Flora Europaea Vol. 3
... As a result, several taxonomic treatments have been proposed in partial monographs and floras (e.g. Watzl, 1910; Stroh, 1942; Borissova, 1955; Hartl, 1966; Walters & Webb, 1972; Elenevsky, 1977; Fischer, 1982; Martínez-Ortega, Sánchez-Agudo & Rico, 2009), which differ substantially regarding taxonomic concepts. Moreover, taxon sampling for previous molecular phylogenetic studies has been limited (Albach, Martínez-Ortega & Chase, 2004c; Albach et al., 2004b; Martínez-Ortega et al., 2004) and the uncertainty regarding the phylogenetic relationships among species remains high. ...
... Veronica dentata F.W.Schmidt and V. austriaca L. s.s. are both hexaploid and are distributed in central and eastern Europe [some authors have included V. dentata or V. jacquinii within the variation of V. austriaca (e.g. Walters & Webb, 1972; Fischer, 2011)]. Morphologically intermediate populations between V. dentata, V. jacquinii and V. orbiculata are found in nature and botanists have interpreted them either as the result of hybridization between species (Lehmann, 1937; Scheerer, 1949) or as transitional forms caused by phenotypic plasticity in one large, variable species (Watzl, 1910). ...
... DNA extractions for V. krylovii were made from herbarium specimens (Table A1 ). Plants were identified following the taxonomy of Borissova (1955), Walters & Webb (1972), Fischer (1991, 2011), Peev (1995) and MartínezOrtega et al. (2009). As a unique taxonomic treatment was not followed, basionyms were used to name the taxa (Supporting Information Table S1). ...
Veronica subsection Pentasepalae is a diploid–polyploid complex of c. 20 species distributed in Eurasia and North Africa, in which species boundaries are difficult to determine. Here, we present the first comprehensive phylogenetic analysis of V. subsection Pentasepalae based on nucleotide sequences [internal transcribed spacer (ITS) and the plastid trnH-psbA and ycf6-psbM spacers] combined with ploidy estimations. Our results support the monophyly of the subsection. Five well-supported clades are recovered in the ITS sequence analyses, corresponding to broad geographical areas. The causes of the extensive incongruence found between the ITS and plastid DNA datasets, namely incomplete lineage sorting and/or hybridization and polyploidization, are discussed. Most of the diploids traditionally recognized based on morphological characters and one tetraploid are each recovered as monophyletic by the ITS sequence analyses. The Balkan species V. kindlii is resurrected. DNA ploidy level for V. teucrioides is reported here for the first time (2x). Diploid populations have been found for V. orbiculata, which was previously thought to be only tetraploid. Past contact in the amphi-Adriatic area between V. orsiniana and V. orbiculata is suggested. Finally, molecular analyses show that diploid V. jacquinii and diploid V. orbiculata are unrelated. This study contributes to our understanding of the evolutionary history of polyploid complexes, especially those in southern Europe, and highlights the importance of using multiple lines of evidence to investigate species boundaries in such actively diversifying groups. © 2015 The Linnean Society of London, Botanical Journal of the Linnean Society, 2015, 00, 000–000.
... 20-Aedo, 2005 Bolòs & Vigo, 2001, 1996; Pedrol, 1997; Rivas-Martínez et al., 2001. 25-Morales, 1986 Tutin et al., 1972. 26-Morales, 1986 Rivas-Martínez et al., 1980 Tal y como se especifica en la ficha general del grupo 2, la evaluación del estado de funcionalidad de los sistemas dunares y de su vulnerabilidad habitualmente se realiza mediante el control de un conjunto de parámetros o variables representativas (Bodéré et al., 1991; Wlilliams et al., 1993 Wlilliams et al., a y b, 1994 Wlilliams et al., , 2001 García-Mora et al., 2001; MartínezVázquez et al., 2006; Martín-Prieto et al., 2007). ...
Matorrales esclerófilos o laurifolios de los cordones dunares más estabilizados de la parte interna de las dunas grises o de los claros de enebrales, sabinares y pinares.
El tipo de hábitat de interés comunitario está presente en los ecosistemas dunares de las costas mediterráneas y atlánticas (de Cádiz a Galicia). Se trata de matorrales de talla variable, florísticamente muy diversos. En el litoral atlántico, la especie más característica es Corema album, acompañada por especies como Ulex europaeus subsp. latebracteatus (costa gallega) o Halimium halimifolium y H. calycinum (costa gaditano-onubense). Entre estos matorrales atlánticos destacan, por su riqueza y por la presencia de elementos de interés biogeográfico, los del suroeste ibérico, con Stauracanthus genistoides, Lavandula stoechas subsp. lusitanica, Ulex australis, Armeria velutina (incluida en el anexo II de la Directiva de Hábitats), Thymus tomentosus, T. camphoratus, etc.
Teucrium teresianum, from southern Spain, is here newly described, illustrated, and compared with the most closely related species of the genus. The species is characterized by leaves that are 3(−4)-whorled, oblong to oblong-spathulate, and narrowly and progressively cuneate, and a calyx that is tubular-campanulate, not flattened in the proximal part and with mucronate lobes, and creamy corolla.
Veronica repens in Corsica is shown to be a subspecies within V.serpyllifolia and is endemic there. A detailed study of the indumentum, together with considerations on palynology, seed morphology, karyology and phytogeography, has shown that the Spanish populations usually assigned to ‘V.repens’ are different from that subsp. repens at species level and belong to ‘V.langei’. This species, endemic to Spain, consist of two varieties: V.langei var. langei and V.langei var. neuadensis. ‘V.humifusa’ (= V.tenella= V.balcanica = V.apennina) is a high mountain ecotype of V.serpyllifolia meriting only varietal rank: V.serpyllifolia subsp. serpyllifolia var. humifusa. A taxonomic key to all taxa is supplied, as well as nomenclatural considerations, typification of several names, descriptions, drawings and distribution maps. A new subsection in Veronica sect. Veronicastrum is proposed.
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