Shuo Wang

East China Normal University | ECNU · College of Life Science

Ontogenetic variation is documented within many dinosaur species, but extreme ontogenetic changes are rare among dinosaurs, particularly among theropods. Here, we analyze 19 specimens of the Jurassic ceratosaurian theropod Limusaurus inextricabilis, representing six ontogenetic stages based on body size and histological data. Among 78 ontogenetic c...

Significance We identified truncation of tooth development during postnatal ontogeny in two theropod dinosaurs, a caenagnathid oviraptorosaur and the Early Cretaceous bird Sapeornis . Developmental and paleontological evidence each suggests dental reduction and beak evolution are coupled, and a sequence of common morphologies is identified that cha...

The rachises of extant feathers, composed of dense cortex and spongy internal medulla, are flexible and light, yet stiff enough to withstand the load required for flight, among other functions. Incomplete knowledge of early feathers prevents a full understanding of how cylindrical rachises have evolved. Bizarre feathers with unusually wide and flat...

Recent discoveries of exquisitely preserved nonavialan and avialan theropod dinosaurs have not only prompted studies of theropod tooth morphologies, but have also provided information about the origin and early evolution of avian beaks. Recent studies on beak morphologies and morpho-genesis in Darwin's finches have greatly improved our understandin...

A new troodontid (LH PV39) recovered from the Upper Cretaceous Wulansuhai Formation, Inner Mongolia, China, is described, highlighting the dorsoventrally compressed sacral centra. The completely fused neurocentral junctions indicate that LH PV39 had reached adulthood at the time of death, but its size is nevertheless 20% smaller than that of the sy...

Recent studies have significantly advanced our understanding of the evolutionary processes and developmental mechanisms behind feather branching patterns. However, little attention has been given to the tissue differentiation of branches associate with this process. Here, we describe eight feathers preserved in Burmese ambers, characterized by ribb...

A dietary shift from carnivory to insectivore has been proposed to explain the dramatic morphological evolution of alvarezsaurians, particularly the adaptation related to the manual digital reduction and body size miniaturization. However, based solely on morphological shifts, this hypothesis lacked direct dietary evidence to support either carnivo...

Tooth attachment and replacement play significant roles in the feeding ecology of polyphyodont vertebrates, yet these aspects have remained largely unexplored in non-avialan paravians including troodontids. Here, we describe a new troodontid species, Urbacodon norelli sp.n., recovered from the Upper Cretaceous Iren Dabasu Formation of Inner Mongoli...

A new small-bodied troodontid (LH PV39) recovered from the Upper Cretaceous Wulansuhai Formation, Suhongtu, Inner Mongolia, China, is described. The new specimen preserves six postaxial cervical vertebrae, five completely fused sacral and four posterior caudal vertebrae in addition to two manual unguals. The completely fused neurocentral junctions...

The evolution of flight in feathered dinosaurs and early birds over millions of years required flight feathers whose architecture features hierarchical branches. While barb-based feather forms were investigated, feather shafts and vanes are under-studied. Here, we take a multidisciplinary approach to study their molecular control and bio-architectu...

Graphical Abstract Highlights d Two new alvarezsaurian dinosaurs are described from Northwest China d They are intermediate between Late Jurassic and Late Cretaceous alvarezsaurians d They showcase the evolution of highly specialized alvarezsaurian forelimb d Specialized alvarezsaurian forelimb morphology evolved slowly, in a mosaic fashion In Brie...

The bone-beds of the Upper Cretaceous Wangshi Group in Zhucheng, Shandong, China are rich in fossil remains of the gigantic hadrosaurid Shantungosaurus. Here we report a new oviraptorosaur, Anomalipes zhaoi gen. et sp. nov., based on a recently collected specimen comprising a partial left hindlimb from the Kugou Locality in Zhucheng. This specimen'...

Among the characters of caenagnathid oviraptorosaurians, the lateral occlusal grooves and ridges on the occlusal surface of the jaw bones often receive special attention. Recent studies demonstrated that ontogenetic edentulism is present in caenagnathids, and therefore the lateral occlusal grooves and ridges are vestigial alveoli and interdental se...

In order to determine the incubation temperature of eggs laid by non-avian dinosaurs, we analysed the oxygen isotope compositions of both eggshell carbonate (δ¹⁸Oc) and embryo bone phosphate (δ¹⁸Op) from seven oviraptorosaur eggs with preserved in ovo embryo bones. These eggs come from the Upper Cretaceous Nanxiong Formation of Jiangxi Province, Ch...

Oviraptorids, like many other dinosaurs, clearly had a complex pattern of skeletal growth involving numerous morphological changes. However, many ontogenetic skeletal changes in oviraptorids were previously unclear due to the lack of well preserved specimens that represent very young developmental stages. Here we report three elongatoolithid dinosa...

The oviraptorosaurian theropod Caenagnathasia was first described from the Late Cretaceous (Turonian) vertebrate assemblage from the Bissekty Formation of the Kyzylkum Desert, Uzbekistan. Here we report a partial pair of oviraptorosaurian fused lower jaws, comprising the symphysial region, from the Upper Cretaceous (Campanian) Iren Dabasu Formation...

This study is based on the newly collected juvenile specimens of Limusaurus inextricabilis and four undescribed oviraptorids from China. Limusaurus inextricabilis is a ceratosaur known from the Upper Jurassic Shishugou Formation at Wucaiwan area, Xinjiang Uyghur Autonomous Region. So far, a total of 19 individuals of L. inextricabilis have been rec...

The dromaeosaurid Linheraptor exquisitus was named in 2010 based on a nearly complete skeleton recovered from the Upper Cretaceous Wulansuhai Formation at the Gate Locality, in Bayan Mandahu, western Nei Mongol, China. However, three recent studies regarded L. exquisitus as a subjective junior synonym of Tsaagan mangas, a dromaeosaurid from the Upp...

The dromaeosaurid Linheraptor exquisitus was named in 2010 based on a nearly complete skeleton recovered from the Upper Cretaceous Wulansuhai Formation at the Gate Locality, in Bayan Mandahu, western Nei Mongol, China. However, three recent studies regarded L. exquisitus as a subjective junior synonym of Tsaagan mangas, a dromaeosaurid from the Upp...

Background Fleas, the most notorious insect ectoparasites of human, dogs, cats, birds, etc., have recently been traced to its basal and primitive ancestors during the Middle Jurassic. Compared with extant fleas, these large basal fleas have many different features. Although several fossil species with transitional morphologies filled the evolutiona...

Background: Fleas, the most notorious insect ectoparasites of human, dogs, cats, birds, etc., have recently been traced to its basal and primitive ancestors during the Middle Jurassic. Compared with extant fleas, these large basal fleas have many different features. Although several fossil species with transitional morphologies filled the evolution...

Brain endocasts are rare in the fossil record because they are only preserved under exceptional conditions. An equid brain endocast from the early Pleistocene of Wanrong County, Shanxi Province, China, is reported in this paper. Measuring approximately 140 × 95.2 × 83 mm, the new specimen represents a relatively advanced adult horse brain. Comparis...

Yixianosaurus longimanus is a small theropod from the Lower Cretaceous of western Liaoning Province, China. It was originally suggested to be a derived maniraptoran, but this interpretation was challenged by a recent study that proposed instead that Y. longimanus was a basal maniraptoran. Given that the systematic position of this taxon will affect...

Fleas are a group of highly specialized blood-feeding ectoparasites whose early evolutionary history is poorly known [1, 2]. Although several recent discoveries have shed new light on the origin of the group [3, 4], a considerable gap exists between stem fleas and crown fleas. Here we report a new transitional flea, Saurophthirus exquisitus sp. nov...

This paper describes a new oviraptorid dinosaur taxon, Ganzhousaurus nankangensis gen. et sp. nov., based on a specimen collected from the Upper Cretaceous Nanxiong Formation of Nankang County, Ganzhou City, Jiangxi Province, southern China. This new taxon is distinguishable from other oviraptorids based on the following unique combination of primi...

Here we report a new oviraptorid taxon based on a specimen collected from the Upper Cretaceous Wulansuhai Formation of Bayan Mandahu, Linhe, China. This new taxon

Here we report a new oviraptorid taxon based on a specimen collected from the Upper Cretaceous Wulansuhai Formation of Bayan Mandahu, Linhe, China. This new taxon is distinguishable from other oviraptorid species by the following unique features: the ventral extremity of the large and elongate external naris is located below the mid-height of the p...

Four-Winged Birds? Recently, nonavialan dinosaurs with feathers on their fore- and hindlimbs have been described. Zheng et al. (p. 1309 ) describe eleven basal avialan fossils with clear evidence of feathered hindlimbs. Together these fossils show that early avialans possessed four wings, rather than two. A gradual reduction in hindlimb feathering...

Yixianosaurus longimanus is a small theropod from the Lower Cretaceous of western Liaoning Province, China. It was originally suggested to be a derived maniraptoran, but this interpretation was challenged by a recent study that proposed instead that Y. longimanus was a basal maniraptoran. Given that the systematic position of this taxon will affect...

Numerous feathered dinosaur specimens have recently been recovered from the Middle-Upper Jurassic and Lower Cretaceous deposits of northeastern China, but most of them represent small animals. Here we report the discovery of a gigantic new basal tyrannosauroid, Yutyrannus huali gen. et sp. nov., based on three nearly complete skeletons representing...

Parasite-host associations among insects and mammals or birds are well attended by neontological studies [1]. An Eocene bird louse compression fossil [2, 3] and several flea specimens from Eocene and Oligocene ambers [4-8], reported to date, are exceptionally similar to living louse and flea taxa. But the origin, morphology, and early evolution of...

Some Miocene ratite eggshell fragments from Gashunyinadege (about 17.5 Ma) and Baogedawula (7.11 ± 0.48 Ma), Inner Mongolia, China, are described. The outer surfaces of eggshell frag-ments from Gashunyinadege are smooth with a sting pore pattern, while their entire thickness is around 2.12 mm. In a cross-sectional view, the surface crystal layer is...

Struthio linxiaensis is the oldest fossil ostrich from Asia till today. This paper analyzes the microstructure of the eggshell of Struthio linxiaensis. The SEM and light microscopic results show that a pathological structure occurred associated with the formation of pore canal. The feature of this structure is that crystals of the columnar layer ar...

A Galliformes bird fossil was collected from the early Pleistocene Sanmen Formation in the lower reaches of the Fenhe River, Shanxi Province. It has the following diagnostic features: the cotylae of the inner and outer proximal foramens of the tarsometatarsus are deep; the attachments of the internal ligament are well developed; trochlea metatarsi...