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Publications (172)
Wege, J.A., Brennan, K.G., Bean, A.R., Barrett, R.L., Dillon, S.J. & Barrett, M.D. Stylidium miscellany IV: novel species, recircumscriptions and range extensions for northern Australia. Nuytsia 35: 141–198 (2024). This paper advances knowledge of triggerplant diversity in monsoonal and arid Australia. The following 14 species are newly described:...
Eucalyptus virgata was validly published in 1827, but the name has not been in common usage for several decades. Its identity has been uncertain because the type lacks mature flower buds and fruits, and there was no description provided for bark or habit. The current availability of numerous images of the type has allowed a careful examination of s...
An examination of the type of Leucopogon biflorus R.Br. (current name Styphelia biflora (R.Br.) Spreng.) has shown that it belongs in Styphelia setigera (R.Br.) Spreng., hence S. biflora is placed in the synonymy of S. setigera. The next available name for the taxon previously known as Styphelia biflora is Leucopogon sparsus A.Cunn. ex DC. The new...
The Brachyscome microcarpa complex, including B. whitei, is taxonomically revised. B. discolor C.Stuart ex Benth. is reinstated, B. whitei G.L.Davis (formerly B. whitei subsp. whitei) is retained, and B. microcarpa F.Muell. is retained with a narrower circumscription. B. pedleyi A.R.Bean sp. nov. (formerly B. whitei subsp. lophoptera), B. stenoloba...
The new species Styphelia kinkuna A.R.Bean is described from the coastal lowlands of the Wide Bay district,
between Gympie and Bundaberg, and Mount Walsh near Biggenden. It is illustrated and compared
to related species, and a distribution map is provided
Hydrocotyle elegans R.Br. ex A.Rich. is newly reinstated and described. Eight species names are lectotypified and notes provided. An identification key for the 19 species of Hydrocotyle in Queensland is included.
Four species of Hydrocotyle L. occurring in Queensland and northern New South Wales are newly described, i.e. H. demissa sp. nov., H. fontana sp. nov., H. inops sp. nov., and H. maculosa sp. nov. They are illustrated, diagnosed against related species, and their distributions mapped.
The new name Seringia impressa A.R.Bean is proposed for the species formerly known as Seringia lanceolata Steetz
Neptunia comprises 18 species for Australia and Malesia, including ten species newly named: N. heliophila A.R.Bean, N. hispida A.R.Bean, N. insignis A.R.Bean, N. longipila A.R.Bean, N. paucijuga A.R.Bean, N. proxima A.R.Bean, N. scutata A.R.Bean, N. tactilis A.R.Bean, N. valida A.R.Bean and N. xanthonema A.R.Bean. One taxon below species rank (N. a...
Polyosma in Australia comprises eight endemic species that occur in rainforests along the east coast. All eight species are here fully described, including two new species from the Wet Tropics of Queensland (P. globosa A.R.Bean & P.I.Forst., P. nigrescens A.R.Bean & P.I.Forst., both illustrated with line drawings), with notes on distribution (inclu...
We introduce the AusTraits database - a compilation of values of plant traits for taxa in the Australian flora (hereafter AusTraits). AusTraits synthesises data on 448 traits across 28,640 taxa from field campaigns, published literature, taxonomic monographs, and individual taxon descriptions. Traits vary in scope from physiological measures of per...
Six major botanical trips undertaken by F.M. Bailey between 1873 and 1898 are documented in the colony of Queensland in Australia and British New Guinea on the island of New Guinea. These were to Cardwell in 1873, Tambo in 1876, Trinity Bay in 1877, Roxborough Downs in 1895/96, Torres Strait in 1897 and British New Guinea in 1898. For all journeys...
The genus Lagenophora Cass. is taxonomically revised for New Caledonia with two species recognised. Lagenophora sinuosa Lannuzel, Gâteblé & Jian Wang ter, sp. nov. is endemic to New Caledonia and the other, L. sublyrata (Cass.) A.R.Bean & Jian Wang ter occurs there and in many other countries from the region. Both are fully described and illustrate...
Bean, A.R. (2021). Ehretia dissita A.R.Bean (Ehretiaceae), a new species from tropical rainforest in Queensland, Australia. Austrobaileya 11: 34-40. Ehretia dissita is described as new and compared to its putative closest relative E. membranifolia. A key to the Australian species of Ehretia, and images of domatia, leaf venation and flower types are...
We introduce the AusTraits database - a compilation of measurements of plant traits for taxa in the Australian flora (hereafter AusTraits). AusTraits synthesises data on 375 traits across 29230 taxa from field campaigns, published literature, taxonomic monographs, and individual taxa descriptions. Traits vary in scope from physiological measures of...
The genus Camptacra N.T.Burb. is taxonomically revised. Four species are recognised, including two newly named: C. perdita A.R.Bean and C. robusta A.R.Bean. A lectotype is chosen for Eurybia gracilis Benth. All species are fully described with notes on distribution (including maps), habitat and conservation status. A key to the identification of al...
A morphological reassessment of specimens identified as Styphelia cuspidata (R.Br.) Spreng. (syn. Leucopogon cuspidatus R.Br.) has revealed the existence of three taxa that differ at species rank. In this paper, these three species, all endemic to Queensland, are described, compared and illustrated. Two new species, Styphelia cognata A.R.Bean and S...
Melaleuca oblivia A.R.Bean and M. stenostachya subsp. amplior A.R.Bean are newly described, and M. nervosa subsp. crosslandiana (W.Fitzg.) Barlow ex Craven is reinstated. Distribution maps and illustrations are provided for all taxa discussed. An identification key for all Queensland members of the M. leucadendra group is included.
The broadly circumscribed Olearia elliptica DC. is taxonomically revised and two new species O. fulgens A.R.Bean sp. nov., and O. praetermissa (P.S.Green) A.R.Bean comb. et stat. nov. are recognised. All three species are fully described with an identification key and a distribution map provided. A lectotype is chosen for Olearia elliptica.
The genus Lagenophora Cass. is taxonomically revised for New Guinea with two species recognised. One (Lagenophora sporadica Jian Wang ter & A.R.Bean sp. nov.) is endemic to New Guinea and the other (L. sublyrata (Cass.) A.R.Bean & Jian Wang ter) is widespread in Asia, Australia and Malesia. The species are described and illustrated. Notes are provi...
The genus Lagenophora Cass. is taxonomically revised for Australia with 12 species recognised from Western Australia, South Australia, Queensland, New South Wales, Australian Capital Territory, Victoria and Tasmania. Nine species are endemic to Australia and three also occur elsewhere. Lagenophora adenosa Jian Wang ter & A.R.Bean and L. platysperma...
The genus Lagenophora Cass. is taxonomically revised for Australia with 12 species recognised from Western Australia, South Australia, Queensland, New South Wales, Australian Capital Territory, Victoria and Tasmania. Nine species are endemic to Australia and three also occur elsewhere. Lagenophora adenosa Jian Wang ter & A.R.Bean and L. platysperma...
In this contribution, we treat all the names in the genus Eucalyptus published by F. Dehnhardt, including the much‐discussed E. camaldulensis. Current opinion would suggest that Dehnhardt applied this name to a plant different from that to which the name has commonly been applied. Hence, E. camaldulensis was conserved with a conserved type. Despite...
A detailed study of the plant specimens of Edward Macarthur Bowman is presented, including notes on their quality, numbering and labelling. This is followed by a chronology of Bowman's collecting activities, including maps depicting the routes he took on his major journeys. Samples of his handwriting and specimen labels are included, and a discussi...
Bean, A.R. (2018). Stemodia anisata A.R.Bean (Plantaginaceae), a new species from Queensland and the Northern Territory. Austrobaileya 10(2): 242-246. Stemodia anisata A.R.Bean is described, illustrated and compared to related taxa. A key to the Stemodia species occurring in Queensland is provided.
The genus Argophyllum is taxonomically revised for Australia with eleven species recognised from Queensland and northeast New South Wales. All Australian species are endemic and seven new species are described: A. curtum A.R.Bean & P.I.Forst., A. ferrugineum A.R.Bean & P.I.Forst., A. heterodontum A.R.Bean & P.I.Forst., A. iridescens A.R.Bean & P.I....
This paper provides species descriptions, specimen citations, identification key and illustrations for two species of the genus Iotasperma G.L.Nesom, namely I. australiense G.L.Nesom and I. sessilifolium (F.Muell.) G.L.Nesom. Both species names are lectotypified, and a distribution map is included.
The 14 species of Teucrium L. indigenous to Queensland are listed with full nomenclatural data and notes on distribution and habitat. Five species (T. daucoides A.R.Bean, T. fallax A.R.Bean, T. irroratum A.R.Bean, T. modestum A.R.Bean and T. sagittatum A.R.Bean) are newly described; full descriptions, distribution maps and
illustrations are provide...
In March 2017, more than 100 Lagenophora Cass. herbarium specimens belonging to the Muséum national d'Histoire naturelle, Paris, were destroyed while en route to the Queensland Herbarium, Australia. A lectotype is here selected for Lagenophora sundana Miq.
Labichea mulliganensis A.R.Bean, confined to Mount Mulligan in northern Queensland, is described as new. Illustrations are provided, as are notes on its affinities and conservation status. A key to the identification of all Queensland species of Labichea is included.
Pimelea section Epallage is revised for Queensland and comprises 24 species, with eight species newly described, viz. P. approximans A.R.Bean, P. chlorina A.R.Bean, P. confertiflora A.R.Bean, P. fugiens A.R.Bean, P. gigandra A.R.Bean, P. mollis A.R.Bean, P. plurinervia A.R.Bean and P. rupestris A.R.Bean. New combinations are P. amabilis (Domin) A.R...
A new species, Melaleuca comosa A.R.Bean is described and illustrated. It is known from a single location near Blackall in western Queensland.
Edward Macarthur Bowman was one of a select group of pioneer settlers in Queensland who had a strong scientific interest in native plants, and attempted to make a living from collecting plant specimens and seeds. In this he was unsuccessful, but recently located letters and newspaper articles, transcribed here for the first time, give an insight in...
A lectotype and epitype are here selected for Ixauchenus sublyratus Cass. We have determined that this name is applicable to a taxon of the Lagenophora gracilis species complex. RÉSUMÉ Identité et typification d'Ixauchenus sublyratus Cass. (Asteraceae). Un lectotype et un épitype sont désignés ici pour Ixauchenus sublyratus Cass. Nous avons égaleme...
Bean, A.R. & Jobson, P.C. (2017). Olearia bella A.R.Bean & Jobson and O. orientalis A.R.Bean & Jobson (Asteraceae: Astereae), two new species from Queensland. Austrobaileya 10(1): 102-112. Two species of Olearia of restricted distribution in Queensland, O. bella and O. orientalis, are described as new. They are compared with their closest relatives...
Two species of Olearia of restricted distribution in Queensland, O. bella and O. orientalis, are described as new. They are compared with their closest relatives, O. ferresii (F.Muell.) Benth. and O. macdonnellensis D.A.Cooke respectively. The new species are illustrated and their conservation status is assessed. A distribution map is provided for...
Lomandra decomposita (R.Br.) Jian Wang ter & A.R.Bean is described, illustrated and differentiated from similar and related taxa. The known distribution of the newly reinstated species is from Charters Towers to the islands of the Torres Strait in Queensland. A conservation status of Least Concern is proposed.
A lectotype and epitype are here selected for Ixauchenus sublyratus Cass. We have determined that this name is applicable to a taxon of the Lagenophora gracilis species complex. RÉSUMÉ Identité et typification d'Ixauchenus sublyratus Cass. (Asteraceae). Un lectotype et un épitype sont désignés ici pour Ixauchenus sublyratus Cass. Nous avons égaleme...
Five species of Lagenophora occur in Queensland. These
include the two named species L. gracilis Steetz, a widespread and variable species; and L. stipitata
(Labill.) Druce, known only in Girraween National Park. Three new species are here described; L.
queenslandica Jian Wang ter & A.R.Bean, L. fimbriata Jian Wang ter & A.R.Bean and L. brachygloss...
Eleven Solanum species from the Solanum ferocissimum group are newly described for New Guinea: S. arachnoides A.R.Bean, S. banzicum A.R.Bean, S. exemptum A.R.Bean, S. invictum A.R.Bean, S. malignum A.R.Bean, S. oomsis A.R.Bean, S. ortivum A.R.Bean, S. petilum A.R.Bean, S. phoberum A.R.Bean, S. pluriflorum A.R.Bean and S. scolophyllum A.R.Bean. Five...
Bean, A.R. (2016). Frederick Hamilton Kenny (1859‒1927), an Australian plant collector of note. Austrobaileya 9(4): 546–559. A biography of Frederick Hamilton Kenny, a significant Australian plant collector in the early 20 th century, is presented. Kenny was a medical doctor and surgeon, but also a keen amateur naturalist. He lived and worked in nu...
Bean, A.R. (2016). Two new species of Solanum (Solanaceae) from the Northern Territory, Australia. Austrobaileya 9(4): 524–533. Two new species, Solanum ultraspinosum A.R.Bean and S. apodophyllum A.R.Bean are described and illustrated, and maps of their distribution provided. Both are related to Solanum clarkiae Symon. A key to the species comprisi...
Five species are here recognised for the Solanum petrophilum F.Muell. complex. They are the previously named S. petrophilum F.Muell. and S. eardleyae Symon, and three newly described species: S. lobatum A.R.Bean, S. osteocarpum A.R.Bean and S. pallidifolium A.R.Bean. All taxa are illustrated, described, and their distributions mapped. A key is prov...
The genus Anisomeles R.Br., naturally occurring from southern Asia to northern Australia,
is taxonomically revised, and its phylogenetic placement is discussed. 26 species are recognised,
of which 18 species are newly described, with one new combination. Diagnostic morphological
characters are discussed, and a comprehensive identification key to th...
Olearia cuneifolia A.R.Bean & M.T.Mathieson (Asteraceae: Astereae), is described, illustrated and compared to related taxa. It has a restricted distribution in the Mungallala area of southern Queensland. A conservation status of Endangered is recommended.
Bean, A.R. (2015). A conspectus of Polyscias J.R.Forst. & G.Forst. in Queensland, Australia. Austrobaileya 9(3): 445-456. The 12 indigenous species of Polyscias in Queensland are enumerated, and their nomenclature, distribution and habitat are discussed. Distribution maps and an identification key are provided. Specimen citations of Polyscias zippe...
The new species Ptilotus senarius A.R.Bean, recorded only between Georgetown and Croydon, is described, illustrated, and differentiated from similar taxa. A conservation status of Presumed Extinct is proposed.
In July and August 1918, Cyril Tenison White spent four and a half weeks in Papua New Guinea, during which time he went on two extended excursions and collected 829 botanical specimens, of which at least 722 are known to be extant, representing 562 species. 32 new species and two new varieties were named from specimens collected by White during his...
Four new species of the Solanum ellipticum species group are described, viz. Solanum adoxum A.R.Bean, S. capitaneum A.R.Bean, S. prolatum A.R.Bean and the new combination Solanum chillagoense (Domin) A.R.Bean. The phenetic relationships between these and other taxa of the S. ellipticum group are clarified by the results of a morphometric analysis,...
Pluchea tenuis A.R.Bean is newly described and illustrated. It is known from just one locality on Cape York Peninsula where it is associated with mound springs.
Pluchea macdonnellensis Albr. & A.R.Bean is described and illustrated, with notes on distribution, habitat and relationships with other species of Pluchea.
Potentilla nanopetala A.R.Bean, a new endemic Australian species, is described, illustrated, and its distribution mapped. The name Argentina anserina (L.) Rydb. (formerly Potentilla anserina L.) is accepted. Potentilla crantzii (Crantz) Beck ex Fritsch is removed from the Australian flora; and the genus Fragaria L. reinstated for Australia. A key t...
The informal taxonomic subgroup typified by Solanum sturtianum F.Muell. comprises those species that have stellate hairs, black mature fruits with a thin brittle pericarp, and brown to black seeds. The subgroup is endemic to Australia. Ten species are recognised here: S. sturtianum is maintained, S. morrisonii Domin is reinstated, and eight new spe...
Two new subspecies of Eucalyptus tereticornis Sm., E. tereticornis subsp. rotunda A.R.Bean and E. tereticornis subsp. basaltica A.R.Bean, are described and illustrated. A key to the four subspecies of E. tereticornis is provided, together with a distribution map for the new taxa.
Sphaeromorphaea DC. and Ethuliopsis F.Muell., historically included in Epaltes Cass., are reinstated and taxonomically revised. Sphaeromorphaea occurs in Australia, Indian Subcontinent, Indochina, China, eastern Asia, Malesia, Papuasia, northwestern Pacific and southwestern Pacific, and comprises six species. Three new species; S. ephemera A.R.Bean...
Three new species of Pluchea Cass. are described; P. longiseta A.R.Bean from northern Western Australia, P. mesotes A.R.Bean from the Northern Territory, and P. alata A.R.Bean from central Queensland. All species are illustrated and distribution maps are provided. A revised key to the Australian of Pluchea species is provided.
The taxonomy of Solanum echinatum and its allies is revised. Seven species are enumerated, all endemic to tropical Australia, including four new species: S. fecundum sp. nov., S. lapidosum sp. nov., S. medicagineum sp. nov. and S. rhaphiotes sp. nov. Solanum longissimum is reduced to synonymy, and S. wilkinsii is reinstated. Illustrations are provi...
Stylidium elachophyllum is described as new. It is an ephemeral herb known only from the Hann Tableland in northern Queensland, with a morphological affinity to S. fissilobum F.Muell., S. prophyllum Lowrie & Kenneally and S. oviflorum A.R.Bean. The species is illustrated and diagnosed against related taxa. Its conservation status is assessed and a...
Botanical surveys from prior to 1949 documented the plant species composition of the mulga (Acacia aneura) communities in the eastern Mulga Lands Bioregion of Queensland, Australia. These surveys recorded 282 vascular plant species
including 268 natives and 14 exotics. Since 1949, mulga communities have experienced considerable modification throug...
The endemic Australian genus Nyssanthes is revised, with the recognition of four species, N. erecta R.Br., N. diffusa R.Br., N. impervia A.R.Bean sp. nov. and N. longistyla C.H.Mill. sp. nov. All species are illustrated, and distribution maps provided. Notes for each species are given on conservation status, distribution and habitat, and an identif...
Pterocaulon section Monenteles, comprising all species of the genus from Asia, Malesia, Australia and Melanesia, is revised. 14 species are recognised, including the six new species, P. brachyanthum A.R.Bean, P. ciliosum A.R.Bean, P. discolor A.R.Bean, P. paradoxum A.R.Bean, P. tricholobum A.R.Bean and P. xenicum A.R.Bean, and one new combination,...
The new species Pluchea punctata A.R.Bean and P. xanthina A.R.Bean are described from Queensland, with the latter replacing the misapplied name P. dioscoridis (L.) DC. Illustrations, a distribution map, and notes on the habitat and conservation status are provided for each new species. A note is given regarding the apparent sexual dimorphism of Plu...
The Solanum ellipticum species group is endemic to Australia, and is here considered to comprise 13 species. Solanum lithophilum F.Muell. and S. cleistogamum Symon are reinstated. Solanum aridicola A.R.Bean, S. callosum A.R.Bean, S. emmottii A.R.Bean and S. unispinum A.R.Bean are described as new. The four new species and Solanum lithophilum are il...
For many years, the classification of forest communities and their associated species from airborne data (primarily aerial photography) has largely been through manual interpretation. With the development of digital imagery, options for delineating individual crowns and identifying these to species have become available but automated techniques for...
The taxonomy of the genus Alphitonia Endl. is reassessed for Australia, and five species are recognised. Alphitonia oblata A.R.Bean sp. nov. is described, illustrated and diagnosed against related taxa. Descriptions, distribution maps and miscellaneous notes are provided for all species. Lectotypes are selected for A. excelsa (Fenzl) Benth., A. pet...
A new genus in the family Amaranthaceae, Kelita A.R.Bean, is described. The new genus belongs to Subtribe Aervinae Endl. and is related to Ptilotus R.Br. and Omegandra G.J.Leach & C.C.Towns. It differs from these genera by the denticulate leaf margins, the zygomorphic flowers, the uncinate awns on the tepals, and the thick-walled fruits. The genus...
Eleven names in Eucalyptus (Myrtaceae) are here typified, and the typification is clarified or corrected for a further three names. The names dealt with in this paper include E. tessellaris var. dallachiana, E. brownii, E. cambageana, E. cloeziana, E. dawsonii, E. intertexta, E. malacoxylon, E. melanophloia, E. morrisii, E. nova-anglica, E. raveret...
Twelve names in Eucalyptus (Myrtaceae) are here lectotypified, and the typification is clarified for a further three names. The species involved are indigenous to eastern Australia. Full discussion of relevant type specimens is given, and other nomenclatural notes are included.
Three new species belonging to Stylidium subgenus Andersonia; S. exiguum A.R.Bean, S. notabile A.R.Bean and S. osculum A.R.Bean, are described from the Northern Territory. One new species belonging to Stylidium section Floodia Mildbr., S. centrolepoides A.R.Bean, is described from Queensland. All species are illustrated and a distribution map provi...
A new subspecies, Eucalyptus sideroxylon subsp. improcera A.R.Bean, is described. It is diagnosed against the typical subspecies, and illustrations are provided. It is confined to a small area of the Barakula State Forest in southern Queensland.
The new species Crudia abbreviata A.R.Bean is described and illustrated. It is endemic to Cape York Peninsula, Queensland and considered to be closely related to C. blancoi Rolfe. A key to the Australian species of Crudia is provided.
Solanum pisinnum A.R.Bean sp. nov. and S. orgadophilum A.R.Bean sp. nov. from central Queensland are described and illustrated. Notes on affinities, conservation status, distribution, etymology, habitat and phenology are provided.
Eleven names in Eucalyptus are newly typified, and the typification is clarified for a further four names. The species involved are all indigenous to Queensland or New South Wales. Full discussion of relevant type specimens is provided, and other nomenclatural notes are included. Information provided in recent nomenclatural references for eucalypts...
The original material of Eucalyptus ambigua DC. is conspecific with the type of E. nitida Hook.f. The name E. ambigua is older by almost 30 years, and is therefore the correct name for the Smithton Peppermint. Bentham and Maiden both correctly determined the identity of E. ambigua, but the name has been ignored for the last 100 years. Eucalyptus am...
Solanum sulphureum A.R.Bean sp. nov. is described and illustrated. It is related to S. brownii and other species in the informal S. macoorai group, and is apparently confined to lowland rainforest in the Taree district on the mid north coast of New South Wales. Its conservation status and relationships are discussed.
Four species are recognised in the Eastern grey box group, E. moluccana Roxb., E. albens Benth., E. woollsiana R.T.Baker and E. microcarpa (Maiden) Maiden. E. pilligaensis is reduced to synonymy under E. woollsiana. The taxonomic status of E. microcarpa is discussed. The nomenclature of E. woollsiana is discussed and lectotypes are selected for E....
A new species of Homoranthus is described, illustrated and diagnosed against related species. It is of very restricted distribution near Mundubbera in south-eastern Queensland.
A synopsis is presented for the 31 Ptilotus species accepted by the author, and indigenous in eastern Australia (Queensland, New South Wales, Victoria and Tasmania). Notes on typification, distribution and habitat are given. Two new combinations are made, and 24 names are here formally reduced to synonymy. A lectotype is chosen for P. calostachyus...
The new species, Solanum succosum from Queensland and Northern Territory, is described and illustrated. It is closely allied to S. chippendalei, for which a comparative description is provided. Distribution maps are presented for both species and S. succosum is illustrated.
The name Ammannia triflora Benth. is reinstated for the species referred to in the Flora of Australia as A. pubiflora (Koehne) Hewson.
Erythrina L. is represented in Australia by the following native or naturalised taxa; *E. caffra Thunb., *E. crista-galli L., E. fusca Lour., E. insularis F.M.Bailey, E. numerosa A.R.Bean, *E. speciosa Andrews, *E. × sykesii, E. variegata L., E. vespertilio subsp. biloba (F.Muell.) A.R.Bean and E. vespertilio Benth. subsp. vespertilio. Erythrina fu...
An examination of Floras and related literature from various countries of the world has revealed a wide range of interpretations and concepts for indigenous plants. Nevertheless, an indigenous plant species has been universally defined as one that was not deliberately or accidentally introduced by man. An important recent addendum to the ‘indigenou...
Questions
Questions (3)
This was collected at Townsville, Queensland, Australia. It does not match any of the native species known from Townsville or elsewhere in Queensland, so perhaps it is an introduced species?
I am aware of the common aquatic plant Veronica anagallis-aquatica. But I have seen, on "The Plant List" website, a species called Veronica anagallis listed as an accepted name. Does such a species exist, or is this a contraction or abbreviation for Veronica anagallis-aquatica?
Growing near Emerald, Queensland, Australia. Suspected Amaranthaceae. Herbaceous, maximum height 1.5 metres; leaves glabrous, alternate; inflorescence slender, spike-like; all flowers bisexual, glabrous, in contracted dichasial cymes. Capsule indehiscent, same length as perianth; fertile stamens 5; stigmas 2; ovules 2; seeds solitary, black. Growing on heavy clay soil.