Wen Wen

Wen Wen
Chengdu Institute of Geology and Mineral Resources

MS

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49
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Publications

Publications (49)
Article
Hybodontiform shark teeth, cephalic and fin spines as well as placoid scales are reported here for the first time from the continental, Lower Triassic Dongchuan Formation near the village of Zhaojia (Huize County, Yunnan Province) in southwest China. The isolated teeth are assigned to three unidentified hybodontiforms but also to a new species of P...
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The end-Permian mass extinction led to the disappearance of >81% of marine species and the collapse of marine ecosystems. Despite the progress made in recent years, the severity of the impact of the mass extinction upon some groups still remains unclear. Horseshoe crabs are a typical example among these groups. Here we report a new fossil horseshoe...
Article
Lime mud (i.e., micrite) is a major component of carbonate deposits. Various mechanisms (biotic versus abiotic) have been proposed for the formation of lime mud in Earth’s history. However, the detailed role that microbes play in the nucleation and subsequent precipitation of micrites remains to be resolved. Herein we undertook a detailed geobiolog...
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The Luoping Biota represents of the recovery of marine ecosystems after the Permian–Triassic Mass Extinction. To better understand the palaeoenvironmental background of the Luoping Biota, sedimentary analysis and multi‐geochemical proxies (Mo, U, V, V/Cr, MoEF, UEF, and VEF) were assessed in Member II of the Middle Triassic Guanling Formation of th...
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The morphology and position of elements in the apparatus are keys to resolving the taxonomy, homology, evolutionary relationships, structure, function and feeding patterns among conodont taxa. Fused clusters preserving natural associations between elements provide direct information on element morphology, the positions of elements within the appara...
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The secular change in ⁸⁷Sr/⁸⁶Sr ratio of the Mesozoic recorded the highest value above 0.7083 in Early–Middle Triassic boundary, i.e., the Triassic maximum, suggesting that a major reform in oceanography occurred after the end-Permian mass extinction. We have detected correlative highest ⁸⁷Sr/⁸⁶Sr value from the Triassic carbonate, including our da...
Article
We reconstruct the apparatus architecture of the gondollelid conodont Nicoraella kockeli based on fused clusters from the early Middle Triassic (middle Anisian, Pelsonian) of Luoping County, east Yunnan Province, south‐west China. This material was characterized non‐invasively using synchrotron x‐ray tomographic microscopy and the ensuing data anal...
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The original version of this article unfortunately contained a mistake. The title was incorrect. The corrected one is given below. Palaeoecological Analysis of Trace FossilSinusichnus sinuosusfrom the Middle Triassic Guanling Formation in Southwestern China
Article
The composition of conodont apparatuses is crucial for understanding the feeding mechanisms of these early vertebrates. However, the multielement apparatus reconstructions of most species remain equivocal because they have been inferred from loose element collections, guided by knowledge from rare articulated ‘bedding plane assemblages’ and fused c...
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The Luoping Biota discovered from the early Middle Triassic (Anisian) Guanling Formation of southwestern China represents a fully recovered shallow marine ecosystem, marking the end point of Early Triassic biotic recovery following the end-Permian mass extinction. Contemporaneously preserved are prolific trace fossils, which offer good opportunitie...
Article
The morphology and position of elements in the apparatus are keys to resolving the taxonomy, homology, evolutionary relationships, structure, function and feeding patterns among conodont taxa. Fused clusters preserving natural associations between elements provide direct information on element morphology, the positions of elements within the appara...
Article
A new helminthomorph millipede, Sinosoma luopingense new genus new species, from the Triassic Luoping biota of China, has 39 body segments, metazonites with lateral swellings that bear a pair of posterolateral pits (?insertion pits for spine bases), and sternites that are unfused to the pleurotergites. This millipede shares a number of characters w...
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Four complete platysiagid fish specimens are described from the Luoping Biota, Anisian (Middle Triassic), Yunnan Province, southwest China. They are small fishes with bones and scales covered with ganoine. All characters observed, such as nasals meeting in the midline, a keystone-like dermosphenotic, absence of post-rostral bone, two infraorbitals...
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The Permian–Triassic bottleneck has long been thought to have drastically altered the course of echinoid evolution, with the extinction of the entire echinoid stem group having taken place during the end-Permian mass extinction. The Early Triassic fossil record of echinoids is, however, sparse, and new fossils are paving the way for a revised inter...
Article
Trace fossils have proven useful for studying the timing and process of biotic recovery after the Permian–Triassic Mass Extinction (PTME). Recovery stages are defined by comparing successive ichnoassemblages from the latest Permian to the early Middle Triassic. Lower Triassic trace fossils have been explored in some detail, but those of the lower M...
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Horseshoe crabs are classic “living fossils”, supposedly slowly evolving, conservative taxa, with a long fossil record back to the Ordovician. The evolution of their exoskeleton is well documented by fossils, but appendage and soft-tissue preservation is extremely rare. Here we analyse details of appendage and soft-tissue preservation in Yunnanolim...
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This paper reports a new specimen of decapod from Xinjiangdi Village of Chajiang Town, Luoping County, Yunnan Province. The new material comes from Triassic (Carnian) Xiawa Formation. It is the first discovery of the decapod from the Guanling biota, and also the first report from the Upper Triassic strata in China. The new findings increase the div...
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Fourteen specimens of cyclidan arthropods form the basis for the description of a new genus and species of Anisian (Middle Triassic) specimens, Yunnanocyclus nodosus n. gen., n. sp., from the Middle Member of the Guanling Formation, Luoping Biota in Yunnan Province, China. Specimens preserve dorsal, ventral, and internal aspects of the morphology o...
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The Permo-Triassic mass extinction devastated life on land and in the sea, but it is not clear why some species survived and others went extinct. One explanation is that lineage loss during mass extinctions is a random process in which luck determines which species survive. Alternatively, a phylogenetic signal in extinction may indicate a selection...
Article
Marine reptiles fossils, accompanied by abundant ammonites, brachipods, bivalves, conodonts and some other fossils, were discovered for the first time from the Lower Triassic Luolou Formation in Wangmo area of the Nanpanjiang Basin. Their age belongs to the Spathian substage of the Early Triassic Olenekian stage. This new discovery has a prominent...
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Tiny, pelagic arthropods from the Anisian Luoping Biota exposed in two quarries near Luoping, Yunnan Province, China, represent the numerically most abundant organisms in the assemblage. They form the basis for definition of two, and possibly three, species referred to the order Lophogastrida, family Eucopiidae. Yunnanocopia grandis new genus new s...
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A large assemblage of small specimens of Tridactylastacus sinensis Feldmann, Schweitzer, and Zhang in Feldmann et al., 2012, from the Middle Triassic (Anisian) Luoping Biota, Yunnan Province, China, permitted cladistic analysis confirming their placement within Litogastridae in the superfamily Glypheoidea. Orientation of 86 specimens closely spaced...
Article
Abundant, exceptionally preserved coprolites are documented from the Luoping biota (Anisian, Middle Triassic) of Yunnan Province, southwest China. These coprolites can be categorized into four morphological types: A) bead to ribbon-shaped, B) short to long cylindrical-shaped, C) flattened, disk-like, and D) segmented faeces. Detailed multi-discipli...
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The distribution of all recognized biotic elements was mapped on five extensive surfaces of the Middle Triassic Guanling Formation exposed in quarries near Luoping, Yunnan Province, China. A 0.5 m 2 grid was overlain on the surfaces, and all recognizable fossils were located by placing them in one of nine quadrants within the grid. A total of 240 m...
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The Luoping Biota was discovered in 2007 by the Chengdu Center of China Geological Survey during 1:50000 regional mapping works at Dawazi Village, Luoxiong Town, 15 km southeast of Luoping City, Yunnan Province. The interval containing the Luoping biota is the middle to upper part of Member II of the Guanling Formation. The fossil assemblage of the...
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The presence of gigantic apex predators in the eastern Panthalassic and western Tethyan oceans suggests that complex ecosystems in the sea had become re-established in these regions at least by the early Middle Triassic, after the Permian-Triassic mass extinction (PTME). However, it is not clear whether oceanic ecosystem recovery from the PTME was...
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An early Middle Triassic stromatolite deposit is documented from the Guanling Formation of the Luoping area, Yunnan, SW China. The Luoping stromatolite shows five types of constructional microbial forms in various magnifications: 1) typical stratified columnar structures, up to 20 cm high, with crinkled laminae. Dark coloured laminae, 1 mm thick, a...
Article
The Triassic was a time of turmoil, as life recovered from the most devastating of all mass extinctions, the Permo-Triassic event 252 million years ago. The Triassic marine rock succession of southwest China provides unique documentation of the recovery of marine life through a series of well dated, exceptionally preserved fossil assemblages in the...
Article
The Triassic was a time of turmoil, as life recovered from the most devastating of all mass extinctions, the Permo-Triassic event 252 million years ago. The Triassic marine rock succession of southwest China provides unique documentation of the recovery of marine life through a series of well dated, exceptionally preserved fossil assemblages in the...
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Full-text available
The seas of the Mesozoic (266-66 Myr ago) were remarkable for predatory marine reptiles, but their modes of locomotion have been debated. One problem has been the absence of tracks, although there is no reason to expect that swimmers would produce tracks. We report here seabed tracks made by Mesozoic marine reptiles, produced by the paddles of noth...
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Two new genera, Anisaeger and Distaeger, and three new species, Anisaeger brevirostrus, A. spiniferus, and Distaeger prodigiosus, extend the range of the Aegeridae (Dendrobranchiata, Penaeoidea) into the Middle Triassic (Anisian) of China. Seven decapod crustacean species are now known from the Luoping biota of southern China. Morphological feature...
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Num erous types of geoheritage resources are widely distributed in the Luoping Biota National Geopark. The main parts of the geoheritage resources include exceptionally preserved diverse fossils of the Luoping Biota and fossil-bearing sections, associated with typical and beautiful kant landscape, springs, waterfalls, lakes and other water landscap...
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Exceptionally preserved carbonate reticulated ridge structures are documented from the lower Middle Triassic Guanling Formation of the Luoping area, eastern Yunnan Province, southwestern China. Macrostructures suggest these reticulated ridge structures are morphologically similar to the growth-related structures in ancient and modern microbial mats...
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A new penaeoid shrimp collected from the Middle Triassic Member II of the Guanling Formation in the vicinity of the city of Luxi, Yunnan, southwest China, is a new species, Aeger luxii n. sp. The new species possesses prominent spinose third maxillipeds, which is one of the typical characteristics of Aeger. The new species differs from the type spe...
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Based on newly collected materials from the Lower Triassic Feixianguan Formation of Xiongwu section in Xingyi, Guizhou and the Ximatang Formation of Ximatang section in Qiubei, Yunnan, southern China, nine species belonging to seven genera are described. Two bivalve assemblages are recognized and regionally correlated in South China. The bivalve as...
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The holothurian sclerites fossil was found in the Member II of Guanling Formation of Middle Triassic (Anisian, Pelsonian) from Luoping, Yunnan Province. One species in one genera is described. The discovery enriches the composition of the Luoping biota and provides new materials for the research of the taxonomy and evolution of Triassic holothurian...
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A large collection of macrurous decapod crustaceans is recorded from the middle–late Anisian (Middle Triassic) Guanling Formation in Yunnan Province, China. A remarkable assemblage of over 20,000 vertebrate, invertebrate, and plant fossils collectively referred to as the Luoping Biota has been collected from quarries in the vicinity of the city of...
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new genus of actinopterygian fish from the Anisian (Middle Triassic) of Luoping, Yunnan Province, Southwest China. Acta Palaeontologica Polonica 5X (X): xxx-xxx. doi:10.4202/app.2010.0089 The new neopterygian fish taxon Luoxiongichthys hyperdorsalis gen. et sp. nov. is established on the basis of five specimens from the second member of the Guanlin...
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The fossil record of coelacanths is patchy, with very few taxa known from the Triassic of Asia. We report here two new genera and species of coelacanths from the Luoping Biota, a recently found site of exceptional fossil preservation from Yunnan, South China. The first new taxon, Luopingcoelacanthus eurylacrimalis, is based on four specimens, which...
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The timing and nature of biotic recovery from the devastating end-Permian mass extinction (252 Ma) are much debated. New studies in South China suggest that complex marine ecosystems did not become re-established until the middle-late Anisian (Middle Triassic), much later than had been proposed by some. The recently discovered exceptionally preserv...

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