Shalva Barjadze

• PhD
• Associate Professor at Ilia State University

During investigations of the cave fauna in Georgia, two new taxa of carabid beetles were discovered. Inotrechus usholtaensis sp. nov. from Usholta Cave and Duvalius gregoi sp. nov. from Kvedi II Cave, Racha-Lechkhumi and Kvemo Svaneti region, Georgia, are described, illustrated, and differentiated from morphologically similar species. Identificatio...

A new leech species, Dina kobakhidzei sp. nov., discovered in the Tskhrajvari I Cave in the Racha-Lechkhumi and Lower Svaneti region of western Georgia, is described and illustrated. The study includes a differential diagnosis and an identification key for Dina species known from Georgia.

In this paper, we present a review of stridulating species of aphids of the Macrosiphini genus Uroleucon (Hemiptera, Aphididae) and include 13 species. The most characteristic feature of the reviewed species is the presence of unusual aphids’ peg-like sensilla distributed on the hind tibia, which are rubbed against the surface of the siphunculi. A...

The new aphid species-Aphis (Bursaphis) kazdagensis sp. nov. living on Papaver pilosum subsp. strictum (Papaveraceae) in Çanakkale Province, Türkiye, is described based on the apterous viviparous females. Diagnosis of the subgenus Bursaphis with respect to other species is given. Aphis popovi Mordvilko, 1932 is transferred from the subgenus Aphis t...

Georgia's rich biodiversity, as a key part of the Caucasus hotspot, makes it an ideal destination for scientific research. Nevertheless, knowledge about the species diversity of different animal groups in this area remains limited. In this article we provide information on the nine centipede species (Henia brevis (Silvestri, 1896), H. hirsuta Verho...

Background Honey bees are the principal commercial pollinators. Along with other arthropods, they are increasingly under threat from anthropogenic factors such as the incursion of invasive honey bee subspecies, pathogens and parasites. Better tools are needed to identify bee subspecies. Genomic data for economic and ecologically important organisms...

This poster explains about integrative taxonomy of H. nitidus (Collembola) from the caves of Slovakia, several Balkan countries, Georgia, and Iran.

This conference paper explains about Integrative taxonomy of H. nitidus (Collembola) from the caves of Slovakia, several Balkan countries, Georgia, and Iran.

Eight aphid species - Aphis hederae Kaltenbach, 1843, A. mirifica (Börner, 1950), A. origani Passerini, 1860, A. umbrella (Börner, 1950), Chaitophorus horii Takahashi, 1939, Hyadaphis coriandri (Das, 1918), Uroleucon jaceae (Linnaeus, 1758) and U. inulicola (Hille Ris Lambers, 1939) are recorded for the first time from Armenia. Additionally, Aphis...

Eight aphid species - Aphis hederae Kaltenbach, 1843, A. mirifica (Börner, 1950), A. origani Passerini, 1860, A. umbrella (Börner, 1950), Chaitophorus horii Takahashi, 1939, Hyadaphis coriandri (Das, 1918), Uroleucon jaceae (Linnaeus, 1758) and U. inulicola (Hille Ris Lambers, 1939) are recorded for the first time from Armenia. Additionally, Aphis...

Plutomurus weinerae sp. nov. from Usholta Cave, Racha-Lechkhumi and Kvemo Svaneti region, Georgia, and P. danialensis sp. nov. from Danial Cave, Mazandaran province, Iran are described, illustrated and differentiated from other morphologically similar congeners. New species are distinguishable from similar species by head and abdominal macrochaetot...

The new aphid species Metopeurum caucasicum sp. nov. living on Achillea millefolium and five species of Tanacetum, namely T. abrotanifolium, T. balsamita, T. balsamitoides, T. canescens and T. vulgare (Asteraceae) in Armenia and Georgia, is described based on the apterous viviparous females. Diagnosis to other species of the genus is given. Some di...

The new aphid species Metopeurum caucasicum sp. nov. living on Achillea millefolium and five species of Tanacetum, namely T. abrotanifolium, T. balsamita, T. balsamitoides, T. canescens and T. vulgare (Asteraceae) in Armenia and Georgia, is described based on the apterous viviparous females. Diagnosis to other species of the genus is given. Some di...

An annotated list of the Georgian harvestmen was prepared based on literature and unpublished collection data, in which 48 species belonging to 19 genera within six families are included. Twenty-seven species are endemic to the Caucasus Ecoregion, while 10 species are endemic to Georgia. Five species are highly specialized cavernicolous ones and ar...

Here we present a description of a new aphid species from the subgenus Asterobium Hille Ris Lambers, 1938 of the genus Macrosiphoniella Del Guercio, 1911 in the tribe Macrosiphini Wilson, 1910 (Aphididae). The new species—Macrosiphiniella (Asterobium) herczeki sp. nov. living on Galatella villosa (Asteraceae) in Tolbuchin region, Bulgaria, is descr...

The present paper is devoted to the description of three new species of the genus Acanthophorella Antić & Makarov, 2016 from Georgia, western Caucasus: A. devi Antić sp. nov. and A. valerii Antić sp. nov., both presumed troglobionts and each from a single cave, and the epigean A. aurita Antić sp. nov. The troglobiotic Acanthophorella barjadzei Anti...

The Caucasus ecoregion is distinguished by its biodiversity. The flora and fauna of the ecoregion became the study area of many zoologists and botanists. Georgia, as a part of it, is important for its valuable and impressive diversity. However, despite many years of scientific research, certain groups, among them centipedes, are not so well studied...

This paper presents a checklist of aphids of Armenia. Nowadays, 203 species of aphids belonging to 83 genera, 10 subfamilies, two families and two superfamilies are known, of which nine species are recognized in Armenia for the first time: Aphis cephalariae Barjadze, A. chloris Koch, A. solanella Theobald, A. urticata Gmelin, A. wellensteini (Börne...

A new species Centromerus georgicus sp. n. collected in a Georgian cave (Caucasus) is described based on male and female specimens, diagnosed, and illustrated. New records of some poorly known species (Tegenaria pontica Haritonov, 1947, Dysdera anatoliae Deeleman-Reinhold, 1988 and Leptonetela caucasica Dunin 1990), as well as the description of th...

Two new Caucasian species of the genus Niphargus Schiödte, 1849 (Crustacea: Amphipoda: Niphargidae), N. rachalechkhumensis sp. nov. and N. tvishiensis sp. nov., are described from the stygobiotic habitats of the Racha-Lechkhumi and Kvemo Svaneti karst systems in Western Georgia. Both newly described species belong to the southwestern Caucasian “Nip...

Two species of leeches were described from Georgia in the past, Dina ratschaensis Kobakhidze, 1958 from the Racha-Lechkhumi and Kvemo Svaneti region and Trocheta ariescornuta Grosser, Barjadze & Maghradze, 2021 from the Samegrelo-Zemo Svaneti region. Both species were the only known typical representatives of cave leeches in Georgia. Recently, two...

This case study is aimed to investigate the invertebrate fauna and distribution features of the faunal and floral diversity in seven habitats represented on the territory of the Saskhori limestone quarry and its adjacent territories (Eastern Georgia, Caucasus). Totally, 122 species of invertebrates and 131 species of plants were registered. The iso...

Specimens of Deuteraphorura collected in 11 Georgian caves were analysed morphologically and molecularly based on the COI gene barcode region. Two molecular delimitation methods revealed four species (MOTUs); however, only two of them were distinguished morphologically and are described in this paper as new to science. Both new species, D. colchisi...

A cavernicolous harvestman species, Nemaspela kotia sp. nov., is described from the Kotia Cave (=Kotiasklde Cave) in Zemo Imereti plateau, Imereti Region, West Georgia. Its males lack a cheliceral apophysis, a feature shared with the Georgian cave-dwelling Nemaspela femorecurvata Martens, 2006 and N. prometheus Martens, Maghradze & Barjadze, 2021....

Chaetophiloscia hastata Verhoeff, 1928 is recorded from Saskhori limestone quarry (Mtskheta municipality, eastern Georgia). This is the first confirmed record of the genus and species for the Georgian isopod fauna. The species is fully illustrated to facilitate its identification.

Six aphid species – Aphis armata Hausmann, 1802; Capitophorus bulgaricus Tashev, 1964; Macrosiphoniella nikolajevi Kadyrbekov, 1999; Macrosiphum cerinthiacum Börner, 1950; Uroleucon cirsicola (Holman, 1962) and U. rapunculoidis (Börner, 1939) are recorded for the first time from Georgia, and Aphis sanguisorbicola Takahashi, 1966, is recorded for th...

Cryptops (Cryptops) datviensis sp. nov. from Datvi Cave in Georgia, is the first troglobiotic species of the genus Cryptops Leach, 1814 to be described from the Caucasus. The new species morphologically resembles the common epigean Caucasian species Cryptops caucasius Verhoeff, 1934, but differs in typical troglomorphic features, such as elongation...

Apterous viviparous ♀♀ of Lipaphis (Lipaphidiella) holmani sp. nov., living on Lepidium virginicum L. (Brassicaceae) in South Korea, are described and differences with all known species belonging to the subgenus Lipaphidiella Doncaster, 1954 and Lepidium-feeding Lipaphis spp. are presented. In addition, the poorly known Lipaphis sisymbrii Bozhko, 1...

Geoica inbari sp. nov., living on Pistacia palaestina Boiss. (Anacardiaceae), is described based on apterous fundatrigeniae, fall migrants, embryos of apterous exules within fall migrants (fundatrispuriae) and the first instar larva of an apterous exule borne by a fall migrant, collected in the Mount Hermon area of Northern Israel. The new species...

Here, we present the descriptions of a new aphid species in the genus Pleotrichophorus Börner, 1930 (Hemiptera: Aphididae: Macrosiphini), found by Kenneth L. Hibbard, inspection supervisor for the Florida Department of Agriculture and Consumer Services, Division of Plant Industry. It is associated with Euthamia graminifolia (L.) Nutt. (Asteraceae)...

Brachyunguis dendrostellerae Kadyrbekov, 2014 and B. monstratus Kadyrbekov, 1999, both living on Diarthron lessertii (Wikstr.) Kit Tan. (Thymelaeaceae), are recorded for Iranian aphid fauna for the first time. The hitherto unknown alate viviparous females of B. dendrostellerae are described, and the little-known apterous viviparous females are rede...

Plutomurus kharagauliensis sp. nov. from Kozmani Cave, Kharagauli Municipality, Georgia, and P. ruseki sp. nov. from the surface and subterranean habitats in different regions of Slovakia are described, illustrated and differentiated from other morphologically similar congeners. Both species are unique by dorsal macrochaetotaxy of head and abdomen....

Prometheus Cave is one of the largest caves in Georgia among the local six show caves. Before opening the cave as a tourist attraction, no research was conducted on the cave to study the invertebrate community living there, despite the cave's status as a natural monument. Before our study, only 22 species of invertebrates were known from Prometheus...

Findings of 6th Eurospeleo Protection Symposium on monitoring EU Habitat 8310 "Caves not open to the public"

This manuscript summarizes the outcomes of the 6th EuroSpeleo Protection Symposium. Special emphasis was laid on presenting and discussing monitoring activities under the umbrella of the Habitats Directive (EU Council Directive 92/43/EEC) for habitat type 8310 "Caves not open to the public" and the Emerald Network. The discussions revealed a high l...

An integrative approach resulted in a description of a new species of stygobiotic shrimps of the genus Xiphocaridinella Sadowsky, 1930 (Crustacea: Decapoda: Atyidae) from the southern part of the Racha-Lechkhumi and Kvemo Svaneti Region of the Western Georgia (SW Caucasus). The area and caves, from which this species is recorded will be flooded dur...

A new dipluran species, Plusiocampa (Plusiocampa) imereti Sendra & Barjadze sp. nov., from the deep zone in three caves in the Imereti region, Georgia, is described. This new troglobitic Plusiocampa is an addition to four others known Diplura from around the Black Sea region, two Dydimocampa and two Plusiocampa s. str. The present study also provid...

en This paper provides an overview on troglobitic springtails found in European caves, including a checklist at species level. The paper also reviews what is currently known about Collembola, which occur in caves of the most important mountain ranges in Europe. Only troglobitic species were included since many troglophiles were of uncertain ecologi...

Globochthonius satapliaensis (Schawaller & Dashdamirov, 1988) (n. comb.) original-ly described as Chthonius (Chthonius) satapliaensis from a single female collected in the Sataplia II Cave, Imereti Region (Georgia) is redescribed and depicted based on specimens of both sexes from caves of Sataphlia-Tskaltubo karst massif and Zemo Imereti Plateau. I...

A new leech species Trocheta ariescornuta n. sp. is described and illustrated here based on two specimens found in Motena Cave in western Georgia, Caucasus. This species differs from the known congeners and other similar Erpobdellids of the region, including Georgian cavernicolous leech - Dina ratschaensis Kobaķhidze, 1958 in a complex of external...

Two highly specialized endemic troglobiotic harvestman species of the genus Nemaspela Šilhavý, 1966 are described. N. melouri sp. nov. from Melouri Cave and N. prometheus sp. nov. from Prometheus Cave (Sataplia-Tskaltubo karst massif, Imereti region, western Georgia), respectively. Despite the fact that the entrances of the caves are positioned onl...

Georgia is located on the southern slopes of the Great Caucasus Mountain Range, as part of the Caucasus Eco‐region, is considered one the 35 worldwide significant biodiversity hotspots based on the species richness and the significant level of species endemism. One of the threats that may be considered in respect to forest habitat types of high con...

Here, we present descriptions of two new aphid species of the genus Uroleucon Mordvilko, 1914 (Hemiptera: Aphididae: Macrosiphini), which are associated with the plant genus Grindelia Willd. (Asteraceae). An apterous viviparous female of Uroleucon (Lambersius) robinsoni sp. nov. from Oregon and apterous and alate viviparous females of Uroleucon (La...

A new species of the aphid genus Uroleucon from Lebanon, with notes on the systematic positions of U. altaicum, U. bielawskii and U. mulgedii (Hemiptera: Aphididae) Abstract Here we present a description of a new aphid species - Uroleucon (Uroleucon) lebanonense sp. nov. associated with Tragopogon coloratus (Asteraceae) in Lebanon. Apterous and ala...

Here we present a review of the 18 species of the Macrosiphini genus Macrosiphoniella (Hemiptera: Aphididae) known to occur in the United States of America, representing two subgenera, Macrosiphoniella and Phalangomyzus Börner, 1939. We report key features for all species including colour in life and pigmentation on slides of apterous and alate viv...

In two European species of the genus Plutomurus Yosii, P. carpaticus Rusek & Weiner and P. unidentatus (Börner), several diagnostic morphological characters are specified and illustrated: body chaetotaxy, number of internal spine-like chaetae on tibiotarsi and number of teeth on claw. The contribution is based on the study of type specimens of both...

A new species of aphid, Xerobion eteriae sp. n., is described based on apterous and alate viviparous females, living on Acantholimon (Plumbaginaceae) in Iran. Differences from the most similar species of the genus are presented. http://www.zoobank.org/urn:lsid:zoobank.org:pub:A3FAD52D-ED85-4BD1-8A28-EEFD42E62E67

Plutomurus jordanai sp. nov. from Zeda Kvilishori Cave (Tskaltubo Municipality, Imereti region, Western Georgia) is described, illustrated and differentiated from other morphologically similar species‒‒P. kelasuricus Martynova, 1969 and P. eristoi Barjadze, Baquero, Soto-Adames, Giordano & Jordana, 2016. In addition, morphological characters omitte...

Apterous viviparous females of Aphidura jimoi sp. nov. living on Dianthus sp. in East Georgia, are described and differences with the most similar species of the genus—A. naimanica Kadyrbekov, 2013, are presented. Additional data are provided for apterous viviparous females of A. pakistanensis Nieto Nafría, Mier Durante & Remaudière, 2013, based on...

A survey of gall-inducing aphids on elm and poplar trees was carried out during 2017 in Razavi Kho-rasan province, NE Iran. As a result, 15 species of gall-inducing aphids from 5 genera, all belonging to the subfamily Eriosomatinae, were recorded on 6 host plant species. The collected species included the genera Eriosoma, Kaltenbachiella, Pemphigus...

During a biospeleological survey of caves of Imereti region (Georgia), a new species of the beetle genus Bergrothia Reitter, B. barbakadzei sp. nov. was discovered in three caves: Datvis (Bear), Prometheus and Melouri caves (Sataplia–Tskaltubo karst massif, Tskaltubo district). The new species is described, illustrated and differentiated from other...

Information about the cave invertebrates of Georgia, Caucasus, is summarised, resulting in 43 troglo- and 43 stygobiont taxa reported from 64 caves. Species distribution analyses were conducted for 61 caves harbouring 58 invertebrate taxa, with the majority of caves (39) located in Apkhazeti (north-western Georgia). In 22 caves from central-west Ge...

We present the first records from Iran of Macrosiphoniella (Asterobium) davazhamci Holman & Szelegiewicz, 1974 and M. (Ramitrichophorus) nikolajevi Kadyrbekov, 1999 (Hemiptera: Aphididae: Aphidinae) from samples collected by the late Prof. G. Remaudière. Previously, Macrosiphoniella davazhamci was known from Kazakhstan, Mongolia and China, whereas...

Inbaria n. gen. is proposed based on type species Geoica swirskii Remaudière & Inbar, 2004, resulting in a new combination, Inbaria swirskii (Remaudière & Inbar, 2004) n. comb. Inbaria easily differs from all Pistacia gall-producing aphids of the tribe Fordini by the size of the primary and secondary rhinaria in fall migrants. Inbaria n. gen. is di...

Two new species of Plutomurus, P. pichkhaiai sp. nov. from Garakha and Letsurtsume caves and P. shurubumuensis sp. nov. from Shurubumu Cave (Chkhorotsku district, Western Georgia) are described, and illustrated. The new species are very similar to P. kelasuricus from the Tsebelda karst massif formation in Apkhazeti, Georgia, but differ in having tw...

Ericaphisvoegtlinisp. n. living on Chamaebatiafoliolosa (Rosaceae) in California is described based on apterous and alate viviparous females. The new species differs from all other species of the genus Ericaphis Börner, 1939 in several important morphological characters including very long and rigid dorsal setae and distinctly swollen siphunculi wi...

One springtail species, Heteraphorura iranica Kaprus’, Shayanmehr et Kahrarian, 2017 is recorded from the Caucasus for the first time. A key to the Holarctic species of the genus Heteraphorura Bagnall, 1948 including H. magnina (Wray, 1950) is given.

The aphid genus Myzaphis van der Goot, 1913 from the tribe Macrosiphini is revised to include eight species. Apterous and alate viviparous females, known fundatrices and known sexual morphs (oviparous females and males) of Myzaphis bucktoni, M. juchnevitschae, M. rosarum, M. tianshanica and M. turanica are re-described and illustrated. Lectotype an...

Apterous and alate viviparous females of Hyadaphis bupleuriphila sp. n. living on Bupleurum sp. (Apiaceae) and further undetermined Apiaceae are described from the Ankara province in central Turkey. The new species is morphologically similar to Hyadaphis coerulescens (Narzikulov, 1965), H. ferganica Mukhamediev & Akhmedov, 1979 and H. foeniculi (Pa...

The hitherto unknown oviparous females and alate males of M. dzhibladzeae Barjadze, 2010, living on Euphorbia macroceras Fish. & Mey. and Euphorbia sp. (Euphorbiaceae), are described from resort Bakhmaro (Chokhatauri district, Guria region, Western Georgia). The life cycle of this species is established. Apterous and alate viviparous females and ov...

Apterous viviparous females of Neoamphorophora ledi (Wahlgren, 1938) living on Rhododendron degronia-num Carr. (Ericaceae) on Honshu island, Japan, are redescribed and the hitherto unknown alate viviparous females are here described. Rhododendron degronianum is a new host plant species for this aphid. This species is recorded for aphid fauna of Eas...

Apterous and alate viviparous females and the alate males of Macrosiphum eastopi Barjadze & Blackman sp. n. living on Oberna multifida Ikonn. (=Silene multifida Rohrb.) (Caryophyllaceae) are described from the Republic of Georgia (Caucasus). Type specimens are deposited at the Institute of Zoology, Ilia State University (IZISU), Tbilisi, Georgia; t...

Apterous viviparous females of Neoamphorophora ledi (Wahlgren, 1938) living on Rhododendron degronianum Carr. (Ericaceae) on Honshu island, Japan, are redescribed and the hitherto unknown alate viviparous females are here described. Rhododendron degronianum is a new host plant species for this aphid. This species is recorded for aphid fauna of East...

Apterous and alate viviparous females of the sage aphid, Aphis passeriniana (Del Guercio, 1900) (Hemiptera: Aphididae), were collected on Garden sage, Salvia officinalis L. (Lamiaceae) in a cultivated sage field in Bornova, Izmir Province, Turkey. The aphid is recorded for the first time from Turkey. Brief information about the morphology, biology...

The aphid genus Myzaphis van der Goot, 1913 from the tribe Macrosiphini is represented by eight species worldwide. Members of this genus are characterized by rather small, elongate oval body and characteristic rugose or wrinkled dorsal cuticle. They live on undersides of leaves of Potentilla and Rosa spp. (Rosaceae), are monoecious holocyclic, rare...

Nine aphid species are recorded for the first time from the following countries in the Palearctic region: four species from Iran, three species from Georgia and one species each from Germany and the Netherlands. Information on biology and geographical distribution for each species is also provided. Besides, three host plant species and one new host...

Adult female individuals of the nematode Mesomermis devii sp. nov. (Nematoda: Mermithidae) have been described. The host organism is not determined. The individuals were isolated from moist guano obtained in the dark zone of Devi Cave (West Georgia). The new species is characterized by the following: rounded and symmetrical apical end of the head;...

Two new species of the genus Plutomurus, P. revazi sp. nov. from Prometheus and Satsurblia caves and P. eristoi sp. nov. from Satevzia Cave are described, illustrated and differentiated from other morphologically closely related species. A high variability in the number of teeth in the claw, unguiculus and mucro of P. revazi sp. nov. demonstrate th...

Faunal surveys were performed in Mtirala National Park (Georgia, Caucasus). Oribatid mites were collected from four microhabitats (soil, moss, litter, decaying wood) at 15 locations from 250 to 1000 m a.s.l. Forest types included walnut (Juglans regia) plantations, mixed chestnut (Castanea sativa) forests, and beech (Fagus orientalis) forests. Our...

Results of grapevine yellows disease (GY) studies in 2005-2015 are reported. Based on symptoms and Dienes’ staining method the disease was detected in Kartli, Kakheti and Guria regions on Vitis vinifera (L.) Rkatsiteli, Saperavi, Shavkapito, Tavkveri, Aladasturi, Kachichi, Ganjuri, Chardonnay, V. labrusca (L.) cultivar Isabella, species hybrid cult...

Four species of springtails and four species of oribatid mites were identified as new records for the Georgian fauna from different regions of the country: Coecobrya tenebricosa (Folsom, 1902); Entomobrya muscorum (Nicolet, 1842) Rondani, 1861; Heteromurus major (Moniez, 1889); Pseudosinella octopunctata Börner, 1901; Austrocarabodes ensifer (Selln...

Aphis polii sp. n. living on Teucrium polium L. (Lamiaceae) is described, based on apterous and alate viviparous females and oviparous females. It has been collected from seventeen localities in mainland France, Corsica, Italy, Ukraine, Lebanon and Iran. The new species is similar to two other Aphis species that feed specifically on plants of the g...

Cryptophagus axillaris Reitter, 1875 from Savekuo Cave is recorded for the beetle fauna of the Caucasus for the first time, while C. lycoperdi (Scopoli, 1763) from Bichvinta (=Pitsunda) is a new record for Georgian beetle fauna. Twenty five species of the Cryptophagus beetles are recorded in Georgia. Sampling data and distribution map for the genus...

Seven aphid species are recorded from the countries of the Middle East and the Caucasus for the first time, of which three aphid species are new records for Turkey, two species for Lebanon, and one species each for Iran and Georgia. New host plant genera are reported for Aphis cephalariae Barjadze, Anoecia vagans (Koch), and Macrosiphum symphyti Ba...

The ant genus Myrmoxenus consists of about ten socially parasitic species including active slave-makers and workerless “degenerate slave-makers”. Myrmoxenus tamarae was previously known only from type material, two workers collected at Daba, Georgia and nothing was known about its life history, colony structure or the morphology of its sexuals. An...

Zelkovaphis caucasica is a little-known Caucasian species of leaf-rolling aphids on Zelkova carpinifolia, and was not included in a previous morphological phylogenetic analysis of the tribe Eriosomatini. We redescribe little-known emigrants of Z. caucasica and describe first instars borne by the emigrants, with keys to species for these morphs of t...

Annotated list of the invertebrates in the caves of Racha region (Western Georgia) is made based on proper and literature data. 73 species of the invertebrates are recorded in the karst caves, 29 species of which belong to the oribatid mites, 12 species-to collembolans, 8 species – to spiders and insects, 5 species – to myriapods, 4 species – to mo...

The ant genus Myrmoxenus consists of about ten socially parasitic species including active slave-makers and workerless "degenerate slave-makers". Myrmoxenus tamarae was previously known only from type material, two workers collected at Daba, Georgia and nothing was known about its life history, colony structure or the morphology of its sexuals. An...

Apterous and alate viviparous females of Protaphis kvavadzei sp. n. on Eryngium campestre (Apiaceae) and Aphis matricariae sp. n. on Matricaria sp. (Asteraceae) from Turkey are described. The new species are differentiated from other aphids colonising these host plants and from species with similar morphology.

the study of specimens collected by the first author, a new species of Argonychiurus from a Georgian cave is described.

Rhododendraphis tuatayae gen. n., sp. n. living on Rhododendron sp. (Ericaceae) in Turkey is described based on apterous viviparous females. The new genus is morphologically most similar to Rostratusaphis Fang and Qiao, 2009, but differs in several important characters. Differences between the new genus and other aphid genera with a spinulose secon...

The collembolan species Plutomurus kelasuricus Martynova, 19695. Martynova, E. (1969): Springtails of the family Tomoceridae (Collembola) from the fauna of the USSR [in Russian]. Entomological Review, 43, 299–314.View all references is redescribed and illustrated based on material sampled in Georgian caves. Differences from the morphologically simi...

Winged morphs of aphids were investigated in 2008-2009 in G�lc�k Natural Park (Isparta Province, Turkey). Nineteen species of aphids were caught in 3 Malaise traps in 3 localities of the above mentioned territory. Most of the aphid species (12 species) were members of the subfamily Aphidinae, subfamilies Calaphidinae and Eriosomatinae were each rep...

Pemphigus saliciradicis (Hemiptera: Aphididae: Eriosomatinae) lives on roots of Salix spp. (Salicaceae) and Vitis vinifera (Vitaceae) in Holarctic region. Morphometric data for eight morphological characters, measured from 10 field collected samples (50 specimens) were analysed by principal component analysis (PCA) and canonical discriminant analys...

The greenbug, Schizaphis graminum (Rondani), is a major pest of cereals in some parts of the world and is of particular concern because it can be resistant to some insecticides and overcome the resistance of crops. In the UK, it has never been found on crops, but two rather little-known and closely-related species (Schizaphis holci and Schizaphis a...

Seven alate females collected on Rhus sp. by Dr. Murat Aslan in Turkey are re-studied. Originally considered as an undescribed species, the specimens are determined as Baizongia pistaciae (L.). Misidentification of the host plant probably contributed to lead on the initial confusion. Morphological measurements and ratios of diagnostic characters fo...

Fundatrices of Zelkovaphis caucasica (Dzhibladze 1960) are described and illustrated. A key to the fundatrices of the genus Zelkovaphis species is given. A transfer experiment was carried out to establish the life cycle of Z. caucasica. In addition, the life cycle of this species was studied in field conditions. The secondary host plant was establi...

Alate viviparous females of Wahlgreniella nervata (Gillette) were found on a Damask rose (Rosa damascena Mill.) plantation in Isparta Province, Turkey. The aphid is recorded for the first time from Turkey. Besides, Damask rose is a new host for this species. Brief information about this aphid including a key to the alate females of aphid species kn...

A new species, Plutomurus birsteini sp. n., from Georgian caves is described and illustrated. It is similar to Plutomurus baschkiricus (Skorikow, 1899). Differences between the species are discussed. A key to the genus Plutomurus species found in the Caucasus is provided.