TAXONOMIC REMARKS ON THE ENDEMIC FLORA OF SILA MASSIF (S ITALY)

Abstract

Within floristic investigations on Sila Massif, taxonomical considerations and new contributions on the endemic taxa occurring in this territory are given. Taxonomical considerations on the endemic taxa occurring in Sila are given. The flora of Sila results rich in endemisms exclusive of its territory, such as Astragalus calabricus, Armeria brutia, Avenula praetutiana subsp. rigida, Anthemis triumphetti var. briquettii, Anthemis hydruntina subsp. silensis, Centaurea sarfattiana, Genista silana, Koeleria splendens subsp. brutia, Luzula calabra, Cardamine silana, Knautia dinarica subsp. silana, ecc. In order to contribute to the study of endemic element of Sila territory two new species and a new combination and status are described.

Full text

S.
Brullo,
C. Gangale
& D.
Uzunov
Taxonomic
remarks on the
endemic
flora ofthe Sila Massif
(S
Italy)
Bnllo. S., cdgale, C. & trnno\ D.: Tdonoùic reDdkr on dre endeni. nora ofthe Sila
Moùntaiù (S
ltaly) Bocconea2t:2Ú-222
200?. ISSN 1120 4060.
withjn llonsric investisltions on the Sila Mastî (S
Ìraly), tsxononical ùn.jdèútjons drd new
conhibùlions
on tùe endenics of this tenilory aÈ given. The flora oî sila has t3 exclusivc
cndemics and ìncludes
turther 55 erdenics wirh a wider disribùtion iangè, B of vhjch arc
Esnicted to thè silicsùs Calabrian nassifs. tn ordú lo conrribuie to thc k owl€dee
of tùe
endenics of úc sihn noiq tyo ncw lpeci€s (,.I/.,,.d?n ie,oreanf úd AUiun iutianuùJ
re h€È dcscribed and a new conbinarìon
b pruporco.
Introduction
Withjn taxonomical investigations
or the flora oî Sourhem Italv. a contdbution on rhe
enderìljc flom of Sila is presented.
The
Sila Massil located in Northem
Calabda
and rcaching 1927
n asl
(Morre
Botte
Donato),
represents a teÌitory ofrema*able phfogeographic interesr becaùse
ofjts geo!
ogy, geogaphical position and bioclimatic fearúes. This area, rogether
with other S
Calabdan nountains and NE Sicily, belongs ro the Calabrian Arc. à! Ercinic ùnit dorni-
Ìlated
by siliceous substraia, mosdy
gaútes, mica-schists
and kinzigftjc gneiss.
According
to Rivas Maúinez & Loidi turegui (1999.),
the bìoclimate
of the upper parrs
of Sila are ro
be included
within the supla-tempemte
submeditenanean
themotype and
marginally also
to the supramediteffanearl
one, with subhurnid
or humid
ombrorwes.
The Íìountain
vegetation of Sila,
between I000-1927
m asl,
jncludes
by beechwoods.
often nixed to Calahian pine (Pr"&r xÌsza subsp. .alaAltca) forests, thomy cushiontitre
shrub communities, meadows
and wetlands.
In th€ submontane
belt, nnging befiveel 700-
1000(1200) m, deciduous oah,voods dominate
the ìandscape.
The flola of Sila has been studied
by many authon (Solla
1896;
Longo 1903, 1905;
Trotter 191l;
Crande 1913; Fiori 1919,
1922; AIbo
1935; Sarfani I959,
1965; cenrite
&
Martini 1974; Cesca 1981, 1982; Ballelli
& Venanzoni 1993;
Cesca & peruzzì
2002i
Bmllo
& al. 2002, 2004; Maùold
& al. 2003,
etc.). On the
basis ofthese studies, the
Silan
flora consists of about 1000 taria at specific and subspecific level, 7% of which are
endemics. Tn
the
present papq taxonomic
considerations
on th€ endcmic
flora ofthe Silan
Massif arc
given,
with parlìcular
attention
to some
cdrical taxa_

2r4 BmUo & al.: Ta-xonomic
rema*s on the endemic flora of the Sila ...
Resulh ard discussion
For tbe mountain belt (over 1000 ln asl) ofthe Silan teritory 68 endemic
species or
subspecies are rccorded. The nost paf (47) are, either, ApeDninic/S Apennìnic endemics
or resirìcted to the mountains ofcalabda and Nofhem Sicily.
Twenty one taxa are endem-
ic to the Calabrian siliceous
massifs and thiteen ofthese are exclusively found
on the Sila
(Fig. 1). Most of ihem are charactedstic elements of dry and open
places
occuring on the
Silan
plateau
between 1200
1
700
m asl,
and
they
are I Asbagalus calabricus, Armería
btu-
tia, Arenula pruetutia ú SÌJbsp. tigida, A thenís hydru îíM sùbsp. síle sis, Cenraurca
safaíidna, Genista
silana, Koelería splendens subsp.
,ruttd, Krautia dinaríca $bsp.
rt/drd. Other species
are frequenr at ùe fringe of pine and beech
forests
(,ldezodalp"r
tenorearus,A iunjulia un, Anthemis triunpheîîi srubsp. btíqueîiù or l]|'wetlands
(Lazula
calabtu, Cdtddnine síla a\.
Some ofthese
iaxa have been rccently desúibed (Brulo & al. 2002q 2004; Maúold &
al. 2003). and two of them new to science are
mesenled
here.
30,0
25'
a3
15,0
10,0
5,8
0t cd, si CSilSi CSlt, Si,
sa
Iig.1. Chorclogic spechun ofiheerdeftic
taxdofthe
Silùflora:Sila exclusive ofthe Sila
Massii
Cal
- Caldb.ia. Sì
- SìciU CSIt
- C and S Itrlr Sa
- Sardinia.

215
Bocconea
21 2007
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ean-elliprici",
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grandurar
wi'i:,t',tl i.- iiU,1i#
,i;
I il1"Jiliffi"JiJT|' i"l'.Hl"Ji','',"$ sra.ndard
crLpìrar-'uborbicur'r,
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hairs
on rl'e
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and
cldq r')-rolm
il'g'*,"g'ì-iò -. r"'gike;l e-10
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20-50
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'ntn'
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i:"#ffilti i',:;r", "": rr i. rocariled
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,he
\oulhem sira
at
,he ma-rgiJì
of
prne
"."Ut "i'" a",."t",rcas. al 1000-1r00
m alhtude
ll occrxs
on
silrL€ou'
sub\trala
-Til"""nXl,t"'J*.*" . AdenocaryB
'ompti':'ttus(L
) Gav
soup'
a Medíteranean
e1e
menueoletntea,n
tiatvU),'l óivonilr' lreJIC Presl
4 connut
d tus r't\'s A btutÙ'
Bnrllo. be Marco
& SrracL'a
dnd'4 'dannlc's Brullo
DcMarco& sÚaruca(l,rurrod
al.2001).
A itn i(lianrn Brullo Cargrle & U4nov tp nov
troó: r alabriu.
Sila
Grecd.
(dmpi luogo
la Jrudr pcr il 'aîtudno
""'i';:1".1
ìÀ3-ir,. cil"od o,,sdl/iHurorvpc:(
Al:
l5or\oe':c\r'cLI,
ìr'\-
ScaDus
qlrbru5.
20-45 cm alrus
ad I 2-2 J u'que ldgini' loìiorlun Iectu\ -l orra ) "
u,que
na lO cm longa.
pluncosrdu
lnUore'cenlia
e\paba Jrltura l) /) Ionous
o.li".rri.
'*.q,rir,,.
*.'0
r40'
".m
ronsi'
spa'a
oe'-N:5,,1fì:l;ìl"l;"T:;*:iì"
!.s-l
z .]n
ro"ei'
PengoniuLn
albo-lrnde.
sub(
""i "-U,".""-
"i"'"*, "a apicen
rotundatis,
'i.s-S
S mm tongls'
2J mrn
latis Stanina
.'-""|"rc'".ìJ".". r'r"rn*ribus
albi' esenoribu'
ì 5
2 nm lonÈi'
iDìerioribus
) s-l nÙr
il;;:.;";;"1',;;;"",'. err,p,r,s
rd apicem
rorundaús
r-r
2 mm
rootsrs
ovari!'m
""r'J'a'i"".,'""u-p"ptl1osum
supeme
?Tila
o:il"1'f:ì1'";"'":H,
*"-", *
*i::T:'"TL3#":J,1î,13;liJ'l-ii.il";'
"""t,
20
45 cm
high.
covered
bv
rear
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r- il iJ or'r'
r"ng'r,
r.u'"' sr' $een
'emicvlifflrical
c:5rre:p
ro
r0,cjn
Ìone.
SDarbe
persi.renl.
srtl 2ur'qual \alve' longer
lhen
umbel
lhe lafger./-Derveo
G
rt i. ìonq.,t
" ""tt., < Der\ed
2 s-b cm lons lofloresccnce
(xpdn'e orrrur rr- r

216 Brullo
& al.:
Taxonomic
remarks
on the endemic flora ofthe Sila ...
FiE.2. Adeúa.arput
teturcarlr Bnllo, Gùgale & UaDo! (l),,1. ònftb tstullo. De Marco
&
Siracxsa(2):A- noùer. B-bud. C - open caÌ)a, D bmcreolcs,
E bncts, F stùda|d, C wirys.
H keel I staninaltube. J
- pisLll.
K - leavcs.
t'1,

Bocconca
21 2007 2t'7
flowered; pedicels
unequal,
flexuous, 8-10(40) mm long. Perigon subcylhdrical, with
tepals subequal,
white-greenish with
green mibrid,
oblong
to oblong-obovate.
munded at
apex,4.5-5.5 mm
long,2-3 nrm wide. Stamens simple,
inclùdcd, wilh filaments white,
sub-
ulatc, the outermost
1.5 2 nÌr long, th€ innemost 2.5-3
mn long, connate at base
into an
annulus
0.6-0.8 nìm high;
anthen elliptical,
pale yello\ rounded
at apex 1-1.2
: 0.7 0.9
lnm. Ovary subcylinddcal,
grecnish, scabrid
papillose
3bove,2.5-3 x 1.5 1.8 mm. Style
white,
0.,1-0.8 Ìnm
long. ùpsule obovoid,
geen
4.5 x 4 mm.
lconognphy:
Fig. 3.
Etymoiogy: The species
is dedicated to Prof. Giuliaro Cesca.
fbr his significant contli-
bution to the investigation of úe Calabrian l1ora
Ecology
and distribution: It gows in thc undeÌwood of OrercK crrrs forests at about
800 m altitude
on siliceors substrata
(schists).
It occurs on lhe NE slope of Sila Massif
(Sila
Gr€ca), where
it is very rare and localized.
A very localized and rarc
cndemic species, belonging to the A. secr.
Codoîaprasun, )s
well differentiaied
fionr the other known species of the
A. pani.ulatun grcnp and
ir paî-
ticular it shows
close relations
mainly with L te uíllorun ÎeI', occúi.1tg
h the ltalian
Peninsula
(Brulìo & al, 2002a).
Othet rpecies
endeni. to Sila teftitot!:
Anthenís htdruntína H. Groves subsp. srle"rrr (Fiori) Brullo, Gangale
& Uzunov
A. h)drunnna bsp. hfdnlnrna
is rcstricted to few localities ofPuglia
and Basilicata
(S Ir.ìly), where it occurs
on calcareous rocky
places,
while subsp.
st1",sts is an orophilous
raror, linked 1() siliceous substrata
and Ìocalized in a small area ofthe
eastem slope olthe
Silan Massif(Brullo
& al. 2004).
Coto tríu fetti (L.) t. Cay subsp.
ódqÍertt (Fiorì) Brullo, Gansale
& Uzxnov, comb. &
Type: Calabria Sila a S. Giol.anni in Fiore, \8/611899
, Fiori sub
A. tinttrtia laî clisco
'/?a (Lectoinc: FI!).
Basion.: A. tinctoia
L. var òrtqrerti Fiori, Nuov.
Fl. Anal. Itat. 2: 649
(1927).
S1n.:
A. nnaoria \ar dísuidea Fiori, N. Giom. Bot. lial. 7:
270
(
1900), non DC.
( I
83 7);
A. îriumfeui
\ar. clhc.ùLea
Fiori, ìn Fìori e Paoletti, Fl. Anal.
Ital.
3:
260
(1903) p. p.
Tnistaxon
differs ftom
the type, widely spread in S Eulope, by tne capituladiscoid
and
achenes wiih a well developed coronule.
Anneiú brutíî Btúlo. Gansale & Uzunov
Specics
belongiry to thc TyÍhenian element, lite lm,€r" neó'rdealir (Guss.)
Boiss.
ùom Sicily,
,.1. avlorioflrdrd Brullo.
Scelsi & SparÌpinato
Èom Aspfomontc,l.
ra'do,
SFengel 1ìom Sardinia,l. 'xrltrceps Wallr. from Corsica and,4.87d.i1ll
Ten. from Ceùf.tl
aùd
Soùthem Apennines.
up Ìo Mt. Pollìno
(Brullo
& al.2004).
AÍragalus calabrícus
F isch.
It belongs
to the cycle of Astrugalus thracictts
Gt1seb.,
reFesented by tÀ{.ì djstributed
in the E-Mediterranean
are3.

218 Brullo & dl.: la\onomrc remarks
on the endemic Ilora ofrhe Sila...
lit
F19.3.A tun iulidrútuBtullo,
Gangale
& Uzùnov: A- habit, B flowcr C - opcn
lerigoD
aDd sta-
mens, D - anthe.. E
- ovary
F - capsule.

Bocconea
21 2007 2t9
A'enuta praetutina (Paîl ex Arcans.)
Pignatti
subsp
/ts'74 (Sadatti) Brulìo' Gangale
&
Alenula
praetutiana
is aiApenrÌìnic endemic,
belongiry to the cvcle of'4 v"6t'olo''
wjdely sFead
in C and S Europe
(Brulo & aI 2004) tt is represented
bv two subspec'es:
a calcicoto,rs
one,
lsubsp praetutis a\ hà\lug a dlscontinùou djstribution
on the whole
Apenninic
range
and
a silicicolous
one
(subsp. ,,gÌda), ocoùrring
on the Sila Massif:
Centaúrca
s,líattianaBîu[o' Gangale & Uzunov
It is closelyrelated
to C.
danld, belonging
to the sect P}a/obÚ (Cass
) DC ' which is
represented
i; C and S Italy by manifold ldta, mainlv chasmophiloÙs
or linted to shrub-
cornmrmities
(Brullo & aÌ 2001,2004)
Cafllamine sila a Marhold
& PelÎ,Y
This ta,xon
is hexaploid,
lìke some
populations of C raphanilolía PoÚrÎ
' bvf morpho'
logìcally shows
closer
reìationships
with the diploid Balkan
populatìon of C dcltr Griseb'
(Marhold & al. 2003)
Ge isto sitanaBrl!,llo,Garigale
& Spampinato
It is vicadated
in S Calabria
(Aspromonte)
by G. Dalia 8ru11o'
Scelsi
& Spampinato'
both species
closely rclated ro G. anglica L., a tvpical atlantic etem€nt
distdbuted
m
Nolthen and
Central
Eùope (Brullo & al 2002)'
Knduria dínortca snbsp
sílnn
4 (Grande) Ehrend
Accor{:ling
to EhreDdorfet
(1975\ Knautia àíMica is rep'esented
bv subsp dj'dlird'
distdbuted
; the Balkanic Peninsula'
and subsp st/anú exclusive
oî the Silan Msssit'
which djff€rs ftom the
tlpe mainlv in the
pub€scence of lower slem intemodes
and
in the
longer
petiole of rhe leaves
Koeleria sptende
s s:ubsp
ó'rfia B llo. Gangale
& Uzunov
A silicicolous
taron morphologically
and ecologicallv
wel diftèrentiated
from the other
known subspecies,
which are
nomallv lirted to limestones;
in particular
'(o"le7'd tplet-
,/e'?J subsp.
spleazlem
occÌns
in N Sicih whÌle l( sptende$ s\tbsp stuttdiforu (Beftol
ex Schultis)
ionin is distribùted
in the rest of the calcareous
mountains
ofthe Apennircs
(B 11o & al. 2004).
L zula calabra
TeD
This ta.\on, belonging to the group of EÙoasiatìc Llauld cdmpeltris LaÎtl' vas
describe<rbyienore(isz9)"erpaludíhdSila€'AccorlingtoMìgliaccio(1964)'l
cat
ahra úîîeîs lrom L. campertír subsp rùlgdrr.!, occurring
in Sìla
te itorv too' in laxer
anthela,
longer capitula
and shofer capsule,
stamens'
srylc
and
tePls, aÙd
in the
coloÌrf of
"up"nt" una
t"par". r. ."loòr'a occurs on wetlands
of mounlain
bell of Sila' espccìallv
ìn
swamps,
,Va'das
srcra communities
and along
the border
oî small steams
jn open
places'
whìIe1. canpzsrts is linkcd to dry $asslaùd li is a member
of igrophvlous
commÙìrtles
h elonging
to Luzul o c a L ab
rue
-
N a
ft I e f u n -

220 Brullo
& al.r
Tà\onomic remarks
on the endemic floÈ of the
Sila
.-
Oth€r endemic speci€s occuning on Sila bui ranging. as well, on other Calabrian
siliceous massifs are,4r*emt calabrica
(4rar.) Brullo, Scelsi & Spampinato closcly
rèlàfed Io A. cretica L., Hypericun calabrícuù Spreng.
geographic
vicarìant ofl1. ódlód-
îu,i Iàc.t,
Lereschia thonarti
(Ten.)
Boiss. belonging
io a taxonomically
isolaled mono-
t'plc eent)s,
Solddnella calaólera Krcss
close to ihe SE European S. lirlngdrrcd Simonkai,
Liùodotun brutloi Bzrtolo & Pulvirenti showing n,lorc ancesrral îeatures Ídthin the
genus,
Canlanine battaglíae Cesca & Peruzi related to SW Eltîopean C. heptaphyla (Yrll.) O.
E. Schulz, Sa/r,r r]rrd Brullo & Spampinato, a Calabrian vicariant ofs. trtdadrd L., and
Buglossoides
calabla (Tcn.) Johnston morphologically
well differentiated
fioln B. ?ùr
Durocaerulea L.\ lohnsron .
Besides. well reFesented is the set of Siculo-Calabrian
eùdcmics
tl1at €mphrìsizes the
clos€st
floristic affinities of Sila with Siciliaù flora
rather than
the r€st
of ApeÌmines.
Among these, therc are Caràxd zea,rodrrrrì Guss., Sìlene sicula Ucna, Ranunc ú,
úlptunontdnus Huter, Par4a & Pjgo, Cirsiun w is-demaniiLojac., nola meusanensis
(W.
Bccker) Brullo, PD"s ztgld Amold subsp- .alaAltca (Land., E. M\\ray, Bdrbdred sìculo
C. Presl, Epìpacris s.huberrlorum BaÍolo, Pulvirenti & Robatsch, etc.
Thc
present
study is a 1ìIst contribxtion for fhc chaEcterisation ofthe cndcmic element
of the Silan flora. Flrlther research
on the qualitative and quantitative
features of the
endemics will contrìbute to the interpreradon ofthe Silan and S Calabdan flola genesis
and
ph},togcography.
List ofthe endemic sDecies to the Sila MÀssif
Acer bbelìi'len;ALer edpolitanunTeÍ.; Achillea
tenorii Chàîde;
Acinos
grunate
sìs
(Boiss. & Reuter) Pignatti subsp. delrsnsN (Strobl) Pìgnattia
Ade"ocarpe, terorcunu!
Brullo, Gangale & Uz\tlo\; Afusa tenolei Prcsl;
A iun julianum
Brullo, Cangale &
Uz,r\ov; Anthemís hydrun rlra Groves subsp. st/ezrir (Fiod) Brullo, Gangale & Uzunov;
A îhemi, nontaM L. súbsp. calabrica Aîcàng.; Arabis rosea DC., Arneria brutid Brù11o,
Gangale & Uzunov;lrerx kìa wù)bilk Ten; Astrugalus
caldhrítus Fiotl; Are uld
prae-
rrlrdrd (Parl.) Pign. subsp.
/igiira (Sarfatti)
Brullo, Ga1\gale & Urr.\loy; Balbalea siculd
Pîesl; Bustossoìdes calabf" (Ten.)
Johnston;
-Bùrlrm
petrdeumTen; Campa xla rricoca-
lycina Tct\.; Cdnlanìne bdttdsliae Cescz & Peî!zz|; Carclanine
silana Ma*ioìd & Pemy;
Cdnluus .hlysdcarrhus Ten.: Cdrlina rchtudensis G]uss.; Centawea centdurium L.;
Cenhurca sarfattiana BÍjlo, Gangale & Uzunov; Celdrtium
gra ulatun (ùtter,P. &R.)
Chiov.; Cerdstium s.ard ííTeI'.; Cirsìuù wltis'.len nit Lojao.i Cora |riumfetti (L.) J. Gay
st)bsp. briqueîtii (Fiori) Brullo, Gangale & Uzunov; Dir thus rulrurius Grss. & Ten.
subsp.
wlr,/tur; ,i8?/a/N nicrantha P.oú1: ELhinops
si.ulut Stîobl: Epipdctis ùeidioÈ
dl,i H. Baumann & R. Loîenz; Epipactis schubertionn Bafolo, Pulvirenti & Robatsch;
Euphorbia dmygdalotules L. subsp. dr'órrcrld Meusel: Euphorhia con ioú|a L.; Gagea
chrysdntha
(lan) Schultes; Gali,n aetni.um Biy.; Genistd tilaM Brullo, Gangale &
Spàrnp:nato:
He ehoru! bo.corci Tèn. sxbsp.
irîemedius (Guss.)
Creuter & Bùrdet;
Hieruciun virydurca Coss.;
Hypericum
caldbticun Spîengel Ktlautia caUcina (Ptesl)
Gr\ss.;
Knautia .línaritd (Murb.) BorMs subsp. stldna (Grande) Eltrend..; Koelería splen
/ers C. Presl snbsp. òruria Brullo, Gangale & Uzxnot Larbrrus
iot lanii (Ten.\
Ces., Pass.

221
Bocconea
2l - 2007
& Grb.; Leonto.ton inteme.liue HrtteÎ, P &' R.; Lercschia r'onaili (Ten ) Boiss ì
i^.i" ^ t'*tt"i s*.lo & PulvÍerti; Linaria puryurca (L) M|llel. Luzuk calabra
lr:,.; t*uta 'iebe,i tarsch subsp rrcrld (Parl ) Pisnatii. Paeonia 'ndscut't (L) ]l.{iltet
subs;. '1ljrtt (Biv.) Cullen &' Hew.; Petrorhasia sarty'asa (L ) Linl subsp aasPar'rdtt
1Cuss.) rignatti; Phteum ambiguum Îcn Pìnpinelt'l anisoides Br]gallti; Pinus nigra
l-.ri ""ù"p. "rlrr"-, ir-and.)
E Munav;Rarmztus aspro'nonta
u\ Hrter' P &R;
Ranuncutus'thomasii'Îet; Sdl i/,lld Bmllo & Spampinato; Se'e'io ram';trc"t Huet;
Sitene echindta Orú; Silene siculaucr'a Soldane a calabreUaKrcss; Teucrium sicuLu
R^fri\.; Thalictruù catabrkun Spîeflgel TfifoLíum
brutium Ten; nci't.co sertùa
S$ensel nola aerhnensis
Parl sllbsp. neria,edtr (Vr'' Becker) Merxn & Lippert
A1bo,
G 1935:
Notizie ed oseNàzioni st la floF delve6Ùte setientrionale-oriertalè
della Sila
Nnovo
Giom Bot ltal 42:
623-6ó4.
Ball€1li.
S. & Venanzod,
R. 1993:
Conbrbuto
alta conÒsceMa
delÌa flora e dellà vegctÚionc di ùn
--__' t"o.* Oa Monte Gdiglione (Parco
Nazionàle della Calabrià "Sila Piccola") Atti 5
Workshop
hogetto Slrategico
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e Terilorio ncl Mezzogiomo
(l) Amalfi' 28
30
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Bnlto, S..'De
Mùo, G & Si.eùsa,
G 2001: Taxonomical
notes
Òn the
geúls
'4d€'d'alP&!
DC'
(Les\ninosae) in lr^ly Bocconea
13: 425-436
-. Gùgalq C. & Span nato,
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$ Ge'dra argli'a L speclc
coúllex'
lnfom. Boi ltal 33(2):491-499
-, &Uzunov.
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úugl:elmo.
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& ralmen.
i 20n'd
Os'e^a/ioÙ
ra\oúoniche
e
(arioloeicl'e
"ullc
s;ecie
del
cic lo J
i ,a /r"a ra
ntuldtun L in ltara ldom Bor'lral
llt2r:50050h
.S".il,t.a Sp^1inr,o
C 200l Ld
\cgera/ione delr'A:prornonre
RefCro(!'abt' -
C*n, C'lS&, io"hil*i ulla
conosceúa
delle
ltìdtc di Crlabna:
1 9 GioD Bot Iú1'
115(6):
387-390.
1982: Cdiribùti
alla conosena
delle
liúte di Calabria:
I0-13
- Gìom
Bo| Itaì lr6(l): 182-186
A peruzzi,
r-. ZOOZ:
e new ryecies
of CalddDD' suògen'
De"fdlta (Cru'Élae)' apoerdmic Ù
Calabrìa
(Southen Italv) Plaúi
Biosvl. 136(3): 313_320'
Ehrolalorfer,
F. 1975:
Tuonomic nÒ1es on ,(,a,tia L. Itn: HcFvood H V (cd
)' FÌon F:lúopaeà
Notul;e
Systenancae
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l7l Bor'
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Note
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crlabrcse
e
incaú NÙovo G1om
Bo1
ltal 26:124-l4r'
ló22: Nuove
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Gdtile, S. & Manini.
E 1974:
Novità
della flora
silana Webbia
29:
113 122
G@de,
L. 1913:Nole
dì tloiìsnca
SilÙa BulL
OIto
Bot- NaPÒlì
3:91-110
i"rg..'e. rm:, eppo"f *lh vegetúìonc di alcunc
]Òcalib della calabÌÌa cileriore AÙ Bot
Roma
t:85101
1905:
Conlribuioùe
aÌla
flora calabresc.
E$cmÌore
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3: l-12
Marhold,
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ó9-70
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dell'Allopiùo
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ni""rltJir"", S. A U.iai Afiegui, J 1999:
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Geobot.
13:41-47

222 Brùìlo & al.: Taxonomic remarks on the endemic
flom ofthe Sila ...
Sarfani, G 1959: Podromo della flo.À della SiÌa
(Calabrìa)
lere L webbia 15(1): 169 248.
1965: Podromo della flora della Sila (calabria.) parre
tr- webbia 20(2):355425.
Solla, R. L 1896: OseNàzioúi botúiche dllmte uM escùrsiorc in provincia di Cosena.
Malpighia l0r 168J95, 105-223-
Tenore, M. 1829:
Prodromo della Flora Napoliîùa. supllemdto IIl. Napoli-
Trotter, A. l9l1: Notizie ed osserzio{i sùIa flora montam della Calabria. Nùovo Giom. Bot.Ii.l
r8t 243-278.
Address ofthe uuthors:
Salvatore Brullo, Dipdtimdto di Botúica, UniveBifa di Crtania,
via A. Longo 19.
I-95125 Catùi4 ltaly. E'mil: ònrlo@dipbor.unict.it.
Cameú Gdgale & Dimitù Uanov, Museo di Storia
Naturale d€lla Calabrin ed
OdÒ
Botmico. UriveBita della Calàbris. I-87030 Arcàvacata
di R€lde (CS), ItaÌy E
mils: csaryale@inwird.it, d@nov@libero.it.