Wouter Holleman

• National Research Foundation, South Africa

Estuaries, at the confluence of marine and freshwater systems, are mostly of geologically recent origin and as such make excellent models for understanding recent speciation events. Using molecular approaches, we compared genetic diversity and demographic histories in 2 closely related southern African klipfish species, the marine Clinus supercilio...

Caesio xanthalytos, a new species of fusilier, closely related to Caesio caerulaurea Lacepède 1801, is described from several specimens from various localities on the east coast of Africa, and from southern Madagascar. While very similar in colour pattern to C. caerulaurea, its lateral line lies within a dominant, composite yellow stripe, whereas i...

The Clinus superciliosus complex comprises six closely related species. Three of the species were originally recognized by Gilchrist and Thompson as two distinct species, Clinus superciliosus (Linnaeus, 1758) and Clinus ornatus Gilchrist & Thompson, 1908, and a variety, Clinus superciliosus var. arborescens Gilchrist & Thompson, 1908. A number of o...

The fishes of the blennioid family Tripterygiidae of the Red Sea are reviewed. Twelve species are recognised: Norfolkia brachylepis, Helcogramma obtusirostris, H. steinitzi and nine species of Enneapterygius. Enneapterygius qirmiz sp. nov. is described for a species originally described, but not named, by Clark (1980), of which the specimens on whi...

Fifteen species of the tripterygiid fish genus Helcogramma are recognised from the Western Indian Ocean (including Sri Lanka and the southeastern coast of India). Helcogramma shinglensis Lal Mohan is recognised as a valid species and four species are described as new: Helcogramm aalkamr , which is similar to H. chica Rosenblatt, is known from the C...

Three species of Helcogramma from the Indian Ocean that form a monophyletic group are recognised: Helcogramma steinitzi Clark (Red Sea and Gulf of Oman), H. rosea n. sp. (Andaman Sea and Sri Lanka) and H. microstigma n. sp. (East African coast and adjacent islands). Helcogramma microstigma is distinguished from the other two species by a head profi...

The fishes of the tripterygiid genus Enneapterygius of the western Indian Ocean (excluding Sri Lanka) are reviewed. Four new species, Enneapterygius elaine , Enneapterygius gruschkai , Enneapterygius genamaculatus and Enneapterygius kosiensis are described and several species are redescribed. Enneapterygius elaine is known only from Rodrigues, E. g...

Ucla xenogrammus, a monotypic genus of tripterygiid fishes, with is widespread in the Indo-West Pacific, is formally described, the unpublished names given by Richard Rosenblatt in 1959 being retained. Ucla xenogrammus is distinguisehd from all other tripteryggid fishes by its protruding lower jaw, depressible acicular teeth and grooved scales in t...

The genus Norfolkia is revised and three species are recognised: N. brachylepis (Schultz) (= N.springeri Clarke) which is widespread throughout the tropical Indo-west Pacific, N. squamiceps (McCulloch & Waite) from the southern Barrier Reef, Norfolk and Lord Howe Islands and N. thomasi Whitley from the Barrier reef and south-central Pacific.

Ceratobregma helenae is described as a new genus and species of fish of the family Tripterygiidae occuring in the eastern Indian, western and central Pacific oceans. The Australian species Vauclusella acanthops Whitley, 1965 is redescribed and assigned to Ceratobregma.

A new species of Helcogramma, H. fuscopinna, which ranges across the Indo-west Pacific (excluding the Red Sea) from as far south as Durban to southern Japan, is described. Two new species of Enneapterygius, E. clarkae from the western Indian Ocean including the Red Sea, and E. ventermaculus from Zululand, Aden and Pakistan, are described. The possi...