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Diving Beetles (Coleoptera: Dytiscidae, Noteridae) of the Southwestern Anatolian Region of Turkey

Authors:
Suat Kiyak at Gazi University
Suat Kiyak
Mustafa Cemal DARILMAZ at Aksaray University
Mustafa Cemal DARILMAZ
Ali Salur at Hitit University
Ali Salur
Savaş Canbulat at Kastamonu University
Savaş Canbulat
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Abstract and Figures

Thirty species of Dytiscidae and one species of Noteridae were collected from 80 sampling stations in various aquatic habitats of the southwestern Anatolian region of Turkey (provinces of Antalya, Aydin, Afyon, Burdur, Denizli, Isparta and Muğla) between 2000 and 2002. Species distributions within Turkey are presented as a catalogue of provincial records. Seventeen species are here recorded for the first time from the southwestern Anatolian region. The presence of the three following species in Turkey is confirmed: Dytiscus semisulcatus MULLER, Eretes griseus (FABRICIUS), Hydaticus ponticus SHARP. Several species known previously from only a few localities in Turkey have been shown to have much wider distributions in the country. In accordance with the present study and literature data 49 species of Dytiscidae and 3 species of Noteridae are known from the southwestern Anatolian region.
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_____________ Mun. Ent. Zool. Vol. 2, No. 1, January 2007___________
103
DIVING BEETLES (COLEOPTERA: DYTISCIDAE,
NOTERIDAE) OF THE SOUTHWESTERN ANATOLIAN
REGION OF TURKEY
Suat Kıyak*, Mustafa Darılmaz*, Ali Salur**
and Savaş Canbulat***
*Gazi University, Sciences and Arts Faculty, Department of Biology, 06500-
Ankara/TURKEY. E-mail: skiyak@gazi.edu.tr, mdarilmaz@yahoo.com
**Hitit University, Sciences and Arts Faculty, Department of Biology, 19030,
Çorum/TURKEY E-mail: alisalur@gmail.com
***Sakarya University, Sciences and Arts Faculty, Department of Biology,
Sakarya/TURKEY E-mail: scanbulat@sakarya.edu.tr
[Kıyak, S., Darılmaz, M., Salur, A. & Canbulat, S. 2007. Diving beetles
(Coleoptera: Dytiscidae, Noteridae) of the southwestern Anatolian region of Turkey.
Munis Entomology & Zoology 2 (1): 103-114]
ABSTRACT: Thirty species of Dytiscidae and one species of Noteridae were collected from
80 sampling stations in various aquatic habitats of the southwestern Anatolian region of
Turkey (provinces of Antalya, Aydın, Afyon, Burdur, Denizli, Isparta and Muğla) between
2000 and 2002. Species distributions within Turkey are presented as a catalogue of
provincial records. Seventeen species are here recorded for the first time from the
southwestern Anatolian region. The presence of the three following species in Turkey is
confirmed: Dytiscus semisulcatus MULLER, Eretes griseus (FABRICIUS), Hydaticus
ponticus SHARP. Several species known previously from only a few localities in Turkey
have been shown to have much wider distributions in the country. In accordance with the
present study and literature data 49 species of Dytiscidae and 3 species of Noteridae are
known from the southwestern Anatolian region.
KEY WORDS: Coleoptera, diving beetles, Dytiscidae, Noteridae, faunistics, Turkey
The Dytiscidae and Noteridae water beetles are commonly called the
diving beetles, since their hind legs are usually highly modified for
swimming. Both families are cosmopolitan in distribution; although in
contrast to the latter occur primarily in the tropical regions of the world
(Zalat et. al., 1999).
Thirtytwo species of Dytiscidae and three species of Noteridae have
been recorded from the studied region in the following published
works: Balfour-Browne (1963), Guéorguiev (1968, 1981), Fery (1991),
Fery & Nilsson (1993), Shaverdo (2004), Barclay et al. (2001), and Fery
et al. (2001, 2005).
The purpose of this paper is to present a checklist of the diving
beetles of the southwestern Anatolian region of Turkey, combining
literature data and the results of our collecting trips between 2000-
2002.
MATERIALS AND METHOD
This study is based on 208 specimens of adult dytiscids and noterids
collected in the southwestern Anatolian region of Turkey (Fig. 1;
_____________ Mun. Ent. Zool. Vol. 2, No. 1, January 2007___________
104
Antalya, Aydın, Afyon, Burdur, Denizli, Isparta and Muğla) between
2000 and 2002. Specimens were collected from various aquatic
habitats, using a ladle or dip net having a 1 mm mesh size. The beetles
were killed in 70 % ethanol and in the laboratory were cleaned of debris
with a small paintbrush. Aedeagophore was dissected under the
stereomicroscope and left in 10 % KOH solution for about 1–2 hours.
Samples were identified using keys by Zaitzev (1972), Franciscolo
(1979) and Nilsson & Holmen (1995). Materials have been deposited in
the Zoological Museum of the Gazi University (=GUZM), Ankara,
Turkey.
In the checklist nomenclature is based on Nilsson (2005, 2006).
Family NOTERIDAE
Subfamily Noterinae Thomson, 1860
Tribe Noterini Thomson, 1860
Canthydrus diophthalmus (Reiche & Saulcy, 1855)
Literature records from Turkey: Antalya (Barclay et al., 2001).
Noterus clavicornis (De Geer, 1774)
Material examined: Antalya: 1 ex. Kalkan (Yeşilköy), 36º 17’ N 29º 19’ E, 8
m, 27.05.2001; Aydın: 1 ex. Söke (Avşar-Azap Lake), 37º 35’ N 27º 26’ E, 37 m,
23.06.2001.
Literature records from Turkey: Aksaray, Ankara, Balıkesir, Bilecik, Bolu,
Isparta, İzmir, Kayseri, Konya, Manisa (Balfour-Browne, 1963; Guéorguiev,
1968, 1981; Darılmaz & Kıyak, 2006).
Remarks: New to Antalya and Aydın.
Noterus crassicornis (O.F.Müller, 1776)
Literature records from Turkey: Adana, Isparta (Guéorguiev, 1981).
Family DYTISCIDAE
Subfamily Agabinae Thomson, 1867
Tribe Agabini Thomson, 1867
Agabus (Acatodes) congener (Thunberg, 1794)
Material examined: Antalya: 1 ex., Beşkonak (Köprü river), 37º 07’ N 31º 12’
E, 112 m, 25.06.2000; Denizli: 1 ex., Çameli (Güre Mountain, vicinity of
Kınıkyeri), 37º 11’ N 29º 25’ E, 1580 m, 27.06.2001.
Literature records from Turkey: Kars (Guéorguiev, 1981).
Remarks: This species is new to the south and west Anatolia regions.
Agabus (Agabus) zimmermanni Scholz, 1920
Literature records from Turkey: Aydın (Guéorguiev, 1981).
Agabus (Gaurodytes) biguttatus (Olivier, 1795)
Literature records from Turkey: Adana, Aksaray, Ankara, Bilecik, Bursa,
Çankırı, Elazığ, Gaziantep, Gümüşhane, Isparta, İzmir, Kastamonu, Sakarya,
Trabzon, Yozgat (Guéorguiev, 1981; Darılmaz & Kıyak, 2006).
_____________ Mun. Ent. Zool. Vol. 2, No. 1, January 2007___________
105
Agabus (Gaurodytes) bipustulatus (Linnaeus, 1767)
Material examined: Antalya: 1 ex., Korkuteli, (vicinity of Söğütcük), 37º 02’
N 30º 24’ E, 904 m, 26.05.2002; Denizli: 1 ex., Çivril (Gökgöl), 38º 11’ N 30º
03’ E, 835 m, 20.04.2002; Isparta: 1 ex., Aksu (Anamas plateau, waterbed), 37º
49’ N 31º 13’ E, 1896 m, 14.07.2000; 1 ex., Yalvaç (Sultan mountains-brook),
38º 15’ N 31º 22’ E, 1565 m, 15.09.2000; 1 ex., Sütçüler (between İncidere and
Kasımlar, spring water), 37º 34’ N 31º 07’ E, 1450 m, 23.08.2001; Muğla: 2 ex.,
between Yeniköy and Makıköy, 37º 02’ N 27º 42’ E, 350 m, 17.04.2000; 12 ex.,
Köyceğiz (Yayla village, Gölgeli Mountains), 37º 03’ N 28º 47’ E, 1730 m,
17.07.2000; 1 ex., Ula (Gölcük town, accumulation of brook), 37º 07’ N 28º 31’
E, 690 m, 18.07.2000.
Literature records from Turkey: Adana, Ankara, Bursa, Isparta, İzmir,
Kars, Kastamonu, Kayseri, Konya, Trabzon, Van (Guéorguiev, 1968, 1981).
Remarks: New to Antalya, Denizli and Muğla.
Agabus (Gaurodytes) conspersus (Marsham, 1802)
Material examined: Afyon: 1 ex., Başmakçı (Akpınar, the brook flows to
Acıgöl), 37º 50’ N 29º 59’ E, 850 m, 20.06.2002; Antalya: 3 ex., Korkuteli (the
brook flows to Korkuteli dam), 37º 05’ N 30º 08’ E, 1074 m, 22.04.2001; 2 ex.,
Elmalı (Armutlu village, brook), 36º 33’ N 29º 43’ E, 1130 m, 22.04.2001;
Burdur: 1 ex., Karamanlı dam and its around), 37º 25’ N 29º 49’ E, 1200 m,
20.06.2002; 1 ex., Çavdır (Kozağacı), 37º 07’ N 29º 41’ E, 1035 m, 22.06.2002;
Denizli: 1 ex., Buldan (Süleymanlı lake), 38º 03’ N 28º 46’ E, 1175 m,
19.05.2001; 1 ex., Çameli (Güre mountain, vicinity of Kınıkyeri), 37º 11’ N 29º
25’ E, 1580 m, 27.06.2001; Isparta: 1 ex., Yalvaç (Sultan Mountains, between
Bağkonak and Cankurtaran, the brook flows to Akşehir lake), 38º 14’ N 31º 21’
E, 1657 m, 27.06.2000; 1 ex., between Aksu and Yenişarbademli , 37º 43’ N 31º
16’ E, 1753 m, 14.07.2000; 2 ex., between Gelendost and Afşar (the brook flows
Eğridir lake), 38º 07’ N 30º 59’ E, 65 m, 20.06.2001.
Literature records from Turkey: Adana, Bursa, Isparta, İzmir, Konya
(Guéorguiev, 1968, 1981).
Remarks: New to Afyon, Antalya, Burdur and Denizli.
Agabus (Gaurodytes) didymus (Olivier, 1795)
Material examined: Antalya: 1 ex., Korkuteli (the brook flows to Korkuteli
dam), 37º 05’ N 30º 08’ E, 1074 m, 22.04.2001; Isparta: 1 ex., between Aksu
and Karağı (Aksu stream and dam lake), 37º 47’ N 31º 06’ E, 1305 m,
28.06.2001.
Literature records from Turkey: Afyon, Balıkesir, İzmir, Manisa
(Guéorguiev, 1981).
Remarks: New to Antalya and Isparta.
Agabus (Gaurodytes) dilatatus (Brullé, 1832)
Literature records from Turkey: Adana, Ankara, Bursa, Gümüşhane,
Isparta, İzmir, İzmit, Konya, Rize, Trabzon, Van (Balfour-Browne, 1963;
Guéorguiev, 1968, 1981).
Agabus (Gaurodytes) faldermanni Zaitzev, 1927
Material examined: Isparta: 1 ex., Yalvaç (Sultan Mountains, between
Bağkonak and Cankurtaran, the brook flows to Akşehir lake), 38º 14’ N 31º 21’
E, 1657 m, 27.06.2000; 1 ex., Muğla: Milas (between Taşlı and Kızılağaç), 37º
18’ N 27º 39’ E, 50 m, 17.04.2000.
_____________ Mun. Ent. Zool. Vol. 2, No. 1, January 2007___________
106
Literature records from Turkey: Diyarbakır, Konya, Nevşehir
(Guéorguiev, 1968, 1981; Nilsson, 1992).
Remarks: This species is new to the south and west Anatolia regions.
Agabus (Gaurodytes) guttatus guttatus (Paykull, 1798)
Material examined: Antalya: 1 ex., Korkuteli (Öküzgözü I dam), 36º 53’ N
30º 02’ E, 1200 m, 22.04.2001; 2 ex., Elmalı (vicinity of Çobanisa village), 36º
52’ N 30º 01’ E, 1170 m, 26.05.2002; Denizli: 2 ex., Çameli (Güre mountain,
vicinity of Kınıkyeri), 37º 11’ N 29º 25’ E, 1580 m, 27.06.2001.
Literature records from Turkey: Bolu, Kastamonu (Guéorguiev, 1981).
Remarks: This species is new to the south and west Anatolia regions.
Agabus (Gaurodytes) nebulosus (Forster, 1771)
Material examined: Afyon: 1 ex., Dinar (Elmalı village, Pınarbaşı lake), 38º
04’ N 30º 16’ E, 1113 m, 22.06.2000; Antalya: 1 ex., Elmalı (Göltarla, Avlan
lake), 37º 35’ N 29º 57’ E, 1200 m, 26.05.2002; Aydın: 1 ex., Söke (Yenidoğan
road), 37º 37’ N 27º 22’ E, 15.04.2000; Burdur: 3 ex., Bucak (Onaç dam), 37º
39’ N 30º 34’ E, 831 m, 25.05.2002; Denizli: 1 ex., Çivril (between İnceköy and
Seraserli), 38º 15’ N 29º 47’ E, 845m, 28.05.2002; Muğla: 8 ex., Yatağan
(vicinity of Bozarmut, Alişar brook), 37º 17’ N 28º 10’ E, 400 m, 27.05.2002.
Literature records from Turkey: Adana, Bursa, Isparta, İstanbul, İzmir,
Sinop, Trabzon (Guéorguiev, 1981).
Remarks: New to Afyon, Antalya, Aydın, Burdur, Denizli and Muğla.
Ilybius fuliginosus fuliginosus (Fabricius, 1792)
Material examined: Isparta:2 ex., Aksu (between Aksu and Yılanlı), 37º 48’
N 31º 00’ E, 1215 m, 28.06.2001; Konya: 2 ex., Taşkent (Belpınarı), 36º 54’ N
32º 20’ E, 1738 m, 25.05.2001.
Literature records from Turkey: Bingöl, Kars, Mersin (Guéorguiev, 1981).
Remarks: New to the southwestern Anatolian region.
Ilybius jaechi (Fery & Nilsson, 1993)
Literature records from Turkey: Aydın, Tekirdağ (Fery & Nilsson, 1993).
Platambus maculatus (Linnaeus, 1758)
Material examined: Burdur: 2 ex., Karamanlı (vicinity of Kılavuz village, the
brook flows to Karataş lake), 37º 22’ N 29º 52’ E, 1062 m, 23.06.2000; Muğla:
2 ex., Fethiye (south of Karahasan, Esen brook), 36º 56’ N 29º 38’ E, 1390 m,
20.08.2001; 3 ex., Fethiye (vicinity of Seki, Esen brook), 36º 49’ N 29º 33’ E,
1130 m, 20.08.2001; 10 ex., Kemer (Seki brook), 36º 50’ N 29º 34’ E, 1150 m,
13.09.2001.
Literature records from Turkey: Adana, Bilecik, Kars (Guéorguiev, 1981).
Remarks: New to the southwestern Anatolian region.
Subfamily Colymbetinae Erichson, 1837
Tribe Colymbetini Erichson, 1837
Colymbetes fuscus (Linnaeus, 1758)
Material examined: Aydın: 2 ex., Söke (Yenidoğan road), 37º 37’ N 27º 22’
E, 15.04.2000; 1 ex., Söke (Büyük Menderes river), 37º 30’ N 27º 20’ E, 19 m,
23.06.2002; Burdur: 1 ex., Yeşilova (Bayındır village), 37º 39’ N 29º 45’ E, 1030
_____________ Mun. Ent. Zool. Vol. 2, No. 1, January 2007___________
107
m, 20.06.2002; Konya: 1 ex., between Taşkent and Alanya (20 km), 36º 51’ N
32º 31’ E, 1740 m, 20.04.2001.
Literature records from Turkey: Afyon, Aksaray, Erzurum, İzmir,
Nevşehir (Guéorguiev, 1981; Darılmaz & Kıyak, 2006).
Remarks: New to Aydın, Burdur and Konya.
Subfamily Dytiscinae Leach, 1815
Tribe Aciliini Thomson, 1867
Graphoderus cinereus (Linnaeus, 1758)
Material examined: Afyon: 2 ex., Dinar (vicinity of Elmalı village, Pınarbaşı
lake), 38º 04’ N 30º 16’ E, 1113 m, 22.06.2000; 1 ex., Dinar (Elderesi, Pınarbaşı
lake), 38º 04’ N 30º 16’ E 1020 m 28.05.2002.
Literature records from Turkey: Ağrı Mountain (Guéorguiev, 1981).
Remarks: This species is new to the south and west Anatolia regions.
Tribe Cybistrini Sharp, 1882
Cybister (Cybister) tripunctatus lateralis (Fabricius, 1798)
Material examined: Aydın: 1 ex., Söke (Avşar, Azap lake), 37º 35’ N 27º 26’
E, 37 m, 23.06.2001.
Literature records from Turkey: Adana, İzmir, Mersin (Guéorguiev, 1981).
Remarks: New to the southwestern Anatolian region.
Cybister (Scaphinectes) lateralimarginalis torquatus (Fischer von
Waldheim, 1829)
Material examined: Denizli: 4 ex., Çivril (vicinity of Beydilli village, Işıklı
lake), 38º 11’ N 30º 03’ E, 920 m, 22.06.2000; 1 ex., Çivril (Bucak, Işıklı lake),
38º 14’ N 29º 51’ E, 838 m, 26.06.2001; 1 ex., Çardak (Gemiş, Acıgöl), 37º 46’ N
29º 50’ E, 860 m, 20.06.2002; Burdur: 2 ex., Yazır lake, 36º 59’ N 29º 44’ E,
1515 m, 23.06.2000.
Literature records from Turkey: Adana, Eskişehir, İzmir, Konya, Mersin,
Van (Guéorguiev, 1981).
Remarks: New to the southwestern Anatolian region.
Tribe Dytiscini Leach, 1815
Dytiscus circumflexus Fabricius, 1801
Material examined: Aydın: 1 ex., Söke (Yenidoğan road), 37º 37’ N 27º 22’ E,
15.04.2000.
Literature records from Turkey: Adana, Balıkesir, Iğdır, Isparta, Kars,
Kütahya (Guéorguiev, 1981).
Remarks: New to Aydın.
Dytiscus dimidiatus Bergsträsser, 1778
Literature records from Turkey: Manisa (Guéorguiev, 1981).
Remarks: This species is also known in Samsun province (ponds near the
village of Çarşamba, 11.07.1992, 2 specimens, leg. M. Toledo) from Turkey
(pers. comm.).
_____________ Mun. Ent. Zool. Vol. 2, No. 1, January 2007___________
108
Dytiscus marginalis marginalis Linnaeus, 1758
Material examined: Antalya: 1 ex., Beşkonak (between Çaltepe and
Değirmenözü, Kanlı brook), 37º 20’ N 31º 13’ E, 492 m, 26.06.2000; 1 ex.,
Alanya (Güney village, Zeytinbükü brook), 36º 40’ N 31º 53’ E, 125 m,
11.07.2000; Denizli: 2 ex., Çivril (Gökgöl), 38º 11’ N 30º 03’ E, 835 m,
20.04.2002; Isparta: 4 ex., Yalvaç (Sultan mountains), 38º 15’ N 31º 22’ E,
1565 m, 15.09.2000; Konya: 1 ex., Hadim (vicinity of Ardıçpınarı fountain), 36º
53’ N 32º 23’ E, 1930 m, 11.07.2001; Muğla: 3 ex., Ula (Gölcük), 37º 07’ N 28º
31’ E, 690 m, 18.07.2000.
Literature records from Turkey: Adapazarı, Ankara, Iğdır, Kars, Kayseri,
Nevşehir (Guéorguiev, 1968, 1981).
Remarks: This species is new to the south and west Anatolia regions.
Dytiscus semisulcatus O.F. Müller, 1776
Material examined: Isparta: 1 ex., Aksu (Karağı village), 37º 45’ N 31º 07’ E,
1210 m, 10.08.2000; Muğla: 2 ex., Köyceğiz (Yayla village, Gölgeli mountains),
37º 03’ N 28º 47’ E, 1730 m, 17.07.2000.
Literature records from Turkey: Without detailed locality data
(Guéorguiev, 1981; Nilsson, 2005).
Remarks: This species is confirmed for Turkey and the first detailed records
are given.
Tribe Eretini Crotch, 1873
Eretes griseus (Fabricius, 1781)
Material examined: Antalya: 1 ex., Serik (Kırbaş village, water
accumulation), 37º 09’ N 30º 49’ E, 57 m, 19.08.2001.
Literature records from Turkey: Without detailed locality data (Miller,
2002; Nilsson, 2005).
Remarks: This species is confirmed for Turkey and the first detailed records
are given.
Tribe Hydaticini Sharp, 1882
Hydaticus (Guignotites) ponticus Sharp, 1882
Material examined: Aydın: 1 ex., Nazilli (Esenköy, Akçay), 37º 47’ N 28º 18’
E, 75 m, 12.09.2001; Burdur: 1 ex., Çavdır (Yamadı village), 37º 07’ N 29º 36’ E,
960 m, 22.06.2002.
Literature records from Turkey: Without detailed locality data
(Guéorguiev, 1981; Nilsson, 2005).
Remarks: This species is confirmed for Turkey and the first detailed records
are given.
Hydaticus (Hydaticus) transversalis laevisculptus Zaitzev, 1910
Material examined: Denizli: 1 ex., Acıpayam (Yeniköy, water channel), 37º
22’ N 29º 25’ E, 863 m, 07.08.2000.
Literature records from Turkey: Adana, Kütahya (Guéorguiev, 1981).
Remarks: New to the southwestern Anatolian region.
_____________ Mun. Ent. Zool. Vol. 2, No. 1, January 2007___________
109
Subfamily Hydroporinae Aubé, 1836
Tribe Bidessini Sharp, 1882
Bidessus anatolicus anatolicus Wewalka, 1972
Literature records from Turkey: Adana, Antalya (Guéorguiev, 1981;
Biström, 1987).
Bidessus calabricus Guignot, 1957
Literature records from Turkey: Ankara, Antalya, Çanakkale, İstanbul,
İzmir, İzmit, Muğla (Fery, 1991).
Bidessus delicatulus (Schaum, 1844)
Literature records from Turkey: Ankara, Muğla, İzmir (Guéorguiev, 1981;
Fery, 1991).
Bidessus nasutus Sharp, 1887
Literature records from Turkey: Konya, Isparta (Guéorguiev, 1981).
Hydroglyphus geminus (Fabricius, 1792)
Material examined: Muğla: 4 ex., border of Muğla (Gazeller, Akçay), 37º 20’
N 28º 43’ E, 440 m, 22.06.2001.
Literature records from Turkey: Adana, Aksaray, Ankara, Antalya, Aydın,
Balıkesir, Bolu, Bursa, Edirne, Gümüşhane, İçel, İzmir, Kastamonu, Kayseri,
Kilis, Konya, Manisa, Nevşehir, Tuz Gölü, Trabzon (Guéorguiev, 1981; Darılmaz
& Kıyak, 2006).
Remarks: New to Muğla.
Tribe Hydroporini Aubé, 1836
Deronectes parvicollis (Schaum, 1864)
Material examined: Isparta: 1 ex., Yalvaç (the northwest of Sultan
mountains), 38º 15' N 31º 18' E, 1438º m, 19.07.2001.
Literature records from Turkey: Adana, Bilecik, Erzurum, Niğde, Yozgat
(Guéorguiev, 1981; Fery et. al., 2001).
Remarks: New to the southwestern Anatolian region.
Deronectes sahlbergi Zimmermann, 1932
Literature records from Turkey: İzmir, Bilecik, Muğla (Guéorguiev, 1981;
Fery et. al., 2001).
Deronectes wittmeri Wewalka, 1971
Literature records from Turkey: Antalya (Guéorguiev, 1981).
Graptodytes veterator behningi Zaitzev, 1927
Literature records from Turkey: Afyon, Ankara, Balıkesir, Bingöl, Bursa,
Erzurum, Konya, İzmir (Guéorguiev, 1981).
Hydroporus bodemeyeri Ganglbauer, 1900
Literature records from Turkey: Adana, Adapazarı, Anlara, Bilecik, Bolu,
Isparta, İzmir, Muğla (Guéorguiev, 1981).
_____________ Mun. Ent. Zool. Vol. 2, No. 1, January 2007___________
110
Hydroporus kozlovskii Zaitzev, 1927
Literature records from Turkey: Antalya, Artvin, Bingöl, Erzincan,
Gümüşhane, Kars, Muş, Rize (Guéorguiev, 1981; Shaverdo, 2004).
Hydroporus marginatus (Duftschmid, 1805)
Material examined: Muğla: 1 ex., Köyceğiz (Yayla village, Gölgeli
mountains), 37º 03’ N 28º 47’ E, 1730 m, 17.07.2000.
Literature records from Turkey: Ankara, Kars, Konya, Sivas, Trabzon
(Guéorguiev, 1981).
Remarks: This species is new to the south and west Anatolia regions.
Hydroporus planus (Fabricius, 1781)
Material examined: Antalya: 1 ex., Elmalı (Armutlu village), 36º 33’ N 29º
43’ E, 1130 m, 22.04.2001.
Literature records from Turkey: Adana, Balıkesir, Bursa, Erzincan,
İstanbul, Kars, Trabzon (Guéorguiev, 1981).
Remarks: New to the southwestern Anatolian region.
Hydroporus pubescens (Gyllenhal, 1808)
Literature records from Turkey: Adana, Aksaray, Antalya, Bilecik, Bursa,
Erzincan, Gümüşhane, İzmir, Manisa, Niğde, Ordu, Sakarya, Trabzon
(Guéorguiev, 1981; Darılmaz & Kıyak, 2006).
Nebrioporus (Nebrioporus) suavis (Sharp, 1882)
Material examined: Antalya: 6 ex., İbradı (Ürünlü village, Altınbeşik cave),
37º 02’ N 31º 36’ E, 800 m, 03.08.2000; 1 ex., Kaş (exit road of Gömbe) 36º
30’ N 29º 35’ E, 1275 m, 05.08.2000; Isparta: 1 ex., Yalvaç (Sultan mountains,
Bağkonak), 38º 13’ N 31º 17’ E, 1260 m, 19.07.2001; Muğla: 1 ex., Dalaman
(between Yanıklar and İnlice), 22.05.2000.
Literature records from Turkey: Adana, Amasya, Ankara, Antalya,
Balıkesir, Bilecik, Bursa, Eskişehir, Gümüşhane, İstanbul, İzmir, Konya, Ordu,
Sakarya, Sinop, Trabzon (Guéorguiev, 1981).
Remarks: New to Isparta and Muğla.
Nebrioporus (Nebrioporus) turca (Seidlitz, 1887)
Material examined: Antalya: 1 ex., Finike (between Yazır and Kiltepe,
Akçay), 36º 34’ N 29º 57’ E, 21.05.2000; 1 ex., between Bağlıağaç village and
Kayadibi village, 22.05.2000; Isparta: 1 ex., Centrum, (vicinity of Güneyce) 37º
39’ N 30º 43’ E, 666 m, 23.08.2001; Konya: 1 ex., Hadim (vicinity of
Ardıçpınarı fountain), 36º 53’ N 32º 23’ E, 1930 m, 11.07.2001
Literature records from Turkey: Adana, Antalya, Elazığ, İzmir, Trabzon
(Guéorguiev, 1968, 1981).
Remarks: New to Isparta and Konya.
Oreodytes davisii davisii (Curtis, 1831)
Material examined: Antalya: 2 ex., Elmalı (Akçay, exit road of Avşar village),
36º 33’ N 29º 43’ E, 1116 m, 21.05.2000.
Literature records from Turkey: Erzurum (Erman & Erman, 2002).
Remarks: The first record of O. davisii davisii from Turkey was given by
Erman & Erman (2002). In this study the second record is given for Turkey.
Also this species is new to the south and west Anatolia regions.
_____________ Mun. Ent. Zool. Vol. 2, No. 1, January 2007___________
111
Scarodytes halensis halensis (Fabricius, 1787)
Material examined: Antalya: 4 ex., Alanya (Yeşilöz brook), 36º 22’ N 32º 11’
E, 20 m, 26.05.2001; 7 ex., between Gazipaşa and Kocadere, 36º 15’ N 32º 19’
E, 30 m, 26.05.2001; 1 ex., Alanya (Demirtaş, Serde stream), 36º 24’ N 32º 10’
E, 10 m, 27.06.2002; 8 ex., Gazipaşa (Gazipaşa stream), 36º 15’ N 32º 19’ E, 25
m, 27.06.2002; Isparta: 1 ex., Sütçüler (the south of Sipahiler), 37º 38’ N 30º
58’ E, 1209 m, 26.06.2000; 1 ex., Sütçüler (Sipahiler, Taşlıkkır brook), 37º 38’
N 30º 59’ E, 1185 m, 15.07.2000; 6 ex., Yenişarbademli (Gedikli, edge of
Beyşehir lake), 37º 56’ N 31º 18’ E, 1140 m, 02.08.2000; 3 ex., Aksu (Karağı
village, brook), 37º 45’ N 31º 07’ E, 1220 m, 14.09.2000.
Literature records from Turkey: Afyon, Aksaray, Ankara, Bursa, Eskişehir,
Gümüşhane, Isparta, İçel, İzmir, Kayseri, Van, Yozgat (Guéorguiev, 1968, 1981;
Darılmaz & Kıyak, 2006).
Remarks: New to Antalya.
Tribe Hydrovatini Sharp, 1882
Hydrovatus cuspidatus (Kunze, 1818)
Literature records from Turkey: Afyon, Adana (Guéorguiev, 1981).
Tribe Hygrotini Portevin, 1929
Hygrotus (Coelambus) confluens (Fabricius, 1787)
Literature records from Turkey: Adana, Ankara, Bolu, Eskişehir, Isparta,
İzmir, Kayseri, Konya, Mersin, Nevşehir (Guéorguiev, 1981).
Hygrotus (Coelambus) lernaeus (Schaum, 1857)
Literature records from Turkey: Adana, Afyon, Ankara, Antalya, Aydın,
Diyarbakır, Elazığ, Erzurum, Eskişehir, Kayseri, Kocaeli, Konya, Muğla, Niğde,
Samsun, Sivas, Van, Yozgat (Guéorguiev, 1968, 1981; Fery et. al., 2005).
Hygrotus (Hygrotus) inaequalis (Fabricius, 1777)
Literature records from Turkey: Bolu, Erzurum, Isparta, Konya, Manisa,
Nevşehir (Balfour-Browne, 1963; Guéorguiev, 1981).
Subfamily Laccophilinae Gistel, 1856
Tribe Laccophilini Gistel, 1856
Laccophilus hyalinus hyalinus (De Geer, 1774)
Material examined: Antalya: 1 ex., Finike (between Yazır and Kiltepe,
Akçay), 36º 34’ N 29º 57’ E, 21.05.2000; 5 ex., Elmalı (Akçay, exit road of Avşar
village), 36º 33’ N 29º 43’ E, 1116 m, 21.05.2000; 4 ex., Manavgat (Boztepe,
Alara stream), 36º 39’ N 31º 39’ E, 20 m, 21.04.2001; 1 ex., Alanya (Yeşilöz
stream), 36º 22’ N 32º 11’ E 20 m, 26.05.2001; 1 ex., Konyaaltı (Boğaz stream),
36º 51’ N 30º 37’ E, 15 m, 26.05.2001; 1 ex., Kalkan (Yeşilköy), 36º 17’ N 29º 19’
E, 8 m, 27.05.2001; 1 ex., Korkuteli (Söğütçük village), 37º 01’ N 30º 20’ E, 930
m, 26.06.2002; 1 ex., Manavgat (Hocalar village, Sarısu), 36º 52’ N 31º 15’ E, 22
m, 26.06.2002; Burdur: 2 ex., Karamanlı (Kılavuz village), 37º 22’ N 29º 52’ E,
1062 m, 23.06.2000; 4 ex., Gölhisar (Gölhisar lake), 37º 07’ N 29º 36’ E, 960
m, 13.09.2001; Isparta: 1 ex., Aksu (Karağı village, brook), 37º 45’ N 31º 07’ E,
1220 m, 14.09.2000; Konya: 1 ex., between Taşkent and Alanya (20km), 36º 51’
N 32º 31’ E, 1740 m, 20.04.2001.
_____________ Mun. Ent. Zool. Vol. 2, No. 1, January 2007___________
112
Literature records from Turkey: Adana, Afyon, Aksaray, Aydın, Bolu,
Isparta, İzmir, Kırşehir, Kilis, Manisa, Sakarya (Guéorguiev, 1981; Darılmaz &
Kıyak, 2006).
Remarks: New to Antalya, Burdur and Konya.
Laccophilus minutus (Linnaeus, 1758)
Material examined: Antalya: 1 ex., between Bağlıağaç village and Kayadibi
village, 22.05.2000; 1 ex., Manavgat (Boztepe, Alara stream), 36º 39’ N 31º 39’
E, 20 m, 21.04.2001; 1 ex., between Gazipaşa and Kocadere 36º 15’ N 32º 19’ E,
30 m, 26.05.2001; Aydın: 1 ex., Söke (Avşar, Azap lake), 37º 35’ N 27º 26’ E, 37
m, 23.06.2001; Burdur: 2 ex., Karamanlı (Karamanlı dam), 37º 24’ N 29º 49’ E,
1206 m, 27.06.2001; 1 ex., Gölhisar (Gölhisar lake), 37º 07’ N 29º 36’ E, 960 m,
20.08.2001; Denizli: 1 ex., Buldan (Süleymanlı lake), 38º 03’ N 28º 46’ E, 1175
m, 19.05.2001; Isparta: 1 ex., Eğridir (Kovada dam), 37º 20’ N 30º 52’ E, 920
m, 21.06.2001; 1 ex., Gölcük lake, 37º 43’ N 30º 30’ E, 1410 m, 28.06.2001.
Literature records from Turkey: Adana, Afyon, Aksaray, Ankara, Aydın,
Balıkesir, Bolu, Bursa, Gümüşhane, Isparta, İzmir, Kayseri, Konya, Manisa,
Sinop, Sivas, Trabzon (Guéorguiev, 1981; Darılmaz & Kıyak, 2006).
Remarks: New to Antalya, Burdur and Denizli.
Laccophilus poecilus Klug, 1834
Material examined: Antalya: 1 ex., Kalkan (Yeşilköy), 36º 17’ N 29º 19’ E, 8
m, 27.05.2001; Aydın: 1 ex., Söke (Avşar, Azap lake), 37º 35’ N 27º 26’ E, 37 m,
23.06.2001.
Literature records from Turkey: Adana, Antalya, Aydın, Bolu, Isparta,
İzmir, Konya, Manisa, Nevşehir (Guéorguiev, 1981).
ACKNOWLEDGEMENTS
We thank Dr. M. Toledo (Italy) for contribution on distributional
data of D. dimidiatus.
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Fig. 1. Map of the southwestern Anatolian region of Turkey (delimited
by bold line).
... No locality data (Zaitsev 1972;Guéorguiev 1981;Vondel 1992;Nardi 2001 (Zimmermann 1930;Balfour-Browne 1963;Guéorguiev 1968Guéorguiev , 1981Zaitsev 1972;Ienistea 1978;Franciscolo 1979;Holmen 1987;Özemsi and Önder 1988;Darilmaz and Kiyak 2006;Kiyak et al. 2007;Karaman et al. 2008;Incekara et al. 2009 (Guéorguiev 1981;Incekara et al. 2009). ...
... Family DYTISCIDAE Subfamily Agabinae Thomson, 1867 Tribe Agabini Thomson, 1867 Genus Agabus Leach, 1817 Subgenus Acatodes Thomson, 1859 Agabus bergi Zaitzev, 1913 Distribution in Turkey Erzurum (Erman et al. , 2007. ( (Guéorguiev 1981;Kiyak et al. 2007;Erman et al. 2007). ...
... Izmir (Guéorguiev 1981) Agabus zimmermanni Scholz, 1920 Distribution in Turkey Aydin (Zimmermann 1935;Zaitsev 1972;Guéorguiev 1981;Nilsson 1990;Shaverdo, 2002 (Zimmermann 1920(Zimmermann , 1930Bertrand 1928;Guéorguiev 1968Guéorguiev , 1981Ienistea 1978;Franciscolo 1979;Shaverdo 2002;Darilmaz and Kiyak 2006;Erman et al. 2007;Erman and Erman 2008;Karaman et al. 2008 (Zimmermann 1920(Zimmermann , 1934Bertrand 1928;Guéorguiev 1968Guéorguiev , 1981Ienistea 1978;Franciscolo 1979;Kiyak et al. 2007;Erman et al. 2007;Erman and Erman 2008;Karaman et al. 2008). ...
Checklist of Gyrinidae, Haliplidae, Noteridae and Dytiscidae of Turkey (Coleoptera: Adephaga)
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... 6), 10.09.1987, 5 ex. Distribution in Turkey: Burdur, Muğla (Kıyak et al, 2007), Ankara (Hızarcıoğlu et al, 2010), Black Sea Region (Topkara & Balık, 2010), Çorum , ...
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View
... Fery & Hosseinie 1998;Erman & Fery 2000;Fery et al. 2001;Hájek et al. 2005;Erman & Fery 2006;Fery & Erman 2009;Fery 2009) the predaceous water beetle fauna of Turkey is comparably rich and includes many very remarkable and rarely collected endemic species, especially in the genera Hydroporus Clairville, 1806 and Deronectes Sharp, 1882. Despite the fact that southern Turkey has been more seriously investigated in recent years for diving beetles (Kıyak et al. 2007), we report the discovery of an undescribed species of the Holarctic genus Hydroporus. The new species belongs to the puberulus species group sensu Nilsson (2001) and was collected in the Köprülü Canyon in southern Turkey. ...
... The new species belongs to the puberulus species group sensu Nilsson (2001) and was collected in the Köprülü Canyon in southern Turkey. According to present knowledge it is by far the smallest species of the genus recorded from Turkey (Erman & Erman 2008;Karaman et al. 2008;Kıyak et al. 2007). The species was found by the German coleopterist Jens Esser (Berlin, Germany) during a short field trip in spring 2004. ...
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View
... Distribution maps were compiled using data from the collection of the first author, The Global Biodiversity Information Facility (GBIF) and the following literature: BEDEL (1895), FERENCA & TAMUTIS (2015), FOSTER & TAYLOR (2006), FRANSISCOLO (1979), GABRIŠ (2011, KARAMAN et al. (2008), KIYAK et al. (2007), LAMRI et al. (2016, PEDERZANI & SCHIZZEROTTO (1998), PRZEWOŹNY et al. (2009), RYNDEVICH et al. (2014, SALAH & RÉGIL CUETO (2014) andVAFEI et al. (2009). The records on the maps are not exhaustive and only intended as indicative for the overall distribution of both species. ...
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Abstract: Agabus didymus (Olivier, 1795) and A. dilatatus (Brullé, 1832) are newly recorded for Croatia. The new records are given and the ecology and overall distribution of both species is mapped and discussed. With the addition of these two species, the number of species of Dytiscidae known from Croatia increases to 122. Résumé: Agabus didymus (Olivier, 1795) et A. dilatatus (Brullé, 1832), deux espèces nouvelles pour la Croatie sont rapportées. Pour chacune, nous discutons l’écologie et la distribution et les cartes de répartition sont commentées. Ces deux espèces portent le nombre de Dytiscidae de la faune Croate à 122. Samenvatting: Agabus didymus (Olivier, 1795) en A. dilatatus (Brullé, 1832) worden voor het eerst gemeld voor Kroatië. In dit artikel worden ecologie en verspreiding van beide soorten in kaart gebracht en bediscussieerd. De toevoeging van deze twee soorten brengt het totaal aantal soorten Dytiscidae in Kroatië op 122.
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... Distribution in Turkey: without detailed locality (Holmen;Vondel, 1991Vondel, , 1997Darılmaz and Kıyak, 2009a). (Marsham, 1802) Materials Distribution in Turkey: Çağıran, Daltaban, Gümüşhane, İskali, İzmir (Guéorguiev, 1981;Darılmaz and Kıyak, 2009a Distribution in Turkey: Aksaray, Ankara, Antalya, Aydın, Balıkesir, Bilecik, Bolu, Isparta, İzmir, Kayseri, Konya, Manisa, Trabzon (Balfour-Browne, 1963;Guéorguiev, 1981;Özesmi and Önder, 1988;Mermer et al., 1995;Darılmaz andKıyak, 2006a, 2009a;Karaman et al., 2008;Kıyak et al., 2007). Distribution in Turkey: Ankara, Artvin, Aydın, Balıkesir, Bolu, Bursa, Erzurum, Isparta, İzmir, Kocaeli, Konya, Manisa, Rize, Sinop, Sivas, Trabzon, Taurus Mountains (Guéorguiev, 1981;Erman et al., 2007;Karaman et al., 2008;Darılmaz and Kıyak, 2009a). ...
Contribution to the Knowledge on Distribution of the Aquatic Beetles (Ordo: Coleoptera) in the Western Black Sea Region and Its Environs of Turkey
Article
  • Sep 2010
The collected aquatic Coleoptera specimens from inland water of the Western Black Sea Region between 2002-2005 are examined, In this study, 47 taxa belonging to 9 family were determined and their distribution in Turkey are given. Twenty three of them are the first record for the study area.
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Research of aquatic Coleoptera fauna of The Inner Western Anatolia, Part - I (Adephaga)
Article
Full-text available
  • Jan 2015
4327 specimens belonging to the adephagan water beetles were collected by area studies between April-September in 2007-2009 in provinces of Afyon, Denizli, Kütahya and Uşak locate in Inner Western Anatolia of Turkey. As a result of identification of these specimens, 77 species group taxa (72 species and 5 subspecies) of 25 genera of 4 families belong to the order Coleoptera were determined (Gyrinidae: 9 species; Haliplidae: 8 species; Noteridae: 1 species; Dytiscidae: 54 species and 5 subspecies).
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Biogeographic zonation of the Eurasian fresh waters based on the macrobenthic faunas
Article
Full-text available
  • Mar 2010
  • ZH OBSHCH BIOL
Spatial differentiation of the Eurasian freshwater faunas is analyzed based on the original and published data on the aquatic insects, crustaceans, and mollusks (about 8800 species in total). The Hacker-Dice similarity index is employed as a principal criterion of differentiation. The schemes of biogeographic zonation are constructed for the nine large macrobenthic taxa, namely, Odonata, Ephemeroptera, Plecoptera, Hemiptera, Coleoptera, Trichoptcra, Malacostraca, Gastropoda, and Bivalvia. Discussed are principal discordances in distribution of three different ecological-systematic groups of the macrobenthos, namely, limnophylic insects, rheophylic insects, and crustaceans with mollusks. A generalized zonation system of the Eurasian fresh waters is elaborated, which is fundamentally divided into Palaearctic and Oriental Regions. The former is further divided into five subregions: Euro-Ob, Near East, Central Asia, Eastern Siberia, and Japan. The latter is divided into three subregions: Indo-Himalaya, China, and Malay. Preliminary classification of the provinces is also provided. Disagreements between the biogeographic systems of different authors are discussed.
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First records of Oreodytes Seidlitz, 1887 (Dytiscidae, Coleoptera) from Turkey: Oreodytes septentrionalis (Gyllenhal, 1826) and Oreodytes davisii (Curtis, 1831)
Article
Full-text available
  • Jan 2002
Oreodytes septentrionalis (Gyllenhal, 1826) and Oreodytes davisi (Curtis, 1831) are recorded in Turkey for the first time and are reviewed according to our specimens and their distributions around the world.
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Another synonym of Agabus faldermanni ZAITZEV, with a discussion of the guttatus-, nebulosus-and paludosus-groups CColeoDtera: Dvtiscidaei (Coleoptera: Dytiscidae)
Article
Full-text available
  • Jan 1992
Agabus iranicus GUEORGUIEV, 1965, is a junior subjective synonym of Agabus faldermanni ZAITZEV, 1927, syn.nov. A lectotype is designated for Gaurodytes palaestinus ZIMMERMANN, 1934. The distribution of A. faldermanni is mapped. Delimitations of the guttat us-, nebulosus-and paludosus-groups are discussed. It is concluded that A. faldermanni either should be placed in a group of its own or placed in the guttatus-group which necessitates a widening of the definition of the group.
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Revision of the nigrita-group of Hydroporus CLAIRVILLE, 1806 (Insecta: Coleoptera: Dytiscidae)
Article
Full-text available
  • Apr 2004
The six species assigned to the nigrita-group of Hydroporus CLAIRVILLE, 1806 are revised. Hydroporus sabaudus FAUVEL is established as a valid name for the species which has been treated as H. nivalis HEER. Hydroporus nivalis is found to be a senior synonym of H. foveolatus HEER but for matters of stability a proposal for precedence of the name Hydroporus foveolatus over the name Hydroporus nivalis has been submitted to the Commission of Zoological Nomenclature. Hydroporus sabaudus sierranevadensis ssp.n. is described from the Sierra Nevada, Southern Spain. Hydroporus tibetanus ZAITZEV, erroneously attributed to the nigrita-group till now, is transferred to the acutangulus-polaris species complex. Descriptions, distributional data, and some bionomical notes are presented. The great variability of the main characters of the species is discussed. A key to all species of the nigritagroup is presented which will allow the separation of most specimens. The body outlines and aedeagi of the studied species are figured to show the variability of body shape, coloration, and shape of median lobe. Lectotypes are designated for the following nominal species: Hydroporus nigrita (F., 1792), H. trivialis STEPHENS, 1828, H. glabellus THOMSON, 1867, H. subalpinus THOMSON, 1871, H. monilicornis J.SAHLBERG, 1875, H. bisbiguttatus J.SAHLBERG, 1921, H. sabaudus FAUVEL, 1865, H. alticola SHARP, 1882, H. nivalis var. scholzii KOLBE, 1899, H. kozlovskii ZAITZEV, 1927, H. nivalis HEER, 1839, H. foveolatus HEER, 1839, and H. tibetanus ZAITZEV, 1953. Key words: Insecta, Coleoptera, Dytiscidae, Hydroporinae, Hydroporus, nigrita-group, new synonymy, new subspecies, key to species, taxonomy, lectotypes, first records, distribution.
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A Contribution to the knowledge of the Turkish Water Beetles Fauna (Coleoptera)
Article
Full-text available
  • Jan 2006
A total of 28 species of water beetles were collected from 11 sampling stations in the semiarid central Turkey (Aksaray, Kırşehir and Konya provinces) between 2004 and 2005. They belong to the following families (species numbers in parentheses): Gyrinidae (4), Haliplidae (2), Noteridae (1), Dytiscidae (9), Helophoridae (4) and Hydrophilidae (8). One species of Dytiscidae [Oreodytes septentrionalis (Gyllenhal, 1826)] and two species of Hydrophilidae [Enochrus (Enochrus) melanocephalus (Olivier, 1792), Enochrus (Lumetus) fuscipennis (Thomson, 1884)] are reported from Turkey for the second time. Also ten new records are added for the Coleoptera fauna of the Central Anatolia region with this study. For some remarkable species phenologies, habitats and additional distributional notes are given.
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Revision of the Genus Eretes Laporte, 1833 (Coleoptera: Dytiscidae)
Article
  • Dec 2002
  • AQUAT INSECT
The genus Eretes Laporte is revised. Four species are recognized in the genus; 1) E. australis (Erichson) (= E. punctipennis (MacLeay)), 2) E. griseus (Fabricius) (= E. plicipennis (Motschulsky), = E. succinctus (Klug), = E. moroderi Báguena Corella), 3) E. sticticus (Linnaeus) (= E. punctatus (Zoubkoff), = E. helvolus (Klug), = E. occidentalis (Erichson), = E. conicollis (Wollaston), = E. subcoriaceus (Wollaston), = E. subdiaphanus (Wollaston)) and 4) E. explicitus Miller, sp. n. The taxonomic history of the group is reviewed. The tribe and genus are diagnosed and a key to the species is provided. Each species is diagnosed, its general distribution is provided and important morphological features are illustrated. The larva of E. australis is illustrated and used to diagnose the tribe. Evidence for the sister-group relationship between Eretini and Aciliini and for relationships within the tribe is presented based on characters from adult and larval morphology.
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Two new Deronectes SHARP, 1882 (Insecta: Coleoptera: Dytiscidae) and notes on other species of the genus
Article
  • Jan 2001
Two new species of Deronectes (Insecta: Coleoptera: Dytiscidae) are described: D. toledoi sp.n. from north- eastern Turkey and D. elburs sp.n. from northern Iran. Modifications to known keys are presented. Several interesting new records are presented, and a few notes on taxonomy, nomenclature, or biology are included. Deronectes latus (STEPHENS, 1829) is recorded from Greece for the first time.
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A revision of the Agabus chalconatus- and erichsoni-groups (Coleoptera: Dytiscidae), with a proposed phylogeny
Article
  • Jan 1993
Fery, H. & Nilsson, A. N.: A revision of the Agabus chalconatus- and erichsoni-groups (Coleoptera: Dytiscidae), with a proposed phylogeny. Ent. scand. 24: 79-108. Copenhagen, Denmark. April 1993. ISSN 0013-8711. The Holarctic chalconatus- and erichsoni-groups, with 19 and three species respectively, are revised. The following seven new species are described: Agabus balkei from Siberia, Agabus jaechi from Turkey, Agabus larsoni from Canada, Agabus lenkoranensis from Azerbaijan, Agabus pederzanii from Italy, Agabus samokovi from Bulgaria, and Agabus wewalkai from Turkey. Lectotypes are designated for the following nominal species: Dytiscus chalconatus Panzer, 1796, Dytiscus nigroaeneus Marsham, 1802, Colymbetes montanus Stephens, 1828, Agabus neglectus Erichson, 1837, Agabus subtilis Erichson, 1837, Agabus gagates Aubé, 1838, Agabus altaicus Gebler, 1848, Agabus erichsoni Gemminger & Harold, 1868, Gaurodytes lutosus Crotch, 1873, Agabus fuscoaenescens Régimbart, 1877, Agabus aenescens Poppius, 1905, Agabus satunini Zaitzev, 1913, and Agabus melanocornis Zimmermann, 1915. The following new synonyms are established: Agabus altaicus Gebler, 1848 = Agabus subtilis Erichson, 1837; Agabus melanocornis Zimmermann, 1915 = Colymbetes montanus Stephens, 1828; Gaurodytes bulgaricus Csiki, 1943 = Dytiscus conspersus Marsham, 1802; Agabus skiathos Hinterseher, 1981 = Agabus pseudoneglectus Franciscolo, 1972; and Agabus maestroae Fresneda & Hernando, 1987 = Agabus albarracinensis Fery, 1986. Agabus aenescens Poppius, 1905, and A. satunini Zaitzev, 1913, are recognized as valid species (sp. rev.). The chalconatus-group is divided into four subgroups and the phylogeny of the group is reconstructed. The relationships of the chalconatus- and erichsoni-groups with other Agabus species-groups and with Ilybius Erichson are discussed.
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