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Eggli U. & Leuenberger B. E. — Type specimens of Cactaceae names in the Berlin Herbarium (B). Willdenowia

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Abstract

A short history of the Cactaceae holdings of the Berlin herbarium and their fate is provided. While a considerable number of historical spirit specimens have escaped the destruction of the herbarium during World War II, the pre-1943 collection of pressed specimens was lost almost entirely. More than half of the present Cactaceae holdings of B were made on the base of cultivated material. The importance of these "Garden Herbarium" specimens is stressed, especially for succulent plants, where much of the type material, even of recently described taxa, is of insufficient quality or even fragmentary. A complete survey of all extant Cactaceae specimens at B (including the material recently transferred from the former KOELN herbarium) has resulted in a detailed and commented list of type specimens and specimens derived from cultivated descendants of type collections. Nomenclatural and typification problems encountered during this compilation are discussed. A list of photographs preserved at B depicting type material located in other herbaria, is also given. Lectotypes are designated for the names Cereus lamprospermus, C. rhodoleucanthus, C. stenogonus, C. xanthocarpus, Echinocactus umadeave, Echinopsis rhodotricha, Pilocereus setosus, Rebutia steinbachii and Turbinicarpus pseudomacrochele var. lausseri.
De Herbario Berolinensi Notulae No. 48
URS EGGLI & BEAT ERNST LEUENBERGER
Type specimens of Cactaceae names in the Berlin Herbarium (B)
Abstract
Eggli, U. & Leuenberger, B. E.: Type specimens of Cactaceae names in the Berlin Herbarium (B) [De
herbario berolinensi notulae 48]. Willdenowia 38: 213-280. ISSN 0511-9618; © 2008 BGBM
Berlin-Dahlem.
doi:10.3372/wi.38.38116 (available via http://dx.doi.org/)
A short history of the Cactaceae holdings of the Berlin herbarium and their fate is provided. While a
considerable number of historical spirit specimens have escaped the destruction of the herbarium dur-
ing World War II, the pre-1943 collection of pressed specimens was lost almost entirely. More than
half of the present Cactaceae holdings of B were made on the base of cultivated material. The impor-
tance of these “Garden Herbarium” specimens is stressed, especially for succulent plants, where
much of the type material, even of recently described taxa, is of insufficient quality or even fragmen-
tary. A complete survey of all extant Cactaceae specimens at B (including the material recently trans-
ferred from the former KOELN herbarium) has resulted in a detailed and commented list of type
specimens and specimens derived from cultivated descendants of type collections. Nomenclatural
and typification problems encountered during this compilation are discussed. A list of photographs
preserved at B depicting type material located in other herbaria, is also given. Lectotypes are
designated for the names Cer eus lampr os per mus , C. rhodoleucanthus , C. stenogonus, C. xanthocar
-
pus, Echinocactus umadeave, Echinopsis rhodotr i cha, Pilocer eus setosus, Rebutia steinbachii and
Turbinicarpus pseudomacrochele var. lausseri.
Additional key words: cacti, succulents, typification, lectotype, botanical nomenclature, collection
history.
1. Introduction
The importance of type specimens as permanent anchors for the application of botanical names is
universally recognized today, and the numerous initiatives of herbaria to make type specimen in
-
formation available on the Internet underlines the importance of types. Mechanisms for selecting
lecto- or neotypes when the holotype has been lost are an important part of the ICBN (McNeill &
al. 2006). But before a replacement specimen can be designated, the existence (or loss) of the
holotype must be established, and this information is often not easily available, especially for old
names. Typification is only mandatory since 1958 (ICBN Art. 37.1, McNeill & al. 2006), and
names published before this date are often not typified or with insufficient type information.
Willdenowia 38 – 2008 213
Since succulents in general and cacti in particular are generally difficult to handle for the her
-
barium, types are missing for many names in this group. While some authors straight away de
-
clared the preparation of herbarium specimens of cacti as hardly possible (e.g., Backeberg 1962:
3916) or attributed questionable usefulness to type specimens (Ritter 1979: 15), the numerous
existing and often very well prepared and preserved old and new specimens of many collectors
prove the contrary. A particularly practical and successful method to prepare herbarium speci
-
mens of succulents has been described in more detail by Eggli & Leuenberger (1996).
Any typification study necessarily starts by establishing the existence and whereabouts of the
holotype and possible isotypes. This information is not always easily obtained, but (published)
type specimen lists are an important source of information. For cacti, such lists have been com
-
piled, e.g., by Eggli & al. (1996) for taxa described by F. Ritter, Eggli & Leuenberger (2005) for
taxa described by M. Cárdenas, and Eggli (1987) for material from Swiss herbaria.
Berlin is undoubtedly one of the most important institutions regarding the study of succu
-
lents, in particular of cacti, and important taxonomists such as K. Schumann, M. Gürke, F.
Vaupel and E. Werdermann devoted time and interest to the study of cacti. Leuenberger
(1978a-b, 1979) produced a first inventory of type specimens of cacti at the Berlin herbarium, al
-
though limited to the spirit collection. Since much type material that would be expected at B has
been lost (see below), we undertook a project to compile an inventory of all type material pres
-
ently available at B, including a new evaluation of the spirit collections.
2. History of the Berlin Herbarium and its Cactaceae holdings
A detailed history of the Berlin herbarium collections was given by Urban (1916) and Hiepko
(1987). The loss of the greater part of the herbarium collections, library and archives were re-
ported by Pilger (1953) and Sleumer (1949). Hiepko (1987) provided a selective list of important
collections acquired between 1914 and 1986. In this list, a number of collectors known to have
collected Cactaceae are mentioned, e.g., Ekman, Ule, Werdermann, but nothing is known about
the total number of Cactaceae specimens. Hiepko (1987) also provided a summary of collections
still extant from the time before 1943 (including the spirit collection). Traditionally, no acces-
sion numbers were used for the specimens at B, but the complete holdings of the spirit collection
were bar-coded in 2004 and 2005.
2.1. Sheet specimens in the General Herbarium
The herbarium material of Cactaceae was lost in 1943, except for the Willdenow material (Pilger
1953; Hiepko 1987; Leuenberger 2004). The former richness of the Berlin herbarium in neotropi
-
cal type specimens is well documented through the photograph collection of Berlin types, since
2006 available online at http://emuweb.fieldmuseum.org/botany/search_berlin.php (accessed
15.11.2006). Cactaceae, however, are completely lacking among the over 15 000 images made by
Macbride in Berlin between 1929 and 1933 of material collected in Central and South America
from 1778 through May 1930, representing 154 plant families (Grimé & Plowman 1986).
Scattered information on Cactaceae specimens of various collectors deposited at B (e.g.,
Anisits, Ekman, Fiebrig, Glaziou, Grosse, Kurtz, Reiche, Söhrens, Ule, Werdermann) can be
found (among others) in publications of Schumann (1897-98: x, 1899: 133, 1903), Vaupel (1916,
1925, 1926) and Werdermann (1927, 1929, 1931a, 1949).
The tragic fate of the succulent collections was described by Werdermann (1949), but this re
-
port was not fully accurate as far as the spirit material is concerned (Leuenberger 1978, 1979).
Present holdings of Cactaceae in the herbarium were all accessioned from 1944 onwards and
include very few older specimens, e.g., saved duplicates (Werdermann), later donations from
other herbaria and also photographs of some older materials. The presence of a limited number of
Werdermann specimens is due to the fact that the first set was destroyed in 1943, but replaced by
an incomplete set of undistributed duplicates (according to a note in Werdermann’s hand dated
1944). This explains the presence of some (though very few) isotype specimens.
214 Eggli & Leuenberger: Type specimens of Cactaceae names in the Berlin Herbarium
Most specimens in the general herbarium are not yet bar-coded and the sheets are without ac
-
cession number. Bar-coding was started in 2002 with new material and material selected for digi
-
tal imaging.
2.2. Spirit material
The major part of the spirit material was salvaged (Sleumer 1949). It was rearranged and comple
-
mented with a card index in 1961 by F. Butzin (Potztal 1962: 42, 1965: 104). Apart from some
jars sealed with unsuitable cork and paraffin wax in the 1930ies, some with material dried out
and decayed, the Cactaceae collection (c. 600 jars) mostly remained in good condition (Leu
-
enberger 1978, 1979). All specimens were bar-coded in 2005.
The older jars have up to three additional older printed labels with numbers added by hand.
The uppermost is the genus number following Dalla Torre & Harms (1900-07). Another label,
usually at the bottom of the jar, contains the taxon number assigned in Schumann (1897-98,
1903). These numbers were apparently added to the jars after publication of that work.
A third label bears an old catalogue number, but the catalogue was lost in 1943 (Potztal 1962:
42). This number is crossed out and was replaced by a number in red, representing a sequential
jar number (for Cactaceae), the same as used in the card index established by F. Butzin in 1961.
2.3. Boxed material
From a note by Anonymus (1893) and Hirscht (1896) on the existence of such material, includ-
ing a further request of Schumann to members of the cactus society to send dead plants for pres-
ervation in the botanical museum, it can be assumed that a considerable collection existed.
Werdermann (1937: 163) mentioned it as “mummy collection” and later noted that, during his
curatorship (before 1943), dead plants from the garden were critically checked for possibly ap-
propriate and useful parts to be preserved in the herbarium (Werdermann 1949: 36). All this old
boxed material was lost in 1943.
The current holdings were mostly accessioned from c. 1980 onwards. They derive to a great
extent from cultivation in the Botanic Garden Berlin-Dahlem (e.g., from the extensive Parodia
collection of the private collector Hummel, live plants acquired in 1987), but also from other
sources (notably P. J. Braun collections incorporated in 1990).
Most specimens are not yet bar-coded and without accession number. The specimens from
cultivation at B have “Garden Herbarium” numbers (see below).
2.4. The Garden H erbarium
Documentation of the living Cactaceae collection by herbarium specimens and photographs was
initiated by the second author in 1977 during the taxonomic revision of Pereskia, in order to
compare herbarium collections side by side with live material and dried material of the same ac
-
cessions. This was gradually extended to the whole family Cactaceae.
The Garden Herbarium, with precursors dating back to 1952, was started in the current form
with consecutive accession numbers in 1974. The number is preceded by the name of the techni
-
cal assistant preparing the specimens (from 1974 to 1989, Schwerdtfeger 1-25999; from October
1989 until July 2008, Cubr 26000-45595). A few specimens have been bar-coded and have this
additional number as well.
The garden herbarium was originally stored entirely separate from the general herbarium as a
reference collection. For safer storage, it has been kept since 1993 within the general herbarium,
as parallel collection to each phanerogam family, facilitating quick access to the collection of
cultivated plants alone, as well as side by side to the general collection.
Importance of material from cultivation is enhanced by the fact that holotypes of succulent
plants often consist of very limited material, sometimes badly prepared, or only very little mate
-
rial collected, e.g., only areole and flower (Ritter). Thus, cultivated descendants of original intro
-
ductions are of special value, both as specimens (pressed, spirit, boxed), and (alone or additionally)
Willdenowia 38 – 2008 215
as photographs. The large number of specimens prepared on this base is a speciality of the her
-
baria B and ZSS, and such material can be used to “back up” original types. The depreciating
view taken by many concerning cultivated “types” is unwarranted.
Another argument in favour of such material is the fact that cacti are important in cultivation,
and much material was described on the base of cultivated material. “Cultitypes” are therefore
especially important as base for comparison and to document the plasticity of characters in culti
-
vation.
2.5. Material transferred from KOELN
In January 2005 a substantial part the Herbarium of the Arbeitsgruppe Geobotanik und Phyto
-
taxonomie, Botanisches Institut, Universität Köln (KOELN, see Follmann 1998), i.e. primarily
consisting of the lichen collection of G. Follmann, was transferred to Berlin-Dahlem (B) on the
initiative of G. Follmann (see Holmgren & Holmgren 1998-). Simultaneously, another important
collection was moved to Berlin: type specimens (holotypes and isotypes) of 52 names of
Cactaceae and a few additional non-type specimens deposited at KOELN from c. 1977 onwards,
many connected with the work of Lothar Diers. Names based on collections deposited at KOELN
were published by W.-R. Abraham, P. J. Braun, L. Diers, E. Esteves Pereira, G. Heimen and R.
Paul, all in cactophile journals in Germany, The Netherlands and the USA.
These names were originally published with varying indications of depository, e.g., herbar
-
ium of the “Pädagogische Hochschule Rheinland, Abt. Köln” (Gymnocalycium ambatoense,
Piltz 1980), Herbarium of the Succulentarium of the University of Cologne” (Discocactus lin-
daianus, Diers & Esteves 1981), or “Herbario Universitatis Colonia (KOELN)” (sic!, Coleoce-
phalocereus diersianus, Braun & Esteves 1988).
At the time of the transfer from KOELN to B in 2005, the material was kept in boxes labelled
outside as “Collatio Lothar Diers”. Actually it is not a classical, mounted herbarium but each
“specimen” is composed of one to six loosely kept entities: plastic bags with dried bodies of
cacti, entire plants, fragments, fruits and flowers preserved in alcohol in a variety of glass jars or
plastic bottles, paper bags containing fruits or seeds, envelopes containing photographs or (very
few) notes, and reprints or entire journal issues containing the protologues, in a number of cases
with handwritten complementary footnotes (presumably by L. Diers). Each entity is labelled with
only minimal data, usually with name, type designation, collector and number (both normally as
abbreviated field number only, e.g., “HB 80”, for “Heimen & P. J. Braun 80”).
Sorting of the material by name and collection number and the compilation of a detailed
working list of all entities, and comparison with data from IPNI (2004-) and the database of the
“Repertorium Plantarum Succulentarum” maintained by U. Eggli (ZSS) revealed several prob
-
lems:
The type material for a name, consisting of a single collection number, is kept in up to seven
separate entities with differing preservation methods, resulting in problems for storing, bar-cod
-
ing and databasing.
For most type numbers, only part of the material (one to three entities) is marked as holotype
or isotype. Flower samples and seed samples are often not marked. They bear the same collection
number and may, or may not, represent a true part of the type material. Moreover, the samples
are not dated, and their status as type or not-type is sometimes impossible to ascertain with abso
-
lute accuracy.
The samples usually lack any indication of country and collecting locality. With very few ex
-
ceptions, this information is only found in the usually very detailed protologues. Neither collec
-
tion dates nor preservation dates are indicated. In the protologues, details given in the Latin part,
as compared with the German, English or Dutch text, may differ. For many of these names, the
protologue does not give detailed locality data but states that this information is stored together
with the holotype for conservation reasons. Indeed, for many names, a copy of the protologue
with the addition (by hand) of more detailed locality data, was present in the material transferred
from KOELN.
216 Eggli & Leuenberger: Type specimens of Cactaceae names in the Berlin Herbarium
–SomeCactaceae material originally published for KOELN was missing when the collection
was transferred to Berlin in January 2005. A total of 23 specimens, including some taxa de
-
scribed from 1980 onwards, as well as taxa recently published by Diers & al. were lacking, e.g.,
Hildewintera colademononis, Parodia hegeri, Sulcorebutia roberto-vasquezii and S. vargasii. A
search for the lacking specimens through G. Follmann led to a communication from L. Diers (5
December 2005) and in February 2006, to a shipment from L. Diers, Bad Neuenahr, of 11 jars
containing seven of the missing holotypes and four isotypes of species described between 1980
and November 2005 (by Piltz and by Diers). It has to be assumed that these specimens were not
deposited in KOELN despite having been cited for this herbarium. Another specimen, the
holotype of Sulcorebutia vargasii var. viridissima Diers & Krahn, was added by Diers without
prior announcement.
3. Nomenclatural considerations
3.1. Problems associated with garden material and “clonotypes”
For the vast majority of plant taxa, the type material consists of one (or more, then termed
isotypes) field-collected and permanently preserved (usually pressed and dried) specimen(s).
Establishing whether a given herbarium specimen is the type (holotype, or isotype) of a name
should not produce undue problems.
In the case of succulent plants, the situation differs to some degree. More often than not, type
specimens are prepared in a botanical collection from cultivated material. When such type mate-
rial is clearly labelled as to its status (holotype or isotype), no problems arise, but such clear la-
belling is often absent. Further, the availability of a cultivated “type collection” makes it
possible to preserve subsequently additional material. Moreover, the material can be propagated
and distributed to other gardens, or even made available to the horticultural trade. Especially in
the case of cacti, type material has also been derived from collections made by commercial or
semi-commercial collectors, as well as by devoted plant collectors.
As long as all material preserved on the base of living collections is clearly labelled,
especially with the preservation date, there are no unsurmountable problems (apart perhaps those
covered by ICBN Art. 8.2, Ex. 1, explaining one view of what consists “a gathering”, McNeill &
al. 2006). Specimens preserved subsequent to the publication of the protologue are usually and
loosely termed “clonotypes”, a term which is not sanctioned, however, by the ICBN (McNeill &
al. 2006).
For the purpose of the present inventory, we continue the concept already introduced for the
inventory of the Ritter types (Eggli & al. 1996: 41), i.e. we use the term “type collection” to refer
to specimens preserved under the type collection number. Any material that cannot be clearly
identified as holotype or isotype is simply assigned “type collection” status.
For some names, an additional complication has to be accounted for: usually, a collection
number refers to a single collection event at a single clearly delimited locality. Some collectors,
however, used their collection numbers in a different way. Eggli & al. (1996) elucidated the
numbering convention adopted by Friedrich Ritter, whose collection numbers are essentially
taxon numbers, and up to more than a dozen different localities might be covered by a single col
-
lection number. To establish which of the several collections under a given number is the “real”
type is often difficult but crucial. When material so numbered enters into cultivation, there is
usually no direct link back to the locality information, and it is thus impossible to pin-point its
possible type status. For such cases, we use the term “type number”.
It could be argued that it is not worth bothering with such corollary material, i.e. any “clo
-
notypes” and “near-clonotypes” from cultivated material. This would, however, deprive us of
much important information. More often than not, the preserved original material is confined to
the holotype specimen, and this is unfortunately far too often of fragmentary nature, ill-pre
-
served, or perhaps even missing. “Clonotypes” and “near-clonotypes” are then the only possibly
original material to establish the characteristics of a taxon and are thus of special importance,
Willdenowia 38 – 2008 217
even though they are not types in the strict sense, but represent “near-paratypes” in the sense of
the ICBN (McNeill & al. 2006). They are not strict paratypes (Art. 9.5, “a paratype is a specimen
cited in the protologue that is neither the holotype nor an isotype, nor one of the syntypes”) be
-
cause they are not literally cited in the protologues. Because of the importance of these speci
-
mens, the complete “near-type” holdings of the Berlin Garden Herbarium have been included
into our inventory.
3.2. Problems associated with ‘synonymous’ type numbers
While comparing typification information in protologues with information associated with speci
-
mens at B labelled as types, or specimens of suspected type status, problems became apparent
when different collections are cited as holotype and isotype(s). An example is Arrojadoa beateae
P. J. Braun & Esteves, where the holotype is cited as “Esteves Pereira (E) 261” and isotypes (for
B and ZSS) as “Braun (Br) 830”, with the added statement that Braun 830 was collected at the
type locality simultaneously with the holotype.
Another example is Coleocephalocereus braunii: The holotype is “Horst, Braun & Esteves
4” and the collection “Braun 470” is given as isotype, again with an additional statement that this
(and two further topotype collections) were made simultaneously at the type locality.
Such disparate typification details are problematical in several respects, especially also
nomenclaturally: at first glance, it could be argued that the enumeration of more than one collec
-
tion as holo- and isotypes is an infringement of ICBN Art. 9 and 37 (McNeill 2006) and that the
collections are to be treated as syntypes, which would invalidate the names in question. Fortu-
nately, Art. 9.8 allows for a different interpretation, and the use of “isotype” for collections other
than the collection from which the holotype was prepared is to be regarded as an “error to be cor-
rected”. In the cases of Arrojadoa beateae and Coleocephalocereus braunii, the “isotype” speci-
mens Braun 830 and, respectively, Braun 470, are thus simply paratypes. In the literature, some
of these disparate typifications have been accepted at face value, e.g., by Taylor & Zappi (2004)
for Coleocephalocereus braunii, or produced erroneous typification statements, e.g., Eggli &
Taylor (1986) for the same taxon.
4. Notes on suspected types for selected names
For a number of names, typification poses special problems. These cases are shortly discussed
here with special emphasis on types or possible types:
Cereus coryne
There is a spirit specimen (B 810006729) with the following notes on the associated index card:
“eingeordnet Jan. 1900” and “grosses Scheitelstück, wahrscheinlich Originalmaterial”, and “vgl.
MfK 3: 176-177, 1893: redaktionelle Anmerkung von K. Schumann, ‘Pflanze im Hort. Bot.
Berol. Ist vielleicht Original’”. This specimen could indeed represent original material that sur
-
vived in cultivation at B and was prepared in 1900, but there is no unambiguous and direct link to
material from the time of the protologue and the specimen cannot safely be considered as original
material.
Cereus damazioi
There is a spirit specimen (B 810006730), consisting of a flower prepared from cultivated mate
-
rial from Rettig, preserved 12.6.1920, i.e. posterior to the publication of the illegitimate (Art.
52.1) name. It could probably have been derived from cultivated descendants of the original type
material, but there is no positive clear evidence for this and the specimen is not considered further
as original material.
Cereus hassleri
This name is based on two syntypes, Anisits Cact. 13 and Hassler 1716. Berlin has a very com
-
plete spirit specimen of the Anisits syntype, but no material of Hassler 1716. No attempt has been
218 Eggli & Leuenberger: Type specimens of Cactaceae names in the Berlin Herbarium
made to locate specimens of this second syntype but it should be noted that the specimen of this
number at NY is not Cereus hassleri, but has been annotated as Selenicereus setaceus by N. P.
Taylor in 1992 (NY virtual herbarium at http://sciweb.nybg. org/science2/hcol/vasc/index.asp,
accessed Nov. 2006). Lectotypification is deferred until other specimens of this syntype can be
located and compared with the protologue.
Cereus megalanthus
There is a spirit specimen (B 810006764), consisting of a stem piece with spines, annotated as
“Cereus megalantha [sic] K. Sch. Amazonas leg. Ule”. The connection with the type is weak, es
-
pecially because no locality data are given and the flowers noted in the protologue are absent. The
material is best not considered as original material.
Cereus peruvianus var. reclinatus
The type collection of this name is a cultivated accession (number 15/49) in the Huntington Bo
-
tanical Gardens, California, USA. There are no photographs of this accession among the Wer
-
dermann photographs of Cereus preserved at B, but B has photographs of two herbarium
specimens in the Huntington herbarium (HNT), namely HNT 1458 (garden acc. no. 5267,
“Type”, material pressed ex cult. 23.7.1973) and HNT 7342 (garden acc. no. 5267): “Type”, ma
-
terial pressed ex cult. 11.7.1986. It is not established whether this Huntington garden accession
(no. 5267) has any connection with the number of the cultivated type collection (15/49) as given
in the protologue of the name.
Cereus pseudocaesius
This name was published by Werdermann as replacement for Cereus caesius Salm-Dyck (non C.
caesius Salm-Dyck ex Pfeiffer). There are photographs at B of both accession numbers cited by
Werdermann. Since Werdermann’s publication of the name C. pseudocaesius cannot be regarded
as containing a valid typification of the replaced name, they are without importance for our list of
type specimens.
Cereus sclerocarpus
Cereus sclerocarpus is based on 3 syntypes from the Galápagos Islands, Snodgrass & Heller 933
and 934 from P. Christopher, Albemarle Island, and Snodgrass & Heller 935 from Black Bight on
the same island, and all of them are credited to “hb. Berl. & hb. Gr.”. Berlin has a spirit specimen
(B 810006791), consisting of flower fragments, and the original label has the collection number
as “Heller 934, 935”. It is impossible to assign the material to any of the two syntype numbers,
nor to know whether the specimen is indeed a mixture. Hb. Gr” has not yet been identified with a
modern herbarium and nothing is therefore known about the possible whereabouts of the remain
-
ing specimens.
Cereus schmollii
There is a spirit specimen at B (B810006790, dried up), the original label of which carries the in
-
formation “Hahn-Schmoll (cult.) (von Schmoll eingeführt)” / “hat Weingart untersucht”, dated
28.4.1931. It is thus prior to the protologue, but cannot be considered as holotype or isotype since
the protologue does not mention material from Hahn, but refers exclusively to material ex Hort.
Knebel (1929) and ex Hort. Rettig (1931).
Echinopsis korethroides
The type is Stümer s.n. and B has a sheet with a photograph (negative + print by Werdermann, not
mentioned by Leuenberger (1997), labelled as Eps. Lob.? Korethroides Werd. W7”, “Typ”
crossed out, “Fb 896”, cult. BG Berlin. The meaning of the numbers are unclear; “Fb 896” could
probably refer to a colour photograph (“Farbbild”). The material is certainly authentic, but there
are serious doubts whether indeed the type is illustrated.
Willdenowia 38 – 2008 219
Lobivia schreiteri
Leuenberger (1978, 1979) records an isotype specimen as Schreiter s.n., but this is not the type
collection (type = Castellanos 29/41). The specimen is annotated as “mis. R. Schreiter”.
Opuntia lagopus
The type for this name is given as Weberbauer s.n. in the protologue, but the only specimen extant
in the spirit collection at B (where the Weberbauer types of other names are located) has the num
-
ber Weberbauer 1355. Crook & al. (2003: 89) selected a neotype for the name (Webster 26, K) be
-
cause they believed that the original material at B was destroyed but carry on to say that it could
serve as epitype if original material should be discovered subsequently. Even though there is no
locality associated with Weberbauer 1355 on the label, there is little doubt that this specimen is
the holotype of the name. The name is shown as “Opuntia cfr. lagopus” on the original label, and
this might indicate that the specimen is not the type collection. However, we cannot positively es
-
tablish that the label is indeed original, or whether it was produced slightly later than the
protologue, and the “cfr.” (“confer”) was added because the incongruence of the type number
with the protologue. The material of the specimen is without any doubts representing O. lagopus,
and we are strongly inclined to accept this specimens as the holotype of the name. This makes the
neotypification by Crook & al. (l.c.) superfluous.
Opuntia pachypus
The case is similar to Opuntia lagopus, also based on an unnumbered Weberbauer collection.
Again, there is only a single specimen available in the spirit collection at B, which is moreover an-
notated as “Typus” on the original label. Iliff (in Cactaceae Syst. Init. 9: 5. 2000) unnecessarily
designated the plate in the protologue as lectotype, being unaware of the presence of the holotype
at B. The lectotypification made by Iliff (2000) is thus unnecessary.
Trichocereus campos-portoi
Leuenberger (1979) indicates a type for this name in the inventory of the surviving spirit material
at B. This specimen (B 810007164, plus photo in the general herbarium) consists of flower and
fruit and is labelled Trichocer. campos-portoi Werd.” and “Typ”. The type of the name is ex-
plicitely stated to be Werdermann 3994 in the protologue, with further material reported as Cam
-
pos-Porto s.n. The spirit specimen in question is most probably the Campos-Porto collection and
thus actually represents a paratype.
5. Cactaceae types at B
5.1. Introduction
The following list is a commented list of all recognized types and recognized authentic material
in the Berlin herbarium. It is a positive list, and for taxa that are not listed, no material has been
located at Berlin. The list also includes authentic material, i.e. specimens prepared from culti
-
vated parts or descendants of type collections. The nomenclatural base data and notes on lecto-
and neotypifications have been derived from the nomenclatural database maintained at the Suk
-
kulenten-Sammlung Zürich (ZSS).
The Berlin herbarium has a good collection of photographs of type specimens preserved in
other herbaria and these are listed in a separate appendix.
5.2. Conventions adopted
Collection data. Collection data are primarily cited from the relevant protologues and supple
-
mented from data on the herbarium labels, sometimes also from other published and unpublished
sources. Data cited from any of these sources enclosed in double quotation marks, and source(s)
are given in square brackets (e.g. [protologue]). Usually, the data are translated into English,
220 Eggli & Leuenberger: Type specimens of Cactaceae names in the Berlin Herbarium
which is also indicated. Geographical directions (north, south, etc.) are always standardized to
the respective 1-character abbreviations.
Other data. – Further data are given whenever relevant. This refers to Berlin accession numbers,
preservation dates, accession dates, etc. Not all these data are present for all specimens. Standard
herbarium abbreviations for known types are given (first one for the holotype, remaining ones in
alphabetical sequence for isotype(s)).
Examples: (rad), corp, ar, sp = body with areoles and spines and fragments of the root, ic (corp),
fl = illustration of the body, flower
5.3. Alphabetical list of type specimens and other authentic material by basionym
Acanthocalycium glaucum F. Ritter in Taxon 13(4): 143-144. 1964
Notes: When transferred to Echinopsis, the correct name is E. glaucina H. Friedrich & G. D.
Rowley 1974 (non E. glauca (F. Ritter) H. Friedrich & G. D. Rowley 1974).
Type: Ritter 970 loc.1 Argentina: Catamarca: “N of Belén” [protologue, translated], s.a. (type
herbaria: U 117834B) [holotype cited for U in Taxon 12(1): 28. 1963].
B (sheet GH12097) (type number): fl, ic (corp, fl). – Notes: Ex cult. BG Berlin, 22.6.1982, from
material originally raised from seeds received 1960 from H. Winter. Collection number es
-
tablished by inference.
Arequipa spinosissima F. Ritter in Taxon 13(3): 115. 1964
Notes: First mentioned (with Krainz & Ritter as authors) in Backeberg, Cact. 2: 1053. 1959 (RPS
10).
Type: Ritter 196 loc.1 Peru: Arequipa: “Quicacha” [protologue / isotype label at SGO], s.a.
(type herbaria: ZSS AX10497, SGO).
B (sheet GH14195) (type number): ic (corp, ar, sp, fl), fl. Notes: Ex cult. BG Berlin 127-
34-74-80, 7.6.1983, from material originally raised from seeds received 1959 from H. Win
-
ter. Collection number established by inference.
Willdenowia 38 – 2008 221
Abbreviations
adnot. annotation
clono “clonotype”, i.e. cultivated part or descendant of the type collection
cult. cultivation
l.c. loco citato
s.a. sine anno
s.n. sine numero
al bud
ar areole
corp body
fl flower
fr fruit
ic illustrations
+ic for spirit material: separate illustration of the material in the general herbarium
juv juvenile
rad root
sem seed
sp spine
( ) bracketed entries refer to fragmentary material
Armatocereus mataranus F. Ritter in Succulenta (Amsterdam) 46(2): 23, ill. 1967
Notes: First invalidly published (without type) in Succulenta (Amsterdam) 45: 117. 1966.
Type: Ritter 672 loc.1 Peru: Cajamarca: “San Marcos” [protologue / holotype label at U],
6.1957 [collection date according to the original Ritter seed sample at ZSS] (type herbaria: U,
SGO, ZSS [seeds only]).
B (sheet GH31644) (type number): ic (corp), corp, ar, sp. – Notes: Ex cult. BG Berlin 127-38-74-80,
2.8.1993, from material originally raised from seeds received 1961 from H. Winter. Collec
-
tion number established by inference.
Arrojadoa aureispina var. guanambensis P. J. Braun & Heimen in Kakt. And. Sukk. 31(11):
334-337, ill. 1980
Type: Heimen & B raun 80 – Brazil: Bahia: “in the further surroundings of Guanambi” [protolo
-
gue, translated]; “flat rock plates in the shade of a tall scrub forest in a rather moist region”,
400 m, 6.1979 [collection cited as “H/B 80” in the protologue] (type herbaria: B (ex KOELN),
ZSS).
B (ex KOELN) (spirit B 810013100) (holotype): fl, fr. – Notes: Part of the holotype, annotated
as “Glas 1 von 2” [jar 1 of 2]. Collection data given as “HB 80 coll. 6/79 Bras.”.
B (ex KOELN) (spirit B 810013101) (holotype): corp, ar, sp. – Notes: Part of the holotype, an
-
notated as “Holotyp” and “Glas 2 von 2 insges.” [jar 2 of a total of 2 jars]. Collection data
given as “HB 80 coll. 6/79 Bras.”.
B (ex KOELN) (boxed 27654) (isotype): sem. – Notes: Collection data given as “HB 80”.
Arrojadoa beateae P. J. Braun & Esteves in Kakt. And. Sukk. 40(10): 254-256, ill. (pp. 250-
255). 1989
Type: Esteves Pereira 261 – Brazil: Minas Gerais: “Serra do Espinhaço, SW of Montes Claros”
[protologue, standardized]; “in rocks and in quartz sand, in full sun or in the shade of shrubs, tall
grasses and trees”, 1000 m, 7.1987 (type herbaria: UFG, B (ex KOELN)) [the isotypes cited for B
and ZSS in the protologue are paratypes and represent a different collection, although made at the
type locality].
B (ex KOELN) (boxed B 100216251) (isotype): corp, ar, sp.
Arrojadoa canudosensis Buining & Brederoo in Cact. Succ. J. (Los Angeles) 44(3): 111-113, ill.
1972
Type: Horst & Uebelmann HU 251 (1968) Brazil: Bahia: “Canudos” [protologue]; “in the caa
-
tinga bush”, 400 m, 11.6.1968 (type herbaria: U).
B (boxed s.n.) (type number): rad, corp, ar, sp. – Notes: Labelled as “HU 252”, which must be a
slip of the pen for “HU 251”. Leg. P. J. Braun (ex cult.?), labelled as “Typstandort Arrojadoa
canudosensis”.
Arrojadoa dinae var. nana P. J. Braun & Esteves in Kakt. And. Sukk. 42(8): 190-195, ill., SEM-ill.
1991
Type: Braun 415 Brazil: Minas Gerais: “high mountains of the Serra do Espinhaço” [proto
-
logue, translated] / “Montezuma” [isotype labels at B]; “under shrubs or in quartz gravel”
[protologue, translated], 1000 m, 9.1983 (type herbaria: ZSS, B, B (ex KOELN)).
B (boxed s.n.) (isotype): (rad), corp, ar, sp.
B (boxed s.n.) (isotype): (rad), corp, ar, sp, ceph.
B (boxed s.n.) (isotype): corp, ar, sp, ceph.
B (boxed s.n.) (isotype): (rad), corp, ar, sp.
B (boxed s.n.) (isotype): rad, corp, ar, sp.
B (ex KOELN) (spirit B 810013103) (isotype): corp, ar, sp. – Notes: Labelled “Braun 415 Mon
-
tezuma, M. Ger.”
222 Eggli & Leuenberger: Type specimens of Cactaceae names in the Berlin Herbarium
Arrojadoa eriocaulis var. albicoronata Van Heek & al. in Kakt. And. Sukk. 33(11): 224-227, ill.
1982
Type: Van Heek & al. 81/113 Brazil: Minas Gerais: “near Grão Mogol” [protologue, trans
-
lated]; “in sandy-loamy soil between grass and low shrubs in a forested region”, 15.8.1981 [the
collectors' names are not given in the protologue but can be established from circumstantial evi
-
dence] (type herbaria: B (ex KOELN)).
B (ex KOELN) (boxed B 100216252) (holotype): rad, corp, ar, sp. Notes: Simply labelled
“Holotyp” and 81/113”.
B (ex KOELN) (boxed B 100216256) (isotype): rad, corp, ar, sp. – Notes: Labelled “Isotyp” but
without number and stored in a separate bag included in the bag of the holotype.
B (ex KOELN) (spirit B 810013104) (isotype): fl, fr. – Notes: Without type indication nor col
-
lection number, but stored together with the holotype and isotype.
Arrojadoa eriocaulis var. rosenbergeriana Van Heek & W. Strecker in Kakt. And. Sukk. 44(12):
258-262, ill., SEM-ill. 1993
Type: Van Heek & al. 85/216 Brazil: Minas Gerais: “SE of Mato Verde” [protologue, trans
-
lated]; “semi-shade, in sandy-stony soil”, 1985 [the protologue gives only the collection number,
and the collectors' names are established by inference. Detailed locality data deposited together
with the holotype for conservation reasons according to the protologue. No such data were found
with the holotype, however] (type herbaria: B (ex KOELN)).
B (ex KOELN) (spirit B 810013105) (holotype): rad, corp, ar, sp. – Notes: Simply labelled “Ho-
lotyp” and 85/216”.
B (ex KOELN) (boxed B 100216253) (isotype): rad, corp, ar, sp. Notes: Simply labelled as
“Isotyp” and “85/216”.
Arrojadoa horstiana P. J. Braun & Heimen in Kakt. And. Sukk. 32(8): 186-190, ill. 1981
Type: Heimen & Braun 83 – Brazil: Minas Gerais: “Serra do Espinhaço” [protologue] / “moun-
tains E of Mato Verde” [isotype labels at B]; “campo rupestre, mountain forest” [isotype labels at
B, translated], 1000 m, 5.7.1979 [detailed locality data deposited together with the holotype for
conservation reasons according to the protologue, but no such data were found with the holotype]
(type herbaria: B (ex KOELN)).
B (ex KOELN) (spirit B 810013106) (holotype): corp, ar, sp. Notes: Part 1 of 2 of the holo
-
type. Simply labelled “Holotyp” and “HB 83”.
B (ex KOELN) (spirit B 810013107) (holotype): fl. – Notes: Part 2 of 2 of the holotype. Simply
labelled “Holotyp”, “HB 83” and “HB 83 coll. 6/79 Bras.”.
B (boxed s.n.) (isotype): rad, corp, ar, sp. – Notes: Leg. P. J. Braun sub “Braun 83”.
B (boxed s.n.) (isotype): corp, ar, sp. – Notes: Leg. P. J. Braun sub “Braun 83”.
B (ex KOELN) (boxed B 100216254) (isotype): sem. – Notes: Simply labelled “HB 83”.
Arrojadoa rhodantha subsp. reflexa P. J. Braun in Kakt. And. Sukk. 35(2): 34-38, ill. 1984
Type: Braun 68 Brazil: Bahia: “C Bahia, W slopes of the Chapada Diamantina, E of the Rio
Paramirim” [protologue, translated] / “Agua da Seca” [isotype labels at B]; “in sandy soil in the
shade of shrubs and trees” [protologue, translated], 1000 m, 1979 [detailed locality data depos
-
ited together with the holotype for conservation reasons according to the protologue, but no such
data were found with the holotype] (type herbaria: B (ex KOELN), B, ZSS).
B (ex KOELN) (boxed B 100216255) (holotype): corp, ar, sp, fl. – Notes: Simply labelled “Ho
-
lotyp” and Br 68”.
B (boxed s.n.) (isotype): rad, corp, ar, sp.
B (boxed s.n.) (isotype): corp, ar, sp.
B (boxed s.n.) (isotype): rad, corp, ar, sp.
Willdenowia 38 – 2008 223
Arrojadoa rhodantha var. occibahiensis P. J. Braun in Kakt. And. Sukk. 36(6): 114-115, ill.
1985
Type: Horst & Uebelmann HU 616 – Brazil: Bahia: “W of the Rio São Francisco” [protologue,
translated] / “W of Bom Jesus da Lapa” [isotype label at B]; “in dense dry scrub, at the margins of
limestone rocks” [protologue, translated], s.a. (type herbaria: ZSS, B).
B (boxed s.n.) (isotype): corp, ar, sp. – Notes: Leg. P. J. Braun as “Horst & Braun 616” [sphalm.
pro Horst & Uebelmann HU 616].
Arthrocereus itabiriticola P. J. Braun in Kakt. And. Sukk. 37(11): 234-237, ill., SEM-ill. 1986
Type: Horst & Uebelmann HU 330 – Brazil: Minas Gerais: “C Minas Gerais, in the mountains
SW of Belo Horizonte, near the road towards o Paulo” [protologue, translated] / “Betim” [1 of
2 isotype labels at B]; “between itabirit rocks [protologue, translated], 800-1000 m, s.a. (type
herbaria: ZSS, B).
B (boxed s.n.) (isotype): corp, ar, sp. – Notes: Locality given as “Betim”.
B (boxed s.n.) (isotype): corp, ar, sp. Notes: Locality given as “SW Belo Horizonte, Richtung
São Paulo”.
Arthrocereus melanurus var. estevesii Diers & P. J. Braun in Kakt. And. Sukk. 39(5): 100-105,
ill., SEM-ill. 1988
Type: Horst, Braun & Esteves 1 Brazil: Minas Gerais: “high mountain regions of the S Serra do
Espinhaço” [protologue, translated] / “São João del Rey, high mountain regions” [isotype label at
B]; “campo rupestre” [protologue], 1000-1500 m, 9.1983 [detailed locality data deposited to-
gether with the holotype for conservation reasons according to the protologue] (type herbaria: B
(ex KOELN), B, ZSS).
B (ex KOELN) (boxed B 100216257) (holotype): corp, ar, sp. Notes: Part 1 of 3 of the holo-
type. Simply labelled as “Holotyp” and “HBE 1”. A separate print of the protologue with an-
notated locality data is archived separately.
B (ex KOELN) (spirit B 810013108) (holotype): fl. – Notes: Part 2 of 3 of the holotype. Simply
labelled “Holotyp” and “HBE 1”.
B (ex KOELN) (spirit B 810013109) (holotype): fl, fr. – Notes: Part 3 of 3 of the holotype. Sim-
ply labelled “Holotyp” and “HBE 1”.
B (boxed s.n.) (isotype): corp, ar, sp. – Notes: Leg. P. J. Braun sub “Horst, Esteves & Braun 1”.
B (ex KOELN) (boxed B 100216258) (isotype): sem. – Notes: Simply labelled “HBE 1”.
B (ex KOELN) (spirit B 810013110) (isotype): fl. – Notes: Simply labelled “Isotyp” and “HBE 1”.
Austrocephalocereus estevesii subsp. grandiflorus Diers & Esteves in Kakt. And. Sukk. 40(4):
77-81, ill., SEM-ill. 1989
Type: Esteves Pereira 149 Brazil: Goiás: “N of the Serra Geral de Goias, at about latitude 12 de
-
gree” [protologue, translated]; “on naked, dark limestone rocks”, 600-650 m, s.a. [detailed local
-
ity information deposited together with the holotype for conservation reasons according to the
protologue] (type herbaria: B (ex KOELN), ZSS).
B (ex KOELN) (boxed B 100216259) (holotype): corp, ar, sp, ceph. – Notes: Part 1 of 3 of the
holotype. Simply labelled “Holotyp” and “E-149”. A separate print of the protologue with
annotated locality data is archived separately.
B (ex KOELN) (spirit B 810013111) (holotype): fl. – Notes: Part 2 of 3 of the holotype. Simply
labelled “Holotyp” and “E-149”.
B (ex KOELN) (spirit B 810013112) (holotype): fl. – Notes: Part 3 of 3 of the holotype. Simply
labelled “Holotyp” and “E-149”.
B (ex KOELN) (boxed B 100216261) (isotype): sem. – Notes: Simply labelled “E-149”.
B (ex KOELN) (boxed B 100216260) (type collection): corp, ar, sp, ceph. – Notes: A plastic bag
with 2 stem parts, without any label but associated with part 1 (of 3) of the holotype of this
name and thus most probably additional (iso-?) type material.
224 Eggli & Leuenberger: Type specimens of Cactaceae names in the Berlin Herbarium
Austrocephalocereus estevesii subsp. insigniflorus Diers & Esteves in Kakt. And. Sukk. 39(11):
264-268, ill., SEM-ill. 1988
Type: Esteves Pereira 122 Brazil: Minas Gerais: “NW region of Minas Gerais, in the region of
16 deg. latitude” [protologue, translated]; “sunny limestone rocks”, 600-650 m, s.a. [detailed lo
-
cality information deposited together with the holotype for conservation reasons according to the
protologue] (type herbaria: B (ex KOELN)).
B (ex KOELN) (boxed B 100216262) (holotype): corp, ar, sp. – Notes: Simply labelled “Holo
-
typ” and “E-122”. A separate print of the protologue with annotated locality data is archived
separately.
B (ex KOELN) (boxed B 100216263) (isotype): corp, ar, sp. – Notes: Simply labelled “Isotyp”
and “E-122”.
B (ex KOELN) (boxed B 100216264) (isotype): fr, sem. – Notes: Simply labelled “E-122”.
B (ex KOELN) (spirit B 810013113) (isotype): fl. – Notes: Simply labelled “E-122”.
B (ex KOELN) (spirit B 810013114) (isotype): fl. – Notes: Simply labelled “E-122”.
Binghamia climaxantha Werdermann in Repert. Spec. Nov. Regni Veg. 42: 4-5. 1937 (incorrect
name, Art.11.4).
Type: Blossfeld 84 Peru: Lima: near Chosica” [protologue, translated]; “sunny steep slopes”,
1936 (type herbaria: B) [ex cult. BG Berlin (“lebend eingeführt”)].
B (spirit B 810006687) (isotype): corp, ar, sp. Notes: Ex cult. BG Berlin, April 1937. Since
the protologue (p. 5) specifically mentions that both living and herbarium material was re-
ceived from H. Blossfeld, the specimen must be treated as isotype and not as holotype.
Blossfeldia liliputana Werdermann in Kakteenkunde 1937(11): 162-163, ill. 1937
Type: Blossfeld & Marsoner s.n. – Argentina: Jujuy: “near Tumbaya” [protologue, translated],
1500 m, 1936 (type herbaria: B) [ex cult. BG Berlin 1937].
B (spirit B 810006688) (holotype):
rad, corp, ar, (fl). – Notes: Ex cult. BG Berlin 1937. Com-
pletely dried out. For an illustration see Leuenberger & Eggli (1999a: 3, fig. 1); the envelope
contains a flower fragment, no seeds.
Cactus bleo Kunth in Humboldt & al., Nov. Gen. Sp. 6: 69. 1823
Type: Humboldt & Bonpland 1546 Colombia: Bolívar: “in ripa fluminis Magdalenae prope
Badillas” [protologue] / “Río Magdalena (Badillas)” [holotype label at P-Bonpl], 5.1805 [collec
-
tion number and locality according to Leuenberger in Willdenowia 32: 138. 2002] (type herbaria:
P-Bonpl [lecto], B-W, P) [Lectotype designated by Leuenberger in Mem. New York Bot. Gard.
41: 99. 1986; type herbaria according to Leuenberger in Willdenowia 32: 138. 2002].
B (sheet B-W 9442-1) (isotype): fol, (fl).
Cactus caripensis Kunth in Humboldt & al., Nov. Gen. Sp. 6: 66. 1823
Type: Humboldt & Bonpland 1249 – Venezuela: “prope Caripe” [protologue] / “Caripe” [holo
-
type label at P-Bonpl] / “Cumana, in lignis putrid. Caripé” [isotype label at B-W], s.a. [collection
number and locality data according to Leuenberger in Willdenowia 32: 138, 150. 2002] (type her
-
baria: P-Bonpl, B-W [sub Cactus pilosus]) [type herbaria according to Leuenberger in Willde
-
nowia 32: 144. 2002; name attached to the B-W specimen according to Leuenberger in
Willdenowia 34: 314. 2004].
B (sheet B-W 9431-1) (isotype): rad, corp (juv), ar, sp. Notes: Labelled Cactus pilosus (cf.
Leuenberger in Willdenowia 34: 314. 2004).
Cactus horridus Kunth in Humboldt & al., Nov. Gen. Sp. 6: 70. 1823, nom. illeg., Art. 53.1.
Notes: Non Cactus horridus Salisbury 1796, nec Colla 1833 (= Echinocactus horridus Gay
1848). The correct name under Pereskia is P. horrida DC. 1828.
Willdenowia 38 – 2008 225
Type: Humboldt & Bonpland 3594 Peru: “ad flumen Marañón, Provinciae de Jaén de Braca
-
moros” [protologue] / “Marañón” [holotype label at P-Bonpl]; “in collibus siccis” [protologue],
8.1802 (type herbaria: P-Bonpl [lecto], B-W, MO, P) [lectotype designated by Leuenberger in
Mem. New York Bot. Gard. 41: 67. 1986; type herbaria according to Leuenberger in Willdenowia
32: 138. 2002].
B (sheet B-W 9441-1) (isotype): corp, ar, sp, fol.
Cactus icosagonus Kunth in Humboldt & al., Nov. Gen. Sp. 6: 67. 1823
Type: Humboldt & Bonpland 3293 Ecuador: “prope pagum Nabón” [protologue] / “Nabon”
[holotype label at P-Bonpl], s.a. [locality data according to Leuenberger in Willdenowia 32: 150.
2002] (type herbaria: P-Bonpl, B-W) [type herbaria according to Leuenberger in Willdenowia
32: 146. 2002].
B (sheet B-W 9429-1) (isotype): al, fl.
Cactus micranthus Kunth in Humboldt & al., Nov. Gen. Sp. 6: 65. 1823
Type: Humboldt & Bonpland 3494 Peru: “prope Olleros Quitensium” [protologue] / “Olleros”
[holotype label at P-Bonpl], 1330 m, s.a. [collection number according to Leuenberger in
Willdenowia 32: 138, 144. 2002] (type herbaria: P-Bonpl, B-W [sub Cactus variabilis], P) [type
herbaria according to Leuenberger in Willdenowia 32: 145. 2002; name attached to specimen at
B-W according to Leuenberger in Willdenowia 34: 315. 2004].
B (sheet B-W 9433-1) (isotype): corp, ar, sp. Notes: Labelled Cactus variabilis (cf. Leuen-
berger in Willdenowia 34: 315. 2004).
Cactus sepium Kunth in Humboldt & al., Nov. Gen. Sp. 6: 67. 1823
Type: Humboldt & Bonpland 3216 Ecuador: Chimborazo: “ad radicem Chimborazo prope
Riobamba” [protologue] / “Chimborazo” [holotype label at P-Bonpl], 2660 m, s.a. [collection
number according to Leuenberger in Willdenowia 32: 138. 2002] (type herbaria: P-Bonpl, B-W
[iso?, sub Cactus endecagonus]) [type herbaria according to Leuenberger in Willdenowia 32:
148. 2002; status of material at B according to Leuenberger in Willdenowia 34: 313. 2004].
B (sheet B-W 9423-1) (isotype): fl. Notes: Labelled Cactus endecagonus. Type status ques-
tionable, cf. Leuenberger in Willdenowia 34: 313. 2004.
Cactus smithii Alexander in Lloydia 2(3): 200-201. 1939
Type: Smith 3388 Guyana: Rupununi Distr.: “NW slopes of Kanuku Mt., in drainage of Moku-
moku creek (Takutu tributary)” [printed herbarium labels, standardized]; “in dense forest”, 150-
400 m, 31.3.1938 (type herbaria: B, G, K, LE, NY, U, W).
B (sheet s.n.) (isotype): (corp), ar, sp, ceph.
Calymmanthium substerile F. Ritter, Kakt. And. Sukk. 13(2): 25-28, ill. 1962
Notes: First mentioned as nom. nud. in Backeberg, Cact. 2: 887. 1959 (cf. RPS 10).
Type: Ritter 315 loc.1 Peru: Cajamarca; Jaén: “lower Huancabamba gorge” [protologue, trans
-
lated], “Bucara” [Pucará] [specimen data at ZSS], 8.1954 (type herbaria: SGO, U097800B, ZSS).
B (sheet GH20873) (type number): ic (corp). – Notes: Ex cult. BG Berlin, 26.6.1986, from ma
-
terial raised from seeds received 1957 from H. Winter. Collection number established by in
-
ference.
B (sheet GH20873a) (type number): corp, ar, sp. Notes: Ex cult. BG Berlin, 21.7.1993, from
material raised from seeds received 1957 from H. Winter. Collection number established by
inference.
B (sheet GH20873b) (type number): fl, ic (fl). – Notes: Ex cult. BG Berlin, 11.9.1996, from ma
-
terial raised from seeds received 1957 from H. Winter. Collection number established by in
-
ference.
226 Eggli & Leuenberger: Type specimens of Cactaceae names in the Berlin Herbarium
Cephalocereus blossfeldiorum Werdermann in Kakteenkunde 1937: 4-5. 1937
Type: Blossfeld 88 Peru: Cajamarca: “near Huancabamba” [protologue, translated], 1936 (type
herbaria: B) [ex cult. BG Berlin (“Pflanze lebend eingeführt”)].
B (spirit B 810006691) (holotype): corp, ar, sp. – Notes: Ex cult. BG Berlin, s.a.; Leuenberger
(1979) erroneously reports this specimen as consisting of only a flower.
Cephalocereus kanukuensis Alexander in Lloydia 2(3): 200. 1939
Type: Smith 3380 British Guiana: “NW slopes of Kanuku Mts., in drainage of Moku-moku
Creek (Takutu tributary)” [printed herbarium specimen labels]; “in dense forest”, 300 m, 31.3.
1938 (type herbaria: B, G, K, LE, MO, NY, P, U 45282A (Box 911), US).
B (sheet s.n.) (isotype): corp, ar, sp, ic (corp, fl).
Cephalocereus melanostele Vaupel in Bot. Jahrb. Syst. 50 (Beiblatt 111): 12-13. 1913
Notes: Abridged version published in Monatsschr. Kakteenk. 24(11): 154-155. 1914.
Type: Weberbauer 2630 Peru: Lima: “near Chosica, at the railway Lima - Oroya” [protologue,
translated]; “stony ground with very poor vegetation”, 800 m, 15.3.1903 (type herbaria: B).
B (spirit B 810006699) (holotype): corp, ar, sp, ceph, fl.
Cephalocereus purpureus Gürke in Monatsschr. Kakteenk. 18(6): 86. 1908
Notes: Non Cephalocereus purpureus Werdermann 1933.
Type: Ule 4 Brazil: Bahia: “Serra do Sincóra” [protologue]; “on rocks”, 800-1200 m, s.a. (type
herbaria: B).
B (spirit B 810006695) (holotype): corp, ar, sp, ceph.
Cephalocereus ulei Gürke in Monatsschr. Kakteenk. 18(6): 85. 1908
Type: Ule 12 Brazil: Bahia: “on rocks of the Serra do São Ignacio” [protologue, translated], s.a.
(type herbaria: B).
B (spirit B 810006698) (holotype): corp, ar, sp, fl.
Cereus acanthurus Vaupel in Bot. Jahrb. Syst. 50 (Beiblatt 111): 13-14. 1913
Notes: Abridged version published in Monatsschr. Kakteenk. 24(11): 155. 1914.
Type: Weberbauer 1699 Peru: Lima: “near Matucana, a station of the railway from Lima to
Oroya” [protologue, translated]; “among rocks with sparse vegetation”, 2370 m, 1.11.1902 (type
herbaria: B).
B (spirit B 810006701) (holotype): rad, corp, ar, sp, fl.
Cereus acranthus K. Schumann ex Vaupel in Bot. Jahrb. Syst. 50 (Beiblatt 111): 14-15. 1913
Notes: Abridged version published in Monatsschr. Kakteenk. 24(11): 155-156. 1914. Name first
used as Pilocereus acranthus K. Schumann (nom. nud.) by Weberbauer, Pfl.-Welt Peruan.
Anden: t. 5b. 1911.
Type: Weberbauer 1679 Peru: Lima: “near Santa Clara, a station of the railway Lima - Oroya”
[protologue, translated]; stony ground in the Loma formation”, 400-600 m, 26.10.1902 (type
herbaria: B).
B (spirit B 810006700) (holotype): corp, ar, sp, fl, [+ic].
Cereus acutangulus Pfeiffer, Enum. Diagn. Cact.: 107. 1837
Notes: Sometimes also cited from the unpublished source Otto, Cat. hort. Berol. 1833.
Type: Anonymus s.n. in B Garden-Herbarium 15056 Cult.: s.l. [neotype designation], 2.9.1983
(type herbaria: B [neo]) [neotype designated by Leuenberger in Bot. Jahrb. Syst. 123(2): 174.
2001].
B (sheet GH15056) (neotype): fl, fr, sem, ic (corp). – Notes: Ex cult. BG Berlin, s.n., 2.9.1983.
Willdenowia 38 – 2008 227
Cereus adscendens Gürke in Monatsschr. Kakteenk. 18(5): 66-69, ill. 1908
Type: Ule 7072 Brazil: Bahia: “near Tambury” [protologue, translated]; “low ground in the
Caatinga”, 10.1906 (type herbaria: B [lecto], HBG, K [photo]) [lectotype designated by Taylor &
Zappi, Cacti Eastern Brazil: 426. 2004].
B (spirit B 810006702) (lectotype): corp, ar, sp, fl, [+ic]. Notes: In addition, B has a photo
-
graph of the isotype at HBG.
Cereus amazonicus K. Schumann ex Vaupel in Notizbl. Königl. Bot. Gart. Berlin 5(50):
283-284. 1913
Notes: Abridged version with an added illustration published in Monatsschr. Kakteenk. 23(11):
164-167. 1913. Name also mentioned by Ule in Bot. Jahrb. Syst. 40: 412. 1913.
Type: Ule 53 Peru: Loreto: “at the Huallaga [River] near Tarapoto” [protologue], 9.1902 (type
herbaria: B [status?]).
B (spirit B 810006703) (holotype): corp, ar, sp, fl [+ic]. – Notes: The bottle is without collection
number and year, and the treatment of this material as holotype relies on the admittedly
weak fact that this is the only specimen present at B, and that the material is absolutely con
-
gruent with the figure of the type in Monatsschr. Kakteenk. 23: 165. 1913. B also has a pho
-
tograph of a topotype (Ule 6314, “Tarapoto, Oct.1902”, sheet specimen [corp, ar, sp, fl]) at
HBG.
Cereus anisitsii K. Schumann in Monatsschr. Kakteenk. 9(12): 185-186. 1899
Type: Anisits Cact. 19 Paraguay: “at the rivulet Tagatiya-mi, near Porongo” [protologue, trans-
lated]; “on sandy ground”, 25.1.1898 (type herbaria: B).
B (spirit B 810006704) (holotype): corp, ar, sp, fl.
Cereus apiciflorus Vaupel in Bot. Jahrb. Syst. 50 (Beiblatt 111): 15-16. 1913
Notes: Abridged version published in Monatsschr. Kakteenk. 24(11): 156-157. 1914.
Type: Weberbauer 3743 Peru: Ancash; Huari: “above Masin in the valley of the Puccha River”
[protologue, translated]; “in open scrub consisting of shrubs, cacti, agaves, bromeliads and
herbs”, 2500-2600 m, 26.10.1903 (type herbaria: B [fragment only]).
B (spirit B 810006705) (isotype): fl. – Notes: Dried out. The apparent holotype material was illus
-
trated by Werdermann in Kakteenkunde 1940: 5-6, which was subsequently destroyed at B.
Cereus bonplandii Parmentier ex Pfeiffer, Enum. Diagn. Cact.: 108. 1837
Notes: Sphalm. ‘bomplandii’.
Type: Guaglianone & Múlgura 2228 Argentina: Formosa; Pilcomayo: “Parque Nacional Pil
-
comayo, alrededores de Laguna Blanca, hacia el E” [neotype label at B]; “palmar”, 14.12.1988
(type herbaria: SI [neo], B) [neotype designated by R. Kiesling in Darwiniana 34: 391. 1996].
B (sheet s.n.) (isoneotype): (corp, ar, sp), fl, fr, sem.
Cereus brachypetalus Vaupel in Bot. Jahrb. Syst. 50 (Beiblatt 111): 16-17. 1913
Notes: Abridged version published in Monatsschr. Kakteenk. 24(12): 161-162. 1914.
Type: Weberbauer 1549 – Peru: Arequipa: “near Mollendo” [protologue, translated]; “in rocky
sandy ground in the Loma formation”, 150 m, 7.10.1902 (type herbarium: B).
B (spirit B 810006717) (holotype): corp, ar, sp, fl, fr.
Cereus brevistylus K. Schumann ex Vaupel in Bot. Jahrb. Syst. 50 (Beiblatt 111): 17-18. 1913
Notes: Abridged version published in Monatsschr. Kakteenk. 24(12): 162-163. 1914. Name first
used by Weberbauer, Pfl.-Welt Peruan. Anden: 128-129. 1911.
228 Eggli & Leuenberger: Type specimens of Cactaceae names in the Berlin Herbarium
Type: Weberbauer 1414 Peru: Arequipa: “near Yura (station of the railway Arequipa – Puno)”
[protologue, translated]; “on stony ground with scarce vegetation", 2400 m, 31.8.1902 (type her
-
barium: B).
B (spirit B 810006719) (holotype): corp, ar, sp, fl.
Cereus decumbens Vaupel in Bot. Jahrb. Syst. 50 (Beiblatt 111): 18-19. 1913
Notes: Abridged version published in Monatsschr. Kakteenk. 24(12): 163. 1914.
Type: Weberbauer 1550 – Peru: Arequipa: “near Mollendo” [protologue, translated]; “on stony
sandy ground with scattered vegetation”, 50-100 m, 7.10.1902 (type herbaria: B).
B (spirit B 810006731) (holotype): corp, ar, sp, fl [+ic].
Cereus deserticola Werdermann in Notizbl. Bot. Gart. Berlin-Dahlem 10(96): 764-765. 1929
Notes: Sphalm. ‘deserticolus’.
Type: Werdermann 869 p.p. Chile: Antofagasta; Taltal: “near Taltal” [protologue], 500 m,
10.1925 [this is a mixed collection and only the flowers are type of Cereus deserticola, see
Leuenberger & Eggli in Willdenowia 25: 690. 1995] (type herbaria: B [lecto], BM, G, K, U, US)
[lectotype according to Leuenberger & Eggli in Willdenowia 25: 690. 1996].
B (sheet s.n.) (holotype): fl.
Cereus extensus Salm-Dyck ex DC., Prodr. 3: 469. 1828
Type: de Granville & al.10991 French Guiana: “Bassin du Maroni, Gobaya Soula (on the river-
bank bordering Suriname)” [neotype designation] / “Atachi Bacca-Berge, (Camp 1), forest on the
right-hand bank of the Itany River” [neotype label, translated], 100 m, 31.1.1989 (type herbaria:
B [neo]) [neotype (ex cult. BG Berlin as Cubr 33353) designated by Leuenberger in Bot. Jahrb.
Syst. 123: 56. 2001].
B (sheet GH33353) (neotype): corp, ar, sp, fl.
Cereus glaziovii K. Schumann in Fl. Bras. 4: 200-201. 1890
Type: Glaziou s.n. – Brazil: Minas Gerais: “Pico d’Itabora do Campo” [protologue], 20.12.1888
(type herbaria: B, K [photo], SPF [photo]).
B (spirit B 810006739) (holotype): corp, ar, sp, fr [+ic].
Cereus grandicostatus Werdermann in Notizbl. Bot. Gart. Berlin-Dahlem 12: 460-461. 1935
Type: Anonymus s.n. in Huntington BG 9/17 s.l., s.a. (type not preserved) [cult. Huntington
BG].
B (sheet 27600) (type collection): ic (corp, ar, sp). Notes: 6 prints made 1987 from original
Werdermann negatives, labelled “cult. Huntington Botanical Garden 9-17, San Marino, Cali
-
fornia”, “Foto E. Werdermann, 1933”.
B (sheet s.n.) (type collection): corp, ar, sp, fl. – Notes: A total of 5 sheets prepared ex cult. BG
Berlin 1982, 1986, 1992, 1998 and 2001 (from 043-84-74-80, a plant originally received
from “Kalifornien 1966”, thus most probably representing the living original material of the
name cultivated at the Huntington Botanical Garden, San Marino, California).
Cereus hassleri K. Schumann in Monatsschr. Kakteenk. 10(3): 45-46. 1900
Type: Anisits Cact. 13 Paraguay: “at the Estancia Tagatiya” [protologue], 20.1.1898 (type her
-
baria: B [syn], G [isosyn], K [isosyn]) [based on 2 syntypes].
B (spirit B810006743) (syntype): corp, ar, sp, fl. – Notes: This is 1 of 2 syntypes for this taxon.
Cereus hertrichianus Werdermann in Notizbl. Bot. Gart. Berlin-Dahlem 12: 459-460. 1935
Type: Anonymus s.n. in Huntington BG 44/65 s.l., s.a. (type not preserved) [cult. Huntington
BG].
Willdenowia 38 – 2008 229
B (sheet s.n.) (type collection): ic (corp, ar, sp, fl). Notes: 2 sheets with prints of a total of 4
photographs made 1987 from original Werdermann negatives “Foto E. Werdermann 1933”,
“cult. Huntington Botanical Garden 44-65, San Marino, California” (number later added by
hand to type-written label).
Cereus horridus var. alatosquamatus Werdermann in Notizbl. Bot. Gart. Berlin-Dahlem 12:
452. 1935 (incorrect name, Art.11.4).
Type: Anonymus s.n. in Huntington BG 41/75 s.l., s.a. (type not preserved) [cult. Huntington
BG].
B (sheet s.n.) (type collection): ic (corp, ar, sp). – Notes: 2 prints made 1987 from original Wer
-
dermann negatives labelled “Foto E. Werdermann 1933”, “cult. Huntington Botanical Gar
-
den 41-75, San Marino, California”.
Cereus iquiquensis K. Schumann in Monatsschr. Kakteenk. 14(7): 99-100. 1904
Type: Reiche s.n. – Chile: Atacama: “Atacama desert near Iquique” [protologue, translated; lec
-
totype specimen index card], s.a. (type herbaria: B [lecto]) [lectotype designated by Leuenberger
& Eggli in Haseltonia 7: 76. 2000].
B (spirit B 810006746) (lectotype): fl, [+ic].
Cereus lamprospermus K. Schumann in Monatsschr. Kakteenk. 9: 166. 1899
Type: Anisits Cact. 2 – Paraguay: “Río Paraguay, near Fuerte Olympo” [protologue, translated,
standardized], 12.1.1898 (type herbaria: B [lecto]).
B (spirit B810006749) (lectotype): corp, ar, sp, fl, fr, sem, +ic. – Notes: Lectotype (here desig-
nated) of Cereus lamprospermus. Berlin also has a spirit specimen of the remaining syntype,
Anisits Cact. 11 (B810006758, corp, ar, sp, fl, +ic).
B (spirit B810006750) (isolectotype): corp, ar, sp, fl, fr, sem, +ic.
Notes. This name is based on two syntypes, Anisits Cact. 2 (2 spirit specimens at B) and
Anisits Cact. 11 (1 spirit specimen at B). The material of all 3 specimens is compatible with
the protologue and we here select the most complete specimen as lectotype.
Cereus leucostele Gürke in Monatsschr. Kakteenk. 18(4): 53. 1908
Type: Ule 2 – Brazil: Bahia: “in the S catinga near Calderão” [protologue, translated], s.a. (type
herbaria: B).
B (spirit B 810006756) (holotype): corp, ar, sp, ceph, fr, sem?
Cereus longiflorus Alexander in Lloydia 2(3): 199-200. 1939
Type: Smith 3387 – British Guiana: Rupununi Distr.: “NW slopes of Kanuku Mt, in drainage of
Moku-moku Creek (Takutu tributary)” [printed herbarium labels, standardized]; “in dense for
-
est”, 275 m, 31.3.1938 (type herbaria: NY, B [photo], G [photo], K, LE, MO, P, U [photo]).
B (sheet s.n.) (isotype): ic (corp, ar, sp, fl). – Notes: Original photograph and label, accessioned 1956.
Cereus luetzelburgii Vaupel in Z. Sukk. 1(6): 57, ill. 1923
Type: Lützelburg 22 – Brazil: Bahia: “in the Serra das Almas, 1500 m, and the Serra Marsalina,
1400 m” [protologue, translated], 7.1913 (type herbaria: B [fragment]).
B (spirit B 810006759) (holotype): fl. – Notes: Dried out.
Cereus micranthus Vaupel in Bot. Jahrb. Syst. 50 (Beiblatt 111): 19-20. 1913
Notes: Non Rhipsalis micrantha (Kunth) DC 1828. Abridged version published in Monatsschr.
Kakteenk. 24(12): 163-164. 1914.
230 Eggli & Leuenberger: Type specimens of Cactaceae names in the Berlin Herbarium
Type: Weberbauer 1353 Peru: Puno: “near Sandía” [protologue, translated]; “on rocks between
scrub”, 2100 m, 31.7.1902 (type herbaria: B, US).
B (spirit B 810006765) (holotype): corp, ar, sp, fl, fr, sem?. – Notes: Dried out.
Cereus pachyrrhizus K. Schumann, Gesamtbeschr. Kakt., Nachtr.: 33-34. 1903
Type: Anisits Cact. 39 Paraguay: “on the Cerro Noaga” [protologue, translated]; “naked granite
rocks”, 350 m, s.a. (type herbaria: B).
B (spirit B 810006771) (holotype): corp, ar, sp, fr, sem? [+ic].
Cereus penicillatus Gürke in Monatsschr. Kakteenk. 18(5): 70-71. 1908
Type: Ule 7052 Brazil: Bahia; Maracás: “near Calderão” [protologue, translated]; caatinga, s.a.
(type herbaria: B, HBG, K [photo]).
B (spirit B 810006773) (holotype): corp, ar, sp, ceph, fl.
Cereus peruvianus var. persicinus Werdermann in Notizbl. Bot. Gart. Berlin-Dahlem 12: 448. 1935
Type: Anonymus s.n. in Huntington BG 9/7 – s.l., s.a. (type not preserved).
B (sheet s.n.) (type collection): ic (corp, ar, sp, fl). – Notes: Prints made 1987 from negatives in
the Werdermann archives, “Foto E. Werdermann 1933”, cult. Huntington Botanic Garden,
California. Even though the photographs do not show the type number (Huntington acces
-
sion number), they almost certainly illustrate the type of this name.
Cereus phatnospermus K. Schumann in Monatsschr. Kakteenk. 9(12): 186-187. 1899
Type: Anisits Cact. 27 Paraguay: Presidente Hayes: “bei Porongo” [protologue], 18.2.1898
(type herbaria: B).
B (spirit B 810006775) (holotype): corp, ar, sp, (al), +ic.
B (spirit B 810006776) (isotype): corp, ar, sp, fl, fr, sem, +ic. – Notes: The original label gives
Cereus phatnospermus K. Sch. Paraguay leg. Anisits”, without date or collection number.
According to the protologue, Anisits 27 is the only material present, and so it is reasonable to
assume that this specimen is part of the original gathering. The labels of this bottle and the
holotype bottle moreover look very similar, have the same handwriting and are of the same
size. The problem is, however, slightly more complicated as there is yet another specimen of
this taxon at B (B 810006777) labelled though in a different handwriting and on a larger-
sized label – as Anisits 49. This could probably be a later addition to the herbarium, received
from Anisits subsequent to the publication of the protologue of this name.
Cereus piauhyensis Gürke in Monatsschr. Kakteenk. 18(6): 84-85. 1908
Type: Ule 9 – Brazil: Piauí: “on rocks of the Serra Branca” [protologue] [Mpio. São Raimundo
Nonato], 1.1907 (type herbaria: B, K [photo], SPF [photo]).
B (spirit B 810006778) (holotype): corp, ar, sp, fl, [+ic].
Cereus plagiostoma Vaupel in Bot. Jahrb. Syst. 50 (Beiblatt 111): 20-21. 1913
Notes: Abridged version published in Monatsschr. Kakteenk. 24(12): 164-167, ill. 1914.
Type: Weberbauer 3906 Peru: Cajamarca: “near San Miguel” [protologue, translated]; “in open
to almost closed association of herbs (numerous grasses), cacti, bromeliads and shrubs”, 2200 m,
5.5.1904 (type herbaria: B).
B (spirit B 810006780) (holotype): corp, ar, sp, fl, [+ic].
Cereus pseudosonorensis Gürke in Monatsschr. Kakteenk. 20: 147, 173. 1910
Type: Runge s.n. Mexico: Sonora: s.l. [protologue], s.a. (type herbaria: B [status ?]) [ex cult. BG
Berlin].
B (spirit B 810006798) (holotype): ar, sp, fl.
Willdenowia 38 – 2008 231
Notes. This name implicitly includes the type of Cereus sonorensis Runge ex K. Schumann
1901 (see below under the latter name).
Cereus regelii Weingart in Monatsschr. Kakteenk. 20(3): 33. 1910
Type: Anonymus in B Garden Herbarium 12552a Cult., s.l., s.a. (type herbaria: B [neo], SI)
[neotype designated by Leuenberger in Kakt. And. Sukk. 47(2): 35. 1996].
B (sheet GH12552a) (neotype): corp, ar, sp, fl, ic. Notes: Ex cult. BG Berlin 045-88-74-80,
10.8.1994, from cultivated material without known wild origin.
Cereus rhodanthus Gürke in Monatsschr. Kakteenk. 18(5): 69. 1908
Type: Ule 11 Brazil: Piauí: “in the Catinga de São Raimundo” [protologue, translated], s.l. (type
herbaria: B).
B (spirit B 810006785) (holotype): corp, ar, sp, ceph, fl, [+ic].
Cereus rhodoleucanthus K. Schumann in Monatsschr. Kakteenk. 9: 187. 1899
Type: Anisits Cact. 18 Paraguay: “near the Estancia Tagatiya” [protologue, translated],
25.1.1898 (type herbaria: B [lecto]).
B (spirit B810006787) (lectotype): corp, ar, sp, fl, fr, sem, +ic. – Notes: Lectotype (here desig
-
nated) of Cereus rhodoleucanthus. In addition, B has dried out specimens of the remaining
syntypes Anisits Cact. 5 (B810006788, corp, ar, sp, fl, fr, sem, +ic) and Anisits Cact. 24
(B810006786, corp, ar, sp, (fl), +ic).
Notes. This name was based on 5 syntypes, Anisits Cact. 5 (1 spirit specimen present at B,
dried out), Anisits Cact. 18 (1 spirit specimen present at B), Anisits Cact. 24 (1 spirit specimen
present at B, dried out), Anisits Cact. 62 (no material traced), and Anisits Cact. 65 (no material
traced). The specimen of Anisits Cact. 18 is the only good material available for lectotypi-
fication. It is wholly compatible with the protologue and is selected as lectotype here.
Cereus sonorensis Runge ex K. Schumann in Monatsschr. Kakteenk. 11: 135-136, ill. 1901
Notes: Frequently attributed to Gürke in Monatsschr. Kakteenk. 20: 146. 1910.
Type: Runge s.n. Mexico: Sonora: s.l. [protologue], s.a. (type herbaria: B) [ex cult. BG Berlin].
B (spirit B 810006798) (holotype): ar, sp, fl. – Notes: Labelled “Cereus sonorensis Runge ded.
K. Schumann”, but without year. Areole and flower are obviously the material illustrated in
the drawing that accompanies the protologue.
Notes. The first valid description of Cereus sonorensis was published by K. Schumann in Mo
-
natsschr. Kakteenk. 11: 135. 1901, but the name was used before by Runge, probably in a catalogue
(not traced). Gürke (1910) was of the opinion that the original material of Runge represented a dif
-
ferent taxon, for which he published the name C. pseudosonorensis Gürke. However, Gürke based
his new name partly on the Schumann description, partly on additional material. Since the flower
details are, however, copied from Schumann, the type of the Schumann name is implicitly in
-
cluded in Gürke’s concept of Cereus pseudosonorensis, which renders this name illegitimate un
-
der ICBN Art. 52.1. C. pseudosonorensis is thus typified by the type of C. sonorensis.
To make matters worse, Gürke compared his Cereus pseudosonorensis with “Cereus sono
-
rensis”, but from the text it becomes clear that this is not C. sonorensis Runge ex K. Schumann
1901, but a supposedly different taxon. Gürke’s concept is an implicit publication of the name C.
sonorensis Runge ex Gürke 1910, which is illegitimate (Art. 53.1) versus C. sonorensis Runge ex
K. Schumann 1901.
Cereus squamosus Gürke in Monatsschr. Kakteenk. 18: 70. 1908
Type: Ule 10 Brazil: Piauí: “on rocks of the Serra Branca” [protologue, translated], 1.1907
(type herbaria: B, K [photo], SPF [photo]).
B (spirit B 810006799) (holotype): corp, ar, sp, fl, [+ic].
232 Eggli & Leuenberger: Type specimens of Cactaceae names in the Berlin Herbarium
Cereus squarrosus Vaupel in Bot. Jahrb. Syst. 50 (Beiblatt 111): 21-22. 1913
Notes: Abridged version published in Monatsschr. Kakteenk. 24(12): 168. 1914.
Type: Weberbauer 1719 Peru: Junín: “near Tarma” [protologue, translated]; “loamy ground
with scattered vegetation”, 3000-3100 m, 25.11.1902 (type herbaria: B).
B (spirit B 810006801) (holotype): corp, ar, sp, fl.
Cereus stenogonus K. Schumann in Monatsschr. Kakteenk. 9: 165-166. 1899
Type: Anisits Cact. 53 – Paraguay: San Salvador: “near Paso La Cruz” [protologue, translated],
22.2.1898 (type herbaria: B [lecto]) [R. Kiesling in Bol. Soc. Argent. Bot. 24: 383. 1988, men
-
tions type material at NY but the relevant specimen is not one of the syntypes].
B (spirit B810006806) (lectotype): corp, ar, sp, fl, fr, sem, +ic. – Notes: Lectotype (here designa
-
ted) of Cereus stenogonus. In addition, B has one spirit specimen each of the remaining syn
-
types Anisits Cact. 31 (B810006807, corp, ar, sp, fl, (fr), +ic) and Anisits Cact. 54 (B810006804,
corp, ar, sp, (fr), +ic).
Notes. – The name was based on 3 syntypes, Anisits Cact. 31, Anisits Cact. 53 and Anisits Cact.
54, each of which is represented by a spirit specimen at B. All are compatible with the proto
-
logue. Anisits 53 is the most complete specimen, including a fruit and seeds, and is selected here
as lectotype.
Cereus torulosus Werdermann in Notizbl. Bot. Gart. Berlin-Dahlem 12: 457-459. 1935
Type: Anonymus s.n. in Huntington BG 36/4 – s.l., s.a. (type not preserved).
B (sheet s.n.) (type collection): ic (corp, ar, sp, fl). – Notes: A print made 1987 from an original
negative in the Werdermann archives, made by Werdermann in 1933 and showing the culti-
vated type specimen at the Huntington Botanical Garden, California.
Cereus trigonus var. guatemalensis Eichlam ex Weingart in Monatsschr. Kakteenk. 21(5): 69-
70. 1911
Type: cult. Knippel s.n. – s.l., 1911 (type herbaria: B).
B (spirit B 810006825) (holotype): corp, ar, sp. Notes: Original label “ded. Weingart 25.4. 1911”,
thus with nearly absolute certainty the holotype.
Notes. This name was first used by Eichlam (in Monatsschr. Kakteenk. 21(2): 18-21. 1911),
based on Eichlam no. 25, but the name was treated as synonym of Cereus purpusii, and is thus
there invalid under Art. 34.1a. The neotypification by Doweld with Deam 695 (MO) in
Turczaninowia 5: 13. 2002, is unnecessary in the light of the confirmed presence of the holotype
at B. This material was deposited by Weingart either shortly before or concurrently with the
publication of the protologue of the name and is therefore to be treated as the holotype.
Cereus weberbaueri K. Schumann ex Vaupel in Bot. Jahrb. Syst. 50 (Beiblatt 111): 22-23.
1913
Notes: Abridged version published in Monatsschr. Kakteenk. 24(12): 168-169. 1914. Name first
used (without description) in Weberbauer, Pfl.-Welt Peruan. Anden: 128-129. 1911.
Type: Weberbauer 1413 Peru: Arequipa: “near Yura, a station of the railway Arequipa - Puno”
[protologue, translated]; “on stony ground with scattered vegetation”, 2400 m, 31.8.1902 (type
herbaria: B).
B (spirit B 810006830) (holotype): corp, ar, sp, fl, [+ic].
Cereus wittii K. Schumann in Monatsschr. Kakteenk. 10(10): 154-157, ill. 1900
Type: Witt s.n. Brazil: Amazonas: “imported from Manaos” [protologue, translated], 1900
(type herbaria: B).
B (spirit B 810006832) (holotype): corp, ar, sp, fr.
Willdenowia 38 – 2008 233
Cereus xanthocarpus K. Schumann, Gesamtbeschr. Kakt., Nachtr.: 32. 1903
Type: Anisits Cact. 59 Paraguay: “in the Calle Manora” [protologue, translated], s.a. (type her
-
baria: B [lecto]).
B (spirit B810006833) (lectotype): corp, ar, sp, fr, sem, +ic. Notes: Lectotype (here desig
-
nated) of Cereus xanthocarpus.
Notes. This name was based on 2 syntypes, Anisits Cact. 59 and Anisits Cact. 60, from the
same locality. Only Anisits Cact. 59 could be located at B. The material is completely compati
-
ble with the protologue, and the specimen is selected as lectotype here.
Chileorebutia glabrescens F. Ritter in Cactus (Paris) No. 64: suppl.: 1. 1959 (incorrect name,
Art. 11.4).
Notes: Repeated in Cactus (Paris) 15(66): 9-10, ill., 1960.
Type: Ritter 710 loc. 1 – Chile: Región III; Copiapó: “Depart. Copiapó, near coast” [protologue,
translated] / “Arica” [holotype label at U], 9.1959 [collection date according to specimen at U]
(type herbaria: U 098144B).
B (sheet GH33919) (type number): ic (corp, ar, sp, fl), fl. Notes: Ex cult. BG Berlin 127-85-74-80,
19.6.1995, from material raised from seeds originally received from G. Köhres (without year).
B (sheet GH44751 / B100210950) (type number): ic (corp, ar, sp, fl), fl. Notes: Ex cult. BG
Berlin 127-85-74-80, 9.5.2007, from material raised from seeds originally received from
Köhres (without year). Garden Herbarium GH 44751.
Cleistocactus chacoanus var. santacruzensis F. Ritter, Kakt. Südamer. 2: 672, fig. 687. 1980
Notes: Based on the same type collection as Cleistocactus santacruzensis Backeberg, but incor-
rectly designated as illegitimate by Mottram in Cact. Succ. J. (US) 61: 156. 1989.
Type: Ritter 356 loc. 1 Bolivia: Santa Cruz: “tile roofs of department capital Santa Cruz”
[protologue, translated]; “frequent”, 1954 [collection date according to specimen at U] (type her-
baria: U, SGO, ZSS) [type cited for U in Ritter, Kakt. Südamer. 1: iii. 1979].
B (sheet GH10078) (type number): fl. – Notes: Ex cult. BG Berlin, 5.6.1981, from material origi-
nally raised from seeds received from H. Winter. Collection number established by inference.
B (sheet GH35778) (type number): corp, ar, sp, fr, sem. Notes: Ex cult. BG Berlin
001-85-74-10, 28.5.1997, from material originally received 1957 from H. Winter. Collection
number established by inference.
Cleistocactus micropetalus F. Ritter, Kakt. Südamer. 2: 675, figs. 653, 656. 1980
Type: Ritter 830 loc. 1 Bolivia: Chuquisaca; Avilez: “Concepción” [protologue], 1958 (type
herbaria: U 160684B, SGO [status?], ZSS [seeds only, status?]) [type cited for U in Ritter, Kakt.
Südamer. 1: iii. 1979].
B (sheet GH18323) (type number): fl, ic (corp). – Notes: Ex cult. BG Berlin, 5.7.1985, from ma
-
terial originally raised from seeds received 1959 from H. Winter. Collection number estab
-
lished by inference.
B (sheet GH18323a) (type number): corp, ar, sp. Notes: Ex cult. BG Berlin 027-87-74-80,
25.2.1998, from material originally raised from seeds received 1959 from H. Winter. Collec
-
tion number established by inference.
Cleistocactus palhuayensis F. Ritter & Shahori in Ritter, Kakt. Südamer. 2: 688-689, fig. 669,
673. 1980
Notes: Cleistocactus viridiflorus Backeberg 1963 and Cephalocleistocactus pallidus Backeberg
1966 are based on further individuals of the type collection, but it is open to debate whether these
names are validly described; if so, they would have priority.
Type: Ritter 324 loc. 2 Bolivia: La Paz; Muñecas: “lower Río Palhuaya” [protologue, trans
-
lated], 10.1954 (type herbaria: U [not found], ZSS) [type cited for U in Ritter, Kakt. Südamer. 1:
iii. 1979].
234 Eggli & Leuenberger: Type specimens of Cactaceae names in the Berlin Herbarium
B (sheet GH20171) (type number): fl, ic (corp). Notes: Ex cult. BG Berlin 045-36-74-80,
19.6.1986. Collection number inferred by inference.
B (sheet GH20171a) (type number): corp, ar, sp. Notes: Ex cult. BG Berlin 045-36-74-80,
21.7.1993. Collection number inferred by inference.
B (sheet GH20171b) (type number): cor, ar, sp. – Notes: Ex cult. BG Berlin 045-36-74-80, June
1995. Collection number inferred by inference.
Coleocephalocereus braunii Diers & Esteves in Kakt. And. Sukk. 36(2): 28-35, ill. 1985
Type: Horst, Braun & Esteves 4 – Brazil: Espírito Santo: “W part of the state of Espírito Santo”
[protologue] / “E of Pico da Bandeira” [isotype labels at B, translated]; “on very steep gra
-
nitic-gneissic rocks”, 500-600 m, 9.1983 [detailed locality data deposited with the holotype for
conservation reasons according to the protologue. Collection date according to isotype labels at
B] (type herbaria: B (ex KOELN), B).
B (ex KOELN) (spirit B 810013115) (holotype): corp, ar, sp. Notes: Part 1 of 2 of the holo
-
type. Simply labelled “Holotyp” and “HBE 4”. A separate print of the protologue with anno
-
tated locality data is archived separately.
B (ex KOELN) (spirit B 810013116) (holotype): fl, fr. – Notes: Part 2 of 2 of the holotype. Sim
-
ply labelled “Holotyp” and “HBE 4”.
B (boxed s.n.) (isotype): corp, ar, sp, (ceph).
B (boxed s.n.) (isotype): corp, ar, sp, ceph.
B (boxed s.n.) (isotype): rad, corp, ar, sp, ceph.
B (ex KOELN) (boxed B 100216265) (isotype): sem. – Notes: Simply labelled as “HBE 4.
Notes. – Whereas the holotype is stated to be “Horst, Braun & Esteves 4”, the collection “Braun
470” is given as isotype, with an additional statement that this (and two further topotype collec-
tions) were made simultaneously at the type locality, but actually represents an paratype (see
3.2., above).
Coleocephalocereus diersianus P. J. Braun & Esteves in Kakt. And. Sukk. 39(3): 48-53, ill.,
SEM-ill. 1988
Type: Horst, Braun & Esteves 3 Brazil: Minas Gerais: “frontier region to Espírito Santo” [pro-
tologue, translated]; “on steep ‘sugar loaf’ rock walls with some grasses and Aechmea sp.”,
500 m, 1983 [detailed locality data deposited with the holotype for conservation reasons accord
-
ing to the protologue] (type herbaria: B (ex KOELN)).
B (ex KOELN) (spirit B 810013117) (holotype): ar, sp, fl. Notes: Simply labelled “Holotyp”
and “HBE 3”. A separate print of the protologue with annotated locality data is archived sep
-
arately.
B (ex KOELN) (boxed B 100216266) (isotype): sem. – Notes: Simply labelled “HBE 3”.
Coleocephalocereus estevesii Diers in Kakt. And. Sukk. 29(9): 201-205, ill. 1978
Type: Esteves Pereira 66 – Brazil: Minas Gerais; Mpio. Pedra do Indaiá: “S of Belo Horizonte”
[protologue, translated], 900-1000 m, 1974 [information on municipio added from Taylor &
Zappi (2004)] (type herbaria: B (ex KOELN)).
B (ex KOELN) (boxed B 100216267) (holotype): corp, ar, sp. Notes: Simply labelled as
“Holotyp” and “E-66”. A separate print of the protologue with annotated locality data is ar
-
chived separately.
B (ex KOELN) (boxed B 100216268) (isotype): corp, ar, sp. Notes: Simply labelled as “Iso
-
typ” and E-66”.
B (ex KOELN) (boxed B 100216269) (isotype): sem. – Notes: Simply labelled as “E-66”.
B (ex KOELN) (spirit B 810013118) (isotype): fl. – Notes: Simply labelled as “E-66”.
Copiapoa alticostata F. Ritter in Taxon 12(1): 29. 1963
Type: Ritter 717 loc. 1 Chile: Región III; Huasco: “Typuslokalität nördl. Nicolasa, Nordchile”
Willdenowia 38 – 2008 235
[holotype label at U] = type locality N of Nicolasa, N Chile, 10.1962 [collection date according to
specimen at U] (type herbaria: U 145223B, SGO, ZSS [seeds only]) [holotype cited for U in
Taxon 12(1): 28. 1963].
B (sheet GH43998 / B100171592) (type number): ic (juv corp, ar, sp). Notes: Ex cult. BG
Berlin 138-16-74-80, Sept.1991, from material originally raised from seeds received from H.
Winter (s.a.). Collection number established by inference.
Copiapoa calderana F. Ritter in Cactus (Paris) No.65: 197-198, ill. 1959
Type: Ritter 507 loc. 1 – Chile: Región III; Copiapó: “coast N of Caldera” [protologue, transla
-
ted] / “von der Typuslokalität Caldera” [holotype label at U] = from the type locality Caldera,
1.1956 [originally distributed as Copiapoa pepiniana] (type herbaria: U 98048B [lecto], SGO,
ZSS) [type cited for ZSS, but none of the specimens at ZSS seems to qualify as holotype.
Lectotype designated by Doweld in Sukkulenty 4(1-2): 55. 2001].
B (sheet GH44002 / B100171589) (type collection): ic (corp, ar, sp). Notes: Ex cult. BG
Berlin 138-20-74-80, 26.5.2006, from material originally raised from seed received 1959
from H. Winter.
Copiapoa dura F. Ritter in Taxon 12(1): 31. 1963
Notes: Name first mentioned in Backeberg, Cact. 6: 3827. 1962 (as nom. nud.).
Type: Ritter 546 loc. 1 Chile: Región III; Copiapó: “E of Totoral” [protologue, translated], 1956
(type herbaria: U 117603B, SGO [status?], ZSS [seeds only]) [holotype cited for U in Taxon
12(1): 28. 1962].
B (sheet GH44003 / B100171588) (type number): ic (corp, ar, sp). – Notes: Ex cult. BG Berlin
138-26-74-80, 26.5.2006, from material originally raised from seeds received 1959 from H.
Winter.
Copiapoa krainziana F. Ritter in Taxon 12(1): 30-31. 1963
Notes: First mentioned in Backeberg, Cact. 6: 3834. 1962 (as nom. nud.).
Type: Ritter 210 loc. 1 – Chile: Región II; Antofagasta: “mountains at the coast in Dept. Taltal”
[protologue, translated], 21.6.1954 [collection date according to material at ZSS] (type herbaria:
ZSS T4724, SGO, ZSS).
B (sheet GH30589) (type number): ic (corp). – Notes: Ex cult. BG Berlin, 30.11.1992, from ma
-
terial raised from seeds originally received 1957 from H. Winter. Collection number estab
-
lished by inference.
Copiapoa laui Diers in Kakt. And. Sukk. 31(12): 362-365, ill. 1980
Notes: Sphalm. ‘lauii’.
Type: Lau 891 – Chile: Antofagasta: “in the farther surroundings of Esmeralda” [protologue, trans
-
lated]; “in strongly schistose, humus-free soil or schist grit on slightly sloping mountain slopes”,
100 m, 1971 [detailed locality data deposited with the holotype for conservation reasons accord
-
ing to the protologue, but this is not the case] (type herbaria: B (ex KOELN), ZSS [type number]).
B (ex KOELN) (spirit B 810013119) (holotype): rad, corp, ar, sp, fl. – Notes: Simply labelled as
“Holotyp” and “L 891”. A separate print of the protologue (but without annotations as to more
detailed locality data) is archived separately.
B (ex KOELN) (boxed B 100216270) (isotype): rad, corp, ar, sp. Notes: Simply labelled
“Isotyp” and (Lau) L 891”.
B (ex KOELN) (boxed B 100216271) (isotype): sem. Notes: Simply labelled as “L 891” and
“unbeschädige Früchte 35 Korn, 48 Korn” [undamaged fruits 35 grains, 48 grains], but the
plastic bag contains only 8 seeds.
Copiapoa marginata subsp. robustispina Doweld in Sukkulenty 4(1-2): 49. 2002
Notes: Based on Copiapoa bridgesii sensu F. Ritter 1980.
236 Eggli & Leuenberger: Type specimens of Cactaceae names in the Berlin Herbarium
Type: Ritter 245 loc. 1 Chile: Región III; Chañaral: “mountains N of air strip of Chañaral”
[Ritter, Kakt. Südamer. 3: 1060. 1980, translated], 6.1954 (type herbaria: ZSS T4806).
B (sheet GH11982) (type number): fl, ic (corp). – Notes: Ex cult. BG Berlin, 9.6.1982, from ma
-
terial raised from seeds originally received 1957 from H. Winter. Collection number estab
-
lished by inference.
Copiapoa rubriflora F. Ritter in Taxon 12(1): 31. 1963
Type: Ritter 211 loc. 1 Chile: Región II; Antofagasta: “S of Taltal, along the coast” [protologue,
translated], 1.1954 (type herbaria: ZSS T4727 [lecto], SGO) [lectotype designated by Doweld in
Sukkulenty 4(1-2): 55. 2002 (‘2001’)].
B (sheet GH44001 / B100171590) (type number): ic (corp, ar, sp). – Notes: Ex cult. BG Berlin
138-37-74-80, 26.5.2006, from material originally raised from seeds received 1957 from H.
Winter.
Copiapoa serpentisulcata F. Ritter in Cactus (Paris) 15(66): 22-23, ill. 1960
Type: Ritter 246 loc. 1 Chile: Región III; Chañaral: “Chañaral” [protologue] / “N of Chañaral,
mountain tops” [Ritter, Kakt. Südamer. 3: 1093. 1980], 6.1954 (type herbaria: ZSS T4811, SGO,
ZSS) [type cited for Z, later corrected to ZSS].
B (sheet GH21139) (type number): ic (corp). – Notes: Ex cult. BG Berlin, 30.7.1986, from ma
-
terial raised from seeds originally received 1958 from H. Winter. Collection number estab-
lished by inference.
Coryphantha pirtlei Werdermann in Notizbl. Bot. Gart. Berlin-Dahlem 12: 226. 1934
Type: Pirtle s.n. USA: Texas; Starr County: s.l. [protologue], 1931 (type herbaria: B [authentic
material]) [ex cult. BG Berlin 1938].
B (spirit B 810006854) (type collection): corp, ar, sp. Notes: Ex cult. BG Berlin 1938, and
thus later than the protologue.
Discocactus buenekeri W.-R. Abraham in Kakt. And. Sukk. 38(11): 282-285, ill. 1987
Type: Büneker s.n. in Abraham CWRA27 Brazil: Bahia: “N Bahia” [protologue, standardized];
“in pure quartz sand” [protologue, translated], 1000 m, 1985 (type herbaria: B (ex KOELN)).
B (ex KOELN) (spirit B 810013120) (holotype): corp, ar, sp, ceph. Notes: Labelled as “Ho
-
lotyp”; collection number given as “CWRA 27/1”; “1” most probably relating to 1 of 4 spec
-
imens deposited.
B (ex KOELN) (boxed B 100216272) (isotype): fr. – Notes: Labelled “CWRA 27 (3)”, the fig
-
ure “3” most probably relating to the third of 4 specimens deposited.
B (ex KOELN) (boxed B 100216273) (isotype): sem. Notes: Labelled “CWRA 27 (4)”, the
figure “4” most probably relating to the fourth of 4 specimens deposited.
B (ex KOELN) (spirit B 810013121) (isotype): fl. – Notes: Labelled “CWRA 27 (2)”, the figure
“2” most probably relating to the second of 4 specimens deposited.
Discocactus cangaensis Diers & Esteves in Cact. Succ. J. (Los Angeles) 52(3): 107-111. 1980
Type: Esteves Pereira 79 – Brazil: Goiás: “SW to W region of the state of Goiás” [protologue];
“rocky outcroppings rich in iron mineral, together with some grasses in a gritty soil containing
some humus, full sun, rarely in half-shade”, 750 m, 1975 [detailed locality data deposited with the
holotype for conservation reasons according to the protologue] (type herbaria: B (ex KOELN)).
B (ex KOELN) (boxed B 100216274) (holotype): corp, ar, sp, ceph. Notes: Simply labelled
“Holotyp” and “E-79”. A separate print of the protologue with annotated locality data is ar
-
chived separately.
B (ex KOELN) (boxed B 100216275) (isotype): fr, sem. – Notes: Simply labelled “E-79”.
Willdenowia 38 – 2008 237
B (ex KOELN) (boxed B 100216342) (isotype): corp, ar, sp, ceph. Notes: Simply labelled as
“E-79 (a-c) Drei Isotypen”, but without connection of the letters a-c with the 3 plant bodies
preserved.
B (ex KOELN) (spirit B 810013122) (isotype): fl. – Notes: Simply labelled “E-79”.
Discocactus crassispinus P. J. Braun & Esteves in Cact. Succ. J. (Los Angeles) 66(5): 228-235,
ill., SEM-ill. 1994
Type: Esteves Pereira 44 – Brazil: Goiás: “W to SW regions [of Goiás]” [protologue, standard
-
ized] / “table mountains NE of Rio Verde” [isotype label at B]; “very often on the tops of ta
-
ble-mountains, in hard soil of red sandy loam and gravel of ‘pedra canga’”, 780-800 m, 7.1989
(type herbaria: UFG 12349, B, ZSS).
B (boxed s.n.) (isotype): (rad), corp, ar, sp, ceph. – Notes: Leg. P. J. Braun (ex cult.?), without date.
Discocactus crystallophilus Diers & Esteves in Kakt. And. Sukk. 32(11): 258-261, ill. 1981
Type: Esteves Pereira & Guimarães 84 Brazil: Minas Gerais; Mun. Augusto de Lima: “C part of
Minas Gerais, in the plains E and SE of the Represa Tres Marias” [protologue, translated] / “S of
Curvelo” [isotype label at B, translated]; “in sandy soil with smaller and larger quartz pebbles”,
700 m, 1977 [probably recollected in later years under the same collection number. Municipio
name added from Taylor & Zappi (2004] (type herbaria: B (ex KOELN), B).
B (ex KOELN) (boxed B 100216277) (holotype): corp, ar, sp, ceph. Notes: Labelled “Holo-
typ” and “E-84”. A separate print of the protologue with annotated locality data is archived
separately.
B (boxed s.n.) (isotype): rad, corp, ar, sp, ceph, (fl). – Notes: Leg. P. J. Braun (ex cult. ?), with-
out date.
B (ex KOELN) (boxed B 100216278) (isotype): corp, ar, sp, ceph. Notes: Labelled “E-84 (a,
b)” and “2 Isotypen”. The two specimens are, however, not identified with “a” or “b”.
B (ex KOELN) (boxed B 100216279) (isotype): sem. – Notes: Simply labelled “E-84” and “Sa-
men und Früchte” [seeds and fruits], but not fruits are present.
B (ex KOELN) (spirit B 810013123) (isotype): fl. – Notes: Simply labelled “E-84”.
B (ex KOELN) (spirit B 810013124) (isotype): fl. – Notes: Simply labelled “E-84” and “innen”,
without known significance.
Discocactus diersianus Esteves in Cact. Succ. J. (Los Angeles) 51(4): 179-183, ill. 1979
Type: Esteves Pereira 87 Brazil: Goiás: “SE region of the state of Goiás” [protologue, standard
-
ized]; “together with bromeliads on scaled rocks of granite outcrops interspersed with humus, in
full sunlight”, 650 m, s.a. [detailed locality data deposited with the protologue for conservation
reasons according to the protologue] (type herbaria: B (ex KOELN)).
B (ex KOELN) (boxed B 100216280) (holotype): corp, ar, sp, ceph. Notes: Labelled “Holo
-
typ” and “E-87”. A separate print of the protologue with annotated locality data is archived
separately.
B (ex KOELN) (boxed B 100216281) (isotype): corp, ar, sp, ceph. Notes: Labelled “Isotyp”
and “E-87”.
B (ex KOELN) (boxed B 100216282) (isotype): fr, sem. – Notes: Simply labelled “E-87”.
B (ex KOELN) (spirit B 810013125) (isotype): fl. – Notes: Simply labelled “E-87”.
Discocactus estevesii Diers in Cact. Succ. J. (Los Angeles) 50(2): 83-85, 94, ill. 1978
Type: Esteves Pereira 2 Brazil: Goiás: “valley of the Paranaiba river, SE region of Goiás” [pro
-
tologue]; “clay soil with stones between grasses on small rises”, 650 m, s.a. [detailed locality data
deposited with the holotype for conservation reasons according to the protologue] (type herbaria: B
(ex KOELN), ZSS).
238 Eggli & Leuenberger: Type specimens of Cactaceae names in the Berlin Herbarium
B (ex KOELN) (boxed B 100216283) (holotype): corp, ar, sp, ceph. Notes: Labelled “Holo
-
typ” and “E-2”. A separate print, which contrary to the protologue is without annotated lo
-
cality data, is archived separately.
B (ex KOELN) (boxed B 100216284) (isotype): corp, ar, sp, ceph. Notes: Labelled “2 Iso
-
typen” and E-2 (a-c)”, but without connection of the lower-case letters to specimen parts.
B (ex KOELN) (boxed B 100216285) (isotype): sem. – Notes: Simply labelled “E-2”.
B (ex KOELN) (spirit B 810013126) (isotype): fl. – Notes: Simply labelled “E-2”.
Discocactus goianus Diers & Esteves in Kakt. And. Sukk. 31(3): 73-79, ill., SEM-ill. 1980
Type: Esteves Pereira 10 Brazil: Goiás: “C and S part of the state of Goiás” [protologue, trans
-
lated] / “S of Goiánia” [isotype label at B]; “together with bromeliads on sunny rocky places near
rivers”, 650-700 m, s.a. (type herbaria: B (ex KOELN), B).
B (ex KOELN) (boxed B 100216286) (holotype): corp, ar, sp, ceph. Notes: Labelled “Holo
-
typ” and “E-10”. A copy of the protologue with annotated locality data is archived separately.
B (boxed s.n.) (isotype): corp, ar, sp, ceph, (fl). – Notes: Leg. P. J. Braun (ex cult. ?), without
date.
B (ex KOELN) (boxed B 100216287) (isotype): corp, ar, sp, ceph. Notes: Labelled “Isotyp”
and “E-10”.
B (ex KOELN) (boxed B 100216288) (isotype): sem. – Notes: Simply labelled “E-10”.
B (ex KOELN) (spirit B 810013127) (isotype): fl. Notes: Simply labelled “E-10” and “E 10
V”, without known significance of the second annotation.
Discocactus latispinus Buining & Brederoo in Succulenta (Amsterdam) 56(11): 259. 1977
Notes: Again given as new species in Buining, Gatt. Discocactus: 95-100, ill. 1980.
Type: Horst & Uebelmann HU 146 – Brazil: Minas Gerais: “Serra do Cabral” [protologue], s.a.
[collection cited as “H 146” in the protologue] (type herbaria: U, ZSS).
B (boxed s.n.) (type number): corp, ar, sp, ceph. – Notes: Leg. P. J. Braun ex cult. without date,
accessioned 1991. This is very probably a later re-collection of the taxon under the original
collection number, and thus not part of the original introduction (pre-1977).
Discocactus lindanus Diers & Esteves in Cact. Succ. J. (Los Angeles) 53(2): 56-60, ill. 1981
Notes: Sphalm. ‘lindaianus’, and mandatorily correctable under Art. 60.11, Rec. 60C(d).
Type: Esteves Pereira 9 Brazil: Goiás: “E to NE region of the state of Goiás” [protologue] /
“Chapada dos Veadeiros, campo cerrado” [isotype label at B]; “on broken scaly flat rocks, inter
-
spersed with humus, sand and fine gravel”, 1100-1200 m, 1974 [probably recollected under the
same collection number in later years] (type herbaria: B (ex KOELN), B).
B (ex KOELN) (boxed B 100216289) (holotype): corp, ar, sp, ceph. Notes: Labelled as “Holo
-
typ”, collection number given as “E-9”. A copy of the protologue with annotated locality data
is archived separately.
B (boxed s.n.) (isotype): corp, ar, sp, ceph. Notes: Leg. P. J. Braun (ex cult.?), without date,
accessioned 1990.
B (ex KOELN) (boxed B 100216290) (isotype): corp, ar, sp, ceph. – Notes: Originally labelled
“E-9 (a-c)” and “3 Isotypen”, but the letters are not associated with the 3 plant bodies mak
-
ing up the specimen.
B (ex KOELN) (boxed B 100216291) (isotype): sem. – Notes: Collection given as “E-9” on the
original label.
B (ex KOELN) (spirit B 810013128) (isotype): fl. Notes: Collection data given as “E-9” on
the original label.
B (ex KOELN) (spirit B 810013129) (isotype): fl. – Notes: Collection number given as “E-9” on
the original label.
Willdenowia 38 – 2008 239
Discocactus multicolorispinus P. J. Braun & Brederoo in Kakt. And. Sukk. 32(3): 54-59, ill.
1981
Notes: Name doubtfully valid because the type collection is only specified in a very general man
-
ner and without indication of the collector.
Type: Bueneker s.n. Brazil: Minas Gerais: “near a small village on the W slope of the Serra de
Minas in the Serra do Espinhaço” [protologue, translated]; “under low shrubs and hidden in very
stony, quartzite-rich soil”, 1000-1100 m, 1977 [collector name and collection year according to
isotype label at B. Detailed locality date deposited with the holotype for conservation reasons ac
-
cording to the protologue] (type herbaria: U, B, ZSS).
B (boxed s.n.) (isotype): corp, ar, sp, ceph, (fl). Notes: Leg. P. J. Braun (ex cult. ?) without
date; accessioned 1991.
Discocactus prominentigibbus Diers & Esteves in Kakt. And. Sukk. 39(1): 14-19, ill., SEM-ill.
1988
Type: Esteves Pereira 151 – Brazil: Goiás: “between the rivers Rio Tocantins and Rio Araguaia
in the central, somewhat northern region of Goias” [protologue, translated]; “in half shade of
trees and shrubs, or in full sun, in coarse sandy-gravelly soil”, 800 m, s.a. [detailed locality data
deposited together with the holotype for conservation reasons according to the protologue] (type
herbaria: B (ex KOELN)).
B (ex KOELN) (boxed B 100216292) (holotype): corp, ar, sp, ceph. Notes: Labelled “Holo
-
typ” on the original label. Collection data simply given as “E-151”. A separate print with an-
notated locality data is archived separately.
B (ex KOELN) (boxed B 100216293) (isotype): corp, ar, sp, ceph. Notes: Labelled “Isotyp”
on the original label. Collection details simply given as “E-151”. The specimen consists of 4
entire plant bodies.
B (ex KOELN) (boxed B 100216294) (isotype): fr, sem. Notes: Collection details simply
given as “E-151”. With an additional label “Samen + Früchte (leer)”.
B (ex KOELN) (spirit B 810013130) (isotype): fl. Notes: Collection details simply given as
“E-151”.
B (ex KOELN) (spirit B 810013131) (isotype): fl. Notes: Collection details simply given as
“E-151”.
Discocactus pseudolatispinus Diers & Esteves in Kakt. And. Sukk. 38(10): 242-247, ill., SEM-ill.
1987
Type: Esteves Pereira 111 Brazil: Minas Gerais: “central-northern Minas Gerais” [protolo
-
gue] / “Montes Claros - Brasilia, at the artificial lake Tres Marias, campo cerrado” [isotype label
at B], 700-800 m, s.a. [detailed locality information deposited with the holotype for conservation
reasons according to the protologue] (type herbaria: B (ex KOELN), B).
B (ex KOELN) (boxed B 100216295) (holotype): corp, ar, sp, ceph. Notes: Labelled “Holo
-
typ” on the original label. Collection data simply given as “E-111”. A separate print with an
-
notated locality information is archived separately.
B (boxed s.n.) (isotype): corp, ar, sp. Notes: Leg. P. J. Braun ex cult. without year; acces
-
sioned 1990.
B (ex KOELN) (boxed B 100216296) (isotype): fr, sem. – Notes: Collection data simply given
as “E-111”.
B (ex KOELN) (spirit B 810013132) (isotype): fl. Notes: Collection data simply given as
“E-111”.
Discocactus pulvinicapitatus Buining & Brederoo, Gatt. Discocactus: 100-105, ill. 1980
Type: Horst & Uebelmann HU 425 Brazil: Minas Gerais: “near the Rio Jequitai” [protologue] /
“E of Jequitai” [isotype label at B] / “20 km E of Engenheiro Omar, Rio Jequitai” [unrecorded
source], 650 m, s.a. [collection cited as “H 425” in the protologue] (type herbaria: U, B [type
number], ZSS [type number]).
240 Eggli & Leuenberger: Type specimens of Cactaceae names in the Berlin Herbarium
B (boxed s.n.) (type number): corp, ar, sp, ceph, (fl). – Notes: Leg. P. J. Braun (ex cult.?), with
-
out year; accessioned 1991. Probably a later re-collection under the same field number.
B (boxed s.n.) (type number): corp, ar, sp, ceph, (fl). – Notes: Leg. P. J. Braun (ex cult.?), with
-
out year; accessioned 1991. Probably a later re-collection under the original field number.
Discocactus subterraneo-proliferans Diers & Esteves in Kakt. And. Sukk. 31(9): 266-271, ill.
1980
Type: Esteves-Pereira 12 Brazil: Goiás: “central region of the Araguaia valley in the W of the
state of Goiás” [protologue, translated]; “below scattered small trees and shrubs on flat ground in
fine, loose predominantly quartzitic sand with little humus”, 200-250 m, 10.1973 (type herbaria:
B (ex KOELN)).
B (ex KOELN) (boxed B 100216297) (holotype): corp, ar, sp, ceph. Notes: Labelled “Holo
-
typ” on the original label; collection data simply given as “E-12”. A separate print of the
protologue with annotated locality data is archived separately.
B (ex KOELN) (boxed B 100216298) (isotype): corp, ar, sp, ceph. Notes: Labelled “Isotyp”
on the original label; collection data given as “E-12 (b)”, which rises doubt as to the correct
-
ness of the isotype statement. The specimen consists of 2 plant bodies.
B (ex KOELN) (boxed B 100216299) (isotype): fr, sem. – Notes: Collection data simply given
as “E-12”.
B (ex KOELN) (spirit B 810013133) (isotype): fl. Notes: Collection data simply given as
“E-12”.
Discocactus zehntneri var. horstiorum P. J. Braun in Succulenta (Amsterdam) 69(10): 218, ill.
1990
Type: Horst 667 Brazil: Bahia: “N Bahia, high mountains (c.1000 m), in manganese ore”
[protologue, translated] / “Mimoso, high mountains, between mangan ore, caatinga” [isotype la-
bels at B], 1000 m, 1988 (type herbaria: B, B, ZSS (ex KOELN), K).
B (boxed s.n.) (isotype): corp, ar, sp.
B (boxed s.n.) (isotype): (rad), corp, ar, sp, ceph.
B (ex KOELN) (boxed B 100216300) (isotype): corp, ar, sp, ceph. – Notes: Labelled as “Isotyp”
on the original label. Locality data simply given as “Bahia”.
Echinocactus anisitsii K. Schumann, Blüh. Kakt.: t. 4. 1900
Type: Anisits Cact. 21 Paraguay: “at the river Tagatiya-mi” [protologue, translated], 25.1.1898
(type herbaria: B).
B (spirit B 810006860) (holotype): corp, ar, sp, fl, fr, [+ic]. – Notes: Original label: Echinocac
-
tus anisitsii K. Sch. Paraguay leg. Anisits”, i.e. without the collection number that is given in
the protologue, but the contents are compatible with the protologue.
Echinocactus aurantiacus Vaupel in Bot. Jahrb. Syst. 50 (Beiblatt 111): 23-24. 1913
Notes: Abridged version published in Monatsschr. Kakteenk. 24(12): 169-170. 1914.
Type: Weberbauer 3846 Peru: Cajamarca: “near San Pablo” [protologue, translated]; “on
rocks”, 2200-2400 m, 29.4.1904 (type herbaria: B).
B (spirit B 810006865) (holotype): rad, corp, ar, sp, fl.
Echinocactus damsii K. Schumann, Gesamtbeschr. Kakt., Nachtr.: 119-120, ill. 1903
Type: Anisits s.n. – Paraguay: “in the N part” [protologue, translated], s.a. (type herbaria: B).
B (spirit B 81006872) (holotype): rad, corp, ar, sp. Notes: Original label: “Echinocactus
damsii K. Sch. Paraguay Anisits n. 21”. The collection number (“n. 21”) has probably been
added and is in error, as Anisits Cact. 21 is the type of Echinocactus anisitsii.
Willdenowia 38 – 2008 241
Echinocactus grandis Rose in Contr. U.S. Natl. Herb. 10(3): 126. 1909
Type: Pringle 6696 Mexico: Puebla: “limestone hills near Tehuacan, 6500 ft.” [original printed
label], 2000 m, 7.8.1897 (type herbaria: B, ENCB, G, HBG, LE, US, Z).
B (sheet s.n.) (isotype): corp, ar, sp, fl. – Notes: Specimen received from E.
Echinocactus hastifer Werdermann & Bödeker in Notizbl. Bot. Gart. Berlin-Dahlem 11(104):
274-276. 1931
Type: Anonymus s.n. Mexico?: cultivated material without known wild origin, but presumably
Mexico., s.a. (type herbaria: B) [ex cult. BG Berlin, June 1930].
B (spirit B 810006882) (holotype): fl. – Notes: Ex cult. BG Berlin, June 1930.
Echinocactus molendensis Vaupel in Bot. Jahrb. Syst. 50 (Beiblatt 111): 24-25. 1913
Notes: Abridged version published in Monatsschr. Kakteenk. 24(12): 170-171. 1914.
Type: Weberbauer 1506 Peru: Arequipa: “Mollendo” [protologue]; “on stony sandy ground
with poor vegetation” [protologue, translated], 20 m, 3.10.1902 (type herbaria: B).
B (spirit B 810006890) (holotype): rad, corp, ar, sp. – Notes: There is a second specimen at B (B
810006891, dried out and useless) numbered “Weberbauer 1806”, which is presumably a
mis-print for the type number.
Echinocactus myriacanthus Vaupel in Bot. Jahrb. Syst. 50 (Beiblatt 111): 25-26. 1913
Notes: Abridged version published in Monatsschr. Kakteenk. 24(12): 171. 1914.
Type: Weberbauer 4272 Peru: Amazonas; Chachapoyas: “at the wall of the valley of the
Marañón above Balsas” [protologue, translated]; “in grass steppe with few cacti”, 2200 m,
25.6.1904 (type herbaria: B).
B (spirit B 810006893) (holotype): rad, corp, ar, sp, fl.
Echinocactus nidulans Quehl in Monatsschr. Kakteenk. 21: 119. 1911
Type: Anonymus ex hort. Quehl s.n. Mexico?: s.l., s.a. (type herbaria: B [fragmentary flower])
[ex cult. hort. Quehl].
B (spirit B 810006896) (holotype): fl. – Notes: Labelled “Quehl Original 12. Mai 1911”. Dried out.
B (spirit B 810006895) (type collection): fl. – Notes: Labelled as “ded. Quehl 17.2.”, but with
-
out year. Dried out.
Echinocactus peruvianus K. Schumann, Gesamtbeschr. Kakt., Nachtr.: 113-114. 1903
Type: Weberbauer s.n. Peru: Lima: “in the Andes above Lima” [protologue, translated]; “to
-
gether with Opuntia vestita”, 4000 m, s.a. (type herbaria: B).
B (spirit B 810006901) (holotype): rad, corp, ar, sp, fl.
Echinocactus reichei K. Schumann, Gesamtbeschr. Kakt., Nachtr.: 110-111. 1903
Type: Reiche s.n. Chile: s.l. [protologue], s.a. (type herbaria: B) [ex cult. BG Berlin, 18.7.1903].
B (spirit B 810006906) (holotype): corp, ar, sp. – Notes: Labelled as “aus Chili von Reiche”.
Echinocactus umade ave Fric ex Werdermann in Monatsschr. Deutsch. Kakteen-Ges. 3: 223-
226, ill. 1931
Notes: Based on Friesia umadeave Fric (nom. nud.) in Möllers Deutsche Gärtn.-Zeitung 45(4):
42-44. 1930.
Type: Stümer s.n. Argentina: Salta/Jujuy: “Salta & Jujuy”, s.l. [protologue], 1930 (type her
-
baria: B [lecto]) [lectotype designated by Kattermann, Succ. Pl. Res. 1: 136. 1994, but see below]
B (spirit B 810006918) (isotype): fl. – Notes: Ex cult. BG Berlin, without year. Labelled “Origi
-
nal zur Blütenbeschreibung” [original of the flower description]. Dried out.
242 Eggli & Leuenberger: Type specimens of Cactaceae names in the Berlin Herbarium
B (spirit B 810006917) (lectotype): corp, ar, sp. – Notes: Labelled “1931”, which could signify
that the specimen was received 1931, or that it was prepared from cultivation in 1931. Dried
out. Lectotype (here designated) of Echinocactus umadeave.
Notes. The name is based on 2 syntypes, Ritter s.n. and Stümer s.n. Kattermann (1994) se
-
lected the Stümer material as lectotype, but did not indicate which of the 2 specimens present at
B is the actual lectotype. While Kattermann’s selection of Stümer s.n. as lectotype of the name
can stand, it needs to be refined (ICBN Art. 9.14). We select here the specimen B810006917 as
lectotype of Echinocactus umadeave.
Echinocactus violaciflorus Quehl in Monatsschr. Kakteenk. 22: 102. 1912
Type: Anonymus s.n. in De Laet s.n. Mexico: Zacatecas: s.l. [protologue], s.a. (type herbaria: B
[status?, useless fragment]).
B (spirit B 810006919) (type collection): fl. Notes: Labelled “ded. Quehl” but without date.
Dried out and completely useless.
Echinocactus vorwerkianus Werdermann in Backeberg, Neue Kakteen: 101. 1931
Notes: See also Repert. Spec. Nov. Regni Veg. 30: 65. 1932 (Latin diagnosis). Leuenberger
(Willdenowia 8: 631. 1979) listed a specimen at B as type, which could be seen as implicit
(lecto-) typification. However the specimen consists of a flower only, and since the protologue
has only a description of the plant body, but not of the flower, this specimen must be posterior to
the publication of the taxon and can thus not serve as lectotype.
Type: Anonymus s.n. in Vorwerk s.n. – Colombia: “near Sagomosa [sic] NE of Bogotá” [proto-
logue, translated]; “cattle pasture”, 1200 m, s.a. (type herbaria: B†, B [“clono”]) [ex cult. BG
Berlin, without date].
B (spirit B 810006920) (type collection): fl. – Notes: Dried out. ex cult. BG Berlin, without year.
Echinocactus weberbaueri Vaupel in Bot. Jahrb. Syst. 50 (Beiblatt 111): 26-27. 1913
Notes: Abridged version published in Monatsschr. Kakteenk. 24(12): 172. 1914.
Type: Weberbauer 4271 Peru: Amazonas; Chachapoyas: “at the E wall of the valley of the
Marañón, above Balsas” [protologue, translated]; “in grass steppe, sometimes with and some-
times without shrubs, together with Echinocactus myriacanthus (the only other cactus ob
-
served)”, 2000-2100 m, 25.6.1904 (type herbaria: B).
B (spirit B 810006921) (holotype): rad, corp, ar, sp, fl.
Echinopsis backebergii Werdermann in Backeberg, Neue Kakteen: 84, ill. 1931
Notes: See also Repert. Spec. Nov. Regni Veg. 30: 64. 1932 (Latin diagnosis); non Echinopsis
backebergii Fric (nom. inval.).
Type: Backeberg s.n. Bolivia: La Paz: s.l. [protologue], s.a. (type herbaria: B [†; photo]; [lecto,
icon]: Backeberg, Neue Kakteen: ill. p. 83 upper right corner) [lectotype (designated by M.
Hjertson in Cactaceae Syst. Init. 20: 14-15. 2005). Details about the typifications are vague in the
protologue but are clarified in the 1932-publication].
B (sheet s.n.) (isolectotype): ic (corp, ar, sp). Notes: Print and negative. Labelled as “Lob.
backebergii (Werd.) Typ!” in Werdermann’s handwriting. This photograph differs from that
published in the protologue.
Notes. Echinopsis backebergii was based on two collections, viz. material collected by
Backeberg (as indicated by the asterisk that precedes the name, cf. p. 63 with the explanation of
the symbols used) and a herbarium specimen (presumably part of loans Werdermann received
from S during 1928-30, perished in the 1943-fire at B) collected earlier by Dr Asplund (without
number). The two collections must be regarded as syntypes. This and all the other names pub
-
lished as new in Backeberg’s book were again covered by Werdermann (1932), giving Latin di
-
agnoses and sometimes additional information. For E. backebergii, Werdermann now clearly
Willdenowia 38 – 2008 243
gives the collection of Asplund (Asplund 623, 1920) as type, and this is to be regarded as
lectotypification. Because Hjertson (2006) could not find any material of Asplund 623, he desig
-
nated the illustration in Backeberg (1931b: 83, upper right) as lectotype. This habitat photo
-
graph by Backeberg shows a flowering plant in habitat and so Hjertson implicitely selected the
Backeberg syntype as new lectotype. B has a photograph of a cultivated plant of this accession.
Echinopsis calochlora subsp. glaetzleana P. J. Braun & Esteves in Kakt. And. Sukk. 45(8): 158-
161, ill., SEM-ill. 1994
Type: Braun 236 – Brazil: Mato Grosso do Sul: “S Pantanal, NW of Bonito” [protologue, trans
-
lated]; “on rocks” [isotype label at B, translated], 7.1983 (type herbaria: ZSS, B).
B (boxed s.n.) (isotype): corp, ar, sp.
Echinopsis chrysantha Werdermann in Notizbl. Bot. Gart. Berlin-Dahlem 11(104): 264-265.
1931
Type: Anonymus s.n. Argentina: “Prov. Los Andes?”, s.l. [protologue], s.a. (type herbaria: B [†;
photo]; [lecto, icon]: Gartenflora 80: 302, ill. 1931) [ex cult. BG Berlin; lectotype designated by
Hunt & Taylor in Cactaceae Syst. Init. 21: 7. 2006].
B (sheet s.n.) (type collection): ic (corp, ar, sp, fl). Notes: Print and negative. Labelled on
non-original labels “ohne Sammler und Datum, lebend, cult. hort. Berol.”, “Foto des Typus”.
The photograph depicts Echinopsis chrysantha (left plant) and E. hossei (right plant). The
same plant of E. chrysantha is also illustrated with a colour photograph in Blüh. Kakt.: t. 18,
Oct. 1931.
Echinopsis chrysochete Werdermann in Repert. Spec. Nov. Regni Veg. 39: 271-272. 1936
Type: Marsoner s.n. Argentina: Jujuy: s.l. [protologue], 2800 m, 1934 (type herbaria: B [†;
photo]) [ex cult. BG Berlin].
B (sheet s.n.) (type collection): ic (corp, ar, sp). Notes: 2 prints and negative. Labelled on a
non-original label “Lebend, cult. Bot. G.”, and on an original label “gepfropfte Köpfe und
getrocknete Import Typ” in Werdermann’s handwriting. According to H. Blossfeld (pers.
comm. to BEL, 2.8.1979), the type was collected by Marsoner but was also distributed under
the number Blossfeld 6.
Echinopsis densispina Werdermann in Kakteenkunde 8(8): 142-143, ill. 1934
Type: Stümer 222 – Argentina/Bolivia: “N Argentina or adjacent Bolivia”, s.l. [protologue], s.a.
(type herbaria: B [fragment]) [ex cult. BG Berlin, 3.10.1932].
B (spirit B 810006963) (holotype): fl. – Notes: Dried out. ex cult. BG Berlin, 3.10.1932.
Echinopsis hossei Werdermann in Notizbl. Bot. Gart. Berlin-Dahlem 11(104): 265-267. 1931
Type: Hosseus s.n. Argentina: “wahrscheinlich Prov. Cordoba und Los Andes”, s.l. [protolo
-
gue], 1929 (type herbaria: B [fragmentary flower]) [ex cult. BG Berlin, 3.6.1931].
B (spirit B 810006939) (type collection): fl. – Notes: Dried out. ex cult. BG Berlin, 3.6.1931.
B (sheet s.n.) (type collection): ic (corp, ar, sp, fl). – Notes: Labelled “lebend, 1 Exemplar ein
-
geführt, cult. Bot. G., 1930 blühend” [living, 1 individual imported, cultivated Bot. Garden,
flowering 1930], “Foto der Typpflanze” [photograph of the type plant].
Echinopsis marsoneri Werdermann in Repert. Spec. Nov. Regni Veg. 30: 55-56. 1932
Notes: Name also used (nom. inval., Art. 32.1c) by A. V. Fric, Cat. “Akklimatisations- und Ver
-
suchsgarten 1932-33”, [unpaged]. 1933.
Type: Stümer s.n. Argentina: “N Argentina: Gobern. Los Andes” [protologue], 1931 (type her
-
baria: B [†; photo]) [ex cult. BG Berlin c.1932].
244 Eggli & Leuenberger: Type specimens of Cactaceae names in the Berlin Herbarium
B (sheet s.n.) (type collection): ic (corp, ar, sp, fl). Notes: Labelled on a non-original label
“Lebend, cult. hort. Berol. Foto der Typpflanze” [living, cult. Berlin Garden, photograph of
the type plant], “farbige Abb. der Typpfl. bei Werdermann, Blüh. Kakt. t. 26, 1932” [coloured
illustration of the type plant], and in Werdermann’s handwriting “Lob. marsoneri Werd.
Typ!”. The published colour photograph differs from the b/w negative but without doubt illus
-
trates the same individual.
Echinopsis mistiensis Werdermann & Backeberg in Backeberg, Neue Kakteen: 84-85, ill. 1931
Type: Backeberg s.n. – Peru: Arequipa: “volcano Misti near Arequipa” [protologue, translated],
3500 m, s.a. (type herbaria: B†, B [photo of authentic material]) [described from living material].
B (sheet s.n.) (type collection): ic (corp, ar, sp). – Notes: Photograph (negative and print) label
-
led “Lobivia vom Misti Import Bbg.1931”, “Backeberg s.n., lebend, cult. hort. Bot. Berol.”,
“Foto der Typpflanze?”, and thus most probably authentic material.
Echinopsis rhodotricha K. Schumann in Monatsschr. Kakteenk. 10: 147-148. 1900
Type: Anisits Cact. 25 Paraguay: “near the Arroyo La Cruz, 6 km from the now deserted S. Sal
-
vador, at the Río Tagatiya-mi” [protologue, translated], 25.1.1898 (type herbaria: B [lecto]).
B (spirit B810006946) (isolectotype): corp, ar, sp, fl, fr, sem, +ic.
B (spirit B810006945) (lectotype): corp, ar, sp, fl, fr, sem, +ic. – Notes: Lectotype (here desig
-
nated) of Echinopsis rhodotricha. In addition, B has a spirit specimen of the remaining syn
-
type, Anisits Cact.20 (B810006944; corp, ar, sp, (fl, fr), +ic).
Notes. – This name is based on 2 syntypes, Anisits Cact. 20 (1 spirit specimen present at B) and
Anisits Cact. 25 (2 spirit specimens present at B), both from the same locality. All of them agree
with the protologue and we here select the most complete specimen of Anisits Cact. 25 as
lectotype.
Echinopsis scoparia Werdermann in Kakteenkunde 8(8): 144-145, ill. 1934
Type: Stümer s.n. – Argentina: s.l. [protologue], s.a. (type herbaria: B [†; photo]) [ex cult. hort.
Hahn, Lichterfelde, c.1934].
B (sheet s.n.) (type collection): ic (corp, ar, sp, fl). – Notes: Labelled “Lob. scoparia Werd. Typ”
in Werdermann’s handwriting, and on a non-original label, copied from the protologue, “Stümer
s.n. lebend in Sammlung Hahn, Lichterfelde. Foto der Typpflanze” [Stümer s.n., living in the
collection Hahn, Lichterfelde. Photograph of the type plant].
Echinopsis silvatica F. Ritter in Succulenta (Amsterdam) 44(2): 24. 1965
Type: Ritter 782 loc. 1 Bolivia: Tarija; O’Connor: “Tacuarandi” [protologue], s.a. (type her
-
baria: U 160265B, SGO [status ?], ZSS [seeds only]).
B (sheet GH15002) (type number): fl, ic (corp). – Notes: Ex cult. BG Berlin, 9.8.1983, from ma
-
terial raised from seeds originally received 1959 from H. Winter. Collection number estab
-
lished by inference.
Echinopsis violacea Werdermann in Notizbl. Bot. Gart. Berlin-Dahlem 11(104): 262-263. 1931
Type: Stümer s.n. Argentina: Córdoba: s.l. [protologue]; “in rock fissures on grassy slopes”
[protologue, translated], 1000 m, 1930 (type herbaria: B [fragmentary flower]) [ex cult. BG
Berlin 1931].
B (spirit B 810006684) (holotype): fl. – Notes: Dried out. ex cult. BG Berlin 1931.
Epiphyllum chrysocardium Alexander in Cact. Succ. J. (Los Angeles) 28(1): 3-6, ill. 1956
Type: MacDougall A. 198 Mexico: Chiapas: “region known as the Selva negra, between
Tapalapa and Blanca Rosa near Santa Lucía and Rio Napak” [protologe]; “tropical rain-forest”,
9.2.1951 [collection number not cited in the original publication and here inserted from the speci
-
Willdenowia 38 – 2008 245
men at G. Distributed as Univ. California Bot. Gard. 54.1242. Locality given as “Pueblo Nuevo,
Blanca Rosa” by Kimnach, MacDougall collections by name, by locality, by collection number,
1992] (type herbaria: B, G, NY, ZSS).
B (sheet s.n.) (isotype): corp, ar, fl.
Eriocereus spinosissimus Buining & Brederoo in Kakt. And. Sukk. 28(3): 49-51, ill. 1977
Type: Horst & Uebelmann HU 328 Brazil: Mato Grosso: “oberhalb Cuiabá” [protologue],
“30 km towards Chapada de Guimarães, at the mountain” [collectors, pers. comm], s.a. (type her
-
baria: U, B [“clono”]).
B (spirit B 810000104) (type collection): fl. – Notes: Leg. W. Schindhelm ex cult., without date,
accessioned c.1984.
B (sheet GH31420) (type collection): fl. Notes: Ex cult. BG Berlin 225-01-85-30, 2.7.1993.
Material originally from W. Schindhelm, acquired 1970 from W. Uebelmann.
B (sheet GH37443) (type collection): fl. Notes: Ex cult. BG Berlin 225-01-85-30, 5.8.1999.
Material originally ex cult. W. Schindhelm, acquired 1970 from W. Uebelmann.
B (sheet s.n.) (type collection): fl, ic (corp, ar, sp, fl). Notes: Leg. W. Schindhelm ex cult.,
“Uebelmann 1971 Importe”. Accessioned 24.8.1984.
Espostoa lanata subsp. roseiflora J. E. Madsen & Z. Aguirre in Nordic J. Bot. 23(1): 24-25, ill.
2004
Type: Madsen & Galvez 7444 Ecuador: Loja: “La Toma – El Tambo, km 5-11 (along gravel
roads), vicinity of Río Catamayo” [protologue]; “dry rocks with cacti”, 1300-1500 m, 15.10.2000
[Shrubby or arborescent with age, to 5 m high; trunk always evident; flowers 4.5-5 cm long; floral
tube and petaloids purple; fruits (one seen) club-shaped, 3 cm wide, 2.5 cm long, caducous (col-
lected on the ground), dark purple at the apex, light purple at the base, not crowned by floral rem-
nants] (type herbaria: AAU, B, K, LOJA, MO, QCA, QCNE, S, ZSS).
B (sheet s.n.) (isotype): corp, ar, sp, ceph, fl, sem. – Notes: J. Madsen.
Facheiroa estevesii P. J. Braun in Kakt. And. Sukk. 37(4): 74-79, ill., SEM-ill. 1986
Type: Esteves Pereira 186 – Brazil: Bahia: “SW part of the State of Bahia” [protologue] / “Bahia,
E Rio São Francisco” [holotype label] / “S of Malhada” [isotype label at B, standardized]; “lime
-
stone rocks” [protologue], 500-550 m, 1984 [collection date according to isotype label at B.
Detailed locality data deposited with the holotype for conservation reasons according to the
protologue, but no data were found with the holotype] (type herbaria: B (ex KOELN), B, K, UFG,
ZSS) [isotypes at K and UFG according to Taylor & Zappi, Cacti Eastern Brazil: 432. 2004].
B (ex KOELN) (boxed B 100216301) (holotype): corp, ar, sp, ceph. Notes: Labelled “Holo
-
typ” and representing part 1 of 2 of the holotype.
B (ex KOELN) (spirit B 810013134) (holotype): al, fr. – Notes: Labelled as “Holotyp” and rep
-
resenting part 2 of 2 of the holotype.
B (boxed s.n.) (isotype): corp, ar, sp. – Notes: Leg. P. J. Braun (ex cult.?).
Facheiroa tenebrosa P. J. Braun & Esteves in Kakt. And. Sukk. 39(6): 126-131, ill., SEM-ill.
1988
Type: Esteves Pereira 183 Brazil: Bahia: “in the SW frontier region between the states of Minas
Gerais and Bahia” [protologue, translated] / “S Serra do Ramalho” [isotype label at B]; “isolated
limestone rocks” [protologue], 550-600 m, 1984 [collection year according to the isotype label at
B. Detailed locality data deposited together with the holotype for conservation reasons according
to the protologue, but no such data were found] (type herbaria: B (ex KOELN), B, K, UFG, ZSS)
[isotypes at K and UFG according to Taylor & Zappi, Cacti Eastern Brazil: 431. 2004].
B (ex KOELN) (boxed B 100216302) (holotype): corp, ar, sp, ceph. – Notes: Labelled as “Holo
-
typ” on the original label, and representing part 1 of 3 of the holotype; locality simply given
as “Bahia”.
246 Eggli & Leuenberger: Type specimens of Cactaceae names in the Berlin Herbarium
B (ex KOELN) (spirit B 810013135) (holotype): corp, ar, sp, ceph. – Notes: Labelled “Holotyp”
on the original label, and representing part 2 of 3 of the holotype. Collection data simply
given as “E-183”.
B (ex KOELN) (spirit B 810013136) (holotype): fl. Notes: Labelled “Holotyp” on the original
label and representing part 3 of 3 of the holotype. Collection details simply given as “Esteves
183”.
B (boxed s.n.) (isotype): corp, ar, sp, ceph. – Notes: Leg. P. J. Braun (ex cult.?).
Frailea fulviseta Buining & Brederoo in Kakt. And. Sukk. 24(8): 170-172, ill. 1973
Type: Horst & Uebelmann HU 352 – Brazil: Rio Grande do Sul:
”São Francisco de Assis” [protologue], 100-200 m, s.a. [locality given more precisely as
“Tafelberg 5 km N of São Francisco de Assis” by W. Uebelmann (pers. comm.1990)] (type her
-
baria: U).
B (boxed s.n.) (type number): corp, ar, sp. – Notes: Leg. P. J. Braun (ex cult.?). Although the orig
-
inal label says “1972: Brasilien, Rio Grande do Sul, São Francisco de Assis, Typstandort”,
there is some doubt whether this indeed represents the original collection, or a later re-collec
-
tion under the original number.
Frailea m ammifera Buining & Brederoo in Krainz, Kakt., Lief. 50-51: C VIe, ill. 1972
Notes: Publication year sometimes erroneously cited as 1973.
Type: Horst & Uebelmann HU 345 Brazil: Rio Grande do Sul: “N of Dom Pedrito” [protologue,
translated], 250 m, s.a. (type herbaria: ZSS).
B (boxed s.n.) (type number): rad, corp, ar, sp. – Notes: Leg. P. J. Braun (ex cult.?); original la-
bel “Brasilien, Rio Grande do Sul, Typstandort”, and most probably a later re-collection un-
der the original number.
Gymnocalycium ambatoense Piltz in Kakt. And. Sukk. 31(1): 10-13, ill. 1980
Type: Piltz 22/5 Argentina: Catamarca: “Sierra Ambato” [protologue]; “shady places below
grasses and shrubs”, 900-1100 m, 27.7.1976 [detailed locality data deposited with the holotype
according to the protologue, but no such data was found] (type herbaria: B (ex KOELN)).
B (ex KOELN) (spirit B 810013156) (holotype): corp, ar, sp, fl. – Notes: Labelled as “Holotyp”.
Collection data simply given as “P-22/5”. Additional data on the original label: “Ripp.+
Blüt. gleichzeitig fix.” [ribs and flowers simultaneously preserved].
B (ex KOELN) (spirit B 810013157) (isotype): fl, fr, sem. Notes: Labelled as “Isotyp”. Col
-
lection data simply given as “P-22”.
Gymnocalycium bruchii var. brigittae Piltz in Succulenta (Amsterdam) 66(10): 213-216, ill.,
SEM-ill. 1987
Notes: Republished in Gymnos 5(9): 30-34. 1988.
Type: Piltz 214/3 Argentina: Córdoba: “W slopes of the Sierra Grande” [protologue, trans
-
lated]; “stony grassland”, 1800 m, 1.7.1980 (type herbaria: B (ex KOELN)).
B (ex KOELN) (spirit B 810013158) (holotype): corp, ar, sp, fl. – Notes: Labelled as “Holotyp”.
Collection data simply given as “P-214/3”.
B (ex KOELN) (spirit B 810013159) (isotype): fl. – Notes: Labelled as “Isotyp”. Collection data
simply given as “P-214”.
Gymnocalycium cardenasianum F. Ritter in Taxon 13(4): 144. 1964
Notes: First mentioned as nom. nud. in Backeberg, Cact. 3: 1747. 1959.
Type: Ritter 88 loc. 1 Bolivia: Tarija; Mendez: “Carrizal” [protologue], 1953 (type herbaria: U
117818B, SGO, ZSS) [holotype cited for U in Taxon 12(1): 28, 1963].
Willdenowia 38 – 2008 247
B (sheet GH19350) (type number): ic (corp). Notes: Ex cult. BG Berlin, 8.1.1986, from mate
-
rial raised from seeds originally received 1956 from H. Winter. Collection number estab
-
lished by inference.
Gymnocalycium damsii var. multiproliferum P. J. Braun in Kakt. And. Sukk. 42(10): 228-231,
ill. 1991
Type: Braun 230 – Brazil: Mato Grosso do Sul: “SW of the Serra da Bodoquena” [protologue,
translated] / “c. 50 km NE of Porto Murtinho [P. J. Braun, pers. comm] / “E of Porto Murtinho”
[isotype label at B]; “Pantanal” [protologue], 1983 (type herbaria: ZSS, B, ZSS).
B (boxed s.n.) (isotype): corp, ar, sp. Notes: Leg. P. J. Braun (accessioned at B in 1991) sub
Gymnocalycium sp.
B (boxed s.n.) (isotype): corp, ar, sp. – Notes: Received at B as isotype but no original Braun la
-
bel present.
Gymnocalycium pflanzii var. albipulpa F. Ritter, Kakt. Südamer. 2: 660-661, fig. 642. 1980
Type: Ritter 397 loc. 1 Bolivia: s.l. [protologue], s.a. [originally distributed as Gymnocalycium
bolivianum Cárdenas ex F. Ritter (nom. nud.)] (type herbaria: U [not found], SGO [status ?], ZSS
[seeds only]) [type cited for U in Ritter, Kakt. Südamer. 1: iii. 1979].
B (sheet GH16276) (type number): fl, ic (corp). – Notes: Ex cult. BG Berlin, 4.7.1984, from ma
-
terial raised from seeds originally received 1960 from H. Winter. Collection number estab
-
lished by inference.
Gymnocalycium taningaense Piltz in Kakt. And. Sukk. 41(2): 22-26, ill., SEM-ill. 1990
Type: Piltz 212/6 Argentina: Córdoba: “near Taninga” [protologue, translated], 950 m, s.a.
[type number erroneously given as “F 212/6” in the protologue] (type herbaria: B (ex KOELN)).
B (ex KOELN) (spirit B 810013160) (holotype): corp, ar, sp, fl. – Notes: Labelled as “Holotyp”.
Collection data simply gievn as “P-212/6”.
B (ex KOELN) (spirit B 810013161) (isotype): fl, fr, sem. Notes: Labelled “Isotyp”. Collec-
tion data simply given as “P-212”.
Hildewintera colademononis Diers & Krahn in Kakt. And. Sukk. 54(8): 221, ill. 2003 (incorrect
name, Art.11.4).
Type: Krahn 950 – Bolivia: Santa Cruz; Florida: “Cerro El Fraile” [protologue], 2000 (type her
-
baria: B (ex KOELN)).
B (ex KOELN) (spirit B 810013162) (holotype): corp, ar, sp. – Notes: Labelled “Holotyp”. Col
-
lection data simply given as “WK 950”.
B (ex KOELN) (spirit B 810013163) (isotype): fl. Notes: Labelled as “Isotyp”, but collection
data given as “WK 950/A”. This most probably refers to a second individual of the same col
-
lection, as the protologue does not mention more than one locality for the taxon.
Horridocactus andicola F. Ritter in Succulenta (Amsterdam) 1959(7): 97-100, ill. 1959
Type: Ritter 468 loc. 1 Chile: Región V; Los Andes: “Cerro Chivato (zwischen Santiago und
Los Andes)” [holotype label at U] = Cerro Chivato (between Santiago and Los Andes), 5.1955
(type herbaria: U, SGO [status ?], ZSS [seeds only, status ?]) [type cited for ZSS but eventually
found at U].
B (sheet GH22399) (type collection): ic (corp, ar, sp, fl), fl. – Notes: Ex cult. BG Berlin 153-92-74-80,
2.8.1987, from material originally raised from seeds received 1957 from H. Winter. Collec
-
tion number established by inference.
B (sheet GH31084) (type collection): fl. Notes: Ex cult. BG Berlin 153-92-74-80, 27.5.1993,
from material originally raised from seeds received 1957 from H. Winter. Collection number
established by inference.
248 Eggli & Leuenberger: Type specimens of Cactaceae names in the Berlin Herbarium
Horridocactus paucicostatus F. Ritter in Succulenta (Amsterdam) 1959(9): 113-114, ill. 1959
Type: Ritter 521 loc.1 Chile: Región II; Antofagasta: “c. 20 km N of Paposo” [protologue, trans
-
lated], s.a. (type herbaria: U 097984B, SGO [status ?], ZSS [seeds only, status ?]) [Type cited for
ZSS in the protologue, but eventuall found at U. F. Kattermann (Succ.Pl. Res.1: 162, 1994) un
-
necessarily designated the illustration on p. 114 of the protologue as lectotype].
B (sheet GH36449) (type collection): ic (corp, ar, sp, fl), fl. – Notes: Ex cult. BG Berlin 154-09-74-80,
13.5.1998, from material raised from seeds originally received 1957 from H. Winter. Collec
-
tion number established by inference.
Horridocactus paucicostatus var. viridis F. Ritter in Succulenta 1959(9): 113-114. 1959
Type: Ritter 521a loc. 1 Chile: Región II; Antofagasta: “surroundings of Paposo” [protologue],
s.a. (type herbaria: U 087554, SGO [status ?], ZSS [seeds only, status ?]) [type cited for ZSS in the
protologue, but eventually found at U].
B (sheet GH23762) (type collection): ic (corp, ar, sp, fl), fl. Notes: Ex cult. BG Berlin 154-11-74-80,
13.7.1988, from material originally raised from seeds received 1960 from H. Winter. Collec
-
tion number established by inference.
Leocereus bahiensis var. barreirensis P. J. Braun & Esteves in Kakt. And. Sukk. 41(9): 205.
1990
Type: Esteves Pereira 118 – Brazil: Bahia: “W part of the state of Bahia, near the city of Barrei-
ras” [protologue, translated]; “on standstone rocks”, s.a. [probably collected 1979, cf. one of the
isotype labels at B] (type herbaria: UFG 12.360, B, ZSS).
B (boxed s.n.) (isotype): corp, ar, sp. – Notes: Leg. P. J. Braun (ex cult.?).
B (boxed s.n.) (isotype): corp, ar, sp. Notes: Leg. P. J. Braun (ex cult.?); original label: “Bra-
silien, Bahia, Barreiras, Campo Cerrado, 1979?”.
Leocereus bahiensis var. exiguospinus P. J. Braun & Esteves in Kakt. And. Sukk. 41(9): 205.
1990
Type: Esteves Pereira 135 Brazil: Bahia: “in the surroundings of Canabrava” [protologue,
translated]; “caatinga” [isotype label at B], 1981 [probably collected 1981, cf. isotype label at B]
(type herbaria: UFG †, B, ZSS) [holotype reported as missing as of Jan. 1991 by Taylor & Zappi,
Cacti Eastern Brazil: 428. 2004].
B (boxed s.n.) (isotype): corp, ar, sp. – Notes: Leg. P. J. Braun (ex cult.?); original label: “Brasi
-
lien, Bahia, Canabrava, Caatinga, 1981?”.
Leocereus bahiensis var. robustispinus P. J. Braun & Esteves in Kakt. And. Sukk. 41(9): 205.
1990
Type: Braun 702 Brazil: Bahia: “in the N of the state of Bahia, upper parts of the Serra do Iná
-
cio, near Gameleira” [protologue, translated], 7.1986 [collection date according to isotype label
at B] (type herbaria: ZSS, B).
B (boxed s.n.) (isotype): corp, ar, sp, (al). – Notes: Leg. P. J. Braun (ex cult.?).
Leocereus estevesii P. J. Braun in Kakt. And. Sukk. 41(9): 204-205. 1990
Type: Esteves Pereira 207 Brazil: Piauí: “S of Piauí, frontier region towards S Maranhão”
[protologue, translated]; “on heavily weathered sandstone rocks in very dry region” [data associ
-
ated with isotype at ZSS], s.a. [collection date 1984 or 1985 according to isotype at B] (type her
-
baria: UFG 12.380, B, B (ex KOELN), ZSS).
B (boxed s.n.) (isotype): corp, ar, sp. – Notes: Leg. P. J. Braun (ex cult.?), accessioned 1990.
B (ex KOELN) (boxed B 100216303) (isotype): corp, ar, sp. – Notes: Labelled “Isotyp” on the
original label. Locality data simply given as “Piauí”.
Willdenowia 38 – 2008 249
Lobivia amblayensis Rausch in Kakt. And. Sukk. 23(3): 67-88, ill. 1972
Type: Rausch 19 Argentina: Salta: “E of Amblayo” [protologue, translated], 2800 m, s.a. (type
herbaria: W, ZSS).
B (sheet GH31078) (type collection): ic (corp), fl. Notes: Ex cult. BG Berlin 123-18-85-50,
27.5.1993, from material originally received from Wien-Schönbrunn.
Lobivia famatimensis var. sanjuanensis Rausch in Kakt. And. Sukk. 28(4): 75. 1977
Type: Rausch 557 Argentina: San Juan: “near San Juan” [protologue, translated], 1700 m, s.a.
(type herbaria: ZSS).
B (sheet GH29575) (type collection): ic (corp), fl. Notes: Ex cult. BG Berlin 201-17-82-30,
19.5.1992, from material originally received from Wien-Schönbrunn.
Lobivia quiabayensis Rausch in Kakt. And. Sukk. 21(3): 45. 1970
Notes: First published in Kakt. And. Sukk. 19(2): 22-23, ill. 1968, and there doubtfully invalid
(Art. 37.1).
Type: Rausch 205 Bolivia: La Paz: “near Quiabaya” [protologue, translated], 3500 m, s.a. (type
herbaria: W).
B (sheet GH19846) (type collection): ic (corp), fl. Notes: Ex cult. BG Berlin 201-06-82-30,
13.5.1986, from material originally received from Wien-Schönbrunn.
B (sheet GH33656) (type collection): fl. Notes: Ex cult. BG Berlin 201-06-82-30, 4.5.1995,
from material originally received from Wien-Schönbrunn.
Lobivia rauschii Zecher in Succulenta (Amsterdam) 53(3): 42-43, ill. 1974
Notes: When transferred to Echinopsis, the valid name is E. yuquina D. R. Hunt 1991.
Type: Rausch 635 Bolivia: Chuquisaca; Sud-Cinti: “S of Culpina” [protologue, translated] /
“near Yuquina” [W. Rausch, pers. comm.1991], 3200 m, s.a. (type herbaria: ZSS).
B (sheet GH24852) (type collection): ic (corp, ar, sp). Notes: Ex cult. BG Berlin 201-20-82-30,
19.5.1989, from material originally received from Wien-Schönbrunn.
Lobivia tiegeliana var. uriondoensis Rausch in Succulenta (Amsterdam) 58(10): 233-234, ill.
1979
Type: Rausch 90 Bolivia: Tarija: “near Concepcion” [protologue, Latin diagnosis, translated] /
“near Uriondo” [protologue, Dutch description, translated], s.a. (type herbaria: ZSS).
B (sheet GH29921) (type collection): ic (corp, fl), fl. – Notes: Ex cult. BG Berlin 201-02-82-30,
14.7.1992, from material originally received from Wien-Schönbrunn.
Lobivia zecheri Rausch in Succulenta (Amsterdam) 50(8): 146, ill. (p. 141). 1971
Type: Rausch 407 Peru: Ayacucho: “along the road from Ayacucho to Huanta” [protologue,
translated], 2400 m, 1970 (type herbaria: W, ZSS).
B (sheet GH15850) (type collection): ic (corp), fl. Notes: Ex cult. BG Berlin 201-10-82-30,
from material originally received from Wien-Schönbrunn.
B (sheet GH22137) (type collection): ic (corp), fl. Notes: Ex cult. BG Berlin 147-59-74-80,
3.7.1987, from material originally received 1978 from G. Köhres.
Mammillaria aylostera Werdermann in Kakteenkunde 1938: 99, 112. 1938
Type: Ritter s.n. – Mexico: Guerrero: “at the lower Balsas river in the coastal plain” [protologue,
translated]; “in open scrub”, s.a. (type herbaria: B [“clono”]) [ex cult. BG Berlin 14.1.1939].
B (spirit B 810006969) (type collection): corp, ar, sp. Notes: Ex cult. BG Berlin, 14.1.1939,
and thus posterior to the protologue.
250 Eggli & Leuenberger: Type specimens of Cactaceae names in the Berlin Herbarium
Mammillaria beiselii Diers in Kakt. And. Sukk. 30(3): 57-60, ill. 1979
Type: Beisel & Krasucka 100 Mexico: Colima / Michoacán: “near Balmerio la Brisa”; “together
with Aporocactus sp.", 25.4.1978 [detailed locality information supplied directly by the collec
-
tor] (type herbaria: B (ex KOELN), ZSS).
B (ex KOELN) (boxed B 100216304) (holotype): rad, corp, ar, sp. – Notes: Labelled as “Holo
-
typ” on the original label. Collection data simply given as “BK 100”. A copy of the
protologue with annotated locality data is archived separately.
B (ex KOELN) (boxed B 100216305) (isotype): corp, ar, sp. Notes: Labelled as “Isotyp” on
the original label. Collection data simply given as “BK 100”.
B (ex KOELN) (boxed B 100216306) (isotype): sem. – Notes: Collection data simply given as
“BK 100”, and without type annotation.
B (ex KOELN) (spirit B 810013137) (isotype): ar, sp, fl. – Notes: Collection data simply given
as “BK 100”, and without type annotation.
Mammillaria duwei Rogozinski & P. J. Braun in Kakt. And. Sukk. 36(8): 158-164, ill. 1985
Type: Duwe & Rogozinski 1 – Mexico: Guanajuato: “E part of the state of Guanajuato, near the
city of San Luis de la Paz” [protologue]; “on and between limestone rock outcrops in organic
soil”, 1983 (type herbaria: KOELN (missing), B, ZSS).
B (boxed s.n.) (isotype): rad, corp, ar, sp. – Notes: Leg. P. J. Braun (ex cult.?).
Mammillaria hahniana Werdermann in Monatsschr. Deutsch. Kakteen-Ges. 1(4): 77-79, ill.
1929
Type: Schmoll s.n. Mexico: Querétaro: “locality will be announced later” [protologue, trans-
lated] / “Sierra de Jalpan, above 2000 m” [neotype designation], 2000 m, 1928 (type herbaria: B
[neo]) [ex cult. hort. Hahn, June 1931. Neotype (as lectotype, correctable under 9.8) designated
by Hunt & Taylor in Cactaceae Syst. Init. 21: 8. 2006].
B (spirit B 810006971) (neotype): rad, corp, ar, sp.
Notes. The spirit specimen (B810006971) was prepared from cultivated material from the
nursery of Hahn and is dated June 1931. Despite the