i o n:
New record from Europa Island, based on an observation by
P. durville et al. during the expedition MD-11.
Tripterygiidae
Enneapterygius abeli (Klausewitz, 1960) – Yellow triplefin. –
Distribution: New record from Europa Island, based
on specimens collected by R. Fricke on 11–12 Nov. 2011,
St. B28. – M at er ia l: SMF 34782 (3), SMF 34824 (1).
Enneapterygius philippinus (Peters, 1868) – Minute triplefin. –
Distribution: New record from Europa Island, based
on specimens collected by R. Fricke, 11–12 Nov. 2011,
St. B28, and by P. durville, 12 Nov. 2011, St. B25; previ-
ously reported as Enneapterygius sp. by Fourmanoir (1952:
188). – M at er ia l: SMF 34825 (25), SMF 34836 (8).
Enneapterygius ventermaculus Holleman, 1982 – Blotched tri-
plefin. – Distribution: New record from Europa Island,
based on specimens collected by R. Fricke on 9–12 Nov.
2011, St. B28; also observed by P. durville, 11 Nov. 2011,
St. B5. – M at er ia l: SMF 34736 (1), SMF 34783 (4), SMF
34826 (4).
Blenniidae
Antennablennius australis Fraser-Brunner, 1951 – Moustached
rockskipper. – Di st ri bu ti on : New record from Europa
Island, based on specimens collected by P. durville & P.
chaBanet on 12 Nov. 2011, St. B25. – M a t e r i a l : SMF
34828 (2).
Antennablennius bifilum (Günther, 1861) – Horned rockskip-
per. – Distribution: New record from Europa Island,
based on specimens collected by R. Fricke on 11 Nov. 2011,
St. B28. – M at er ia l: SMF 34746 (2).
Aspidontus tractus Fowler, 1903 – Mimic blenny. – D i s t r i b u -
tion: New record from Europa Island, based on an observa-
tion by R. Fricke on 11 Nov. 2011, St. EU5.
Blenniella chrysospilos (Bleeker, 1857) – Red-spotted blenny;
Blennie à taches rouges. – D i s t r i b u t i o n : New record
from Europa Island, based on specimens collected by R.
Fricke on 11–12 Nov. 2011, St. B28, and by P. durville &
P. chaBanet on 12 Nov. 2011, St. B25. – Ma t e r i a l : SMF
34747 (2), SMF 34789 (16), SMF 34829 (2).
Blenniella gibbifrons (Quoy & Gaimard, 1824) – Hump-headed
blenny. – Distribution: New record from Europa Island,
based on specimens collected by R. Fricke on 11–12 Nov.
2011, St. B28, and by P. durville & P. chaBanet on 12 Nov.
2011, St. B25. – M a t e r i a l : SMF 34748 (3), SMF 34790
(1), SMF 34830 (2).
Blenniella periophthalmus (Valenciennes in Cuvier & Valenci-
ennes, 1836) – Blue-dashed rockskipper, bullethead rock-
skipper. – Di st ri b u t i o n: First record from Europa Island
as Alticops periophtalmus by Fourmanoir (1952: 169); re-
cord confirmed by R. Fricke based on a specimen collected
on 9–12 Nov. 2011, St. B28, EU5. – M a te r ia l : SMF 34728
(1), SMF 34749 (4), SMF 34791 (4).
Cirripectes auritus Carlson, 1981 – Blackflap blenny. – Distri-
bution: New record from Europa Island, based on an ob-
servation by P. durville et al. during the expedition MD-11.
Cirripectes castaneus (Valenciennes in Cuvier & Valenciennes,
1836) – Muzzled rockskipper, chestnut blenny. – D i s t r i -
bution: New record from Europa Island, based on an ob-
servation by P. durville et al. during the expedition MD-11.

Fricke et alii, checklist oF the shore Fishes oF euroPa island 269
Cirripectes filamentosus (Alleyne & Macleay, 1877) – Filamen-
tous blenny. – Di s t r i b u t i o n : New record from Europa
Island, based on an observation and a specimen collected by
R. Fricke on 8–9 Nov. 2011, St. B28, B34; also on a speci-
men collected by P. durville & P. chaBanet on 9 Nov. 2011,
St. EU3.
Ecsenius lineatus Klausewitz, 1962 – Linear blenny; Blennie
bigula. – Distribution: New record from Europa Is-
land, based on an observation by R. Fricke on 8 Nov. 2011,
St. B49; also observed at Europa Island by P. durville et al.
as Ecsenius sp. during the expedition MD-11; also observed
by P. durville & P. chaBanet on 7–11 Nov. 2011, St. B5,
EU2, EU3, EU7, photographed at St. EU6, and collected on
11 Nov. 2011, on the north side of the island, 18 m depth.
Ecsenius sp. (black, head light). – D i s t r i b u t i o n : Recorded
from Europa Island, based on an observation by P. durville,
10–12 Nov. 2011, St. B5, B8, B12, B14, EU2, EU5.
Exallias brevis (Kner, 1868) – Shortbodied blenny, leopard
blenny, leopard rockskipper. – D i s t r i b u t i o n : New re-
cord from Europa Island, based on an observation by P. dur-
ville et al. during the expedition MD-11; also observed by P.
durville & P. chaBanet on 9 Nov. 2011, St. EU4.
Istiblennius edentulus (Schneider [ex Forster] in Bloch & Schnei-
der, 1801) – Rippled rockskipper. – D i s t r i b u t i o n : New
record from Europa Island, based on specimens collected by
R. Fricke on 7–12 Nov. 2011, St. B28, B31, B34, B37, EU5,
and by P. durville & P. chaBanet on 12 Nov. 2011, St. B25.
– M at er ia l: SMF 34792 (1), SMF 34831 (1).
Istiblennius steindachneri (Pfeffer, 1893) – Steindachner’s rock-
skipper. – Distribution: New record from Europa Is-
land; previously listed as Alticops oryx (non Rüppell, 1830)
by Fourmanoir (1952: 169).
Plagiotremus rhinorhynchos (Bleeker, 1852) – Twostripe blenny.
– Distribution: New record from Europa Island, based
on an observation by P. durville et al. during the expedition
MD-11, and by R. Fricke on 8 Nov. 2011, St. B34, B49; also
observed by P. durville & P. chaBanet on 9–12 Nov. 2011,
St. B5, B12, B14, EU3, EU4.
Plagiotremus tapeinosoma (Bleeker, 1857) – Piano blenny. –
Distribution: New record from Europa Island, based
on an observation by P. durville et al. during the expedition
MD-11; also observed by P. durville & P. chaBanet on 7–12
Nov. 2011, St. B8, B12, B14, EU3, EU7.
Salarias fasciatus (Bloch, 1786) – Jewelled blenny. – Di s t r i -
bution: New record from Europa Island, based on an ob-
servation by R. Fricke on 11 Nov. 2011, St. EU5.
Callionymidae
Diplogrammus infulatus Smith, 1963 – Sawspine dragonet. –
Distribution: New record from Europa Island, based
on specimens collected by R. Fricke on 11–12 Nov. 2011,
St. B28. – M at er ia l: SMF 34750 (2), SMF 34793 (1).
Eleotridae
Ophiocara porocephala (Valenciennes in Cuvier & Valenciennes,
1837) – Flathead sleeper; Cabot marare (Réunion), éléotris à
tête poreuse (France). – D is t r i b u t i o n : New record from
Europa Island; previously listed as Ophiocara sp. by Four-
manoir (1952: 187).
Gobiidae
Amblygobius semicinctus (Bennett, 1833) – Half-barred goby;
Gobie galonne (Fig. 18). – D i s t r i b u t i o n : New record
from Europa Island, based on an observation by R. Fricke on
10 Nov. 2011, St. B37, and on a photograph by P. durville
taken on 11 Nov. 2011 among the mangroves in the inner la-
goon of Europa Island.
Amblygobius sphynx (Valenciennes in Cuvier & Valenciennes,
1837) – Sphinx goby. – Di s t r i b ut io n : New record from
Europa Island, based on an observation by R. Fricke on 7
Nov. 2011, St. B32.
Bathygobius cotticeps (Steindachner, 1880) – Cheekscaled frill-
goby. – Distribution: New record from Europa Island,
based on a specimen collected by R. Fricke on 10–12 Nov.
2011, St. B28, B50. – M a t e r i a l : SMF 34737 (1), SMF
34753 (3), SMF 34796 (2).
Bathygobius fuscus (Rüppell, 1830) – Dusky frillgoby. – Di s -
t r ib ut i on : First record from Europa Island by Fourmanoir
(1952: 169); record confirmed by R. Fricke based on speci-
mens collected and observed at Europa on 7–12 Nov. 2011,
St. B28, B31, B34, B37, EU5, and collected by P. durville &
P. chaBanet on 12 Nov. 2011, St. B25. – Ma t e r i a l : SMF
34797 (12), SMF 34832 (5).
Cryptocentrus cryptocentrus (Valenciennes in Cuvier & Valen-
ciennes, 1837) – Ninebar prawn goby. – Di s t r i b u t i o n :
New record from Europa Island, based on a photograph by P.
durville taken on 11 Nov. 2011 among the mangroves in the
inner lagoon of Europa Island.
Ctenogobiops maculosus (Fourmanoir, 1955) – Seychelles
shrimpgoby. – Di s t r i b u t i o n : New record from Europa
Island, based on an observation by R. Fricke on 8 Nov. 2011,
St. B34.
Eviota indica Lachner & Karnella, 1980 – Indian Ocean coral
goby. – Distribution: New record from Europa Island,
based on a specimen collected by P. durville & P. chaBanet
on 11 Nov. 2011, on the north side of the island, 18 m depth.
Eviota prasina (Klunzinger, 1871) – Green bubble goby. – D i s-
tribution: New record from Europa Island, based on
specimens collected and observed by R. Fricke on 8–12 Nov.
2011, St. B28, B34, B50, EU5, and collected by P. durville
& P. chaBanet on 12 Nov. 2011, St. B25. – M a t e r i a l :
SMF 34729 (2), SMF 34738 (3), SMF 34755 (9), SMF 34798
(21), SMF 34833 (1).
Eviota sp. 1. – D i s t r i b u t i o n : Observed at Europa Island,
based on specimens collected by P. durville & P. chaBanet
on 8–11 Nov. 2011, St. EU6, and on the north side of the is-
land, 18 m depth, and a specimen collected by R. Fricke on 11
Nov. 2011, St. B28. – M a t e r i a l : SMF 34754 (1).
Eviota sp. 2. – D i s t r i b u t i o n : Observed at Europa Island,
based on a specimen collected by P. durville & P. chaBanet
on 11 Nov. 2011, on the north side of the island, 18 m depth.
Fusigobius inframaculatus (Randall, 1994) – Flag fusegoby. –
Distribution: New record from Europa Island, based on
a specimen collected by P. durville & P. chaBanet on 7 Nov.
2011, St. EU7, observed by P. durville & P. chaBanet on 10
Nov. 2011, St. EU2, and a photograph taken by P. durville
at St. EU6.

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270 Neue Serie 6
Gnatholepis anjerensis (Bleeker, 1851) – Eye-bar goby. – D is -
tribution: New record from Europa Island, based on a
specimen collected by P. durville & P. chaBanet on 11 Nov.
2011, on the north side of the island, 18 m depth, and on a
photograph taken by P. durville at St. EU6.
Gnatholepis cauerensis (Bleeker, 1853) – Dark eye-bar goby. –
Distribution: New record from Europa Island, based on
observation by P. durville et al. during the expedition MD-
11, and on specimens collected by R. Fricke at Europa on
7–11 Nov. 2011, St. B31, B33, B34, B37, B49, B50, EU5, and
by P. durville & P. chaBanet on 7–10 Nov. 2011, St. EU2,
EU7. – M at er ia l: SMF 34739 (1).
Gobiodon citrinus (Rüppell, 1838) – Lemon coral goby. – Di s -
t r ib ut i on : First record from Europa Island by Fourmanoir
(1952: 187).
Koumansetta hectori (Smith, 1957) – Hector’s goby. – Di s tr i-
bution: New record from Europa Island, based on an ob-
servation by P. durville et al. during the expedition MD-11.
Lotilia graciliosa Klausewitz, 1960 – Whitecap goby. – D i s -
tribution: New record from Europa Island, based on an
observation by P. durville & P. chaBanet on 7 Nov. 2011,
St. EU7.
Pleurosicya micheli Fourmanoir, 1971 – Michel’s ghost goby. –
Distribution: New record from Europa Island, based on
a specimen collected by P. durville & P. chaBanet on 7 Nov.
2011, St. EU7.
Priolepis cincta (Regan, 1908) – Girdled goby. – Di s t r i b u -
tion: New record from Europa Island, based on specimens
collected by R. Fricke on 11–12 Nov. 2011, St. B28. – Ma -
t er ia l: SMF 34756 (4), SMF 34799 (6).
Priolepis inhaca (Smith, 1949) – Brick goby. – Di s t ri b u ti o n :
New record from Europa Island, based on a specimen col-
lected by R. Fricke on 11–12 Nov. 2011, St. B28. – Ma te -
r ia l: SMF 34757 (1), SMF 34800 (1).
Silhouettea insinuans Smith, 1959 – Phantom goby. – Di s tr i -
bution: New record from Europa Island, based on a speci-
men collected by P. durville & P. chaBanet on 7 Nov. 2011,
St. EU7.
Trimma anaima Winterbottom, 2000 – Comores goby. – D i s -
tribution: New record from Europa Island, based on
specimens collected by P. durville & P. chaBanet on 7–8
Nov. 2011, St. EU6, EU7.
Trimma sp. [aff. Trimma okinawae (Aoyagi, 1949)]. – D is t ri -
b ut i o n : Observed at Europa Island, based on a photograph
taken by P. durville on 12 Nov. 2011, St. EU5.
Valenciennea strigata (Broussonet [ex Forster], 1782) – Blue-
band goby. – Distribution: New record from Europa Is-
land, based on observations by R. Fricke on 8–11 Nov. 2011,
St. B34, B37, EU5; also observed by P. durville, 12 Nov.
2011, St. EU5.
Vanderhorstia ambanoro (Fourmanoir, 1957) – Ambanoro prawn
goby. – Distribution: New record from Europa Island,
based on observations by R. Fricke on 8–10 Nov. 2011,
St. B34, B37.
Microdesmidae
Nemateleotris magnifica Fowler, 1938 – Fire goby, fire dartfish;
Poisson de feu. – D i s t r i b u t i o n : New record from Eu-
ropa Island, based on an observation by P. durville et al. dur-
ing the expedition MD-11; also on observations by P. dur-
ville & P. chaBanet on 7–12 Nov. 2011, St. B5, B12, B14,
EU2, EU3, EU7.
Ptereleotris evides (Jordan & Hubbs, 1925) – Blackfin dartfish,
scissortail; Flèche noire. – D i s t r i b u t i o n : New record
from Europa Island, based on an observation by P. durville
et al. during the expedition MD-11; also observed by P. dur-
ville & P. chaBanet on 9–10 Nov. 2011, St. EU2, EU3.
Ptereleotris microlepis (Bleeker, 1856) – Blue gudgeon. – D i s -
tribution: New record from Europa Island, based on
observations and specimens collected by R. Fricke on 8–11
Nov. 2011, St. B28, B34. – M a t e r i a l : SMF 34731 (1),
SMF 34765 (1).
Ptereleotris zebra (Fowler, 1938) – Zebra dartfish; Gobie zèbre.
– Distribution: New record from Europa Island, based
on an observation by P. durville et al. during the expedition
MD-11; also observed by P. durville & P. chaBanet on 9
Nov. 2011, St. EU4.
Siganidae
Siganus laqueus Bonde, 1934 – Star-spotted rabbitfish, brown-
spotted spinefoot; Sigan marguerite. – D i s t r i b u t i o n :
New record from Europa Island, based on an observation by
P. durville et al. during the expedition MD-11.
Zanclidae
Zanclus cornutus (Linnaeus, 1758) – Moorish idol; Cocher blanc.
– Distribution: New record from Europa Island, based
on an observation by P. durville et al. at St. 1 (equals St.
EU5 of this paper) during an expedition with R/V Marion
Dufresne on 5 April 2011; also observed by P. durville & P.
chaBanet on 7–12 Nov. 2011, St. B5, B8, B12, EU2, EU3,
EU7, and by R. Fricke on 10–11 Nov. 2011, St. B50, EU5.
Acanthuridae
Acanthurus bariene Lesson, 1831 – Black-spot surgeonfish. –
D i st r ib u ti o n: First record from Europa Island as Acan-
thurus basiene by Fourmanoir (1952: 184), and as Acanthu-
rus bariene by Fourmanoir (1952: 186); record confirmed by
R. Fricke based on an observation at Europa on 7 Nov. 2011,
St. B31, 33.
Acanthurus blochii Valenciennes in Cuvier & Valenciennes, 1835
– Tailring surgeonfish; Chirurgien à queue rayée. – D i s -
tribution: New record from Europa Island, based on
observation by P. durville & P. chaBanet on 9 Nov. 2011,
St. EU3.
Acanthurus dussumieri Valenciennes in Cuvier & Valenciennes,
1835 – Eyestripe surgeonfish; Chirurgien de Dussumier. –
Distribution: New record from Europa Island, based
on an observation by P. durville et al. during the expedition
MD-11; also observed by P. durville & P. chaBanet on 7–11
Nov. 2011, St. B5, B8, EU7.
Acanthurus leucosternon Bennett, 1833 – Powder-blue surgeon-
fish; Chirurgien poudré. – D i s t r i b u t i o n : New record
from Europa Island, based on an observation by P. durville
et al. during the expedition MD-11; also observed by P. dur-
ville & P. chaBanet on 9–12 Nov. 2011, St. B5, B8, B12,
EU3.

Fricke et alii, checklist oF the shore Fishes oF euroPa island 271
Acanthurus lineatus (Linnaeus, 1758) – Lined surgeonfish; Chi-
rurgien à lignes bleues. – D i s t r i b u t i o n : New record
from Europa Island, based on an observation by P. durville
et al. at St. 1 (equals St. EU5 of this paper) during an expe-
dition with R/V Marion Dufresne on 5 April 2011; also ob-
served by P. durville & P. chaBanet on 7–12 Nov. 2011,
St. B5, B8, B12, EU5, EU7, and by R. Fricke on 9–11 Nov.
2011, St. B28, EU5.
Acanthurus nigricauda Duncker & Mohr, 1929 – Epaulette sur-
geonfish; Chirurgien à queue blanche. – D i s t r i b u t i o n :
New record from Europa Island, based on an observation by
P. durville et al. during the expedition MD-11; also observed
by P. durville & P. chaBanet on 9–11 Nov. 2011, St. B14,
EU2, EU3, EU5.
Acanthurus nigrofuscus (Forsskål in Niebuhr, 1775) – Dusky sur-
geonfish; Chirurgien brun. – D i s t r i b u t i o n : First record
from Europa Island as Acanthurus lineolatus by Fourmanoir
(1952: 184); record confirmed by P. durville et al. during the
expedition MD-11; also observed by P. durville & P. cha-
Banet on 9–12 Nov. 2011, St. B8, B12, EU2, EU3, EU4.
Acanthurus tennentii Günther, 1861 – Lieutenant surgeonfish,
doubleband surgeonfish; Chirurgien fer à cheval. – D i st ri -
bution: New record from Europa Island, based on an ob-
servation by P. durville et al. during the expedition MD-11.
Acanthurus thompsoni (Fowler, 1923) – Thompson’s surgeon-
fish; Chirurgien chocolat. – D i s t r i b u t i o n : New record
from Europa Island, based on an observation by P. durville
& P. chaBanet on 7–12 Nov. 2011, St. B5, B8, B12, EU7.
Acanthurus triostegus (Linnaeus, 1758) – Convict surgeonfish;
Cordonnier (Réunion). – D is tr i b u t i o n : First record from
Europa Island by Fourmanoir (1952: 172); record confirmed
by P. durville et al. based on a observation at St. 1 and St. 2
(equals St. EU5 and B34 of this paper) during an expedition
with R/V Marion Dufresne on 5–6 April 2011, an observa-
tion by P. Borsa in the mangrove area on 7 April 2011, and
by R. Fricke based on observations and specimens collected
at Europa on 7–12 Nov. 2011, St. B28, B31, B32, B33, B34,
B37, B49, B50, EU5; also observed by P. durville, 12 Nov.
2011, St. B25. – M a t e r i a l : SMF 34726 (2), SMF 34742
(1), SMF 34785 (1).
Acanthurus xanthopterus Valenciennes in Cuvier & Valenci-
ennes, 1835 – Yellowfin surgeonfish; Chirurgien (Réunion),
chirurgien aile jaune (France). – D i s t r i b u t i o n : First re-
cord from Europa Island as Acanthurus fuliginosus by Four-
manoir (1952: 184); record confirmed by P. durville et al.
during the expedition MD-11; also observed by P. durville
& P. chaBanet on 9–12 Nov. 2011, St. B5, B8, B12, EU3.
Ctenochaetus striatus (Quoy & Gaimard, 1825) – Striated sur-
geonfish; Chirurgien strié. – D i s t r i b u t i o n : New record
from Europa Island, based on an observation by P. durville
et al. during the expedition MD-11; also observed by P. dur-
ville & P. chaBanet on 9–12 Nov. 2011, St. B5, B12, EU3,
EU5, and by R. Fricke on 11 Nov. 2011, St. EU5.
Ctenochaetus truncatus Randall & Clements, 2001 – Goldring
bridletooth. – D i s tr i b ut i o n : New record from Europa Is-
land, based on an observation by P. durville et al. during the
expedition MD-11, and by R. Fricke based on an observation
at Europa on 7–11 Nov. 2011, St. B31, B34, B49, B50, EU5;
also observed by P. durville & P. chaBanet on 7–12 Nov.
2011, St. B5, B8, B12, B14, EU2, EU3, EU7.
Naso brachycentron (Valenciennes in Cuvier & Valenciennes,
1835) – Humpback unicornfish; Nason bossu. – D is tr ib u-
tion: New record from Europa Island, based on an observa-
tion by P. durville et al. during the expedition MD-11; also
observed by P. durville, 10 Nov. 2011, St. B8.
Naso brevirostris (Cuvier, 1829) – Spotted unicornfish, palefin
unicornfish; Nason à rostre court. – D i s t r i b u t i o n : New
record from Europa Island, based on an observation by P.
durville et al. during the expedition MD-11; also observed
by P. durville, 10–11 Nov. 2011, St. B5, B8, B14.
Naso elegans (Rüppell, 1829) – Indian Ocean orangespine uni-
cornfish, elegant unicornfish; Nason bariolé (Fig. 13). – D i s-
tribution: New record from Europa Island, based on an
observation by P. durville et al. during the expedition MD-
11; also observed by P. durville & P. chaBanet on 7–11
Nov. 2011, St. B5, B8, B14, EU2, EU3, EU4, EU7, and by R.
Fricke on 10 Nov. 2011, St. B50.
Naso hexacanthus (Bleeker, 1855) – Sleek unicornfish; Nason
lisse. – Distribution: New record from Europa Island,
based on an observation by P. durville et al. during the ex-
pedition MD-11.
Naso tuberosus Lacepède [ex Commerson], 1801 – Humpnose
unicornfish. – D i s t r i b u t i o n : New record from Europa
Island, based on an observation by P. durville, 10 Nov. 2011,
St. B8.
Naso unicornis (Forsskål in Niebuhr, 1775) – Bluespine unicorn-
fish; Licorne (Réunion), nason à épérons bleus (France). –
Distribution: New record from Europa Island, based on
observations by P. durville et al. at St. 1 and St. 2 (equals
St. EU5 and B34 of this paper) during an expedition with
R/V Marion Dufresne on 5–6 April 2011; also observed by
P. durville & P. chaBanet on 7–11 Nov. 2011, St. B5, B8,
B14, EU2, EU5, EU7, and by R. Fricke on 10–11 Nov. 2011,
St. B37, EU5; record also confirmed based on photograph by
E. BretaGne, taken in November 2011.
Naso vlamingi (Valenciennes in Cuvier & Valenciennes, 1835) –
Bignose unicornfish; Nason zébré. – D i s t ri bu ti on : New
record from Europa Island, based on an observation by R.
Fricke on 9 Nov. 2011, St. B28.
Paracanthurus hepatus (Linné, 1766) – Palette surgeonfish;
Chirugien bleu. – D i s t r i b u t i o n : New record from Eu-
ropa Island, based on a photograph by P. durville taken on
11 Nov. 2011, on the north side of the island below the boat
mooring, at 18 m depth.
Zebrasoma gemmatum (Valenciennes in Cuvier & Valenciennes,
1835) – Gem surgeonfish; Acanthure à pierreries. – Di st ri -
bution: New record from Europa Island, based on a speci-
men collected by P. durville & P. chaBanet on 7 Nov. 2011,
St. EU7.
Zebrasoma scopas (Cuvier, 1829) – Brushtail tang, twotone tang;
Chirurgien à brosses. – D i st r i b ut i o n : New record from
Europa Island, based on an observation by P. durville et al.
during the expedition MD-11; also observed by P. durville
& P. chaBanet on 7–12 Nov. 2011, St. B5, B8, B12, B14,
EU2, EU7.
Zebrasoma velifer (Bloch, 1795) – Desjardin’s sailfin tang; Chi-
rurgien à voile. – D i s t r i b u t i o n : New record from Eu-
ropa Island, based on an observation by P. durville et al. dur-

stuttGarter BeiträGe zur naturkunde a
272 Neue Serie 6
ing the expedition MD-11; also observed by P. durville & P.
chaBanet on 9–11 Nov. 2011, St. B5, B8, B14, EU3.
Sphyraenidae
Sphyraena barracuda (Catesby [ex Edwards], 1771) – Great bar-
racuda; Barracuda (Réunion) (Fig. 19). – D i s t r i b u t i o n :
New record from Europa Island, based on an observation by
P. durville et al. during the expedition MD-11, and an obser-
vation by P. Borsa in the mangrove area on 7 April 2011; pre-
viously listed as Sphyraena sp. by Fourmanoir (1952: 170);
also observed by P. durville & P. chaBanet on 7–12 Nov.
2011, St. B12, EU3, EU7; record confirmed based on photo-
graph by E. BretaGne, taken in November 2011.
Scombridae
Euthynnus affinis (Cantor, 1850) – Eastern little tuna; Bonite la
côte (Réunion), thonine orientale (France). – D i s t r i b u -
tion: New record from Europa Island; previously listed as
Euthynnus alleteratus (non Rafinesque-Schmaltz, 1810) by
Fourmanoir (1952: 179).
Thunnus albacares (Bonnaterre, 1788) – Yellowfin tuna; Thon
jaune (Réunion). – D i st ri bu ti on : New record from Eu-
ropa Island, based on an observation by P. durville et al. dur-
ing the expedition MD-11.
Bothidae
Bothus mancus (Broussonet, 1782) – Peacock flounder; Sole (Ré-
union). – D i st ri bu ti on : First record from Europa Island
by Fourmanoir (1952: 184).
Balistidae
Balistapus undulatus (Park, 1797) – Orange-striped triggerfish;
Baliste ondulé. – D is tr i b u t i on : First record from Europa
Island by Fourmanoir (1952: 181); record confirmed by P.
durville et al. during the expedition MD-11; also observed
by P. durville & P. chaBanet on 7–12 Nov. 2011, St. B5,
B8, B12, B14, EU2, EU3, EU4, EU7, and by R. Fricke on 11
Nov. 2011, St. EU5.
Balistoides conspicillum (Bloch & Schneider, 1801) – Clown
triggerfish; Bourse piastre, bourse pièce (Réunion), baliste
léopard, baliste clown (France). – D is tr ib ut io n: First re-
cord from Europa Island by Fourmanoir (1952: 188); record
confirmed by P. durville et al. during the expedition MD-11;
also observed by P. durville & P. chaBanet on 9–12 Nov.
2011, St. B12, B14, EU3.
Balistoides viridescens (Bloch & Schneider, 1801) – Titan trig-
gerfish; Bourse jaune (Réunion), baliste verdâtre (France). –
D i st ri bu ti on : First record from Europa Island by Four-
manoir (1952: 188); record confirmed by P. durville et al.
during the expedition MD-11, and by R. Fricke on 9–10 Nov.
2011, St. B28, B37; also observed by P. durville & P. cha-
Banet on 9–12 Nov. 2011, St. B5, B8, B12, B14, EU2, EU3.
Melichthys niger (Bloch, 1786) – Black triggerfish; Baliste noir.
– Distribution: New record from Europa Island, based
on an observation by P. durville et al. during the expedition
MD-11; also observed by P. durville & P. chaBanet on 9–11
Nov. 2011, St. B5, B8, B14, EU2, EU3, EU4.
Odonus niger (Rüppell, 1836) – Redtooth triggerfish; Bourse
noire (Réunion). – D i st r ib u ti on : New record from Eu-
ropa Island, based on an observation by P. durville et al. dur-
ing the expedition MD-11.
Pseudobalistes flavimarginatus (Rüppell, 1829) – Yellowface
triggerfish; Baliste face jaune. – D i s t r i b u t i o n : First re-
cord from Europa Island as Pseudobalistes flavomarginatus
by Fourmanoir (1952: 188); record confirmed by P. durville
et al. during the expedition MD-11; also observed by P. dur-
ville, 10 Nov. 2011, St. EU2.
Rhinecanthus aculeatus (Linnaeus, 1758) – Picassofish; Bourse
revolver (Réunion). – D i s t r i b u t i o n : First record from
Europa Island by Fourmanoir (1952: 172); record confirmed
by R. Fricke based on observations and specimens collected
at Europa on 7–12 Nov. 2011, St. B28, B31, B32, B34, B37,
B49, B50, EU5.
Rhinecanthus rectangulus (Lacepède [ex Commerson] in Bloch
& Schneider, 1801) – Wedge picassofish; Bourse (Réunion),
picasso noir (France). – D is t r i b u t i o n : New record from
Europa Island, based on an observation by R. Fricke on 9–11
Nov. 2011, St. B28, EU5; also observed by P. durville, 10–
12 Nov. 2011, St. B25, EU2.
Sufflamen bursa (Lacepède [ex Commerson] in Bloch &
Schneider, 1801) – Boomerang triggerfish; Baliste boomer-
ang. – Distribution: New record from Europa Island,
based on an observation by P. durville et al. during the expe-
dition MD-11; also observed by P. durville & P. chaBanet
on 7–12 Nov. 2011, St. B12, B14, EU2, EU7.
Sufflamen chrysopterum (Bloch & Schneider, 1801) – Halfmoon
triggerfish; Baliste double-queue. – D i s t r i b u t i o n : New
record from Europa Island, based on an observation by P.
durville et al. during the expedition MD-11; also observed
by P. durville & P. chaBanet on 9–10 Nov. 2011, St. B8,
EU3, EU4.
Monacanthidae
Cantherhines dumerilii (Hollard, 1854) – Barred filefish; Bourse
à taches blanches. – D is t r i bu ti o n : New record from Eu-
ropa Island, based on an observation by P. durville et al. dur-
ing the expedition MD-11; also observed by P. durville, 11
Nov. 2011, St. B5, B14.
Cantherhines pardalis (Rüppell, 1837) – Honeycomb filefish;
Bourse rayon de miel. – D is tr i b u t i on : New record from
Europa Island, based on observation by P. durville & P. cha-
Banet on 9–11 Nov. 2011, St. B14, EU2, EU3.
Pervagor janthinosoma (Bleeker, 1854) – Blackbar filefish. –
Distribution: New record from Europa Island, based
on an observation by P. durville et al. during the expedition
MD-11.
Ostraciidae
Ostracion meleagris Shaw in Shaw & Nodder, 1796 – Whitespot-
ted boxfish; Coffre pintade. – D is tr ib u t i on : New record
from Europa Island, based on an observation by P. durville
et al. during the expedition MD-11; also observed by P. dur-
ville & P. chaBanet on 7 Nov. 2011, St. EU7, and on 12 Nov.
2011 at St. EU5.

Fricke et alii, checklist oF the shore Fishes oF euroPa island 273
Tetraodontidae
Arothron meleagris (Anonymus [ex Lacepède, ex Commerson],
1798) – Guineafowl puffer; Compère pintade. – D i s t r i b u -
tion: New record from Europa Island, based on observation
by P. durville & P. chaBanet on 9 Nov. 2011, St. EU3.
Arothron nigropunctatus (Bloch & Schneider, 1801) – Black-
spotted puffer; Compère à taches noires. – D i s t r i b u t io n:
First record from Europa Island by Fourmanoir (1952: 172);
record confirmed by P. durville et al. during the expedition
MD-11.
Arothron stellatus (Anonymus [ex Lacepède, ex Commerson],
1798) – Star puffer; Compère étoilé. – D i s t r i b u t i o n :
First record from Europa Island by Fourmanoir (1952: 172,
pl. 13A); record confirmed by P. durville et al. during the
expedition MD-11.
Canthigaster amboinensis (Bleeker, 1865) – Spotted toby. –
D i s t r i b u t i o n : First record from Europa Island as Can-
thigaster margaritatus (non Rüppell, 1829) by Fourmanoir
(1952: 188).
Canthigaster bennetti (Bleeker, 1854) – Exquisite toby; Canthi-
gaster delicat. – D is t r i b u t i on : New record from Europa
Island, based on an observation by R. Fricke on 10 Nov.
2011, St. B50.
Canthigaster janthinoptera (Bleeker, 1855) – Honeycomb toby.
– Distribution: New record from Europa Island, based
on an observation by P. durville et al. during the expedition
MD-11; also observed by P. durville, 12 Nov. 2011, St. B12.
Canthigaster natalensis (Günther, 1870) – Natal toby. – D i s -
tribution: New record from Europa Island, based on an
observation by R. Fricke on 7–8 Nov. 2011, St. B32, B49.
Canthigaster solandri (Richardson [ex Solander], 1845) – Spot-
ted sharpnose pufferfish; Canthigaster de Solander. – Di s -
tribution: New record from Europa Island, based on a
specimen collected by P. durville & P. chaBanet on 7 Nov.
2011, St. EU7.
Canthigaster valentini (Bleeker, 1853) – Black-saddled toby;
Canthigaster à selle. – D i s t r i b u t i o n : First record from
Europa Island as Canthigaster Valentini by Fourmanoir
(1952: 188); record confirmed by P. durville et al. during the
expedition MD-11; also observed by P. durville & P. cha-
Banet on 7 Nov. 2011, St. EU7, and by R. Fricke on 8 Nov.
2011, St. B49.
4 Fish fauna of Europa Island
The shore fish fauna of Europa Island comprises a to-
tal of 389 species in 62 families. 302 species are recorded
from Europa Island for the first time. No introduced spe-
cies have been found. The largest families are the Labridae
(47 native species, 12.1 % of the native species), Pomacen-
tridae (35 species, 9.0 %), Serranidae (30 species, 7.7 %),
Gobiidae and Acanthuridae (24 species each, 6.2 %),
Chaetodontidae (20 species, 5.1 %), Blenniidae (17 spe-
Tab. 2. Proportion (percent of total) and rank (in parentheses) of the most speciose families of shore shes around islands in the south-
western Indian Ocean, based on the sources cited under Methods and Materials. Only “shore” sh species are included, i. e. marine
species which live in less than 80 m depth and are not found as adults in the oceanic pelagic waters.
Europa
Island
Seychelles Maldives Chagos
Archipelago
Réunion Mauritius Rodrigues
Labridae 12.1 (1) 7.8 (2) 7.2 (2) 8.1 (2) 7.7 (1) 9.8 (1) 7.9 (2)
Pomacentridae 9.0 (2) 4.8 (3) 4.9 (4) 4.9 (5) 5.5 (4) 4.3 (5) 6.7 (3)
Serranidae 7.8 (3) 4.4 (4) 6.4 (3) 6.4 (3) 6.2 (3) 6.7 (2) 5.5 (4)
Gobiidae 6.2 (4) 9.1 (1) 8.2 (1) 12.6 (1) 6.5 (2) 5.7 (3) 12.8 (1)
Acanthuridae 6.2 (4) 3.6 (6) 2.5 (11) 3.9 (7) 3.4 (8) 3.1 (8) 3.5 (8)
Chaetodontidae 5.1 (6) 2.2 (12) 3.2 (8) 3.2 (8) 2.8 (9) 2.9 (9) 3.7 (7)
Blenniidae 4.4 (7) 3.5 (7) 3.4 (7) 3.2 (8) 4.2 (5) 4.5 (4) 3.9 (6)
Lutjanidae 3.9 (8) 2.5 (10) 2.8 (10) 2.6 (11) 2.7 (10) 2.7 (10) 2.8 (9)
Carangidae 2.8 (9) 3.1 (8) 2.0 (14) 1.9 (14) 4.0 (6) 2.9 (9) 2.2 (11)
Balistidae 2.6 (10) 1.5 (15) 1.7 (15) 2.1 (13) 2.2 (12) 1.9 (14) 1.8 (13)
Scaridae 2.6 (10) 2.1 (13) 2.3 (12) 2.8 (10) 1.8 (15) 2.6 (11) 3.5 (8)
Holocentridae 2.3 (12) 2.4 (11) 2.2 (13) 2.8 (10) 2.8 (9) 2.1 (13) 2.0 (12)
Muraenidae 2.3 (12) 2.9 (9) 4.0 (5) 5.4 (4) 3.8 (7) 3.9 (6) 2.6 (10)
Pomacanthidae 2.1 (14) 1.1 (17) 1.3 (17) 0.8 (22) 1.2 (20) 1.2 (18) 0.6 (19)
Tetraodontidae 2.1 (14) 2.4 (11) 2.0 (14) 1.2 (17) 2.6 (11) 2.1 (13) 1.0 (17)
Apogonidae 1.8 (16) 4.1 (5) 3.8 (6) 4.2 (6) 2.2 (12) 2.7 (10) 4.1 (5)
Caesionidae 1.8 (16) 1.1 (17) 1.3 (17) 1.2 (17) 0.9 (25) 0.7 (23) 0.2 (21)
Mullidae 1.8 (16) 2.5 (10) 1.0 (19) 1.4 (16) 2.0 (13) 1.5 (16) 2.6 (10)
Total number of
species 389 905 1013 780 818 940 493

stuttGarter BeiträGe zur naturkunde a
274 Neue Serie 6
cies, 4.4 %), Lutjanidae (15 species, 3.9 %), Carangidae
(11 species, 2.8 %), Balistidae and Scaridae (10 species
each, 2.6 %). It is surprising that relatively few Gobiidae
have been found, and this is not just based on the focus on
visual census, but supported by collecting efforts.
The fish fauna is exclusively marine; no freshwater
fishes have been reported from the island, and there is no
permanent fresh water habitat on the island. The general
impression is that the fish fauna is impoverished, with rela-
tively few species occurring, some of them having a very
high biomass. Europa Island is an extremely poor-nutrient
area, with relatively few plankton in the water. Even the
mangrove area has a clean sand bottom, without any mud
or silt. Though the distance to the African continental shelf
is only 262 km towards the west, and 153 km to the Mada-
gascar island shelf in the east, many species of the conti-
nental type which are adapted to nutrient-rich waters are
not found in Europa Island.
It is difficult to compare the faunas of islands and island
groups in the Western Indian Ocean as the sampling effort
is highly variable from one island to the next. Restrict-
ing the comparison to the coastal species (marine species,
found in less than 80 m, excluding oceanic pelagic species)
there is a similarity in the ranking of the various families
(Tab. 2). The families with the highest species richness
usually include the Labridae, Pomacentridae, Seranidae
and Gobiidae, with different ranks. While at Europa Island
the Labridae was the most speciose family, in other islands
the Gobiidae are the dominant group. The situation at Eu-
ropa Island is apparently not affected by sampling effort, as
even in shallow lagoon habitats a very poor diversity of go-
bies is observed relative to other regions. Several other fish
families appear to be under-represented at Europa Island;
the family Scorpaenidae, for instance, which is usually
among the top 15 in other island groups, is not included in
the top 18 list of Europa Island.
No endemism of fish species has been reported for Eu-
ropa Island. This fact reflects the proximity of the island
to Bassas da India Atoll, another of the Îles Éparses, to the
continental shelf of Mozambique and the island shelf of
Madagascar.
The fish fauna of Europa island is typical for offshore,
low islands in the southwestern Indian Ocean. Zoogeo-
graphically, the main element of the fish fauna of Europa
Island consists of widespread tropical Indo-Pacific spe-
cies (292 species, 75.1 % of the total occurring species),
of which 273 species (70.2 %) have a typical Indo-West
Pacific distribution, and 19 species (4.9 %) an Indo-Pacific
distribution including the East Pacific. A total of 13 spe-
cies (3.3 %) are found worldwide, either circumtropical or
circumtropical including warm temperate zones. An ad-
ditional 76 species (19.5 %) are Indian Ocean endemics,
including 36 western Indian Ocean endemics (9.2 %), and
10 southwestern Indian Ocean endemics (2.6 %). These
figures are similar to the zoogeographical relationships of
the fish fauna of Réunion (see Fricke et al. 2009).
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Authors’ addresses:
Dr. ronald Fricke (corresponding author), Staatliches Museum für Naturkunde, Rosenstein 1, 70191 Stuttgart, Germany;
e-mail: ronald.fricke@smns-bw.de
Dr. Patrick durville, Sarl Aquarium de la Réunion, Port de Plaisance, 97437 Saint-Gilles les Bains, La Réunion, France;
e-mail: aquarium.reunion@wanadoo.fr
Dr. Giacomo Bernardi, Long Marine Laboratory, University of California at Santa Cruz, 100 Shaffer Road, Santa Cruz, CA 95060,
U. S. A.;
e-mail: bernardi@ucsc.edu
Dr. PhiliPPe Borsa, UR 227, Biocomplexité des écosystèmes récifaux, Institut de Recherche pour le Développement (IRD), Centre de
Montpellier, PS2, 911 avenue Agropolis, 34032 Montpellier Cedex, France;
e-mail: philippe.borsa@ird.fr
Dr. Gérard mou-tham, Institut de Recherche pour le Développement (IRD), Centre de Nouméa, 101 Promenade Roger Laroque,
Anse Vata, BP A5, 98848 Noumea, New Caledonia;
e-mail: gerard.mou-tham@ird.fr
Dr. Pascale chaBanet, Institut de Recherche pour le Développement (IRD), BP 172, 97492 Sainte-Clotilde cedex, La Réunion,
France;
e-mail: pascale.chabanet@ird.fr
Manuscript received: 25.VI.2012, accepted: 7.VIII.2012.