Stephen J Richards

Stephen J Richards
South Australian Museum · Herpetology

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241
Publications
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4,560
Citations
Citations since 2017
45 Research Items
1711 Citations
20172018201920202021202220230100200300
20172018201920202021202220230100200300
20172018201920202021202220230100200300
20172018201920202021202220230100200300

Publications

Publications (241)
Article
Full-text available
The mountains of New Guinea are home to species-rich but poorly understood communities of stream or torrent-breeding pelodryadid treefrogs. Here we describe a new species of moderately sized torrent-breeding Litoria from the mountains of Papua Province, Indonesia. The new species is most similar to Litoria dorsivena but differs from that species in...
Article
Full-text available
The Islands of East Melanesia have a unique and highly endemic frog fauna derived entirely from overseas colonisation events. Within East Melanesia New Britain is a notable centre of frog diversity and endemism, with at least 15 endemic species, mostly in the ceratobatrachid genus Cornufer . Here we describe the first endemic pelodryadid treefrog f...
Article
Full-text available
Identifying hotspots of biological diversity is a key step in conservation prioritisation. Melanesia—centred on the vast island of New Guinea—is increasingly recognised for its exceptionally species-rich and endemic biota. Here we show that Melanesia has the world’s most diverse insular amphibian fauna, with over 7% of recognised global frog specie...
Article
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Cryptic ecologies, the Wallacean Shortfall of undocumented species' geographical ranges and the Linnaean Shortfall of undescribed diversity, are all major barriers to conservation assessment. When these factors overlap with drivers of extinction risk, such as insular distributions, the number of threatened species in a region or clade may be undere...
Article
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We describe a new species of the asterophryine microhylid genus Xenorhina from the lowlands of northwestern Papua New Guinea. It is a medium-sized species (SUL of two males 29.2 and 29.9 mm; of four females 29.9–33.0 mm) that can be distinguished from congeners by having a single short, triangular odontoid spike (palatal tooth) on each vomeropalati...
Article
Full-text available
We describe six new species of the microhylid frog genus Xenorhina from the southern slopes of Papua New Guinea’s central cordillera and adjacent lowlands, based on a combination of morphological (including osteology) and bioacoustic features. All of the new species are fossorial or terrestrial inhabitants of tropical rainforest habitats and belong...
Article
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Australia and New Guinea (together referred to as Sahul) were linked by land for much of the late Tertiary and share many biotic elements. However, New Guinea is dominated by rainforest, and northern Australia by savannah. Resolving patterns of biotic interchange between these two regions is critical to understanding the expansion and contraction o...
Article
We show that the Wotjulum frog, Litoria watjulumensis (Copland, 1957), comprises two deeply divergent mitochondrial DNA lineages that are also reciprocally monophyletic for a nuclear gene locus and have discrete distributions. The taxa are differentiated in multivariate analysis of shape but show no appreciable differences in colour and pattern. Th...
Article
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We describe a new species of groundsnake of the genus Stegonotus (Colubridae) from the Purari River basin in Gulf Province, Papua New Guinea. The new species can be most readily distinguished from all other New Guinean Stegonotus by its unique dorsal colour pattern which consists of a dark head and creamy-white anterior one third to two thirds of t...
Article
We describe a new species of large green treefrog from southern Papua Province, Indonesia. Litoria lubisi sp. nov. is placed in the L. infrafrenata Group based on its size, colouration and configuration of hand webbing, but differs from other members of this group in aspects of body size and proportions, extent of hand webbing, colouration and male...
Article
Full-text available
We describe a new species in the Australopapuan pelodryadid frog genus Litoria from upper hill forest (940 m a.s.l.) on the northern slopes of Papua New Guinea’s central cordillera. The new species is moderately small (male body length = 31.9–35.1 mm) and slender (head width/body length = 0.29–0.30), with extensive golden-yellow markings ventrally....
Article
We describe a new species of gecko in the Lepidodactylus novaeguineae Group from Salawati Island, West Papua Province, Indonesia. The new species can be distinguished from all congeners by a unique combination of aspects of body size, shape, colouration, and scalation. The holotype and only known specimen is a mature male with a snout-vent length o...
Article
Systematic investigations of vertebrate faunas from the islands of Melanesia are revealing high levels of endemism, dynamic biogeographic histories, and in some cases surprisingly long evolutionary histories of insularity. The bent-toed geckos in the Cyrtodactylus sermowaiensis Group mainly occur in northern New Guinea and nearby islands, however a...
Article
New Guinea has been considered both as a refuge for mesic rainforest-associated lineages that contracted in response to the late Cenozoic aridification of Australia and as a centre of biotic diversification and radiation since the mid-Miocene or earlier. Here, we estimate the diversity and a phylogeny for the Australo-Papuan forest dragons (Sauria:...
Article
We describe a new species of bent-toed gecko in the genus Cyrtodactylus from hill forest in Southern Highlands Province, Papua New Guinea. Cyrtodactylus manos sp. nov. can be distinguished from all congeners in New Guinea by its small size in combination with aspects of colouration and body and tail scalation. The new species adds to the growing nu...
Article
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The amphibian chytrid fungus Batrachochytrium dendrobatidis (chytrid) has caused the most widespread, disease‐induced declines and extinctions in vertebrates recorded to date. The largest climatically suitable landmass that may still be free of this fungus is New Guinea. The island is home to a sizeable proportion of the world's known frog species...
Article
New Guinea is home to the world’s most diverse insular frog biota, but only a small number of taxa have been included in genetically informed assessments of species diversity. Here we describe two new species of New Guinea treefrog in the genus Litoria that were first flagged during assessments of genetic diversity (DNA barcoding) and are currently...
Article
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A new species of the dicroglossid frog genus Limnonectes is described from recent and historical museum specimens collected in central and southern Laos and northeastern Thailand. Limnonectessavansp. nov. has males that bear a caruncle on top of the head, and most closely resembles L.dabanus from adjacent southern Vietnam and eastern Cambodia. Howe...
Article
A small number of treefrog species (Litoria) from Melanesia are unusual amongst Anura in having distinctive fleshy rostral spikes. Here, we first present an extended description for Litoria pronimia Menzies, a small species that is widespread along the southern edge of the Central Cordillera of New Guinea, and in which males have a long and erectil...
Article
Full-text available
Bironides ypsilon sp. nov. and Nannophlebia ballerina sp. nov. from PNG 331 Odonatologica 46(3/4) Abstract. Two new species of stream-dwelling libellulid dragonflies are described from Gulf Province, Papua New Guinea. Bironides ypsilon sp. nov. and Nannophlebia ballerina sp. nov. are both small black and yellowish green dragonflies that appear to b...
Article
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A new species of damselfly, Rhyacocnemis gassmanni sp. n. from Papua New Guinea, is described and illustrated from both sexes with notes on its habitat and habits. It represents the fourth species of an enigmatic genus, known from only a handful of specimens. The placement of the new species is problematical and is discussed. Introduction The zygop...
Article
Two new species of the genus Idiocnemis Selys, 1886 from southern Papua New Guinea are described: Idiocnemis lakekamuensis sp. nov. from the Lakekamu Basin and I. milou sp. nov. from Lakekamu and the Kikori River lowlands. Males and females are illustrated and compared with other species of the Idiocnemis bidentata group. Both new species are known...
Article
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A series of photographs of the recently described Star Mountains Worm-eating Snake, Toxicocalamus ernstmayri O'Shea et al., 2015, taken at the Ok Tedi Mine in the Star Mountains, North Fly District, Western Province, Papua New Guinea, represents only the second record of this poorly-known species. Toxicocalamus ernstmayri was hitherto only known fr...
Article
The published repository for the holotype of Pseudagrion Jumipennis Polhemus, Michalski & Richards, 2008, is corrected to the South Australian Museum, Adelaide, Australia, not the 'Australian Museum of Natural History, Sydney' as stated in the original description. © 2018 Entomological Society of Queensland. All rights resrerved.
Article
Full-text available
A new species of damsel fly, Nososticta moginae sp. n. from Gulf Province, Papua New Guinea, is described from both sexes and its affinities are discussed. It represents the 80th species of the genus and is currently known only from a single location in the Kikori River Basin, A description of the hitherto unknown female of the morphologically simi...
Article
Regions with complex geological histories present a major challenge for scientists studying the processes that have shaped their biotas. The history of the vast and biologically rich tropical island of New Guinea is particularly complex and poorly resolved. Competing geological models propose New Guinea emerged as a substantial landmass either duri...
Article
We describe a new species of Cornufer, subgenus Batrachylodes, from high-elevation forests of New Britain Island in the Bismarck Archipelago of Eastern Melanesia. The new species, Cornufer exedrus sp. nov., is a biogeographically disjunct member of the Batrachylodes clade, representing the first record of the subgenus from outside of the Solomon Ar...
Article
Two new microhylid frogs in the genera Choerophryne and Oreophryne are described from the Foja Mountains in Papua Province of Indonesia. Both are small species (males 15.9 - 18.5 mm snout-urostyle length [SUL] and 21.3 - 22.9 mm SUL respectively) that call from elevated positions on foliage in primary lower montane rainforest. The new Choerophryne...
Article
Full-text available
To ensure that the name Papuargia stueberi luciedecknerae, described (2017) in Odonatologica 41: 283-291, is available, the type repository, inadequately presented in the original description, is stated along with a diagnosis of the subspecies.
Article
Full-text available
The island of New Guinea has been identified as biologically megadiverse but many taxa are still poorly known. This is especially the case for many of the island’s snakes, which by their very nature can be difficult to collect and study. Here we examine the phylogenetic and phylogeographic structure of a poorly studied snake genus, Stegonotus, focu...
Article
A second species of the hitherto monotypic New Guinean genus Papuargia Lief- tinck, 1938 and a new subspecies of P. stueberi Lieftinck, 1938 are described from Papua New Guinea. These are P. brevistigma sp. nov. and P. stueberi luciedecknerae ssp. nov. The char- acters which define the genus are discussed with special reference to the labium and pe...
Article
Full-text available
We describe three new species of the microhylid frog genus Choerophryne from the mountains and foothills of southern and northeastern Papua New Guinea. All three species lack elongated snouts and all are arboreal, calling from elevated perch sites between ~1 and 10 m above the forest floor. Advertisement calls and habitat preferences are described...
Article
Full-text available
Aims Mountain ranges in the tropics are characterised by high levels of localised endemism, often-aberrant evolutionary trajectories, and some of the world’s most diverse regional biotas. Here we investigate the evolution of montane endemism, ecology and body size in a clade of direct-developing frogs ( Choerophryne, Microhylidae) from New Guinea....
Data
Appendix S3 Treefile for chronogram estimated in BEAST with ancestral states.
Data
Appendix S1 Supplementary tables and figures. Specimen numbers, locality information and GenBank accession numbers for Choerophryne specimens included in analyses (Table S1); GenBank accession details for outgroup samples (Table S2); genetic distance data for species and candidate lineages (Table S3); and summary data on body, elevational distribut...
Data
Appendix S2 BEAST input file for ancestral state analyses
Preprint
Full-text available
Aims. Mountain ranges in the tropics are characterised by high levels of localised endemism, often-aberrant evolutionary trajectories, and some of the world’s most diverse regional biotas. Here we investigate the evolution of montane endemism, ecology and body size in a clade of direct-developing frogs ( Choerophryne, Microhylidae) from New Guinea....
Preprint
Full-text available
Aims. Mountain ranges in the tropics are characterised by high levels of localised endemism, often-aberrant evolutionary trajectories, and some of the world’s most diverse regional biotas. Here we investigate the evolution of montane endemism, ecology and body size in a clade of direct-developing frogs ( Choerophryne, Microhylidae) from New Guinea....
Preprint
Full-text available
Aims. Mountain ranges in the tropics are characterised by high levels of localised endemism, often-aberrant evolutionary trajectories, and some of the world’s most diverse regional biotas. Here we investigate the evolution of montane endemism, ecology and body size in a clade of direct-developing frogs ( Choerophryne, Microhylidae) from New Guinea....
Article
Full-text available
The East Melanesian Islands have been a focal area for research into island biogeography and community ecology. However, previously undescribed and biogeographically significant new species endemic to this region continue to be discovered. Here we describe a phylogenetically distinct (~20% divergence at the mitochondrial ND2 gene) and biogeographic...
Article
Full-text available
Gynacantha nuda sp. nov. is described based on a male from Southern Highlands Province, Papua New Guinea. It is the largest known species of the genus to be reported from New Guinea. Characters of the adult male are illustrated, the affinities of the new species are discussed, and a key is presented to males of the Gynacantha species recorded from...
Article
Two new species of the microhylid genus Oreophryne are described on the basis of material collected from the Kikori River Basin in southern Papua New Guinea. Both species belong to the group of Oreophryne that have a ligamentous connection between the procoracoid and scapula. One of the new species is small (snout-urostyle length in males 19.7 - 23...
Article
Idiocnemis schorri sp. nov. is described from the Hindenburg and Muller Ranges and the Kikori River Basin of southern Papua New Guinea. The new species differs from all congeners by, among other characters, a unique colour pattern on the thorax. Characters of males and females are illustrated and compared to those of similar species from the Idiocn...
Article
Full-text available
We describe a new species of treefrog from northern Australia. Litoria bella sp. nov. is morphologically and genetically most similar to frogs in the L. gracilenta and L. chloris groups but is distinguished from all members in these groups by a combination of a moderately large male body size (34.5-41.8 mm SVL), near-immaculate green dorsum, orange...
Article
Full-text available
We describe a striking new species of the microhylid frog genus Mantophryne from Woodlark Island in Milne Bay Province, Papua New Guinea. It is most similar to M. lateralis but is distinguished from that species by its more slender body, longer shanks, larger discs on the toes, and unique advertisement call. Most known specimens had, in life, a str...
Article
Pseudagrion woodlarkensis sp. nov., a new damselfly species from Woodlark Island, Papua New Guinea, is described and male and female characters are illustrated. The new species differs from all regional congeners by having a predominantly yellow thorax with bold black stripes, and males can be clearly distinguished from all Papuan Pseudagrion speci...
Article
Full-text available
The diverse biota of New Guinea includes many nominally widespread species that actually comprise multiple deeply divergent lineages with more localised histories of evolution. Here we investigate the systematics of the very large geckos of the Cyrtodactylus novaeguineae complex using molecular and morphological data. These data reveal two widespre...
Article
Three distinctive new species of Papuagrion Ris, 1913 are described from a high altitude area (1,770–1,820 m a.s.l.) at the base of the Hindenburg Wall, Western Province, Papua New Guinea. These are P. chrysosoma sp. nov., P marijanma- toki sp. nov. and P. tydecksjuerging sp. nov.; all type material is deposited in the South Australian Museum (SAMA...
Article
We describe a new species in the recently erected microhylid frog genus Pseudocallulops, increasing the number of species in this genus from two to three. The new species differs from congeners in body proportions, color and advertisement calls which consist of a series of 4-5 soft, distinctly pulsed “creaking” notes. It is presently known only fro...
Article
Full-text available
The males and, when available, females of six new species of the damselfly genus Nososticta are described from the upper Fly, Strickland, and Kikori River basins in southern Papua New Guinea. They are Nososticta caelestis sp. nov. (♂ holotype, 10-viii-2014), Nososticta chrismullerisp. nov. (<3 holotype, 04-viii-2013), Nososticta makrodon sp. nov. h...
Article
Full-text available
We describe two new species of small microhylid frogs in the genus Choerophryne from the northern slopes of Papua New Guinea's central cordillera. Choerophryne epirrhina sp. nov. can be distinguished from congeners by the combination of moderately small size (SUL 14.9-15.0 mm), distinctly elongated snout (OHG/SUL 0.09-0.10), first finger without ex...
Article
Full-text available
Two new species of Teinobasis and one new species of Nososticta from the Hindenburg Wall region of western Papua New Guinea are described and illustrated. They are Teinobasis cuneata sp. nov. (Holotype SAMA 07-001421), Teinobasis flavolineata sp. nov. (Holotype SAMA07-001422), and Nososticta oculata sp. nov. (Holotype SAMA 07-001424). The new Teino...
Article
Full-text available
A new genus and species belonging to the damselfly subfamily Idiocnemidinae from Papua New Guinea, Macrocnemis gracilis gen. nov. sp. nov. is described and illustrated. It is the largest known member of the Papuan idiocnemidine radiation, and its affinities to existing genera remain unclear. The new taxon is currently known with certainty only from...
Book
Full-text available
The relatively remote islands of Manus and Mussau, located in the northern portion of the Bismark sea have been long identified as key biodiversity areas in Papua New Guinea and within greater Melanesia. Manus Island has long been known for its endemism and relatively intact forest, while Mussau Island, although relatively unstudied, has been recog...
Article
Full-text available
The males and, when available, females of ten new species and subspecies of Nososticta are described from the Papuan region. They are: Nososticta caerulea sp. nov. (♂ holotype: Papua New Guinea, Upper Sepik Basin, 09-vi-2010, ◃ described), Nososticta finisterrae satisbona ssp. nov. (♂ holotype: Papua New Guinea, Goodenough Island, 26-X-1953), Nosos...
Book
Full-text available
The Hindenburg Wall, along with the Muller Range and Nakanai Mountains, is a part of a proposed UNESCO World Heritage Site called The Sublime Karst of Papua New Guinea (Hamilton-Smith 2006). This survey document reports on a biodiversity assessment undertaken by the Wildlife Conservation Society Papua New Guinea (WCS), financed by the Papua New Gui...
Article
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We present a near comprehensive, densely sampled, multilocus phylogenetic estimate of species relationships within the anuran family Ceratobatrachidae, a morphologically and ecologically diverse group of frogs from the island archipelagos of Southeast Asia and the South-West Pacific. Ceratobatrachid frogs consist of three clades: a small clade of e...
Article
Full-text available
Despite considerable progress in unravelling the phylogenetic relationships of microhylid frogs, relationships among subfamilies remain largely unstable and many genera are not demonstrably monophyletic. Here, we used five alternative combinations of DNA sequence data (ranging from seven loci for 48 taxa to up to 73 loci for as many as 142 taxa) ge...
Article
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Drepanosticta machadoi sp. nov. (Holotype ♂: Dablin Creek, Hindenburg Range) from Papua New Guinea is described. The new species is a predominantly black damselfly, the male with four pale/bright pattern elements on each side of the synthorax, dorsum of segments 9 and 10 largely bright blue, and a uniquely shaped posterior lobe of the pronotum whic...