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DOI: 10.1127/0006-8152/2009/0127-0459 0006-8152/09/0127-0459 $ 03.25
© 2009 E. Schweizerbart’sche Verlagsbuchhandlung, D-70176 Stuttgart
Received January 15, 2009, accepted February 25, 2009
Bot. Jahrb. Syst. 127 4459-471 Stuttgart, 26. Juni 2009
New Allium (Alliaceae) species from Tajikistan, Kyrgyzstan,
and Uzbekistan
By
Reinhard M. Fritsch
With 4 figures and 1 table
Abstract
Fritsch, R. M.: New Allium (Alliaceae) species from Tajikistan, Kyrgyzstan, and Uzbeki-
stan. — Bot. Jahrb. Syst. 127: 459–471. 2009. — ISSN 0006-8152.
Four Allium species from the mountains of Central Asia are described as new to science.
Three species belong to A. subg. Melanocrommyum: A. intradarvazicum (A. sect. Regelo-
prason) and A. khozratense (with unclear affiliation, probably A. sect. Thaumasioprason)
occur in the Tajik Darvaz mountain range, A. chychkanense (A. sect. Longibidentata) in
the Kyrgyz Susamyr massif of the Western Tian-Shan mountain range. The fourth spe-
cies, i. e. A. furkatii (A. subg. Reticulatobulbosa sect. Campanulata), occurs in the Uzbek
part of the Chatkal mountain range. Taxonomic relationships and systematic affiliations
of the new taxa are discussed, and Allium subsect. Longibidentata is raised from subsec-
tional to sectional rank.
Keywords: Allium, taxonomy, Central Asia, new taxa.
Introduction
In the last decades, many Allium taxa were newly reported or newly de-
scribed from the former Soviet part of Central Asia. More than 35 years ago,
Vvedensky & Kovalevskaya (1971) accepted 191 species to occur in this part of
Central Asia. The more recent conspectus of Khassanov (1997) enumerated 207
wild Allium species and 4 subspecies, and the latest version (Khassanov 2008)
lists even 224 species and 20 subspecies. This is certainly not the final number of
Allium species and subspecies growing in this botanically interesting area. Dur-
ing joint taxonomic research missions undertaken with Uzbek botanists in the
eschweizerbartxxx ingenta
460 R. M. Fritsch, New Allium (Alliaceae) species
1990ies (Khassanov & Fritsch 1994, Fritsch et al. 1998) we became aware of
several taxa difficult to associate to known species. One of them was cultivated
in Gatersleben and is described below as a new species.
In the years 2003 to 2006, several research missions were undertaken in
Tajikistan and Uzbekistan focusing in part on mountainous areas which for-
merly had been rarely studied by botanists. These missions were undertaken in
order to find out which wild-growing Allium species are collected by the local
population, and for what kind of purpose. The plants were often collected prior
to (or sometimes after) anthesis, and a remarkable number of them could only
be identified in a preliminary way. After fieldwork the plants were transferred to
newly established Allium collections in the countries where they had been col-
lected (for more details see Keusgen et al. 2006, Keusgen & Fritsch 2008). Later
these cultivated plants were taxonomically analyzed, but not all plants could be
shown to belong to known taxa. From this material, two species are described
here as new to science. The formal descriptions of other newly recognized taxa
belonging to A. subgenera Allium and Polyprason will be published elsewhere
(Fritsch & Khassanov 2008; Fritsch & Friesen, in press).
During a visit to the Institute of Botany in Dushanbe (TAD), two incor-
rectly determined herbarium specimens not belonging to any hitherto known
Tajik species were found. They represent a new species which is also described
below.
Results
1. A. subg. Melanocrommyum (Webb & Berthel.) Rouy
1.1. A. sect. Regeloprason Wendelbo
During the first half of 20th century, all pinkish-flowering members of A.
subsect. Odoratae R.M. Fritsch were named A. winklerianum Regel irrespective
of a conspicuous morphological and ecological diversity. Only Vvedensky
(1963) separated plants from submontane steppe slopes of the Central Hissar
mountain range as A. hissaricum Vved. A second subspecies A. hissaricum subsp.
cisdarvasicum Kamelin & Seisums from southern Tajikistan was mentioned by
Seisums (1994) but never validly published. Later the morphologically distinct
populations from the Fergan mountain range have been described as A. pseudo-
winklerianum R.M. Fritsch & F.O. Khass. (Fritsch 2000), and plants from the
western Alai mountain range having conspicuously narrow leaves as A. sochense
R.M. Fritsch & U. Turakulov (Fritsch et al. 2002).
Recently we were able to verify that plants growing on montane to subalpine
slopes of the Hissar mountain range are morphologically inseparable from
plants growing at the type location of A. winklerianum in the Khobu-Rabot
pass area of the central Darvaz mountain range (Fritsch 2008). However, on
moist slopes in the Khumbov valley less than 20 km apart from the Khobu-
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461R. M. Fritsch, New Allium (Alliaceae) species
Rabot pass plants with similar flowers grew which were generally about twice as
high (Fig. 1A) and showed obliquely directed, much broader, smooth, and semi-
glossy leaves, as well as rotundate-triangular and nearly smooth ovaries (Fig. 1C).
These plants do not belong to A. pseudowinklerianum which is characterized by
thick, erectly standing, glossy leaf blades, and spherical, finely tuberculate ova-
ries. Thus the plants from the Khumbov valley represent a new species.
Allium intradarvazicum R.M. Fritsch, spec. nova — Fig. 1.
Type: Cultivated in Gatersleben no. TAX6452, leg. 14. 05. 2007; bulbs col-
lected in Tajikistan, Darvaz range, Khumbov canyon, canyon Ganjurak oppo-
site to village Gshun, N exposed loamy slope among shrubs and perennials,
2560–2950 m, 38°38’01,8” N, 70°52’06,2” E, 23.07.2005, R.M. Fritsch, M. Keus-
gen, H. Hisoriev & I. Kudratov 6219 (holotype GAT).
Scapus 25 usque 40 cm altus, folia tenuia laevia semi-nitida oblique erecta et
ovaria rotundato-triangularia sublaevia. Differt ab aliis speciebus rubrifloribus
characteribus supra citatis; A. winklerianum statura humiliore et foliis strictis
impolitis asperis, A. pseudowinklerianum foliis crassis strictis nitidis et ovariis
globosis subtiliter tuberculatis.
Bulbs depressed-globose, rarely more than 2 cm wide, outer tunics greyish
brown. Scape cylindrical-conical, slightly flexuose, smooth, (15) 25–40 cm high,
4–5 mm in diameter, vividly green with glaucous bloom, near base brown
flushed. Leaves 1–2, narrowly lanceolate with a linear medium part, gradually
tapering in a hooded tip, lower part ± canaliculate and with fine teeth, upper part
even and smooth, strictly upright, later upper part hanging down, very thin, up-
per side smooth or with fine grooves, lower side with dense fine ribs, 35–60 cm
long, 1–3(4.5) cm wide, vividly green but brown flushed near base, weakly
glossy because of glaucous bloom. Outer sheath leaf delicately membranous,
visible but soon decaying. Spathe membranous, completely split in 2 widely
ovate-triangular parts more or less adpressed to the pedicels, yellowish-brown-
ish with brown veins. Flowers narrowly funnel-shaped, with nice sweet odor,
anthesis in May. Inflorescence initially ovate-fasciculate later widely fasciculate,
moderately many-flowered, (1.5)2–4 cm in diameter. Pedicels thin and wiry,
nearly straight, cylindrical, green. Tepals up to 1/3 of its length basally united,
the free part widely lanceolate, outer ones narrower and initially somewhat
shorter, obliquely forward directed and slightly recurved, arcuately tapering
into an obtuse plicate tip, c. 12 (free part 7–8) mm long, inner tepals 2 mm,
outer ones 3 mm wide, pinkish-carmine with narrow green median vein. Fila-
ments basally united with the tepals and among themselves, free part of outer
filaments 1/3 as long as the free part of tepals, shortly subulate, fleshy, free part
of inner filaments 2/5–3/5 as long as the free part of tepals and ± triangular; ba-
sally more intensely red colored than upper part. Anthers ovately arrow-shaped,
tipped by a short outgrowth of connective, c. 1.5 mm long and 0,8 mm wide,
yellow. Pollen faintly yellow. Ovary long stalked, ovate, 3 mm long and nearly
2 mm in diameter, green (sutures brighter), somewhat glossy, surface cells of up-
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462 R. M. Fritsch, New Allium (Alliaceae) species
Fig 1. Allium intradarvazicum. A: Cultivated plants from type location in anthesis, B:
inflorescence in early stage of anthesis, C: green capsules, D: stamina with anthers, tepals,
and ovary, scale bar = 4 mm.
per part shallowly tuberculate, those of lower part ± smooth, mound of nectary
very small, located near the very base of ovary. Style conically filiform, 1–3 mm
long, basally whitish and pinkish-carmine above. Stigma very shortly incrassate,
whitish. Capsule triangularly globose, valves with irregular ribs, about 6–7 mm
in diameter, opening by narrow clefts. Seeds 2–3 (6) per loculus, 2–2.5 mm long,
c. 1.5–2 mm wide and thick, dull black.
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463R. M. Fritsch, New Allium (Alliaceae) species
Distribution: Tajikistan: Darvaz mountain range, only known so far from
Khumbov valley.
Other material seen: Two accessions cultivated in Gatersleben from bulbs
collected in Tajikistan, Darvaz range, Khumbov canyon: (1) TAX6450 leg.
07.05.2007 (GAT), from Regi Ravon above third bridge, E exposed loamy-stony
slope between shrubs and perennials, 2100 m, 38°37’24,1” N, 70°51’48,7” E,
22.07.2005, R.M. Fritsch, M. Keusgen, H. Hisoriev, I. Kudratov 6216; (2)
TAX6454 leg. 14.05.2007 (GAT), small canyon Osiobdara near village Gshun, S
exposed loamy-stony slope, among high perennials, 2700 m, 38°39’24,1” N,
70°51’20,1” E, 24.07.2005, R. M. Fritsch, M. Keusgen, H. Hisoriev, I. Kudratov
6221.
1.2. A. sect. Thaumasioprason Wendelbo
Vvedensky determined two herbarium specimens kept at TAD as “Allium
angustitepalum Wendelbo aff.”. However, only at a first glance these plants look
like members of A. sect. Megaloprason Wendelbo. On closer examination, the
tepals are not reflexed after anthesis (Fig. 2), and the ovaries show a smooth
surface and a very narrow base. In addition, a pedicel character is very specific:
the ovaries are not only inflated near the base but also at the tip (Fig. 2 inset).
The conical widening of the tip of pedicels directly below the flower is an
extremely common character in the genus Allium, but this kind of spherical in-
flation is not known from any other species. These plants represent a hitherto
undescribed species:
Allium khozratense R.M. Fritsch, spec. nova — Fig. 2.
Typus: Tajikistan, Yugo-vostochn. chast Yuzho-tadzhikistanskoj depressii,
zapadnyj sklon khrebta Khirkh, 2 km yugo-vostochni kishlaka Sarygor, na
galechnikakh [Khozratishoh mountain range SE of Kulyab, W slope of Khirkh
massif 2 km SE of vill. Sarygor, gravel area], 1400 m, 01. VI. 1961, Soskov &
Yunusov 283 (left plant; holotype TAD 12811).
Statura graciliore, scapus usque 25 cm altus foliis anguste-lanceolatis acutis
et brevioribus quam scapus, inflorescentia deinde basi dilabentia, flores kermesi-
ni stellari-infundibuliformes tepalis strictis deinde non reflexis, ovaria subpolita
applanato-obovata basin versus peracutata, pedicelli tenues stricti basi elongato-
obovati et apice sphaerico-inflati. Differt ab speciebus A. sectionis Megalopra-
son tepalis post anthesin non reflexis et ovariis subpolitis non verrucosis, et ab
specie affini Allio scotostemon tepalis lanceolatis subacutis (non elliptico-oblon-
gis obtusis) et filamentis tertia partis brevioribus tepalorum (non equilongis) et
statura graciliore (non compacta humiliore).
Bulbs ovate, c. 1.5 cm in diameter and 2 cm long, outer tunics membranous,
grey up to brown. Scape erect, slightly flexuose, cylindrical, probably smooth,
c. 3 mm in diam., 15–25 cm long. Leaves 2 (3), blade very narrowly lanceolate,
long tapering, 4–8 mm wide, 20–30 cm long. Spathe irregularly split, patent.
Flowers funnel-shaped star-like, anthesis in the beginning of June. Inflorescence
semi-globular, moderately dense, c. 4 cm in diameter, after anthesis (and in the
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464 R. M. Fritsch, New Allium (Alliaceae) species
dry state, Fig. 2) basally disintegrating. Pedicels thin and wiry, nearly straight, at
base long-obovate and at tip somewhat spherically inflated. Tepals lanceolate,
4,5–6 mm long, carmine, after anthesis brownish but direction unchanged. Fila-
ments roughly 2/3 as long as tepals, subulate, inner ones basally slightly wider,
probably whitish. Anthers oblong, about 1 mm long and 0.5 mm wide. Ovary
depressed obovate with 6 shallow furrows, strongly tapering towards base but
not stalked, about 2 mm long and in diameter, surface ± smooth but not glossy.
Style long-conical tapering towards the undivided stigma. Capsule and seeds not
seen.
Fig 2. Allium khozratense, type specimen (holotype: most left plant), inset: ovary, stamina
with anthers, and tepals, scale bar = 2 mm.
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465R. M. Fritsch, New Allium (Alliaceae) species
Distribution: South-East Tajikistan, southern part of Khozratishoh moun-
tain range.
Other material seen: Yu-v. chast tadzhikistan. depressii, 6 km S.-v. kishl. Iel,
k yu.-v. ot kishl. Sary-Mazgar. Skaly [southeastern part of the south-Tajik low-
land, 6 km NE of village Iol SE of village Sary-Mazgar, rocks], 1400–1500 m, 18.
VI. 1961, Yu. D. Soskov & S. Yu. Yunusov 1440 (TAD 12812). Possibly also an-
other sheet may belong here, with inflorescence and flowers untypically devel-
oped in the press: Tajikistan, Gory u k-ka Iol nedaleko ot k-ka Sary-Mazgar,
kamenistaya sklon, [mountains near village Iol not far from village Sary-Mazgar,
stony slope] 14000 (sic!) — 1500 m nad u. m., 06. VI. 1991, A. Seisums (LE, also
labelled as “Holo-Typus A. pandzhi Seisums”, a name never validly pub-
lished).
One may assume that the capsules of A. khozratense do not open very much
but are dropping down with the seeds inside as it is known from A. giganteum
and A. macleanii of A. sect. Compactoprason (Fritsch 1992). All the above men-
tioned specific characters do not allow to assign A. khozratense with certainty to
any of the known sections of A. subg. Melanocrommyum. A certain affinity (but
no close relationship) may exist to A. scotostemon Wendelbo which shows, de-
spite of a much more condensed habit, a similar shape of tepals and filaments,
and shares basally disintegrating inflorescences with all species of A. sect. Thau-
masioprason. Allium gypsaceum Vved. (A. sect. Popovia F.O. Khass. & R.M.
Fritsch) also possesses basally disintegrating inflorescences but completely
different flower characters. Further investigations will be essential in order to
verify whether this affiliation is correct, or if A. khozratense belongs to a not yet
recognized taxonomic group.
1.3. A. sect. Longibidentata (R.M. Fritsch) R.M. Fritsch, stat. et comb. nov.
≡ Allium L. sect. Acmopetala R.M. Fritsch subsect. Longibidentata R.M.
Fritsch in Linzer Biol. Beiträge 26: 974, 1994. — Type: A. fetisowii Regel.
Recent molecular-phylogenetic studies of A. subg. Melanocrommyum (Gu-
rushidze et al. 2008) arranged the species in a basal grade (8 species) and a core
clade with all the remaining species. The divisions within the grade were statisti-
cally well supported. One group represents A. fetisowii in the wide sense con-
sisting of two statistically well supported geographical subgroups (1. west of 76°
E longitude; 2. south of Kirgizian Alatau and Talassian Alatau mountain ranges)
and a third weakly supported geographical subgroup (east of 76° E longitude).
The large genetic distance between these subgroups favors to recognize them at
the level of a section.
Well visible morphological differences between the two “northern” sub-
groups (1. and 3.) were not found yet. They constitute typical A. fetisowii which
grows on grassy steppe slopes. However, plants of the “southern” (2.) subgroup
differed morphologically from them by shorter and broader leaves and by tepals
completely turned backwards. They occur in higher altitudes under more hu-
eschweizerbartxxx ingenta
466 R. M. Fritsch, New Allium (Alliaceae) species
mid conditions among other perennials and small bushes and represent another
taxon:
Allium chychkanense R.M. Fritsch, spec, nova. — Fig. 3.
Type: Cultivated in Gatersleben no. TAX5057, leg. 05.05.1999, (holotype
GAT), plants collected in Kyrgyzstan, Talassischer Alatau, Rechter Hang am
Chichkan-Fluß ca. 15 km unterhalb des Alabel-Passes, trockene Stellen am
Hang, ca. 2200 m, 42°15’ N, 73°00’ E, 03.07.1994, R.M Fritsch, K. Pistrick &
F. O. Khassanov 1206 (GAT, TASH).
Eminet foliis usque 5 cm latis et non plus 25 cm longis, tepalis sub anthesin
perfecte retrorsis, et dentibus lateralibus brevibus variabiliformibus basin fila-
Fig 3. Allium chychkanense. A: Cultivated plants from type location in anthesis, B: inflo-
rescence in full anthesis, C: ovary, stamina with anthers, and tepals, scale bar = 2 mm.
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467R. M. Fritsch, New Allium (Alliaceae) species
mentorum interiorum (et non numquam exteriorum) versus. Allium fetisowii
differt foliis usque 2 cm latis et usque 40 cm longis, dentibus lateralibus basin
interiorum filamentorum versus longe triangularibus. Allium schachimardani-
cum non eminet dentibus basin filamentorum interiorum versus.
Bulbs depressed-globose, 8–25 mm in diameter, 2–20 mm long, inner tunics
membranous, white, outer ones grey to black, decomposing in pieces. Scape
cylindrical, smooth, 4–6 mm in diameter, 40–60 cm long, green, basally purple
flushed. Leaves 1–2, lanceolate, stiff up to low obliquely positioned, ± longitu-
dinally carved, later nearly flat, 20–25 cm long, 1.5–5 cm broad, vividly green,
glaucous, basally violet flushed, upper side with shallow ribs, lower side with
flat and broad ribs, shortly arcuately tapering in a hooded (often reddish) tip,
margin reddish, smooth or very finely toothed. Outer sheath leaf delicate,
quickly decaying. Spathe membranous, nearly completely divided into two
shortly tipped valves, ± reflexed. Anthesis in May. Inflorescence very dense,
many-flowered, initially oval, later globular or somewhat conical. Pedicels thin,
wire-like, straight, 15–25 mm long, green, basally reddish. Flowers initially cup-
shaped, star-like. Tepals lanceolate to triangular, turned backwards but upper
part slightly inverted, shortly tipped, tip claw-like, 6–8 mm long, 1.2–2 mm
wide, rose, the greenish median vein is visible at the outer side only in the distal
part. Filaments approximately as long as the tepals, basally c. 0.5 mm long unit-
ed and shortly triangular widened (inner ones twice as broad as the outer ones),
above subulate, inner (and often also outer) filaments basally with a blunt, ir-
regularly formed tooth at every side, pink, basally mostly whitish. Anthers ob-
long, 1.6–2 mm long, c. 1 mm wide, violet. Pollen yellowish grey. Ovary globu-
lar-pyriform, stipitate, c. 2 mm long and in diameter, surface coarsely tuberculate,
slightly glossy, with shallow longitudinal furrows, greenish with purplish
flushed suturs. Mound of ovary slit-like near the base. Style conical-threadlike,
3–6 mm long, rose, like the undivided stigma. Capsule triangular-pyramidate,
4–5 mm long and in diameter, finely scrobiculate, yellowish-brown, slightly
glossy. Seeds dull black, 1.5–1.8 mm long, c. 1.2 mm wide and thick.
Distribution: moderately dry slopes and river terraces in higher altitudes of
the Susamyr massif, Central Tian-Shan mountain range.
Other material seen: Cultivated in Gatersleben no. TAX5060, leg. 29.05.1996
and 03.05. 1999 (GAT), collected in Kyrgyzstan, Chichkan-Tal ca. 25 km südlich
vom Pass Alabel, schattiger Südhang, Talassischer Alatau, 1750 m, 42°10’ N,
72°52 E, 04.07.1994, R.M Fritsch, K. Pistrick & F. O. Khassanov 1210.
Allium schachimardanicum Vved. shows similar leaves as the new species
but differs in tepals with retrorse basal and introrse upper part and filaments
without side-teeth. It grows on rock terraces in a small area of Alai mountain
range only. This species belongs molecularly to the core clade where also other
subsections of A. sect. Acmopetala were inserted (Gurushidze et al. 2008). Thus
the former A. subsect. Longibidentata cannot longer be inserted under A. sect.
Acmopetala but represents a section of its own to which only A. fetisowii and
A. chychkanense belong.
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468 R. M. Fritsch, New Allium (Alliaceae) species
2. A. subg. Reticulatobulbosa (Kamelin) Friesen sect. Campanulata Kamelin
In the mountains south and east of Tashkent, most often A. jodanthum Vved.
with deep-carmine flowers can be met with, and also two forms of A. barscze-
wskii Lipsky with pinkish -violet (f. violaceum Krassovs.) and white flowers
(f. niveum Krassovs.) are not rare on slopes covered with good soil. On rocky
outcrops and terraces a more delicate, rose-pinkish flowering, and narrow-
leafed species is sometimes found which keys out as A. inconspicuum Vved.
according to Vvedensky & Kovalevskaya (1971). However, the original descrip-
tion of the latter species, based on plants from the Kazakh steppe areas north of
Tashkent, mentions mostly rough leaf margins, light violet flowers, and violet
anthers. Plants from the Kazakh Syrdarya-Karatau mountain range, also keying
out as A. inconspicuum, have smooth leaves, blood-red tepals, and yellow an-
thers (see Fritsch 2008: 77, fig. 30 C) and represent probably another related
taxon. The different key characters of the discussed taxa are compared in Ta-
ble 1. The plants from the Chatkal mountain range (Fig. 4) also do not belong to
A. kysylkumi Kamelin which has pinkish tepals with scarcely darker median
vein, smooth leaves, and as main difference crown-like outgrowths at the top of
ovary. Thus the plants similar to A. inconspicuum represent a new species, a fact
which also could be clearly recognized in cultivated specimens.
Table 1. Key characters of A. inconspicuum, A. barsczewskii, and A. furkatii
Taxon A. inconspicuum A. inconspicuum A. barsczewskii A. furkatii
s. lat. (Karatau) s. str.
leaf blades Smooth rough smooth dentate or shortly
hairy along margin
flower shape urn-shaped (no data) narrowly widely funnel-
funnel-shaped shaped
Tepals blood-red light violet white, pink to pink with purple
purplish pink median vein
filaments pink with blood- (no data) white through- whitish with purple
red distal part out or with distal part
pink distal part
Anthers Yellow violet yellow or purple purple
Allium furkatii R.M. Fritsch, spec. nova — Fig. 4.
Type: Cultivated in Gatersleben no. TAX6360, leg. 07. 05. 2007 (holotype
GAT); plants collected in Uzbekistan, Chatkal range, above village Sukok 50 km
eastern Tashkent, southern rocky slope, steppe meadow, 1380 m, 41°14’54,7” N,
69°49’01,3” E, 20.07.2005, R.M. Fritsch et F.O. Khassanov 4209.
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469R. M. Fritsch, New Allium (Alliaceae) species
Folia angusta marginibus dentatis sive breve pilosis, flores late infundibuli-
formes, et tepala rosea nervo mediano conspicuo purpureo. Ab speciebus affini-
bus differt foliis non glabris, forma et colore florum et tepalis nervo mediano
parum conspicuo.
Bulbs narrowly ovate, c. 5–7 mm in diameter, 1.5–2 cm long, covered by
yellowish-brown reticulate tunics. Scape cylindrical, straight, 20–40 cm long,
2–3 (4) mm in diameter, smooth, green. Leaf sheathes covering the stem up to ¼
of its length, coarse, greenish, basally purple flushed; leaf blades sub-cylindrical,
later semi-cylindrical, 15–25 cm long, 1–2 mm wide, coarse by short cilia along
margins and partly along the longitudinal ribs of lower side; vividly green.
Spathe membranous, very shortly beaked, brownish with carmine flush, veins
Fig 4. Allium furkatii. A: Flowering plants at type location, B: inflorescence at type loca-
tion, C: inflorescence of cultivated plant, D: ovary, stamina with anthers, and tepals, scale
bar = 2 mm.
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470 R. M. Fritsch, New Allium (Alliaceae) species
yellowish-brown. Flowers campanulate, at full anthesis triangular-funnel-
shaped. Inflorescence fasciculate, dense. Pedicels straight, stiff, smooth, shorter
than flowers, after anthesis elongated; green, reddish flushed. Tepals basally
2.5–3.5 mm high connate, free part narrowly lanceolate, ± triangularly tapering
in a short obtuse tip, 6–7 mm long, plicate, pink with purple median vein, outer
tepals c. 2 mm wide, with basal margins covering the margins of inner ones, in-
ner tepals c. 1.5 mm wide. Filaments basally 2.5–3 mm high adnate to the bases
of tepals, fleshy, free part basally connate and triangular widened (inner fila-
ments two times wider than outer ones), whitish, upper part subulate and pink
or purple. Anthers oblong, c. 1 mm long and 0.5 mm wide, purple. Pollen yel-
lowish. Ovary ovate with three shallow longitudinal furrows, c. 2–2.5 mm long,
c. 1.5 mm in diameter, mound of nectary not seen, nearly smooth, green. Style ±
conically filiform, 1–3 mm long, pinkish, with white sub-capitate stigma. Cap-
sule broadly pyriform-triangular, 4–5 mm in diameter.
Distribution: Western Chatkal range, dry and rocky slopes.
Other materials seen: Uzbekistan, Schotterhang oberhalb Taschkent, ca.
1700 m, 15. 6. 1984, P. Gutte (GAT); cultivated in Gatersleben no. TAX5804 leg.
2005, collected by Dr. Maltzev in the vicinity of Parkent (S Tashkent), Chatkal
mountain range, Uzbekistan (GAT).
Acknowledgements
Thanks are due to the directors of the supporting institutions, Prof. Dr. H. Hisoriev
(Institute of Botany of the Tajik Academy of Sciences, Dushanbe), Prof. Dr. O. A. Ashur-
metov (Scientific-Productive Centre “Botanika” of the Uzbek Academy of Sciences,
Tashkent), for technical and organizational support, and Parvina Kurbonova, Hikmat
Hisoriev, and Furkat Khassanov for organizational work and leading participation in the
research missions. In addition, I would like to thank them and Michael Keusgen for de-
tailed and often long discussions on newly collected plants du ring field-work. I would
also like to express my gratitude to the staff of the IPK taxonomic Allium reference col-
lection, Mrs. Ch. Koch, Dr. K. Pistrick, and Mr. K. Hecker, for permanent support and
assistance especially when I took part in the missions. Financial support by the Volks-
wagenStiftung (Hannover, Germany) as part of the “PharmAll” project in the frame of
the program “Zwischen Europa und Orient — Mittelasien/Kaukasus im Fokus der Wis-
senschaft” is gratefully acknowledged.
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Address of the author:
Dr. Reinhard M. Fritsch, Leibniz-Institut für Pflanzengenetik und Kulturpflanzen-
forschung (IPK), Corrensstr. 3, 06466, Gatersleben, Germany.
E-mail: fritsch@ipk-gatersleben.de
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