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Disappearing 'alpine tundra,'Köppen climatic type in the western United States

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1] We examine changes in the areal extent of the Köppen ''alpine tundra'' climate classification type for the mountainous western United States following the work of Kottek et al. (2006). We find a significant decline in the area occupied by this climate category. In the early decades of the 20th century, the mean temperature of the warmest month in the areas of the western U.S. with nominal alpine tundra climates ranged largely between 8.5°Cand9.5°C.Inthelast20years(19872006),risingtemperatureshavecausedasignificantfractionoftheseareastoexceedthe10°Cthresholdforalpinetundraclassification.Theresulthasbeena738.5°C and 9.5°C. In the last 20 years (1987– 2006), rising temperatures have caused a significant fraction of these areas to exceed the 10°C threshold for alpine tundra classification. The result has been a 73% reduction in coverage of this climatic type. The remaining classified alpine tundra in the last 20 years now averages between 9°C–10°C during the warmest month, so that continued warming past the classification threshold, would imply that areas where this climate type is found today in the West will no longer be present.
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Disappearing ‘alpine tundra’ Ko¨ppen climatic type in the western
United States
Henry F. Diaz
1
and Jon K. Eischeid
1
Received 6 July 2007; accepted 27 August 2007; published 27 September 2007.
[1] We examine changes in the areal extent of the Ko¨ppen
‘alpine tundra’ climate classification type for the
mountainous western United States following the work of
Kottek et al. (2006). We find a significant decline in the area
occupied by this climate category. In the early decades of
the 20th century, the mean temperature of the warmest
month in the areas of the western U.S. with nominal alpine
tundra climates ranged largely between 8.5°C and 9.5°C.
In the last 20 years (19872006), rising temperatures have
caused a significant fraction of these areas to exceed the
10°C threshold for alpine tundra classification. The result
has been a 73% reduction in coverage of this climatic type.
The remaining classified alpine tundra in the last 20 years
now averages between 9°C–10°C during the warmest
month, so that continued warming past the classification
threshold, would imply that areas where this climate type is
found today in the West will no longer be presen t.
Citation: Diaz, H. F., and J. K. Eischeid (2007), Disappearing
‘‘alpine tundra’’ Ko¨ppen climatic type in the western United States,
Geophys. Res. Lett., 34, L18707, doi:10.1029/2007GL031253.
1. Introduction
[2] A number of studies have been published in the past
decade that indicate that the world’s mountain regions
comprise an area where the effects of global warming is
likely to be amplified. In particular, many of these studies
have documented an amplification of temperature trends
with height [Diaz and Graham, 1996; Liu and Chen, 2000;
Diaz et al., 2003; Bradley et al., 2004]. To examine changes
in climatic types that may have occurred during the past
century, we make use of the Ko¨ppen climate classification
system, which has been used to evaluate ongoing and future
climate change impacts on characteristic regional climate
types [Lohmann et al., 1993; Fraedrich et al., 2001; Kottek
et al., 2006; see also Dang et al., 2007]. Recent strong
warming trends in the western United States [Mote, 2003]
and associated impacts on vegetation [Cayan et al., 2001;
Breshears et al., 2005; Westerling et al., 2006] and stream-
flow [Stewart et al., 2005; Regonda et al., 2005] suggest
that climatic types in the region may also be changing.
[
3] WehaveusedtheKo¨ppen climate classification
system to evaluate changes in climatic types across the
United States. The classification algorithm was applied to
the PRISM (Parameter-elevation Regressions on Indepen-
dent Slopes Model) gridded data set of surface temperature
at 4-km resolution [Daly et al., 2002]. We have also used
the PRISM temperature data to evaluate maximum and
minimum temperature trends with elevation in the West; a
comparison of means and trends in the PRISM data set with
other data sets is given in the auxiliary material.
1
Here we
report on changes in one particular phenotype—the alpine
tundra, which is equivalent to Ko¨ppen type-E (polar cli-
mates). Briefly, polar climates are defined as occurring if the
mean temperature of the warmest month is less than 10°C.
In the West, only the tundra climate sub-type is present,
which is defined as occurring when the mean temperature of
the warmest month lies in the interval (0°,10°C]. We note
that the percent area coverage of this climate type is rather
small, occupying only about 0.2% of the area of the western
United States, or about 15,000 km
2
. There were 1226 4-km
pixels classified as ‘alpine tundra’ in the 1901 30 period,
whereas in 1987 2006, there were only 336 thus categorized—
a decline of 73%.
[
4] We also calculated linear temperature trends for the
period 1979 2006 for mountainous areas in Colorado, the
Pacific Northwest and the Sierra Nevada, which contain
most of the areas classified as ‘alpine tundra’’. This period
was chosen for two reasons. First, it corresponds to a period
of rapidly warming surface temperatures in the West (see
below). Second, it enabled us to compare elevational differ-
ences in temperature change s in PRISM with changes
derived from two independent data sets: the MSU lower
troposphere satellite-derived temperatures [Spencer et al.,
2006] for the domain of the western US that starts around
1979, and a homog enized version of the SNOTEL temper-
ature record provided by the Natural Resources Conserva-
tion Service, USDA, which also starts around this time.
2. Analysis
[5] To illustrate the recent warming trends across the
entire West, Figure 1 shows mean a nnual temperature
changes for the U.S. region west of about 102°W longitude.
The last 20 years of record (1987 2006) is a period of
consistently higher temperatures—an increase of 1°F
(0.6°C) averaged over the entire region compared to the
long-term (period-of-record) mean. This in turn has resulted
in a dramatic reduction—as much as 73%—in the area
classified as alpine tundra in the West, compared to the first
decades of the 20th century (Figure 2). This relatively large
change in the coverage of alpine tundra climate in the
western U.S. is quite remarkable. We should note that the
mean monthly temperature of the warmest month in most of
the areas we classified as alpine tundra (the key parameter in
1
Auxiliary materials are available in the HTML. doi:10. 1029/
2007GL031253.
GEOPHYSICAL RESEARCH LETTERS, VOL. 34, L18707, doi:10.1029/2007GL031253, 2007
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1
NOAA Earth System Research Laboratory, Boulder, Colorado, USA.
This paper is not subject to U.S. copyright.
Published in 2007 by the American Geophysical Union.
L18707 1of4
Ko¨ppen Type E) in the West is close to the critical threshold
of 10°C. This is illustrated in Figure 3 (left), which shows
the temperature changes in pixels that exceeded the classi-
fication threshold in the 1987 2006 period compared to the
earlier (1901 30) period. Figure 3 (right) also indicates that
below elevations of about 3000 m, the mean temperature of
the warmest month today is quite near the critical 10°C
threshold, and so one would expect that a continuation of
the recent warming trends will cause further reductions in
the areal coverage of this climate type.
[
6] We further illustrate the warming at high elevations in
the West in Figure 4, which show the distribution of linear
trends for annual mean minimum and maximum temper-
atures as a function of ground elevation for areas where the
highest mountain regions in the West are located. The
results are presented as distributions of linear trends com-
puted for successive 250 m elevation bins. In all three cases,
the trend in minimum average temperature increases with
elevation, while the trends in the mean maximum temper-
ature increases with height only in the Colorado Rockies.
The trend in the other two mountain ranges is positive, but
rather more uniform with height. We also note that precip-
itation trends in the West over the period 1979 2006 (not
shown), using a data set of snow water equivalent values
from snow-depth measurements together with collocated
PRISM data show declines during this time. The drier
conditions prevalent over the last few decades in the West
would also have contributed to the decline in the areal
extent of the alpine tundra in this region.
[
7] We also examined spring season (MarchMa y,
MAM) temperature trends for the same three regions
because generally the largest climate trends and impacts
signals in the West reported in the literature are associated
with this transition season, and because a connection can be
made between amplification of the warming trends in the
West with elevation and surface changes related to dimin-
ished western snow packs [Mote et al., 2005]. The results
Figure 2. Distribution of Ko¨ppen classification ‘E’
(tundra climates, E-T) corresponding to the ‘alpine tundra’
climate in the western United States considered in this
study. (top) Areas classified as K-E for the period 1901
1930. (bott om) Same as for Figure 2 (top), except for the
period 1987 2006. These areas are colored red and
comprise all the 4-km pixels in the PRISM data that are
classified as E-T.
Figure 1. Time series of annual mean temperature
anomalies (°C) for the western United States, 1901 2006.
(top) Values computed from divisional data (blue curve)
available from the National Climatic Data Center, NOAA in
Asheville, NC, and from PRISM. (bottom) Difference
between the two curves above. The mean di fference
between the two curves in the period 1901 1930 (1987
2006) is 0.2°C (0.05° C).
L18707 DIAZ AND EISCHEID: CLIMATIC TYPE SHIFT IN THE WEST L18707
2of4
(not shown because of space limitations) indicate that for
the period 19792006, significant increases in MAM max-
imum and minimum temperatures have taken place, with
mean annual minimums displaying substantial amplification
with altitude. Annual average maximums, however, do not
generally exhibit this vertical amplification of the warming.
3. Summary and discussion
[8] Average temperature in the western United States has
risen considerably in the last 20 years—about 0.6°C. This
period is the warmest such interval i n the instrumental
record. Based on the PRISM data set, the warming has
generally been accentuated at the higher elevations (above
about 2000 m) where trends in excess of 1°C are calculated.
Using the Ko¨ppen climate classification system, an area of
the western US of around 20,000 km
2
was typed as alpine
tundra for the period 190130. In contrast, for the period
1987 2006, the same climate type covered only 336 4-km
pixels—a decline in area coverage of about 73%.
[
9] The U.S. Climate Change Science Program recently
took up the question of whether there was agreement
between observations and climate model simulations on
the nature of temperature changes at the surface and in
the troposphere [Karl et al., 2006]. In particular, for global
average temperature, 1979 through 2004, the report con-
cluded that they could not state definitively whether trends
at the surface were different from those in the troposphere.
The results presented here are for the mountainous regions
of the western United States since 1979, where studies have
documented significant changes to snow packs and the
timing of the seasonal melting [e.g., Stewart et al., 2005;
Mote et al., 2005], and where these hydroclimatic changes
have been associated with significant veget atio n-related
impacts [cf. Breshears et al., 2005].
[
10] We have focused here on one particular western U.S.
ecotone—the alpine tundra—as defined by a simple metric,
namely the mean temperature of the warmest month. A
range of different observations indicate that environ mental
aspects of global warming are beginning to show, with
significant impacts already documented in a number of
phenological, biological, and hydrological indicators in
the West. These include earlier blossoming of shrubs in
spring, massive infestations of western forests by insect
pests, intensified wildfire seasons, and earlier spring runoff.
Our results suggest that the prevailing climate associated
with alpine tundra landscapes in the West may soon be out
of equilibrium with present climate and may largely
disappear.
References
Bradley, R. S., F. T. Keimig, and H. F. Diaz (2004), Projected temperature
changes along the American cordillera and the planned GCOS network,
Geophys. Res. Lett., 31, L16210, doi:10.1029/2004GL020229.
Breshears, D. D., et al. (2005), Regional vegetation die-off in response to
global-cha nge-type dro ught, Proc. Natl. Acad. Sci. U. S. A., 102(42 ),
15,144 15,148.
Cayan, D. R., S. A. Kammerdiener, M. D. Dettinger, J. M. Caprio, and D. H.
Peterson (2001), Changes in the onset of spring in the western United
States, Bull. Am. Meteorol. Soc., 82, 399 415.
Daly, C., W. P. Gibson, G. H. Taylor, G. L. Johnson, and P. Pasteris (2002),
A knowledge-based approach to the statistical mapping of climate, Clim.
Res., 22, 99 113.
Dang, H., N. P. Gillett, A. J. Weaver, and F. W. Zwiers (2007), Climate
change detection over different land surface vegetation classes, Int. J.
Climatol., 27, 211 220.
Diaz, H. F., and N. E. Graham (1996), Recent changes in tropical freezing
heights and the role of sea surface temperature, Nature, 383, 152 155.
Diaz, H. F., J. K. Eischeid, C. Duncan, and R. S. Bradley (2003), Variability
of freezing levels, melting season indicators, and snow cover for selected
high-elevation and continental regions in the last 50 years, Clim. Change,
59, 33 52.
Fraedrich, K., F.-W. Gerstengarbe, and P. C. Werner (2001), Climate shifts
during the last century, Clim. Change, 50, 405 417.
Karl, T. R., S. J. Hassol, C. D. Miller, and W. L. Murray (Eds.) (2006),
Temperature trends in the lower atmosphere: Steps for understanding and
reconciling differences, Syn. Assess. Prod. 1.1, 180 pp., Clim. Change Sci.
Program and the Subcomm. on Global Change Res., Washington, D. C.
Kottek, M., J. Grieser, C. Beck, B. Rudolf, and F. Rubel (2006), World map
of the Ko¨ppen-Geiger climate classification updated, Meteorol. Z., 15,
259 263.
Figure 3. (left) Distribution of mean annual temperature of the warmest month for pixels that were classified as ‘alpine
tundra’ in 19011930 (blue curve) and for the subset of pixels that exceeded the 10°C threshold, and hence were no longer
classified as such in 19872006. (right) Temperature distribution of pixels that are still classified as alpine tundra in both
periods. Note the elimination of this category at t he lowest elevation bin. Solid dots represent the median value in each
250-meter bin.
L18707 DIAZ AND EISCHEID: CLIMATIC TYPE SHIFT IN THE WEST L18707
3of4
Liu, X., and B. Chen (2000), Climatic warming in the Tibetan Plateau
during recent decades, Int. J. Climatol., 20, 1729 1742.
Lohmann, R., R. Sausen, L. Bengtsson, U. Cubasch, J. Perlwitz, and
E. Roeckner (1993), The Ko¨ppen climate classification as a diagnostic
tool for general circulation models, Clim. Res., 3, 177 193.
Mote, P. W. (2003), Trends in temperature and precipitation in the Pacific
Northwest during the twentieth century, Northwest Sci., 77, 271 282.
Mote, P. W., A. F. Hamlet, M. P. Clark, and D. P. Lettenmaier (2005),
Declining mountain snowpack in western North America, Bull. Am. Me-
teorol. Soc., 86, 39 49.
Regonda, S., B. Rajagopalan, M. Clark, and J. Pitlick (2005), Seasonal
cycle shifts in hydr oclimatology over the western U.S., J. Clim., 18,
372 3384.
Spencer, R. W., J. R. Christy, W. D. Braswell, and W. B. Norris (2006),
Estimation of tropospheric temperature trends from MSU channels 2 and 4,
J. Atmos. Oceanic Technol., 23, 417 423.
Stewart, I. T., D. R. Cayan, and M. D. Dettinger (2005), Changes toward
earlier streamflow timing across western North America, J. Clim., 18,
11361155.
Westerling, A. L., H. G. Hidalgo, D. R. Cayan, and T. W. Swetnam (2006),
Warming and earlier spring increase in U.S. forest wildfire activity,
Science, 313, 940 943.
H. F. Diaz and J. K. Eischeid, NOAA Earth System Research Laboratory,
325 Broadway, Boulder, CO 80305, USA. (henry.f.diaz@noaa.gov)
Figure 4. Distribution of annual mean minimum and maximum temperature trends expressed as total trend for the period
1979 2006, in °C, calculated and plotted in successive 250-meter intervals. The median of the distribution of linear trends
is plotted along with the approximate 5 and 95% cumulative distribution values (small crosses). (top) Colorado Rockies,
(middle) Pacific Northwest, (bottom) Sierra Nevada mountains.
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