Vinay Sahu

Vinay Sahu
  • M.Sc., Ph.D. (Environ. Sci.), Fellow of Association for plant Taxonomy, Fellow of Indian Botanical Society
  • Senior Technical officer at National Botanical Research Institute

About

106
Publications
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Introduction
Assistant Professor AcSIR CSIR- National Botanical Research Institute, Rana Pratap Marg, Lucknow, UP India, 226018 Current area of work 1) Exploration and survey of Bryophytes and Herbarium enrichment 2) In vitro culture and conservation of rare and potential bryophytes 3) Prospection of Bryophytes for antimicrobial and medicinal properties
Current institution
National Botanical Research Institute
Current position
  • Senior Technical officer
Additional affiliations
June 2001 - present
National Botanical Research Institute
Position
  • Senior Technical Officer
June 2001 - present
National Botanical Research Institute
Position
  • Senior Technical officer

Publications

Publications (106)
Article
The present study deals with the investigation of moss flora of Govind Wild Life Sanctuary (GWLS), Uttarkashi, Uttarakhand. It includes an account of 216 species of mosses belonging to 103 genera of 32 families. Families Pottiaceae and Bryaceae seem to be more dominant in the region with 32 species, followed by Brachytheciaceae and Thuidiaceae. Bot...
Article
This study addresses the pressing issue of high arsenic (As) contaminations, which poses a severe threat to various life forms in our ecosystem. Despite this prevailing concern, all organisms have developed some techniques to mitigate the toxic effects of As. Certain plants, such as bryophytes, the earliest land plants, exhibit remarkable tolerance...
Article
Full-text available
The present paper provides an account of 61 taxa of liverworts and three hornwort species observed during recent survey of Nagaland. Out of these, 46 taxa (44 liverworts, 2 hornworts) are being reported for the first time from Nagaland. Cephaloziella herzogiana (Pande and Srivastava) Udar and Kumar, and Porella caespitans (Steph.) S. Hatt. var. cae...
Article
Present study reveals the occurrence of 15 taxa of the genus Fissidens Hedw. from Assam. Eight species viz. Fissidens biformis Mitt., F. bryoides Hedw., F. ceylonensis Dozy. & Molk., F. curvatoinvolutus Dixon., F. flaccidus Mitt., F. griffithii Gangulee, Thwait ex Mitt., F. pellucidus Hornsch. and F. zollingeri Mont. are new to Assam, while F. cras...
Article
New records of 18 species of mosses and liverworts are presented, with detailed notes on their taxonomy, distribution, and ecological characteristics. These findings provide valuable insights for advancing research in biogeography, ecology, and conservation.
Article
Full-text available
Marchantia polymorpha L. (Marchantiaceae), a thalloid liverwort, is cosmopolitan in distribution. It has various medicinal properties by exhibiting antimicrobial activity, cytotoxic activity, antifungal activity, antipy-retic activity, and diuretic activity. In the present study, chemical profiling of M. polymorpha by using GCÀMS has been carried o...
Article
A study on the quantitative estimation of diversity in 32 selected grids in the bryophyte-rich localities of Assam, North East India has been carried out. Assessment was done in 160 macroplots of 10 m  10 m, randomly esta-blished in the forest and within each macroplot, five microquadrats (10 cm  10 cm) were placed at differ-ent habitats, viz. sa...
Article
Present study on liverworts diversity of some underexplored areas of Manipur has revealed the occurrence of 23 genera and 32 species of liverworts belonging to 17 families. Fourteen species have been reported for the first time from Manipur while three Indian endemic species (Cephalozia kashyapii Udar, Microlejeunea indica (Udar et U. S. Awasthi) Y...
Article
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Plagiochila pseudopoeltii, a leafy liverwort is reported here for the first time from Kashmir Himalaya. The species was collected from Dachigam National Park, Srinagar district, and grows on moist soil as well as on the tree trunk. A detailed illustrated account of the species has been provided to validate the new distribution record and facilitate...
Article
Full-text available
During an investigation on mosses of some underexplored regions of Nagaland, 121 taxa of mosses belonging to 74 genera and 29 families have been identified. Atrichum crispulum Schimp. & Besch. and Plagiothecium neckeroideum var. niitakayamae (Toyama) Z. Iwats. are new records for India. Five taxa namely Fissidens crassinervis var. laxus (Sull & Les...
Article
The present paper deals with the investigation of moss taxa of some underexplored regions of Manipur, Northeast India. During the assessment of moss diversity in various localities of Manipur a total 126 species of mosses belonging to 84 genera in 34 families have been investigated and enumerated. Ten species, namely Amblystegium serpens,Entodontop...
Book
Full-text available
The book provides an original account of the mosses of Govind wildlife Sanctuary, Uttarakhand and has 211 taxa belonging to 113 genera and 33 families supported by 110 color plates. This work includes six species new to the bryflora of India, 3 species new to Himalaya, 28 species new to Western Himalaya and 32 species new to Uttarakhand. The study...
Article
On the basis of compilation of earlier reports and results of fresh exploration, the present work provides a list of 113 bryophyte taxa from Mount Abu, Rajasthan including 24 taxa as new to the Mount Abu area. Out of these, 14 are new records to Rajasthan state, while 07 taxa are new to Central Indian bryo-geographical region.
Article
Pohlia leucostoma (Bosch & Sande Lac.) M. Fleisch. has been identified and described as a new addition to the bryoflora of India from Govind Wildlife Sanctuary, Uttarkashi, Uttarakhand. The taxon is characterized by acrocarpous, bright green, glossy plants with linear and filiform leaf propagules with 1-2 peg-like leaf primordia, forming clusters i...
Article
Full-text available
New records of 48 species of mosses and liverworts are presented, with detailed notes on their taxonomy, distribution, and ecological characteristics. These findings provide valuable insights for advancing research in biogeography, ecology, and conservation.
Article
Full-text available
Plagiobryum zieri (Dicks. ex Hedw.) Lindb. inhabitant of higher altitude has been recorded for the first time from Uttarkashi, Uttarakhand of the Western Himalaya earlier known from Sikkim (Eastern Himalaya). About 25 species of Plagiobryum are known worldwide. A morpho-taxonomic account of Western Himalayan plants of Plagiobryum zieri is provided.
Article
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During study on the family Mniaceae in Darjeeling and its neighbouring areas, three genera and six species (Mnium lycopodioides, Orthomnion bryoides, Plagiomnium acutum, P. confertidens, P. rhynchophorum and P. succulentum) have been identified. Of these Plagiomnium acutum is reported here for the first time from eastern Himalaya. A detailed morpho...
Article
During the course of intensive exploration of the Govind Wild Life Sanctuary, Uttarakhand, Western Himalaya, some interesting specimens of genus Paraleucobryum (Lindb. ex Limpr.) Loeske have been identified. A critical morpho-taxonomic study of these plants has revealed its close resemblance with P. enerve (Thed.) Loeske; however, it differs from t...
Article
Full-text available
Mylia taylorii (Hook.) Gray, a rare taxon has been collected from Singalila National Park, Darjeeling (West Bengal), in the subalpine region, on way to Phalut from Sandakphu (ca 3657 m). A detailed morpho-taxonomic and illustrated account of the Indian plants is provided. Keywords: Eastern Himalaya, India, Liverwort, Mylia, Rare.
Article
Leaf cell papillae and mamillae and peristome teeth of selected moss species of Mount Abu (Rajasthan) were examined using SEM and results are provided.
Article
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Ellis L.T., Almeida L.A., Peralta D.F., Bačkor M., Baisheva E.Z., Bednarek-Ochyra H., Burghardt M., Czernyadjeva I.V., Kholod S.S., Potemkin A.D., Erdağ A., Kırmacı M., Fedosov V.E., Ignatov M.S., Koltysheva D.E., Flores J.R., Fuertes E., Goga M., Guo S.-L., Hofbauer W.K., Kurzthaler M., Kürschner H., Kuznetsova O.I., Lebouvier M., Long D.G., Mamon...
Article
Abstract: The diversity of the liverwort genus Porella is studied in the Govind Wildlife Sanctuary (GWLS), Uttarakhand. A total of 12 species and infraspecific taxa are described, all being reported for the first time from the area. A working key is provided for identification along with the taxonomic description of each of them. As Porella is repr...
Article
Full-text available
During the present study an effort has been made to propagate (in-vitro) the endangered and endemic Indian liverwort Stephensoniella brevipedunculata Kashyap using different culture media under controlled Laboratory conditions. Axenic cultures of the taxon have been established using tubers as explants. Seven combinations of media with Full Strengt...
Article
We here report, a total of thirty bryophyte (sixteen liverworts and fourteen mosses) taxa from the Tawang District, Arunachal Pradesh, India. Of these, Heterophyllium affine (Hook.) M. Fleisch. is reported here as new to India while the eight liverworts [Anastrophyllum minutum (Schreb.) R.M. Schust., Frullania retusa Mitt., F. tagawana (S. Hatt. &...
Article
During investigation on the bryoflora of Govind Wild Life Sanctuary, Uttarakhand, India, a rare liverwort, Plagiochila ghatiensis Stephani (Plagiochilaceae), with known restricted occurrence within south India, has been identified amongst the bryoflora of Himalaya, north India. The taxon is characterized by filiform, flagilleferous, microphyllous,...
Article
Full-text available
The growth pattern of Marchantia polymorpha subsp. ruderalis, using gemmae as explant, in different culture media in combination with auxin and cytokinins has been studied. The plant growth was maximum in ½ KNOP’s followed by KNOP’s, Hoagland and Murashige & Skoog media. The normal growth of gemmae was inhibited at higher concentrations of hormone...
Article
ABSTRACT: During investigation on the bryophytes of The Govind Wildlife Sanctuary. Grimmia nepalensis has been identified for the first time from this region. It is a new record for India and has been collected after about 150 years since its original collection from Nepal. The species is endemic to the Himalaya. KEYWORDS: Grimmia nepalensis, Govin...
Article
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In vitro propagation of Splachnum sphaericum Hedw. has been carried out to study its growth pattern and morphogenetic attributes. Plants of this species were freshly collected from Se La Pass, Tawang, Arunachal Pradesh. Axenic culture of the species has been established using spores as explants. Half Knop's macronutrients, Half Knop's + Vitamins, H...
Article
Anthelia julacea (L.) Dumort., a less known plant of Indian bryoflora, has been rediscovered from Himachal Pradesh and Uttarakhand after a gap of around eight decades. The study confirms its presence in the country and also makes its first ever report from Uttarakhand.
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A total of 30 taxa of bryophytes are reported for the first time from Tawang district of Arunachal Pradesh, India, including 10 taxa as new to Arunachal Pradesh.
Article
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The study compares the effect of Arsenic (As) on some physiological metabolites and anti-oxidant enzymes in two thalloid liverworts Marchantia paleacea Bertol. and Marchantia papillata subsp grossibarba (Stephani) Bischel, having different morphological characteristics. The plants were administered with different concentrations of As ranging from 5...
Article
Gymnomitrion crenatilobum Grolle, earlier reported from Nepal and known from the type only, is reported from Uttarakhand, India. The plants are clearly distinguishable based on a 2-3 cells wide hyaline leaf margin which is crenulate in apical half of leaves and smooth in basal half.
Article
Kurzia trichoclados (K. Müller) Grolle, is a disjunct liverwort known from Europe, North America and Southeast Asia. Its distribution in South Asia has now been expanded to include India.
Article
During an investigation on the bryophytes of western Himalaya, Lejeunea aquatica Horik. was identified from Lohaghat area (Uttarakhand) and is reported as a new addition to the Indian bryoflora. Prior to this study, it was known only from China, Japan and Taiwan. The species is characterized by its monoicous sexuality, small lobules which is 1/5–1/...
Article
Present study provides an account of 72 taxa of mosses belonging to 24 families. During the study five mosses Fissidens polysetulus C. Muell. ex Gangulee & Norkett, Ectropothecium kerstanii Dix. & Herz., Isopterygium serrulatum Fleisch., Weissia edentula Mitt., Rhachithecium perpusillum (Thwait & Mitt.) Broth. are new to western Himalaya, while 10...
Article
Abstract Govind Wildlife Sanctuary is very rich in Biodiversity. During recent investigation on the bryoflora of this area, Campylopus schmidii (Mu¨ll. Hal.) A. Jaegr. (Dicranaceae) has been identified, which is a new addition to the bryoflora of Indian Himalaya. It was previously reported from South India. In India, the genus is represented by 10...
Article
Full-text available
Abstract During an investigation on the bryophytes of Gangetic plains, Archilejeunea minutilobula Udar et U. S. Awasthi has been identified at Farakka, district Murshidabad (West Bengal) which is a new addition to the Gangetic plains and West Bengal. Previously it was known from South India (Mettupalayam, Mahabaleshwar) only. The taxon belongs to s...
Article
In-vitro propagation of three species of genus Bryum, viz. B. argenteum Hedw., B. billardieri Schwaegr. and B. pallescens Schleich ex Schwaegr. has been carried out to study their growth patterns and morphogenetic attributes. Plants of all the three species were freshly collected from Govind Wildlife Sanctuary (Uttarakhand). Axenic cultures of thes...
Article
The Govind Wild Life Sanctuary (GWLS), established in 1955, is situated in Uttarkashi district of Uttarakhand. The whole area of the Sanctuary is covered with light to heavy snow and is very rich in biodiversity. During the course of extensive bryological exploration of this area, specimens of Tetralophozia filiformis (Steph.) Urmi have been collec...
Article
Rawat K. K., Sahu V., Verma P. K. & Asthana A. K. 2015. Horikawaella S. Hatt. & Amakawa (Marchantiophyta: Solenostomataceae) from Sikkim, India. Geophytology 45(1): 67-70. Genus Horikawaella S. Hatt. & Amakawa, a rare taxon of Indian bryoflora, is endemic to Himalaya and is represented in India by single species H. subacuta (Herzog) S. Hatt. & Amak...
Article
Govind Wild Life Sanctuary in the Western Himalayas with its diverse vegetation types provides excellent habitats and microclimates for a lush growth of bryophytes. During investigations on the bryophytes of this region Plagiothecium euryphyllum (Cardot et Thér.) Z. Iwats. and P. cavifolium (Brid.) Z. Iwats. have been identified for the first time...
Article
Full-text available
Abstract: An investigation of the Bryophytes from Terai region of Uttar Pradesh has revealed the occurrence of 29 species of bryophytes: 21 species belonging to 16 genera of 11 Families of Mosses; 6 species belonging to 3 genera of 3 families of Liverworts and 2 species belonging to 1 genus of 1 family of Hornworts. This includes several new report...
Article
Abstract Rudrabhisek is a tradition of worshipping Lord Shiva in his Rudra Avatar. This pooja gives wealth, prosperity, purify the bed Karma & give the shield in life for different types of diseases. Serva Aushidi (Plant materials for overcoming diseases) is offered to Lord Shiva for defending devotees from various kinds of diseases. This Serva Aus...
Article
Govind Wild Life Sanctuary situated in Uttarkashi District, Uttarakhand harbours a rich biodiversity. During investigation on the bryophytes of this region, Fabronia pusilla Raddi (Family Fabroniaceae) and Didymodon michiganensis (Steere) Saito (Family Pottiaceae) have been identified in this reserve area for the first time. Earlier, F. pusilla was...
Article
The Govind Wildlife Sanctuary is situated in Uttarakashi district of Uttarakhand. The area is very rich in biodiversity in general and bryodiversity in particular. During recent investigation on the bryoflora of this area, two species of mosses viz., Cyathophorella burkillii (Dixon) Broth. (Family-Hypopterygiaceae) and Symphyodon erraticus (Mitt.)...
Article
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alpestris (Thed.) Schimp. Contributor: L. Thouvenot Andorra: Encamp, Ensagents, 42u319070N, 1u389410E, 2520 m a.s.l., on the top of granitic rock in alpine meadow, 17 July 2006, leg. L. Thouvenot 3086 (BCB 58238). According to Casas (2005) this species was pre-viously unrecorded in Andorra (Eastern Pyrenees) although it is known in the neighbouring...
Article
During the present study Methanolic extracts of 9 thalloid liverworts were analyzed by HPLC which contain Gallic acid, Protocatechuic acid, Chlorogenic acid, Caffeic acid, Rutin, Ferulic acid, Quercetin and Kaempferol known as antioxidants and also have some role in toxicity against cancer cells. Plants were grown in moss House for ex situ conserva...
Article
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Abstract: The present study deals with the investigation of Pogonatum perichaetiale subsp. thomsonii from Watan Village, Pithoragarh. The important characteristics of this species are plants simple, leaves stiff, tufted and forming a bud like structure when dry, margin sharply toothed in upper part of the leaves, costa ends in a sharp awn like poin...
Article
An enumeration of 30 taxa of liverworts belonging to 13 families and single taxon of hornwort of Pithoragarh and its neighbouring areas (Mayawati-Lohaghat) has been made. One liverwort, Lejeunea aquatica Horik. has been recorded as new to India, four liverworts Frullania neurota Taylor, F. companulata Sde., Lejeunea brittoniae (Evans) Grolle,and L....
Article
ABSTRACT During investigation on the bryophytes of Govind Wild life Sanctuary, a rare moss Hedwigia ciliata (Hedw.) Ehrh. Ex P. Beauv. var. ciliata has been identified from this area. This species is found mainly in the Europe. Taxon has been reported first time from Uttarakhan, however, earlier it was merely reported from North Western Himalaya. T...
Article
During the present study in vitro propagation of acrocarpous moss: Anomobryum filiforme var. concinnatum (Spruce.) Aman. has been carried out on different culture media: Full strength Knop's macronutrients, half-strength Knop's macronutrients, Nitsch's trace elements + 10 ppm ferric citrate and Hoagland media with the aim to establish stable in vit...
Article
During investigation on the bryophytes of Darjeeling and its neighbouring areas, 6 species of Pogonatum (Pogonatum patulum (Harv.) Mitt., Pogonatum cirratum (Sw.) Brid. Pogonatum neesii (C. Muell.) Dozy. Pogonatum microstomum (Schwaegr.) Brid. Pogonatum perichaetiale (Mont.) Jaeg. Pogonatum urnigerum (Hedw.) P. Beauv.) have been identified. These a...
Article
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Liochlaena subulata (A. Evans) Schljakov, a species with fertile plants, is reported from Singalila National Park (Darjeeling) which is new to eastern Himalaya. Earlier it was known from western Himalaya and South India. It belongs to family Jungermanniaceae and characteristically possesses flagelliform and gemmiparous shoots developing near stem a...
Article
Full-text available
The present study deals with the investigation of bryophytic flora of Mukteshwar (Uttarakhand). The present account deals with the study of 38 species belonging to 31 genera of 21 families. The mosses are represented by 21 species of 20 genera and 12 families, while Liverworts are represented by 15 species of 10 genera and 8 families. Hornworts con...
Article
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Stichococcus bacillaris (Green Alga) growth was observed as a contaminant on two taxa of bryophytes, a thalloid liverwort Lunularia cruciata (L.) Dumort. ex. Lindb. and a hornwort Folioceros physocladus Bharad. ex Schiffn. et Pande, growing on soil in pots in acclimatization chamber of Moss House at National Botanical Research Institute, Lucknow, I...
Article
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Growth of nine species of cyanobacteria was observed on eight species of bryophytes (thalloid liverworts and mosses) growing in the polyhouse, Bryophyte conservatory (Moss House), at National Botanical Research Institute, Lucknow, India. Bryophytes were cultured in the laboratory and transferred to polyhouse for hardening and acclimatization. Cyano...
Article
The present study deals with the investigation of bryophyte flora of Jim Corbett National Park. It includes an account of 58 species belonging to 42 genera of 25 families. The mosses are represented by 38 species of 28 genera belonging to15 families, while liverworts are represented by 18 species of 14 genera belonging to 8 families and hornworts a...
Article
During an investigation on the bryophytes of Darjeeling and its neighbouring areas Polytrichastrum formosum (Hedw.) G.L. Smith has been identified from Sandakphu region, which is being reported for the first time from Indian territory. The plants of this species are distinctly characterized by its large, robust size; tomentose stem part below, usua...
Article
Full-text available
The Govind Wildlife Sanctuary is situated in Uttarakashi district of Uttarakhand. The area is very rich in biodiversity in general and bryodiversity in particular. During recent investigation on the bryoflora of this area, two species of mosses viz., Cyathophorella burkillii (Dixon) Broth. (Family-Hypopterygiaceae) and Symphyodon erraticus (Mitt.)...
Article
Full-text available
Thal loid liverwort Plagiochasma appendiculatum has been exposed to different concentrations of lead in the laboratory to observe the effect of Pb on Chlorophyll a, Chlorophyll b, Carotenoid and Protein contents of the plant. A decrease in the photosynthetic pigments and protein contents of the plants was observed with the increase in the concentra...
Article
Plagiochasma appendiculatum L. & L. (Aytoniaceae), Conocephalum conicum (L.) Necker (Conocephalaceae), Bryum argenteum Hedw. (Bryaceae), and Mnium marginatum (With.) P. Beauv. (Mniaceae) are bryophytes (liverworts and mosses) used by traditional healers for the treatment of burn, cuts, wounds, and skin disorders. This study evaluated the antibacter...
Article
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During a recent investigation on the Mosses of Meghaiaya (eastern Himalaya), plants of Erpodium domingense (Sprengel.) Muller Hal. have been identified from Shillong for the first time which is a new addition to the bryoflora of India. The taxon belongs to the family Erpodiaceae and is characterised by leaves arranged in more than four rows, densel...
Article
Full-text available
An attempt has been made to study in vitro growth and multiplication of epiphytically growing moss Brachymenium capitulatum on different culture media (Hogland, Half strength Knop's solution, Murashige and Skoog medium with Sucrose, Murashige and Skoog medium with Calcium chloride, Inositol and Thiamine, Murashige and Skoog medium with Calcium chlo...
Article
Full-text available
A recent investigation on the bryophytes of Darjeeling has revealed the occurrence of 12 taxa of hornworts (anthocerotes) in this region. Of these, Anthoceros punctatus, Folioceros assamicus, Phaeoceros himalayensis, Notothylas indica, N. khasiana, N. levieri are new to Darjeeling, while Anthoceros alpinus, a critically endangered species, is new t...
Article
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The capability of bryophytes to accumulate heavy metals, indicate their effectiveness for heavy metal pollution monitoring. They possess no real root system or continuous cuticle layer and absorb water and dissolved elements as atmospheric depositions directly through their entire body surface. Their cell walls are characterized by high ion exchang...
Article
The present investigation on the bryoflora of western Himalaya revealed the occurrence of three moss taxa, viz., Entodon concinnus subsp. caliginosus, Entodon luteonitens and Hypnum subimponens subsp. ulophyllum, which are new additions to the western Himalaya. Entodon concinnus subsp. caliginosus and H. subimponens subsp. ulophyllum have been disc...

Questions

Questions (4)
Question
My Plants more closely related to Ectropothecium.
Plants yellowish brown with orange tinge, prostrate, main stem up to 15 mm long, branches 3-7 mm long, leaves closely imbricate, falcate on one side, 0.7 mm long and 0.80 mm long and 0.2-0.24 mm wide, ovate, revoulte at apex on one or both sides, costa two short or faint. Leaf apical cells 20-28 µm long and 4 µm wide, linear rhomboid, median leaf cells 20-40µm long and 4 µm wide, basal alar cells short quadrate to rectangular, 8-16µm long and 4-8µm wide.
Question
Plants green above, brownish below, branching dichotomously, shoots 2 -2.5 cm long, densely covered with falcatosecund leaves, 5-7 mm long, lanceolate, at base 0.8-1 m wide, canaliculate. Leaf tip entire or minutely dentate at apex. Costa occupying more than ¾ of the leaf width. Leaf tip setaceous which is formed by the costa in the upper part. Alar slightly bulging out with large deep brown cells, 40-44 µm long and 20-24 µm wide, rectangul;ar. Basal cells 32-40 µm long and 8 µm wide, rectangular with porose incrassate walls.
This plant belongs to Family Dicranaceae. It may be Campylopus or Dicranum. Leaf section ?
Question
Can anybody help me to identify this moss or suggest some references. 
Plant yellowish brown, erect, 8-10 m long, unbranched. Leaves closely imbricate at apex and distantly arranged at base, erectopatent, ovate oblong, 1.28-1.5 mm long and 0.68-0.80 mm wide, margin entire or crenulate at apex. Leaf obtuse or apiculate at apex. Leaf costa ending just below leaf apex, 80 – 100 µm wide at base. Leaf apical cells 16-28  µm long and 16-28  µm wide, basal cells 20 - 44 µm long and 16-20  µm wide.
Explanation of fig 1- A: Plant; B: Leaf; C: median cells; D: apical cells; E: basal cells.
Question
Plants prostrate with erect long olive green branches, brownish below. Plants 4.0-5.5 cm long. Stem in cross section rounded, 280 - 320 µm long, cortical cells thick walled 8-12 µm wide, medullary cells 16-28 µm wide, polygonal. Central strand absent. Leaves appressed to stem when dry, closely imbricate, ovate lanceolate, 1.8-2 mm long and 0.8-1 mm wide. Leaves end in a hyaline tip with dentate margins and elongated cells covered by papillae, margin revolute at base. Costa absent. Leaf apical cells incrassate, sinuose, short quadrate to rectangular in shape, 8-12 µm long and 8 µm wide, papillate, median leaf cells short quadrate to rectangulate, incrassate, sinuose, papillate. Leaf cells at base in the middle part, linear, rhomboidal, incrassate, sinuose, papillate, 40-88 µm long and 4-8 µm wide. The larger cells near base are more sinouse and more papillose. Alar cells short quadrate to rectangular, thick walled, 8-12 µm wide.

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