Project

Angiosperm Phylogeny Poster (APP)

Goal: Educational posters giving a clear overview of the hereditary relationships among angiosperms - the evolution of flowering plants - phylogeny with characteristics of the major lineages - regularly updated - now available in 31 languages.

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Project log

Theodor C. H. Cole
added a research item
Dashzeveg N, Cole TCH, Hilger HH , Magsar U, Enkhtaivan E, Batlai O (2022) Plant phylogeny posters – useful digital teaching tools for botanical education. Mongolian Journal of Biological Sciences 20(1): 69-72 >>>>>>> Visual documentation and presentation are essential for botanical education. Wallcharts and posters depicting plant structures and evolutionary relationships as phylogenetic trees have served as essential tools in teaching. With the advancement of digital technologies, new approaches to a traditional concept have become feasible. The Plant Phylogeny Poster project is a collaborative initiative providing clear and concise overviews of orders, families and genera of various groups of plants. The global project now involves around 150 botanists from all around the world with currently 400 posters in 35 languages (as of March 2022). All posters are freely available online and regularly updated, providing an extensive portfolio of versatile new teaching resources for educators. Key words: Angiosperm, Tracheophyte, Bryophyte, Teaching Tool
Theodor C. H. Cole
added an update
"However, there are a few changes in relationships and new families/orders that have been made since APG IV. The latter are listed immediately below, while in the summary they are indicated by asterisks. Some of these changes are new families that reflect recent collections, e.g. the remarkable Tiganophytaceae, from the Namib area and in Brassicales, while others, like Oncothecales, a very small order that may well be sister to all other core asterids/gentianids, are the results of changed ideas of relationships that are being suggested by comprehensive analyses using nuclear genomes. This latter kind of change in particular is being made only when support for the changed relationships is strong, otherwise confusion will result. However, now that PAFTOL is off the ground, with an immediate goal of including all genera of flowering plants (W. J. Baker et al. 2021a, b: see also Seed Plant Tree of Life), who knows what the future holds? Note that many relationships along the spine of the Pentapetalae well up into the asterids are subject to change! All changes are being carefully justified.
MAJOR ADDITIONS/CHANGES SINCE 2016
Brassicales: add Tiganophtyaceae.
Santalales: add Mystropetalaceae.
Core Asterids/gentianids: add Oncothecales: Oncothecaceae.
Lamiales: add Wightiaceae
CAMPANULID/ASTERID II: Add Cardiopteridales: [Cardiopteridaceae + Stemonuraceae]
Add: Desfontainiales: Columelliaceae"
quoted from Stevens, P. F. (2001 onwards). Angiosperm Phylogeny Website, http://www.mobot.org/MOBOT/research/APweb/
These changes, among others, will be implemented in the next update of the "Angiosperm Phylogeny Poster"
 
Theodor C. H. Cole
added an update
A recent plastid phylogenomic analysis (PPA II) is largely confirming previous results of plastid DNA phylogenetic reconstructions in angiosperm relationships. One statement there: "Thus, we suggest separating Dasypogonales from Arecales, Sabiales from Proteales, Huales from Oxalidales, and Oncothecales from Icacinales, ...." Li HT, Luo Y, Gan L et al. 2021. Plastid phylogenomic insights into relationships of all flowering plant families. BMC Biology 19:232; https://doi.org/10.1186/s12915-021-01166-2
We, the authors of the "Angiosperm Phylogeny Poster", have decided to wait for more nuclear DNA data before making a next update to our poster (based on PPA I).
 
Sergei L. Mosyakin
added a research item
Historically, wallcharts and posters created by botanical illustrators, often highly skilled artists, have played an important role in teaching botany at the university level. Large-scale panels and posters can visualize complex interrelationships and entire stories in a clear and appealing overview in graphs, tables, and diagrams. Carrying this concept of educational tools into the electronic era, the Plant Phylogeny Poster project uses this approach for displaying evolutionary relationships in systematic botany. The Angiosperm Phylogeny Poster (APP) displays, as phylogenetically arranged clades, the orders and families of flowering plants (with orders hyperlinked to APweb, Stevens, 2001-onwards), the Tracheophyte Phylogeny Poster (TPP) families and genera of ferns and gymnosperms, and the Bryophyte Phylogeny Poster (BPP) orders and families of liverworts, mosses, and hornworts. The portfolio currently also includes about 30 posters on individual orders and families of angiosperms. Each group within these evolutionary trees is matched with essentially relevant morphological features, biogeographic occurrences, and other information in compactly condensed text blocks. All posters are freely available online, some in more than 30 languages, coauthored by a team of more than 130 botanists. The posters are regularly updated, current literature is cited. The project is expanding steadily and rapidly.
Theodor C. H. Cole
added a research item
Cole TCH , Hilger HH, Stevens PF, Gavrilović M, Janaćković P (2021) Filogenija skrivenosemenica – Sistematika cvetnica • hipotetičko stablo na osnovu molekularno-filogenetičkih podataka (2019) • filogenija, klasifikacija i karakteristike prvenstveno prate APweb, APG i PPA (Plastid Phylogenomic Angiosperm: Li HT et al. 2019) • 64 reda i oko 420 familija trenutno prepoznato od strane APG IV (neke manje familije su isključene) • suprotno od APG, ali u saglasnosti sa APweb-om i drugim originalnim izvorima, ovde je prepoznato nekoliko familija unutar Boraginales • dužina grana proizvoljna, ne izražava stvarnu vremensku skalu • navedeni karakteri nužno ne pripadaju svim članovima odgovarajuće klade • položaj raznih karaktera na stablu je provizoran • za karakteristike familija videti: FGVP, Kubitzki K, ed. (1990 ff) • redovi (i više kategorije) su povezani sa odgovarajućim stranicama na APweb-u • brojevi u sivoj boji postavljeni ispred redova se odnose na familije, rodove i vrste * Ceratophyllum: takozvani tepali bi mogli biti brakteje (pitanje koje je predmet tekućih istraživanja i razmatranja) • Ovaj poster je sada dostupan na 31 jezika (molimo vidite stranice autora na ResearchGate-u)
Theodor C. H. Cole
added a research item
COLE TCH , HILGER HH, STEVENS PF, VLADIMIROV V, STOYANOV S (2021) ФИЛОГЕНИЯ НА ПОКРИТОСЕМЕННИТЕ, Систематика на Покритосеменните (Цветните) растения. Българска версия на: COLE TCH, HILGER HH, STEVENS PF (2020) Angiosperm Phylogeny Poster – Flowering Plant Systematics ● Хипотетично филогенетично дърво, основано на молекулярните филогенетични данни (2020) ● филогенията, класификацията и белезите във висока степен следват интернет страницата (APweb) и групата (APG) за филогения на Покритосеменните растения (Angiospermae) ● 64 разреда и около 420 семейства в момента признати от APGIV (изключени са само някои малки семейства) ● в противоречие с APG, но в съответствие с APweb и други авторитетни източници, тук приемаме няколко семейства в Boraginales ● дължините на разклоненията не отразяват действителните мащаби на времето ● изброените белези не се отнасят непременно за всички членове на съответния клад (клъстер). ● позицията на различни белези върху дървото е несигурна ● за характеристиките на семействата вж. FGVP, Kubitzki K., ed. (1990+) ● разредите (и по-високите нива) са свързани със съответните страници на APweb ● числата, посочени в сиво до разредите, се отнасят за броя на семействата, родовете и видовете ● Постерът вече е достъпен на 30 езика (моля, вж. страниците на авторите в ResearchGate)
Theodor C. H. Cole
added an update
List of all 228 posters (as of Feb. 2021) with hyperlinks to each poster on ResearchGate * click on the ‘x’ within a box to open the respective page of the poster on ResearchGate from which it can be downloaded
 
Theodor C. H. Cole
added an update
Two recent nDNA/transcriptome analyses of asterids by Stull et al.* and Zhang et al.** are pointing to a possible position of Oncothecaceae as sister to all other core asterids and an immediate sister relationship between Ericales and Cornales (then best referred to as: EC clade) – and that the families currently assigned to Icacinales and Aquifoliales are not monophyletic. Further studies will be needed to confirm these hypotheses. In a further nuclear/transciptome study by Yang et al.***, an attempt was made to elucidate the general "deep" backbone of the angiosperm tree, resulting in the proposed topology: monocots, magnoliids, Chloranthales, Ceratophyllales, eudicots. However, the conflicting results in this study lead those same authors to state that a "fully bifurcating tree may be an inadequate representation of the early radiation of angiosperms".
Thus, we have decided to retain the present topology of the "Angiosperm Phylogeny Poster" which greatly follows PPA (cpDNA) and wait for further developments.
* Stull et al. (2020) AJB 107(5): 790-805;
** Zhang et al. (2020) MBE, doi.org/10.1093/molbev/msaa160;
*** Yang et al. (2020) Plant Comm., doi.org/10.1016/j.xplc.2020.100027
 
Theodor C. H. Cole
added a research item
Seit Jahrhunderten bemühen sich Botaniker, „Ordnung ins System“ zu bringen – für die Blütenpflanzen ist man diesem Ziel nun sehr nah. Ein Konsortium international führender Pflanzensystematiker hat die Ergebnisse der molekularen Phylogenie der vergangenen drei Jahrzehnte zusammengefasst und unter Berücksichtigung morphologisch-anatomischer und chemotaxonomischer Gesichtpunkte daraus die Verwandtschaftsbeziehungen der Blütenpflanzen reinterpretiert. Es resultiert ein evolutionär begründeter Stammbaum, der auf der Ebene der Hauptgruppen (Ordnungen und Familien) nun ein hohes Maß an Stabilität aufweist.
Theodor C. H. Cole
added a research item
COLE TCH , HILGER HH, STEVENS PF, ENROTH J (2019) KOPPISIEMENISTEN FYLOGENIA JA SYSTEMATIIKKA • hypoteettinen puu perustuu molekyyliaineistoon lähinnä plastideista (toukokuu 2019) • fylogenia, luokittelu ja ominaisuudet: APweb, APG ja PPA (Plastid Phylogenomic Angiosperm, ks. Li HT ym. 2019) • APG IV: n mukaan lahkoja on 64 ja heimoja n. 420 (muutama pieni heimo puuttuu tästä puusta) • kuten APweb ja muut keskeiset lähteet, sisällytämme Boraginales-lahkoon enemmän heimoja kuin APG • haarojen pituudet eivät osoita aikaskaalaa • luetellut ominaisuudet eivät välttämättä esiinny kaikissa kladin jäsenissä • joidenkin ominaisuuksien sijainti puussa on epävarma • heimotason ominaisuudet, ks. FGYP, Kubitzki K., toim. (1990-) • lahkot (ja niitä korkeammat taksonit) on linkitetty APwebin vastaaviin sivuihin • harmaalla fontilla olevat luvut lahkojen vieressä tarkoittavat heimoja, sukuja ja lajeja • Ceratophyllum: oletetut verholehdet saattavat olla tukilehtiä • Posteri on nyt käännetty 29 kielelle (löytyvät kirjoittajien sivuilta ResearchGatessa)
Theodor C. H. Cole
added a research item
КОУЛ TКГ, ХИЛЬГЕР XГ, СТИВЕНС ПФ, ТУРГИНОВ OT (2020) ЁПИҚ УРУҒЛИ ЎСИМЛИКЛАР ФИЛОГЕНИЯСИ (ШАЖАРАСИ), Ўзбекча версияси: КОУЛ TКГ, ХИЛЬГЕР XГ, СТИВЕНС ПФ, (2020) Ёпиқ уруғли ўсимликлар филогенияси плакати – гулли ўсимликлар тизими, Uzbek version of: COLE TCH, HILGER HH, STEVENS PF (2020) Angiosperm Phylogeny Poster – Flowering Plant Systematics • Молекуляр филогенетик маълумотларга асосланган гипотетик филогенетик дарахт (шажара) (2019) • Филогения таснифлар ва белгилар асосан APweb, APG ва PPA бўйича берилган (Li HT ва бошқалар, 2019) • APG IV тизимида 64 тартиб ва тахминан 420 оила жамланган (айрим кичик оилалар инобатга олинмаган) • APG дан фарқ қилган ҳолда, APweb ҳамда бошқа манбаалар сингари биз Boraginales тартибидаги биз бир нечта оилаларни тан оламиз • APG дан фарқ қилган ҳолда, лекин The Caryophyllales Global Synthesis Initiative (Hernández-Ledesma et al. 2015) биз Chenopodiaceae оиласини Amaranthaceae оиласидан алоҳида оила сифатида тан оламиз • дендрограмма шажарасидаги шохларнинг узунлиги эхтимолий бўлган ва кладларнинг ажралишининг ҳақиқий вақтини белгиламайди • бу ерда келтирилган белгилар тегишли кладларнинг барчасида бўлмаслиги
Theodor C. H. Cole
added 2 research items
COLE TCH, HILGER HH, STEVENS PF, MAJESKÝ Ľ, MÁRTONFI P (2020) FYLOGENÉZA KRYTOSEMENNÝCH RASTLÍN, Slovenský preklad COLE TCH, HILGER HH, STEVENS PF (2020) Angiosperm Phylogeny Poster – Flowering Plant Systematics • hypotetický fylogram odvodený z molekulárno-fylogenetických dát (2020) • fylogenéza, klasifikácia, ako aj jednotlivé znaky vychádzajú z APweb-u a APG systému a PPA (Li HT et al. 2019) • poster zobrazuje 64 radov a približne 420 čeľadí v súčastnosti rozoznávaných systémom APG IV (niektoré menšie čeľade neboli do tejto verzie zahrnuté) • na rozdiel od APG, no v súlade s APweb-om a ostatnými zdrojmi, je tu rozoznávaných viacero čeľadí v rámci radu Boraginales • dĺžka vetiev je arbitrárna a nevyjadruje skutočnú časovú divergenciu • uvádzané znaky nemusia byť nevyhnutne prítomné pri všetkých členoch danej skupiny • pozícia viacerých znakov na zobrazenom fylograme je neistá • pre charakterizáciu jednotlivých čeľadí pozri: FGVP, Kubitzki K, ed. (1990 ff) • rady (a vyššie vetvy) sú prepojené so stránkami „APweb“
COLE TCH, HILGER HH, STEVENS PF, MAJESKÝ Ľ, HRONEŠ M (2020) FYLOGENEZE KRYTOSEMENNÝCH ROSTLIN, Český překlad COLE TCH, HILGER HH, STEVENS PF (2020) Angiosperm Phylogeny Poster – Flowering Plant Systematics • hypotetický strom odvozený z molekulárně-fylogenetických dat (2020) • fylogeneze, klasifikace a znaky vychází z APwebu, APG systému, a PPA (Plastid Phylogenomic Angiosperm: Li HT et al. 2019) • na plakátu je vyobrazeno 64 řádů a přibližně 420 čeledí v současnosti rozeznávaných systémem APG IV (některé z menších čeledí zde nejsou zahrnuty) • na rozdíl od systému APG, ale v souladu s APwebem a dalšími zdroji, je zde rozeznáváno několik čeledí v rámci řádu Boraginales • délka větví je arbitrární a nevyjadřuje časovou divergenci • uvedené znaky nemusí byt nevyhnutelně přítomny u všech zástupců dané skupiny • pozice vícero znaků na uvedeném stromě je nejistá • pro charakterizaci jednotlivých čeledí viz: FGVP, Kubitzki K, ed. (1990 ff) • řády (a vyšší linie/skupiny) jsou propojeny se stránkami APweb
Theodor C. H. Cole
added an update
Two recent phylogenomic studies based on nuclear genome data1,2 suggest slightly different topologies in some parts of the tree, especially immediately basal to rosids and asterids. These vary with respect to the strictly plastome-based PPA tree. We are waiting for further studies before making changes to the topology of the Angiosperm Phylogeny Poster (APP).
1Zhao T et al. (2020) Novel phylogeny of angiosperms inferred from whole-genome microsynteny analysis. bioRxiv, doi: http://dx.doi.org/10.1101/2020.01.15.908376.
2One Thousand Plant Transcriptomes Initiative (2019) One thousand plant transcriptomes and the phylogenomics of green plants. Nature 574: 679-685, https://doi.org/10.1038/s41586-019-1693-2
 
Theodor C. H. Cole
added a research item
The Angiosperm Phylogeny Poster – Flowering Plant Systematics is an educational poster presenting an overview of the evolutionary relationships among flowering plants, based on APG IV/APweb and in this update chiefly following the topology suggested by plastid phylogenomic relationships (see: Li HT et al. 2019. Origin of angiosperms and puzzle of the Jurassic gap. Nature Plants 5: 461–470; Fig. 2 therein). The phylogenetic tree depicts the 64 orders and the majority of the 416 families, listing the main apo- and plesiomorphies as well as diagnostic and nondiagnostic anatomical, morphological, and phytochemical features for orders and higher ranks within the tree along with the number of families, genera, and species for each order. The intuitive color-coding facilitates memorization and teaching. The poster is now available in 29 languages (all downloadable from ResearchGate).
Theodor C. H. Cole
added an update
We are in the process of updating all posters in the APP series to chiefly comply with the recently published plastid phylogenomic angiosperm (PPA) tree: Li HT et al. (2019) Origin of angiosperms and the puzzle of the Jurassic gap, Nature Plants 5: 461–470, Fig. 2 therein. Major changes include:
1) Petrosaviales sister to Asparagales
2) Dilleniales sister to superrosids
3) Asterales + Escalloniales clade
4) Santalales (rather than Berberidopsidales) sister to remaining superasterids
5) COM clade: Oxalidales (Celastrales + Malpighiales)
6) resolution of the polytomy in Lamiids
 
Theodor C. H. Cole
added an update
We are proud to present to the community with the Vietnamese version of the APP by Tran The Bach of the INSTITUTE OF ECOLOGY AND BIOLOGICAL RESOURCES, VAST, Hanoi, now up and running! ... this is the 25th language of the globally successful Angiosperm Phylogeny Poster (APP) involving a team of 31 internationally renowned scholars, all researchers and teachers of systematic botany!
 
Theodor C. H. Cole
added a research item
COLE TCH , HILGER HH, STEVENS PF, TRAN TB (2019) Poster PHÁT SINH CHỦNG LOẠI CỦA THỰC VẬT HẠT KÍN Bản poster này hiện có 25 ngôn ngữ (tham khảo trang các tác giả trên ResearchGate) ... now following PPA!
Theodor C. H. Cole
added an update
Here, Akitoshi Iwamoto is teaching the Japanese APP in a FLO-RE-S "Flower Morphology Course" in Santiago de Chile in 2018. He places the vials with samples of specimens on the respective branches ... inspiring!
 
Theodor C. H. Cole
added 3 research items
COLE TCH, HILGER HH, STEVENS PF, KRIGAS N, KAGIAMPAKI A (2019) Αφίσα φυλογένεσης των Αγγειοσπερμων - Συστηματική των ανθοφόρων φυτών; Greek version of: Angiosperm Phylogeny Poster - Flowering Plant Systematics (2019) ... now following PPA. Improved 2021!
NEW UPDATE! The ANGIOSPERM PHYLOGENY POSTER (APP) – Flowering Plant Systematics is an educational tool presenting an overview of the evolutionary relationships among flowering plants according to APG IV, APweb, and PPA (Li HT et al. Nature Plants 2019). The phylogenetic tree depicts the 64 orders and the majority of the 416 families, listing the main apomorphies and plesiomorphies as well as diagnostic and nondiagnostic anatomical, morphological, and phytochemical features for orders and higher levels within the tree along with the number of families, genera, and species for each order. The intuitive color-coding facilitates memorization and teaching. Translations of the poster by internationally renowned botanists are now available in 29 languages. Hyperlinks to APweb (Peter F. Stevens, Missouri Botanical Garden) are provided for the orders and higher ranks. This is one in a series of three educational posters on the phylogeny of land plants: Poster 1: "Angiosperms", Poster 2: "Tracheophytes: Lycophytes, Ferns, Gymnosperms", Poster 3: "Bryophytes: Liverworts, Mosses, Hornworts".
نسالة كاسيات البذور Arabic version of: Angiosperm Phylogeny Poster - Flowering Plant Systematics, 2019 (now following PPA) - cite as: COLE TCH, HILGER HH, STEVENS PF, FENNANE M (2019) نسالة كاسيات البذور تصنيف النباتات المزهرة
Theodor C. H. Cole
added a research item
COLE TCH, HILGER HH, STEVENS PF (2019) Stammbaum der Blütenpflanzen - Phylogenie und Systematik der Angiospermen, 2019. Aktualisierte Version des Lehr- und Lernposters zur Phylogenie der Blütenpflanzen gemäß APG IV, APweb und PPA (Li HT et al. 2019). Merkmale wurden aktualisiert und optimiert; die Anzahl der Arten, Gattungen und Familien erscheinen links neben den einzelnen Ordnungen; zusätzliche Familien wurden aufgenommen. Dies ist Teil einer Trilogie zur Phylogenie der Landpflanzen: Poster 1: "Bryophyten", Poster 2: Tracheophyten, Poster 3: Blütenpflanzen. Die drei Lehr- und Lernposter werden regelmäßig aktualisiert und bilden den derzeitigen Stand der Forschung ab.
Theodor C. H. Cole
added an update
A recent paper by Chen JH et al. (2018) in Nature Plants provides further evidence for the sister position of magnoliids in relation to monocots + eudicots >>> DOI: 10.1038/s41477-018-0323-6
... while Chaw et al. (2018) in BioRxiv find monocots as sister to the rest >>>> DOI: https://doi.org/10.1101/371112
Soltis and Soltis comment on this in Nature Plants 5: 6–7 (2019)
 
Theodor C. H. Cole
added an update
In a recent major plastome multi-gene analysis of monocots, Givnish et al. (AJB, 2018) propose "to recognize Dasypogonales as its own order rather than as part of an extraordinarily anomalous, expanded order of "palms", and that the next update of the APG schema should be emended accordingly". Further, they propose to add Thismiaceae to Dioscoreales.
 
Theodor C. H. Cole
added 3 research items
Poster: Filogenesi delle Angiosperme – Sistematica delle piante a fiore (2019). The Italian version of the Angiosperm Phylogeny Poster – Flowering Plant Systematics by Theodor C. H. Cole (Heidelberg University, Germany) and Hartmut H. Hilger (Free University Berlin, Germany), and Peter F. Stevens (Missouri Botanical Garden) translated by Federico Selvi (University of Florence, Italy). Cite as: Cole TCH, Hilger HH, Stevens PF, Selvi F (2019) Poster: Filogenesi delle Angiosperme – Sistematica delle piante a fiore (Italian version of the Angiosperm Phylogeny Poster – Flowering Plant Systematics). Poster: Filogenesi delle Angiosperme – Sistematica delle piante a fiore (2019). La versione italiana del Poster Angiosperm Phylogeny – Flowering Plant Systematics di Theodor C. H. Cole (Heidelberg University, Germany), Hartmut H. Hilger (Free University Berlin, Germany) e Peter F. Stevens (Missouri Botanical Garden), tradotto da Federico Selvi (Università di Firenze, Italia). Citazione: Cole TCH, Hilger HH, Stevens PF, Selvi F (2019) Poster: Filogenesi delle Angiosperme – Sistematica delle piante a fiore (versione italiana del Poster Angiosperm Phylogeny – Flowering Plant Systematics). ... now following PPA !
Póster: Filogenia de las Angiospermas – Sistemática de las plantas con flores (2019). The Spanish version of the "Angiosperm Phylogeny Poster – Flowering Plant Systematics" by Theodor C. H. Cole (Heidelberg University, Germany) and Hartmut H. Hilger (Free University Berlin, Germany), and Peter F. Stevens (MoBot, St. Louis, Missouri) translated by Diego Medan (University of Buenos Aires, Argentina). Cite as: Cole TCH, Hilger HH, Stevens PF, Medan D (2019) Póster: Filogenia de las Angiospermas – Sistemática de las plantas con flores (Spanish version of the Angiosperm Phylogeny Poster – Flowering Plant Systematics) - updated to PPA >>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>> Póster: Filogenia de las Angiospermas – Sistemática de las plantas con flores (2019). Versión en español de "Angiosperm Phylogeny Poster– Flowering Plant Systematics" por Theodor C. H. Cole (Heidelberg University, Alemania), Hartmut H. Hilger (Free University Berlin, Alemania) y Peter F. Stevens (MoBot, St. Louis, Missouri), traducido por Diego Medan (University of Buenos Aires, Argentina) actualizado de acuerdo con la APG IV. Cite: Cole TCH, Hilger HH, Stevens PF, Medan D (2019) Póster: Filogenia de las Angiospermas – Sistemática de las plantas con flores (Spanish version of the Angiosperm Phylogeny Poster – Flowering Plant Systematics) - updated to PPA
Theodor C. H. Cole
added a research item
Phylogénie des Angiospermes – Systématique des Plantes à Fleurs, 2020 (French version of: Angiosperm Phylogeny Poster - Flowering Plant Systematics), citez comme: COLE TCH, HILGER HH, STEVENS PF, BACHELIER JB (2020) Phylogénie des Angiospermes - Systématique des Plantes à Fleurs (version française du Angiosperm Phylogeny Poster) ... adapted to PPA
Theodor C. H. Cole
added an update
The Angiosperm Phylogeny Poster now in 24 languages by a team of 30 collaborators worldwide!
 
Theodor C. H. Cole
added a research item
COLE TCH, HILGER HH, STEVENS PF, KAINULAINEN K (2019) Angiosperm-Fylogeni – Blomväxternas släktträd, Svensk version av: COLE TCH, HILGER HH, STEVENS PF (2019) Angiosperm Phylogeny Poster – Flowering Plant Systematics • Hypotetiskt släktträd baserat på molekylärfylogenetiska data (2019) • Fylogeni, klassifikation och beskrivningar främst tagna från APweb, APG och PPA • 64 ordningar och ungefär 420 familjer erkänns i APG IV (några av de mindre familjerna har exkluderats här) • I motsatts till APG, men i enlighet med APweb och andra referenser, så erkänner vi här flera familjer inom Boraginales • Grenlängderna är medvetet ritade så att de inte står i korrelation till tid • De karaktärer som listas här återfinns inte nödvändigtvis hos alla taxa inom kladen • Karaktärernas positioner på trädet är osäkra • För familjebeskrivningar se: FGVP, Kubitzki K, ed. (1990 ff) • Ordningar (och högre taxonomiska ranger) har länkats till motsvarande hemsidor på APweb • De nummer som anges i grå text brevid namnen på ordningarna återspeglar antalet familjer, släkten och arter * Ceratophyllum: det som har betraktats som tepaler skulle kunna vara braktéer (detta är en fråga som undersöks och debatteras) - Denna affisch finns nu tillgänglig på 25 språk (se författarnas hemsidor på ResearchGate) ... now following PPA!
Theodor C. H. Cole
added a research item
وعائی تخم کا نسلی ارتقاء COLE TCH, HILGER HH, STEVENS PF, ABID R (2019) Urdu version of: COLE TCH, HILGER HH, STEVENS PF (2019) Angiosperm Phylogeny Poster – Flowering Plant Systematics (now following PPA) by Prof. Dr. Rubina Abid, University of Karachi, Pakistan • Hypothetical tree based on molecular phylogenetic data (2019) • phylogeny, classification, and features chiefly follow APweb, APG, and PPA (Li HT et al. 2019) • 64 orders and around 420 families currently recognized by APG IV (some minor families excluded here) • contrary to APG, but in compliance with APweb and other seminal sources, we here recognize several families within Boraginales • branch lengths deliberate, not expressing actual time scale • the characters listed do not necessarily apply to all members of the according clade • position of various characters on the tree uncertain • for family characteristics see: FGVP, Kubitzki K, ed. (1990 ff) • orders (and higher ranks) are linked to the according pages on APweb • This poster is now available in 25 languages (please refer to the authors' pages on ResearchGate) ... now following PPA!
Theodor C. H. Cole
added a research item
Ишлэл: КОУЛ TКГ, ХИЛЬГЕР ХГ, СТИВЕНС ПФ, ОЮУНЦЭЦЭГ Б, НЯМБАЯР Д, УРГАМАЛ М (2019) Далд үртний филогени – Цэцэгт ургамлын ангилалзүй, Mongolian version of: Cole TCH, Hilger HH, Stevens PF (2019) Angiosperm Phylogeny Poster – Flowering Plant Systematics ... now following PPA!
Theodor C. H. Cole
added a research item
COLE TCH, HILGER HH, STEVENS PF, KUMAR S, SINGH P: अावृतबीजीय पौधों की फलोभिद्की : पुष्पीय पादप वर्गिकी Hindi-Devanagari Version by the Botanical Survey of India, Kolkata, of: Cole TCH, Hilger HH, Stevens PF, Angiosperm Phylogeny Poster (APP) 2019 ... now chiefly complying with the topology of PPA (Li HT et al. 2019)
Theodor C. H. Cole
added 2 research items
COLE TCH, HILGER HH, STEVENS PF, TRIBOUN P (2019) วิวัฒนาการชาติพันธุ์พืชดอก อนุกรมวิธานพืชดอก, Thai version of COLE TCH, HILGER HH, STEVENS PF (2019) Angiosperm Phylogeny Poster – Flowering Plant Systematics ... now complying with PPA (Li HT et al. 2019)
COLE TCH, HILGER HH, STEVENS PF, BARINA Z (2019) A Zárvatermők Törzsfejlődésének Posztere – A virágos növények rendszerezése Hungarian version of: COLE TCH, HILGER HH, STEVENS PF (2019) Angiosperm Phylogeny Poster – Flowering Plant Systematics • a hipotetikus törzsfa molekuláris filogenetikai adatokon alapul (2019) • a törzsfejlődés, a rendszertan és a jellemzők döntően az APweb, APG és PPA (Li HT et al. 2019) alapján • az APG IV jelenleg 64 rendet és 420 családot tárgyal (néhány kisebb család itt hiányzik) • az APG-vel szemben, de az APweb-bel és más forrásokkal összhangban itt számos családot a Borágóvirágúak rendjében tárgyalunk • a törzsfa ágainak hosszúsága nem arányos az időskálával • a felsorolt jellemzők nem feltétlenül igazak az adott klád minden tagjára • különféle jellemzők fán belüli helyzete bizonytalan • családjellemzések itt érhetők el: FGVP, Kubitzki K, ed. (1990 ff) • a rendek (és efeletti kategóriák) az APweb megfelelő oldalaihoz kapcsolódnak • a rendek melletti szürke számsorok családokra/nemzetségekre/fajokra vonatkoznak * Ceratophyllum: az ún. leplek murvák lehetnek (további kutatásokat igényel és vita tárgya). Ez a poszter 25 nyelven érhető el (kérjük, hivatkozzon a szerzők ResearchGate-oldalára)
Theodor C. H. Cole
added a research item
COLE TCH, HILGER HH, STEVENS PF, GUDŽINSKAS Z (2019) Gaubtasėklių filogenija – žiedinių augalų sistema, Lithuanian version of: Cole TCH, Hilger HH, Stevens PF (2019) Angiosperm Phylogeny Poster – Flowering Plant Systematics, ... NOW FOLLOWING PPA ! • hipotetinis medis pagrįstas molekulinės filogenijos duomenimis (2019) • filogenija, klasifikacija ir požymiai pateikti pagal APweb, APG ir PPA (Li HT et al. 2019) duomenis • dabar APG IV (kai kurios nedidelės šeimos čia neįtrauktos) pripažįstamos 64 eilės ir maždaug 420 šeimų • kitaip negu APG, bet remiantis APweb ir kitais šaltiniais, čia pripažįstamos kelios Boraginales eilės šeimos • šakų ilgis pasirinktas laisvai, jis neatitina laiko skalės • išvardyti požymiai nebūtinai tinka visiems tam tikrai šakai priklausantiems atstovams • daugelio požymių pasiskirstymas iki šiol neaiškus • šeimų charakteristikas galima rasti: FGVP, Kubitzki K, ed. (1990–) • eilės (ir aukštesnio rango grupės) susietos laikantis APweb pateiktos schemos • pilki skaičiai, parašyti šalia eilių, reiškia šeimų, genčių ir rūšių skaičių toje eilėje ; Plakatas parengtas 25 kalbomis (juos galima rasti ResearchGate svetainėje esančiose autorių paskyrose)
Theodor C. H. Cole
added a research item
КOУЛ TКГ, ХИЛЬГЕР XГ, СТИВЕНС ПФ, ГОДИН BH (2022) ФИЛОГЕНИЯ ПОКРЫТОСЕМЕННЫХ # Russian version of: Cole TCH, Hilger HH, Stevens PF (2019) Angiosperm Phylogeny Poster – Flowering Plant Systematics * This poster is a revision based on an earlier Russian-language version by Sergei L. Mosyakin and Anastasiya V. Odintsova, still available from those authors
Theodor C. H. Cole
added 2 research items
КОУЛ TКГ, ХІЛЬГЕР XГ, СТІВЕНС ПФ, МОСЯКІН CЛ, ОДІНЦОВА AB (2019) ФІЛОГЕНІЯ ПОКРИТОНАСІННИХ – Систематика квіткових рослин, Ukrainian version of: COLE TCH, HILGER HH, STEVENS PF (2019) Angiosperm Phylogeny Poster – Flowering Plant Systematics - Порядки і групи вищого рангу тепер мають лінки на відповідні сторінки на APweb, APG и PPA (Li HT et al. 2019). Цей постер тепер опубліковано 25 мовами (див. сторінки авторів на ResearchGate) ... now following PPA!
:به صورت زیر ارجاع داده شود COLE TCH, HILGER HH, STEVENS PF, SALMAKI Y, ZARRE S (2019) Angiosperm Phylogeny Poster – Systematics of Flowering Plants [Farsi Translation] of کل، تی.سی.اچ.، هيلگر، اچ.اچ.، استيونز، پی.اف.، سلمکی، ی.، زارع، ش. (١۳۹٧) پوستر تبارزایی نهاندانگان – سازگان شناسی گياهان گل دار ... now following PPA!
Theodor C. H. Cole
added a research item
COLE TCH, HILGER HH, STEVENS PF, BARANOW P (2019) FILOGENEZA ROŚLIN OKRYTOZALĄŻKOWYCH – Systematyka Roślin Kwiatowych, Polskie tłumaczenie COLE TCH, HILGER HH, STEVENS PF (2019) Angiosperm Phylogeny Poster – Flowering Plant Systematics ... NOW FOLLOWING PPA! • hipotetyczne drzewo oparte na danych molekularnych (2019) • filogeneza, klasyfikacja i cechy głownie w oparciu o APweb, APG i PPA (Li HT et al. 2019) • 64 rzędy i około 420 rodzin aktualnie uznawanych przez APG IV (niektóre mniejsze rodziny pominięto) • w przeciwieństwie do APG, jednak zgodnie z APweb i innymi źródłami, wyróżniono kilka rodzin w rzędzie Boraginales • długość gałęzi nie wyraża rzeczywistej skali czasowej • wymienione cechy niekoniecznie dotyczą wszystkich przedstawicieli danego kladu • umiejscowienie pewnych cech na drzewie jest niepewne • charakterystyka rodzin: FGVB, Kubitzki K, red. (1990–) • nazwy rzędów i taksonów wyższej rangi są powiązane z odpowiednimi stronami APweb • Różowe: pochodzenie przedstawicieli grupy jest kontrowersyjne • liczby zapisane szarą czcionką obok nazw rzędów, odnoszą się do liczby rodzin, rodzajów i gatunków
Theodor C. H. Cole
added an update
We want to point to an interesting recent paper by a group around Hong Ma (Fudan Univ., Shanghai) in New Phytologist (early online, March 2017):
Zeng L, Zhang N, Zhang Q, Endress PK, Huang J, Ma H
Resolution of deep eudicot phylogeny and their temporal diversification using nuclear genes from transcriptomic and genomic datasets
doi: 10.1111/nph.14503
The tree based on 161 genes in Fig. 3 presents strong support for i) Dilleniales as sister to Caryophyllales, ii) Santalales basal to, consecutively, Vitales and Saxifragales (followed by the classical Rosids), while iii) the position of Berberidopsidales there is weakly supported.
The according changes will be implemented in the poster if these relationships prove to hold.
 
Theodor C. H. Cole
added a research item
Theodor C. H. Cole
added a research item
Japanese version of the Angiosperm Phylogeny Poster, 2019, by Akitoshi Iwamoto, Kanagawa University, Yokohama. 引用方法: コール TCH, ヒルガー HH, スティーブンス PF, 岩元明敏 (2019) 被子植物の系統ポスター – 被子植物の分類 • オリジナル版: COLE TCH, HILGER HH, STEVENS PF (2019) Angiosperm Phylogeny Poster - Flowering Plant Systematics • 分子データに基づく現段階での系統樹 (2019) ... now following PPA !
Theodor C. H. Cole
added a research item
Indonesian translation of the 2019 version of the Angiosperm Phylogeny Poster. Translation by Sofi Mursidawati, M.Sc., Center for Plant Conservation, Bogor Botanic Gardens, Indonesian Institute of Sciences (LIPI), Bogor, INDONESIA. Dikutip dari: COLE TCH, HILGER HH, STEVENS PF, MURSIDAWATI S (2019) FILOGENI ANGIOSPERMA - Sistematika Tumbuhan Berbunga (versi Bahasa Indonesia Angiosperm Phylogeny Poster - Systematics of Flowering Plants)... now following PPA!
Theodor C. H. Cole
added 2 research items
Portuguese Version of the ”Angiosperm Phylogeny Poster” - citar: COLE TCH, HILGER HH, STEVENS PF, CARVALHO FA (2019) Filogenia das Angiospermas – Sistemática das Plantas com Flores (Portuguese translation of: Angiosperm Phylogeny Poster – Flowering Plant Systematics) - updated to PPA
Theodor C. H. Cole
added 8 research items
Chinese translation by Chen-Kun Jiang of the ”Angiosperm Phylogeny Poster - Flowering Plant Systematics 2019” by Cole TCH, Hilger HH, and Stevens PF - now adapted to PPA (Li HT et al. 2019). Our thanks and appreciation go to Prof. Guang-Yuan Rao, Peking University, 海淀区颐和园路5号, 北京大学, 生命科学学院. This is the ONLY authorized and qualified Chinese translation of the Angiosperm Phylogeny Poster. There are other versions of this poster on the net that we, the authors, have NEITHER endorsed NOR that we have been consulted about.
Cole TCH, Hilger HH, Stevens PF, Doğru-Koca A (2019) Kapalı Tohumluların Filogenisi – Çiçekli Bitkiler Sistematiği, Turkish version of: Cole, Hilger, Stevens (2019) Angiosperm Phylogeny Poster – Flowering Plant Systematics ... now following PPA!
Theodor C. H. Cole
added a project goal
Educational posters giving a clear overview of the hereditary relationships among angiosperms - the evolution of flowering plants - phylogeny with characteristics of the major lineages - regularly updated - now available in 31 languages.