Zhi-Shun Song

• PhD
• Professor (Associate) at Jiangsu Second Normal University, Nanjing, China

Zhi-Shun Song [songzs@ioz.ac.cn]; Ai-Ping Liang * [liangap@ioz.ac.cn] — Abstract The dictyopharid planthopper genus Miasa Distant, 1906 (Orthopagini) is revised to include three known species, M. producta (Lethierry, 1888), M. smaragdilinea (Walker, 1857), and M. wallacei Muir, 1923, and two new species M. borneensis sp.n. and M. nigromacu­ lata sp...

The Austro-Oriental planthopper genus Dictyomorpha Melichar, 1912 (Hemiptera: Fulgoroidea: Dictyopharidae) is revised to contain three previously described and four undescribed species: D. elongata Melichar, 1912 (Papua New Guinea, Indonesia); D. furca sp. nov. (China); D. hectica Haupt, 1926 (Philippines); D. laosensis sp. nov. (Laos); D. moluccan...

The dictyopharid planthopper tribe Aluntiini s.l. is revised and reclassified into two tribes: Aluntiini s.s. and Arjunini Song & Szwedo trib. nov. The tribe Aluntiini s.s. includes five genera: Aluntia Stål, 1866; Dendrophora Melichar, 1903 stat. rev.; Dictyomorpha Melichar, 1912; Indodictyophara Liang & Song, 2012; and Madagascaritia Song & Liang...

A phylogenetic analysis is conducted for all 15 of the Oriental genera of Orthopagini (OGO), and representatives in Arjunini, Dictyopharini, and Hastini, so as to investigate the monophyly and phylogenetic relationships of the OGO. Although the length, thickness, and curvature of the cephalic process vary dramatically in different taxa of Orthopagi...

The Neotropical planthopper genus Trigava O'Brien, 1999 (Hemiptera, Fulgoromorpha, Dictyopharidae, Nersiini) is revised. Four species are included: T. brachycephala (Melichar, 1912) (the type species, from Peru), T. obrieni Song, Malenovský & Deckert, sp. nov. (from Brazil), T. peruensis Song, O'Brien & Bartlett, sp. nov. (from Peru), and T. recurv...

The second genus and species of the planthopper family Katlasidae, Dumpyawnus hpungwanus gen. et sp. nov., is described from mid-Cretaceous Kachin (Burmese) amber. It can be definitely attributed to Katlasidae mainly based on its tegminal structure (e.g., costal area absent, clavus closed, tegmen widened at membrane, multiplied forked of ScP+RA, MP...

A new species of Mimarachnidae, Cretodorus multifoveatus sp. nov., is described from the mid-Cretaceous amber of northern Myanmar. This is the fourth species of the genus Cretodorus which mainly differs from the three previously described species in the number of sensory pits on the head and pronotum, and the CuA of the hind wing with three termina...

Laiyangella zhangi gen. et sp. nov., the first planthopper from the Lower Cretaceous Laiyang formation, Shandong Province of China, is described. Although exhibiting a cixioid venation that would seem to indicate its placement with or near the Cixiidae, the well-preserved impression of the genitalia of this female specimen is not of the orthopteroi...

A new extinct genus and species, Zhangpumazus sheni n. gen. n. sp., of the planthopper family Caliscelidae Amyot and Audinet-Serville, 1843, is described from mid-Miocene Zhangpu amber. This is the second fossil record of this family and the first formally described planthopper species from Zhangpu biota. This new genus and species is apparently cl...

The second genus and species of recently established planthopper family Inoderbidae, Ingensala xiai gen. et sp. nov., is described based on a well-preserved specimen from mid-Cretaceous Kachin (Burmese) amber, and it can be definitely attributed to Inoderbidae mainly based on its head structure, pronotum and mesonotum without median and lateral car...

The second species of Saltissus, as the first mimarachnid planthopper, is herein described from the upper Weald Clay Formation, Smokejacks brickworks, England. Saltissus fennahi Luo, Liu et Jarzembowski sp. nov. can be distinguished from the type species Saltissus eskovi Shcherbakov, 2007 due to the tegmen being more than 20 mm long, stem Pc+CP mer...

A new species, Dorytocus jiaxiaoae Song, Szwedo & Bourgoin sp. nov., from the mid-Cretaceous Burmese amber is described and illustrated. This species represents the first record of the adult of the extinct family Dorytocidae. The supplementary description for the fourth-instar nymph of Dorytocus ornithorhynchus Emeljanov & Shcherbakov is provided....

The tropiduchid genus Catullioides Bierman, 1910 is redescribed and illustrated. Catullioides includes two species, C. rubrolineata Bierman, 1910 (the type species) and C. taishunensis Zhu, Wang & Song, sp. nov. A key to the species of the genus is provided.

In this study, we sequenced and annotated the complete mitochondrial genome of Sogatella kolophon (Kirkaldy) (Hemiptera: Delphacidae). The mitogenome was 16,708 bp in length, containing 13 protein-coding genes (PCGs), 22 tRNA genes, two rRNA genes and a control region. All PCGs started with ATN, except for atp6 and nad5, which used noncanonical cod...

Four species in four genera of Dictyopharidae (Hemiptera: Fulgoromorpha) are recorded from Nepal, including Dictyohimalaya nepalensis Song & Liang, gen. & sp. nov. which is described and illustrated herein, Orthopagus lunulifer Uhler, 1897, Raivuna sp., and Tenguna medogensis Song & Liang, 2007. An identifi cation key to the species of Dictyopha-ri...

A new genus and species, Ayaimatum trilobatum gen. et sp. nov. Jiang and Szwedo, is described based on a planthopper preserved in mid-Cretaceous amber from Kachin State, northern Myanmar, and assigned to the Cretaceous planthopper family Mimarachnidae. A short overview of fossil record of the Mimarachnidae is given. The taxonomic diversity and morp...

A new dictyopharid genus Neonersia Song & Deckert, gen. nov. is described here based on Dictyophora [sic] fugax Melichar, 1912 (previously also placed in the genus Nersia Stål, 1862) from Cameroon. The new genus is placed in the tribe Orthopagini. It may be easily distinguished from all other Orthopagini genera by carinate tegulae.

The genus Chiltana Shakila-Mushtaq & Akbar, 1995 is redescribed and redefined based on the types and new material from Pakistan. Chiltana includes two species, C.acarinatasp. n. and C.baluchi Shakila-Mushtaq & Akbar, 1995 (the type species), both from Chiltan, Balochistan, Pakistan. A key to the species of the genus is provided. Nomenclatorial rema...

The Oriental and eastern Palaearctic planthopper genus Orthopagus Uhler, 1897 (Hemiptera, Fulgoromorpha, Dictyopharidae, Dictyopharinae, Orthopagini) is revised. Six species are included: O.bartletti Song, Malenovský & Deckert, sp. n. (described from India), O.exoletus (Melichar, 1903), comb. n. , stat. rev. (material studied from India and Sri Lan...

The Oriental and eastern Palaearctic planthopper genus Orthopagus Uhler, 1897 (Hemiptera, Fulgoromorpha, Dictyopharidae, Dictyopharinae, Orthopagini) is revised. Six species are included: O.bartletti Song, Malenovský & Deckert, sp. n. (described from India), O.exoletus (Melichar, 1903), comb. n. , stat. rev. (material studied from India and Sri Lan...

The Oriental genus Protolepta Melichar is revised to include two species: P. turbata (the type species) and P. linnavuorii sp. nov., both from Sulawesi, Indonesia. Redescription is provided for P. turbata together with habitus photographs and detailed illustrations of the male genitalia for the first time. A key to the species of the genus and a di...

The Afrotropical planthopper genus Centromeriana Melichar, 1912 (Hemiptera, Fulgoromorpha, Dictyopharidae, Dictyopharinae, Orthopagini) is revised. Four species are included: C. jocosa (Gerstaecker, 1895) (the type species, with confirmed records from Cameroon, Equatorial Guinea and Gabon), C. lindbergae sp. nov. (described from Sierra Leone), C. r...

The Afrotropical planthopper genus Fernandea Melichar, 1912 (Hemiptera: Fulgoromorpha: Dictyopharidae: Dictyopharinae: Orthopagini) is revised to include two species: F. conradti Melichar, 1912 (the type species), with material studied from Cameroon, Equatorial Guinea (Bioko island) and Togo, and F. latifemorata sp. nov., described as new from main...

The dictyopharid planthopper genus Anasta Emeljanov from the Australasian region is revised to include six species including a new one: A. australiaca (Lallemand, 1935) comb. nov. (North Australia), A. lobosa sp. nov. (Papua New Guinea), A. minuta (Lallemand, 1935) comb. nov. (Timor, Indonesia), A. prognatha (Distant, 1906) (North Australia, Papua...

The dictyopharid planthopper genus Anasta Emeljanov from the Australasian region is revised to include six species including a new one: A. australiaca (Lallemand, 1935) comb. nov. (North Australia), A. lobosa sp. nov. (Papua New Guinea), A. minuta (Lallemand, 1935) comb. nov. (Timor, Indonesia), A. prognatha (Distant, 1906) (North Australia, Papua...

The dictyopharid planthopper genus Doryphorina Melichar, 1912 is revised on the base of examination of the type specimens deposited in Museum and Institute of Zoology, Polish Academy of Sciences in Warsaw. The genus includes three known species D. stali Melichar, 1912, D. minor Fennah, 1978, and D. subdeflexa Fennah, 1978. The rank of the two latte...

The Oriental genus Leprota Melichar, 1912, is formally resurrected from synonymy with Saigona Matsumura, 1910, and re-established here as a valid genus. Its type species L. melichari Fennah, 1963, is redescribed and illustrated based on the newly designated lectotype, including a first description of male genitalia. A new Leprota species is describ...

The Oriental planthopper genus Metaurus Stål, 1866 is revised to include the type species Metaurus reticulatus Stål, 1866, a new related species Metaurus ramusitis sp. nov. and an unnamed species. Photographs and descriptions of the species are provided together with structural illustrations of male and female genitalia.

The Microcoryphia fauna of Vietnam remains inadequately studied. Only three species were described before the present study: Pedetontus (Verhoeffilis) bouvieri (Silvestri, 1906), P. (V.) meridionalis Mendes, 1981 (Machilidae), and Machilellus heteropalpus Mendes, 1981 (Meinertellidae). Two new species Machilontus (s. str.) phami, new species, and M...

The monotypic genus Pibrocha Kirkaldy, 1902, known only from Sri Lanka in the Oriental region, is closely related to Dorysarthrus Puton, 1895 from southwestern Asia and northern Africa (Palaearctic region). The genusis revised to include a first description of the male genital structures and a discussion of relationships between Pibrocha, Dorysarth...

A new bristletail species, Machilontus (s. str.) medogensis Song & Huang, sp. nov., is described and illustrated from Medog, Tibet, southwestern China. The new species not only represents the northernmost record of the genus Machilontus Silvestri, 1912, but also represents the first record of the family Meinertellidae Verhoeff, 1910 in China. Check...

There has been confusion regarding the taxonomy and distribution of species in the dictyopharid planthopper genus Putala Melichar, 1903. Based on critical review and examination of parts of type material, Putala is revised and restricted in southern Asia of the Oriental region. Three Putala species are recognized in the genus, including a new speci...

Two new Oriental dictyopharid genera are described from Sri Lanka and southern India, respectively. Truncatomeria gen. nov. is established based on a single known species from Sri Lanka. Its type species, Dictyophora [sic] viridistigma Kirby, 1891 was first described in Dictyophara and then assigned to Centromeria Stål by Distant (1906). However, i...

The dictyopharids planthopper genus Dictyophara Germar, 1833 from China is revised and restricted in Palaearctic Region of northern China. Only four recognized Dictyophara species are: D. europaea (Linnaeus, 1767), D. koreana Matsumura, 1915, D. nekkana Matsumura, 1940 and D. pannonica (Germar, 1830), which the former two are recorded from China fo...

Paralectotypes of three Aphrophora species, A. flavipes Uhler, 1896, A. intermedia Uhler, 1896, and A. major Uhler, 1896 (Hemiptera: Cercopoidea: Aphrophoridae), described by P. R. Uhler (1896) from Japan, recently found in the Natural History Museum in London are documented. The dorsal habitus images, taxonomic notes, host plant and distribution i...

The Oriental dictyopharid planthopper genus Tenguna, Matsumura, 1910 is redescribed, including its type species and a new species: Tenguna medogensis sp. nov. (S.W. China: Xizang). The discovery of the new species broadens our knowledge of the morphology and biogeography of the genus Tenguna and also represents the first record of the planthopper f...

The Oriental dictyopharid planthopper genus Tenguna, Matsumura, 1910 is redescribed, including its type species and a new species: Tenguna medogensis sp. nov. (S.W. China: Xizang). The discovery of the new species broadens our knowledge of the morphology and biogeography of the genus Tenguna and also represents the first record of the planthopper f...

The Oriental and eastern Palaearctic planthopper genus Saigona Matsumura, 1910 is revised to include nine species including five new species: S. capitata (Distant, 1914) (Indo-China, S.W. China: Yunnan), S. fulgoroides (Walker, 1858) (S. China, Sumatra, Borneo), S. fuscoclypeata sp. nov. (C. China: Shaanxi, Hubei and Gansu), S. henanensis sp. nov....

The Oriental and eastern Palaearctic planthopper genus Saigona Matsumura, 1910 is revised to include nine species including five new species: S. capitata (Distant, 1914) (Indo-China, S.W. China: Yunnan), S. fulgoroides (Walker, 1858) (S. China, Sumatra, Borneo), S. fuscoclypeata sp. nov. (C. China: Shaanxi, Hubei and Gansu), S. henanensis sp. nov....

A new dictyopharid planthopper species, Sphenocratus xinjiangensis Liang, sp. nov. (subfamily Orgeriinae), is described and illustrated from Xinjiang in northwestern China. This represents the first authentic record of the genus Sphenocratus Horvath, 1910 as well as of the subfamily Orgeriinae in China. The new species extends the range of the genu...

A new dictyopharid planthopper species, Sphenocratus xinjiangensis Liang, sp. nov. (subfamily Orgeriinae), is described and illustrated from Xinjiang in northwestern China. This represents the first authentic record of the genus Sphenocratus Horváth, 1910 as well as of the subfamily Orgeriinae in China. The new species extends the range of the genu...

The genus Dictyopharina Melichar, 1903, and its type species, D. viridissima Melichar, 1903 from Sri Lanka and India, are redescribed and illustrated. Two new species, D. octaprotrusa sp. nov. and D. sichuanensis sp. nov., are added to the genus from southern China, and they represent the first record of the genus from China. A key to the species o...

A new species, Allopsontus (Anisopsontus) huashanmendesi, sp. nov. is described from Mt. Huashan, Shaanxi Province, China. A key to the species of Allopsontus (Anisopsontus) is provided.