Stephen J Maxwell

Stephen J Maxwell
Verified
Stephen verified their affiliation via an institutional email.
Verified
Stephen verified their affiliation via an institutional email.
  • AdvDipA, BA, BA/BSc, MSc, MPhil, PhD
  • Doctorial Student at James Cook University

About

82
Publications
59,382
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243
Citations
Introduction
I have a passion for taxonomic practice, and have a great deal of affection for the classics. Currently I am wallowing in governance structures of universities and their decline from which I am aiming to achieve a second PhD. I hold 4 undergrad degrees (majors: Ancient Greek literature/ ancient & modern history/ sociology/evolution/ biodiversity/ geomorphology), and 2 masters, a MSc (zoology and ecology) and MPhil (research -taxonomy), as well as a PhD (Systematics).
Current institution
James Cook University
Current position
  • Doctorial Student
Additional affiliations
January 2023 - present
James Cook University
Position
  • Post Grad
Description
  • I am a Doctorial Candidate.
March 2009 - November 2012
Academic Escorts
Position
  • Consultant
Description
  • Convening academic writing workshops, tutoring (Contracted)
July 2007 - December 2008
University of New England
Position
  • Fellow
Description
  • Administration, mediation, OH&S, academic mentoring (Contracted)
Education
February 2019 - December 2022
James Cook University
Field of study
  • Systematics
February 2016 - November 2018
James Cook University
Field of study
  • Research -Taxonomy
February 2014 - November 2015
James Cook University
Field of study
  • Zoology/Ecology

Publications

Publications (82)
Article
Full-text available
A new species of volute, Cymbiola distructa n. sp., has been collected by the Arafura Sea Prawn Fishery, northern Australia. This species can be distinguished from others in Cymbiola by its larger size, angled spire, and the outer lip form. C. distructa lacks the posteriorly inwardly recurving lobe often associated with the outer lip in adults of C...
Article
Full-text available
This paper presents a new species of Amoria from the Ashmore Reefs, Western Australia, a species which has been long been confused by collectors with Amoria spenceriana (Glatiff, 1908) of the Ashmore Banks, off Far North Queensland. Amoria textilata n. sp. shows differences in morphological characteristics and geographic separation from similar spe...
Conference Paper
Full-text available
This study reports on useful tracking components of the Qualtrics survey instrument, and their application to higher degree research and survey validity. Qualtrics captures the unique survey responses from an individual as coming from a specific device or computer with its own unique internet protocol (IP) address. This IP address is expressed as a...
Article
Full-text available
This study advances knowledge in the AI field. It provides deep insight into current industry generative AI inclusion systems. It shows both literature and practical leading industry operations can link, overlap, and complement each other when it comes to AI and understanding its complexities. It shows how to structurally model and link AI inclusio...
Article
Full-text available
There is concern over potential toxic elements (PTEs) impacting river ecosystems due to human and industrial activities. The river’s water, sediment, and aquatic life are all severely affected by the release of chemical and urban waste. PTE concentrations in sediment, water, and aquatic species from river ecosystems are reported in this review. Amo...
Article
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The invasive Asian clam species, Corbicula fluminea, has significant ecological and societal implications at both local and international levels due to its nutritional aspects. C. fluminea from four urban rivers in Bangladesh exhibited negative allometric growth and degree of contamination with potentially toxic elements (PTEs), which posed a conce...
Article
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A new species of Turricula from the northwest continental shelf of Western Australia is described. Turricula infida Yao and Maxwell, nov. sp. can be differentiated from other Turricula by the sculpture of the upper whorls with its distinctive subsutural nodulations. This study indicates that further work is needed in Turricula.
Article
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This paper introduces a new species of Dominus from Indonesia, Dominus vinki nov. sp., and is compared with the recently described Dolomena likuii Zhu, 2023 and other species in the genus. Dominus vinki nov. sp. differs from other members of the Dominus complex in morphological form and range. Dominus was erroneously subjugated under Dolomena by Ra...
Article
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A review of the Relegamoria molleri (Iredale, 1936) complex was undertaken using newly sourced material from a number of private research collections and Atlas of Living Australia ("ALA") database, which contains the Australian and Queensland Museums holdings and records. After a detailed review of several hundred specimens including morphological...
Article
Full-text available
Lambis vertriesti ranges from the Philippines into Indonesia. Significant morphological features differentiate L. vertriesti from Lambis lambis and the synonymisation of those species by Alf and Wienke (2023) is unwarranted. Lambis species in general are highly plastic in form, and therefore comparative studies should be based on type material for...
Conference Paper
Full-text available
The shift towards a corporate model for universities and the role of technology has shifted the power structures and operations within universities. In this paper, we examine the university governance changes and structures and how these have shifted the internal focus of those institutions. Historical arguments that the use of technology would red...
Conference Paper
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The COVID pandemic has driven innovations that have made information technology all-pervasive in the lives of most individuals. This systematic review seeks to determine the key themes that reflect these advances. The demographics of the publications reviewed were also examined. A total of 84 peer-reviewed and conference proceedings publications we...
Conference Paper
Full-text available
The shift in artificial intelligence (AI) from mildly-intelligent data manipulation to complex AI that fuses global data, plus sensory capture, along with interpretation, interpersonal and environmental assessment and virtual realities systems, and that then network-collaborates, self-learns and continually coalesces with internet of everything (Io...
Conference Paper
Full-text available
The inclusion of artificial intelligence (AI) can alter firm competitiveness through data analysis and strategic planning modelling. In this paper we demonstrate that firm AI inclusion is literature supported as a stage-wise, developmental, intelligences-driven, change process. Identified benefits that AI inclusion may bring to the firm: (1) a know...
Article
Full-text available
The consumption of contaminated finfish from the polluted river channel of Turag‐Tongi‐Balu, Kamarpara site, Dhaka poses significant health hazards to humans. We used mass spectrometry on chemically digested liquid samples from five fish species from Turag‐Tongi‐Balu to estimate the concentrations of 10 elements (Cr, Mn, Ni, Cu, Zn, As, Se, Cd, Fe,...
Article
Full-text available
The genus Neodilatilabrum contains taxa that are both morphologically stable across their ranges, while others are highly plastic, making the choice of discerning characteristics critical to achieving taxonomic stability. A new species of Neodilatilabrum from Japan is presented here. Neodilatilabrum tasminae n. sp. lacks the axial shoulder rib on t...
Article
Full-text available
A new species of Miliariconus from tropical Western Australia, is described based on morphological differences with known taxa from that region. The new species differs from known Australian species in sculptural form and colour. This species further highlights the endemism of much of the new taxa that is being discovered in remote northern Austral...
Article
Full-text available
This study examined the level of perceived responsibility junior and senior psychiatric nurses have for human resources and governance in Saudi Arabia. Bullying is a significant issue in nursing and an entrenched cultural practice that highlights a failure in governance and human resource responsibilities. A total of 90 responses (43.1%) to a 5...
Article
Full-text available
As ecologically and economically, marine gastropod mollusca are highly significant in the benthic ecosystems. Among marine gastropods, Umbonium vestiarium (Linnaeus, 1758) is a key stone species in marine ecosystems. Adult Umbonium vestiarium from coastal areas in Bangladesh including Cox’s Bazar, Sonadia and the St. Martin’s Islands typically rang...
Conference Paper
Full-text available
We demonstrate that the use of visual cues in reminder emails can elicit increased response rates. Survey demographics and response rates were drawn from an ongoing survey being conducted within a not-for-profit organisation in Australia. The use of animals ("fur babies") has an impact on the younger and older demographics and those without a unive...
Thesis
Full-text available
This dissertation presents a classical revision of Strombus urceus Linné, 1758 post Abbott 1960 (Mollusca, Neostromboidae, Strombidae) and has resolved this monospecific group into twelve species. This involved a review and the presentation of novel theories in the areas of speciation, hybridisation and clade recognition. The species concept was re...
Article
Full-text available
This paper presents a new putative inter-generic Harpago chiragra x Solidistrombus latissimus hybrid from Đảo Phú Quý, Vietnam. Hybridisation is not uncommon with the Strombidae, with 12 inter-generic and 11 infra-generic hybrids known in the literature. Understanding the rates and extent of hybridisation within an organism complex facilitates a gr...
Article
Full-text available
The Molluscan family Strombidae Rafinesque, 1815 is taxonomically diverse, with a widespread global distribution. However, the population structure of many of these taxa remains enigmatic. There is a growing interest in exploring population structures within species across their distributions to understand factors affecting morphological diversity....
Article
Full-text available
This article examines the legal status of nomenclature codes and their application to denomination in certain forms of intellectual property, in particular plant breeder's rights and patents for micro‐organisms. Its objective is to show how rigidity and the absence of appeals against nomenclature decisions can have adverse effects on the rights and...
Article
Full-text available
Canarium (Canarium) incisum and Canarium (Canarium) esculentum are small members of the molluscan Strombidae family. Little is known of their population structure. Therefore, we explored this using samples from a population of each. The first sample from Corong Corong Beach, El Nido, Philippines, consisted of 81 adult C. incisum, of which 33 were f...
Article
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[This corrects the article DOI: 10.3389/fpls.2021.748562.].
Article
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In this study, the extant Doxanderina species are addressed. Species contained within Doxander are examined based on type material, and a new species from Queensland is recognised. Comparisons are made based on morphology. This revision recognises six species in the Doxander complex: D. campbellii, D. entropi, D. japonicus, D. queenslandicus nov. s...
Article
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This iconography explores the variability and ranges of members of the Ministrombus and introduces three new species bringing the total to six in the genus. Two of these species are alluded to in previous works but were never described. M. athenius is recircumscribed and the type locality is shifted to reflect this change in taxonomic position. Thi...
Article
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This paper defines the new crown clades Neoxenophoroidae new clade and Neoaporrhaoidae new clade to separate the Neostromboidae, which contains Strombidae Rafinesque, 1815, Rostellariidae Gabb, 1868, and Seraphsidae Gray, 1853, from their sister families Aporrhaidae Gray, 1850, Struthiolariidae Gabb, 1868 and Xenophoridae Troschel, 1852. There is s...
Article
Full-text available
This paper describes a new endemic species from the central Philippines, Lambis vertriesti n. sp. and this brings the total number of Lambis species to twelve. The news species can be differentiated form other Lambis by its unique morphology. The lectotypes for Lambis lambis (Linné, 1758) and Lambis millepeda (Linné, 1758) are also designated from...
Article
Full-text available
The propensity for hybridisation is well established in Strombidae. I propose that there are three forms of hybridisation: first the true hybrids; second reticulatory hybrids; and third diversificatory hybrids. The illustrated putative true hybrid between Conomurex decorus and Conomurex luhuanus is known from a single example from central Indonesia...
Article
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A new species of Muricidae, Chicoreus (Triplex) tangaroai is described from the Samoan Infraprovince. This new taxa is compared with C. (T.) thomasi (Crosse, 1872) and C. (T.) lorenzi Houart, 2009, related species in the Marquesan Molluscan Province.
Article
Full-text available
This part of the Canarium (Canarium) urceus-complex (Linnaeus, 1758) review after Abbott’s revision (Abbott, 1960) revision examines material from the Andaman Sea. At present, material from that region has been synonymised under the name Canarium (Canarium) urceus (Linnaeus, 1758). Canarium (Canarium) andamanense new species is known from the Andam...
Article
Full-text available
Modelling sexual dimorphism has important implications for understanding the evolution of size relationships among and within organisms. We present a composite model for the regulation and evolution of sex-specific inter-population shell size in a family of herbivorous marine molluscs, the Strombidae. In particular, this model postulates that gene...
Article
Full-text available
Cryptic species comprise two or more taxa that are grounded under a single name because they are more-or-less indistinguishable morphologically. These species are potentially important for detailed assessments of biodiversity, but there now appear to be many more cryptic species than previously estimated. One taxonomic group likely to contain many...
Article
Full-text available
Presented herein is the first verifiable records of Neodilatilabrum Dekkers, 2008 in Australia. The two examples of Neodilatilabrum robustum (Sowerby, 1875) come from Point Cartwright and Dingo Beach, Queensland. These specimens represent an anomaly, being morphologically similar to a localised South China Sea population. The possible modalities to...
Article
The investigation of sex bias and pseudohermaphroditism is a relatively unexplored area of research in the Neostromboidae. Here, we inform on these phenomena in a population of Gibberulus dekkersi Maxwell, Hernandez Duran, Rowell & Rymer, 2021 on Green Island, Great Barrier Reef. We collected 598 individuals, and noted the size of the cluster in wh...
Article
Full-text available
This part of the Canarium (Canarium) urceus (Linné, 1758) revision after Abbott (1960) revision examines the northwest Pacific specimens, which are currently synonymised under Canarium (Canarium) urceus (Linné, 1758). Canarium (Canarium) nipponium n. sp. is known from the Ryukyuan Subprovince, Mariana Islands and Palau. The new species differs from...
Article
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This part of the Canarium (Canarium) urceus (Linné, 1758) after Abbott (1960) revision examines the Australian species of that complex. Currently, there is one recognised species, Canarium (Canarium) orrae (Abbott, 1960), which is divided herein into two species, with the description of Canarium (Canarium) darwinense n. sp. from the Van Diemen Gulf...
Article
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Page 406, Taxonomy, line 6 : The “Seraphsidae Gray, 1853” should read “Seraphsidae Jung, 1974”. The authors thank Aart Dekkers for pointing out this error. Page 425, The type genus for Pseudoterebellinae new subfamily is “Pseudoterebellum new genus” and not “Terebellum Röding, 1798”
Article
Full-text available
This paper provides the International Code of Phylogenetic Nomenclature RegNum repository registration numbers for the clades defined in The Festivus. The definitions are based on the current understanding of the internal resolution within Stromboidae, and maybe amended as further taxa are resolved. This set of registration references reflects the...
Article
Full-text available
Abstract.—The gastropod family Strombidae has sparked the recent interest of taxonomists as early revisions of the family are re-examined, with a plethora of new species and genera being described. This has brought a greater understanding of the level of diversity within the family, which has assisted in conceptualizing its evolutionary intergeneri...
Article
Full-text available
This revision demonstrates that an integrated taxonomic approach to classical taxonomic practice can lead to increased internal cladistic resolution within a clade, including the recognition of new taxa at all nomenclatural levels. In particular, this revision has two aims: 1) to complete an α-taxonomic revision of Seraphsidae (Stromboidea); and 2)...
Article
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A definition for differing terminal taxa in nomenclature is needed to make them practically relevant. The International Commission of Zoological Nomenclature (1999) provides for two levels: species and subspecies. At present species and subspecies are used interchangeably and arbitrarily. We examine both concepts, with a focus on practical applicati...
Article
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This study provides a checklist of the distribution and relative abundance of Strombidae from the nearshore environment of Green Island, Queensland, Australia. Historical records indicate that this island has not been surveyed for at least half a century. We used an opportunistic sighting survey method, where we walked the path of the receding tida...
Article
Abbott (1960) was too conservative in his assessment of species delimitations. Here, we evaluate the taxonomy of the marine gastropod Laevistrombus canarium (Linnaeus, 1758), and determine that there are two other extant species within this genus that are very similar, of which one occurs in the northwest Pacific and the other in the eastemmost par...
Article
Full-text available
The taxonomy of Stromboideans has, historically, been simplistic. However, recent revisions have seen new taxa introduced to distinguish relationships between species clusters. We discuss these numerous advancements in Stromboidean systematics, and describe two new subfamilies here. The key diagnostic characteristic, the basal peg on the first late...
Article
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This revision of the genus Canarium Schumacher, 1817 after Abbott (1960) advances our understanding of the phylogeny of Strombidae. Morphological characters were used to generate a phylogeny using maximum likelihood and including all of the recognised species. This resulted in the recognition of one tree, and within that tree the existing genera Ca...
Article
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This study introduces four new species within the Canarium urceus complex. Canarium daveyi nov. sp. and the sympatric C. geelvinkbaaiensis nov. sp. from the region surrounding Geelvink Bay in northeastern Indonesia, C. youngorum nov. sp. from the island of northeastern Papua New Guinea, and finally Canarium manintveldi nov. sp from the southern Sou...
Article
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Many valid species can hide buried within prior taxonomic revisions. These need to be examined before the taxonomist embarks on the challenging task of naming new taxa. In this study we reexamine the synonyms complied by Abbott (1960) under Canarium urceus (Linné, 1758), examining each taxon for morphological clarity and distinctiveness. After cons...
Article
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Canarium labiatum is a small gastropod of the Strombidae family that is commonly encountered in the inter-tidal zones of tropical Queensland, Australia, yet little is known of its population structure. A targeted survey of the Canarium labiatum population on Green Island, located near Cairns, Queensland, was conducted on 12 August 2015. Ninety adul...
Article
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This paper presents a new Altivasum found off Jurien Bay, Western Australia at 60 m. This new species expands our understanding of the distribution, and in particular extends the northern range of Altivasum, in the Southwest Marine Region. Altivasum pauladellaboscae n. sp. is more rhomboidal than A. hedleyi Maxwell and Dekkers, 2019, which is elong...
Article
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A new strombid genus (Spinatus gen. nov.) is described from the Miocene of Indonesia. Two new species of Spinatus from the middle and upper Miocene of Indonesia are described: S. echinatus spec. nov. and S. wonosariensis spec. nov. The former is selected as the type species of Spinatus. A third species, Strombus (Strombus) tjilonganensis Martin, 18...
Article
Full-text available
Strombus urceus Linné, 1758 is a gastropod species that is one of the most variable and well documented through the centuries. However, we found the present identity of Strombus urceus misleading. Abbott (1960) designated the type locality, and confirmed type specimen, based on the microfiche of the Uppsala University, Museum of Evolution Zoology S...
Article
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The resurrection of essentialist arguments, in the context of the modern evolutionary synthesis, highlights the need for a break from historical definitions of 'kind' and 'essence' in order to bring a new paradigm in which these terms are used to conceptualise and understand evolutionary processes. The definition of 'essence' is herein divided into...
Article
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The phylogeny of the American Strombidae the genus Lobatus is limited to the extant Lobatus raninus and several fossil precursors, the genera Macrostrombus, Aliger and Titanostrombus are re-installed as valid genera. The genus Persististrombus which was used as a sink for a plethora of species, is limited to the extant Persististrombus granulatus,...
Article
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This paper presents an examination of the relationships between Dolomena, Doxander, Mirabilistrombus, Neodilatilabrum and Labiostrombus using character state analysis and maximum parsimony. Hypothesized relationships are presented through the introduction of definitions that bridge the understanding of the evolutionary relationships with the applie...
Preprint
Full-text available
This preprint presents the results of an examination of the relationships between Dolomena, Doxander, Mirabilistrombus, Neodilatilabrum and Labiostrombus using character state analysis and maximum parsimony. Hypothesized relationships are presented through the introduction of definitions that bridge the understanding of the evolutionary relationshi...
Article
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Speciation as a consequence of lineage reticulation is not uncommon. A taxonomic and nomenclatural issue arises when a putative hybrid becomes established and is, therefore, in contention for species recognition. While giving a unique name to a hybrid may be acceptable under the codes that govern nomenclature, this does not address issue of whether...
Article
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This paper uses the now accepted tool of phototyping in conjunction with extensive published anecdotal evidence to describe a new carnivorous Marsupialia. This new taxon differs from all known Marsupialia in having reduced incisors and greatly developed canines. Bicingulatus ninjabearus new species is purported to be an aggressive mimic, with the p...
Article
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This paper addresses the taxonomy irregularity concerning Altivasum typicum Hedley, 1916 fide Dekkers and Maxwell, 2018 that has historically been synonymized under Altivasum flindersi Verco, 1914. We introduce the erection of Altivasum hedleyi n. sp. correcting this taxonomical conundrum that originated from the Hedley (1916) paper. Furthermore, t...
Article
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The paper recongising a new crown clade within Stromboidea contains typographic errors that occurred during copy editing: “Neostromboidea” should read “Neostromboidae” throughout. Neostromboidae is an epifamily.
Article
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This paper defines a new crown clade Neostromboidea to separate the Strombidae, Rostellariidae, and Seraphsidae from their sister families Struthiolariidae and Aporrhaidae. There is significant value to understanding evolutionary processes within Stromboidea to recognise the universal similarity in the position of the eye on the end of peduncles an...
Preprint
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A new species of Terebellum is published in Visaya. We present a proof print of the plate issued with the description.
Article
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A new species of Seraphsidae Gray, 1853 , Terebellum simoni n.sp. is described from the Phillipines
Article
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A new species of Serapsidae Gray, 1853 , Terebellum simoni n.sp. is described from the Philippines based on conchological differences with T. terebellum (Linnaeus, 1758) and especially the recently described T. hubrechti Poppe & Tagaro, 2016
Article
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Many Australian small molluscan taxa are often overlooked, falling by the wayside and therefore, and remaining relatively unknown to the wider collecting community. This paper redescribes and illustrates Retizafra intricata Hedley, 1912, a species that has not been illustrated within the literature for over a century. Many species are listed in dat...
Article
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Zootaxa 4555 (4): 491-506 https://www.mapress.com/j/zt/ Article 491 https://doi.org/10.11646/zootaxa.4555.4.2 http://zoobank.org/urn:lsid:zoobank. Abstract Here we evaluate the taxonomy of the marine gastropod genus Laevistrombus Abbott, 1960 and determine that there are five extant species within this genus, three of which occur in the southwest P...
Article
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There has been confusion on the status of Terebellum delicatum Kuroda & Kawamoto in Kawamoto & Tanabe 1956, which has often erroneously been referred to as a colour form of Terebellum terebellum Linné 1758. The taxon T. delicatum is revised, giving precision regarding authorship, and a translation of the original Japanese description with a further...
Article
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The Paraseraphs Jung 1974 is an extinct group of gastropods that existed during the Late Paleocene and Eocene of Europe and the Caribbean. P. cantamessae sp. nov. is described; it differs structurally from its sympatric sister taxon and spatiotemporally from other members of the clade.
Article
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This paper seeks to correct the erroneous taxonomy associated with the current understanding of the Altivasum flindersi (Verco, 1914) complex based on type revision, morphological differences and geographic separation. Structurally, there are three distinct species that can be differentiated, Altivasum flindersi (Verco, 1914), A. typicum Hedley, 19...
Article
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This paper highlights a putative inter-generic Conomurex x Gibberulus hybrid from the Sodwana Bay area, North Natal, Republic of South Africa. Putative hybridisation in the Strombidae is becoming ever more recognised at the intra-generic level. Much rarer are the inter-generic hybrids. The new putative hybrid indicates that inter-generic hybridisat...
Article
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One of the problems with undertaking revisions within Strombidae Rafinesque, 1815 is the tendency for chaining within the genus Strombus Linne´, 1758. Strombus is often used taxonomically as a holder for confusing taxa acting in sensu lato (s.l.). This paper addresses one of the chaining issues with Strombus with the erection of Striatostrombus gen....
Article
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A new species of Mitridae, Domiporta valdacantamessae, is described from Dingo Beach, Queensland, Australia. The shell shows similarities with other Queensland Domiporta species: D. and D. praestantissima Röding 1798, however the new species can be differentiated based on the clathrate micro-sculpture. At present, this species is only known from Qu...
Article
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In general, sex-ratios in herbivorous gastropods show a strong female bias, particularly in the family Strombidae of the Indo-Pacific. Previous studies of Indo-Pacific strombs point to an underlying female bias in most species. However, the full spatial extent and potential mechanisms driving this phenomenon are currently unknown. Therefore, we s...
Article
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A new species of Vasticardium from tropical Queensland, Australia, is described based on morphological differences with known taxa from that region. The new species differs in having acute ribs, a character it shares only with Vasticardium angulata Lamarck, 1819, which typically has coarser ribbing and which lacks colour within the shell interior.

Questions

Questions (2)
Question
The idea of a deep homology has greatly influenced the development of new taxonomic theory. In particular, Conway Morris (2015, p. 3) conceptionalisaton of evolution as “endlessly exploratory, probing the vast spaces of biological hyperspace”. However, the cleverness of Runes is the concept of convergences, these are not used as means of deconstructing Darwinism, rather stretching a band around clustering of favourable evolutionary potentialities.
The text extols through with examples of how laws of natural selection and survival of the fittest draws the evolution of taxa inextricably towards the idea of a concentrated peak of benefit. If we consider the different phyla, Conway Morris presents convergences between many. In doing so he argues for a fundamentally guiding pattern or evolutionary string upon which taxa revolve, the structural essence of Maxwell et al (2020).
If we accept Conway Morris at face value then all life has at it core a level of predestination. At this point Morris becomes dangerous to the neo-Darwinian orthodoxy of randomness in the face of mathematical impossibility, then even Dawkins said he does not do numbers, as if to ask for blind faith to the ultimate answer. So sadly I predict the ideas contained within the “Runes of Evolution” are for a time when there is balance in the debate, and where maybe we argue for the attributing the world around us to an overarching ground plan, call it intelligent design if you must, but to whom or who to attribute? May I suggest Τύχη?
Morris, S.C. (2015) The ruins of evolution: How the Universe became self-ware. Tempeton Press.
Question
"The choice of species rank over form or
subspecies for this taxon reflects a more
nuanced understanding of the role of infraspecific
ordering. We consider forms to reflect a
consistent variant within a wider population. ...
Páll-Gergely et al. (2019) argued that the rank
of subspecies was arbitrarily applied based on
“human factors”. These factors reflect the
choices that the taxonomist has to make with
regard to the differentiating of taxa in terms of
morphology, homology and the pre-existing
taxonomic hypotheses (Páll-Gergely et al. 2019).
However, Páll-Gergely et al. (2019) offer no
practical solution to how subspecies should be
identified; rather they are seeking a rule to
“prohibit taxonomic decisions resulting in
uneven subspecies rates across taxonomic
groups.” This raises the serious question of what
is a “subspecies”, and in particular once you
move away from the biological species concept
how do you demarcate between subspecies and
what is considered a full species. We argue that
subspecies should be restricted to cryptic
species, where the difference between taxa are
grounded on the unobservable genetic distance;
there is no morphological difference and
typically no test for biological isolation between
isolated populations or their clines. That is, we
argue that the rank of subspecies should be
applied to reflect genetic differences within a
species complex, rather than used to distinguish
unique taxonomic entities with observable
differences. These are species. Subspecies
therefore, is a rank that should be restricted to
cryptic species. This approach would provide a
level of taxonomic stability to the species rank
and at the same time address the issues
identified in Páll-Gergely et al. (2019). Where a
taxon can be readily identified based on
observable differences we argue, as the case of
the species herein, that the rank of species is
justified." (Maxwell and Dekkers 2019, Festivus, 51(3), 171- 176).

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