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September 2018 - December 2018
Publications
Publications (48)
Selaginella subg. Heterostachys, especially S. sect. Heterostachys and S. sect. Tetragonostachyae, represents the most taxonomically difficult subgenus of Selaginella in China. Here, Selaginella austro-orientalis, a new species in S. sect. Heterostachys from Southeast China is described and illustrated based on morphological and phylogenetic studie...
A new spikemoss species, Selaginella wuyishanensis , is described and illustrated based on materials collected from Fujian Province, East China. The new species can be distinguished from S. lutchuensis Koidzumi and S. albociliata P. S. Wang by its leaves with extremely long cilia (up to 8 mm) and distinctly white margins, ovate ventral sporophylls,...
A new fern species, Athyrium aberrans (Athyriaceae), currently only known from southeastern Xizang, China is described. Morphologically, A. aberrans is superficially similar to A. falcatum in the habit and pinna shape, but the new species can easily be distinguished from the latter by having purplish petioles and rachises, a bulbil on the rachis be...
Lycopodiales are one of the oldest lineages of any living vascular plants and contain about 400 species distributed on all continents except Antarctica but with its highest diversity in tropical regions. Following a recent molecular study, here we classify
Lycopodiales into two families: Huperziaceae and Lycopodiaceae, and Lycopodiaceae are further...
As one of the earliest land plant lineages, Selaginella is important for studying land plant evolution. It is the largest genus of lycophytes containing 700–800 species. Some unique characters of Selaginella plastomes have been reported, but based only on 20 species. There have been no plastome phylogenies of Selaginella based on a relatively large...
The Old World fern genera Hypodematium and Leucostegia had long been placed in the families Dryopteridaceae and Davalliaceae, respectively, before the advent of molecular phylogenetics. Recent molecular studies confirmed the recognition of the family Hypodematiaceae composed of these two genera, but the relationships within each of these two genera...
Lycopodiaceae are one of the oldest lineages of any living vascular plants and contain about 400 species distributed on all continents except Antarctica, with its highest diversity in tropical regions. Previous studies on the morphology, anatomy, and molecular systematics of Lycopodiaceae have made substantial progress in understanding the diversit...
Based on morphological and phylogenetic studies, Selaginella qingchengshanensis, a new species in S. sect. Heterostachys from Sichuan, China is described and illustrated. Morphologically, the new species is similar to S. amblyphylla, but has sterile leaves with denticulate to slightly ciliate margins, dorsal leaves with long arista (as long as leav...
We describe a new species of Asplenium (Aspleniaceae) from Vietnam as A. quangbinhense. Morphologically, it is most similar to A. oldhamii. The new species is closely related to A. pseudopraemorsum, and a morphological comparison is presented between the two species. Three species, A. cyrtosorum, A. maguanense, and A. pseudopraemorsum, are reported...
Cheilanthes qiaojiaensis, a new fern of Pteridaceae subfamily Cheilanthoideae, is described and illustrated. Morphologically, C. qiaojiaensis is similar to C. chusana and Aleuritopteris rufa in having plants hairiness and laminae elliptic-lanceolate, but the new species has plants covered with dense hairs throughout, petiole and rachis flattened to...
The complete chloroplast genome of Aristolochia delavayi was determined in this study. The chloroplast genome consists of 160,344 bp, with a typical circular structure including a pair of inverted repeats of 25,454 bp separated by a large single-copy region and a small single-copy region of 89,502 and 19,795 bp, respectively. The plastome contains...
A new species, Selaginella coriaceifolia, is described and illustrated from central Vietnam. The new species can be distinguished from other species in Asia by its habit epilithic, main stem nearly isotomously branched, leaves strongly leathery, ventral leaves imbricate on stem and branch, dorsal leaves with reflexed arista at apex, and sporophylls...
Lepisoroid ferns (tribe Lepisoreae, Polypodiaceae) are arguably one of the most confusing fern groups in Polypodiaceae in terms of delimitation of genera largely because of their simple morphology. Previous molecular studies either had very small taxon sampling of the non-Lepisorus genera and did not well resolve the relationships among these gener...
The taxonomy of the cliff fern family Woodsiaceae has been controversial with 22–59 species in 1–7 genera recognized in various classifications. This is mainly due to limited taxon sampling (up to 33 accessions representing up to 21 species) and poor resolution in molecular studies so far. In the present study, DNA sequences of five plastid markers...
As an ancient lineage of ferns, Ophioglossaceae are evolutionarily among the most fascinating because they have the highest chromosome count of any known organism as well as the presence of sporophores, subterranean gametophytes, eusporangiate sporangia without annuli, and endophytic fungi. Previous studies have produced conflicting results, identi...
Rubus is one of the most species-rich and most taxonomically challenging genera in Rosaceae. Pollen morphology of Rubus species has been described by many researchers but never in a comprehensive study. In this study, pollen morphology of 155 species and 13 varieties representing all 12 subgenera of Rubus was examined using scanning electron micros...
The newly defined fern genus Leptochilus contains about 50 species occurring in subtropical to tropical Asia and adjacent Pacific islands. The circumscription and phylogeny of the genus have been ambiguous and its species had been included in various genera such as Colysis, Dendroglossa, Kontumia, Microsorum, and Paraleptochilus. Previous molecular...
Arachniodes (Dryopteridaceae) is one of the most confusing and controversial fern genera in terms of its circumscription, nomenclature, and taxonomy. Estimates of species number range from 40 to 200. Previous molecular works included only 2–17 accessions representing 2–12 species of Arachniodes and allied genera, leaving most of the Asian species r...
Three new species of the fern genus Arachniodes (Dryopteridaceae) are described from Vietnam: A. daklakensis, A. longicaudata, and A. quangnamensis. The first two occur in Central Highlands and Central Vietnam, respectively, while the last is found in the north (Bavi, Langson and Tamdao) as well as in the central Vietnam (Nghean, Quangbinh, Quangtr...
Nine new species of Hymenasplenium (Aspleniaceae) from Asia are here described based on both morphology and our recent molecular phylogenetics of the genus. Of the nine new species, H. chingii, H. denticulatum, H. sinense, H. speluncicola, and H. wangpeishanii are from southern China, whereas H. distans, H. ngheanense, H. phamhoanghoi, and H. quang...
A new fern species, Hymenasplenium perriei (Aspleniaceae), from very shaded and wet rocks in southeastern Fiji, is described and illustrated based on our morphological and recent phylogenetic studies. The new species is distinguished by its phyllopodia 2–3(–10) mm apart, marginal teeth irregular and usually with notches, veins usually terminating i...
The fern genus Hymenasplenium (Aspleniaceae) is one of the two genera in the family. It is generally recognized among modern pteridologists. However, its infrageneric relationships and species diversity have been unclear and controversial. The molecular studies so far have had small taxon and character sampling. In the present study, DNA sequences...
We performed phylogenetic analyses to resolve the relationships of Pteridrys and related taxa based on six plastid markers (atpA, atpB, matK & rps16-matK, rbcL, rps4 & rps4-trnS, and trnL & trnL-F) and nuclear pgiC. We included 195 accessions representing approximately 147 species in 38 genera, and seven of the nine families in Polypodiineae (eupol...
A new fern species, Hymenasplenium hastifolium (Aspleniaceae), from a limestone cave in western Guangxi, China, is described and illustrated. Morphological affinities of genus Hymenasplenium are also compared and discussed. The new species is distinguished by the shape of lamina, the venation, and the sorus distribution. Hymenasplenium hastifolium...
The Old World fern genus Pyrrosia is one of the largest genera in Polypodiaceae. It is estimated to contain up to 100 species distributed in tropical and subtropical South and East Asia, from New Zealand to Siberia and from Africa to Polynesia. The monophyly of Pyrrosia has frequently been questioned, previous molecular analyses have produced confl...
The fern genus Pteridrys is characterized by having free veins and a tooth on each sinus between two pinna or pinnule lobes. This genus is currently known from nine to ten species. Here we add three new species of Pteridrys from northern Vietnam found in an expedition in 2013.
The Old World fern genus Pyrrosia (Polypodiaceae) offers a rare system in ferns to study morphological evolution because almost all species of this genus are well studied for their morphology, anatomy, and spore features, and various hypotheses have been proposed in terms of the phylogeny and evolution in this genus. However, the molecular phylogen...
A new combination, Pyrrosia annamensis (Christ) Li Bing Zhang, X.M.Zhou & C.W.Chen, for a Southeast Asian species formerly treated in Niphobolus is made. We also lectotypify Cyclophorus rhomboidalis Bonaparte.
One earlier known species of Lycopodium from China, “L. neopungens H.S. Kung & Li Bing Zhang”, is described as Spinulum lioui Li Bing Zhang & H.He (sp. nov.).
Tectaria (Tectariaceae) is one of the most confusing fern genus in terms of its circumscription and phylogeny. Since its original description, a number of genera had been moved into or related with this genus, while others had been segregated from it. Tectaria is also among the largest fern genera, comprising 150–210 mostly tropical species. Previo...
A new name of the lycophyte genus Huperzia (Huperziaceae), H. wusugongii Li Bing Zhang, X.G. Xu & X.M. Zhou, from west China is proposed to replace the later homonym, H. rubricaulis S.K. Wu & X. Cheng [‘rubicaulis’], of H. rubricaulis (Alderwerelt van Rosenburgh) Holub.
A new combination in the lycophyte genus Dendrolycopodium (Lycopodiacae) from Lycopodium, D. verticale (Li Bing Zhang) Li Bing Zhang & X.M.Zhou, for a Chinese species is made.
As the second most genera-rich fern family, Pteridaceae contain more than 1000 species contributing to ca. 10% of extant leptosporangiate fern diversity. The subfamily Pteridoideae is one of the five subfamilies often recognized. The circumscription of Pteridoideae has not been clear. A large number of species have not yet been included in any mole...
Phylogeny has long informed pteridophyte classification. As our ability to infer evolutionary trees has improved, classifications aimed at recognizing natural groups have become increasingly predictive and stable. Here, we provide a modern, comprehensive classification for lycophytes and ferns, down to the genus level, utilizing a community-based a...
The circumscription and the phylogeny of the fern family Tectariaceae have been controversial. Previous molecular studies have supported the monophyly of this family, with 4-5 genera. However, these studies were exclusively based on plastid markers and relatively small sampling, especially of the non-Tectaria genera. In the present study, DNA seque...
Background:
Polystichum (Dryopteridaceae) is probably the third largest fern genus in the world and contains ca. 500 species. Species of Polystichum occur on all continents except Antarctica, but its highest diversity is found in East Asia, especially Southwest China and adjacent regions. Previous studies typically had sparse taxon sampling and us...
The largest genus of seed-free vascular plants Selaginella alone constitutes the family Selaginellaceae, the largest of the lycophyte families. The genus is estimated to contain ca. 800 species distributed on all continents except Antarctica, with the highest species diversity in tropical and subtropical regions. The monophyly of Selaginella has ra...
Using light microscope (LM) and scanning electron microscope (SEM), the megaspores and microspores of 77 samples representing ca. 70 species of Selaginella from China are observed. Combing previous studies, the spore morphology of nearly all documented Selaginella species from China were reviewed. Based on the morphological characteristics in megas...
A new lycophyte species, Selaginella chuweimingii, from central Yunnan, China is described and illustrated. Selaginella chuweimingii is similar to S. sinensis in the habit and the morphology of strobili and sporophylls, but the new species can easily be distinguished from the latter by having ovate or oblong-ovate ventral leaves, extremely asymmetr...
The lycophyte genus Selaginella alone constitutes the family Selaginellaceae, the largest of the lycophyte families. The genus is estimated to contain 700–800 species distributed on all continents except Antarctica, with highest species diversity in tropical and subtropical regions. The monophyly of Selaginella in this broad sense has rarely been d...
A global plastid phylogeny of the brake fern genus Pteris (Pteridaceae) and related genera in the Pteridoideae Abstract The brake fern genus Pteris belongs to the Pteridaceae subfamily Pteridoideae. It contains 200–250 species distributed on all continents except Antarctica, with its highest species diversity in tropical and subtropical regions. Th...
Pellaea barklyae, a fern endemic to the Seychelles, is transferred to the genus Pteris based on recent molecular studies. We also neotypified Adiantum pallens Swartz, a species endemic to Madagascar, Mauritius, and La Réunion.
The brake fern genus Pteris belongs to the Pteridaceae subfamily Pteridoideae. It contains 200–250 species distributed on all continents except Antarctica, with its highest species diversity in tropical and subtropical regions. The monophyly of Pteris has long been in question because of its great morphological diversity and because of the controve...
Acronema crassifolium, a distinct new species endemic to Yunnan province, China, is described and illustrated. It is characterized by its thickly papery, ternate, abaxially dark purple leaves, terminal umbels with 8-13 rays, and absent calyx teeth. The pollen morphology of the new species is also described in this study.
Projects
Projects (2)
Systematics and Evolution Study of Vascular Plants: Phylogeny, Cassification, Revision
To some extent, phylogeny has always informed the classification of pteridophytes. Today, as our ability to infer evolutionary trees continues to improve, our classifications aimed at recognizing natural groups are becoming increasingly predictive and stable.
We now feel the time is right to update the classification of ferns and lycophytes, following the main tenets of the Smith et al.2006 and related work, but utilizing a community-based approach. We have therefore established the Pteridophyte Phylogeny Group (PPG), following loosely the model established by those working on flowering plants.
see http://botany.si.edu/PPG/
