Article

Vegetational and environmental history during the Holocene in the Esbjerg area, west Jutland, Denmark

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Abstract

A pollen diagram from a site in the Esbjerg area, western Denmark, is used for reconstruction of the Holocene vegetational and environmental history there. During the Atlantic there was a parallel development of the landscape to that of other areas in Jylland (Jutland). From the late Neolithic onwards the development took its own course related to the approaching North Sea, which periodically inundated parts of the Esbjerg area. The record reflects landscape development in a formerly marine valley where sediments seem to be missing from parts of the Bronze Age and the early Iron Age. Consequently the landscape development during these times is only reflected in glimpses in the vegetation record, which shows gradually more open woodland and increasing human impact. During the late part of the Iron Age, Viking period and Middle Ages, the woodland was diverse in taxa but became increasingly open, finally reaching a stage during which there may have been too little wood even for daily use. At the same time the use of the land intensified. During the Sub-Atlantic, the Esbjerg area offered good natural resources with extensive grazing areas in the marine marshes in addition to good possibilities for farming and use of the woodland on higher ground, but devastating floods occurred.

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Holocene sediments of three closed Danish lake basins (Solsø, Skånsø, Kragsø) were used for the inference of post-glacial vegetational dynamics in former heathland areas in northern West Jutland, Denmark. The sites were selected to represent the major geomorphological units of West Jutland. The Holocene history of each lake basin was investigated by mapping of sediment distribution, analysis of loss-on-ignition, coarse inorganic matter, humus content, mineral magnetics, δ13C, pollen and selected other microfossils. These techniques were supplemented by plant macrofossil analysis at one site. Holocene terrestrial vegetational development was inferred at each site from analyses of pollen and microscopical charred particles. Chronologies were provided by numerous 14C dates. Stratigraphies of wet ground and terrestrial pollen and spore types were zooned by stratigraphically constrained cluster analysis. Based on the resultant site pollen asemblage zones (site PAZ), regional PAZ were proposed. Using modern analogues, Holocene floristic richness was estimated from pollen richness in the microfossil assemblages. The results support the hypothesis that disturbance is one of the most important mechanisms behind the maintenance of floristic richness. In particular, the response of estimated floristic richness to the intensity of vegetational fires followed the predictions of the Intermediate Disturbance Hypothesis. A period of elevated palynological richness and inferred vegetational disturbance was identified at all sites between 6000 and 5200 BC (calendar years). Using correspondence analysis (CA), the major gradient in the terrestrial pollen sequences was identified as a light-shade gradient, and CA first axis sample scores were used as a supplement to standard AP/NAP pollen ratios as an indicator of the shade-tolerance/light-demand of Holocene terrestrial plant communities. In spite of different vegetational developments since 4000 BC, the timing of major changes towards more light-demanding vegetation types were broadly synchronous at the three sites. Using chord distance as a dissimilarity index, rates of palynological change suggest that the interval between 8000 and 7500 BC (calendar years) was the period of most rapid vegetational change during the Holocene, both in terrestrial as well as lacustrine ecosystems. While climatic forcing of the rapid events around 8000 BC is hypothesised, the synchronous timing of relatively rapid inferred change in lake and terrestrial vegetation around AD 600 may reflect changes in climate as well as in land-use. Reducdancy analysis was used to develop a model between fire intensity (inferred from microscopical charred particles) and vegetational response, as reflected by pollen assemblages. Formulated at one site and tested at the two other sites, the model explains regional Calluna-heathland expansions as a result of vegetational burning. Similarly, declines in heathland cover are explained by lack of maintenance by fire. Regional vegetational development in northern West Jutland is reconstructed and special consideration is given to heathland history. The Holocene heathland development is interpreted as resulting from its importance for grazing. It is hypothesized that on poor soils, Calluna-dominated heathland was a better grazing resource than grass-dominated pasture, due to the winter-grazing offered by Calluna and the low palatibility of dominant grasses on poor soils. This hypothesis is relevant for the explanation of the variation in timing of heathland expansions on the different soil types represented by the study sites.
Article
Coastal dunefields have developed on the west coast of Jutland in Denmark over the past 5000 years. The dunefields are situated in a temperate climate zone with frequent high energy wind events. Dunefield development was characterized by repeated periods of transgressive dune formation punctuated by periods of dune stabilization and soil formation. The chronology of dunefield evolution is based on Accelerator Mass Spectrometry (AMS) radiocarbon dating of peaty palaeosols (24 samples) and Optically Stimulated Luminescence (OSL) dating of aeolian sand deposits (19 samples). These dates indicate that the completeness of the stratigraphic record varies considerably, but that the timing of aeolian activity events was identical in the dunefields examined here. Initiation of aeolian activity occurred around 2200 BC, 800 BC, AD 100, AD 1050–1200, and between AD 1550 and 1650. Proxy-climate data from bogs in southwestern Sweden suggest that these periods of dunefield activity were initiated during wet/cool summers. Most likely these climatic situations were associated with a more frequent passage of cyclones across Denmark in the summer seasons (increased storminess) causing aeolian sand movement and dune formation. The continued removal of vegetation in Jutland between 4000 BC and AD 1850 allowed the aeolian sand, when first activated, to move more and more easily across the coastal plains; the final phase of aeolian sand movement between AD 1550–1650 and 1900 had catastrophic implications for many coastal inhabitants in the region.
Article
Four elm declines were found in a pollen diagram from a small lake in northwest Denmark. Matching consecutive radiocarbon dates with the dendro-chronological calibration curve indicated a reservoir effect of 120 years, and dates for the four elm declines were obtained (4530, 4130, 3870, 3410 cal B.C.). The occurrence of apophytes (native plants encouraged by human activities) and increased vegetation diversity during the four elm declines indicates human disturbance. The first and second elm declines coincide with traces of early agriculture in northern Germany. The third and fourth elm declines are contemporary with the transitions to the Early Neolithic and the Middle Neolithic Funnel Beaker Culture in Denmark. The possible influence of outbreaks of elm disease is discussed.
Article
The coastal dunefield at Vejers (west coast of Jutland), which is now stabilized by vegetation, covers an area of approximately 120 km2 and occurs in a very high-energy wind regime. Along the coastline a narrow belt of 5–15 m high dune ridges occur. The dune ridges are gradually replaced inland by 10–20 m high parabolic or irregular dune forms. In the central part of the dunefield a large aeolian sand plain (ca. 50 km2) occurs at the windward side of a large and up to 20 m high parabolic dune.The 3D structure of the dunefield deposits have been studied by geomorphological analysis, sedimentological facies analysis of borings, trenches and natural exposures, and most importantly by georadar mapping. The aeolian deposits which overlie a middle to late Holocene barrier spit depositional system can be divided into a Lower unit that drapes the underlying barrier system topography, and an Upper unit that includes the present dunes. The boundary between the two aeolian units is a well-developed Phragmites peat. The Lower aeolian unit is composed of two aeolian subunits separated by an organic-rich horizon. Also the Upper unit is composite and composed of a basal aeolian sand cover and overlying dune or sand plain deposits. The dune deposits locally are composed of up to four depositional subunits separated by immature soils. The base of the Lower aeolian unit formed around 300 A.D., whereas the Phragmites peat at the base of the Upper aeolian unit has been dated to ca. 1000 A.D. From historical sources we know that the present dunefield primarily formed between 1550 and 1850 A.D.The sand content in the Upper unit is estimated to 550 × 106 m3, which yields sand transport rates between 25 m3 (m width)−1 yr−1 (accumulation in 1000 years), and 83 m3 (m width)−1 yr−1 (accumulation in 300 years). The sand in the dunefield originated from beach deposits. A large but pulsating supply of sand was supplied to the beaches by southwards running coastal currents.The composition of the Vejers dunefield deposits indicate that periods of dunefield growth alternated with periods of dunefield stabilization during the last ca. 1700 years. The final and most important phase of dunefield growth took place during ‘the Little Ice Age’. This period was characterized by an overall cold and stormy climate and a relative low sea level, and resulted in an increased availability of sand in the shorezone and a high influx of sand into the dunefield. Short periods of decreased storminess are recorded by the immature soils. Dunefield growth prior to 1000 A.D. was also linked to cold and stormy climatic intervals.
Article
The decline in Ulmus pollen frequencies that occurred ca. 5000 14C years ago before present (BP) is a key biostratigraphic marker horizon in northwest European pollen diagrams, although its causes are still a subject of debate. To investigate this event further, fungal spore analyses were carried out across the Ulmus decline at Moel y Gerddi, north Wales, United Kingdom. The Ulmus decline was in three phases, with a primary decline with low cereal and Rumex pollen records as the only agricultural indicators. This was followed by a more significant decline, with general forest opening and the grassland/pasture indicator Plantago lanceolata. A third, less significant, decline was again accompanied by cereal-type pollen. Tree pollen frequencies subsequently recovered, with cereal-type pollen remaining well represented. The fungal data recorded woodland taxa and a background level of the obligate dung fungus Sporormiella, a proxy for local herbivore abundance. Sporormiella frequencies increased greatly after the main Ulmus decline, around which there were high percentages of the ascospores of the wood rot fungus Kretzschmaria deusta. The neoecology of Kretzschmaria deusta, and the behavior of its spore curve suggests the colonisation of local populations of already severely wounded trees. At this site Ulmus, and perhaps Tilia were infected at a time of markedly increased inferred herbivore concentrations. Neolithic farming techniques could have provided mortally wounded trees while enhancing livestock grazing, although the role of disease must also be considered. Elevated Kretzschmaria deusta values may be of ecological significance in the interpretation of the causes and nature of the Ulmus decline and similar forest disturbance events, and the indicator role of this fungus in forest paleoecology requires further study.
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