Xabier Pereda Suberbiola

University of the Basque Country | UPV/EHU · Geology

Late Cretaceous marine ecosystems saw a major adaptive radiation of mosasaurids, which evolved highly disparate jaw and tooth morphologies to feed on different prey. A striking pattern seen in mosasaurids was a high diversity of durophagous forms. Durophagy likely evolved several times independently in the mosasaurine genera Globidens, Prognathodon...

Skull remains of an ornithopod dinosaur, identified as MDS-LTASI, from the Los Terreros-Altollano site in Salas de los Infantes (Burgos, Spain) are described. The site is located in the upper Barremian-Aptian Castrillo de la Reina Formation (Cameros Basin). The fossils are part of a single individual and consist of a predentary, a dentary and a set...

In recent years the Middle Atlas of Morocco has become an area of interest for the study of dinosaurs in northern Africa. The Boulahfa locality, near Boulemane, has produced a diverse dinosaur assemblage from the Middle Jurassic of the El Mers Group. Fossil remains of sauropods and thyreophorans, such as ankylosaurs (Spicomellus) and stegosaurs (Ad...

A new member of Spinosauridae from the Enciso Group (uppermost Barremian–lower Aptian) from Igea (La Rioja, Spain) is here erected on the basis of axial, pelvic girdle, and hindlimb elements that exhibit a unique combination of characters. Riojavenatrix lacustris gen. et sp. nov. is one of the latest Iberian and European spinosaurid taxa. It retain...

In the Late Cretaceous, northern and southern hemispheres evolved distinct dinosaurian faunas. Titanosaurians and abelisaurids dominated the Gondwanan continents; hadrosaurids, ceratopsians and tyrannosaurs dominated North America and Asia. Recently, a lambeosaurine hadrosaurid, Ajnabia odysseus, was reported from the late Maastrichtian phosphates...

Un nouveau spécimen de dinosaure thyréophore a été découvert dans les couches rouges du Bathonien-Callovien (Formation d’El Mers III) dans la bordure sud-occidentale du synclinal d’El Mers au sud de Boulemane (Moyen Atlas plissé, Maroc). Le Moyen Atlas est connu depuis longtemps par sa richesse en restes d’os et en traces de pas de dinosaures et co...

The end of the Cretaceous saw the evolution of endemic dinosaur faunas on different landmasses, driven by continental fragmentation. Understanding the evolution of these biogeographic patterns is important for understanding the evolution of Mesozoic ecosystems. However, the faunas of the southern land masses remain understudied relative to the inte...

New theropod tracks found in the Papo Seco Formation (lower Barremian, Lusitanian Basin) are presented. In 2022, thirteen theropod tracks were identified on the lowermost bed of this formation, preserved as natural cast infillings on the bedding surface. The tracks are attributed to the ichnogenus cf. Megalosauripus isp. It is suggested that they w...

New theropod tracks found in the Papo Seco Formation (lower Barremian, Lusitanian Basin) are presented. In 2022, thirteen theropod tracks were identified on the lowermost bed of this formation, preserved as natural cast infillings on the bedding surface. The tracks are attributed to the ichnogenus cf. Megalosauripus isp. It is suggested that they w...

Among the many valuable uses of the artificial intelligence in the field of museums, it may assist the classification of out-of-context items. This paper deals with the problem of the identification of dinosaur teeth (a type of fossil that is usually found scattered), which can be tackled by means of multivariate algorithms (such as the principal c...

Among the many valuable uses of the artificial intelligence in the field of museums, it may assist the classification of out-of-context items. This paper deals with the problem of the identification of dinosaur teeth (a type of fossil that is usually found scattered), which can be tackled by means of multivariate algorithms (such as the principal c...

In the last decades, the Middle Atlas region has become an important area for the study of Moroccan’s Mesozoic vertebrates, especially the Boulemane-El Mers region, rich in diverse dinosaur bones and tracks. The first dinosaur skeletal fossils have been found in Middle Jurassic (El Mers I Formation) sites of this area (Termier et al., 1940; Lappare...

Mosasaurids, a clade of specialized marine squamates, saw a major adaptive radiation in the Late Cretaceous, evolving a wide range of body sizes, shapes, and specialized tooth morphologies. The most diverse known mosasaurid faunas come from the late Maastrichtian phosphates of Morocco. Here, we report an unusual new mosasaurid, Stelladens mysterios...

Equoid perissodactyls were diverse in the Eocene of the Iberian Peninsula, and those from the central and western Iberian basins (Mondego, Oviedo, Almazán, Duero, and Miranda-Treviño) and the eastern Ebro Basin (Ulldemolins area) were endemic, with the presence of six genera and 19 species (mainly Palaeotheriidae sensu stricto) that are unknown in...

Early Cretaceous ornithopod fossil bones from Portugal are reported only from the Papo-Seco Formation (lower Barremian) at Cabo Espichel, south of Lisbon. Ornithopod and other vertebrate remains from the Papo-Seco Formation occur in limestones, marls, sandstones and conglomerates in a succession interpreted as deposited in lagoonal and estuarine en...

Early Cretaceous ornithopod fossil bones from Portugal are reported only from the Papo-Seco Formation (lower Barremian) at Cabo Espichel, south of Lisbon. Ornithopod and other vertebrate remains from the Papo-Seco Formation occur in limestones, marls, sandstones and conglomerates in a succession interpreted as deposited in lagoonal and estuarine en...

The most representative ankylosaurian remains from Argentina have been found in sediments of the Allen Formation (Campanian–Maastrichtian) in Salitral Moreno, Río Negro Province. Several authors have discussed the identity and history of these remains. In this study, we review all published material along with some new remains in order to summarize...

Ornithopod remains are reported from the Papo-Seco Formation (lower Barremian, Early Cretaceous) at Cape Espichel, south of Lisbon, in western central Portugal. The fossil material comes from the Boca do Chapim, Praia do Areia do Mastro and Praia do Guincho palaeontological sites. Some fossils are known from the 19th century and the beginning of th...

We described a dinosaur tracksite found in the uppermost part of the Areia do Mastro Formation (lowermost Barremian, Lower Cretaceous), located at 1.5 km north of Cabo Espichel (Sesimbra, Portugal). The studied tracks are distributed in a heavily trampled limestone bed which crops out alongside the rocky beach. The studied trampled surface is highl...

The early evolution of thyreophoran dinosaurs is thought to have occurred primarily in northern continents since most evidence comes from the Lower and Middle Jurassic of Europe and North America. The diversification into stegosaurs and ankylosaurs is obscured by a patchy fossil record comprising only a handful of fragmentary fossils, most with unc...

Entre los materiales de derrubio extraídos de las areniscas de la segunda secuencia que forman los materiales de las facies Weald (Cretácico inferior de Aldea de Losilla, Valencia), se encontró una espina caudal de relativa longitud atribuible a un dinosaurio estegosaurio. Se trata del primer hallazgo en España de un resto de estegosaurio y una de...

A B S T R A C T The Cretaceous-Paleogene (K-Pg) transition saw mass extinctions in terrestrial and marine ecosystems. Terrestrial vertebrate diversity patterns across the K-Pg boundary have seen extensive study, but less is known about marine vertebrates. We describe a new mosasaurid from the latest Maastrichtian phosphatic beds of Morocco, showing...

Albian dinosaur tracks from the Monte Grande Formation (Basque-Cantabrian Basin) are described. Sedimentary succession shows seven different facies associations (FA) with four track-bearing levels (levels 1-4) of a braid delta system. The FA4, a proximal crevasse subdelta, preserves in levels 2 and 3 theropod, possibly sauropod and undetermined din...

Thyreophora is a diverse clade of herbivorous ornithischian dinosaurs composed of stegosaurs, ankylosaurs, and basal forms. Thyreophorans have an extensive fossil record, spanning from the Lower Jurassic to the Upper Cretaceous, which is best known from well-preserved specimens in Laurasia. In contrast, the remains are extremely rare and mostly par...

Spinosaurid dinosaurs show a wide geographical and temporal distribution, being present in Gondwanan and Laurasian landmasses, at least, during the Lower Cretaceous and the Cenomanian. European spinosaurids are more diverse than previously thought with five, and probably six, currently recognised genera. The revision of CPI 477, a fragment of a lef...

The family Palaeotheriidae (Perissodactyla, Equoidea) was one of the most abundant and successful Eocene mammalian groups from western Europe. Equoid perissodactyls were diverse in the Iberian Peninsula, with the presence of six genera and 19 species (mainly palaeotheres) which are unknown elsewhere in Europe. In this work, an overview of the Eocen...

Dinosauro teropodoak erregistro fosilean nahiko urriak dira eta askotan aurkitutako fosil bakarrak hortz isolatuak dira. Teropodoen hortzekin, ordea, azterketa paleontologiko anitz egin daitezke, metodologia kualitatibo eta kuantitatibo mota desberdinak erabiliz. Hortzetatik abiatuz, teropodoen biosistematika burutu eta talde honen erregistro fosil...

Two new ornithopod natural casts are reported from the Praia do Areia do Mastro site, at Espichel Cape (near Sesimbra), western central mainland Portugal (western Iberia). In this Tracksite two geological formations occur: Areia do Mastro Formation and Papo-Seco Formation (lower Barremian). This locality (Praia do Areia do Mastro) occurs in a sedim...

The climatic cooling that began in the late middle Eocene and culminated in the Eocene-Oligocene transition meant major changes in Palaeotheriidae (Perissodactyla, Mammalia) biodiversity in Europe and could have caused the appearance of new dietary strategies. This work is the first to study the spatiotemporal response of one palaeotheriid genus (P...

Se describe un fémur y se hace una síntesis del material de estegosaurios hallado hasta el momento en los alrededores de Aras de Alpuente (Valencia), en las localidades de Losilla y Cerrito del Olmo. Se indica la situación estratigráfica de las mencionadas localidades y se establece su relación con los restantes yacimientos de la zona que han propo...

Se analizan las faunas de reptiles marinos del Cretácico final (Santoniense a Maastrichtiense) de Europa, en el margen norte del mar de Tetis mediterráneo, desde un punto de vista sistemático y bioestratigráfico. Se establece una correlación entre las principales formaciones y las zonas de belemnites. Las faunas de reptiles marinos de la provincia...

A total of 227 theropod teeth have so far been recovered from the upper Campanian Laño site (northern Iberian Peninsula). The teeth were studied for their qualitative and quantitative features. From the theropod sample found at Laño, seven morphotypes attributed to five taxa are identified: a medium to large abelisaurid (Arcovenator sp.) and four s...

In this work seventeen new maxillary and mandibular dental pieces of the endemic palaeotheriids Leptolophus sp., Iberolophus jimenezi, Plagiolophus mazateronensis, Palaeotherium giganteum, Franzenium durense and Cantabrotherium casanovasae from the late middle Eocene (Bartonian, MP 16) of Mazaterón (Soria, Almazán Basin) are described. The palaeoth...

Theropod behaviour and biodynamics are intriguing questions that paleontology has been trying to resolve for a long time. The lack of extant groups with similar bipedalism has made it hard to answer some of the questions on the matter, yet theoretical biomechanical models have shed some light on the question of how fast theropods could run and what...

At least 700 crocodylomorph trace fossils were discovered in 2021 at Ribeira do Chapim tracksite (Cabo Espichel, Sesimbra, SW Portugal) in one limestone bed on top of the Areia do Mastro Formation (lower Barremian), deposited in a carbonate shallow lagoon environment, under a tropical climate. The footprints have several sizes. The arrangement of s...

Ornithopod remains are reported from the Papo-Seco Formation, at Cabo Espichel area (Sesimbra County), in western central Portugal, south of Lisbon. The material was collected in three palaeontological sites: Boca do Chapim, Areia do Mastro and Praia do Guincho. We present new remains and the review of ornithopod already described from the Papo-Sec...

Ornithopod remains are reported from the Papo-Seco Formation, at Cabo Espichel area (Sesimbra County), in western central Portugal, south of Lisbon. The material was collected in three palaeontological sites: Boca do Chapim, Areia do Mastro and Praia do Guincho. Vertebrate fossils from Cabo Espichel are known since the 19th century. Dinosaur and cr...

The revision and recent discoveries of new theropod material in the Iberian Peninsula shows that spinosaurids were more common and diverse than previously expected with the description of Vallibonavenatrix, the identification of Baryonyx and the probable spinosaurid Camarillasarus. Findings of spinosaurid remains in the Enciso Group (upper Barremia...

Se describen por primera vez restos fósiles de cocodrilo del Mioceno Inferior (Formación Tudela, Rambliense, Biozonas MN2b-3) de las Bardenas Reales de Navarra, región situada en el sector noroccidental de la Depresión del Ebro. El material consiste en fragmentos craneales y mandibulares, dientes, huesos del esqueleto axial y apendicular y del escu...

Theropod behaviour and biodynamics are intriguing questions that paleontology has been trying to resolve for a long time. The lack of extant groups with similar bipedalism has made it hard to answer some of the questions on the matter, yet theoretical biomechanical models have shed some light on the question of how fast theropods could run and what...

During the Eocene, in the European archipelago, the most abundant and diverse components of the perissodactyl fauna were the palaeotheres (Family Palaeotheriidae, Equoidea, Mammalia). In western Iberian Eocene fossil record (Duero, Almazán, Oviedo and Miranda-Treviño basins), a wide diverse and endemic association of palaeotherids and lophiodontids...

The middle and late Eocene perissodactyl fossil record of western Iberia is characterized by an endemic fauna, which is clearly different from those of northeastern Iberia and other areas of the European archipelago. In this work, we describe three endemic equoid taxa belonging to the family Palaeotheriidae. Two come from the late Eocene site of Za...

El hallazgo más antiguo conocido de un dinosaurio en España es un supuesto diente de terópodo del Jurásico Superior de Asturias. Fue descrito en 1858 como un diente de tiburón por el ingeniero de minas alemán Guillermo Schulz y asignado en 1873 al dinosaurio Megalosaurus por el geólogo navarro Justo Egozcue. Se desconoce el paradero actual de este...

Since the second half of the 20th Century, the study of fossil tracks in South America has steadily increased. A large number of tetrapod ichnogenera is currently known mainly from Argentina, Bolivia and Brazil. In this study, we present a new record consisting of several trackways that we refer to cf. Tetrapodosaurus, representing the first explic...

The Quintanilla la Ojada section (Basque-Cantabrian Region, northern Spain) has yielded two assemblages of Late Cretaceous vertebrates, deposited during the Maastrichtian in coastal environments and related to a transgressive lag at the base of the Valdenoceda Formation. Numerous teeth of Elasmobranchii and Actinopterygii are the most prevailing fo...

Lorente, F. 2007. Icnitas de posibles dinosaurios tireó-foros del Jurásico Inferior (Alto Atlas, Goulmina, Marruecos). �Possible thyreophoran dinosaur tracks from the �Possible thyreophoran dinosaur tracks from the Possible thyreophoran dinosaur tracks from the Lower Jurassic (High-Atlas, Goulmima. Morocco).�� .�� Revista Española de Paleontología,...

New ornithopod remains are reported from the Papo-Seco Formation (Lower Barremian, Lower Cretaceous) at the Espichel Cape, western central Portugal, south of Lisboa. Marine, lagoonal and estuarine sedimentary rocks, consisting in limestones, marls, sandstones and conglomerates, have yielded fossil remains of dinosaurs and other vertebrates since th...

Remains of Garrigatitan meridionalis nov. gen. et sp. were found in two bonebeds of sequence 2 from the upper Campanian site of Velaux-La Bastide Neuve (Aix-en-Provence Basin, Bouches du Rhône department). The vertebrate assemblage is dominated by dinosaurs, including the titanosaur Atsinganosaurus velauciensis. Garrigatitan meridionalis presents t...

The Late Cretaceous saw distinctly endemic dinosaur faunas evolve in the northern and southern hemispheres. The Laurasian continents of North America and Asia were dominated by hadrosaurid and ceratopsian ornithischians, with tyrannosaurs as apex predators. In Gondwanan communities, including Africa, South America, India and Madagascar, titanosauri...

A track-bearing slab (MNHN. F. AC10007) from the Stolling Formation of Hessberg (Hildburghausen, Thuringia, central Germany) is housed in the Muséum national d'Histoire naturalle of Paris, France, Since 1835.

Twelve isolated spinosaurid teeth from the Enciso Group of the eastern Cameros Basin (La Rioja, Spain) have been studied. The fossil material has been found in five sites near the town of Igea, where a fluvio-lacustrine system with marine influence was developed during the Barremian-early Aptian. Besides the study of the qualitative and quantitativ...

The study of palaeopathology provides valuable information about injury and behaviour in extinct organisms. Appendicular pathologies are interesting as they directly affect mobility and therefore the ability of an animal to survive. Here, the injuries recorded in the left pes of the neornithischian Othnielosaurus consors are described. The implicat...

Despite continuous improvements, our knowledge of the palaeoneurology of sauropod dinosaurs is still deficient. This holds true even for Titanosauria, which is a particularly speciose clade of sauropods with representatives known from numerous Cretaceous sites in many countries on all continents. The data currently available regarding the palaeoneu...

A palynological study of the Upper Cretaceous vertebrate site of Chera (Valencia, Spain) has been carried out. The occurrence of diversified Normapolles including various species of Vancampopollenites, Papillopollis and Pseudoromeinipollenites suggest a late Campanian–?early Maachtrichtian age. The palynological assemblages suggest an azonal vegeta...

The western Pyrenees is a territory with a remarkable geodiversity and an important fossil record, which constitutes a large archive of palaeobiodiversity. With respect to the vertebrate fossil record, the western Pyrenees (Basque-Cantabrian Region, Pamplona Basin and the north-western sector of the Ebro Basin) are relevant both from the point of v...

Despite continuous improvements, our knowledge of the palaeoneurology of sauropod dinosaurs is still deficient. This holds true even for Titanosauria, which is a particularly speciose clade of sauropods with representatives known from numerous Cretaceous sites in many countries on all continents. The data currently available regarding the palaeoneu...

The Upper Cretaceous site of Velaux-La Bastide Neuve (Bouches-du-Rhône Department, France) is well-known for its dinosaur specimens, more specifically the titanosaurian remains. The taxon Atsinganosaurus velauciensis is represented by several partially articulated skeletons found in this upper Campanian site; some of this material was described by...

Titanosaurs were the last-surviving group of sauropod dinosaurs, having persisted until the end-Cretaceous. Since the early 21st century, knowledge of this diverse and cosmopolitan group of dinosaurs has greatly improved, although many aspects of their biology and evolutionary history remain poorly understood. In particular, knowledge of Late Creta...

New ornithopod remains are reported from the Papo-Seco Formation (Lower Barremian, Lower Cretaceous) at the Espichel Cape, situated about 40 km south of Lisbon, western central Portugal. Marine, lagoonal and estuarine sedimentary records, consisting in limestones, marls, sands and conglomerates, have yielded fossil remains of dinosaurs and other ve...

We report new ornithopod remains from the Papo-Seco Formation at the Espichel Cape (40 km south of Lisbon, western central Portugal). The Papo-Seco Formation, stratigraphically located between the Areias do Mastro Formation and the Boca do Chapim Formation and assigned to the Lower Barremian (Lower Cretaceous). This study is based on ornithopod bon...