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Checklist of the Albanian mollusc fauna

Authors:

Abstract

Abstract: As the result of 27 Albanian collecting trips, performed within the framework of the long-term Balkan exploration project of the Hungarian Natural History Museum, two new taxa - Gyralina (Gyralina) lunikense n. sp. and Zebrina detrita sallake n. ssp. are described, occurrence of further 38 taxa, namely Radomaniola curta (KÜSTER, 1853), Belgrandia ionica (SCHÜTT, 1980), Bythinella cf. drimica RADOMAN, 1976, Grossuana euxina (A. J. WAGNER, 1828), Paladilhiopsis cf. serbica (PAVLOVIĆ, 1913), Ohridohoratia sturanyi (WESTERLUND, 1902), Pyrgohydrobia grochmalickii (POLIŃSKI 1929), Valvata relicta (POLIŃSKI, 1929), Valvata stenotrema POLIŃSKI, 1929, Gyraulus albidus RADOMAN, 1953, Gyraulus trapezoides POLIŃSKI, 1929, Bithynia radomani GLÖER & PESIĆ, 2007, Carychium tridentatum (RISSO, 1826), Carychium minimum O. F. MÜLLER, 1774, Myosotella myosotis (DRAPARNAUD, 1801), Planorbis presbensis STURANY, 1894, Lauria sempronii (CHARPENTIER, 1837), Vertigo pusilla O. F. MÜLLER, 1774, Vertigo pygmaea (DRAPARNAUD, 1801), Columella edentula (DRAPARNAUD, 1805), Pupilla sterrii (VOITH, 1840), Acanthinula aculeata (O. F. MÜLLER, 1774), Gittenbergia sororcula (BENOIT, 1859), Multidentula squalina (L. PFEIFFER, 1848), Albinaria senilis inconstans (MOUSSON, 1859), Montenegrina dofleini occidentalis NORDSIECK, 1977, Montenegrina perstriata crassa ERŐSS & SZEKERES, 1999, Vitrea pygmaea (O. BOETTGER, 1880), Oxychilus inopinatus (ULICNÝ, 1887), Lehmannia nyctelia (BOURGUIGNAT, 1861), Limax conemenosi O. BÖTTGER, 1882, Limax graecus SIMROTH, 1889, Malacolimax mrazeki (SIMROTH, 1904), Arion fasciatus (NILSSON, 1823), Candidula rhabdotoides (A. J. WAGNER, 1828), Pseudotrichia rubiginosa (ROSSMÄSSLER, 1838), Liburnica albanograeca (SUBAI, 1995) and Pisidium personatum MALM, 1855 are verified. New distribution data are given for Cochlostoma tessellatum tepelenum FEHÉR, ERŐSS & VARGA, 2001, Cochlostoma auritum lamellatum (WESTERLUND, 1885), Radomaniola albanica RADOMAN, 1973, Gyralina circumlineata (L. PFEIFFER, 1846), Gyralina nopcsai RIEDEL, FEHÉR & ERŐSS, 1999, Chondrula lugorensis A. J. WAGNER, 1914, Medora proxima remota POLIŃSKI, 1924, Montenegrina apfelbecki okolensis SZEKERES, 2006, Montenegrina helvola ornata ERŐSS & SZEKERES, 1999, Monacha emmigrata senitschika HAUSDORF, 1996, Metafruticicola occidentalis SUBAI, 1999, and Cattania maranajensis (A. J. WAGNER, 1914).
Schr. Malakozool. 25
22-38 Cismar, Ostholstein, 15.9.2009 ISSN 0936-2959
Checklist of the Albanian mollusc fauna
by Z
OLTÁN FEHÉR and ZOLTÁN PÉTER ERŐSS,
Budapest
22
23
Neritidae
Theodoxus fluviatilis (Linnaeus, 1758) + + +
Theodoxus varius (Menke, 1828) + + +
Cochlostomatidae
Cochlostoma scalarinum (A. & J.B. Villa, 1841)
12
+ +
13
Cochlostoma gracile gracile (L. Pfeiffer, 1849)
14
+
Cochlostoma gracile martensianum (Möllendorff, 1873) + +
+
Cochlostoma kleciaki arnautorum (A.J. Wagner, 1906) + + +
Cochlostoma sturanii mnelense (A.J. Wagner, 1914) + +
Cochlostoma sturanii scalariniformis (A.J. Wagner, 1906) + +
+
15
*Cochlostoma tergestinum (Westerlund, 1878)
16
+ +
Cochlostoma auritum lamellatum (Westerlund, 1885)
[C. a. alatum (A.J. Wagner, 1906)]
17
+ + +
Cochlostoma auritum gjonii Fehér, 2004
Cochlostoma georgi (A.J. Wagner, 1906) + +
18
+
Cochlostoma hoyeri hoyeri (Polinski, 1922) + + +
Cochlostoma hoyeri csikii Fehér, 2004
Cochlostoma hoyeri lillae Fehér, 2004
Cochlostoma fuchsi Fehér, 2004
Cochlostoma pinteri pinteri Fehér, 2004
Cochlostoma pinteri hanswagneri Fehér, 2004
Cochlostoma pinteri erossi Fehér, 2004
Cochlostoma roseoli roseoli (A.J. Wagner, 1901) + +
Cochlostoma roseoli kiriense (A.J. Wagner, 1906) + +
Cochlostoma roseoli prekalense (Polinski, 1924) + +
Cochlostoma roseoli rioliense (A.J. Wagner, 1914) + +
Cochlostoma roseoli scutariense (A.J. Wagner, 1906) + +
+
Cochlostoma tessellatum tessellatum (Rossmässler, 1837)
19
+ +
Cochlostoma tessellatum excisum (Mousson, 1859) + +
+
Cochlostoma tessellatum tepelenum Fehér, Erıss & Varga, 2001
+ +
Aciculidae
Platyla banatica (Rossmässler, 1842) + + +
Platyla corpulenta Subai, 2009
Platyla pinteri (Subai, 1976) + + +
Platyla wilhelmi (A.J. Wagner, 1910) + + +
Renea kobelti albanica Boeters, Gittenberger & Subai, 1989 + + +
Viviparidae
Viviparus contectus (Millet, 1813) + + +
*Viviparus janinensis (Mousson, 1859) +
20
21
Viviparus mamillatus (Küster, 1852) + + +
Viviparus viviparus (Linnaeus, 1758) + + +
12
First Albanian record: Fehér et al. (2001)
13
"Lives probably not in today's Albania"
14
Fehér et al. (2001) reported on Albanian populations that look more like the nominate subspecies.
15
Mentioned as C. sturanyi.
16
Most probably does not live in Albania.
17
For nomenclatural notes see Fehér (2004).
18
Despite Fauna Europaea lists this species also from Yugoslavia, it is known only from Albania.
19
First Albanian record: Fehér et al. (2001).
20
Synonym for V. mamillatus.
21
"occurrence in Albania needs confirmation"
24
Melanopsidae
Amphimelania holandrii (C. Pfeiffer, 1828) + + +
Pomatiidae
Pomatias elegans (O.F. Müller, 1774) + + +
Bithyniidae
*Bithynia tentaculata (Linnaeus, 1758)
22
+ + +
Bithynia radomani Glöer & Pesić, 2007
23
*Bithynia montenegrina (Wohlberedt, 1901)
24
*Bithynia schwabii Frauenfeld, 1865
25
+ +
*Bithynia walderdorffii Frauenfeld, 1865
26
+ +
Hydrobiidae
Adriohydrobia gagatinella (Küster, 1852) + +
27
Belgrandia ionica (Schütt 1980)
28
+
Bythinella cf. drimica Radoman, 1976
29
+
Grossuana euxina (A.J. Wagner, 1828)
30
+
Horatia klecakiana Bourguignat, 1887 +
31
*Hydrobia ventrosa (Montagu, 1803) + +
32
*Hydrobia vitrea (Risso, 1826) + +
Malaprespia albanica Radoman, 1973 + + +
33
Ohridohoratia polinskii (Radoman, 1960) + + +
34
Ohridohoratia pygmaea (Westerlund, 1902) + + +
Ohridohoratia sturanyi (Westerlund, 1902)
35
+
Ohrigocea stankovici (Hadžišce, 1959) + + +
Radomaniola curta (Küster, 1853)
36
+
Radomaniola albanica Radoman, 1973
38
+ +
+
37
Paladilhiopsis cf. serbica (Pavlovic, 1913)
39
+
Parabythinella malaprespensis Radoman, 1973 + + +
22
Probably all records refer to B. radomani.
23
First Albanian record: Fehér & Erıss (2009)
24
Mentioned by Reischütz et al. (2008b), probably refers to B. radomani.
25
Probably does not live in Albania.
26
Probably does not live in Albania.
27
Mentioned as A. consociella with the note "occurrence in Albania needs confirmation". There is a lot in the
collection of the Hungarian Natural History Museum, collected in Himara (South Albania) by the Albanian
entomologist X. Murraj, which is labelled as "Adriohydrobia consociella", but it was revised by M. Bodon and
proved to be Radomaniola albanica.
28
First Albanian record: Fehér & Erıss (2009).
29
First Albanian record: Fehér & Erıss (2009).
30
= G. serbica sensu Radoman. First Albanian record: Fehér & Erıss (2009).
31
"Lives probably not in today's Albania"
32
Dhora & Welter-Schultes (1996) reported this species from three Albanian localities. Two of them are
inland and therefore, these records are surely incorrect, since H. ventrosa lives always in brackish water.
The third locality is mapped in the coast, thus this record is not impossible, but needs confirmation (M.
Bodon personal communication).
33
Dhora & Welter-Schultes (1996) indicate Albaniana albanica as a separate species, according to the
Fauna Europaea, M. albanica and A. albanica are synonyms.
34
Mentioned as Polinskiola polinskii
35
First Albanian record: Reischütz et al. (2008b), see main text for further notes and distribution data.
36
First Albanian record: Fehér & Erıss (2009), see maint text for details.
37
Mentioned by Dhora & Welter-Schultes (1996) as Orientalina, but being a junior homonym, that name must
be replaced (see main text for details). Dhora & Welter-Schultes (1996) treat "Orientalina albanica" as the
subspecies of "O. curta" and they map only the published "O. albanica" records.
38
Possibly a synonym for "Hydrobia haesitans Westerlund, 1881", see the main text for details.
39
First Albanian record: Fehér & Erıss (2009). See the text for details.
25
Prespolitorea malaprespensis Radoman, 1973 + + +
Pseudohoratia ohridana (Polinski, 1929)
40
+
Vinodolia (Anagastina) scutarica (Radoman, 1973) + + +
Zaumia kusceri (Hadžišce, 1959) + + +
Emmericiidae
*Emmericia patula (Brumati, 1838) +
41
Pyrgulidae
Ochridopyrgula macedonica (Brusina, 1896) + + +
Chilopyrgula sturanyi (Brusina, 1896) + + +
Pyrgula annulata dalmatica Schütt, 1968 + + +
42
Pyrgohydrobia grochmalickii (Polinski 1929)
43
+
Valvatidae
Valvata cristata O.F. Müller, 1774 + + +
Valvata piscinalis (O.F. Müller, 1774) + + +
Valvata rhabdota Sturany, 1894 +
44
Valvata relicta (Polinski, 1929)
45
+
Valvata stenotrema Polinski, 1929
46
+
Carychiidae
47
Carychium tridentatum (Risso, 1826)
48
+
Carychium minimum O.F. Müller, 1774
49
+
Ellobiidae
Myosotella myosotis (Draparnaud, 1801) +
50
Acroloxidae
Acroloxus lacustris (Linnaeus, 1758) + + +
Lymnaeidae
Lymnaea stagnalis (Linnaeus, 1758) + + +
Radix auricularia (Linnaeus, 1758) + + +
Radix ampla (W. Hartmann, 1821) + + +
Radix balthica (Linnaeus, 1758) + +
Radix labiata (Rossmässler, 1835) + + +
51
Radix relicta Polinski, 1929 + +
Stagnicola corvus (Gmelin, 1791) + + +
Stagnicola palustris (O.F. Müller, 1774) + + +
Galba truncatula (O.F. Müller, 1774) + + +
Physidae
Aplexa hypnorum (Linnaeus, 1758) + + +
Physella acuta (Draparnaud, 1805) + + +
40
First Albanian record: Fehér & Drimmer (2004).
41
"occurrence in Albania needs confirmation"
42
Mentioned as P. annulata.
43
First Albanian record: Fehér & Erıss (2009).
44
"occurrence in Albania needs confirmation"
45
First Albanian record: Fehér & Erıss (2009).
46
First Albanian record: Fehér & Erıss (2009).
47
Carychium spp. have not been known from Albania to now, first records are presented in this study. Our
records are from the northern part of Albania. Since C. hellenicum Bank & Gittenberger, 1985 is known from
Epirus and Korfu Island, this species probably lives in South Albania.
48
First Albanian record: Fehér & Erıss (2009).
49
Reischütz et al. (2008b) have found subrecent shells. First verified Albanian record: Fehér & Erıss (2009).
50
"occurrence in Albania needs confirmation". See the main text for a recent distribution record.
51
Mentioned as R. peregra.
26
Physa fontinalis (Linnaeus, 1758) + + +
Planorbidae
Ancylus fluviatilis O.F. Müller, 1774 + + +
Ancylus tapirulus Polinski, 1929
52
+
Gyraulus acronicus (A. Férussac, 1807) + +
Gyraulus albidus Radoman, 1953
53
+
Gyraulus albus (O.F. Müller, 1774) + + +
Gyraulus laevis (Alder, 1838) + + +
Gyraulus crista (Linnaeus, 1758) + + +
Gyraulus crenophilus Hubendick & Radoman, 1959 + + +
Gyraulus fontinalis Hubendick & Radoman, 1959 + + +
Gyraulus trapezoides Polinski, 1929
54
+
Bathyomphalus contortus (Linnaeus, 1758) + + +
*Anisus vortex (Linnaeus, 1758) + + +
55
*Anisus vorticulus (Troschel, 1834) + + +
56
Anisus spirorbis (Linnaeus, 1758)
57
+
Planorbis carinatus O.F. Müller, 1774 + + +
Planorbis planorbis (Linnaeus, 1758) + + +
Planorbis presbensis Sturany, 1894
58
+
Ferrissia clessiniana (Jickeli, 1882) + + +
59
Planorbarius corneus corneus (Linnaeus, 1758) + +
Planorbarius corneus cf. grandis (Dunker, 1850)
60
+
Planorbarius corneus arabatzis Reischütz, Reischütz & Fischer,
2008
61
+
*Planorbella anceps (Menke, 1830)
62
Succineidae
52
Fauna Europaea list this species only from Macedonia. First Albanian record: Reischütz et al. (2008b).
Albrecht et al. (2006) studied recently the molecular phylogeny of the Ancylus species of the Lake Ohrid,
mostly based on Macedonian material. They have found that A. fluviatilis does not occur in the lake, just A.
tapirulus, A. scalariniformis, A lapicidus and an undescribed fourth Ancylus species.
53
First Albanian record: Fehér & Erıss (2009).
54
First Albanian record: Fehér & Erıss (2009).
55
Listed by Dhora & Welter-Schultes (1996) without any distribution map and specific distribution data.
According to their reference list, Dhora & Welter-Schultes (1996) included this species in the Albanian fauna
list on the basis of a roughly drawn distribution map of Meier-Brook (1983: p. 92).
56
Listed by Dhora & Welter-Schultes (1996) without any distribution map and specific distribution data.
According to their reference list, Dhora & Welter-Schultes (1996) included this species in the Albanian fauna
list on the basis of a roughly drawn distribution map of Meier-Brook (1983: p. 92).
57
First Albanian record: Reischütz et al. (2008b).
58
First Albanian records: Fehér & Drimmer (2004) and Fehér & Erıss (2009).
59
Mentioned as F. wautieri
60
According to the Fauna Europaea, P. c. grandis occurs only in Greece. In the Prespa Lakes, P. corneus
populations are significantly different from the nominate form and resemble to the figure in the original
description of Planorbis grandis. Reischütz et al. (2008a) believes that two distinct subspecies occur in the
Prespa region: P. corneus grandis in the Small Prespa Lake and P. corneus arabatzis in the Great Prepa
Lake. The distinction of their new subspecies was based on slight shell morhological differences. As the two
lakes are divided only by a narrow isthmus and they are connected by a creek, it can be supposed
reasonably that P. corneus populations of the two lakes are in connection. In view of the above, it is
questionnable and needs further confirmation whether or not the the Great Prespa populations belong to a
separate taxon.
61
See the above note.
62
The abnormal P. corneus form in the Great Prespa Lake, was misidentified and incorrectly reported as P.
anceps by Erıss et al. (2005). See the main text for details.
27
Oxyloma (Oxyloma) elegans (Risso, 1826) + + +
63
*Succinella oblonga (Draparnaud, 1801) + + + (!)
64
Cochlicopidae
Cochlicopa lubrica (O.F. Müller, 1774) + + +
Cochlicopa lubricella (Porro, 1838) + + +
Azecidae
Hypnophila pupaeformis (Cantraine, 1835) + + +
Hypnophila polita (Porro, 1838) +
65
Lauriidae
Lauria cylindracea (Da Costa, 1778) + + +
Lauria sempronii (Charpentier, 1837)
66
+
Orculidae
Pagodulina kaeufeli Klemm, 1939 + + +
67
Pagodulina subdola gracilior Pilsbry, 1926 + + +
Sphyradium doliolum (Bruguière, 1792) + + +
Orcula schmidtii schmidtii (Küster, 1843) + + +
Orcula wagneri wagneri Sturany, 1914 + + +
Argnidae
Agardhiella truncatella (L. Pfeiffer, 1841) +
Agardhiella skipetarica
69
(A.J. Wagner, 1914) [A. truncatella
skipetarica]
+
+
68
*Agardhiella lamellata (Clessin, 1887)
70
+ +
Agardhiella zoltanorum Subai, 2008
Agardhiella extravaganta Subai, 2008
Strobilopsidae
Spelaeodiscus albanicus (A.J. Wagner, 1914) + + +
Valloniidae
Vallonia costata (O.F. Müller, 1774) + + +
Vallonia enniensis (Gredler, 1856) + + +
Vallonia excentrica Sterki, 1893 + + +
Vallonia pulchella (O.F. Müller, 1774) + + +
Acanthinula aculeata (O. F. Müller, 1774)
71
+
72
Gittenbergia sororcula (Benoit, 1859)
73
+
74
*Plagyrona placida (Shuttleworth, 1852)
75
+
Pupillidae
63
Dhora & Welter-Schultes (1996) list Succinea pfeifferi Rossmassler, 1835 separately, which is however a
synonym for this species.
64
Due to a misinterpretation of Bába (1982), this species was falsely included in the Albanian list by Dhora &
Welter-Schultes (1996).
65
"lives very probably in Albania"
66
First Albanian record: Fehér & Erıss (2009).
67
Mentioned as P. subdola kaeufeli.
68
Dhora & Welter-Schultes (1996) indicates this two species as "syn/ssp.". In their map (p. 120) only one
locality (the type locality of A. skipetarica) is indicated.
69
According to Subai (1999a and 2008) this is a distict species.
70
Does not live in Albania. Found in Romania and in Hungary (fluvial flotsam) up to now.
71
First Albanian record: Fehér & Erıss (2009).
72
"occurrence in Albania needs confirmation"
73
First Albanian record: Fehér & Erıss (2009).
74
"occurrence in Albania needs confirmation"
75
Mentioned by Reischütz et al. (2008b).
28
*Pupilla alpicola (Charpentier, 1837)
76
+ + +
*Pupilla triplicata (S. Studer, 1820) + + + (!)
77
Pupilla sterrii (Voith, 1840)
78
+
Pyramidulidae
Pyramidula cephalonica (Westerlund, 1898) + + +
Pyramidula pusilla (Vallot, 1801) + + +
Chondrinidae
Granopupa granum (Draparnaud, 1801) + + +
*Granaria frumentum illyrica (Rossmässler, 1835) [G. illyrica] +
79
+
Granaria frumentum hungarica (M. von Kimakowicz, 1890) +
80
Granaria frumentum ssp. n.
81
+
Chondrina arcadica abundans (Westerlund, 1894) + +
Chondrina arcadica clienta (Westerlund, 1883) + +
+
Chondrina spelta spelta (Beck, 1837)
82
+
Chondrina spelta fuchsi H. Nordsieck, 1970 + +
Chondrina spelta serbica H. Nordsieck, 1970 + +
+
Rupestrella philippii (Cantraine, 1841) + + +
Rupestrella rhodia (Roth, 1839) + + +
Vertiginidae
Vertigo pusilla O.F. Müller, 1774
83
+ + + (!)
84
Vertigo pygmaea (Draparnaud, 1801)
85
+
*Vertigo antivertigo (Draparnaud, 1801)
86
+
Columella edentula (Draparnaud, 1805)
87
+
Truncatellina callicratis (Scacchi, 1833) + + +
Truncatellina claustralis (Gredler, 1856) + + +
Truncatellina cylindrica (A. Férussac, 1807) + + +
Truncatellina rothi (Reinhardt, 1916) + + +
Enidae
Ena concolor (Westerlund, 1887) + + +
Ena montana (Draparnaud, 1801) + + +
Ena subtilis subtilis (Rossmässler, 1837) + +
Ena subtilis corcyrensis (Mousson, 1859) + +
+
Jaminia quadridens (O.F. Müller, 1774) + + +
76
Soós (1924) reports on the occurrence of this species in the Korab Mts., that record, however, possibly
based on misidentified P. sterrii material.
77
Due to a misinterpretation of Bába (1982), this species was included in the Albanian list by Dhora &
Welter-Schultes (1996), but there are no confirmed Albanian record up to now.
78
First Albanian record: Fehér & Erıss (2009).
79
Fauna Europaea treats this taxon as a distinct species.
80
Fauna Europaea treats this taxon as a synonym for the nominate subspecies.
81
According to a recent revision of the Balkan species of Granaria, "G. illyrica" is the subspecies of G.
frumentum, G. f. hungarica is a valid subspecies and only two G. frumentum subspecies occur in Albania: G.
f. hungarica and a new form (Fehér et al. submitted).
82
Although Fauna Europaea does not indicate it, the nominate subspecies lives in Albania too (see
Nordsieck 1970)
83
First verified Albanian records are given in the Fehér & Erıss (2009).
84
This species was included in the Albanian checklist of Dhora & Welter-Schultes (1996) due to a
misinterpretation of Bába (1982).
85
First Albanian record: Reischütz et al. (2008b), further distribution data are given in the Fehér & Erıss
(2009).
86
A subrecent occurrence was reported by Reischütz et al. (2008b), recent occurrence needs confirmation.
87
First Albanian record: Fehér & Erıss (2009).
29
Pseudochondrula seductilis (Rossmässler, 1837) + + +
Multidentula cf. squalina (L. Pfeiffer, 1848)
88
Merdigera obscura (O.F. Müller, 1774) + + +
Mastus grandis grandis (Mousson, 1859) + + +
89
Chondrula consentanea (Westerlund, 1887) +
90
Chondrula lugorensis A.J. Wagner, 1914 + + +
Chondrula macedonica A.J. Wagner, 1914 + + +
Chondrula microtragus (Rossmässler, 1839) + + +
Chondrula quinquedentata (Rossmässler, 1837) + +
Chondrula tridens (O.F. Müller, 1774) + +
+
91
Napaeopsis cefalonicus (Mousson, 1859) +
Napaeopsis merditanus merditanus (Sturany, 1907) + +
Napaeopsis merditanus latifianus (Sturany, 1907) + +
Napaeopsis merditanus winneguthi (Sturany, 1907) + +
Napaeopsis merditanus zebianus (Sturany, 1907) + +
+
92
Zebrina detrita (O.F. Müller, 1774) + + +
Zebrina detrita sallake ssp. n.
93
Clausiliidae
Sciocochlea llogaraensis (Reischütz & Reischütz, 2009)
Bulgarica vetusta gracilior S.H.F. Jaeckel, 1954 + +
Bulgarica vetusta nannodes (A.J. Wagner, 1912) + +
+
Vestia roschitzi apragmosyne (A.J. Wagner, 1919) + + +
Alinda biplicata biplicata (Montagu, 1803)
94
[Balea biplicata] +
Alinda biplicata byshekensis Nordsieck, 2008
Alinda biplicata distincta (Sturany, 1894)
95
+ +
*Alinda biplicata labiata (Westerlund, 1885)
96
+ +
*Alinda biplicata metriotes (A.J. Wagner, 1919)
97
+ +
+
Balea serbica serbica (Möllendorff, 1873) + + +
98
Laciniaria plicata (Draparnaud, 1801) + + +
Clausilia dubia vindobonensis A. Schmidt, 1856 + +
99
Macrogastra ventricosa (Draparnaud, 1801)
100
Strigilodelima conspersa (L. Pfeiffer, 1848)
101
+ +
Strigilodelima pentheri (A.J. Wagner, 1919) + + +
* Albinaria maculosa (Deshayes, 1835)
[Albinaria s. schuchii (Rossmässler, 1836)]
102
+ + +
88
First Albanian record: Fehér & Erıss (2009).
89
Mentioned as M. pupa.
90
"occurrence in Albania needs confirmation"
91
Dhora & Welter-Schultes (1996) treat C. quinquedentata as a "syn/ssp." of C. tridens.
92
Dhora & Welter-Schultes (1996) treat merditanus, latifianus, winneguthi and zebianus as "syn/ssp." of the
N. cefalonicus. Fauna Europaea indicates N. cefalonicus only from Greece.
93
Described in the Fehér & Erıss (2009).
94
Systematics and distribution of this species has been revised by Nordsieck (2008).
95
Though Dhora & Welter-Schultes (1996) do not go into the particulars at the subspecies level generally,
but for this subspecies they note that "B. b. distincta does not live in Albania". But see Nordsieck (2008).
96
According to Nordsieck (2008), this is a synonym of the nominate subspecies ("oreinos form").
97
According to Nordsieck (2008), this is a synonym of the nominate subspecies ("oreinos form").
98
Dhora & Welter-Schultes (1996) treat this taxon as a "syn/ssp." of B. fallax.
99
"needs confirmation"
100
First Albanian record: Dhora & Welter-Schultes (1999a).
101
Fauna Europaea lists this species only for Macedonia. It is very common in Albania.
102
Nordsieck (1974) indicates A. schuchii from Albania (collected near Sarande by Edlauer). Despite our
efforts, we could not confirm this occurrence to now. Nordsieck (1995) proposed to use the name schuchii
30
Albinaria scopulosa epirotes H. Nordsieck, 1974 + +
Albinaria scopulosa winneguthi (A.J. Wagner, 1914) + +
+
Albinaria senilis inconstans (Mousson, 1859)
103
+
Agathylla biloba biloba (A.J. Wagner, 1914) + +
Agathylla biloba dabovici Brandt, 1961 + +
+
Agathylla biloba martae Eröss & Szekeres, 1999 + +
Agathylla neutra neutra (Westerlund, 1898) + + +
104
Agathylla neutra merditana (A.J. Wagner, 1914) + + +
Medora proxima remota (Polinski, 1924) + + +
Triloba sandrii (Küster, 1844) + + +
Triloba thaumasia thaumasia (Sturany, 1907) + +
Triloba thaumasia dardanorum (A.J. Wagner, 1919) + +
Triloba thaumasia korabensis (A.J. Wagner, 1919) + +
+
*Herilla bosniensis (L. Pfeiffer, 1868) + +
105
Herilla illyrica diabasis (A.J. Wagner, 1919) + +
Herilla illyrica oribates (Sturany, 1909) +
+
Herilla jabucica interrupta Nordsieck, 1971
106
+
Herilla ziegleri klemmi Nordsieck, 1971
107
+ +
Protoherilla baleiformis (O. Boettger, 1909) + + +
Protoherilla pseudofallax (Nordsieck, 1972) + +
Montenegrina apfelbecki apfelbecki (Sturany, 1907) + + +
Montenegrina apfelbecki caesia Fehér & Szekeres, 2006
108
Montenegrina apfelbecki csikii Erıss & Szekeres, 2006
109
Montenegrina apfelbecki okolensis Szekeres, 2006
110
Montenegrina apfelbecki wagneri Szekeres, 2006
111
Montenegrina dofleini occidentalis Nordsieck, 1977
112
+
Montenegrina dofleini plenostoma Fehér & Szekeres, 2006
Montenegrina drimmeri Fehér & Szekeres, 2006
Montenegrina ersekensis ersekensis H. Nordsieck, 1996
113
+ + +
Montenegrina ersekensis nobilis Erıss & Szekeres, 2006
114
instead of maculosa because several authors had misunderstood the name maculosa before, and he
considered it as a nomen oblitum, but this view is not tenable (the name was not forgotten, only
misinterpreted). Since Nordsieck did not put his proposal into a frame of nomenclatural rules, we have to use
the name maculosa which has priority (Welter-Schultes personal communication).
103
First Albanian record: Fehér & Erıss (2009).
104
Mentioned by Dhora & Welter-Schultes (1996) as A. merditana scraparana Nordsieck 1996. See
Nordsieck (2001) for details.
105
Dhora & Welter-Schultes (1996) lists this species without a distribution map. Though there are 3
subspecies indicated for Albania in the table of Jaeckel et al. (1957), but this is probably a misprint. To our
present knowledge, H. bosniensis does not occurr in Albania.
106
First Albanian record: Dhora & Welter-Schultes (1999a).
107
First Albanian record: Nordsieck (1971).
108
Nordsieck (2007) mentions as the subspecies of M. janinensis.
109
Nordsieck (2007) mentions as the subspecies of M. janinensis.
110
Nordsieck (2007) mentions as the subspecies of M. janinensis, later, proposed as the synonym of "M. j.
caesia" (Nordsieck 2009).
111
Probably, in the origibal description (Erıss et al. 2006) the type locality (Mali Thate) was incorrectly
identified with Maja e Madhe in North Albania. The type locality is more probably identical with Mali i Thatës
in Southeast Albania. In Nordsieck's (2009) opinion, this taxon is the synonym of M. janinensis fagorum.
112
First Albanian record: Fehér & Erıss (2009).
113
Nordsieck (2007) mentions as the subspecies of M. irmengardis, and later (Nordsieck 2009) as the
subspecies of M. skipetarica. See note at M. perstriata skipetarica.
114
Its subspecies rank was questionned by Nordsieck (2009).
31
Montenegrina fuchsi fuchsi R.A. Brandt, 1961 + + +
Montenegrina fuchsi klemmi R.A. Brandt, 1962 + + +
Montenegrina fuchsi muranyii Feher & Szekeres, 2006
Montenegrina helvola helvola (Küster, 1853) + +
Montenegrina helvola pageti R.A. Brandt, 1962 + +
+
Montenegrina helvola carinata Erıss & Szekeres, 1999 + +
Montenegrina helvola magna Fehér & Szekeres, 2006
Montenegrina helvola ornata Eröss & Szekeres, 1999 + +
Montenegrina irmengardis flava Erıss & Szekeres, 2006
Montenegrina irmengardis remota Feher & Szekeres, 2006
Montenegrina irmengardis rugosa Feher & Szekeres, 2006
Montenegrina irmengardis timeae Erıss & Szekeres, 2006
Montenegrina irmengardis welterschultesi Feher & Szekeres,
1999
+ + +
115
Montenegrina janinensis coerulescens H. Nordsieck, 1996 + +
Montenegrina janinensis fagorum H. Nordsieck, 1974 + +
Montenegrina janinensis tomorosi R.A. Brandt, 1961 + +
+
Montenegrina janinensis ampla Feher & Szekeres, 2006
Montenegrina janinensis soosi Erıss & Szekeres, 2006
Montenegrina laxa laxa (Küster, 1860) + +
Montenegrina laxa chiasma H. Nordsieck, 1972 + +
Montenegrina laxa iba H. Nordsieck, 1972 + +
Montenegrina laxa miraka H. Nordsieck, 1996 + +
+
Montenegrina laxa disjuncta Fehér & Szekeres, 2006
Montenegrina laxa errans Erıss & Szekeres, 2006
Montenegrina laxa kontschani Erıss & Szekeres, 2006
Montenegrina minuscula Erıss & Szekeres, 2006
Montenegrina perstriata callistoma Fehér & Szekeres, 2006
Montenegrina perstriata crassa Erıss & Szekeres, 1999
116
+
Montenegrina perstriata gracilis Erıss & Szekeres, 2006
Montenegrina perstriata nana Fehér & Szekeres, 2006
Montenegrina perstriata skipetarica (Soós, 1924)
117
+ + +
Montenegrina perstriata tenebrosa Nordsieck, 2009
Montenegrina sattmanni sattmanni Nordsieck, 1988
118
+
Montenegrina sattmanni costulata Erıss & Szekeres, 2006
Montenegrina sattmanni fusca Fehér & Szekeres, 2006
119
Montenegrina subcristata wohlberedti (Möllendorff, 1899) + + +
*Cochlodina commutata (Rossmässler, 1836)
120
+ +
Cochlodina laminata albanica S.H.F. Jaeckel, 1956 + + +
Charpentieria lamellata (Rossmässler, 1836) + + +
121
Charpentieria pantocratoris margaritifera (H. Nordsieck, 1996) + + +
115
The record of M. irmengardis by Dhora & Welter-Schultes (1996) was based on this subspecies.
116
First Albanian record: Fehér & Erıss (2009). Nordsieck (2009) questionned its subspecies rank.
117
In Nordsieck's (2009) opinion, this subspecies belongs to M. irmengardis rather than to M. perstriata. As
skipetarica is an older name than irmengardis, he suggest to use this specific name for each M. irmengardis
subspecies.
118
First Albanian record: Erıss et al. (2006).
119
In Nordsieck's (2009) opinion it belongs to "M. skipetarica" rather than to M. sattmanni. See note at M.
perstriata skipetarica.
120
Most probably does not live in Albania.
121
Mentioned as Siciliaria lamellata.
32
Charpentieria splendens (H. Nordsieck, 1996) + + +
122
Charpentieria stigmatica stigmatica (Rossmässler, 1836) + + +
Charpentieria stigmatica sturmii (L. Pfeiffer, 1848) + + +
Papillifera bidens (Linnaeus, 1758) + +
123
Delima bilabiata (J.A. Wagner, 1829) + +
Delima montenegrina (L. Pfeiffer, 1848) + +
*Delima binotata (Rossmässler, 1836) + +
124
*Delima blanda (Rossmässler, 1836) + +
Ferussaciidae
Cecilioides acicula (O.F. Müller, 1774) + + +
Cecilioides tumulorum (Bourguignat, 1856) + + +
Subulinidae
*Rumina decollata (Linnaeus, 1758) + +
125
Oleacinidae
Poiretia compressa (Mousson, 1859) + + +
Poiretia cornea (Brumati, 1838) + + +
Poiretia delesserti (Bourguignat, 1852) + + +
Helicodiscidae
Lucilla singleyana (Pilsbry, 1889)
126
+
Punctidae
Punctum pygmaeum (Draparnaud, 1801) + + +
Euconulidae
Euconulus fulvus (O.F. Müller, 1774) + + +
Gastrodontidae
Zonitoides nitidus (O.F. Müller, 1774) + + +
Zonitidae [incl. Pristilomatidae, Oxychilidae, Zonitidae]
Vitrea botterii (L. Pfeiffer, 1853) + + +
Vitrea contracta (Westerlund, 1871) + + +
Vitrea illyrica (A.J. Wagner, 1907) + + +
Vitrea pygmaea (O. Boettger, 1880)
127
+
Vitrea selecta L. Pintér, 1972 + + +
Vitrea sturanyi (A.J. Wagner, 1907) + +
128
Vitrea subrimata (Reinhardt, 1871) + + +
Gyralina circumlineata (L. Pfeiffer, 1846) + + +
Gyralina gjirokastrana Riedel & Welter-Schultes, 1996 + + +
Gyralina candida candida (A.J. Wagner, 1909) + + +
Gyralina nopcsai Riedel, Feher & Eröss, 1999 + +
Gyralina lunikense sp. n.
129
Gyralina tarabosensis (Riedel, 1970) + + +
Gyralina korabensis (Riedel, 1970) +
130
Aegopinella minor (Stabile, 1864) + + +
Oxychilus camelinus (Bourguignat, 1852) + + +
122
Mentioned as S. pantocratoris splendens.
123
Mentioned as P. papillaris.
124
"does not live in today's Albania"
125
"occurrence in Albania needs confirmation"
126
First Albanian record: Reischütz et al. (2008b)
127
First Albanian record: Fehér & Erıss (2009).
128
"occurrence in Albania needs confirmation"
129
Described in the Fehér & Erıss (2009).
130
"Lives probably in today's Albania"
33
[Schistophallus camelinus]
Oxychilus cyprius (L. Pfeiffer, 1847) [Schistophallus cyprius] + + +
Oxychilus depressus (Sterki, 1880) [Mediterranea depressa] + + +
Oxychilus hydatinus (Rossmässler, 1838)
[Mediterranea h. hydatina]
+ + +
Oxychilus juliae Riedel, 1990 [Mediterranea juliae] + +
Oxychilus planorbis (Möllendorff, 1899) [Mediterranea planorbis]
+ +
+
131
Oxychilus inopinatus (Ulicný, 1887)
132
[Mediterranea inopinata] +
[Morlina glabra nitidissima
(Mousson, 1859)]
+ +
Oxychilus glaber
(Rossmässler, 1835)
[Morlina glabra striaria
(Westerlund, 1881)]
+ +
+
Oxychilus draparnaudi (H. Beck, 1837) + + +
Carpathica stussineri (A.J. Wagner, 1895) + + +
Daudebardia brevipes (Draparnaud, 1805) + + +
Daudebardia rufa (Draparnaud, 1805) + + +
Paraegopis albanicus (Rossmässler, 1836) + + +
Paraegopis bizonus A.J. Wagner, 1914 + + +
*Paraegopis mauritii (Westerlund, 1886) +
133
Paraegopis skipetaricus A.J. Wagner, 1914 + + +
134
*Aegopis laughofferi A.J. Wagner, 1914
135
[A. septentrionalis laughofferi]
136
+ +
137
*Aegopis acies (Férussac, 1832)
138
+
139
*Aegopis verticillus (Lamarck, 1822)
140
+
141
Allaegopis amphikypellon Riedel, 1982 + + +
Allaegopis skanderbegianus (Polinski, 1924) + + +
*Allaegopis transiens (Mousson, 1859)
142
+
Vitrinidae
Oligolimax annularis (S. Studer, 1820) + + +
Oligolimax apatelus (Soós, 1924) + + +
143
Vitrina pellucida (O.F. Müller, 1774) + + +
Semilimacella bonellii reitteri (O. Boettger, 1880) + + +
*Semilimacella carniolica (O. Boettger, 1884) + +
144
Milacidae
Tandonia albanica (Soós, 1924) + + +
Tandonia sowerbyi (A. Férussac, 1823) + + +
Limacidae
131
Listed together as "complex"
132
First Albanian record: Fehér & Erıss (2009).
133
"occurrence in Albania needs confirmation"
134
"taxonomic revision is needed"
135
Most probably does not live in Albania.
136
According to Fauna Europaea, the nominate subspecies of this taxon is A. septentrionalis (Kobelt, 1899).
Kobelt's name, however, can not be used, because that is a junior homonym of Zonites septentrionalis
Bourguignat, 1870. Thus probably laughofferi is the oldest available name for the species.
137
"taxonomy and distribution not clear"
138
Most probably does not live in Albania.
139
"occurrence in Albania needs confirmation"
140
Most probably does not live in Albania.
141
"occurrence in Albania needs confirmation"
142
"occurrence in Albania needs confirmation"
143
"systematic position not clear"
144
"systematic position not clear"
34
Lehmannia marginata (O.F. Müller, 1774) + + +
Lehmannia szigethyae Wiktor, 1975 + + +
Lehmannia nyctelia (Bourguignat, 1861)
145
+
Limacus flavus (Linnaeus, 1758) + + +
Limax cephalonicus Simroth, 1886 + + +
Limax cinereoniger Wolf, 1803 + + +
Limax conemenosi O. Böttger, 1882
146
+
Limax graecus Simroth, 1889
147
+
Limax maximus Linnaeus, 1758 + + +
Limax wohlberedti Simroth, 1900 + + +
Malacolimax mrazeki (Simroth, 1904)
148
+
Agriolimacidae
Deroceras agreste (Linnaeus, 1758) + + +
Deroceras reticulatum (O.F. Müller, 1774) + + +
*Deroceras sturanyi (Simroth, 1894) + +
149
Deroceras turcicum (Simroth, 1894) + + +
*Gigantomilax csikii Soós, 1924 +
150
Arionidae
Arion subfuscus (Draparnaud, 1850)
[A. fuscus (O.F. Müller, 1774)]
+ + +
Arion circumscriptus Johnston, 1828 + + +
Arion fasciatus (Nilsson, 1823)
151
+
*Arion rufus (Linnaeus, 1758) +
152
Helicodontidae
Lindholmiola corcyrensis (Rossmässler, 1838) + + +
153
Lindholmiola lens (Férussac, 1832) + +
*Helicodonta laughofferi A.J. Wagner, 1912 + +
154
Helicodonta obvoluta albanica A.J. Wagner, 1914 + + +
Cochlicellidae
Cochlicella acuta (O.F. Müller, 1774) + + +
Cochlicella barbara (Linnaeus, 1758) + + +
Hygromiidae
Xeropicta derbentina (Krynicki, 1836) + + +
Semifruticicola serbica A.J. Wagner, 1914
[Semifruticicola costulata serbica A.J. Wagner, 1914]
155
+ +
156
145
First Albanian record: Fehér & Erıss (2009).
146
First Albanian record: Fehér & Erıss (2009).
147
First Albanian record: Fehér & Erıss (2009).
148
First Albanian record: Fehér & Erıss (2009).
149
"occurrence in Albania needs confirmation"
150
"probably belongs to Deroceras, but doubtful"
151
First Albanian record: Fehér & Erıss (2009).
152
"Lives probably not in today's Albania"
153
Dhora & Welter-Schultes (1996) refer to this species as Lindholmiola girva (Frivaldszky, 1835), because
they, as well as e.g. Irikov & Mollov (2006) treat these names as synonyms. In Subai's opinion – which is
suppported by molecular phylogenetic data (see Fehér & Erıss 2009) – L. girva and L. corcyrensis are
morphologically distinguishable, distict species; and only the latter one occurs in Albania.
154
"Lives probably not in today's Albania"
155
According to Fauna Europaea, the nominate subspecies of this taxon is Helix costulata Brancsik, 1897,
however that is a homonym of Helix costulata Pfeiffer, 1828. Therefore serbica seems to be the oldest
available name for the species.
156
"systematic position not clear"
35
Trochulus erjaveci (Brusina, 1870)
157
[Trichia erjaveci] + + +
Candidula castriota Soós, 1924 + + +
158
Candidula rhabdotoides (A.J. Wagner, 1928)
159
+
Xerotricha conspurcata (Draparnaud, 1801) + + +
Cernuella cisalpina (Rossmässler, 1837)
160
+ +
Cernuella virgata (Da Costa, 1778)
161
+ + +
*Hiltrudia mathildae (Westerlund, 1881)
162
+ + +
Hiltrudia kusmici (Clessin, 1887)
163
+
Metafruticicola occidentalis Subai, 1999 + +
Monachoides fallax (A.J. Wagner, 1914) + +
Monachoides incarnatus (O.F. Müller, 1774) + + +
Pseudotrichia rubiginosa (Rossmässler, 1838)
164
+ + +
Xeromunda vulgarissima (Mousson, 1859) + + +
Xerolenta obvia (Menke, 1828) + + +
Xerocrassa muehlfeldtiana (Rossmässler, 1837) + +
165
Trochoidea pyramidata (Draparnaud, 1805) + + +
Trochoidea trochoides (Poiret, 1789) + + +
Monacha cartusiana (O.F. Müller, 1774) + + +
Monacha subobstructa (Bourguignat, 1855)
[Monacha claustralis (Menke, 1828)]
166
+ + +
167
Monacha cf. dofleini (P. Hesse, 1928)
168
+
Monacha emigrata sattmanni Hausdorf, 1996 + + +
Monacha emigrata senitshika Hausdorf, 1996 + + +
Monacha frequens (Mousson, 1859) + + +
Monacha microtricha S.H.F. Jaeckel, 1954 + +
169
Monacha parumcincta (Menke, 1828) + +
170
Helicidae
Theba pisana pisana (O.F. Müller, 1774) + + +
Helix cincta O.F. Müller, 1774 + +
171
Helix dormitoris Kobelt, 1898 + + +
157
Dhora & Welter-Schultes (1996) indicate the subspecies T. e. floerickei (Kobelt, 1898) for Albania, which
is treated as a synonym by Fauna Europaea.
158
"systematic position is not clear"
159
First Albanian record: Fehér & Erıss (2009).
160
According to Fauna Europaea, this is the senior synonym of C. jonica (Mousson, 1854), and therefore
listed for Albania. See also the note for C. virgata.
161
Dhora & Welter-Schultes (1996) treat as the senior synonym of C. jonica (Mousson, 1854).
162
This species does not live in Albania. See notes for H. kusmici.
163
See notes and some Albanian records in the Fehér & Erıss (2009).
164
First verified Albanian record: Fehér & Erıss (2009)
165
"indeterminated sp."
166
According to F. Welter-Schultes (personal communication) the name claustralis can not be used for this
species. Although Hausdorf (2000: 82) argued that Mousson (1854, describing a species from Greece)
"adopted" the name claustralis from Menke (1828, published in synonymy) and used it for a valid species
(Code Art. 11.6), this was incorrect. Mousson (1859: 254) described a new species and had no idea that
Menke (1828: 12) had mentioned such a name as a synonym in a list of names. This species should than
called Monacha subobstructa (Bourguignat, 1855), because this is the next available name according to
Hausdorf (2000).
167
Mentioned as M. dissimulans Pintér, 1968.
168
First mentioned for Albania by Reischütz et al. (2008b)
169
"systematic position not clear"
170
"occurrence in Albania needs confirmation"
171
"occurrence in Albania needs confirmation"
36
Helix lucorum Linnaeus, 1758 + + +
Helix philibinensis Rossmassler, 1839 +
*Helix oestreichi Kobelt, 1902 + +
172
Helix secernenda Rossmässler, 1847 + + +
Helix vladika (Kobelt, 1898) [H. vladica] + + +
Cornu aspersum (O.F. Müller, 1774) + + +
Eobania vermiculata (O.F. Müller, 1774) + + +
Cepaea vindobonensis (C. Pfeiffer, 1828) + + +
Codringtonia neocrassa Zilch, 1952
173
+ +
174
[Chilostoma apfelbecki bindzaensis (Sturany, 1909)]
175
+ + +
Cattania inflata (Kobelt, 1876) [Chilostoma inflata] + + +
Cattania maranajensis (A.J. Wagner, 1914) [Chilostoma
maranajensis]
+ + +
*Cattania jakupicae (Urbanski, 1979) [Chilostoma phocaea
jakupicae]
176
+ +
Dinarica albanica (S.H.F. Jaeckel, 1954)
[Chilostoma stenomphala albanica]
+ +
177
Dinarica petrovici (A.J. Wagner, 1914) [Chilostoma petrovici] + + +
Dinarica pouzolzi montenegrina (Rossmässler, 1836)
[Chilostoma p. montenegrina]
+ +
*Dinarica pouzolzi pouzolzi (Deshayes, 1830) [Chilostoma p.
pouzolzi]
178
+ +
+
Dinarica serbica (Kobelt, 1872) [Chilostoma serbica] + + +
Josephinella (?) korabensis (Subai, 1997)
179
Josephinella (?) zebiana (Sturany, 1907) [Chilostoma zebiana]
180
+ + +
Josephinella byshekensis (Knipper, 1941) [Chilostoma
byshekensis]
+ + +
*[Chilostoma edlaueri (Knipper, 1941)]
181
+ + +
Josephinella fuchsi (Knipper, 1939) [Chilostoma fuchsi] + + +
*Josephinella pindica (O. Boettger, 1886)
[Chilostoma subzonata pindica]
+ + +
182
Liburnica albanograeca (Subai, 1995) [Chilostoma
albanograeca]
183
+
184
Liburnica dochii (Sturany, 1907) [Chilostoma dochii] + + +
Liburnica dunjana (Knipper, 1941) [Chilostoma dunjana] + + +
Liburnica harpya (Subai, 1995) [Chilostoma harpya] + + +
Liburnica hoffmanni walteri (O. Boettger, 1886) [Chilostoma h.
walteri]
+ + +
Liburnica setigera setigera (Rossmässler, 1836) [Chilostoma s.
setigera]
+ + +
172
"systematic position of oestreichi is not clear, possibly H. secernenda or H. philibinensis"
173
First Albanian record: Dhora & Welter-Schultes (1999b).
174
"lives possibly in South Albania"
175
According to Subai (personal communication) its systematic position on the generic level is still doubtful.
176
Most probably does not live in Albania.
177
"systematic position not clear"
178
According to Subai (personal communication) does not live in Albania.
179
According to Subai (personal communication) the genus level position of this taxon is still doubtful.
180
According to Subai (personal communication) the genus level position of this taxon is still doubtful.
181
According to Subai (personal communication) this is a synonym for J. byshekensis.
182
Due to a misinterpretation of Subai (1995), Dhora & Welter-Schultes (1996) erroneously included this
species in the Albanian list. Most probably however, this species does not live in Albania.
183
First Albanian record: Fehér & Erıss (2009).
184
"probably lives in South Albania"
37
Superba grisea Subai & Fehér, 2006
Superba kulmakana Subai & Fehér, 2006
Superba skipetarica skipetarica (Subai, 1995) [Chilostoma
skipetaricus]
+ + +
Superba skipetarica asketa Subai & Fehér, 2006
*Superba reischuetzi Subai, 1990 [Chilostoma reschuetzi]
185
+ + +
*Arianta arbustorum (Linnaeus, 1758)
186
+ +
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... The species reaches the Black Forest in Germany and Poland in the north, and northern Greece at the southern limit of its range. It also occurs in the mountain areas of Albania (Fehér & erőss 2009) According to welteR-schultes (2012), this species is mainly found in moderately humid and rocky forests, but also in various other habitat types such as open grassy mountain slopes above the tree line or in habitats with scattered shrubs or trees. It also inhabits rocks rich in crevices and colonises retaining walls of dry stones (Rüetschi et al. 2012) or ruins of mediaeval castles (Juřičková & kučera 2004). ...
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Summary 1. Zebra mussels and their relatives (Dreissena spp.) have been well studied in eastern, central and western Europe as well as in North America, because of their invasiveness and economic importance. Much less is known about the biology and biogeography of indigenous (endemic) taxa of Dreissena, in the Balkans. A better knowledge of these taxa could help us (i) understand the factors triggering invasiveness in some taxa and (ii) identify other potentially invasive species. 2. Using a phylogenetic approach (2108 base pairs from three gene fragments), Dreissena spp. from natural lakes in the Balkans were studied to test whether invasive Dreissena populations occur in such lakes on the Balkan Peninsula, whether Dreissena stankovici really is endemic to the ancient Lakes Ohrid and Prespa, and to infer the phylogenetic and biogeographical relationships of Balkan dreissenids. 3. No invasive species of Dreissena, such as Dreissena polymorpha, were recorded. The supposedly ‘endemic’D. stankovici is not restricted to the ancient Lakes Ohrid and Prespa, but is the most widespread and dominant species in the west-central Balkans. Its southern sister taxon, Dreissena blanci, occurs sympatrically with D. stankovici in Lakes Prespa, Mikri Prespa and Pamvotis. Both species are classified into the subgenus Dreissena (Carinodreissena) of which the subgenus Dreissena (Dreissena) (which includes the invasive D. polymorpha) is the sister taxon. Dreissena blanci and D. stankovici are considered to represent distinct species. 4. On a global scale, the two Balkan species have small ranges. An early Pliocene time frame for the divergence of the subgenera Carinodreissena and Dreissena is discussed, as well as potential colonization routes of the most recent common ancestor of Carinodreissena spp. 5. The ambiguous taxonomy of dreissenids in the Balkans is addressed. As nominal D. blanci presbensis from Lake Prespa has nomenclatural priority over D. stankovici, the correct name for the latter taxon should be Dreissena presbensis.
169 "systematic position not clear" 170 "occurrence in Albania needs confirmation" 171 "occurrence in Albania needs confirmation" 172 "systematic position of oestreichi is not clear, possibly H. secernenda or H. philibinensis" 173 First Albanian record
  • Albania By Reischütz
First mentioned for Albania by Reischütz et al. (2008b) 169 "systematic position not clear" 170 "occurrence in Albania needs confirmation" 171 "occurrence in Albania needs confirmation" 172 "systematic position of oestreichi is not clear, possibly H. secernenda or H. philibinensis" 173 First Albanian record: Dhora & Welter-Schultes (1999b). 174 "lives possibly in South Albania"
) its systematic position on the generic level is still doubtful. 176 Most probably does not live in Albania. 177 "systematic position not clear" 178 According to Subai
  • According
  • Subai
According to Subai (personal communication) its systematic position on the generic level is still doubtful. 176 Most probably does not live in Albania. 177 "systematic position not clear" 178 According to Subai (personal communication) does not live in Albania.
Mollusc cenoses from different environments in Albania
  • D H Welter-Schultes
DHORA, DH. & WELTER-SCHULTES, F. W. (1999a): Mollusc cenoses from different environments in Albania.Schriften zur Malakozoologie 13: 61-66. Cismar/Ostholstein.