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A review of the genus Lepidosperma Labill. (Cyperaceae:
Schoeneae)
Russell L. Barrett
A,B,C,E
and Karen L. Wilson
D
A
Botanic Gardens and Parks Authority, Kings Park and Botanic Garden, West Perth, WA 6005, Australia.
B
School of Plant Biology, Faculty of Natural and Agricultural Sciences, The University of Western Australia,
Crawley, WA 6009, Australia.
C
Western Australian Herbarium, Department of Environment and Conservation, Locked Bag 104,
Bentley Delivery Centre, WA 6983, Australia.
D
National Herbarium of NSW, Royal Botanic Garden, Mrs Macquaries Road, Sydney, NSW 2000, Australia.
Email: karen.wilson@rbgsyd.nsw.gov.au
E
Corresponding author. Email: russell.barrett@bgpa.wa.gov.au
Abstract. Species diversity in the genus Lepidosperma Labill. is much greater than previously thought. On the basis of
morphological and molecular data, we currently recognise 73 named species (mainly in Australia), with many more
species yet to be described. As a precursor to a complete revision, we review the names published in Lepidosperma.
All published names at infrageneric, specific and infraspecific rank are typified and their current taxonomic status is
indicated. Brief distribution notes are given for the 73 named species recognised. We also give a list of names referrable to
other genera. A summary of the taxonomic history of the genus is provided, as well as notes on the specimens collected
by early collectors in Australia. Three new combinations are made in Lepidosperma: L. asperatum (Kük.) R.L.Barrett,
L. neozelandicum (Kük.) R.L.Barrett & K.L.Wilson and L. rigidulum (Kük.) K.L.Wilson. L. sanguinolentum K.L.Wilson is a
nomen novum based on L. drummondii var. floribundum Kük. Lectotypes are designated for eight infrageneric names
and for 39 specific and infraspecific names, including the following: L. angustifolium Hook.f., L. angustatum R.Br.,
L. angustatum var. curvispiculum Benth., L. australe (A.Rich.) Hook.f., L. benthamianum C.B.Clarke, L. brunonianum
Nees, L. brunonianum var. binuciferum Kük., L. canescens Boeckeler, L. carphoides Benth., L. concavum var. pyramidatum
Benth., L. confine Nees, L. costale Nees, L. costale var. densispicatum Kük., L. drummondii Benth., L. effusum Benth.,
L. forsythii A.A.Ham., L. gladiatum Labill., L. globosum Labill., L. inops F.Muell. ex Rodway, L. laterale var. angustum
Benth., L. laterale var. majus Benth., L. leptophyllum Benth., L. leptostachyum Benth., L. leptostachyum var.
asperatum
Kük., L. muelleri Boeckeler, L. neesii Kunth, L. perplanum Guillaumin, L. perteres C.B.Clarke, L. pruinosum Kük.,
L. pruinosum var. rigidulum Kük., L. quadrangulatum A.A.Ham., L. resinosum var. pleianthemum Kük., L. scabrum Nees,
L. scabrum var. effusum Benth., L. sieberi Kunth, L. squamatum Labill., L. tenue Benth., L. viscidum R.Br. and L. viscidum
var. subpyramidale Kük. Twenty-two excluded names are listed and new combinations are provided in Tricostularia for
L. aphyllum R.Br. and L. exsul C.B.Clarke. A lectotype is selected for L. pauciflorum F.Muell. (= Tricostularia pauciflora
(F.Muell.) Benth.).
Received 13 October 2011, accepted 29 June 2012, published online 15 August 2012
Introduction
Species diversity in the genus Lepidosperma Labill. is much
greater than previously thought. On the basis of morphological
and molecular data, we currently recognise 73 named species
(mainly in Australia), with many more species yet to be described.
As a precursor to a taxonomic revision of the genus, for the Flora
of Australia project, we review the names currently accepted as
applying to taxa in this genus (Barrett 2012), as well as provide a
list of excluded names considered referable to other genera.
All published names at infrageneric, specific and infraspecific
rank are typified and their current taxonomic status is indicated.
Images of type specimens are included and brief distribution
notes are given for the 73 named species as an interim aid to
identifying this difficult group.
Lepidosperma (Labillardière 1805) was erected for seven
species cited as being collected from Tasmania (but at least
one was actually from Western Australia). Brown (1810)
enumerated 19 species from across southern Australia. Further
taxa were described by Richard (1832, under Vauthiera A.
Rich.), Kunth (1837), Nees (in Lehmann 1844, under
Machaerina Vahl), Nees von Esenbeck (
1846), Hooker (1853,
1858), Steudel (1855), Boeckeler (1874) and Mueller (1875).
The Australian taxa were then reviewed by Bentham (1878),
who was the first to propose an infrageneric classification
Journal compilation Ó CSIRO 2012 www.publish.csiro.au/journals/asb
CSIRO PUBLISHING
Australian Systematic Botany, 2012, 25, 225–294
Review
http://dx.doi.org/10.1071/SB11037
Full version available from www.publis.csiro.au
Acknowledgements
Russell L. Barrett was the recipient of an Australian Postgraduate Award.
Additional travel funding was provided to R. L. Barrett by the School of Plant
Biology, The University of Western Australia. Aspects of this work were
supported by Alcoa World Alumina (Australia), Worsley Alumina Pty Ltd,
Landcare Services and the Friends of Kings Park. Karen Wilson
acknowledges the support of the Australian Biological Resources Study
when she was Australian Botanical Liaison Officer at the Royal Botanic
GardensKew(1988–1989)andfor a research granttoK.Wilsonand J. J. Bruhl
(2009). She was the recipient of a travel grant from the Friends of the Botanic
Gardens to visit BM, K and LD in 2008. We thank directors of the
following herbaria for permission to reproduce images of type specimens
held in their collections: B, BM, BRI,DNA, FI,G, HO, K, L,LD, MEL,NSW,
P, PERTH. Images from collections at K were reproduced with the
permission of the Board of Trustees, Royal Botanic Gardens, Kew. Images
from MEL were reproduced with permission from the State Botanical
Collection, National Herbarium of Victoria (MEL). Images from P are
from the Muséum National d’Histoire Naturelle collection, Paris. All
images are not to be further reproduced without permission from the
respective herbaria. We thank Zoe Davies, Matt Barrett, Jeremy Bruhl,
Gillian Craig, Rob Davis, Kingsley Dixon, Marco Duretto, Alex George,
Steve Hopper, Greg Keighery, Karina Knight, Siegfried Krauss, Neville
Marchant, Guiseppe Messina, Kelly Shepherd, Cate Tauss, Kevin Thiele,
Stephen Vlahos, Mark Wallace, Juliet Wege, David Willyams, Alastair
Wilson, Paul Wilson and Peter Wilson for their support and advice on
aspects of this research. Jeremy Bruhl is thanked for photographing
specimens at E and FI while serving as ABLO at K and for images of
material on loan to NE from B. John McNeill is thanked for providing
nomenclatural advice on the publication of Machaerina tetragona. Two
anonymous reviewers are thanked for comments that significantly
improved the manuscript. We are grateful to directors, curators and staff
for assisting in our visits to herbaria and organising loans and images of
specimens, including Juergen Kellermann, Rosemary Taplin, Helen Vonow,
Andrea Ramsay, Robyn Barker, Bill Barker and Graham Bell at AD; Ludwig
Martins for his patient assistance and Robert Vogt at B; Rob Huxley and Roy
Vickery at BM; Ailsa Holland, Ron Booth and Gordon Guymer at BRI;
Brendan Lepschi and Jo Palmer at CANB; Giorgio Padovani and Piero
Cuccuini at FI; Nicholas Fumeaux at G; Wieland Kirschner at HBG;
Marco Duretto and Alex Buchanan at HO; Juliet Wege and Jeremy Bruhl
while serving as ABLO at K; David Simpson at K; J. Veldkamp, Peter
Audiffred and Gerard Thijsse at L; Ingvar Kärnefelt, Patric Frodin and Arne
Thell at LD; Josephina Milne, Valantino Stajsic, Wayne Gebert, Neville
Walsh, Catherine Gallagher and Jill Thurlow at MEL; Jeremy Bruhl and Ian
Telford at NE; Barbara Briggs and Peter Wilson (for nomenclatural advice),
Leonie Stanberg and Clare Herscovitch at NSW; Wayt Thomas at NY;
Grégoire Flament and Philippe Morat at P; Karina Knight, Kevin Thiele,
Kelly Shepherd, Juliet Wege, Meriel Falconer, Susan Carroll, Cheryl Parker,
Skye Coffey and Louise Biggs at PERTH; and Mark Strong at US.
References
Bailey FM (1902) ‘The Queensland flora, part V. Loranthaceae to
Lemnaceae.’ (H. J. Diddams and Co.: Brisbane)
Barker RM (2005) James Drummond’s plant collections today – a global
dispersal. In ‘The Drummond symposium: a review of the work of James
Drummond, the first Government Botanist in Western Australia, held at
Toodyay,WA,by the Toodyay Naturalists’ Club,27August 2004’. Curtin
University of Technology. Department of Environmental Biology
Bulletin no. 27. (Ed. SJJF Davies) pp. 40–57. (Curtin University of
Technology. Department of Environmental Biology: Perth)
Barrett RL (2007a) New species of Lepidosperma (Cyperaceae)
associated with banded ironstone in southern Western Australia.
Nuytsia 17,37–60.
Barrett RL (2007b) Lepidosperma gahnioides, a new species of Cyperaceae
from the Ravensthorpe Range, Western Australia. Nuytsia 17,61–66.
Barrett RL (2012) Systematic studies in Cyperaceae tribe Schoeneae:
Lepidosperma and allied genera. PhD Thesis. The University of
Western Australia, Perth.
Barrett RL. Description of six species of Lepidosperma based on type
specimens. Nuytsia, in press.
Bebber DP, Carine MA, Wood JRI, Wortley AH, Harris DJ, Prance GT,
Davidse G, Paige J, Pennington TD, Robson NKB, Scotland RW (2010)
Herbaria are a major frontier for species discovery. Proceedings of the
National Academy of Sciences of the United States of America 107,
22 169–22 171. doi:10.1073/pnas.1011841108
Bennett EM, Dundas P (1988) ‘The bushland plants of Kings Park Western
Australia.’ (Kings Park Board: Perth)
Bentham G (1863) ‘Flora Australiensis: a description of the plants of the
Australian Territory. Vol. I. Ranunculaceae to Anacardiaceae.’ (Lovell
Reeve: London)
Bentham G (1878) ‘Flora Australiensis: a description of the plants of the
Australian Territory. Vol. VII. Roxburghiaceae to Filices.’ (Lovell Reeve:
London)
Blackall WE, Grieve BJ (1974) ‘How to know Western Australian
wildflowers. Parts I, II, III.’ (University of Western Australia Press: Perth)
Blake ST (1943) Critical notes on the Gramineae and Cyperaceae of South
Australia with descriptions of new species. Transactions of the Royal
Society of South Australia 67,42–61.
Blake ST (1949) Notes on Australian Cyperaceae, VII. Proceedings of the
Royal Society of Queensland 60,45–53.
Boeckeler JO (1874) Lepidosperma.In‘Die Cyperaceen des königlichen
Herbarium zu Berlin’. Linnaea 38, 315–338.
Boeckeler JO (1896) Diagnosen neuer Cyperaceen. Allgemeine Botanische
Zeitschrift für Systematik, Floristik, Pflanzengeographie 2, 109–113.
Botanical Museum Berlin-Dahlem (1999) List of families including extant
collections of the Botanical Museum Berlin-Dahlem (B) from the time
before 1943. Available at http://www.bgbm.fu-berlin.de/BGBM/
research/colls/herb/phanerog.htm [Verified December 2010].
Brown R (1810) ‘Prodromus florae Novae-Hollandiae et insulae van-
Diemen.’ (Taylor: London)
Bruhl JJ (1995) Sedge genera of the world: relationships and a new
classification of the Cyperaceae. Australian Systematic Botany 8,
125–305. doi:10.1071/SB9950125
Brummitt RK (2011) Report of the Nomenclature Committee for Vascular
Plants: 62. Taxon 60, 226–232.
Buchanan AM (1990) Ronald Campbell Gunn (1808–1881). In ‘ History of
systematicbotany in Australasia’. (Ed.PSShort)pp. 179–192. (Australian
Systematic Botany Society: Melbourne)
Burbidge NT, Gray M (1970) ‘Flora of the Australian Capital Territory.’
(Australian National University Press: Canberra)
Carr SGM, Carr DJ (1976) The French contribution to the discovery of
Australia and its flora. Endeavour 35(124), 21–26.
doi:10.1016/0160-9327(76)90058-2
Chapman AD (1991) ‘Australian plant name index, K–P.’
Australian Flora
and Fauna series number 14. (Australian Biological Resources Study:
Canberra) Available at http://www.anbg.gov.au/cpbr/databases/apni.
html [Verified March 2012].
Cheeseman TF (1906) ‘Manual of the New Zealand Flora.’ (Government
Printer: Wellington, New Zealand)
ClarkeCB(1894) Cyperaceae.In ‘Conspectus florae Africae. Vol.5.’ (EdsTA
Durand, H Schinz) pp. 526–692. (Jardin Botanique de l’Etat: Bruxelles)
Clarke CB (1897–1898) Cyperaceae. In ‘Flora Capensis: being a systematic
description of the plants of the Cape Colony, Caffraria, & Port Natal (and
neighbouring territories)’. (Ed. WT Thiselton-Dyer) pp. 149–310.
(L. Reeve & Co.: Kent, UK)
Clarke CB (1908) New genera and species of Cyperaceae. Kew Bulletin
Additional Series 8,1–196.
A review of the genus Lepidosperma Labill. (Cyperaceae: Schoeneae) Australian Systematic Botany 289
Crisp MD (1983) Plantae Preissianae types at Lund. Australian Systematic
Botany Society Newsletter 36,4–6.
Dashorst GRM, Jessop JP (2006) ‘Plants of the Adelaide plains and hills.’
(The Board of the Botanic Gardens of Adelaide & State Herbarium:
Adelaide)
Diels L (1906) ‘Vegetation der Erde. VII. Die Pflanzenwelt von West-
Australien südlich des Wendekreises.’ (W. Engelmann: Leipzig,
Germany). [Translated to English by BJ Grieve, BB Lamont and EO
Hellmuth and republished in Conservation Science Western Australia 6
(2): 1–373 (2007).]
Dietrich FC (1881) Franz Wilhelm Sieber – ein Beitrag zur Geschichte der
Botanik vor sechzig Jahren. In ‘Jahrbuch des Königlichen Botanischen
Gartens und des Botanischen Museums zu Berlin’. (Ed. AW Eichler)
pp. 278–306. (Royal Botanical Museum Berlin-Dahlem: Berlin)
[F. W. Sieber: a contribution to the history of botany sixty years ago.
In ‘ Yearbook of the Royal Botanical Museum Berlin-Dahlem for 1881’.
English translation by A Wilson, T Schuster and S Stark at NSW.]
Ducker SC (1990) Early Austrian influence on Australian botany. In ‘History
of systematic botany in Australasia’. (Ed. PS Short) pp. 297–304.
(Australian Systematic Botany Society: Melbourne)
Duyker E (2003) ‘Citizen Labillardière – a naturalist’s life in revolution and
exploration (1755–1834).’ (Miegunyah Press: Melbourne)
Eckardt T, Hiepko P (1970) Missing Willdenow specimens. Taxon 19,
951–952.
Edgar E (1970) Cyperaceae. In ‘Flora of New Zealand. Vol. II. Indigenous
Trachyophyta. Monocotyledones except Gramineae’. (Eds LB Moore, E
Edgar) pp. 167–285. (A. R. Shearer, Government Printer: Wellington,
New Zealand)
Elliot WR, Jones DL (1993) ‘Encyclopaedia of Australian plants suitable for
cultivation. Vol. 6, K–M.’ (Thomas C. Lothian: Melbourne)
Erickson R (1969) ‘The Drummonds of Hawthornden.’ (Lamb Paterson Pty
Ltd: Perth)
Ewart AJ (1930) ‘Flora of Victoria.
’ (Government Printer for the University
Press: Melbourne)
George AS (2009) ‘Australian botanist’s companion.’ (Four Gables Press:
Perth)
Germishuizen G, Meyer NL (2003) (Eds) ‘Plants of southern Africa: an
annotated checklist.’ Strelitzia 14. (National Botanical Institute: Pretoria,
South Africa)
Goetghebeur P, Vande Plassche D (1986) On the use and citation of true and
false homonyms. Taxon 35, 321–323. doi:10.2307/1221279
Govaerts R, Simpson DA, Bruhl JJ, Egorova T, Goetghebeur P,
Wilson KL (2007) ‘World checklist of Cyperaceae.’ (Kew Publishing:
London)
Guillaumin A (1938) Matériaux pour la flore de la Nouvelle-Calédonie.
L. Révision des Cypéracées. Bulletin de la Société Botanique de
France 85,37–47.
Hamilton AA (1910) A new species of Lepidosperma [N.O. Cyperaceae]
from the Port Jackson district; with some miscellaneous botanical
notes. Proceedings of the Linnean Society of New South Wales 35,
411–415.
Hamilton AA (1920) Notes from the Botanic Gardens, Sydney. Proceedings
of the Linnean Society of New South Wales 45, 260–264.
Harris SA, Buchanan A, Connolly A (2001) ‘One hundred islands: the flora of
the Outer Furneaux.’ (Tasmanian Department of Primary Industries,
Water and Environment: Hobart)
Hiepko P (1970) List of missing Willdenow specimens with
herbarium number and names of donors. Taxon 19, 952–953.
doi:10.2307/1218327
Hiepko P(1987) The collections of the Botanical Museum Berlin-Dahlem (B)
and their history. Englera 7, 219–252.
Hodgon J, Bruhl JJ, Wilson KL (2006) Systematic studies in Lepidosperma
(Cyperaceae: Schoeneae) with particular reference to L. laterale.
Australian Systematic Botany 19, 273–288. doi:10.1071/SB06007
Holmgren PK, Holmgren NH (1998) Index Herbariorum: a global directory
of public herbaria and associated staff. New York Botanical Garden’s
Virtual Herbarium. Available at http://sweetgum.nybg.org/ih/ [Veri
fied
March 2012].
Home RW, Lucas AM, Maroske S, Sinkora DM, Voigt JH (2002) (Eds)
‘Regardfully yours: selected correspondence of Ferdinand von Mueller,
vol. 2: 1860–1875.’ (Peter Lang: Bern, Switzerland)
Hooker JD (1853) ‘Flora Novae-Zelandiae, I.’ (Reeve: London)
Hooker JD (1858) ‘The botany of the Antarctic voyage of H.M. Discovery
Ships ‘Erebus’ and ‘Terror’, in the years 1839–1843, under the command
of Captain Sir James Clark Ross. Vol. II. Flora Tasmaniae.’ (Reeve:
London)
Horner F (1995) ‘Looking for La Pérouse – D’Entrecasteaux in Australia and
the South Pacific 1792–1793.’ (Miegunyah Press: Melbourne)
Jessop JP (Ed.) (1978) ‘Flora of South Australia. Vol. 1.’ 3rd edn.
(Government Printer: Adelaide)
Jessop JP, Weber JZ (1986) Cyperaceae. In ‘Flora of South Australia. Vol. 4.’
4th edn. (Eds JP Jessop, HR Toelken) pp. 2001–2053. (South Australian
Government Printing Division: Adelaide)
Kern JH (1974) Cyperaceae. Flora Malesiana Series I 7, 435–753.
Kirkbride JH (2007) A 19th century Brazilian botanical dictionary. Taxon 56,
927–937. doi:10.2307/25065874
Koyama T (1956) Taxonomic studies of the Cyperaceae V. x 12. A new
classification of the genus Cladium in a wide meaning. Botanical
Magazine, Tokyo 69,59–67.
KoyamaT(1961)Taxonomicstudiesof Cyperaceae 12. Naturaliste Canadien
88, 237–252.
Kraehenbuehl DN (1986) History of botany in South Australia (1800–1955).
In ‘
Flora of SouthAustralia. I.Lycopodiaceae–Rosaceae’. (Eds JP Jessop,
HR Toelken) pp. 13–39. (South Australian Government Printing
Division: Adelaide)
Kükenthal G (1941a) Vorarbeiten zu einer Monographie der
Rhynchosporoideae.X.Repertorium Novarum Specierum Regni
Vegetabilis 50,19–50. doi:10.1002/fedr.19410500104
Kükenthal G (1941b) Vorarbeiten zu einer Monographie der
Rhynchosporoideae. XI. Repertorium Novarum Specierum Regni
Vegetabilis 50, 112–128. doi:10.1002/fedr.19410500606
Kükenthal G (1942) Vorarbeiten zu einer Monographie der
Rhynchosporoideae. XIII. Repertorium Novarum Specierum Regni
Vegetabilis 51, 139–193. doi:10.1002/fedr.19420510803
Kükenthal G (1944) Vorarbeiten zu einer Monographie der
Rhynchosporoideae. XV. Repertorium Novarum Specierum Regni
Vegetabilis 53, 187–219. doi:10.1002/fedr.19440530304
Kunth CS (1837) ‘Enumeratio plantarum omnium hucusque cognitarum,
secundum.’ (J.G. Cottae: Stuttgart, Germany)
Labillardière JJHd (1805) ‘Novæ Hollandiæ plantarum specimen. Vol. 1, part
2.’ [1966 reprint]. (J. Cramer: Lehre, Germany)
Lander NS (1987) Asteraceae specimens collected by Johann August
Ludwig Preiss. Kingia 1,9–19.
le Moore SM (1902) Some new species from Australia. The Journal of Botany
– British and Foreign 40,25–30.
Lehmann JGC (1844) ‘Novarum et minus cognitarum stirpium pugillus
Vol. 8.’ (J.A. Meissner: Hamburg, Germany)
Levyns MR (1950) Cyperaceae. In ‘Flora of the Cape Peninsula’. (Eds
RS Adamson, TM Salter) pp. 97–132. (Juta & Co.: Cape Town, South
Africa)
Link JHF, Otto CF (1820) ‘Icones plantarum selectum. Part 1’ pp. 69–
80
(J. H. F. Link and C. F. Otto: Berlin)
Marchant NG (1990) The Western Australian collecting localities of
J. A. L. Preiss. In ‘History of systematic botany in Australasia’. (Ed.
PS Short) pp. 131–135. (Australian Systematic Botany Society:
Melbourne)
McGillivray DJ (1975) Johann August Ludwig Preiss (1811–1883) in
Western Australia. Telopea 1,1–18.
290 Australian Systematic Botany R. L. Barrett and K. L. Wilson
McNeill J, Barrie FR, Burdet HM, Demoulin V, HawksworthDL, Marhold K,
Nicolson DH, Prado J, Silva PC, Skog JE, Wiersema JH, Turland
NJ (2006) (Eds) ‘International code of botanical nomenclature
(Vienna Code) adopted by the Seventeenth International Botanical
Congress Vienna, Austria, July 2005.’ Regnum Vegetabile Vol. 146
(A.R.G. Gantner Verlag: Ruggell, Liechenstein) Available at http://
ibot.sav.sk/icbn/main.htm [Verified April 2009].
Moore DT (2005) Robert Brown on HMS Investigator 1801–1805: an
overview of the natural history collecting localities. In ‘ Matthew
Flinders and his scientific gentlemen. The expedition of HMS
Investigator to Australia, 1801–05’. (Eds JA Wege, AS George,
J Gathe, KL Lemson, K Napier) pp. 49–65. (Western Australian
Museum: Perth)
Morris DI (1994) Lepidosperma.In‘The student’s flora of Tasmania.
Part 4B. Angiospermae: Alismataceae to Burmanniaceae.’ (Eds
WM Curtis, DI Morris) pp. 120–127, 425. (St David’s Park
Publishing: Hobart)
Mueller FJH (1855). Annual report from the GovernmentBotanist for the year
1854. Victoria. Parliamentary Papers Votes and Proceedings of the
Legislative Council 1854. (A. No. 10), pp. 1–20. (Government Printer:
Melbourne)
Mueller FJHv (1875) ‘Fragmenta phytographiae Australiae. Vol. 9.’
(Melbourne Printers: Melbourne)
Mueller FJHv (1888) ‘Key to the system of Victorian plants. Dichotomous
arrangement of the orders, genera and species of the native plants, with
annotations of primary distinctions and supporting characteristics.’
(Government Printer: Melbourne)
Nees von Esenbeck CG (1846) Lepidosperma.In‘Plantae Preissianae
sive enumeratio plantarum, quas in Australasia Occidentali et
Meridionali-occidentale annis 1838–1841 collegit Ludovicus Preiss,
Ph. Dr. Vol. 2(1)’. (Ed. JGC Lehmann) pp. 89–93. (Meissner:
Hamburg, Germany)
Nelson EC (1974) The locations of collection and collectors of specimens
described by Labillardière in ‘Novae Hollandiae Plantarum Specimen’:
additional notes. Papers and Proceedings of the Royal Society of
Tasmania 108, 159–170.
Nelson EC (1975) The collectors and type locations of some of Labillardière’s
‘Terra Van-Leuwin
’ (Western Australia) specimens. Taxon 24, 319–336.
doi:10.2307/1218341
Orchard AE (1999) A history of systematic botany in Australia. In ‘Flora of
Australia. Vol. 1. Introduction’. 2nd edn. (Ed. AE Orchard) pp. 11–103.
(ABRS: Canberra; and CSIRO Publishing: Melbourne)
Orchard AE, Wilson AJG (2004) Cassinia glossophylla Cass. is Ozothamnus
leptophyllus (G.Forst.) Breitw. & J.M.Ward (Asteraceae). Australian
Systematic Botany 17, 567–570. doi:10.1071/SB04033
Pinto J de A (1873) ‘Diccionario de botanica brasileira ou compendio dos
vegetaes do Brasil, tanto indigenas como acclimados revista por uma
commissão da Sociedade Vellosiana, e approvada pela Faculdade de
Medicina da Corte.’ (Typographia-Perseveran¸ca: Rio de Janeiro)
Potts BM, Kantvilas G, Jarman J (Eds) (2006) ‘Janet Somerville’s botanical
history of Tasmania 1642–1820.’ (University of Tasmania and
Tasmanian Museum and Art Gallery: Hobart)
Raymond M (1961) Revision of the Rhynchosporoideae of Indo-China and
Thailand. Naturaliste Canadien 88,8–24.
RaynalJ (1972) Notes cypérologiques:17.RévisiondesCladium P. Browne s.
lat. (Cyperaceae) de Madagascar et des Mascareignes. Adansonia. Sér. 2
12, 103–112.
Richard A (1832) ‘Essai d’une Flore de la Nouvelle-Zélande.’ (Tastu: Paris)
Ridley HN (1915) An expedition to Mount Menuang Casing, Selangor.
XIII. The botany of Gunong Tahan, Pahang. Journal of the Federated
Malay States Museums 6, 127–202.
Ridley HN (1925) ‘The flora of the Malay Peninsula. Vol.
V–Monocotyledones (concluded), Gymnospermeae, General indices.’
(L. Reeve & Co.: London)
Rodway L (1893) Notes on some plants new to Tasmania. Papers and
Proceedings of the Royal Society of Tasmania 1892,92–93.
Rodway L (1894) Botanical notes. Papers and Proceedings of the Royal
Society of Tasmania 1893, 179–187.
Rodway L (1895) Botanical notes. Papers and Proceedings of the Royal
Society of Tasmania 1894–1895, 102–104.
Rodway L (1903) ‘The Tasmanian Flora.’ (Government Printer: Hobart)
Roemer JJ, Schultes JA (1817) ‘Systema vegetabilium. Vol. 2.’ (J.G. Cottae:
Stuttgart, Germany)
Ross JH (1995) Some insights into F. J. H. Mueller’s modus operandi.
Australian Systematic Botany Society Newsletter 84,4–6.
Ross JH (2006) A conspectus of the Western Australian Bossiaea species
(Bossiaeeae: Fabaceae). Muelleria 23,15–143.
Rye BL (1987) Cyperaceae. In ‘Flora of the Perth region’. (Eds NG Marchant,
JR Wheeler) pp. 870–906. (Western Australian Herbarium: Perth)
Schrader HA (1821) Analecta ad floram Capensem. Sect. II. Cyperaceae.
Gramineae. Göttingische gelehrte Anzeigen 3, 2065–2079.
Schrader HA (1832) ‘Analecta ad Floram Capensum. Vol. 1.’ (Dietrich:
Göttingen, Germany)
Schultes JA (1824) ‘Mantissa in volumen secundum systematis
vegetabilium.’ (J.G. Cottae: Stuttgardt)
Sprengel KPJ (1824) ‘Systema Vegetabilium.’ 16th edn. Vol. 1. (Dietrich:
Göttingen, Germany)
Stafleu F, Cowan R (1976) ‘Taxonomic Literature – A Selective Guide to
Botanical Publications and Collections with Dates, Commentaries and
Types.’ 2nd edn. Regnum Vegetabile series, vol. I. A–G. (Bohn,
Scheltema and Holkema: Utrecht, The Hague)
Stafleu F, Cowan R (1979) ‘Taxonomic Literature – A Selective Guide to
Botanical Publications and Collections with Dates, Commentaries and
Types.’ 2nd edn. Regnum Vegetabile series, vol. II. H–Le. (Bohn,
Scheltema and Holkema: Utrecht, The Hague)
Stafleu F, Cowan R (1981) ‘Taxonomic Literature – A Selective Guide to
Botanical Publications and Collections with Dates, Commentaries and
Types.’ 2nd edn. Regnum Vegetabile series, vol. III. Lh–O. (Bohn,
Scheltema and Holkema: Utrecht, The Hague)
Stafleu F, Cowan R (1983) ‘Taxonomic Literature – A Selective Guide to
Botanical Publications and Collections with Dates, Commentaries and
Types.’ 2nd edn. Regnum Vegetabile series, vol. IV. P–Sak. (Bohn,
Scheltema and Holkema: Utrecht, The Hague)
Stafleu F, Cowan R (1985) ‘Taxonomic Literature – A Selective Guide to
Botanical Publications and Collections with Dates, Commentaries and
Types.’ 2nd edn. Regnum Vegetabile series, vol. V. Sal–Ste. (Bohn,
Scheltema and Holkema: Utrecht, The Hague)
Stafleu F, Cowan R (1986) ‘Taxonomic Literature – A Selective Guide to
Botanical Publications and Collections with Dates, Commentaries and
Types.’ 2nd edn. Regnum Vegetabile series, vol. VI. Sti–Vuy. (Bohn,
Scheltema and Holkema: Utrecht, The Hague)
Stafleu F, Cowan R (1988) ‘Taxonomic Literature – A Selective Guide to
Botanical Publications and Collections with Dates, Commentaries and
Types.’ 2nd edn. Regnum Vegetabile series, vol. VII. W–Z. (Bohn,
Scheltema and Holkema: Utrecht, The Hague)
Stafleu F, Cowan R, Mennega EA (1992) ‘Taxonomic Literature –
A Selective Guide to Botanical Publications and Collections
with Dates, Commentaries and Types.’ 2nd edn. Regnum Vegetabile
series, suppl. I. A–Ba. (Bohn, Scheltema and Holkema: Utrecht, The
Hague)
StafleuF, CowanR,Mennega EA (1993)‘TaxonomicLiterature – A Selective
Guide to Botanical Publications and Collections with Dates,
Commentaries and Types.’ 2nd edn. Regnum Vegetabile series, suppl.
II. Be–Bo. (Bohn, Scheltema and Holkema: Utrecht, The Hague)
Stafl
eu F, CowanR, Mennega EA (1995)‘Taxonomic Literature – A Selective
Guide to Botanical Publications and Collections with Dates,
Commentaries and Types.’ 2nd edn. Regnum Vegetabile series, suppl.
III. Br–Ca. (Bohn, Scheltema and Holkema: Utrecht, The Hague)
A review of the genus Lepidosperma Labill. (Cyperaceae: Schoeneae) Australian Systematic Botany 291
StafleuF, CowanR,Mennega EA (1997)‘TaxonomicLiterature – A Selective
Guide to Botanical Publications and Collections with Dates,
Commentaries and Types.’ 2nd edn. Regnum Vegetabile series, suppl.
IV. Ce–Cz. (Bohn, Scheltema and Holkema: Utrecht, The Hague)
StafleuF, CowanR,Mennega EA (1998)‘TaxonomicLiterature – A Selective
Guide to Botanical Publications and Collections with Dates,
Commentaries and Types.’ 2nd edn. Regnum Vegetabile series, suppl.
V. Da–Di. (Bohn, Scheltema and Holkema: Utrecht, The Hague)
Stanley TD, Ross EM (1989) ‘Flora of south-eastern Queensland. Vol. 3.’
(Queensland Department of Primary Industries: Brisbane)
Stearn WT (1992) ‘Botanical Latin.’ (Timber Press: Portland, OR)
Steinberg CH (1977) The collectors and collections in the Herbarium Webb.
Webbia 32,1–49.
Steudel EG(1855) ‘Synopsis plantarum Glumacearum. Pars II. Cyperaceae et
familiae affines, Restiaceae, Eriocauloneae, Xyrideae, Desvauxieae,
Junceae.’ (J.B. Metzler: Stuttgart, Germany)
Strong MT (1997) Machaerina (Cyperaceae) in South America. Novon 7,
308–319. doi:10.2307/3391947
Vahl M (1805) ‘Enumeratio plantarum, vel ab alium vel ab ipso observatum,
cum earum differentiis specificis, synonymis selectis et descriptionibus
succinctis.’ Vol. 2. (J.H. Schubothe: Copenhagen)
Vallance TG, Moore DT, Groves EW (2001)‘Nature’s investigator. The diary
of Robert Brown in Australia, 1801–1805.’ (Australian Biological
Resources Study: Canberra)
Wakefield NA (1953) Two new species of Lepidosperma. The Victorian
Naturalist 70,75–78.
Western Australian Herbarium (1998) ‘FloraBase – The Western Australian
Flora.’ Department of Environment and Conservation. Available at http://
fl
orabase.dec.wa.gov.au/ [Verified 8 June 2011].
Wheeler JR, Graham L (2002) Cyperaceae. In ‘Flora of the South West.
Bunbury, Augusta, Denmark. Flora of Australia Supplementary Series
12’. (Ed. JR Wheeler) pp. 263–298. (Australian Biological Resources
Study: Canberra; Conservation and Land Management: Como; and
University of Western Australia Press: Perth)
Willis JH, Pearson D, Davis MT, Green JW (1986) Australian plants:
collectors and illustrators 1780s–1980s. Western Australian Herbarium
Research Notes 12,1–111.
Wilson KL (1980) Notes on some Australian species of Cyperaceae. Telopea
1, 457–467.
Wilson KL (1981a) Revision of the genus Mesomelaena (Cyperaceae).
Telopea 2, 181–195.
Wilson KL (1981b) A synopsis of the genus Scirpus sens. lat. (Cyperaceae) in
Australia. Telopea 2, 153–172.
Wilson KL (1981c) Lepidosperma.In‘ Flora of Central Australia’. (Ed. JP
Jessop) pp. 513. (Australian Systematic Botany Society/A.H. &
A.W. Reed: Sydney)
Wilson KL (1993) Cyperaceae. In ‘Flora of New South Wales. Vol. 4’. (Ed.
GJ Harden) pp. 293–396. (University of New South Wales Press:
Sydney)
Wilson KL (1994a) New taxa and combinations in the family Cyperaceae in
eastern Australia. Telopea 5, 589–625.
Wilson KL (1994b) Cyperaceae. In ‘Flora of Victoria. Vol. 2. Ferns and allied
plants, conifers and monocotyledons’. (Eds NG Walsh, TJ Entwisle)
pp. 238–356. (Inkata Press: Melbourne)
Wilson KL (1997) New species of Cyperaceae in Western Australia. Nuytsia
11, 269–282.
Wilson KL, Morris DI (1993) Lepidosperma curtisiae (Cyperaceae), a new
Australian species. Telopea 5, 423–425.
292 Australian Systematic Botany R. L. Barrett and K. L. Wilson
