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Typification of Sicilian Helichrysum (Compositae) Revisited

November 2007
Taxon 56(4):1285-1288

DOI:10.2307/25065922

Authors:

Mariam Aghababyan

Freie Universität Berlin

Werner Greuter

University of Palermo

Pietro Mazzola

University of Palermo

Francesco Maria Raimondo

University of Palermo

Typification of the 13 validly published, pre-1958 names of Helichrysum taxa based on Sicilian material is reassessed. Five of the types designated by Galbany-Casals & al. in their recent studies are confirmed (in some cases with additional precision), two neotypes proposed by them are superseded by lectotypes, and six lectotypes are newly designated. The correct name of the species known as H. rupestre, an illegitimate name, is H. panormitanum (or, if taken in a wide sense, H. pendulum).

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Aghababyan & al. • Typification of Sicilian HelichrysumTA XO N 56 (4) • November 2007: 1285–1288

INTRODUCTION

As a forerunner to their recent taxonomic revision

and nomenclatural conspectus of Mediterranean Heli-

chrysum sect. Stoechadina (Galbany-Casals & al., 2006a,

b), Galbany-Casals & al. (2006c) typified a number of

relevant names. In some cases, their conclusions differ

from those we had reached in the context of our current

project on the typification of names of Sicilian taxa of

Compositae. Furthermore, some names relevant in our

context either remained untypified or were omitted alto-

gether from their accounts. The following note is offered

to comment, correct and complement their Sicilian type

designations. To the best of our knowledge, it takes into

account all validly published names based on Sicilian

material that pertain to the genus Helichrysum, except

those which, having been published later than 1957, are

holotypified. Our list does not include Helichrysum scan-

dens Guss. (Fl. Sicul. Syn. 2: 465. 1844), said by Galbany-

Casals & al. (2006b: 64) to have been described from

“Cassaro, e Ferla” in Sicily, because its lectotype must

be a specimen of a Sieber gathering from “Chalepha”

[Halepa] in West Crete, which, being cited in the proto-

logue, has syntype status (Arts. 9.4 & 9.10 of the ICBN,

McNeill & al., 2006).

TYPIFICATION

Gnaphalium cespitosum C. Presl in Presl & Presl, Delic.

Prag.: 98. 1822 (non Lam., Encycl. 2: 742. 1788) ≡

Gnaphalium siculum Spreng., Syst. Veg. 3: 476. 1826

≡ Helichrysum elegans C. Presl, Fl. Sicul.: xxix. 1826

[non G. Don 1825] ≡ Helichrysum cespitosum DC.,

Prod r. 6: 182. 1838, nom illeg. ≡ Helichrysum sicu-

lum (Spreng.) Boiss., Fl. Orient. 3: 229. 1875 [non

Jord. & Fourr. 1868] [= Helichrysum stoechas subsp.

barrelieri (Ten.) Nyman] – Lectotype (designated

here): “Helichrysum elegans Presl, Gnaphalium el-

egans [sic!] Presl. Habitat: Pascua siccissima montis

la Nivera ad S. Martinum prope Panormum. Floruit

Junio 1817, leg. Presl ” (PR, photo!) [handwritten

data are all by Čelakovský; original label not pre-

served].

Not in Galbany-Casals & al. (2006c), not typified by

Galbany-Casals & al. (2006a, b).

No original material could be traced in PRC (J.

Stepánek, pers. comm.). In PR, the second likely loca-

tion of Presl specimens, a single sheet exists, searched

out for us by O.

ida who most helpfully provided colour

photographs at various scales. The traditional interpreta-

tion of Presl’s name is thereby confirmed.

The nomenclature of the East Mediterranean repre-

sentative of the Helichrysum stoechas complex, treated

as a separate species by many, as a mere synonym of H.

stoechas (L.) Moench by Galbany-Casals & al. (2006a, b)

and as a subspecies by us, has been unravelled by Greuter

& Rechinger (1967). Our present nomenclatural assess-

ment of the names involved agrees with the synonymy

given there. An additional example in the Vienna Code

(Art. 7 Ex. 5) confirms that H. cespitosum DC., based on

the illegitimate Gnaphalium cespitosum C. Presl and itself

illegitimate because Gnaphalium

siculum

Spreng. 1826

is cited in synonymy, does not have the same type as the

latter because Candolle’s use of the epithet cespitosum is

equivalent to definite indication of a different type. H.

cespitosum DC., although illegitimate, takes precedence

over its coetaneous homonym H. cespitosum (Lam.) DC.

(Prodr. 6: 173. 1838), as Buek (1840) retained it while

Typification of Sicilian Helichrysum (Compositae) revisited

Mariam Aghababyan1, Werner Greuter2, Pietro Mazzola1 & Francesco Maria Raimondo1

1 Dipartimento di Scienze Botaniche dell’Università, via Archirafi 38, 90123 Palermo, Italy.

m_agababian@hotmail.com (author for correspondence)

2 Botanischer Garten und Botanisches Museum Berlin-Dahlem, Freie Universität Berlin, Königin-Luise-

Str. 6– 8, 14195 Berlin, Germany

Typification of the 13 validly published, pre-1958 names of Helichrysum taxa based on Sicilian material is

reassessed. Five of the types designated by Galbany-Casals & al. in their recent studies are confirmed (in

some cases with additional precision), two neotypes proposed by them are superseded by lectotypes, and six

lectotypes are newly designated. The correct name of the species known as H. rupestre, an illegitimate name,

is H. panormitanum (or, if taken in a wide sense, H. pendulum).

KEYWORDS: Gnaphalium, Helichrysum, nomenclature, Sicily, typification

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TA XO N 56 (4) • November 2007: 1285–1288Aghababyan & al. • Typification of Sicilian Helichrysum

renaming the latter H. barbellatum H. Buek (see ICBN

Art. 53.6).

Gnaphalium pendulum C. Presl in Presl & Presl, Delic.

Prag.: 97. 1822 ≡ Helichrysum pendulum (C. Presl)

C. Presl, Fl. Sicul.: xxix. 1826 – Lectotype (designated

here): “Helichrysum pendulum Pr., Gnaphalium pen-

dulum Pr. del. Pendulum in praeruptis mont. Scalune

Nebrodum. ђ Jul. 1817” [original label in Presl’s own

handwriting] (PR 616046, photo!).

Not in Galbany-Casals & al. (2006c), not typified by

Galbany-Casals & al. (2006a, b).

No original material could be traced in PRC (J.

Stepánek, pers. comm.). In PR, three sheets exist of what

appears to be a single gathering, of which O.

ida provided

excellent colour photographs at various scales. One speci-

men (sheet PR 616045) is from the Sternberg herbarium

and lacks an original label. The two other sheets were

separated upon being mounted and had a single label with

them (sheet PR 616047 has a photocopy of that same la-

bel).

Gnaphalium rupestre Raf., Précis Découv. Somiol.: 41.

1814 (non Pourr. in Hist. & Mém. Acad. Roy. Sci.

Toulouse 3: 320. 1788) ≡ Helichrysum rupestre DC.,

Prod r. 6: 182. 1838, nom. illeg. [= Helichrysum pan-

ormitanum Guss.] – Neotype (designated by Gal-

bany-Casals & al. in Taxon 55: 492. 2006): Palermo,

in rupibus calcareis, V, Todaro 551 (PAL No. 8720;

isoneotypes: FI, K, P, PH No. 1029697).

Galbany-Casals & al. (2006c: 492) correctly state that

Rafinesque’s name is an illegitimate later homonym of

Gnaphalium rupestre Pourr. (a synonym of Helichrysum

stoechas (L.) Moench). They nevertheless accepted H.

rupestre DC., apparently overlooking the fact that Can-

dolle cited G. pendulum C. Presl in synonymy, the epi-

thet of which he should have adopted. H. rupestre DC.

is, therefore, illegitimate, even though still typified by

the type of G. rupestre Raf. (see the preceding comments

under G. cespitosum).

Helichrysum panormitanum is treated by Galbany-

Casals & al. (2006a, b) as a synonym of the widely defined

species they call H. rupestre (and shou ld instead have

named H. pendulum); pending further study of that com-

plex, we prefer to maintain the two entities, and several

non-Sicilian ones similarly merged by Galbany-Casals &

al. (2006a, b), as distinct if closely related species.

Helichrysum cespitosum var. longifolium Guss., Fl. Sicul.

Syn. 2: 468. 1844 [=Helichrysum stoechas subsp.

barrelieri (Ten.) Nyman] – Lectotype (designated

here): specimen bearing three labels: “Giugno, Pal-

agonia”, “Palagonia”, and “7c. Helichrysum caespi-

tosum c. longifolium Guss. syn. 2 p. 468, Junio, Julio

ђ, in collibus calcareis vel vulcanicis apricis.” [manu

Gussone], herb. siculum Gussonei (NAP). Mounted

together, on the same sheet, with a different specimen,

labelled “Tineo, Nº 4” by Gussone.

Not in Galbany-Casals & al. (2006a, c), not typified

by Galbany-Casals & al. (2006b).

Helichrysum errerae Tineo, Pl. Rar. Sicil. 2: 27. 1846

– Lectotype (designated here): “Helichrysum Errerae

Tin., Pantellaria” [manu Tineo], herb. siculum Gus-

sonei (NAP), superseding the neotype proposed by

Galbany-Casals & al. (2006c: 494).

Galbany-Casals & al. (l.c.) commented that no origi-

nal material of Tineo is extant at either PAL or FI. There

are, however, three specimens in Gussone’s Sicilian her-

barium (kept in NAP as a separate collection) that must

be considered for lectotypification purposes. They are in

all probability duplicates of the same gathering, made by

the physician Alfonso Errera on the island of Pantelleria

and sent by him to Tineo. The first, here designated as

lectotype, has a label in the handwriting of Vincenzo

Tineo, the author of the name. The second is labelled,

in Gussone’s handwriting, “Helichrysum Errerae Tin.,

Pantelleria a Zineddi dietro l’isola, Tin.”, which cor re-

sponds almost literally with the locality statement in

Tineo’s protologue. The third lacks an original label, but

has a label transcript in the hand of Grande and dated

1916, reading “Elichrysum Errerae Tin.!!, Loc. Pantel-

leria, legit Tineo!”.

Helichrysum nebrodense Heldr. in Ann. Accad. As-

piranti Naturalisti Napoli 1: 286. 1843 – Lectotype

(designated here): “Helichrysum nebrodense ? Nob.

inedit., in rupibus calcareis prope Isnello, 12 Jun.

184 0. Theod. de Heldreich” [manu Heldreich], herb.

siculum Gussonei (NAP), superseding the neotype

proposed by Galbany-Casals & al. (2006c: 496).

Galbany-Casals & al. (l.c.) include NAP among the

herbaria in which no original Heldreich material could

be traced. They were misinformed. Details on Held-

reich’s Sicilian journey in 1840–1841, which explain why

Gussone’s herbarium is a likely place for specimens then

collecte d to be fou nd, will be published separately by one

of us (P. Mazzola, in prep.).

Helichrysum panormitanum Gu ss., Fl. Sicul. Syn. 2: 467.

184 4 ≡ Helichrysum panormitanum var. angustifo-

lium Guss., Fl. Sicul. Syn. 2: 467. 1844 (homotypic

by type designation) – Lectotype (Galbany-Casals &

al., 2006b: 70, as “holotype”; again designated here):

“Gnaphalium panormitanum Nob., In rupibus cal-

careis prope Panormum, scala di Maseddu” [manu

Tineo], “Tineo” [manu Gussone], herb. siculum Gus-

sonei (NAP).

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Aghababyan & al. • Typification of Sicilian HelichrysumTA XO N 56 (4) • November 2007: 1285–1288

Galbany-Casals & al. (2006b: 70) cited the above

specimen as “holotype”. Gussone in the protologue rec-

ognised two named varieties in his new species, and al-

though from the context it is plausible to infer that he

considered the first of them, var. angustifolium, as typical,

he did not make this explicit. Whereas we agree that the

above “holotype” specimen is the appropriate choice, we

prefer to consider it as lectotype. Under the Saint Louis

Code it was debatable whether or not a “holotype” state-

ment, even if published after 2000, was still acceptable

as equivalent to effective lectotype designation (as we

believe it should). In the Vienna Code, an additional clari-

fication (Art. 9 Note 4) tilts the balance the other way. We

hope that a future Congress may restore the unconditional

precedence of Art. 9.8 over 9.21, so as not to encourage

authors to designate lectotypes which are probably holo-

types, or resort to the nonsensical statement “holotype,

designated here”.

Helichrysum panormitanum var. latifolium Guss., Fl.

Sicul. Syn. 2: 467. 1844 [= Helichrysum panormi-

tanum Guss.].

The lectotype designation by Galbany-Casals & al.

(2006c: 497) reads: “Bagheria a Capo Zafferano, 1916,

Grande s.n. (NAP)”. It seems curious at first sight that a

specimen collected in 1916 should be original material for

a name published in 1844. Nevertheless the designation

stands, but must be amended to read: “Elichrysum panor-

mitanum Tin. b. latifolium Guss.!! [scripsit] Grande, 1916”;

“Maggio, Bagheria a Capo Zafferano” [manu Gussone],

[Gussone], herb. siculum Gussonei (NAP).

Helichrysum pendulum var. compactum Guss., Fl. Sicul.

Syn. 2: 467. 1844 [= Helichrysum pendulum (C.

Presl) C. Presl] – Lectotype (designated here): speci-

men bearing two labels: “Luglio, Madonie”, and “6b.

Helichrysum pendulum b. compactum Supl. syn. 2

p. 467, Junio, Julio ђ, in rupibus calcareis montosis.”

[manu Gussone], herb. siculum Gussonei (NAP).

Not in Galbany-Casals & al. (2006a, b), not typified

by Galbany-Casals & al. (2006c).

Helichrysum porcarii Tineo ex Lojac. in Natural. Sicil.

2: 182. 1883 [= Helichrysum pendulum (C. Presl) C.

Presl] – Lectotype (designated here): “Helichrysum

porcari Tin., Agosto, Madonie, Salto della Botte,

Porcari ” [manu Porcari], herb. siculum Gussonei

(NAP).

Not in Galbany-Casals & al. (2006a–c).

Note: Described by Lojacono under the heading

“Species dubiae”. We consider this placement as an ex-

pression of taxonomic doubt that does not affect the valid

publication of the name. Lojacono’s locality, “Madonie

al Passo della Botte”, is the same as on our specimen.

Lojacono adds a concrete specimen citation, “Tin. ined.

in Herb. H. Reg. Bot. Pan.”, but no such specimen is

present nowadays in the historical herbarium in PAL.

We consider our lectotype specimen to be a duplicate of

the lost holotype.

Helichrysum siculum Jord. & Fourr., Brev. Pl. Nov. 2: 67.

1868 ≡ Helichrysum italicum subsp. siculum (Jord.

& Fourr.) Galbany & al. in Canad. J. Bot. 84: 1225.

2006 – Lectotype (designated by Galbany-Casals &

al., in Canad. J. Bot. 84: 1225. 2006: Sicily, Gussone

ex herb. Al. Jordan 187 (LY; isolectotype: MPU).

Helichrysum stramineum Guss., Fl. Sicul. Syn. 2: 467.

184 4 [= Helichrysum panormitanum Guss.] – Lec-

totype (designated by Galbany-Casals & al., in Taxon

55: 499. 2006): Sferracavallo, Tineo, [herb. siculum

Gussonei] (NAP).

The Sicilian herbarium of Gussone is kept in NAP as

a separate, closed collection.

Helichrysum wickstromii Tineo ex Lojac. in Natural. Sicil.

2: 182. 1883 [= Helichrysum pendulum (C. Presl)

C. Presl] – Lectotype (designated here): “Giugno 49,

Elichrysum Wickströmii Tin. ined., Pizzuta” [manu

Tineo], herb. siculum Gussonei (NAP).

Not in Galbany-Casals & al. (2006a, c), treated as

an unpublished label name by Galbany-Casals & al.

(2006b).

As is the case with Helichrysum porcarii (q.v.), H.

wickstromii was placed under “Species dubiae” by Lo-

jacono. It was described from “Pizzuta” with an added

specimen citation “Tin. ined. l.c.” [i.e., Herb. H. Reg. Bot.

Pan.], but again, no such specimen is present nowad ays in

the historical herbarium in PAL. The lectotype designated

here is most likely a duplicate of the lost holotype, as

is a second sheet, also in Gussone’s Sicilian herbarium,

which lacks an original label but bears a label copy signed

“Grande, 1916”, with the text “Helichrysum pendulum

Presl!, loc. Pizzuta, legit Tineo”.

ACKNOWLEDGEMENTS

We thank Dr Galbany-Casals for kindly providing digital

photographs of several original specimens, including those she

de signa ted as lect o- or n eot ypes . Dr O. Šíd a an d Dr J. Štěpánek

went out of their way to search for Presl types in their respective

herbaria in Prague. Dr Paolo Caputo monitored and assisted our

search of the Sicilian herbarium of Gussone at Naples. Professor

John McNeill devoted his experience and energy to the discus-

sion of tricky nomenclatural questions and skilfully revised the

present manuscript.

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Full-text available

Jul 2008

Some taxa of Sicilian Helichrysum related to sect. Stoeachadina have an uncertain systematic position; moreover, they are rare and endangered. In order to preserve these taxa in a genebank, a strategy to keep their genetic variation as high as possible is required. The high informative content of AFLP markers was used to assess genetic variations within and between populations and their relationships. The results show that all populations are genetically similar, even though there are two distinct groups: the first is the population referred to H. pendulum, and the second is represented by other populations belonging to several different specific and intraspecific taxa. The H. pendulum population is homogeneous, and characterised by the lowest genetic variation among all the populations considered. The second group shows a greater variation as evidenced by the presence of three subgroups: (i) populations referred to H. panormitanum var. panormitanum and H. panormitanum var. stramineum, (ii) populations referred to H. errerae var. errerae and H. errerae var. messerii, (iii) a population referred to H. hyblaeum, which, despite morphological and chorological differences, is similar to the other populations of the second group from a genetic stand point. Each population of the above-mentioned taxa has local fragment markers whose frequency is useful to determine how many plants should be sampled in order to collect the seeds necessary to preserve the population's genetic diversity in a genebank.

Names of Italian vascular plants published by Michele Lojacono Pojero

Article

Full-text available

Dec 2014

Michele Lojacono Pojero (1853-1919) is among the most prolific Italian authors of plant names of his era. A list is presented of the names of 581 new Italian (almost invariably Sicilian) vas-cular plant taxa he published (303 species, 272 varieties, 6 formae), with reference to existing lectotype designations.

Polyploidy and new chromosome counts in Helichrysum (Asteraceae, Gnaphalieae)

Article

Nov 2008
BOT J LINN SOC

Mitotic chromosome numbers are reported for 31 populations representing 28 taxa of Helichrysum. Twelve are new and eight others provide confirmation of a unique previous reference. A new chromosome number, 2n = 42, is reported for H. odoratissimum. Polyploidy is confirmed as the most significant evolutionary trend in chromosome number within the genus. Chromosome data agree with trends observed in phylogenetic studies: a South African and diploid origin of the genus, followed by a radiation and diversification in southern Africa and several migrations towards the north of the African continent, the Mediterranean basin and Asia. Expansion and diversification of the genus have been accompanied by several genome duplications which have led to the acquisition of the tetraploid, hexaploid and octoploid levels, all in several independent events. Both autopolyploidy and allopolyploidy are suggested as probable speciation agents within the genus. © 2008 The Linnean Society of London, Botanical Journal of the Linnean Society, 2008, 158, 511–521.

T E X Lecture Notes in Computer Science

Article

Jul 1993

. This paragraph shall summarize the contents of the paper in short terms. 1 Fixed-Period Problems: The Sublinear Case With this chapter, the preliminaries are over, and we begin the search for periodic solutions : : : 1.1 Autonomous Systems In this section we will consider the case when the Hamiltonian H(x) : : : The General Case: Nontriviality. We assume that H is (A1 ; B1 )-subquadratic at infinity, for some constant : : : Notes and Comments. The first results on subharmonics were : : : Proposition4. Assume H 0 (0) = 0 and H(0) = 0. Set : : : Proof of proposition. Condition (8) means that, for every ffi 0 ? ffi , there is some " ? 0 such that : : : ut Example 1 (External forcing). Consider the system : : : Corollary 5. Assume H is C 2 and (a 1 ; b 1 )-subquadratic at infinity. Let : : : Lemma 6. Assume that H is C 2 on IR 2n nf0g and that H 00 (x) is : : : Theorem7 (Ghoussoub-Preiss). Let X be a Banach Space and Phi : X ! IR : : : Definition8. We shall say that ...

International Code of Botanical Nomenclature (Vienna Code)

Book

Full-text available

Oct 2006

available online: http://www.iapt-taxon.org/historic/2006.htm

Flora der Insel Kythera, gleichzeitig Beginn einer nomenklatorischen Überprüfung der griechischen Gefäßpflanzenarten

Article

Jan 1967

Typification of Names in Gnaphalium L. and Helichrysum Mill. (Asteraceae), and Some Taxonomic Notes

Article

May 2006

As a part of a research study on the systematics of the genus Helichrysum in the Mediterranean and Irano-turanian area, 53 new typifications of names in Gnaphalium and Helichrysum belonging to taxa in Helichrysum sect. Stoechadina, H. sect. Helichrysum and H. sect. Virginea are presented, along with some additional comments on the nomenclature of the names involved and on the taxonomy of the taxa to which they apply. Twenty-three previously published typifications of names of taxa belonging to these sections are also listed and, in some cases, commented upon. The accepted taxa of Helichrysum for which we propose typification of accepted names or synonyms, or upon which we make comments, are, alphabetically: H. angustifolium (Lam.) DC., H. arenarium (L.) Moench, H. arenarium subsp. aucheri (Boiss.) P. H. Davis & Kupicha, H. armenium DC., H. crassifolium (L.) D. Don, H. doerfleri Rech. f., H. errerae Tineo, H. graveolens (M. Bieb.) Sweet, H. italicum (Roth) G. Don, H. italicum subsp. microphyllum (Willd.) Nyman, H. orientale (L.) Gaertn., H. pallasii (Spreng.) Ledeb., H. pamphylicum P. H. Davis & Kupicha, H. picardii Boiss., H. plicatum DC., H. pygmaeum Post, H. rupestre DC., H. sanguineum (L.) Kostel., H. sibthorpii Rouy, H. stoechas (L.) Moench, H. taenari Rothm. and H. virgineum DC.

Conspectus of Helichrysum Mill. sect. Stoechadina (DC.) Gren., Godr. (Asteraceae, Gnaphalieae)

Article

Jan 2006

Es presenta un conspectus de Helichrysum sect. Stoechadina (DC.) Gren. and Godr. Es reconeixen vuit espècies i cinc subespècies. Per cada taxó acceptat es relacionen tots el sinònims trobats a la bibliografia, a partir de l’estudi dels protòlegs, el material tipus i els nombrosos exemplars d’herbari que comprenen tota l’àrea de distribució i la variabilitat de cada espècie. S’aporten notes sobre l’àrea de distribució. Quatre noms han estat lectotipificats y es presenten també referencies sobre lectotipificacions prèvies.

Conspectus of 'Helichrysum' Mill. sect. Stoechadina (DC.) Gren. & Godr. (Asteraceae, Gnaphalieae)

Article

Jan 2007

Es presenta un conspectus de Helichrysum sect. Stoechadina (DC.) Gren. & Godr. Es reconeixen vuit espècies i cinc subespècies. Per cada taxó acceptat es relacionen tots el sinònims trobats a la bibliografia, a partir de l'estudi dels protòlegs, el material tipus i els nombrosos exemplars d'herbari que comprenen tota l'àrea de distribució i la variabilitat de cada espècie. S'aporten notes sobre l'àrea de distribució. Quatre noms han estat lectotipificats y es presenten també referencies sobre lectotipificacions prèvies. A conspectus of Helichrysum sect. Stoechadina (DC.) Gren. & Godr. is provided. Eight species and five subspecies are recognized. For each accepted taxa all synonyms found in the literature are listed, based on the study of the protologues, the type material, and numerous herbarium specimens comprising the whole distribution area and the variability of each species. Notes on the distribution area are provided. Four names are lectotypified, and references to previous lecotypifications are also presented.

@BULLET Typification of Sicilian Helichrysum LITERATURE CITED

Aghababyan

Aghababyan & al. @BULLET Typification of Sicilian Helichrysum LITERATURE CITED

Jan 2006
489-501

M Galbany-Casals
Sáez
Ll
C Benedí
C E Jarvis

Galbany-Casals, M., Sáez, Ll., Benedí, C. & Jarvis, C.E. 2006c. Typification of names in Gnaphalium L. and Heli chrysum Mill. (Asteraceae), and some taxonomic notes. Taxon 55: 489–501.

Conspectus of Helichrysum sect

Jan 2006
59-81

M Galbany-Casals
Ll Sáez
C Benedí

Galbany-Casals, M., Sáez, Ll. & Benedí, C. 2006b. Conspectus of Helichrysum sect. Stoechadina (DC.) Gren. & Godr. (Asteraceae, Gnaphalieae). Orsis 21: 59-81.

Genera, species et synonyma Candolleana alphabetico ordine disposita, seu index generalis et specialis ad A. P. Decandolle Prodromum systematis naturalis regni vegetabilis, pars 2 (continens tomos operis Candolleani quintum, sextum et sectionem priorem septimi)

Jan 1840

H W Buek

Buek, H.W. 1840. Genera, species et synonyma Candolleana alphabetico ordine disposita, seu index generalis et specialis ad A. P. Decandolle Prodromum systematis naturalis regni vegetabilis, pars 2 (continens tomos operis Candolleani quintum, sextum et sectionem priorem septimi). Nauck, Berlin.

Typification of Sicilian Helichrysum LITERATURE CITED

Aghababyan

Aghababyan & al. • Typification of Sicilian Helichrysum LITERATURE CITED

Typification of Sicilian Helichrysum (Compositae) Revisited

Abstract

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