Samuel Zamora

Samuel Zamora
Instituto Geológico y Minero de España-CSIC

About

171
Publications
87,864
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2,121
Citations
Citations since 2017
78 Research Items
1365 Citations
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2017201820192020202120222023050100150200250300
2017201820192020202120222023050100150200250300
2017201820192020202120222023050100150200250300

Publications

Publications (171)
Article
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Decapod crustaceans are an important component in modern ecosystems with a rich fossil record spanning more than 400 million years. Last summer (19th–24th June 2022) the 8th Symposium on Fossil Decapod Crustaceans took place in Zaragoza (Spain), a meeting covering all aspects of studies on fossil decapod crustaceans, including information from mode...
Article
Full-text available
Eubrachyurans, or ‘higher’ true crabs, are the most speciose group of decapod crustaceans and have a rich fossil record extending into the Early Cretaceous. However, most extant families are first found in the fossil record in the Palaeogene, and particularly in the Eocene. Unfortunately, fossils of many early eubrachyuran groups are often fragment...
Article
Decapod crustaceans are rare in Early Cretaceous invertebrate assemblages from the Iberian Peninsula; however, current knowledge does allow differences in taxonomic composition between different basins and subbasins to be recognised. In order to understand the distribution of Barremian–Aptian decapod crustaceans of Iberia better, seven new localiti...
Article
Full-text available
Asturicystis' havliceki Fatka & Kordule from the middle Cambrian of Bohemia (Czech Republic) is re-described based on its type material. Several features, including the extension of the food grooves and presence of ventral swellings suggest that 'A.' havliceki does not belong to Asturicystis, and is placed in the new genus Bohemiacinctus. To ascert...
Article
Full-text available
2023. Filling the Silurian gap of solutan echinoderms with the description of new species of Dehmicystis from Spain. Acta Palaeontologica Polonica 68 (2): 185-192. Solutans were among the most enigmatic pre-radial and asymmetric echinoderms. A new species Dehmicystis ariasi sp. nov. is described from the upper part of the Llagarinos Formation, lowe...
Article
Full-text available
Until now, the earliest fossil echinoderms have been predominantly known from the early Cambrian of Laurentia and West Gondwana. Here, we describe Sprinkleoglobus extenuatus gen. et sp. nov. from the Chengjiang biota (Cambrian Series 2, Stage 3) of Yunnan Province, China. The overall profile of the theca and the morphology of the ambulacra clearly...
Article
Full-text available
A highly diversified fauna of hermit crabs associated with reef environments from the Serraduy Formation (lower Eocene) in the southern Pyrenees (Huesca, Spain) is described. Other European Eocene outcrops have yielded paguroids associated with a single environment; however, the studied association represents one of the highest paguroid diversities...
Article
A new echinoid belonging to the genus Habanaster is described from the Western Pyrenees (Pamplona basin, Spain). Definition of Habanaster itzae nov. sp. is based on 16 well-preserved specimens that were collected from Lutetian levels (middle Eocene) of the Anotz Formation. These levels have been interpreted as deposited in low energy, hemipelagic c...
Article
About 40 million years after the Cambrian Explosion, the Great Ordovician Biodiversification Event (GOBE) represents a second and dramatic burst in marine biodiversity, with major changes in the structure of ecosystems and the progressive replacement of the distinctive Cambrian Evolutionary Fauna by the Paleozoic Evolutionary Fauna. However, the GO...
Article
The Roda Formation (early Eocene Ypresian, Huesca, Aragón, northern Spain) records successive levels including rich decapod faunas relevant for the understanding of Ypresian decapod assemblages. Asthenognathus fernandezin. sp. is described from one of these levels, representing the oldest report of a fossil Asthenognathinae, and the first report fr...
Article
Full-text available
Millericrinids constitute an order of extinct articulate crinoids that range from the Middle Triassic to the Late Cretaceous. Based on partially articulated material comprising calyces, columns, and holdfasts, six species (one of them new) belonging to five different genera are described from the Late Jurassic of Spain for the first time. They incl...
Article
The exquisitely preserved, diverse and abundant fossil assemblages yielded by the ‘echinoderm meadows’ of the Tafilalt region of the eastern Anti-Atlas represent a new Konservat-Lagerstätte, one of the few exceptionally preserved Late Ordovician open marine faunas found globally, giving us an insight into the radiation of life during the later phas...
Article
Full-text available
Echinoderms are characterized by a distinctive high-magnesium calcite endoskeleton as adults, but elements of this have been drastically reduced in some groups. Herein, we describe a new pentaradial echinoderm, Yorkicystis haefneri n. gen. n. sp., which provides, to our knowledge, the oldest evidence of secondary non-mineralization of the echinoder...
Article
The extensive, predominantly siliciclastic deposits of the Upper Ordovician of the Tafilalt have long been the subject of scientific investigation. In the past 25 years, intensified collecting for commercial purposes has resulted in the discovery of several exceptionally-preserved faunas ( Konservat-Lagerstätten ) in the Tafilalt region, preserving...
Article
In this paper, we analyse the upper Barremian-lowermost Aptian Alacón Formation (Oliete subbasin, Maestrazgo Basin) of Spain to understand the progressive opening of a shallow-marine bay found on the northeastern margin of Iberia. The succession consists of four transgressive-regressive sequences (S1S4) bounded by widespread unconformities caused b...
Article
Full-text available
A B S T R A C T The cyrtocrinid crinoid Proholopus holopiformis (Remeš, 1902) is described from the upper Albian succession that may be included in the Albeniz unit to the east of Iruñea-Pamplona (Navarre, Western Pyrenees, Spain). Although based on partially disarticulated material, this taxon preserve calyx, stem, attachment structure, and brachi...
Article
Full-text available
A highly diverse fauna of dromioid brachyurans from the Serraduy Formation (lower Eocene) in the western Pyrenees (Huesca, Spain) is described and illustrated. Recorded taxa are Mclaynotopus longispinosus new genus new species, Torodromia elongata n. gen. n. sp., Basidromilites glaessneri n. gen. n. sp., ? Basidromilites sp., Sierradromia gladiator...
Article
Full-text available
A new genus and two new species of fossil hermit crabs (Anomura, Paguroidea) are described from the southern Pyrenean basins (Huesca, NE Spain). Parapetrochirus nov. with P. robustus n. gen., n. sp., as type species, comes from the lower Eocene (Ypresian) Roda Formation. It preserves both chelipeds exhibiting a notable heterochely, a larger left ch...
Article
Full-text available
The Sobrarbe-Pirineos UNESCO Global Geopark, located in the Central Pyrenees, is a region of remarkable geodiversity that includes extensive Eocene fossil-bearing sites and constitutes an important archive of paleobiodiversity. The Sobrarbe-Piri-neos Geopark hosts outcrops of Eocene formations bearing an unusual abundance and diversity of fossils f...
Article
Modern reefs are considered important hot spots of biodiversity, but the analysis of the distribution of the invertebrate fauna across different reefal domains in ancient ecosystems can be challenging, because the fossil record is usually affected by strong taphonomic biases. The lower Eocene coral reef in the well-exposed outcrops of Ramals (Tremp...
Article
A b s t r a c t Decapod crustaceans from the Forcall Formation (Aptian, Lower Cretaceous) of the Oliete Sub-basin (Maestrazgo Basin) are recorded, specimens having been collected from the localities of Josa and Obón (Teruel) in the Aragonese branch of the Iberian Cordillera, northeast Spain. The glypheid lobster Atherfieldastacus magnus is by far t...
Article
Full-text available
Semi-articulated paguroids are rare fossils, and there are only few records from Cenozoic strata. Here we present a new and exceptionally preserved hermit crab (Diogenidae) from the Eocene of Huesca (Spain) that preserves the anterior part of the carapace, together with appendages. Diogenes augustinus sp. nov. represents one of most completely pres...
Article
Full-text available
Reconstructing the function and behaviour of extinct groups of echinoderms is problematic because there are no modern analogues for their aberrant body plans. Cinctans, an enigmatic group of Cambrian echinoderms, exemplify this problem: their asymmetrical body plan differentiates them from all living species. Here, we used computational fluid dynam...
Article
A new decapod crustacean assemblage associated with late Eocene coral reef deposits in northeast Spain (southern Pyrenees) is recorded; it includes Gemmellarocarcinus riglosensis sp. nov., Daira corallina sp. nov., Lobogalenopsis joei sp. nov., Liopsalis cf. anodon (Bittner, 1875) and Gale nopsis crassifrons A. Milne-Edwards, 1865. The genera Gemme...
Article
A new exceptionally-preserved Cambrian biota, with fossils belonging to multiple phyla, has recently been found from the middle part of the Longha Formation, near Fulu Village, southeastern Yunnan, South China, and is named the Fulu biota. Groups recovered so far include polymerid trilobites, agnostoids, large bivalved arthropods, bradoriid arthrop...
Article
Quantifying morphological evolution is key to determining the patterns and processes underlying the origin of phyla. We constructed a hierarchical morphological character matrix to characterize the radiation and establishment of echinoderm body plans during the early Paleozoic. This showed that subphylum-level clades diverged gradually through the...
Article
Full-text available
A new exceptionally-preserved Cambrian biota, with fossils belonging to multiple phyla, has recently been found from the middle part of the Longha Formation, near Fulu Village, southeastern Yunnan, South China, and is named the Fulu biota. Groups recovered so far include polymerid trilobites, agnostoids, large bivalved arthropods, bradoriid arthrop...
Article
Full-text available
Featherstars, comatulid crinoids that shed their stalk during their ontogeny, are the most species‐rich lineage of modern crinoids and the only ones present in shallow water today. Although they are of considerable palaeontological interest as a ‘success story’ of the Mesozoic Marine Revolution, their fossil record is relatively species‐poor and fr...
Article
Rhenopyrgids are rare, turreted edrioasterid edrioasteroids from the lower Paleozoic with a distinctive and apparently conservative morphology. However, new, well-preserved rhenopyrgid edrioasteroid material from Canada, along with a review of described taxa, has revealed broader structural diversity in the oral surface and enabled a re-evaluation...
Chapter
The Anti-Atlas contains a thick, volcanic-free Ordovician succession that originally deposited in a passive-margin basin. Three main sedimentary packages are bounded by major unconformities: (i) the Tremadocian–Floian Lower Fezouata and Upper Fezouata formations, which unconformably overlie a palaeorelief of Cambrian rifting volcanosedimentary comp...
Chapter
Full-text available
Progressive opening of the Rheic Ocean led to the drifting away of one or several ribbon terranes, generally ascribed to Avalonia, and inaugurated a passive margin stage on the newly formed margin of NW Gondwana. In Iberia, which remained on the Gondwanan side of the ocean, the rift to drift transition is recorded in the Ossa Morena Zone in latest...
Article
A new Lower Devonian fauna from the Iberian Chains (NE Spain) is described. Specimens have been collected from the shaley intervals of the Mariposas Formation dated as early Emsian. These include the camerates Acanthocrinus carsli n. sp., Platyhexacrinus santacruzensis n. sp., Culicocrinus breimeri n. sp., Camerata indeterminate, and an indetermina...
Article
A taxonomic study of the trilobites found in the Marianian parastratotype of the Iberian Chains, north‐east Spain, results in the recognition of the new redlichiid Redlichia isuelaensis sp. nov., the ellipsocephalid Lusatiops ribotanus, and the saukiandids Onaraspis garciae sp. nov., Paulaspis tiergaensis gen. et sp. nov. and Luciaspis matiasi gen....
Article
Full-text available
We describe Aragolambrus collinsin. gen., n. sp. from the Eocene (Ypresian) of Huesca, Aragón, northern Spain, which represents the second report of a fossil Parthenopidae from Spain and the first from the Eocene. Aragolambrus collinsin. gen. n. sp. is considered one of the oldest parthenopid crabs, and important morphological features include the...
Article
Hexedriocystis is a very unusual echinoderm with a puzzling combination of morphological features. The two described species H. inexpectatus and H. mimus occur in Upper Ordovician siliciclastic rocks from Morocco, North Africa. They show a striking combination of isorophid edrioasteroid, paedomorphic crinoid and blastozoan characteristics including...
Article
New Upper Ordovician Echinoderm–Lagerstätten from the Anti-Atlas of Morocco have provided large collections of blastozoan echinoderms. The echinosphaeritid Echinosphaerites dianae n. sp. preserves the oral region, stem and brachioles, characteristics absent in many species of the same genus. Homocystites adidiensis n. sp. is represented by almost 5...
Preprint
Repaired fossil skeletons provide the opportunity to study predation rates, repair mechanisms, and ecological interactions in deep time. Trilobites allow the study of repaired damage over long time periods and large geographic areas due to their longevity as a group, global distribution, and well-preserved mineralized exoskeletons. Repair frequenci...
Poster
Full-text available
Latest findings on our study about the evolution of pentaradiality in echinoderms, presented in a poster here.
Article
Full-text available
Los paleontólogos investigadores han utilizado el epíteto latinizado aragonensis o aragonense para nombrar a cuarenta (40) de las mil diez (1010) especies de fósiles que han descrito por primera vez en territorio aragonés. De forma similar, han utilizado en doce (12) ocasiones un lexema que se refiere a Aragón o a alguna cuestión relacionada con Ar...
Article
Full-text available
Post-Palaeozoic crinoids from northeast Spain ranging from the Ladinian (Middle Triassic) to the Ilerdian (lower Ypresian, early Eocene) are documented. Here we provide the first attempt to reconstruct the environmental distribution of these crinoids based on relatively complete material (mostly cups). Triassic forms are dominated by encrinids from...
Article
Full-text available
Coronates are widely distributed in Ordovician and Silurian rocks from Gondwana but have never been reported from Africa. A single theca of Mespilocystites sp. is the first representative of this blastozoan clade from the African continent, extending its geographical range. Previous reports of Mespilocystites include the Czech Republic, France, Ita...
Article
Edrioasteroids from the Katian, Lower Ktaoua and Upper Tiouririne formations provide new data on several previously described species and two new species of edrioasteroids. Ontogenetic and palaeoecological data are provided for Moroccopyrgus matacarros . Well-preserved oral areas and internal views of the oral frame show Euryeschatia reboulorum to...
Article
Full-text available
A new brachiopod species, Westonia mardini, from the Furongian of Turkey and a new occurrence of Westonia urbiona from the Cambrian Series 3 of Iberian Peninsula are reported. These new finds of ‘westoniids’ collected in Gondwana allow the discussion of the functional morphology of their terrace ridges. This structure enabled an effective and rapid...
Article
Full-text available
Clades that represent a new 'Bauplan' have been hypothesised to exhibit more variability than more derived clades. Accordingly, there is an expectation of greater variation around the time of the origin of a clade than later in its evolutionary history. This 'canalisation' has been tested in terms of morphological disparity (interspecific variation...
Data
All data collected and used in analysis using the R script included
Data
Summary of all species sampled, their age, taxonomic affiliations and the proxies of variation Grey font denote species with less than 5 individuals, which were excluded from the intraspecific variation analyses.
Data
R code R code—intraspecific variation of primibrachials in the Phanerozoic.
Conference Paper
Fossil decapods from the Forcall Formation (Lower Cretaceous, Aptian) of the Oliete Sub-basin (Maestrazgo Basin) are reported. Specimens have been collected from the localities of Josa and Obón (Teruel) in the Aragonese branch of the Iberian Cordillera, NE Spain. The taxon Atherfieldastacus magnus (M'Coy, 1849) is by far the most common species in...
Article
Full-text available
Repaired fossil skeletons provide the opportunity to study predation rates, repair mechanisms, and ecological interactions in deep time. Trilobites allow the study of repaired damage over long time periods and large geographic areas due to their longevity as a group, global distribution, and well-preserved mineralized exoskeletons. Repair frequenci...
Book
Full-text available
The book “Paleotipos aragoneses” consists of an extensive list containing all fossil taxa, of a genus or lower taxonomic level, whose type localities lie in the territory of Aragon. The development of such research included a meticulous search of literature. It was possible to review four hundred original texts in which the new taxa were establishe...
Article
Full-text available
Echinoderms are characterized by a calcite endoskeleton with a unique microstructure, which is optimized for multiple functions. For instance, some light-sensitive ophiuroids (Ophiuroidea) and asteroids (Asteroidea) possess skeletal plates with multi-lens arrays that are thought to act as photosensory organs. The origins of these lens-like microstr...
Article
The Cambrian, characterized by the global appearance of diverse biomineralized metazoans in the fossil record for the first time, represents a pivotal point in the history of life. This period also documents a major change in the nature of the sea floor: Neoproterozoic-type substrates stabilized by microbial mats were replaced by unconsolidated sof...
Article
Moroccodiscus smithi represents a new cyclocystoid genus and species based on moldic specimens from the Middle Ordovician Taddrist Formation (Darriwilian) of SE Morocco. This represents the earliest articulated member of the Cyclocystoidea and is the first complete cyclocystoid described from the Ordovician of Gondwana, as well as the first cyclocy...
Article
A Cambrian immigration event of paradoxidid trilobites has traditionally marked some regional lower–middle Cambrian boundaries in the Acado-Baltic subprovince (including Baltica and the Mediterranean and Avalonian margins of West Gondwana). The oldest paradoxidine species in Morocco and the Iberian Peninsula have been used as a chronostratigraphic...
Article
Progress in echinoderm paleobiology - Volume 91 Issue 4 - Samuel Zamora, Imran A. Rahman
Article
Trilobites were capable of enroling in different ways based on the flexible articulation of thoracic segments and associated interlocking devices; the type of enrolment (spiral or sphaeroidal) is thought to have largely depended on the coaptative devices that each trilobite used to enclose the body. Based on X-ray microtomography scans of complete...